treatments-xml/data/FD/3C/10/FD3C1063FFA03A3CC0C43BEDFEA3FE6A.xml
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<document id="95B0241D8C24D52DEA4D24C2E8A9FCE3" ID-CLB-Dataset="6002" ID-DOI="10.11646/zootaxa.4980.2.7" ID-GBIF-Dataset="3de41046-fff8-458c-8b77-9b0b6dd51632" ID-ISSN="1175-5326" ID-Zenodo-Dep="4889226" ID-ZooBank="CFB3B4E3-83BF-484E-B5BE-68FE1BD215D1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622537339575" checkinUser="plazi" docAuthor="Muthukatturaja, Marimuthu &amp; Balasubramanian, Chellaiah" docDate="2021" docId="FD3C1063FFA03A3CC0C43BEDFEA3FE6A" docLanguage="en" docName="zootaxa.4980.2.7.pdf" docOrigin="Zootaxa 4980 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Caenis maduraiensis Balasubramanian and Muthukatturaja 2021, new species" docType="treatment" docVersion="8" lastPageNumber="371" masterDocId="0105681BFFA13A39C0533B32FF94FFE3" masterDocTitle="A new species of Caenis Stephens, 1835 (Ephemeroptera: Caenidae) from Tamil Nadu, Southern India" masterLastPageNumber="372" masterPageNumber="366" pageNumber="367" updateTime="1698995776350" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="52B43E035D82ACE0C09EDC68764209C1">A new species of Caenis Stephens, 1835 (Ephemeroptera: Caenidae) from Tamil Nadu, Southern India</mods:title>
</mods:titleInfo>
<mods:name id="64EAB0CCEBFC8039D441A46F29A95678" type="personal">
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<mods:namePart id="80DF5AAB19ABE69E5E139D65CF46BE39">Muthukatturaja, Marimuthu</mods:namePart>
<mods:nameIdentifier id="E5021E85602E7DAFEE81CDB0530F7F83" type="ORCID">0000-0001-5796-0211</mods:nameIdentifier>
<mods:affiliation id="BFE32C0E8033A1035DBAF09E4B5E3DA4">Department of Zoology, Thiagarajar College, Madurai- 625009, Tamil Nadu, India &amp; drmkraja 83 @ gmail. com; https: // orcid. org / 0000 - 0001 - 5796 - 0211</mods:affiliation>
<mods:nameIdentifier id="774128855489335DBF83FD8BC69EED38" type="email">drmkraja83@gmail.com</mods:nameIdentifier>
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<mods:namePart id="AC4B0AD2CF1A70BCB132FB0B8A88979F">Balasubramanian, Chellaiah</mods:namePart>
<mods:affiliation id="37FA58DE14A7DD6A8DC4FDF0ED1CD409">Department of Zoology, Thiagarajar College, Madurai- 625009, Tamil Nadu, India</mods:affiliation>
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<mods:date id="E26FA267DD1EB41332FB7B857951BB89">2021</mods:date>
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<mods:number id="1FDEC4EE74F9B3E9AF14C436E3123FCF">2021-06-01</mods:number>
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<treatment id="FD3C1063FFA03A3CC0C43BEDFEA3FE6A" ID-DOI="http://doi.org/10.5281/zenodo.4889680" ID-GBIF-Taxon="182958725" ID-Zenodo-Dep="4889680" LSID="urn:lsid:plazi:treatment:FD3C1063FFA03A3CC0C43BEDFEA3FE6A" httpUri="http://treatment.plazi.org/id/FD3C1063FFA03A3CC0C43BEDFEA3FE6A" lastPageId="5" lastPageNumber="371" pageId="1" pageNumber="367">
<subSubSection id="3D8FF2FEFFA03A38C0C43BEDFBB1FF1A" box="[151,1061,223,250]" pageId="1" pageNumber="367" type="nomenclature">
<paragraph id="752AA175FFA03A38C0C43BEDFBB1FF1A" blockId="1.[151,1061,223,286]" box="[151,1061,223,250]" pageId="1" pageNumber="367">
<heading id="2E621619FFA03A38C0C43BEDFBB1FF1A" bold="true" box="[151,1061,223,250]" fontSize="11" level="1" pageId="1" pageNumber="367" reason="1">
<emphasis id="47E17D67FFA03A38C0C43BEDFBB1FF1A" bold="true" box="[151,1061,223,250]" pageId="1" pageNumber="367">
<taxonomicName id="B295DAF6FFA03A38C0C43BEDFC1CFF1A" authority="Balasubramanian and Muthukatturaja" authorityName="Balasubramanian and Muthukatturaja" authorityYear="2021" box="[151,904,223,250]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="maduraiensis" status="sp. nov.">
<emphasis id="47E17D67FFA03A38C0C43BEDFE01FF19" bold="true" box="[151,405,223,250]" italics="true" pageId="1" pageNumber="367">Caenis maduraiensis</emphasis>
Balasubramanian and Muthukatturaja
</taxonomicName>
,
<taxonomicNameLabel id="5CD2C01CFFA03A38C3C63BEDFBB1FF1A" box="[917,1061,223,249]" pageId="1" pageNumber="367" rank="species">new species</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="3D8FF2FEFFA03A38C0C43A31FECCFEFD" box="[151,344,259,286]" pageId="1" pageNumber="367" type="description">
<paragraph id="752AA175FFA03A38C0C43A31FECCFEFD" blockId="1.[151,1061,223,286]" box="[151,344,259,286]" pageId="1" pageNumber="367">
(
<figureCitation id="EDAEBDF0FFA03A38C0CC3A31FE9FFEFE" box="[159,267,259,285]" captionStart="FIGURES 14" captionStartId="2.[151,264,1663,1687]" captionTargetBox="[151,1436,381,1640]" captionTargetId="figure-120@2.[151,1436,381,1640]" captionTargetPageId="2" captionText="FIGURES 14. Caenis maduraiensis sp. nov. dorsal habitus: 1, Mature nymph; 2, Male imago; 3, Female imago; 4, Collection site." figureDoi="http://doi.org/10.5281/zenodo.4889228" httpUri="https://zenodo.org/record/4889228/files/figure.png" pageId="1" pageNumber="367">Figs. 13</figureCitation>
,
<figureCitation id="EDAEBDF0FFA03A38C1443A31FEC4FEFD" box="[279,336,259,286]" captionStart-0="FIGURES 511" captionStart-1="FIGURES 1218" captionStart-2="FIGURES 1930" captionStartId-0="3.[151,264,1762,1786]" captionStartId-1="4.[151,264,1962,1986]" captionStartId-2="5.[151,264,1589,1613]" captionTargetBox-0="[153,1434,500,1727]" captionTargetBox-1="[181,1405,195,1927]" captionTargetBox-2="[161,1427,423,1558]" captionTargetId-0="figure-184@3.[153,1434,489,1739]" captionTargetId-1="figure-17@4.[171,1416,181,1939]" captionTargetId-2="figure-122@5.[151,1436,417,1566]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." captionText-1="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." captionText-2="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi-0="http://doi.org/10.5281/zenodo.4889230" figureDoi-1="http://doi.org/10.5281/zenodo.4889234" figureDoi-2="http://doi.org/10.5281/zenodo.4889236" httpUri-0="https://zenodo.org/record/4889230/files/figure.png" httpUri-1="https://zenodo.org/record/4889234/files/figure.png" httpUri-2="https://zenodo.org/record/4889236/files/figure.png" pageId="1" pageNumber="367">530</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="3D8FF2FEFFA03A38C0C43A79FCBDFD82" pageId="1" pageNumber="367" type="materials_examined">
<paragraph id="752AA175FFA03A38C0C43A79FF46FE16" blockId="1.[151,1437,331,2014]" pageId="1" pageNumber="367">
<emphasis id="47E17D67FFA03A38C0C43A79FED2FE86" bold="true" box="[151,326,331,357]" pageId="1" pageNumber="367">Type material.</emphasis>
<materialsCitation id="C5FDAB28FFA03A38C11D3A79FE46FE4E" ID-GBIF-Occurrence="3305364302" collectingDate="2019-08-09" collectorName="Balasubramanian &amp; Muthukatturaja " country="India" latitude="9.914166" location="Madurai district" longLatPrecision="21" longitude="78.18111" municipality="Vandiyur" pageId="1" pageNumber="367" specimenCount="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus id="AA2E1FD7FFA03A38C11D3A79FE22FE86" box="[334,438,331,357]" pageId="1" pageNumber="367" type="holotype">Holotype</typeStatus>
(in ethanol): male imago (reared from nymph) (
<collectorName id="D860C4A3FFA03A38C3893A79FBADFE86" box="[986,1081,331,357]" pageId="1" pageNumber="367">Reg.No.</collectorName>
SRC/ ZSI-I/E 469) from
<collectingCountry id="0D82E1E5FFA03A38C50F3A79FA0CFE86" box="[1372,1432,331,357]" name="India" pageId="1" pageNumber="367">India</collectingCountry>
,
<collectingMunicipality id="954E3B0FFFA03A38C0C43A5DFF6AFE6A" box="[151,254,367,393]" pageId="1" pageNumber="367">Vandiyur</collectingMunicipality>
pond,
<location id="704AF7AEFFA03A38C11E3A5DFD92FE6A" LSID="urn:lsid:plazi:treatment:FD3C1063FFA03A3CC0C43BEDFEA3FE6A:704AF7AEFFA03A38C11E3A5DFD92FE6A" box="[333,518,367,393]" country="India" latitude="9.914166" longLatPrecision="21" longitude="78.18111" municipality="Vandiyur" name="Madurai district" pageId="1" pageNumber="367" stateProvince="Tamil Nadu">Madurai district</location>
,
<collectingRegion id="B7516F97FFA03A38C2423A5DFD41FE69" box="[529,725,367,394]" country="India" name="Tamil Nadu" pageId="1" pageNumber="367">Tamil Nadu state</collectingRegion>
,
<date id="012B87B5FFA03A38C2B13A5DFCCAFE6A" box="[738,862,367,393]" pageId="1" pageNumber="367" value="2019-08-09">
<collectingDate id="116F7E5DFFA03A38C2B13A5DFCCAFE6A" box="[738,862,367,393]" pageId="1" pageNumber="367" value="2019-08-09">
9.viii.
<quantity id="B26D0C90FFA03A38C3733A5DFCCAFE6A" box="[800,862,367,393]" metricMagnitude="6" metricUnit="m" metricValue="2.019124" pageId="1" pageNumber="367" unit="m" value="2019124.0">2019</quantity>
</collectingDate>
</date>
, 124 m,
<geoCoordinate id="10A1C7B2FFA03A38C3913A5DFBC2FE69" box="[962,1110,367,394]" degrees="09" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="367" precision="15" seconds="51" value="9.914166">09°5451” N</geoCoordinate>
,
<geoCoordinate id="10A1C7B2FFA03A38C4303A5DFB67FE69" box="[1123,1267,367,394]" degrees="78" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="367" precision="15" seconds="52" value="78.18111">78°1052” E</geoCoordinate>
,
<collectorName id="D860C4A3FFA03A38C5523A5DFF53FE4E" pageId="1" pageNumber="367">Balasubramanian</collectorName>
&amp;
<collectorName id="D860C4A3FFA03A38C0BA3AA1FE46FE4E" box="[233,466,403,429]" pageId="1" pageNumber="367">Muthukatturaja Cols</collectorName>
</materialsCitation>
.
<materialsCitation id="C5FDAB28FFA03A38C1883AA6FF5AFE16" ID-GBIF-Occurrence="3305364303" collectingDate="2019-08-09" collectorName="Balasubramanian &amp; Muthukatturaja" country="India" latitude="9.914166" location="Madurai district" longLatPrecision="21" longitude="78.18111" municipality="Vandiyur" pageId="1" pageNumber="367" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu" typeStatus="paratype">
<typeStatus id="AA2E1FD7FFA03A38C1883AA6FDDCFE4D" box="[475,584,404,430]" pageId="1" pageNumber="367" type="paratype">Paratypes</typeStatus>
(in ethanol):
<specimenCount id="63936AFCFFA03A38C28D3AA6FCCCFE4E" box="[734,856,403,430]" pageId="1" pageNumber="367" type="female">one female</specimenCount>
imago (reared from nymph) (
<collectorName id="D860C4A3FFA03A38C4F33AA1FB6BFE4E" box="[1184,1279,403,429]" pageId="1" pageNumber="367">Reg.No.</collectorName>
SRC/ ZSI-I/E 470) and 5 mature nymphs (
<collectorName id="D860C4A3FFA03A38C19F3A85FDBFFE32" box="[460,555,439,465]" pageId="1" pageNumber="367">Reg.No.</collectorName>
SRC/ ZSI-I/E 471), same data as holotype.
<collectorName id="D860C4A3FFA03A38C45F3A85FB5AFE32" box="[1036,1230,439,465]" pageId="1" pageNumber="367">Balasubramanian</collectorName>
&amp;
<collectorName id="D860C4A3FFA03A38C4A33A85FF5AFE16" pageId="1" pageNumber="367">Muthukatturaja Cols</collectorName>
</materialsCitation>
.
</paragraph>
<paragraph id="752AA175FFA03A38C0943ACDFCBDFD82" blockId="1.[151,1437,331,2014]" pageId="1" pageNumber="367">
<emphasis id="47E17D67FFA03A38C0943ACDFDB3FDFA" bold="true" box="[199,551,511,537]" pageId="1" pageNumber="367">
Additional material (
<collectionCode id="138439B0FFA03A38C1923ACDFD8EFDFA" box="[449,538,511,537]" pageId="1" pageNumber="367">ZMTC</collectionCode>
).
</emphasis>
<specimenCount id="63936AFCFFA03A38C27D3ACDFD01FDFA" box="[558,661,511,538]" pageId="1" pageNumber="367" type="male">156 male</specimenCount>
imago (reared from nymph),
<specimenCount id="63936AFCFFA03A38C3B23ACDFBCAFDFA" box="[993,1118,511,538]" pageId="1" pageNumber="367" type="female">254 female</specimenCount>
<materialsCitation id="C5FDAB28FFA03A38C4363ACDFCB1FD82" ID-GBIF-Occurrence="3305364301" collectingDate="2019-08-19" collectorName="Balasubramanian &amp; Muthukatturaja Cols" country="India" latitude="9.914166" location="Madurai district" longLatPrecision="21" longitude="78.18111" municipality="Vandiyur" pageId="1" pageNumber="367" specimenCount="1" stateProvince="Tamil Nadu">
imago (reared from nymph) and 245 mature nymphs,
<collectingCountry id="0D82E1E5FFA03A38C1E03911FE7BFDDE" box="[435,495,547,573]" name="India" pageId="1" pageNumber="367">India</collectingCountry>
,
<collectingMunicipality id="954E3B0FFFA03A38C1A83911FDF6FDDE" box="[507,610,547,573]" pageId="1" pageNumber="367">Vandiyur</collectingMunicipality>
pond,
<location id="704AF7AEFFA03A38C2FD3911FCF1FDDE" LSID="urn:lsid:plazi:treatment:FD3C1063FFA03A3CC0C43BEDFEA3FE6A:704AF7AEFFA03A38C2FD3911FCF1FDDE" box="[686,869,547,573]" country="India" latitude="9.914166" longLatPrecision="21" longitude="78.18111" municipality="Vandiyur" name="Madurai district" pageId="1" pageNumber="367" stateProvince="Tamil Nadu">Madurai district</location>
,
<collectingRegion id="B7516F97FFA03A38C3233911FBA5FDDD" box="[880,1073,547,574]" country="India" name="Tamil Nadu" pageId="1" pageNumber="367">Tamil Nadu state</collectingRegion>
,
<date id="012B87B5FFA03A38C46E3911FB52FDDE" box="[1085,1222,547,573]" pageId="1" pageNumber="367" value="2019-08-19">
<collectingDate id="116F7E5DFFA03A38C46E3911FB52FDDE" box="[1085,1222,547,573]" pageId="1" pageNumber="367" value="2019-08-19">
19.viii.
<quantity id="B26D0C90FFA03A38C4DB3911FB52FDDE" box="[1160,1222,547,573]" metricMagnitude="6" metricUnit="m" metricValue="2.019124" pageId="1" pageNumber="367" unit="m" value="2019124.0">2019</quantity>
</collectingDate>
</date>
, 124 m,
<geoCoordinate id="10A1C7B2FFA03A38C5743911FF3AFD81" degrees="09" direction="north" minutes="54" orientation="latitude" pageId="1" pageNumber="367" precision="15" seconds="51" value="9.914166">09°5451” N</geoCoordinate>
,
<geoCoordinate id="10A1C7B2FFA03A38C0EA3975FEDCFD81" box="[185,328,583,610]" degrees="78" direction="east" minutes="10" orientation="longitude" pageId="1" pageNumber="367" precision="15" seconds="52" value="78.18111">78°1052” E</geoCoordinate>
,
<collectorName id="D860C4A3FFA03A38C1073975FD82FD82" box="[340,534,583,609]" pageId="1" pageNumber="367">Balasubramanian</collectorName>
&amp;
<collectorName id="D860C4A3FFA03A38C2683975FCB1FD82" box="[571,805,583,609]" pageId="1" pageNumber="367">Muthukatturaja Cols</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3D8FF2FEFFA03A3CC0943959FCCAFEA2" lastPageId="5" lastPageNumber="371" pageId="1" pageNumber="367" type="diagnosis">
<paragraph id="752AA175FFA03A38C0943959FD95FC46" blockId="1.[151,1437,331,2014]" pageId="1" pageNumber="367">
<emphasis id="47E17D67FFA03A38C0943959FED6FD66" bold="true" box="[199,322,619,645]" pageId="1" pageNumber="367">Diagnosis.</emphasis>
<taxonomicName id="B295DAF6FFA03A38C11A395EFDA2FD66" authority="Muthukatturaja &amp; Balasubramanian, 2021" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2021" box="[329,566,619,646]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="367" phylum="Arthropoda" rank="species" species="maduraiensis" status="sp. nov.">
<emphasis id="47E17D67FFA03A38C11A395EFDA2FD66" box="[329,566,619,646]" italics="true" pageId="1" pageNumber="367">Caenis maduraiensis</emphasis>
</taxonomicName>
<emphasis id="47E17D67FFA03A38C26D3959FD0DFD66" bold="true" box="[574,665,619,645]" pageId="1" pageNumber="367">
<taxonomicNameLabel id="5CD2C01CFFA03A38C26D3959FD0DFD66" box="[574,665,619,645]" pageId="1" pageNumber="367" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: Nymph: (i) anterior margin of labrum with row of long bipectinate setae laterally and a group of highly fringed setae behind emargination; (ii) fore femora with transverse row of bifid spines; middle and hind femora with sparse bifid spines; (iii) hind claw long, slender and hooked with row of denticles; (iv) abdominal tergum II with pointed medial projection and pointed teeth on the posterior margin. Imago: (i) prosternal triangle with slightly concave sides, tip acutely pointed; (ii) tarsomere I extremely short; (iii) tarsomere II apically with tongue-shaped projections on single side; (iv) apical part of forceps narrowed to the tip, with a long apical spine, and few trichomes on surface; (v) cross veins of forewing arranged singly except in R
<subScript id="E911A330FFA03A38C5553864FA9AFC87" attach="left" box="[1286,1294,854,868]" fontSize="6" pageId="1" pageNumber="367">1</subScript>
and R
<subScript id="E911A330FFA03A38C5083864FAF7FC87" attach="left" box="[1371,1379,854,868]" fontSize="6" pageId="1" pageNumber="367">4</subScript>
; (vi) MP forked at near 1/3 of distance from base to margin; (vii) fore tibia two times longer than femur and (viii) female subanal plate apically truncated.
</paragraph>
<paragraph id="752AA175FFA03A3BC094389DFC67FE86" blockId="1.[151,1437,331,2014]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="368" pageId="1" pageNumber="367">
<emphasis id="47E17D67FFA03A38C094389DFEEBFC2A" bold="true" box="[199,383,943,969]" pageId="1" pageNumber="367">Mature nymph</emphasis>
(in 95% ethanol) (
<figureCitation id="EDAEBDF0FFA03A38C233389DFD24FC2A" box="[608,688,943,969]" captionStart="FIGURES 14" captionStartId="2.[151,264,1663,1687]" captionTargetBox="[151,1436,381,1640]" captionTargetId="figure-120@2.[151,1436,381,1640]" captionTargetPageId="2" captionText="FIGURES 14. Caenis maduraiensis sp. nov. dorsal habitus: 1, Mature nymph; 2, Male imago; 3, Female imago; 4, Collection site." figureDoi="http://doi.org/10.5281/zenodo.4889228" httpUri="https://zenodo.org/record/4889228/files/figure.png" pageId="1" pageNumber="367">Figs. 1</figureCitation>
,
<figureCitation id="EDAEBDF0FFA03A38C291389DFD6AFC29" box="[706,766,943,970]" captionStart-0="FIGURES 511" captionStart-1="FIGURES 1218" captionStart-2="FIGURES 1930" captionStartId-0="3.[151,264,1762,1786]" captionStartId-1="4.[151,264,1962,1986]" captionStartId-2="5.[151,264,1589,1613]" captionTargetBox-0="[153,1434,500,1727]" captionTargetBox-1="[181,1405,195,1927]" captionTargetBox-2="[161,1427,423,1558]" captionTargetId-0="figure-184@3.[153,1434,489,1739]" captionTargetId-1="figure-17@4.[171,1416,181,1939]" captionTargetId-2="figure-122@5.[151,1436,417,1566]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." captionText-1="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." captionText-2="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi-0="http://doi.org/10.5281/zenodo.4889230" figureDoi-1="http://doi.org/10.5281/zenodo.4889234" figureDoi-2="http://doi.org/10.5281/zenodo.4889236" httpUri-0="https://zenodo.org/record/4889230/files/figure.png" httpUri-1="https://zenodo.org/record/4889234/files/figure.png" httpUri-2="https://zenodo.org/record/4889236/files/figure.png" pageId="1" pageNumber="367">519</figureCitation>
): Body length (excluding cerci)
<quantity id="B26D0C90FFA03A38C42A389DFA93FC29" box="[1145,1287,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.4" metricValueMin="3.1" pageId="1" pageNumber="367" unit="mm" value="3.25" valueMax="3.4" valueMin="3.1">3.13.4 mm</quantity>
; cerci length 3.54.0 mm; median caudal filament
<quantity id="B26D0C90FFA03A38C26E38E1FD5CFC0D" box="[573,712,979,1006]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.5" metricValueMin="4.2" pageId="1" pageNumber="367" unit="mm" value="4.35" valueMax="4.5" valueMin="4.2">4.24.5 mm</quantity>
; length of head
<quantity id="B26D0C90FFA03A38C32E38E1FC43FC0D" box="[893,983,979,1006]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="367" unit="mm" value="0.4">0.4 mm</quantity>
; width of head
<quantity id="B26D0C90FFA03A38C4D638E1FB4BFC0D" box="[1157,1247,979,1006]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="367" unit="mm" value="0.7">0.7 mm</quantity>
; antennae length
<quantity id="B26D0C90FFA03A38C0C438C5FF66FBF1" box="[151,242,1015,1042]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="367" unit="mm" value="1.3">1.3 mm</quantity>
; length of fore femora/ tibia/ tarsi/ claw (mm) 0.7/0.5/0.3/0.2; length of middle femora/ tibia/ tarsi/ claw (mm) 0.6/0.37/0.25/0.18 and length of femora/ tibia/ tarsi/ claw (mm) viz., 0.65/0.6/0.4/0.15. General body coloration yellowish brown dorsally and pale ventrally, mature specimens generally paler; legs pale brown. Head brownish yellow; scape and pedicel of antennae pale brown without hairs, flagellum translucent; labrum (
<figureCitation id="EDAEBDF0FFA03A38C4A03F51FAA2FB9D" box="[1267,1334,1123,1150]" captionStart="FIGURES 511" captionStartId="3.[151,264,1762,1786]" captionTargetBox="[153,1434,500,1727]" captionTargetId="figure-184@3.[153,1434,489,1739]" captionTargetPageId="3" captionText="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." figureDoi="http://doi.org/10.5281/zenodo.4889230" httpUri="https://zenodo.org/record/4889230/files/figure.png" pageId="1" pageNumber="367">Fig. 5</figureCitation>
) broadly emarginated, dorsum with long setae, anterior margin with row of bipectinate setae longer laterally and a group of highly fringed setae behind emargination; mandibles (
<figureCitation id="EDAEBDF0FFA03A38C2BF3F99FCC7FB25" box="[748,851,1195,1222]" captionStart="FIGURES 511" captionStartId="3.[151,264,1762,1786]" captionTargetBox="[153,1434,500,1727]" captionTargetId="figure-184@3.[153,1434,489,1739]" captionTargetPageId="3" captionText="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." figureDoi="http://doi.org/10.5281/zenodo.4889230" httpUri="https://zenodo.org/record/4889230/files/figure.png" pageId="1" pageNumber="367">Figs. 6, 7</figureCitation>
) with broad incisors, outer margin of mandibles with long thin setae, left mandible with 7 thin setae below the molar, with elevated hump between prostheca and molar; maxillae (
<figureCitation id="EDAEBDF0FFA03A38C1543FC1FEE5FAEE" box="[263,369,1267,1293]" captionStart="FIGURES 511" captionStartId="3.[151,264,1762,1786]" captionTargetBox="[153,1434,500,1727]" captionTargetId="figure-184@3.[153,1434,489,1739]" captionTargetPageId="3" captionText="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." figureDoi="http://doi.org/10.5281/zenodo.4889230" httpUri="https://zenodo.org/record/4889230/files/figure.png" pageId="1" pageNumber="367">Figs. 8, 9</figureCitation>
) with 3 prominent teeth on galealacinia, ventral base of inner margin of galea with tuft of thin setae, distal portion of cardo with cluster of bipectinate stout setae, maxillary palp 3 segmented, segment I broad, segment I and II subequal in length, segment III longer, inner margin of segment III with row of 13 bipectinate setae; hypopharynx (
<figureCitation id="EDAEBDF0FFA03A38C1683E6DFE1BFA9A" box="[315,399,1375,1401]" captionStart="FIGURES 511" captionStartId="3.[151,264,1762,1786]" captionTargetBox="[153,1434,500,1727]" captionTargetId="figure-184@3.[153,1434,489,1739]" captionTargetPageId="3" captionText="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." figureDoi="http://doi.org/10.5281/zenodo.4889230" httpUri="https://zenodo.org/record/4889230/files/figure.png" pageId="1" pageNumber="367">Fig. 10</figureCitation>
): anterior margin of lingua with long setae, superlingua with small hair-like setae at anterior margin; labium with glossae (
<figureCitation id="EDAEBDF0FFA03A38C1B83EB1FDA8FA7E" box="[491,572,1411,1437]" captionStart="FIGURES 511" captionStartId="3.[151,264,1762,1786]" captionTargetBox="[153,1434,500,1727]" captionTargetId="figure-184@3.[153,1434,489,1739]" captionTargetPageId="3" captionText="FIGURES 511. Caenis maduraiensis sp. nov. mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium." figureDoi="http://doi.org/10.5281/zenodo.4889230" httpUri="https://zenodo.org/record/4889230/files/figure.png" pageId="1" pageNumber="367">Fig. 11</figureCitation>
) ovate with stout spines at outer margin, mesal margin of glossae with thin setae, inner and outer margin of paraglossae with thin spines directed anteriorly, labial palp 3 segmented, segment II slightly longer than segment I, segment III short; outer margin of segment II with stout spines, outer margin of segment I with row of bipectinate setae, segment III triangular with 2 rows of stout setae ventrally, inner margin of segment III with row of stout spines. Thorax brownish yellow; lateral margins of pronotum convex, anterolateral margins with pair of spine-like setae and wider than posterolateral margins.
<emphasis id="47E17D67FFA03A38C3A63D0AFBBEF9B2" box="[1013,1066,1592,1617]" italics="true" pageId="1" pageNumber="367">Legs</emphasis>
pale brown covered with scales, tibiae and tarsi light pale. Foreleg (
<figureCitation id="EDAEBDF0FFA03A38C2713D69FDE3F996" box="[546,631,1627,1653]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="1" pageNumber="367">Fig. 12</figureCitation>
): trochanter with transversal row of stout spines; outer margin of femur with few setae and spines, near ½ of dorsal surface with transversal row of bifid spines, inner margin with few stout spines, ¼ of dorsal surface with few long hairs; tibia with 3 bifid spines at mesal margin, inner distal margin with long bipectinate setae; few long hairs on tibia; tarsi with row of stout spines on inner margin, mesal margin with bipectinate spine; few long hairs on tarsi; claw long, slender and slightly hooked. Middle leg (
<figureCitation id="EDAEBDF0FFA03A38C4923DD9FA81F8E6" box="[1217,1301,1771,1797]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="1" pageNumber="367">Fig. 13</figureCitation>
): trochanter with two bifid spines; outer margin of femora with long spines, ½ of inner margin with stout spines and remainder with bifid spines, dorsal surface with sparse bifid spines; inner margin of tibia with row of stout spines; ½ of tarsi with short bipectinate setae at mesal margin, claw long, slender and slightly hooked. Hindleg (
<figureCitation id="EDAEBDF0FFA03A38C4923C65FA82F892" box="[1217,1302,1879,1905]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="1" pageNumber="367">Fig. 14</figureCitation>
): trochanter with bifid spine; inner and outer margin of femur with long spines, ½ of dorsal surface with few bifid spines; inner margin of tibia with row of stout spines, outer margin with thin setae; tarsi similar to midleg; claw (
<figureCitation id="EDAEBDF0FFA03A38C5523CADFAC7F859" box="[1281,1363,1951,1978]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="1" pageNumber="367">Fig. 15</figureCitation>
) long, slender and hooked with row of denticles. Abdomen. Terga VIIX yellowish brown; tergum II with pointed medial projection and pointed teeth on the posterior margin; posterior margin of terga VIIIX with row of long setae; abdominal sterna IIIIX pale brown. Gills (
<figureCitation id="EDAEBDF0FFA33A3BC2D73B89FC84FF35" box="[644,784,187,214]" captionStart-0="FIGURES 1218" captionStart-1="FIGURES 1930" captionStartId-0="4.[151,264,1962,1986]" captionStartId-1="5.[151,264,1589,1613]" captionTargetBox-0="[181,1405,195,1927]" captionTargetBox-1="[161,1427,423,1558]" captionTargetId-0="figure-17@4.[171,1416,181,1939]" captionTargetId-1="figure-122@5.[151,1436,417,1566]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." captionText-1="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi-0="http://doi.org/10.5281/zenodo.4889234" figureDoi-1="http://doi.org/10.5281/zenodo.4889236" httpUri-0="https://zenodo.org/record/4889234/files/figure.png" httpUri-1="https://zenodo.org/record/4889236/files/figure.png" pageId="2" pageNumber="368">Figs. 1619</figureCitation>
): pale brown; gill I (
<figureCitation id="EDAEBDF0FFA33A3BC3AA3B89FBD8FF35" box="[1017,1100,187,214]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="2" pageNumber="368">Fig. 16</figureCitation>
) 2-segmented, distal segment 4 times longer than proximal segment; gill II (
<figureCitation id="EDAEBDF0FFA33A3BC2FA3BEDFD68FF1A" box="[681,764,223,249]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="2" pageNumber="368">Fig. 17</figureCitation>
) brown, covered with scales; margins evenly covered with long setae, inner margin with row of fringed setae; gills 36 (
<figureCitation id="EDAEBDF0FFA33A3BC3013A31FC25FEFE" box="[850,945,259,285]" captionStart="FIGURES 1218" captionStartId="4.[151,264,1962,1986]" captionTargetBox="[181,1405,195,1927]" captionTargetId="figure-17@4.[171,1416,181,1939]" captionTargetPageId="4" captionText="FIGURES 1218. Caenis maduraiensis sp. nov. mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III." figureDoi="http://doi.org/10.5281/zenodo.4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="2" pageNumber="368">Figs. 18</figureCitation>
,
<figureCitation id="EDAEBDF0FFA33A3BC3ED3A31FC4FFEFE" box="[958,987,259,285]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="2" pageNumber="368">19</figureCitation>
) roughly triangular and with more than 15 multifid fimbriae with more than 2 branches; pigmented areas of the gill surface alternating with non-pigmented transparent tracheae. Cerci pale brown with whorls of setae in each segment.
</paragraph>
<caption id="21EAF1FDFFA33A3BC0C43D4DFF7BF958" ID-DOI="http://doi.org/10.5281/zenodo.4889228" ID-Zenodo-Dep="4889228" httpUri="https://zenodo.org/record/4889228/files/figure.png" pageId="2" pageNumber="368" startId="2.[151,264,1663,1687]" targetBox="[151,1436,381,1640]" targetPageId="2">
<paragraph id="752AA175FFA33A3BC0C43D4DFF7BF958" blockId="2.[151,1437,1663,1723]" pageId="2" pageNumber="368">
<emphasis id="47E17D67FFA33A3BC0C43D4DFDE3F974" bold="true" box="[151,631,1663,1687]" pageId="2" pageNumber="368">
FIGURES 14.
<taxonomicName id="B295DAF6FFA33A3BC1103D4DFD8BF974" authority="Muthukatturaja &amp; Balasubramanian, 2021" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2021" box="[323,543,1663,1687]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="368" phylum="Arthropoda" rank="species" species="maduraiensis" status="sp. nov.">
<emphasis id="47E17D67FFA33A3BC1103D4DFD8BF974" bold="true" box="[323,543,1663,1687]" italics="true" pageId="2" pageNumber="368">Caenis maduraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5CD2C01CFFA33A3BC2753D4DFDE3F974" box="[550,631,1663,1687]" pageId="2" pageNumber="368" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
dorsal habitus: 1, Mature nymph; 2, Male imago; 3, Female imago; 4, Collection site.
</paragraph>
</caption>
<paragraph id="752AA175FFA33A3AC0943DDBFE06FE32" blockId="2.[151,1437,1769,2012]" lastBlockId="3.[151,1437,151,465]" lastPageId="3" lastPageNumber="369" pageId="2" pageNumber="368">
<emphasis id="47E17D67FFA33A3BC0943DDBFECCF8E0" bold="true" box="[199,344,1769,1795]" pageId="2" pageNumber="368">Male Imago</emphasis>
(in alcohol) (
<figureCitation id="EDAEBDF0FFA33A3BC1A73DDBFDD0F8E0" box="[500,580,1769,1795]" captionStart="FIGURES 14" captionStartId="2.[151,264,1663,1687]" captionTargetBox="[151,1436,381,1640]" captionTargetId="figure-120@2.[151,1436,381,1640]" captionTargetPageId="2" captionText="FIGURES 14. Caenis maduraiensis sp. nov. dorsal habitus: 1, Mature nymph; 2, Male imago; 3, Female imago; 4, Collection site." figureDoi="http://doi.org/10.5281/zenodo.4889228" httpUri="https://zenodo.org/record/4889228/files/figure.png" pageId="2" pageNumber="368">Figs. 2</figureCitation>
,
<figureCitation id="EDAEBDF0FFA33A3BC2023DDBFD08F8E0" box="[593,668,1769,1795]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="2" pageNumber="368">2027</figureCitation>
): body length,
<quantity id="B26D0C90FFA33A3BC3163DDBFC45F8E7" box="[837,977,1769,1796]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.5" metricValueMin="2.2" pageId="2" pageNumber="368" unit="mm" value="2.35" valueMax="2.5" valueMin="2.2">2.22.5 mm</quantity>
; cerci length,
<quantity id="B26D0C90FFA33A3BC4223DDBFB68F8E7" box="[1137,1276,1769,1796]" metricMagnitude="-3" metricUnit="m" metricValue="8.95" metricValueMax="9.1" metricValueMin="8.8" pageId="2" pageNumber="368" unit="mm" value="8.95" valueMax="9.1" valueMin="8.8">8.89.1 mm</quantity>
; terminal filament,
<quantity id="B26D0C90FFA33A3BC08D3C3FFEFDF8CB" box="[222,361,1805,1832]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="9.9" metricValueMin="9.6" pageId="2" pageNumber="368" unit="mm" value="9.75" valueMax="9.9" valueMin="9.6">9.69.9 mm</quantity>
; length of head
<quantity id="B26D0C90FFA33A3BC24D3C3FFD10F8CB" box="[542,644,1805,1832]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="368" unit="mm" value="0.15">0.15 mm</quantity>
, width of head
<quantity id="B26D0C90FFA33A3BC3673C3FFC19F8CB" box="[820,909,1805,1832]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="368" unit="mm" value="0.5">0.5 mm</quantity>
; length of antenna
<quantity id="B26D0C90FFA33A3BC4363C3FFB2AF8CB" box="[1125,1214,1805,1832]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="2" pageNumber="368" unit="mm" value="0.5">0.5 mm</quantity>
; length of forewing
<quantity id="B26D0C90FFA33A3BC0C43C03FEAAF8AF" box="[151,318,1841,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.95" metricValueMax="2.05" metricValueMin="1.85" pageId="2" pageNumber="368" unit="mm" value="1.95" valueMax="2.05" valueMin="1.85">1.852.05 mm</quantity>
; width of forewing 1.0
<quantity id="B26D0C90FFA33A3BC2013C03FD3FF8AF" box="[594,683,1841,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="368" unit="mm" value="1.3">1.3 mm</quantity>
; dorsal diameter of eye
<quantity id="B26D0C90FFA33A3BC3EF3C03FB82F8AF" box="[956,1046,1841,1868]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="368" unit="mm" value="0.1">0.1 mm</quantity>
; distance between compound eyes
<quantity id="B26D0C90FFA33A3BC0C43C67FE9AF893" box="[151,270,1877,1904]" metricMagnitude="-4" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="368" unit="mm" value="0.125">0.125 mm</quantity>
; length of segments in foreleg femur/ tibia/ tarsus (mm): 0.45/0.9/0.6; length of tarsal segments 1 to 5 (mm): 0.03/0.28/0.12/0.11/0.06. Head (
<figureCitation id="EDAEBDF0FFA33A3BC20B3C4BFD38F870" box="[600,684,1913,1939]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="2" pageNumber="368">Fig. 20</figureCitation>
) pale brown, anterior margin and area around antennal base pale brown; scape and pedicel of antennae pale brown, flagellum translucent. Thorax yellowish brown; prosternal triangle with slightly concave sides, tip acutely pointed (
<figureCitation id="EDAEBDF0FFA33A3BC2A93CF3FCDAF838" box="[762,846,1985,2011]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="2" pageNumber="368">Fig. 21</figureCitation>
). Legs: foreleg (
<figureCitation id="EDAEBDF0FFA33A3BC4543CF3FB17F83F" box="[1031,1155,1985,2012]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="2" pageNumber="368">Fig. 2325</figureCitation>
) pale brown, middle and hindleg translucent; fore femur brown, tibia washed brown and two times longer than femur; tarsomere I extremely short; tarsomere II apically with tongue-shaped projections on single side; tarsomeres IIIV provided with short spines (
<figureCitation id="EDAEBDF0FFA23A3AC0BF3BEDFEE2FF1A" box="[236,374,222,249]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="3" pageNumber="369">Figs. 2425</figureCitation>
); segment V short. Wings: Forewings (
<figureCitation id="EDAEBDF0FFA23A3AC37B3BEDFCEDFF1A" box="[808,889,223,249]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="3" pageNumber="369">Fig. 22</figureCitation>
) oval, with few cross veins; cross veins arranged singly except in R
<subScript id="E911A330FFA23A3AC1313A27FEFEFEC0" attach="left" box="[354,362,277,291]" fontSize="6" pageId="3" pageNumber="369">1</subScript>
and R
<subScript id="E911A330FFA23A3AC1E03A27FE2FFEC0" attach="left" box="[435,443,277,291]" fontSize="6" pageId="3" pageNumber="369">4</subScript>
; two cross veins at near ½ of R; MP forked at near 1/3 from base; CuA not forked. Veins C and Sc brown basally and washed brown distally, remaining veins translucent.Abdomen: tergum IVI pale brown remaining terga translucent; sterna whitish. Genitalia (
<figureCitation id="EDAEBDF0FFA23A3AC2AD3A79FCC4FE85" box="[766,848,331,358]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="3" pageNumber="369">Fig. 26</figureCitation>
) pale brown, penis lobes completely fused and produced laterally; styliger sclerite broad with short apophyses; forceps slender, apical part narrowed to the tip, with a long apical spine, and many trichomes on surface (
<figureCitation id="EDAEBDF0FFA23A3AC2873AA1FCBDFE4E" box="[724,809,403,429]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="3" pageNumber="369">Fig. 27</figureCitation>
); cerci and terminal filament translucent with whorls of setae in each segment.
</paragraph>
<caption id="21EAF1FDFFA23A3AC0C43DD0FE3FF8FD" ID-DOI="http://doi.org/10.5281/zenodo.4889230" ID-Zenodo-Dep="4889230" httpUri="https://zenodo.org/record/4889230/files/figure.png" pageId="3" pageNumber="369" startId="3.[151,264,1762,1786]" targetBox="[153,1434,500,1727]" targetPageId="3">
<paragraph id="752AA175FFA23A3AC0C43DD0FE3FF8FD" blockId="3.[151,1436,1761,1822]" pageId="3" pageNumber="369">
<emphasis id="47E17D67FFA23A3AC0C43DD0FD1CF919" bold="true" box="[151,648,1761,1786]" pageId="3" pageNumber="369">
FIGURES 511.
<taxonomicName id="B295DAF6FFA23A3AC1033DD0FDB9F91A" authority="Muthukatturaja &amp; Balasubramanian, 2021" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2021" box="[336,557,1761,1786]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="369" phylum="Arthropoda" rank="species" species="maduraiensis" status="sp. nov.">
<emphasis id="47E17D67FFA23A3AC1033DD0FDB9F91A" bold="true" box="[336,557,1761,1786]" italics="true" pageId="3" pageNumber="369">Caenis maduraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5CD2C01CFFA23A3AC2663DD0FD1CF919" box="[565,648,1762,1786]" pageId="3" pageNumber="369" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mature nymph dorsal view: 5, Labrum; 67, Mandibles; 89, Maxillae; 10, Hypopharynx; 11, Labium.
</paragraph>
</caption>
<paragraph id="752AA175FFA23A3CC0943C7EFCCAFEA2" blockId="3.[151,1437,1868,2039]" lastBlockId="5.[151,1437,151,393]" lastPageId="5" lastPageNumber="371" pageId="3" pageNumber="369">
<emphasis id="47E17D67FFA23A3AC0943C7EFEE5F885" bold="true" box="[199,369,1868,1894]" pageId="3" pageNumber="369">Female Imago</emphasis>
(in alcohol) (
<figureCitation id="EDAEBDF0FFA23A3AC2543C7EFDC1F885" box="[519,597,1868,1894]" captionStart="FIGURES 14" captionStartId="2.[151,264,1663,1687]" captionTargetBox="[151,1436,381,1640]" captionTargetId="figure-120@2.[151,1436,381,1640]" captionTargetPageId="2" captionText="FIGURES 14. Caenis maduraiensis sp. nov. dorsal habitus: 1, Mature nymph; 2, Male imago; 3, Female imago; 4, Collection site." figureDoi="http://doi.org/10.5281/zenodo.4889228" httpUri="https://zenodo.org/record/4889228/files/figure.png" pageId="3" pageNumber="369">Figs. 3</figureCitation>
,
<figureCitation id="EDAEBDF0FFA23A3AC2313C7EFD39F885" box="[610,685,1868,1894]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="3" pageNumber="369">2830</figureCitation>
): body length,
<quantity id="B26D0C90FFA23A3AC3023C7EFC4EF884" box="[849,986,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="4.449999999999999" metricValueMax="4.6" metricValueMin="4.3" pageId="3" pageNumber="369" unit="mm" value="4.449999999999999" valueMax="4.6" valueMin="4.3">4.34.6 mm</quantity>
; cerci length,
<quantity id="B26D0C90FFA23A3AC4253C7EFB6BF884" box="[1142,1279,1868,1895]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="3" pageNumber="369" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.23.5 mm</quantity>
; terminal filament, 4.0
<quantity id="B26D0C90FFA23A3AC15C3C42FEF3F868" box="[271,359,1904,1931]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="369" unit="mm" value="4.3">4.3 mm</quantity>
; length of head,
<quantity id="B26D0C90FFA23A3AC2723C42FDEDF868" box="[545,633,1904,1931]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="369" unit="mm" value="0.3">0.3 mm</quantity>
, width of head,
<quantity id="B26D0C90FFA23A3AC37F3C42FC11F868" box="[812,901,1904,1931]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="369" unit="mm" value="0.8">0.8 mm</quantity>
; length of antenna,
<quantity id="B26D0C90FFA23A3AC4323C42FB2EF868" box="[1121,1210,1904,1931]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="369" unit="mm" value="0.6">0.6 mm</quantity>
; length of forewing,
<quantity id="B26D0C90FFA23A3AC0C43CA6FEB6F84C" box="[151,290,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="3.35" metricValueMax="3.5" metricValueMin="3.2" pageId="3" pageNumber="369" unit="mm" value="3.35" valueMax="3.5" valueMin="3.2">3.23.5 mm</quantity>
; width of forewing,
<quantity id="B26D0C90FFA23A3AC25B3CA6FD07F84C" box="[520,659,1940,1967]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="3" pageNumber="369" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.41.7 mm</quantity>
; distance between compound eyes,
<quantity id="B26D0C90FFA23A3AC4753CA6FB19F84C" box="[1062,1165,1940,1967]" metricMagnitude="-4" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="369" unit="mm" value="0.58">0.58 mm</quantity>
; dorsal diameter of eye,
<quantity id="B26D0C90FFA23A3AC0C43C8AFE94F830" box="[151,256,1976,2003]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="3" pageNumber="369" unit="mm" value="0.12">0.12 mm</quantity>
; length of segments in foreleg femur/tibia/tarsus (mm): 0.52/0.73/0.63; length of tarsal segments (mm): 0.035/0.35/0.12/0.15/0.075. Head (
<figureCitation id="EDAEBDF0FFA23A3AC24C3CEEFDE5F815" box="[543,625,2012,2038]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="3" pageNumber="369">Fig. 28</figureCitation>
) pale brown; anterior margin and area around antennal base pale brown; scape and pedicel of antennae pale brown, flagellum translucent. Thorax yellowish brown; prosternal triangle with concave sides, tip acutely pointed (
<figureCitation id="EDAEBDF0FFA43A3CC27A3B89FDEBFF36" box="[553,639,187,213]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="5" pageNumber="371">Fig. 29</figureCitation>
). Legs: foreleg pale brown remainders translucent; tarsomere II much longer than others. Wings: veins C and Sc brown basally and washed brown distally, remaining veins translucent. Abdomen: terga IVI pale brown remaining terga translucent; sterna whitish; cerci and median terminal filament translucent; posterior margin of sternum IX truncated (
<figureCitation id="EDAEBDF0FFA43A3CC2AE3A15FCC5FEA2" box="[765,849,295,321]" captionStart="FIGURES 1930" captionStartId="5.[151,264,1589,1613]" captionTargetBox="[161,1427,423,1558]" captionTargetId="figure-122@5.[151,1436,417,1566]" captionTargetPageId="5" captionText="FIGURES 1930. Caenis maduraiensis sp. nov. Mature nymph: 19, Gill VI. Male imago: 20, Head; 21, Prosternal triangle; 22, Forewing; 23, Foreleg; 24, Tarsomeres IIIII; 25, Foretarsus detail, 26, Genitalia, 27, Forceps detail. Female imago: 28, Head; 29, Prosternal triangle; 30, subanal plate." figureDoi="http://doi.org/10.5281/zenodo.4889236" httpUri="https://zenodo.org/record/4889236/files/figure.png" pageId="5" pageNumber="371">Fig. 30</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="21EAF1FDFFA53A3DC0C43C98FC8BF805" ID-DOI="http://doi.org/10.5281/zenodo.4889234" ID-Zenodo-Dep="4889234" httpUri="https://zenodo.org/record/4889234/files/figure.png" pageId="4" pageNumber="370" startId="4.[151,264,1962,1986]" targetBox="[181,1405,195,1927]" targetPageId="4">
<paragraph id="752AA175FFA53A3DC0C43C98FC8BF805" blockId="4.[151,1436,1962,2023]" pageId="4" pageNumber="370">
<emphasis id="47E17D67FFA53A3DC0C43C98FD0FF820" bold="true" box="[151,667,1962,1987]" pageId="4" pageNumber="370">
FIGURES 1218.
<taxonomicName id="B295DAF6FFA53A3DC1333C99FDAAF821" authority="Muthukatturaja &amp; Balasubramanian, 2021" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2021" box="[352,574,1962,1987]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="370" phylum="Arthropoda" rank="species" species="maduraiensis" status="sp. nov.">
<emphasis id="47E17D67FFA53A3DC1333C99FDAAF821" bold="true" box="[352,574,1962,1987]" italics="true" pageId="4" pageNumber="370">Caenis maduraiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="5CD2C01CFFA53A3DC2143C99FD0FF820" box="[583,667,1963,1987]" pageId="4" pageNumber="370" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
mature nymph all drawings in dorsal view: 12, Foreleg; 13, Midleg; 14, Hindleg; 15, Hind tarsal claw; 16, Gill I; 17, Gill II; 18, Gill III.
</paragraph>
</caption>
<subSubSection id="3D8FF2FEFFA43A3CC0943A79FEA3FE6A" pageId="5" pageNumber="371" type="etymology">
<paragraph id="752AA175FFA43A3CC0943A79FEA3FE6A" blockId="5.[151,1437,151,393]" pageId="5" pageNumber="371">
<emphasis id="47E17D67FFA43A3CC0943A79FEDAFE86" bold="true" box="[199,334,331,357]" pageId="5" pageNumber="371">Etymology.</emphasis>
The species name
<taxonomicName id="B295DAF6FFA43A3CC2713A79FD2DFE86" authorityName="Muthukatturaja &amp; Balasubramanian" authorityYear="2021" box="[546,697,331,357]" class="Insecta" family="Caenidae" genus="Caenis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="371" phylum="Arthropoda" rank="species" species="maduraiensis">
<emphasis id="47E17D67FFA43A3CC2713A79FD2DFE86" box="[546,697,331,357]" italics="true" pageId="5" pageNumber="371">maduraiensis</emphasis>
</taxonomicName>
refers to the locality, Madurai,
<collectingRegion id="B7516F97FFA43A3CC4443A79FB08FE86" box="[1047,1180,331,357]" country="India" name="Tamil Nadu" pageId="5" pageNumber="371">Tamil Nadu</collectingRegion>
where the new species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>