treatments-xml/data/66/34/87/663487D6FFA8FFD5FCB6FBCDFC39CBD2.xml
2024-06-21 12:38:50 +02:00

235 lines
20 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document ID-DOI="http://doi.org/10.5281/zenodo.6492996" ID-ISSN="1982-0224" ID-Zenodo-Dep="6492996" approvalRequired="1" approvalRequired_for_originalDoi="1" checkinTime="1630380785404" checkinUser="carolina" docAuthor="A, Pablo Lehmann, Mayer, Fernanda &amp; Reis, Roberto E." docDate="2010" docId="663487D6FFA8FFD5FCB6FBCDFC39CBD2" docLanguage="en" docName="NeotropIchthyol.2010.7.pdf" docOrigin="Neotropical Ichthyology 8 (1)" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Otocinclus mimulus Axenrot &amp; Kullander 2003" docType="treatment" docVersion="2" lastPageId="4" lastPageNumber="61" masterDocId="9A0DFFAEFFABFFD1FFD5FFFDFFE9CC56" masterDocTitle="Re-validation of Otocinclus arnoldi Regan and reappraisal of Otocinclus phylogeny (Siluriformes: Loricariidae)" masterLastPageNumber="68" masterPageNumber="57" pageId="3" pageNumber="60" updateTime="1650995928801" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Re-validation of Otocinclus arnoldi Regan and reappraisal of Otocinclus phylogeny (Siluriformes: Loricariidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>A, Pablo Lehmann</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mayer, Fernanda</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reis, Roberto E.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="volume">
<mods:number>8</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>57</mods:start>
<mods:end>68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6492996" ID-Zenodo-Dep="6492996" LSID="urn:lsid:plazi:treatment:663487D6FFA8FFD5FCB6FBCDFC39CBD2" httpUri="http://treatment.plazi.org/id/663487D6FFA8FFD5FCB6FBCDFC39CBD2" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
<subSubSection pageId="3" pageNumber="60" type="nomenclature">
<paragraph blockId="3.[867,1393,1072,1127]" pageId="3" pageNumber="60">
<emphasis bold="true" pageId="3" pageNumber="60">
<taxonomicName authority="Axenrot &amp; Kullander, 2003" authorityName="Axenrot &amp; Kullander" authorityYear="2003" box="[867,1393,1072,1095]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
<emphasis bold="true" box="[867,1086,1072,1095]" italics="true" pageId="3" pageNumber="60">Otocinclus mimulus</emphasis>
Axenrot &amp; Kullander, 2003
</taxonomicName>
<figureCitation box="[1099,1161,1104,1127]" captionStart="Fig" captionStartId="7.[129,170,1761,1784]" captionTargetBox="[166,1413,174,1750]" captionTargetId="figure-102@7.[166,1413,174,1750]" captionTargetPageId="7" captionText="Fig. 4. Color variation of paratypes Otocinclus mimulus. All from NRM 42332, (a) SL = 34.4, (b) 31.2, (c) 35.1, and (d) 34.4 mm." pageId="3" pageNumber="60">Fig. 4</figureCitation>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="60" type="description">
<paragraph blockId="3.[809,1452,1168,1351]" pageId="3" pageNumber="60">
<taxonomicName authority="Axenrot &amp; Kullander, 2003: 251" authorityName="Axenrot &amp; Kullander" authorityPageNumber="251" authorityYear="2003" box="[809,1388,1168,1191]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
<emphasis box="[809,1024,1168,1191]" italics="true" pageId="3" pageNumber="60">Otocinclus mimulus</emphasis>
<materialsCitation accessionNumber="NRM 43480" box="[1031,1388,1168,1191]" collectionCode="MNHNP, NRM" country="Paraguay" county="rio Parana" latitude="-25.670277" location="Colonel Patricio Colman" longLatPrecision="21" longitude="-55.09778" municipality="Estancia Maria Belen" pageId="3" pageNumber="60" specimenCount="2" stateProvince="Caaguazu" typeStatus="holotype">Axenrot &amp; Kullander, 2003: 251</materialsCitation>
</taxonomicName>
. Type locality:
<collectingCountry box="[937,1042,1200,1223]" name="Paraguay" pageId="3" pageNumber="60">Paraguay</collectingCountry>
:
<collectingRegion box="[1054,1327,1200,1223]" country="Paraguay" name="Caaguazu" pageId="3" pageNumber="60">Departamento Caaguazú</collectingRegion>
:
<collectingCounty box="[1339,1452,1200,1223]" pageId="3" pageNumber="60">río Paraná</collectingCounty>
drainage, small stream (arroyo) at
<collectingMunicipality box="[1199,1429,1232,1255]" pageId="3" pageNumber="60">Estancia María Belén</collectingMunicipality>
,
<quantity metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="60" unit="km" value="8.0">8 km</quantity>
from
<location LSID="urn:lsid:plazi:treatment:663487D6FFA8FFD5FCB6FBCDFC39CBD2:EB42601BFFA8FFD2FC70FB0DFB4EC951" box="[933,1191,1264,1287]" country="Paraguay" county="rio Parana" latitude="-25.670277" longLatPrecision="21" longitude="-55.09778" municipality="Estancia Maria Belen" name="Colonel Patricio Colman" pageId="3" pageNumber="60" stateProvince="Caaguazu">Colonel Patricio Colman</location>
;
<geoCoordinate box="[1199,1322,1264,1287]" degrees="25" direction="south" minutes="40" orientation="latitude" pageId="3" pageNumber="60" precision="15" seconds="13" value="-25.670277">25°4013”S</geoCoordinate>
<geoCoordinate box="[1327,1447,1264,1287]" degrees="55" direction="west" minutes="5" orientation="longitude" pageId="3" pageNumber="60" precision="15" seconds="52" value="-55.09778">55°552”W</geoCoordinate>
.
<typeStatus box="[839,944,1296,1319]" pageId="3" pageNumber="60">Holotype</typeStatus>
: MNHNP uncatalogued (ex
<accessionNumber box="[1247,1387,1296,1319]" httpUri="http://www.ncbi.nlm.nih.gov/protein/NRM43480" pageId="3" pageNumber="60">NRM 43480</accessionNumber>
),
<quantity metricMagnitude="-2" metricUnit="m" metricValue="3.41" pageId="3" pageNumber="60" unit="mm" value="34.1">34.1 mm</quantity>
SL [
<typeStatus box="[928,1030,1328,1351]" pageId="3" pageNumber="60">Holotype</typeStatus>
originally cited as MNHNP uncat.].
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60" type="diagnosis">
<paragraph blockId="3.[809,1452,1392,1959]" lastBlockId="4.[136,779,1837,1956]" lastPageId="4" lastPageNumber="61" pageId="3" pageNumber="60">
<emphasis bold="true" box="[809,925,1392,1415]" pageId="3" pageNumber="60">Diagnosis.</emphasis>
<taxonomicName authorityName="Axenrot &amp; Kullander" authorityYear="2003" box="[929,1138,1392,1415]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="mimulus">
<emphasis box="[929,1138,1392,1415]" italics="true" pageId="3" pageNumber="60">Otocinclus mimulus</emphasis>
</taxonomicName>
is distinguished from all other
<taxonomicName box="[809,1004,1424,1447]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[809,923,1424,1447]" italics="true" pageId="3" pageNumber="60">Otocinclus</emphasis>
species
</taxonomicName>
except
<taxonomicName authorityName="Regan" authorityYear="1909" box="[1082,1190,1424,1447]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
<emphasis box="[1082,1101,1424,1447]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1113,1190,1424,1447]" italics="true" pageId="3" pageNumber="60">arnoldi</emphasis>
</taxonomicName>
, by having five branched rays in the pectoral-fin (
<emphasis box="[1074,1096,1456,1479]" italics="true" pageId="3" pageNumber="60">vs</emphasis>
. six branched pectoral-fin rays), and except from
<taxonomicName authorityName="Regan" authorityYear="1909" box="[995,1108,1488,1511]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
<emphasis box="[995,1014,1488,1511]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1029,1108,1488,1511]" italics="true" pageId="3" pageNumber="60">arnoldi</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[1164,1294,1488,1511]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
<emphasis box="[1164,1183,1488,1511]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1198,1294,1488,1511]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
</taxonomicName>
by having the prootic involved in the contact with the hyomandibular articular condyle (
<emphasis box="[1005,1027,1552,1575]" italics="true" pageId="3" pageNumber="60">vs</emphasis>
. only compound pterotic contacting the hyomandibular articular condyle). It is distinguished from
<taxonomicName authorityName="Regan" authorityYear="1909" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
<emphasis box="[1426,1445,1584,1607]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[809,888,1616,1639]" italics="true" pageId="3" pageNumber="60">arnoldi</emphasis>
</taxonomicName>
by having an elevated patch of enlarged odontodes on the postero-dorsal parieto-supraoccipital tip and typically having 16-17 caudal vertebrae (see
<tableCitation box="[1191,1273,1680,1703]" captionStart="Table 2" captionStartId="5.[129,192,178,201]" captionText="Table 2. Frequency distribution of variable counts of Otocinclus species. Caudal vertebrae counted in clear and stained specimens only. *Five specimens counted and 13 from radiographs by S.O. Kulander; **Data from radiographs by S. O. Kulander." pageId="3" pageNumber="60">Table 2</tableCitation>
;
<emphasis box="[1284,1306,1680,1703]" italics="true" pageId="3" pageNumber="60">vs</emphasis>
. having a not elevated patch of enlarged odontodes on the parietosupraoccipital and having 15 caudal vertebrae). It is further distinguished (except for
<taxonomicName authorityName="Steindachner" authorityYear="1877" box="[1071,1168,1776,1799]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="affinis">
<emphasis box="[1071,1090,1776,1799]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1102,1168,1776,1799]" italics="true" pageId="3" pageNumber="60">affinis</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Steindachner" authorityYear="1915" box="[1179,1310,1776,1799]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="hasemani">
<emphasis box="[1179,1198,1776,1799]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1209,1310,1776,1799]" italics="true" pageId="3" pageNumber="60">hasemani</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[1321,1444,1776,1799]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
<emphasis box="[1321,1340,1776,1799]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1351,1444,1776,1799]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Regan" authorityYear="1909" box="[809,921,1808,1831]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
<emphasis box="[809,828,1808,1831]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[842,921,1808,1831]" italics="true" pageId="3" pageNumber="60">arnoldi</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Cope" authorityYear="1894" box="[980,1085,1808,1831]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="flexilis">
<emphasis box="[980,999,1808,1831]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1013,1085,1808,1831]" italics="true" pageId="3" pageNumber="60">flexilis</emphasis>
</taxonomicName>
) by possessing an iris operculum, and (except for
<taxonomicName authorityName="Steindachner" authorityYear="1877" box="[964,1056,1840,1863]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="affinis">
<emphasis box="[964,983,1840,1863]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[993,1056,1840,1863]" italics="true" pageId="3" pageNumber="60">affinis</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[1102,1221,1840,1863]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
<emphasis box="[1102,1121,1840,1863]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1131,1221,1840,1863]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
</taxonomicName>
) by having an elevated patch of enlarged odontodes on the postero-dorsal parietosupraoccipital tip. It is also distinguished from all remaining
<taxonomicName box="[809,1004,1936,1959]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="undetermined">
<emphasis box="[809,924,1936,1959]" italics="true" pageId="3" pageNumber="60">Otocinclus</emphasis>
species
</taxonomicName>
(except for
<taxonomicName authorityName="Schaefer" authorityYear="1997" box="[1127,1252,1936,1959]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="xakriaba">
<emphasis box="[1127,1146,1936,1959]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1158,1252,1936,1959]" italics="true" pageId="3" pageNumber="60">xakriaba</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Regan" authorityYear="1909" box="[1263,1372,1936,1959]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="arnoldi">
<emphasis box="[1263,1282,1936,1959]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[1294,1372,1936,1959]" italics="true" pageId="3" pageNumber="60">arnoldi</emphasis>
</taxonomicName>
, and
<taxonomicName authorityName="Cope" authorityYear="1894" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" lastPageId="4" lastPageNumber="61" order="Siluriformes" pageId="3" pageNumber="60" phylum="Chordata" rank="species" species="flexilis">
<emphasis box="[1426,1445,1936,1959]" italics="true" pageId="3" pageNumber="60">O</emphasis>
.
<emphasis box="[136,205,1837,1860]" italics="true" pageId="4" pageNumber="61">flexilis</emphasis>
</taxonomicName>
) by having a lateral trunk coloration composed of either a row of 3-6 distinct dark blotches or a distinct dark stripe extending from the compound pterotic to the base of the caudal fin, or a diffuse mixture of those two color patterns (
<figureCitation box="[673,737,1933,1956]" captionStart="Fig" captionStartId="7.[129,170,1761,1784]" captionTargetBox="[166,1413,174,1750]" captionTargetId="figure-102@7.[166,1413,174,1750]" captionTargetPageId="7" captionText="Fig. 4. Color variation of paratypes Otocinclus mimulus. All from NRM 42332, (a) SL = 34.4, (b) 31.2, (c) 35.1, and (d) 34.4 mm." pageId="4" pageNumber="61">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption pageId="4" pageNumber="61" startId="4.[136,174,1757,1780]" targetBox="[174,1421,174,1742]" targetPageId="4">
<paragraph blockId="4.[136,1459,1757,1812]" pageId="4" pageNumber="61">
<emphasis bold="true" box="[136,202,1757,1780]" pageId="4" pageNumber="61">Fig. 2.</emphasis>
Color variation of
<taxonomicName authorityName="Cope" authorityYear="1894" box="[391,575,1757,1780]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="flexilis">
<emphasis box="[391,575,1757,1780]" italics="true" pageId="4" pageNumber="61">Otocinclus flexilis</emphasis>
</taxonomicName>
.
<emphasis bold="true" box="[586,616,1757,1780]" pageId="4" pageNumber="61">(a)</emphasis>
and
<emphasis bold="true" box="[663,694,1757,1780]" pageId="4" pageNumber="61">(b)</emphasis>
: MCP 17414, SL = 37.6 and 44.6 mm respectively.
<emphasis bold="true" box="[1211,1240,1757,1780]" pageId="4" pageNumber="61">(c)</emphasis>
and
<emphasis bold="true" box="[1286,1318,1757,1780]" pageId="4" pageNumber="61">(d)</emphasis>
: MCP 18307, SL = 37.2 and 36.8 mm respectively.
</paragraph>
</caption>
<subSubSection pageId="4" pageNumber="61" type="distribution">
<paragraph blockId="4.[816,1459,1837,1924]" pageId="4" pageNumber="61">
<emphasis bold="true" box="[816,963,1837,1860]" pageId="4" pageNumber="61">Distribution.</emphasis>
<taxonomicName authorityName="Axenrot &amp; Kullander" authorityYear="2003" box="[967,1178,1837,1860]" class="Actinopterygii" family="Loricariidae" genus="Otocinclus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="61" phylum="Chordata" rank="species" species="mimulus">
<emphasis box="[967,1178,1837,1860]" italics="true" pageId="4" pageNumber="61">Otocinclus mimulus</emphasis>
</taxonomicName>
is only known from the río Mondai in
<collectingCountry box="[940,1042,1869,1892]" name="Paraguay" pageId="4" pageNumber="61">Paraguay</collectingCountry>
, a left-bank tributary of the lower rio Paraná (
<figureCitation box="[901,963,1901,1924]" captionStart="Fig" captionStartId="8.[136,174,752,775]" captionTargetBox="[156,753,185,739]" captionTargetId="figure-1800@8.[144,771,174,745]" captionTargetPageId="8" captionText="Fig. 5. Geographic distribution of Otocinclus flexilis (triangles) and O. arnoldi (dots). Open dot represents paratypes of O. mimulus. One symbol may cover more than one lot or locality." pageId="4" pageNumber="61">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>