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<mods:title id="903F852D0DBF7A17C38DF204323DF3D8">Taxonomic revision of Guapinannus Wygodzinsky, 1951 (Hemiptera: Schizopteridae), with description of 19 new species</mods:title>
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<mods:affiliation id="C4ADE99FD43CD18F3869BD4AD8070395">Department of Entomology, University of California, Riverside, 900 University Avenue, 92521 Riverside, CA, USA sara. frankenberg @ ucr. edu; http: // orcid. org / 0000 - 0002 - 7673 - 492 X &amp; https: // orcid. org / 0000 - 0002 - 2141 - 9447</mods:affiliation>
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<taxonomicName id="AFD24D4BFFCDFF93FF226EB4E23EFA4A" authority="Wygodzinsky, 1951" authorityName="Wygodzinsky" authorityYear="1951" box="[151,565,1410,1437]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="genus">
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<emphasis id="5AA6EADAFFCDFF93FF226EB4E136FA4B" bold="true" box="[151,317,1411,1437]" italics="true" pageId="2" pageNumber="263">Guapinannus</emphasis>
<bibRefCitation id="0C434B39FFCDFF93FEF16EB5E23EFA4A" author="Wygodzinsky, P." box="[324,565,1410,1436]" pageId="2" pageNumber="263" pagination="259 - 270" refId="ref19918" refString="Wygodzinsky, P. (1951) Descripcion de generos y especies nuevos de la familia Cryptostemmatidae (Hemiptera). Revista Brasileira de Biologia, 11, 259 - 270." type="journal article" year="1951">Wygodzinsky, 1951</bibRefCitation>
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<paragraph id="686D36C8FFCDFF93FF226E90E100FA17" blockId="2.[151,565,1410,1473]" box="[151,267,1447,1473]" pageId="2" pageNumber="263">
(
<figureCitation id="F0E92A4DFFCDFF93FF2A6E90E109FA17" box="[159,258,1447,1473]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStartId-0="4.[151,250,1955,1979]" captionStartId-1="10.[151,250,1929,1953]" captionStartId-2="15.[151,250,1926,1950]" captionStartId-3="18.[151,250,1651,1675]" captionStartId-4="20.[151,250,1688,1712]" captionStartId-5="21.[151,250,1490,1514]" captionStartId-6="23.[151,250,1943,1967]" captionStartId-7="24.[151,250,1053,1077]" captionStartId-8="24.[151,250,2012,2036]" captionTargetBox-0="[152,1434,851,1929]" captionTargetBox-1="[167,1421,181,1905]" captionTargetBox-2="[168,1420,181,1903]" captionTargetBox-3="[151,1437,181,1627]" captionTargetBox-4="[153,1433,183,1662]" captionTargetBox-5="[153,1435,183,1464]" captionTargetBox-6="[219,1368,193,1906]" captionTargetBox-7="[224,1363,181,1028]" captionTargetBox-8="[151,1435,1137,1988]" captionTargetId-0="figure-369@4.[151,1436,849,1931]" captionTargetId-1="figure-24@10.[167,1421,181,1905]" captionTargetId-2="figure-17@15.[168,1420,181,1903]" captionTargetId-3="figure-24@18.[151,1437,181,1627]" captionTargetId-4="figure-24@20.[151,1436,181,1664]" captionTargetId-5="figure-17@21.[151,1435,181,1465]" captionTargetId-6="figure-17@23.[205,1383,181,1919]" captionTargetId-7="figure-24@24.[224,1363,181,1028]" captionTargetId-8="figure-44@24.[151,1435,1137,1988]" captionTargetPageId-0="4" captionTargetPageId-1="10" captionTargetPageId-2="15" captionTargetPageId-3="18" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="23" captionTargetPageId-7="24" captionTargetPageId-8="24" captionText-0="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." captionText-1="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-2="FIGURE 3. Dorsal habitus of Guapinannus graziae, sp. n., G. minutus, sp. n., G. orbiculatus, sp. n., G. plurilobus, sp. n., G. policis, sp. n., G. robustus, sp. n., and G. sinuosus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-3="FIGURE 4. Dorsal habitus of Guapinannus tatumbia, sp. n., G. tenuis, sp. n., G. tergus, sp. n., G. trilobus, sp. n., and G. uncus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-4="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." captionText-5="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." captionText-6="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." captionText-7="FIGURE 8. Female Guapinannus sp. (UCR_ENT 00101286), genitalic structures including ovipositor and spermathecal gland, and spermathecal duct." captionText-8="FIGURE 9. Known distributions of Guapinannus spp." figureDoi-0="http://doi.org/10.5281/zenodo.4691722" figureDoi-1="http://doi.org/10.5281/zenodo.4691724" figureDoi-2="http://doi.org/10.5281/zenodo.4691726" figureDoi-3="http://doi.org/10.5281/zenodo.4691730" figureDoi-4="http://doi.org/10.5281/zenodo.4691734" figureDoi-5="http://doi.org/10.5281/zenodo.4691736" figureDoi-6="http://doi.org/10.5281/zenodo.4691739" figureDoi-7="http://doi.org/10.5281/zenodo.4691741" figureDoi-8="http://doi.org/10.5281/zenodo.4691743" httpUri-0="https://zenodo.org/record/4691722/files/figure.png" httpUri-1="https://zenodo.org/record/4691724/files/figure.png" httpUri-2="https://zenodo.org/record/4691726/files/figure.png" httpUri-3="https://zenodo.org/record/4691730/files/figure.png" httpUri-4="https://zenodo.org/record/4691734/files/figure.png" httpUri-5="https://zenodo.org/record/4691736/files/figure.png" httpUri-6="https://zenodo.org/record/4691739/files/figure.png" httpUri-7="https://zenodo.org/record/4691741/files/figure.png" httpUri-8="https://zenodo.org/record/4691743/files/figure.png" pageId="2" pageNumber="263">Figs 1–9</figureCitation>
)
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<typeStatus id="B769886AFFCDFF93FF226EDCE0C7F9D5" box="[151,204,1515,1539]" pageId="2" pageNumber="263">Type</typeStatus>
species:
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<taxonomicName id="AFD24D4BFFCDFF93FE996EDBE2C6F9D5" authority="Wygodzinsky, 1951" authorityName="Wygodzinsky" authorityYear="1951" box="[300,717,1515,1539]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="5AA6EADAFFCDFF93FE996EDBE1F4F9D5" box="[300,511,1515,1539]" italics="true" pageId="2" pageNumber="263">Guapinannus bierigi</emphasis>
<bibRefCitation id="0C434B39FFCDFF93FDB16EDCE2C6F9D5" author="Wygodzinsky, P." box="[516,717,1515,1539]" pageId="2" pageNumber="263" pagination="259 - 270" refId="ref19918" refString="Wygodzinsky, P. (1951) Descripcion de generos y especies nuevos de la familia Cryptostemmatidae (Hemiptera). Revista Brasileira de Biologia, 11, 259 - 270." type="journal article" year="1951">Wygodzinsky, 1951</bibRefCitation>
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<paragraph id="686D36C8FFCDFF93FF226D06E25BF929" blockId="2.[151,1437,1585,2008]" pageId="2" pageNumber="263">
<emphasis id="5AA6EADAFFCDFF93FF226D06E11FF99D" bold="true" box="[151,276,1585,1611]" pageId="2" pageNumber="263">Diagnosis:</emphasis>
Recognized among 
<taxonomicName id="AFD24D4BFFCDFF93FDB46D06E2ACF99D" authorityName="Reuter" authorityYear="1891" box="[513,679,1585,1611]" class="Insecta" family="Schizopteridae" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="family">Schizopteridae</taxonomicName>
by the relatively small eyes (
<figureCitation id="F0E92A4DFFCDFF93FC4A6D06E474F99A" box="[1023,1151,1585,1612]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="2" pageNumber="263">Fig. 5C, D</figureCitation>
), four-segmented labium (
<figureCitation id="F0E92A4DFFCDFF93FF156D62E0FCF9B9" box="[160,247,1621,1647]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="2" pageNumber="263">Fig. 1A</figureCitation>
), flat and wide forewings (
<figureCitation id="F0E92A4DFFCDFF93FDAB6D62E28AF9B9" box="[542,641,1621,1647]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="4.[151,250,1955,1979]" captionStartId-1="10.[151,250,1929,1953]" captionStartId-2="15.[151,250,1926,1950]" captionStartId-3="18.[151,250,1651,1675]" captionTargetBox-0="[152,1434,851,1929]" captionTargetBox-1="[167,1421,181,1905]" captionTargetBox-2="[168,1420,181,1903]" captionTargetBox-3="[151,1437,181,1627]" captionTargetId-0="figure-369@4.[151,1436,849,1931]" captionTargetId-1="figure-24@10.[167,1421,181,1905]" captionTargetId-2="figure-17@15.[168,1420,181,1903]" captionTargetId-3="figure-24@18.[151,1437,181,1627]" captionTargetPageId-0="4" captionTargetPageId-1="10" captionTargetPageId-2="15" captionTargetPageId-3="18" captionText-0="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." captionText-1="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-2="FIGURE 3. Dorsal habitus of Guapinannus graziae, sp. n., G. minutus, sp. n., G. orbiculatus, sp. n., G. plurilobus, sp. n., G. policis, sp. n., G. robustus, sp. n., and G. sinuosus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-3="FIGURE 4. Dorsal habitus of Guapinannus tatumbia, sp. n., G. tenuis, sp. n., G. tergus, sp. n., G. trilobus, sp. n., and G. uncus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi-0="http://doi.org/10.5281/zenodo.4691722" figureDoi-1="http://doi.org/10.5281/zenodo.4691724" figureDoi-2="http://doi.org/10.5281/zenodo.4691726" figureDoi-3="http://doi.org/10.5281/zenodo.4691730" httpUri-0="https://zenodo.org/record/4691722/files/figure.png" httpUri-1="https://zenodo.org/record/4691724/files/figure.png" httpUri-2="https://zenodo.org/record/4691726/files/figure.png" httpUri-3="https://zenodo.org/record/4691730/files/figure.png" pageId="2" pageNumber="263">Figs 1–4</figureCitation>
), unique forewing venation with long fracture in the costal margin, long rc1, rc, and tc cells (
<figureCitation id="F0E92A4DFFCDFF93FECC6D4EE1DFF945" box="[377,468,1657,1683]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="2" pageNumber="263">Figs 1B</figureCitation>
, 
<figureCitation id="F0E92A4DFFCDFF93FE556D4EE1FBF945" box="[480,496,1657,1683]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="2" pageNumber="263">2</figureCitation>
, 
<figureCitation id="F0E92A4DFFCDFF93FE496D4EE22AF942" box="[508,545,1657,1684]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="2" pageNumber="263">6A</figureCitation>
), C+S, with An1/2 claval process except 
<taxonomicName id="AFD24D4BFFCDFF93FC5B6D4DE453F945" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1006,1112,1657,1684]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="263" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCDFF93FC5B6D4DE453F945" box="[1006,1112,1657,1684]" italics="true" pageId="2" pageNumber="263">G. dispar</emphasis>
</taxonomicName>
<emphasis id="5AA6EADAFFCDFF93FBEA6D4EE495F945" bold="true" box="[1119,1182,1657,1683]" pageId="2" pageNumber="263">
<taxonomicNameLabel id="419557A1FFCDFF93FBEA6D4EE495F945" box="[1119,1182,1657,1683]" pageId="2" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F0E92A4DFFCDFF93FB196D4EE4FEF945" box="[1196,1269,1657,1683]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="2" pageNumber="263">Figs 2</figureCitation>
, 
<figureCitation id="F0E92A4DFFCDFF93FAB46D4EE543F942" box="[1281,1352,1657,1684]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="2" pageNumber="263">6A, D</figureCitation>
), better developed in males than in females (
<figureCitation id="F0E92A4DFFCDFF93FD9B6DAAE275F961" box="[558,638,1693,1719]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="2" pageNumber="263">Fig 1B</figureCitation>
), males with wing organ associated with An1 on corium adjacent to the claval process (
<figureCitation id="F0E92A4DFFCDFF93FEF16DF6E1A2F90D" box="[324,425,1729,1755]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="10.[151,250,1929,1953]" captionStartId-1="15.[151,250,1926,1950]" captionStartId-2="18.[151,250,1651,1675]" captionTargetBox-0="[167,1421,181,1905]" captionTargetBox-1="[168,1420,181,1903]" captionTargetBox-2="[151,1437,181,1627]" captionTargetId-0="figure-24@10.[167,1421,181,1905]" captionTargetId-1="figure-17@15.[168,1420,181,1903]" captionTargetId-2="figure-24@18.[151,1437,181,1627]" captionTargetPageId-0="10" captionTargetPageId-1="15" captionTargetPageId-2="18" captionText-0="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-1="FIGURE 3. Dorsal habitus of Guapinannus graziae, sp. n., G. minutus, sp. n., G. orbiculatus, sp. n., G. plurilobus, sp. n., G. policis, sp. n., G. robustus, sp. n., and G. sinuosus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-2="FIGURE 4. Dorsal habitus of Guapinannus tatumbia, sp. n., G. tenuis, sp. n., G. tergus, sp. n., G. trilobus, sp. n., and G. uncus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi-0="http://doi.org/10.5281/zenodo.4691724" figureDoi-1="http://doi.org/10.5281/zenodo.4691726" figureDoi-2="http://doi.org/10.5281/zenodo.4691730" httpUri-0="https://zenodo.org/record/4691724/files/figure.png" httpUri-1="https://zenodo.org/record/4691726/files/figure.png" httpUri-2="https://zenodo.org/record/4691730/files/figure.png" pageId="2" pageNumber="263">Figs 2–4</figureCitation>
, 
<figureCitation id="F0E92A4DFFCDFF93FE016DF6E1F0F90A" box="[436,507,1729,1756]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="2" pageNumber="263">6A, D</figureCitation>
), 2-2-3 tarsal formula in both sexes, male genitalia simple (
<figureCitation id="F0E92A4DFFCDFF93FB206DF6E4D2F90A" box="[1173,1241,1729,1756]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="2" pageNumber="263">Fig. 7</figureCitation>
) and female with well-developed ovipositor (
<figureCitation id="F0E92A4DFFCDFF93FE7E6DD2E22DF929" box="[459,550,1765,1791]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="2" pageNumber="263">Figs 1C</figureCitation>
, 
<figureCitation id="F0E92A4DFFCDFF93FD866DD2E248F929" box="[563,579,1765,1791]" captionStart="FIGURE 8" captionStartId="24.[151,250,1053,1077]" captionTargetBox="[224,1363,181,1028]" captionTargetId="figure-24@24.[224,1363,181,1028]" captionTargetPageId="24" captionText="FIGURE 8. Female Guapinannus sp. (UCR_ENT 00101286), genitalic structures including ovipositor and spermathecal gland, and spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.4691741" httpUri="https://zenodo.org/record/4691741/files/figure.png" pageId="2" pageNumber="263">8</figureCitation>
).
</paragraph>
<paragraph id="686D36C8FFCDFF92FF726C3EE3D2F9FB" blockId="2.[151,1437,1585,2008]" lastBlockId="3.[151,1437,150,2013]" lastPageId="3" lastPageNumber="264" pageId="2" pageNumber="263">
<emphasis id="5AA6EADAFFCDFF93FF726C3EE1C7F8F5" bold="true" box="[199,460,1801,1827]" pageId="2" pageNumber="263">Re-description: Male:</emphasis>
Ovoid (greatest length to width ratio 1.57–1.85) to elongate ovoid (length to width ratio 3.4), total body length 
<quantity id="AF2A9B2DFFCDFF93FE3A6C1AE238F89E" box="[399,563,1837,1864]" metricMagnitude="-3" metricUnit="m" metricValue="1.36" metricValueMax="2.06" metricValueMin="0.66" pageId="2" pageNumber="263" unit="mm" value="1.36" valueMax="2.06" valueMin="0.66">0.66–2.06 mm</quantity>
; length from posterior margin of pronotum to posterior wing margin (
<quantity id="AF2A9B2DFFCDFF93FA9E6C1AE0CCF8BA" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.49" metricValueMin="0.51" pageId="2" pageNumber="263" unit="mm" value="1.0" valueMax="1.49" valueMin="0.51">0.51–1.49 mm</quantity>
). 
<emphasis id="5AA6EADAFFCDFF93FF6F6C66E1CFF8BD" bold="true" box="[218,452,1873,1899]" pageId="2" pageNumber="263">General coloration:</emphasis>
uniformly pale or a mix of pale and darker brown or reddish brown. 
<emphasis id="5AA6EADAFFCDFF93FB726C66E530F8BD" bold="true" box="[1223,1339,1873,1899]" pageId="2" pageNumber="263">Vestiture:</emphasis>
Dorsum with dense setae, setae long or short, setae on posterior margin of forewing long or extremely long. 
<emphasis id="5AA6EADAFFCDFF93FA956C42E0EBF865" bold="true" pageId="2" pageNumber="263">Structure: Head:</emphasis>
Eye ranging from small (less than one quarter head height) to relatively large (almost half as high as head) (
<figureCitation id="F0E92A4DFFCDFF93FF156C8AE110F80E" box="[160,283,1981,2008]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="2" pageNumber="263">Fig. 5C, D</figureCitation>
); ocellus adjacent to eye (
<figureCitation id="F0E92A4DFFCDFF93FDF56C8AE291F80E" box="[576,666,1981,2008]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="2" pageNumber="263">Fig. 5D</figureCitation>
), at 11 o’clock position seen in lateral view and about twice the size of one ommatidium; four-segmented labium with segments 1, 2 and 3 of similar length and 4 about 2.5–3 times as long as any other segment (
<figureCitation id="F0E92A4DFFCCFF92FE786B8CE22CFF03" box="[461,551,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Fig. 1A</figureCitation>
); muscle scars present on the frons and vertex present (
<figureCitation id="F0E92A4DFFCCFF92FB2D6B8CE4FEFF03" box="[1176,1269,187,213]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Figs 1A</figureCitation>
, 
<figureCitation id="F0E92A4DFFCCFF92FAB76B8DE52DFF03" box="[1282,1318,186,213]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="3" pageNumber="264">5C</figureCitation>
). 
<emphasis id="5AA6EADAFFCCFF92FA8F6B8CE597FF03" bold="true" box="[1338,1436,187,213]" pageId="3" pageNumber="264">Thorax:</emphasis>
Collar in dorsal view narrow (
<figureCitation id="F0E92A4DFFCCFF92FE456BE8E241FF2F" box="[496,586,222,249]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="3" pageNumber="264">Fig. 5C</figureCitation>
); proepisternal lobe inflated (
<figureCitation id="F0E92A4DFFCCFF92FC2E6BE8E3FDFF2F" box="[923,1014,222,249]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="3" pageNumber="264">Fig. 5D</figureCitation>
); separation of anterior and posterior lobes (dorsal view) marked by weak depression (
<figureCitation id="F0E92A4DFFCCFF92FD036A34E306FECB" box="[694,781,258,285]" captionStart="FIGURE 5" captionStartId="20.[151,250,1688,1712]" captionTargetBox="[153,1433,183,1662]" captionTargetId="figure-24@20.[151,1436,181,1664]" captionTargetPageId="20" captionText="FIGURE 5. Scanning electron micrographs of head, thoracic structures, and tarsi of the male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Head, frontal view. B: Head (expanded), frontal view. C: Head and pronotum, dorsal view, showing collar and vestiture on pronotum. D: Head and pronotum, lateral view, showing collar and vestiture on pronotum. E: Hind left leg, ventral view, showing claws and tarsomeres. F: Thorax, lateral view, of ventral portion of head and prothorax, showing scutellum." figureDoi="http://doi.org/10.5281/zenodo.4691734" httpUri="https://zenodo.org/record/4691734/files/figure.png" pageId="3" pageNumber="264">Fig. 5C</figureCitation>
); anterior pronotal lobe with muscle scars, margin concave or straight; posterior margin slightly concave to almost straight; pits on pronotum ranging from small to large, evenly distributed or grouped (
<figureCitation id="F0E92A4DFFCCFF92FE5A6A7CE24CFEB3" box="[495,583,331,357]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Fig. 1A</figureCitation>
); scutellum in dorsal view abruptly or gradually narrowed to tip, tip pointed, rounded, or slightly inflated (
<figureCitation id="F0E92A4DFFCCFF92FE6B6A58E24EFE5F" box="[478,581,367,393]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="10.[151,250,1929,1953]" captionStartId-1="15.[151,250,1926,1950]" captionStartId-2="18.[151,250,1651,1675]" captionTargetBox-0="[167,1421,181,1905]" captionTargetBox-1="[168,1420,181,1903]" captionTargetBox-2="[151,1437,181,1627]" captionTargetId-0="figure-24@10.[167,1421,181,1905]" captionTargetId-1="figure-17@15.[168,1420,181,1903]" captionTargetId-2="figure-24@18.[151,1437,181,1627]" captionTargetPageId-0="10" captionTargetPageId-1="15" captionTargetPageId-2="18" captionText-0="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-1="FIGURE 3. Dorsal habitus of Guapinannus graziae, sp. n., G. minutus, sp. n., G. orbiculatus, sp. n., G. plurilobus, sp. n., G. policis, sp. n., G. robustus, sp. n., and G. sinuosus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-2="FIGURE 4. Dorsal habitus of Guapinannus tatumbia, sp. n., G. tenuis, sp. n., G. tergus, sp. n., G. trilobus, sp. n., and G. uncus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi-0="http://doi.org/10.5281/zenodo.4691724" figureDoi-1="http://doi.org/10.5281/zenodo.4691726" figureDoi-2="http://doi.org/10.5281/zenodo.4691730" httpUri-0="https://zenodo.org/record/4691724/files/figure.png" httpUri-1="https://zenodo.org/record/4691726/files/figure.png" httpUri-2="https://zenodo.org/record/4691730/files/figure.png" pageId="3" pageNumber="264">Figs 2–4</figureCitation>
). 
<emphasis id="5AA6EADAFFCCFF92FDED6A59E2D8FE5E" bold="true" box="[600,723,366,392]" pageId="3" pageNumber="264">Forewing:</emphasis>
With outline elongate oval or broad anteriorly; with (
<taxonomicName id="AFD24D4BFFCCFF92FA9E6A58E59DFE5F" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1323,1430,367,393]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FA9E6A58E59DFE5F" box="[1323,1430,367,393]" italics="true" pageId="3" pageNumber="264">G. dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FF226AA4E0DDFE7B" bold="true" box="[151,214,403,429]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FF226AA4E0DDFE7B" box="[151,214,403,429]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<figureCitation id="F0E92A4DFFCCFF92FF576AA4E12FFE7B" box="[226,292,403,429]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
) or without (all other species) distinct honey-comb pattern; membrane (cub, dc1, rc2–3 and areas distal to these cells) contributing about one third or less to forewing length; posterior distal margin rounded or square; clavus twice as long as wide; vein-tracing areoles along margins of a combination of scc, rc1, rc1, rc2, and tc on proximal part of wing absent or present (
<figureCitation id="F0E92A4DFFCCFF92FDEA6AC8E2D2FDCF" box="[607,729,511,537]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="3" pageNumber="264">Fig. 6A, C</figureCitation>
); C+Sc narrow (
<figureCitation id="F0E92A4DFFCCFF92FC276AC8E3DDFDCF" box="[914,982,511,537]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC576AC8E472FDCF" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[994,1145,511,537]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC576AC8E472FDCF" box="[994,1145,511,537]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FB336AC8E4CDFDCF" bold="true" box="[1158,1222,511,537]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FB336AC8E4CDFDCF" box="[1158,1222,511,537]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), fairly broad (
<figureCitation id="F0E92A4DFFCCFF92FADA6AC8E0ACFDEB" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<emphasis id="5AA6EADAFFCCFF92FF006914E11DFDEB" box="[181,278,547,573]" italics="true" pageId="3" pageNumber="264">G. clava</emphasis>
, 
<emphasis id="5AA6EADAFFCCFF92FE906914E16CFDEB" bold="true" box="[293,359,547,573]" pageId="3" pageNumber="264">sp. n.</emphasis>
), broad (
<figureCitation id="F0E92A4DFFCCFF92FE656914E21DFDEB" box="[464,534,547,573]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FD916914E28EFDEB" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[548,645,547,573]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="falcis" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FD916914E28EFDEB" box="[548,645,547,573]" italics="true" pageId="3" pageNumber="264">G. falcis</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FD206914E2DDFDEB" bold="true" box="[661,726,547,573]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FD206914E2DDFDEB" box="[661,726,547,573]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), or very broad (
<figureCitation id="F0E92A4DFFCCFF92FC2F6914E3D4FDEB" box="[922,991,547,573]" captionStart="FIGURE 3" captionStartId="15.[151,250,1926,1950]" captionTargetBox="[168,1420,181,1903]" captionTargetId="figure-17@15.[168,1420,181,1903]" captionTargetPageId="15" captionText="FIGURE 3. Dorsal habitus of Guapinannus graziae, sp. n., G. minutus, sp. n., G. orbiculatus, sp. n., G. plurilobus, sp. n., G. policis, sp. n., G. robustus, sp. n., and G. sinuosus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691726" httpUri="https://zenodo.org/record/4691726/files/figure.png" pageId="3" pageNumber="264">Fig. 3</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC5B6914E464FDEB" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1006,1135,547,573]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="lutuosus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC5B6914E464FDEB" box="[1006,1135,547,573]" italics="true" pageId="3" pageNumber="264">G. lutuosus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FBCA6914E4CAFDEB" bold="true" box="[1151,1217,547,573]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FBCA6914E4CAFDEB" box="[1151,1217,547,573]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), either of uniform width (
<figureCitation id="F0E92A4DFFCCFF92FF506970E122FDB7" box="[229,297,583,609]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FE816970E1C1FDB7" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[308,458,583,609]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FE816970E1C1FDB7" box="[308,458,583,609]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE626970E21DFDB7" bold="true" box="[471,534,583,609]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE626970E21DFDB7" box="[471,534,583,609]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or wider at widest part of wing (
<figureCitation id="F0E92A4DFFCCFF92FC326970E3C1FDB7" box="[903,970,583,609]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC606970E45DFDB7" box="[981,1110,583,609]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="auriculu">
<emphasis id="5AA6EADAFFCCFF92FC606970E45DFDB7" box="[981,1110,583,609]" italics="true" pageId="3" pageNumber="264">G. auriculu</emphasis>
</taxonomicName>
s, 
<emphasis id="5AA6EADAFFCCFF92FBDB6970E4A7FDB7" bold="true" box="[1134,1196,583,609]" pageId="3" pageNumber="264">sp. n.</emphasis>
); R flattened in distal part (black arrow in 
<figureCitation id="F0E92A4DFFCCFF92FECE695CE1DAFD53" box="[379,465,619,645]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Fig. 1B</figureCitation>
); distal process of R pronounced (white arrow in 
<figureCitation id="F0E92A4DFFCCFF92FC49695CE45BFD53" box="[1020,1104,619,645]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Fig. 1B</figureCitation>
) or weakly developed; An1/2 claval process well developed in males (except 
<taxonomicName id="AFD24D4BFFCCFF92FD0569B8E31DFD7E" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[688,790,655,681]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="tenuis" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FD0569B8E31DFD7E" box="[688,790,655,681]" italics="true" pageId="3" pageNumber="264">G. tenuis</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC9169B8E36FFD7F" bold="true" box="[804,868,655,681]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC9169B8E36FFD7F" box="[804,868,655,681]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), overlapping base of An1 wing organ, apex acute and in lateral view pointing dorsad (
<figureCitation id="F0E92A4DFFCCFF92FD936984E280FD1B" box="[550,651,691,717]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStartId-0="10.[151,250,1929,1953]" captionStartId-1="15.[151,250,1926,1950]" captionStartId-2="18.[151,250,1651,1675]" captionTargetBox-0="[167,1421,181,1905]" captionTargetBox-1="[168,1420,181,1903]" captionTargetBox-2="[151,1437,181,1627]" captionTargetId-0="figure-24@10.[167,1421,181,1905]" captionTargetId-1="figure-17@15.[168,1420,181,1903]" captionTargetId-2="figure-24@18.[151,1437,181,1627]" captionTargetPageId-0="10" captionTargetPageId-1="15" captionTargetPageId-2="18" captionText-0="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-1="FIGURE 3. Dorsal habitus of Guapinannus graziae, sp. n., G. minutus, sp. n., G. orbiculatus, sp. n., G. plurilobus, sp. n., G. policis, sp. n., G. robustus, sp. n., and G. sinuosus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." captionText-2="FIGURE 4. Dorsal habitus of Guapinannus tatumbia, sp. n., G. tenuis, sp. n., G. tergus, sp. n., G. trilobus, sp. n., and G. uncus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi-0="http://doi.org/10.5281/zenodo.4691724" figureDoi-1="http://doi.org/10.5281/zenodo.4691726" figureDoi-2="http://doi.org/10.5281/zenodo.4691730" httpUri-0="https://zenodo.org/record/4691724/files/figure.png" httpUri-1="https://zenodo.org/record/4691726/files/figure.png" httpUri-2="https://zenodo.org/record/4691730/files/figure.png" pageId="3" pageNumber="264">Figs 2–4</figureCitation>
, 
<figureCitation id="F0E92A4DFFCCFF92FD236984E2D7FD1B" box="[662,732,691,717]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="3" pageNumber="264">6A, D</figureCitation>
); An1 wing organ on corium strongly inflated in all males, with or without (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FEE069E0E192FD27" box="[341,409,727,753]" captionStart="FIGURE 4" captionStartId="18.[151,250,1651,1675]" captionTargetBox="[151,1437,181,1627]" captionTargetId="figure-24@18.[151,1437,181,1627]" captionTargetPageId="18" captionText="FIGURE 4. Dorsal habitus of Guapinannus tatumbia, sp. n., G. tenuis, sp. n., G. tergus, sp. n., G. trilobus, sp. n., and G. uncus, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691730" httpUri="https://zenodo.org/record/4691730/files/figure.png" pageId="3" pageNumber="264">Fig. 4</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FE1369E0E204FD26" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[422,527,727,753]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="tergus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FE1369E0E204FD26" box="[422,527,727,753]" italics="true" pageId="3" pageNumber="264">G. tergus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FDAB69E0E255FD27" bold="true" box="[542,606,727,753]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FDAB69E0E255FD27" box="[542,606,727,753]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) notch articulating with claval process, wing organ rounded, elongate, or acute, with or without median notch; scc about 1.5 times longer than wide (
<figureCitation id="F0E92A4DFFCCFF92FC4569CCE442FCC0" box="[1008,1097,763,790]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="3" pageNumber="264">Fig. 6A</figureCitation>
) (except ~2 times longer than wide in 
<taxonomicName id="AFD24D4BFFCCFF92FF466828E18CFCEE" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[243,391,799,825]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="castigatus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FF466828E18CFCEE" box="[243,391,799,825]" italics="true" pageId="3" pageNumber="264">G. castigatus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE236828E1DDFCEF" bold="true" box="[406,470,799,825]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE236828E1DDFCEF" box="[406,470,799,825]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); scc undivided except in 
<taxonomicName id="AFD24D4BFFCCFF92FD4A6828E361FCEF" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[767,874,799,825]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FD4A6828E361FCEF" box="[767,874,799,825]" italics="true" pageId="3" pageNumber="264">G. dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FCCD6828E3B3FCEF" bold="true" box="[888,952,799,825]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FCCD6828E3B3FCEF" box="[888,952,799,825]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="F0E92A4DFFCCFF92FC7D6828E406FCEF" box="[968,1037,799,825]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
); rc1 about 3 times as long as wide except in 
<taxonomicName id="AFD24D4BFFCCFF92FEB36874E17AFC8B" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[262,369,835,861]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FEB36874E17AFC8B" box="[262,369,835,861]" italics="true" pageId="3" pageNumber="264">G. dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE356874E1CAFC8B" bold="true" box="[384,449,835,861]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE356874E1CAFC8B" box="[384,449,835,861]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, ranging from parallel-sided to slightly or much wider anteriorly; rc about 4 or 5 times as long as wide; tc 3 times or more than 3 times as long as wide; part of An1 and Cu forming posterodistal margin of tc of similar width as other veins (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FDCC68BCE2B5FC73" box="[633,702,907,933]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FD7968BCE321FC72" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[716,810,907,933]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="artus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FD7968BCE321FC72" box="[716,810,907,933]" italics="true" pageId="3" pageNumber="264">G. artus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC8C68BCE371FC73" bold="true" box="[825,890,907,933]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC8C68BCE371FC73" box="[825,890,907,933]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or thicker (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FBF668BCE483FC73" box="[1091,1160,907,933]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FB2068BCE4FDFC73" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1173,1270,907,933]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="falcis" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FB2068BCE4FDFC73" box="[1173,1270,907,933]" italics="true" pageId="3" pageNumber="264">G. falcis</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FAB068BCE54DFC73" bold="true" box="[1285,1350,907,933]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FAB068BCE54DFC73" box="[1285,1350,907,933]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); distal margin of rc1, rc, and tc cells double s-shaped (
<figureCitation id="F0E92A4DFFCCFF92FD166898E2EDFC1F" box="[675,742,943,969]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FD446898E383FC1F" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[753,904,943,969]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FD446898E383FC1F" box="[753,904,943,969]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC216898E3D8FC1F" bold="true" box="[916,979,943,969]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC216898E3D8FC1F" box="[916,979,943,969]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), s-shaped (
<figureCitation id="F0E92A4DFFCCFF92FBEF6898E490FC1F" box="[1114,1179,943,969]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Fig. 1</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FB1D6898E51EFC1F" authorityName="Wygodzinsky" authorityYear="1951" box="[1192,1301,943,969]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="bierigi">
<emphasis id="5AA6EADAFFCCFF92FB1D6898E51EFC1F" box="[1192,1301,943,969]" italics="true" pageId="3" pageNumber="264">G. bierigi</emphasis>
</taxonomicName>
), or angular s-shaped (
<figureCitation id="F0E92A4DFFCCFF92FEBD68E4E140FC3B" box="[264,331,979,1005]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<emphasis id="5AA6EADAFFCCFF92FEE268E4E1C8FC3B" box="[343,451,979,1005]" italics="true" pageId="3" pageNumber="264">
<taxonomicName id="AFD24D4BFFCCFF92FEE268E4E1B4FC3B" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[343,447,979,1005]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">G. dispar</taxonomicName>
,
</emphasis>
<emphasis id="5AA6EADAFFCCFF92FE7F68E4E203FC3B" bold="true" box="[458,520,979,1005]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE7F68E4E203FC3B" box="[458,520,979,1005]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); rc2–3 almost triangular (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FCDD68E4E3A0FC3B" box="[872,939,979,1005]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC0368E4E446FC3B" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[950,1101,979,1005]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC0368E4E446FC3B" box="[950,1101,979,1005]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FBEC68E4E493FC3B" bold="true" box="[1113,1176,979,1005]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FBEC68E4E493FC3B" box="[1113,1176,979,1005]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), trapezoidal (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FAC568E4E0ACFBC7" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FF0668C0E143FBC6" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[179,328,1015,1041]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="castigatus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FF0668C0E143FBC6" box="[179,328,1015,1041]" italics="true" pageId="3" pageNumber="264">G. castigatus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FEE068C0E195FBC6" bold="true" box="[341,414,1014,1041]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FEE068C0E19EFBC6" box="[341,405,1014,1041]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
)
</emphasis>
, or rectangular (
<figureCitation id="F0E92A4DFFCCFF92FDE268C0E297FBC7" box="[599,668,1015,1041]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FD1D68C0E319FBC7" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[680,786,1015,1041]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FD1D68C0E319FBC7" box="[680,786,1015,1041]" italics="true" pageId="3" pageNumber="264">G. dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC9568C0E354FBC7" bold="true" box="[800,863,1015,1041]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC9568C0E354FBC7" box="[800,863,1015,1041]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
); cub oval, almost reaching wing margin or distant form wing margin (except elongate in 
<taxonomicName id="AFD24D4BFFCCFF92FDF36F2CE2BAFBE3" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[582,689,1051,1077]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="5AA6EADAFFCCFF92FDF36F2CE2BAFBE3" box="[582,689,1051,1077]" italics="true" pageId="3" pageNumber="264">G. dispar</emphasis>
</taxonomicName>
; 
<figureCitation id="F0E92A4DFFCCFF92FD0A6F2CE308FBE3" box="[703,771,1051,1077]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
); dc1 with basal portion fairly narrow and elongate (
<figureCitation id="F0E92A4DFFCCFF92FAE66F2CE59CFBE3" box="[1363,1431,1051,1077]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FF226F08E125FB8F" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[151,302,1087,1113]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FF226F08E125FB8F" box="[151,302,1087,1113]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE8F6F08E172FB8F" bold="true" box="[314,377,1087,1113]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE8F6F08E172FB8F" box="[314,377,1087,1113]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or relatively wide and short (
<figureCitation id="F0E92A4DFFCCFF92FD726F08E301FB8F" box="[711,778,1087,1113]" captionStart="FIGURE 2" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[167,1421,181,1905]" captionTargetId="figure-24@10.[167,1421,181,1905]" captionTargetPageId="10" captionText="FIGURE 2. Habitus in dorsal, lateral, and ventral view of Guapinannus anaticulus, sp. n., and in dorsal view of G. artus, sp. n., G. auriculus, sp. n., G. castigatus, sp. n., G. clava, sp. n., G. dispar, sp. n., and G. falcis, sp. n. Arrows and stars indicate diagnostic features listed in the species diagnoses." figureDoi="http://doi.org/10.5281/zenodo.4691724" httpUri="https://zenodo.org/record/4691724/files/figure.png" pageId="3" pageNumber="264">Fig. 2</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FCA06F08E39DFB8F" box="[789,918,1087,1113]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="auriculu">
<emphasis id="5AA6EADAFFCCFF92FCA06F08E39DFB8F" box="[789,918,1087,1113]" italics="true" pageId="3" pageNumber="264">G. auriculu</emphasis>
</taxonomicName>
s, 
<emphasis id="5AA6EADAFFCCFF92FC1B6F08E3E7FB8F" bold="true" box="[942,1004,1087,1113]" pageId="3" pageNumber="264">sp. n.</emphasis>
); M beyond Cu short. 
<emphasis id="5AA6EADAFFCCFF92FB536F08E515FB8F" bold="true" box="[1254,1310,1087,1113]" pageId="3" pageNumber="264">Legs</emphasis>
: males and females with tarsal formula 2-2-3, with claws long except short in 
<taxonomicName id="AFD24D4BFFCCFF92FC3E6F54E3E4FBAB" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[907,1007,1123,1149]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC3E6F54E3E4FBAB" box="[907,1007,1123,1149]" italics="true" pageId="3" pageNumber="264">G dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC4B6F54E4BFFBAA" bold="true" box="[1022,1204,1122,1149]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC4B6F54E434FBAB" box="[1022,1087,1123,1149]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
Genitalia
</emphasis>
(
<figureCitation id="F0E92A4DFFCCFF92FB706F54E502FBAB" box="[1221,1289,1123,1149]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
): Pygophore without lobe on right side (
<figureCitation id="F0E92A4DFFCCFF92FE716FB0E203FB77" box="[452,520,1159,1185]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FDA66FB0E2A1FB77" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[531,682,1159,1185]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FDA66FB0E2A1FB77" box="[531,682,1159,1185]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FD026FB0E2FDFB77" bold="true" box="[695,758,1159,1185]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FD026FB0E2FDFB77" box="[695,758,1159,1185]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), with small (
<figureCitation id="F0E92A4DFFCCFF92FC256FB0E3DFFB77" box="[912,980,1159,1185]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC556FB0E430FB76" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[992,1083,1159,1185]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="artus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC556FB0E430FB76" box="[992,1083,1159,1185]" italics="true" pageId="3" pageNumber="264">G. artus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FBFD6FB0E48CFB77" bold="true" box="[1096,1159,1159,1185]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FBFD6FB0E48CFB77" box="[1096,1159,1159,1185]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or large lobe (
<figureCitation id="F0E92A4DFFCCFF92FA876FB0E57EFB77" box="[1330,1397,1159,1185]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FA346FB0E10AFB13" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="auriculus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FA346FB0E10AFB13" italics="true" pageId="3" pageNumber="264">G. auriculus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FEA56F9CE15BFB13" bold="true" box="[272,336,1195,1221]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FEA56F9CE15BFB13" box="[272,336,1195,1221]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), or with slightly extended right margin (
<figureCitation id="F0E92A4DFFCCFF92FC916F9CE362FB13" box="[804,873,1195,1221]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FCC36F9CE400FB12" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[886,1035,1195,1221]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="castigatus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FCC36F9CE400FB12" box="[886,1035,1195,1221]" italics="true" pageId="3" pageNumber="264">G. castigatus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FBAC6F9CE452FB13" bold="true" box="[1049,1113,1195,1221]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FBAC6F9CE452FB13" box="[1049,1113,1195,1221]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Vesica about one half loop (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FF626FF8E110FB3F" box="[215,283,1231,1257]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FE926FF8E1B7FB3E" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[295,444,1231,1257]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="castigatus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FE926FF8E1B7FB3E" box="[295,444,1231,1257]" italics="true" pageId="3" pageNumber="264">G. castigatus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE7C6FF8E202FB3F" bold="true" box="[457,521,1231,1257]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE7C6FF8E202FB3F" box="[457,521,1231,1257]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), between half and one loop (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FC256FF8E3DFFB3F" box="[912,980,1231,1257]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC556FF8E47CFB3F" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[992,1143,1231,1257]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="anaticulus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC556FF8E47CFB3F" box="[992,1143,1231,1257]" italics="true" pageId="3" pageNumber="264">G. anaticulus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FB306FF8E4CEFB3F" bold="true" box="[1157,1221,1231,1257]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FB306FF8E4CEFB3F" box="[1157,1221,1231,1257]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), or one loop (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FF226FC4E0D7FADB" box="[151,220,1267,1293]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FF5F6FC4E18AFADB" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[234,385,1267,1293]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="plurilobus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FF5F6FC4E18AFADB" box="[234,385,1267,1293]" italics="true" pageId="3" pageNumber="264">G. plurilobus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE256FC4E1DAFADB" bold="true" box="[400,465,1267,1293]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE256FC4E1DAFADB" box="[400,465,1267,1293]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), either not reaching right pygophore margin or surpassing margin, at midpoint thick (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FF626E20E110FAE7" box="[215,283,1303,1329]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FE926E20E196FAE6" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[295,413,1303,1329]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="graziae" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FE926E20E196FAE6" box="[295,413,1303,1329]" italics="true" pageId="3" pageNumber="264">G. graziae</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FE1E6E20E1E0FAE7" bold="true" box="[427,491,1303,1329]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FE1E6E20E1E0FAE7" box="[427,491,1303,1329]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), very thick (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FD0F6E20E2F6FAE7" box="[698,765,1303,1329]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FCBC6E20E381FAE7" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[777,906,1303,1329]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="robustus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FCBC6E20E381FAE7" box="[777,906,1303,1329]" italics="true" pageId="3" pageNumber="264">G. robustus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC2C6E20E3D2FAE7" bold="true" box="[921,985,1303,1329]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC2C6E20E3D2FAE7" box="[921,985,1303,1329]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), or more slender (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FB526E20E521FAE7" box="[1255,1322,1303,1329]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FA836E20E59DFAE7" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1334,1430,1303,1329]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="falcis" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FA836E20E59DFAE7" box="[1334,1430,1303,1329]" italics="true" pageId="3" pageNumber="264">G. falcis</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FF226E0CE0D3FA83" bold="true" box="[151,216,1339,1365]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FF226E0CE0D3FA83" box="[151,216,1339,1365]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), tapering to blunt or acute tip, tip itself blunt, bottle-opener shaped (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FB9C6E0CE466FA83" box="[1065,1133,1339,1365]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FBCE6E0CE4D4FA82" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1147,1247,1339,1365]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="uncus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FBCE6E0CE4D4FA82" box="[1147,1247,1339,1365]" italics="true" pageId="3" pageNumber="264">G. uncus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FB5A6E0CE53BFA83" bold="true" box="[1263,1328,1339,1365]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FB5A6E0CE53BFA83" box="[1263,1328,1339,1365]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), slightly s-shaped (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FEF16E68E181FAAF" box="[324,394,1375,1401]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FE2D6E68E1F2FAAF" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[408,505,1375,1401]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="falcis" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FE2D6E68E1F2FAAF" box="[408,505,1375,1401]" italics="true" pageId="3" pageNumber="264">G. falcis</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FDBC6E68E240FAAF" bold="true" box="[521,587,1375,1401]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FDBC6E68E240FAAF" box="[521,587,1375,1401]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), rounded (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FCBE6E68E35AFAAF" box="[779,849,1375,1401]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FCEA6E68E3B6FAAE" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[863,957,1375,1401]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="artus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FCEA6E68E3B6FAAE" box="[863,957,1375,1401]" italics="true" pageId="3" pageNumber="264">G. artus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC786E68E404FAAF" bold="true" box="[973,1039,1375,1401]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC786E68E404FAAF" box="[973,1039,1375,1401]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), or fairly straight (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FA986E68E578FAAF" box="[1325,1395,1375,1401]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FA346E68E0F0FA4B" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="tatumbia" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FA346E68E0F0FA4B" italics="true" pageId="3" pageNumber="264">G. tatumbia</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FEBD6EB4E14DFA4B" bold="true" box="[264,326,1411,1437]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FEBD6EB4E14DFA4B" box="[264,326,1411,1437]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Anophoric ridge with process present (e.g., 
<figureCitation id="F0E92A4DFFCCFF92FC8A6EB4E38AFA4B" box="[831,897,1411,1437]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FC396EB4E3ECFA4A" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[908,999,1411,1437]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="artus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FC396EB4E3ECFA4A" box="[908,999,1411,1437]" italics="true" pageId="3" pageNumber="264">G. artus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FC416EB4E439FA4B" bold="true" box="[1012,1074,1411,1437]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FC416EB4E439FA4B" box="[1012,1074,1411,1437]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
) or absent (
<figureCitation id="F0E92A4DFFCCFF92FB016EB4E4F3FA4B" box="[1204,1272,1411,1437]" captionStart="FIGURE 7" captionStartId="23.[151,250,1943,1967]" captionTargetBox="[219,1368,193,1906]" captionTargetId="figure-17@23.[205,1383,181,1919]" captionTargetPageId="23" captionText="FIGURE 7. Male genitalic features of Guapinannus spp., dorsal view, showing species-diagnostic differences in vesica, right and left parameres, and anophoric process." figureDoi="http://doi.org/10.5281/zenodo.4691739" httpUri="https://zenodo.org/record/4691739/files/figure.png" pageId="3" pageNumber="264">Fig. 7</figureCitation>
, 
<taxonomicName id="AFD24D4BFFCCFF92FAB76EB4E59DFA4A" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[1282,1430,1411,1437]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="species" species="castigatus" status="sp. nov.">
<emphasis id="5AA6EADAFFCCFF92FAB76EB4E59DFA4A" box="[1282,1430,1411,1437]" italics="true" pageId="3" pageNumber="264">G. castigatus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCCFF92FF226E90E0DDFA17" bold="true" box="[151,214,1447,1473]" pageId="3" pageNumber="264">
<taxonomicNameLabel id="419557A1FFCCFF92FF226E90E0DDFA17" box="[151,214,1447,1473]" pageId="3" pageNumber="264" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
), process short or long, with single lobe curved or relatively straight, or with multiple lobes, tip pointed, blunt, or combination of pointed and blunt. Right paramere curved or straight, tapering towards apex or broad throughout; basal process of paramere broad, narrow, thumb-shaped, or duck head-shaped. Left paramere straight or curved, tapering towards apex or broad throughout, basal process broad or narrow.
</paragraph>
<paragraph id="686D36C8FFCCFF92FF726D00E208F94F" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="264">
<emphasis id="5AA6EADAFFCCFF92FF726D00E123F987" bold="true" box="[199,296,1591,1617]" pageId="3" pageNumber="264">Female:</emphasis>
As male, but with less well developed An1/2 claval process and without An1 wing organ on corium (
<figureCitation id="F0E92A4DFFCCFF92FF156D6CE0F3F9A3" box="[160,248,1627,1653]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Fig. 1B</figureCitation>
). 
<emphasis id="5AA6EADAFFCCFF92FEBA6D6DE18CF9A2" bold="true" box="[271,391,1626,1652]" pageId="3" pageNumber="264">Genitalia:</emphasis>
with well-developed ovipositor (
<figureCitation id="F0E92A4DFFCCFF92FCB76D6CE355F9A3" box="[770,862,1627,1653]" captionStart="FIGURE 1" captionStartId="4.[151,250,1955,1979]" captionTargetBox="[152,1434,851,1929]" captionTargetId="figure-369@4.[151,1436,849,1931]" captionTargetPageId="4" captionText="FIGURE 1. Guapinannus bierigi, female holotype. A: Head with rostrum extended forward and pronotum showing muscle scars on anterior lobe, dorsal view. B: Left forewing, dorsal view. C-D: Abdomen dorsum and ventrum, respectively." figureDoi="http://doi.org/10.5281/zenodo.4691722" httpUri="https://zenodo.org/record/4691722/files/figure.png" pageId="3" pageNumber="264">Figs 1C</figureCitation>
, 
<figureCitation id="F0E92A4DFFCCFF92FCD86D6CE376F9A3" box="[877,893,1627,1653]" captionStart="FIGURE 8" captionStartId="24.[151,250,1053,1077]" captionTargetBox="[224,1363,181,1028]" captionTargetId="figure-24@24.[224,1363,181,1028]" captionTargetPageId="24" captionText="FIGURE 8. Female Guapinannus sp. (UCR_ENT 00101286), genitalic structures including ovipositor and spermathecal gland, and spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.4691741" httpUri="https://zenodo.org/record/4691741/files/figure.png" pageId="3" pageNumber="264">8</figureCitation>
). Reservoir of spermatheca kidney-shaped with short spermathecal duct (
<figureCitation id="F0E92A4DFFCCFF92FE056D48E1FEF94F" box="[432,501,1663,1689]" captionStart="FIGURE 8" captionStartId="24.[151,250,1053,1077]" captionTargetBox="[224,1363,181,1028]" captionTargetId="figure-24@24.[224,1363,181,1028]" captionTargetPageId="24" captionText="FIGURE 8. Female Guapinannus sp. (UCR_ENT 00101286), genitalic structures including ovipositor and spermathecal gland, and spermathecal duct." figureDoi="http://doi.org/10.5281/zenodo.4691741" httpUri="https://zenodo.org/record/4691741/files/figure.png" pageId="3" pageNumber="264">Fig. 8</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="20C86543FFCCFF92FF726D95E21AF86F" pageId="3" pageNumber="264" type="distribution">
<paragraph id="686D36C8FFCCFF92FF726D95E172F89B" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="264">
<emphasis id="5AA6EADAFFCCFF92FF726D95E16AF96A" bold="true" box="[199,353,1698,1724]" pageId="3" pageNumber="264">Distribution.</emphasis>
Species of 
<taxonomicName id="AFD24D4BFFCCFF92FE686D94E27FF96B" authorityName="Wygodzinsky" authorityYear="1951" box="[477,628,1699,1725]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCCFF92FE686D94E27FF96B" box="[477,628,1699,1725]" italics="true" pageId="3" pageNumber="264">Guapinannus</emphasis>
</taxonomicName>
are documented from Central and South America, ranging from 
<collectingRegion id="AA16F82AFFCCFF92FA8C6D94E597F96B" box="[1337,1436,1699,1725]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="264">Veracruz</collectingRegion>
in 
<collectingCountry id="10C57658FFCCFF92FF066DF0E102F937" box="[179,265,1735,1761]" name="Mexico" pageId="3" pageNumber="264">Mexico</collectingCountry>
to the southern part of 
<collectingCountry id="10C57658FFCCFF92FDBE6DF0E258F937" box="[523,595,1735,1761]" name="Brazil" pageId="3" pageNumber="264">Brazil</collectingCountry>
. The greatest species diversity is found in Central America and the northern areas of South America. Specimens have been collected in tropical wet or moist forest, including cloud forest and mixed hardwood forest, from elevation ranging from 5 meters above sea level in 
<collectingCountry id="10C57658FFCCFF92FB9F6C38E488F8FF" box="[1066,1155,1807,1833]" name="Panama" pageId="3" pageNumber="264">Panama</collectingCountry>
to more than 200 meters in 
<collectingRegion id="AA16F82AFFCCFF92FF016C04E11AF89B" box="[180,273,1843,1869]" country="Mexico" name="Chiapas" pageId="3" pageNumber="264">Chiapas</collectingRegion>
, 
<collectingCountry id="10C57658FFCCFF92FEA96C04E17EF89B" box="[284,373,1843,1869]" name="Mexico" pageId="3" pageNumber="264">Mexico</collectingCountry>
.
</paragraph>
<paragraph id="686D36C8FFCCFF92FF726C61E21AF86F" blockId="3.[151,1437,150,2013]" pageId="3" pageNumber="264">
<emphasis id="5AA6EADAFFCCFF92FF726C61E1A3F8A7" bold="true" box="[199,424,1878,1905]" pageId="3" pageNumber="264">Collecting method.</emphasis>
The majority of specimens for which collection method was recorded were collected using leaf litter sifting and Berlese and Winkler extraction methods. A small number of specimens are derived from Malaise and flight intercept traps.
</paragraph>
</subSubSection>
<subSubSection id="20C86543FFCCFF95FF726CF5E2A1FCEF" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264" type="discussion">
<paragraph id="686D36C8FFCCFF95FF726CF5E4EBFDCF" blockId="3.[151,1437,150,2013]" lastBlockId="4.[151,1437,151,825]" lastPageId="4" lastPageNumber="265" pageId="3" pageNumber="264">
<emphasis id="5AA6EADAFFCCFF92FF726CF5E147F80A" bold="true" box="[199,332,1986,2012]" pageId="3" pageNumber="264">Discussion.</emphasis>
The forewing shape, structure with distinct claval suture, and venation set apart 
<taxonomicName id="AFD24D4BFFCCFF92FB7C6CF4E56BF80B" authorityName="Wygodzinsky" authorityYear="1951" box="[1225,1376,1987,2013]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCCFF92FB7C6CF4E56BF80B" box="[1225,1376,1987,2013]" italics="true" pageId="3" pageNumber="264">Guapinannus</emphasis>
</taxonomicName>
from all other genera of 
<taxonomicName id="AFD24D4BFFCBFF95FED16BA0E205FF67" authorityName="Reuter" authorityYear="1891" box="[356,526,151,177]" class="Insecta" family="Schizopteridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="family">Schizopteridae</taxonomicName>
. While 
<taxonomicName id="AFD24D4BFFCBFF95FDD56BA0E312FF66" authorityName="Wygodzinsky" authorityYear="1950" box="[608,793,151,176]" class="Insecta" family="Dipsocoridae" genus="Humpatanannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FDD56BA0E312FF66" box="[608,793,151,176]" italics="true" pageId="4" pageNumber="265">Humpatanannus</emphasis>
</taxonomicName>
and related genera have not been included in phylogenetic analyses, genitalic features are strikingly different in the two groups, and they are unlikely to be close relatives. The non-genitalic morphology of species of 
<taxonomicName id="AFD24D4BFFCBFF95FD386BE8E32FFF2F" authorityName="Wygodzinsky" authorityYear="1951" box="[653,804,223,249]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FD386BE8E32FFF2F" box="[653,804,223,249]" italics="true" pageId="4" pageNumber="265">Guapinannus</emphasis>
</taxonomicName>
is fairly uniform. Most species are mid-sized and ovoid with some variation, with only few species being distinctly smaller or bigger or markedly elongated ovoid. Some forewing venation features are fairly distinctive and useful as diagnostic characters at the species level, among them the width of the C+Sc and the relative proportion of certain cells. As in many other genera of 
<taxonomicName id="AFD24D4BFFCBFF95FB716A7CE566FEB3" authorityName="Reuter" authorityYear="1891" box="[1220,1389,331,357]" class="Insecta" family="Schizopteridae" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="family">Schizopteridae</taxonomicName>
, the bulk of diagnostic features is derived from male genitalic structures, in particular the length, curvature, and tip shape of the vesica, shape of the anophoric process, where present, and shape of the pygophore that is expanded into a distinct lobe on the right side in some species. We described as new two species for which only female specimens are known (
<taxonomicName id="AFD24D4BFFCBFF95FEAE6AECE18EFE23" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[283,389,475,501]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCBFF95FEAE6AECE18EFE23" box="[283,389,475,501]" italics="true" pageId="4" pageNumber="265">G. dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCBFF95FE276AECE1DBFE23" bold="true" box="[402,464,475,501]" pageId="4" pageNumber="265">
<taxonomicNameLabel id="419557A1FFCBFF95FE276AECE1DBFE23" box="[402,464,475,501]" pageId="4" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="AFD24D4BFFCBFF95FDB06AECE274FE22" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" box="[517,639,475,501]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="minutus" status="sp. nov.">
<emphasis id="5AA6EADAFFCBFF95FDB06AECE274FE22" box="[517,639,475,501]" italics="true" pageId="4" pageNumber="265">G. minutus</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCBFF95FD396AECE2C0FE23" bold="true" box="[652,715,475,501]" pageId="4" pageNumber="265">
<taxonomicNameLabel id="419557A1FFCBFF95FD396AECE2C0FE23" box="[652,715,475,501]" pageId="4" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
). Both are distinctive based on non-genitalic morphology and 
<taxonomicName id="AFD24D4BFFCBFF95FA346AECE0D4FDCF" authority="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch, 2021" authorityName="Frankenberg &amp; Knyshov &amp; Hoey-Chamberlain &amp; Weirauch" authorityYear="2021" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="species" species="dispar" status="sp. nov.">
<emphasis id="5AA6EADAFFCBFF95FA346AECE0D4FDCF" italics="true" pageId="4" pageNumber="265">G. dispar</emphasis>
</taxonomicName>
, 
<emphasis id="5AA6EADAFFCBFF95FF586AC8E127FDCF" bold="true" box="[237,300,511,537]" pageId="4" pageNumber="265">
<taxonomicNameLabel id="419557A1FFCBFF95FF586AC8E127FDCF" box="[237,300,511,537]" pageId="4" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
in addition is geographically isolated from all other known species of 
<taxonomicName id="AFD24D4BFFCBFF95FBF76AC8E4D2FDCF" authorityName="Wygodzinsky" authorityYear="1951" box="[1090,1241,511,537]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FBF76AC8E4D2FDCF" box="[1090,1241,511,537]" italics="true" pageId="4" pageNumber="265">Guapinannus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="686D36C8FFCBFF95FF726914E2A1FCEF" blockId="4.[151,1437,151,825]" pageId="4" pageNumber="265">
As part of this study we discovered a so far undocumented structure on the forewings that is restricted to males that we interpret as a novel male-specific wing organ. This wing organ is located on the posterior margin of the corium, associated with An1, consist of a strongly inflated semicircular or elongate area and in most species features a median notch. SEM observation of this wing organ in one species (
<figureCitation id="F0E92A4DFFCBFF95FC2669B8E3E1FD7C" box="[915,1002,655,682]" captionStart="FIGURE 6" captionStartId="21.[151,250,1490,1514]" captionTargetBox="[153,1435,183,1464]" captionTargetId="figure-17@21.[151,1435,181,1465]" captionTargetPageId="21" captionText="FIGURE 6. Scanning electron micrographs of wings and genitalic structures of male of Guapinannus clava, sp. n. (UCR_ENT 00090715). A: Right forewing, dorsal view. B: Right hindwing, dorsal view. C: Right forewing, showing vein tracing areoles, dorsal view. D: Right forewing, showing An1/2 claval process; pronounced An1 wing organ on corium, dorsal view. E: Left forewing, showing wing coupling, dorsal view. F: Pygophore and genitalic structures, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4691736" httpUri="https://zenodo.org/record/4691736/files/figure.png" pageId="4" pageNumber="265">Fig. 6D</figureCitation>
) shows a tuft of hairlike structures that may represent a trichome. Based on our observations, males of all species of 
<taxonomicName id="AFD24D4BFFCBFF95FC446984E483FD1B" authorityName="Wygodzinsky" authorityYear="1951" box="[1009,1160,691,717]" class="Insecta" family="Dipsocoridae" genus="Guapinannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FC446984E483FD1B" box="[1009,1160,691,717]" italics="true" pageId="4" pageNumber="265">Guapinannus</emphasis>
</taxonomicName>
possess this wing organ. Among closely related taxa (based on 
<bibRefCitation id="0C434B39FFCBFF95FDFF69E0E330FD27" author="Knyshov, A. &amp; Weirauch, C. &amp; Hoey-Chamberlain, R." box="[586,827,727,753]" pageId="4" pageNumber="265" refId="ref19634" refString="Knyshov, A., Weirauch, C. &amp; Hoey-Chamberlain, R. (2020) Phylogenetic relationships and revised classification of the true bug infraorder Dipsocoromorpha (Insecta: Hemiptera: Heteroptera), Cladistics. [early view] https: // doi. org / 10.1111 / cla. 12435" type="book" year="2020">
Knyshov 
<emphasis id="5AA6EADAFFCBFF95FD0269E0E2E2FD27" box="[695,745,727,753]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
., 2020
</bibRefCitation>
), 
<taxonomicName id="AFD24D4BFFCBFF95FCFA69E0E3E2FD27" authorityName="Weirauch, Knyshov, &amp; Hoey-Chamberlain" authorityYear="2020" box="[847,1001,727,753]" class="Insecta" family="Schizopteridae" genus="Caucanannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FCFA69E0E3E2FD27" box="[847,1001,727,753]" italics="true" pageId="4" pageNumber="265">Caucanannus</emphasis>
</taxonomicName>
also have a male-specific wing organ, although in a different position on the forewing (
<bibRefCitation id="0C434B39FFCBFF95FD7269CCE3C8FCC3" author="Weirauch, C &amp; Hoey-Chamberlain, R. &amp; Knyshov, A." box="[711,963,763,789]" pageId="4" pageNumber="265" pagination="95 - 111" refId="ref19862" refString="Weirauch, C, Hoey-Chamberlain, R. &amp; Knyshov, A. (2020) Four new genera of Schizopteridae (Hemiptera: Heteroptera) from the Afrotropical and Neotropical regions. Zootaxa, 4768 (1), 95 - 111. https: // doi. org / 10.11646 / zootaxa. 4768.1.6" type="journal article" year="2020">
Weirauch 
<emphasis id="5AA6EADAFFCBFF95FC8869CCE37BFCC3" box="[829,880,763,789]" italics="true" pageId="4" pageNumber="265">et al</emphasis>
., 2020
</bibRefCitation>
), while wing organs have so far not been documented in 
<taxonomicName id="AFD24D4BFFCBFF95FEF16828E20EFCEF" authorityName="Wygodzinsky" authorityYear="1950" box="[324,517,799,825]" class="Insecta" family="Dipsocoridae" genus="Luachimonannus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FEF16828E20EFCEF" box="[324,517,799,825]" italics="true" pageId="4" pageNumber="265">Luachimonannus</emphasis>
</taxonomicName>
and 
<taxonomicName id="AFD24D4BFFCBFF95FD8E6828E2A8FCEF" authorityName="Miyamoto" authorityYear="1960" box="[571,675,799,825]" class="Insecta" family="Schizopteridae" genus="Kokeshia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="265" phylum="Arthropoda" rank="genus">
<emphasis id="5AA6EADAFFCBFF95FD8E6828E2A8FCEF" box="[571,675,799,825]" italics="true" pageId="4" pageNumber="265">Kokeshia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>