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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1056.66553" ID-GBIF-Dataset="6f05d14c-7490-40a6-b34a-8e4bd3431df6" ID-PMC="PMC8384827" ID-Pensoft-Pub="1313-2970-1056-17" ID-Pensoft-UUID="9F460C6A018E5D5B889A40EAE1F34989" ID-PubMed="34497473" ID-ZooBank="B62820E7CE524501802CD8E864C7199A" ModsDocID="1313-2970-1056-17" checkinTime="1629294920043" checkinUser="pensoft" docAuthor="Liu, Zhiping &amp; Cuccodoro, Giulio" docDate="2021" docId="602F4690185057AFB7C03D6DE2D14DA1" docLanguage="en" docName="ZooKeys 1056: 17-34" docOrigin="ZooKeys 1056" docPubDate="2021-08-17" docSource="http://dx.doi.org/10.3897/zookeys.1056.66553" docTitle="Megarthrus hemipterus" docType="treatment" docVersion="4" id="9F460C6A018E5D5B889A40EAE1F34989" lastPageNumber="17" masterDocId="9F460C6A018E5D5B889A40EAE1F34989" masterDocTitle="Megarthrus of China. Part 4. The M. hemipterus complex (Coleoptera, Staphylinidae, Proteininae), with description of a new species from Yunnan Province" masterLastPageNumber="34" masterPageNumber="17" pageNumber="17" updateTime="1668150849830" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Megarthrus of China. Part 4. The M. hemipterus complex (Coleoptera, Staphylinidae, Proteininae), with description of a new species from Yunnan Province</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Liu, Zhiping</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3308-3274</mods:nameIdentifier>
<mods:affiliation>Institute of Entomology, College of Plant Protection, Southwest University, Chongqing 400715, China</mods:affiliation>
<mods:nameIdentifier type="email">lzpzena@126.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Cuccodoro, Giulio</mods:namePart>
<mods:affiliation>Museum d'histoire naturelle, Case postale 6434 CH- 1211, Geneve 6, Switzerland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1056</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>17</mods:start>
<mods:end>34</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1056.66553</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1056.66553</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1056-17</mods:identifier>
<mods:identifier type="ZooBank">B62820E7CE524501802CD8E864C7199A</mods:identifier>
<mods:identifier type="Pensoft-UUID">9F460C6A018E5D5B889A40EAE1F34989</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="184587225" LSID="urn:lsid:plazi:treatment:602F4690185057AFB7C03D6DE2D14DA1" httpUri="http://treatment.plazi.org/id/602F4690185057AFB7C03D6DE2D14DA1" lastPageNumber="17" pageId="0" pageNumber="17">
<subSubSection pageId="0" pageNumber="17" type="nomenclature">
<paragraph pageId="0" pageNumber="17">
<taxonomicName LSID="602F4690-1850-57AF-B7C0-3D6DE2D14DA1" authority="(Illiger, 1794)" baseAuthorityName="Illiger" baseAuthorityYear="1794" class="Insecta" family="Staphylinidae" genus="Megarthrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megarthrus hemipterus" order="Coleoptera" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="hemipterus">Megarthrus hemipterus (Illiger, 1794)</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="description">
<paragraph pageId="0" pageNumber="17">
<figureCitation captionStart="Figures 7–9" captionStartId="F3" captionText="Figures 7 - 9. Megarthrus hemipterus (Illiger): habitus, male: dorsal 7 ventral 8 and lateral 9 views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1056.66553.figures7-9" httpUri="https://binary.pensoft.net/fig/578545" pageId="0" pageNumber="17">Figs 7-9</figureCitation>
<figureCitation captionStart="Figures 10–18" captionStartId="F4" captionText="Figures 10 - 18. Megarthrus dentipes Bernhauer (10, 13, 16) M. flavolimbatus Cameron (11, 14, 17) M. hemipterus (Illiger) (12, 15, 18) male: aedeagus in ventral (10 - 12) and lateral (13 - 15) views female: genital segments, dorsal part (16 - 18) in ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17">, 12</figureCitation>
<figureCitation captionStart="Figures 10–18" captionStartId="F4" captionText="Figures 10 - 18. Megarthrus dentipes Bernhauer (10, 13, 16) M. flavolimbatus Cameron (11, 14, 17) M. hemipterus (Illiger) (12, 15, 18) male: aedeagus in ventral (10 - 12) and lateral (13 - 15) views female: genital segments, dorsal part (16 - 18) in ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17">, 15</figureCitation>
<figureCitation captionStart="Figures 10–18" captionStartId="F4" captionText="Figures 10 - 18. Megarthrus dentipes Bernhauer (10, 13, 16) M. flavolimbatus Cameron (11, 14, 17) M. hemipterus (Illiger) (12, 15, 18) male: aedeagus in ventral (10 - 12) and lateral (13 - 15) views female: genital segments, dorsal part (16 - 18) in ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17">, 18</figureCitation>
<figureCitation captionStart="Figure 41" captionStartId="F9" captionText="Figure 41. Distribution map in China of Megarthrus dentipes Bernhauer, M. flavolimbatus Cameron, M. hemipterus (Illiger) and Megarthrus panda sp. nov. Scale bar: 300 km." figureDoi="10.3897/zookeys.1056.66553.figure41" httpUri="https://binary.pensoft.net/fig/578551" pageId="0" pageNumber="17">, 41</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="reference_group">
<paragraph pageId="0" pageNumber="17">
<taxonomicName authorityName="Illiger" authorityYear="1794" class="Insecta" family="Staphylinidae" genus="Silpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Silpha hemiptera" order="Coleoptera" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="hemiptera">Silpha hemiptera</taxonomicName>
Illiger, 1794: 597;
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.79957" author="Cuccodoro, G" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="17" pagination="475 - 524" refId="B3" refString="Cuccodoro, G, 1996. Revision of the Japanese rove-beetles of the genus Megarthrus (Coleoptera, Staphylinidae, Proteininae). Revue suisse de Zoologie 103 (2): 475 - 524, DOI: https://doi.org/10.5962/bhl.part.79957" title="Revision of the Japanese rove-beetles of the genus Megarthrus (Coleoptera, Staphylinidae, Proteininae)." url="https://doi.org/10.5962/bhl.part.79957" volume="103" year="1996">Cuccodoro 1996</bibRefCitation>
: 485 (new synonymy, Japanese records);
<bibRefCitation DOI="https://doi.org/10.1080/00222939700770761" author="Cuccodoro, G" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="17" pagination="1347 - 1415" refId="B6" refString="Cuccodoro, G, Loebl, I, 1997. Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae). Journal of Natural History 31: 1347 - 1415, DOI: https://doi.org/10.1080/00222939700770761" title="Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae)." url="https://doi.org/10.1080/00222939700770761" volume="31" year="1997">
Cuccodoro and
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
1997
</bibRefCitation>
(synonymic framework, detailed redescription, general distribution, the first record for China);
<bibRefCitation DOI="https://doi.org/10.4039/n11-004" author="Cuccodoro, G" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="17" pagination="317 - 331" refId="B7" refString="Cuccodoro, G, Kim, HM, Ahn, KJ, 2011. Megarthrus of Korea, with description of a new species (Coleoptera: Staphylinidae: Proteininae). The Canadian Entomologist 143: 317 - 331, DOI: https://doi.org/10.4039/n11-004" title="Megarthrus of Korea, with description of a new species (Coleoptera: Staphylinidae: Proteininae)." url="https://doi.org/10.4039/n11-004" volume="143" year="2011">Cuccodoro et al. 2011</bibRefCitation>
(synonymic framework, photograph of dorsal habitus, records in Korea).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="diagnosis">
<paragraph pageId="0" pageNumber="17">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="17">
For detailed morphology see
<bibRefCitation DOI="https://doi.org/10.1080/00222939700770761" author="Cuccodoro, G" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="17" pagination="1347 - 1415" refId="B6" refString="Cuccodoro, G, Loebl, I, 1997. Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae). Journal of Natural History 31: 1347 - 1415, DOI: https://doi.org/10.1080/00222939700770761" title="Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae)." url="https://doi.org/10.1080/00222939700770761" volume="31" year="1997">
Cuccodoro and
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
(1997)
</bibRefCitation>
. Combined length of head, pronotum, and elytra 1.6-1.9 mm; maximal pronotal width = 0.9-1.1 mm. Body and appendages (Figs
<figureCitation captionStart="Figures 7–9" captionStartId="F3" captionText="Figures 7 - 9. Megarthrus hemipterus (Illiger): habitus, male: dorsal 7 ventral 8 and lateral 9 views. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1056.66553.figures7-9" httpUri="https://binary.pensoft.net/fig/578545" pageId="0" pageNumber="17">7-9</figureCitation>
) rust brown; head markedly darker with frons slightly paler than vertex. Anterior frontal margin slightly carinate, evenly convex in dorsal view.
</paragraph>
<caption doi="10.3897/zookeys.1056.66553.figures7-9" httpUri="https://binary.pensoft.net/fig/578545" pageId="0" pageNumber="17" start="Figures 7–9" startId="F3">
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Figures 7-9.</emphasis>
<taxonomicName baseAuthorityName="Illiger" baseAuthorityYear="1794" class="Insecta" family="Staphylinidae" genus="Megarthrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megarthrus hemipterus" order="Coleoptera" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="hemipterus">
<emphasis italics="true" pageId="0" pageNumber="17">Megarthrus hemipterus</emphasis>
</taxonomicName>
(Illiger): habitus, male: dorsal
<emphasis bold="true" pageId="0" pageNumber="17">7</emphasis>
ventral
<emphasis bold="true" pageId="0" pageNumber="17">8</emphasis>
and lateral
<emphasis bold="true" pageId="0" pageNumber="17">9</emphasis>
views. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">
<emphasis italics="true" pageId="0" pageNumber="17">Male</emphasis>
.
</emphasis>
Protibia fairly straight and slightly enlarged toward apex, evenly, with adventral side markedly depressed transversely at middle. Mesotrochanter with about a dozen of peg-like setae arranged in two rows. Mesofemur slightly arcuate and slightly swollen. Mesotibia subangulate, bearing peg-like setae arranged in one row. Metatrochanter and metafemur slightly swollen; posterior margin of metatrochanter evenly arcuate; inner margin of metafemur slightly concave in ventral view, forming sharp ridge on entire length. Metatibia swollen, at middle forming conspicuous tooth-like process projecting above flattened apical portion of metatibia; metatibial peg-like setae arranged in 1-2 rows on apical third, the latter group of 15-20, discontinuously with more than 12 peg-like setae arranged in two rows on distal side of tooth-like process; proximal side of tooth-like process broad and flat, bearing 4-10 scattered peg-like setae. Aedeagus (Figs
<figureCitation captionStart="Figures 10–18" captionStartId="F4" captionText="Figures 10 - 18. Megarthrus dentipes Bernhauer (10, 13, 16) M. flavolimbatus Cameron (11, 14, 17) M. hemipterus (Illiger) (12, 15, 18) male: aedeagus in ventral (10 - 12) and lateral (13 - 15) views female: genital segments, dorsal part (16 - 18) in ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17">12</figureCitation>
,
<figureCitation captionStart="Figures 10–18" captionStartId="F4" captionText="Figures 10 - 18. Megarthrus dentipes Bernhauer (10, 13, 16) M. flavolimbatus Cameron (11, 14, 17) M. hemipterus (Illiger) (12, 15, 18) male: aedeagus in ventral (10 - 12) and lateral (13 - 15) views female: genital segments, dorsal part (16 - 18) in ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17">15</figureCitation>
) with ventral wall not narrowed at base and gradually narrowed to mucronate apex in ventral view, with ventral fairly straight to apex in lateral view.
</paragraph>
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">
<emphasis italics="true" pageId="0" pageNumber="17">Female</emphasis>
.
</emphasis>
Gonocoxal plate with lateral portions of dorsobasal margin straight to middle portion forming small blunt process slightly projecting anterad. Dorsal part of genitalia with arcuate sclerite wider at middle (Fig.
<figureCitation captionStart="Figures 10–18" captionStartId="F4" captionText="Figures 10 - 18. Megarthrus dentipes Bernhauer (10, 13, 16) M. flavolimbatus Cameron (11, 14, 17) M. hemipterus (Illiger) (12, 15, 18) male: aedeagus in ventral (10 - 12) and lateral (13 - 15) views female: genital segments, dorsal part (16 - 18) in ventral view. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17">18</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1056.66553.figures10-18" httpUri="https://binary.pensoft.net/fig/578546" pageId="0" pageNumber="17" start="Figures 10–18" startId="F4">
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Figures 10-18.</emphasis>
<taxonomicName authorityName="Bernhauer" authorityYear="1938" class="Insecta" family="Staphylinidae" genus="Megarthrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megarthrus dentipes" order="Coleoptera" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="dentipes">
<emphasis italics="true" pageId="0" pageNumber="17">Megarthrus dentipes</emphasis>
</taxonomicName>
Bernhauer (
<emphasis bold="true" pageId="0" pageNumber="17">10, 13, 16</emphasis>
)
<taxonomicName lsidName="M. flavolimbatus" pageId="0" pageNumber="17" rank="species" species="flavolimbatus">
<emphasis italics="true" pageId="0" pageNumber="17">M. flavolimbatus</emphasis>
</taxonomicName>
Cameron (
<emphasis bold="true" pageId="0" pageNumber="17">11, 14, 17</emphasis>
)
<taxonomicName lsidName="M. hemipterus" pageId="0" pageNumber="17" rank="species" species="hemipterus">
<emphasis italics="true" pageId="0" pageNumber="17">M. hemipterus</emphasis>
</taxonomicName>
(Illiger) (
<emphasis bold="true" pageId="0" pageNumber="17">12, 15, 18</emphasis>
) male: aedeagus in ventral (
<emphasis bold="true" pageId="0" pageNumber="17">10-12</emphasis>
) and lateral (
<emphasis bold="true" pageId="0" pageNumber="17">13-15</emphasis>
) views female: genital segments, dorsal part (
<emphasis bold="true" pageId="0" pageNumber="17">16-18</emphasis>
) in ventral view. Scale bars: 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="comparisons">
<paragraph pageId="0" pageNumber="17">Comparisons and diagnostic notes.</paragraph>
<paragraph pageId="0" pageNumber="17">
See above under
<taxonomicName lsidName="M. dentipes" pageId="0" pageNumber="17" rank="species" species="dentipes">
<emphasis italics="true" pageId="0" pageNumber="17">M. dentipes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="materials_examined">
<paragraph pageId="0" pageNumber="17">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="17">
(
<specimenCount type="generic">4 specimens</specimenCount>
):
<materialsCitation collectingDate="2001-06-15" collectorName="J. Cooter, P. Hlava" country="China" county="Dongling Mts" elevation="1400" latitude="39.97" location="Liu Lang Yu" longLatPrecision="699" longitude="115.43" municipality="Xiaolongmen" specimenCount="1" specimenCount-female="1" stateProvince="Beijing">
<emphasis bold="true" pageId="0" pageNumber="17">
<collectingCountry name="China">China</collectingCountry>
:
<collectingRegion country="China" name="Beijing">Beijing</collectingRegion>
</emphasis>
: ca
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
,
<collectingCounty>Dongling Mts</collectingCounty>
,
<collectingMunicipality>Xiaolongmen</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:602F4690185057AFB7C03D6DE2D14DA1:2308C46AEEE790B98DB6079BC100CF35" country="China" county="Dongling Mts" latitude="39.97" longLatPrecision="699" longitude="115.43" municipality="Xiaolongmen" name="Liu Lang Yu" stateProvince="Beijing">Liu Lang Yu</location>
,
<geoCoordinate degrees="39.97" direction="north" orientation="latitude" precision="555" value="39.97">39.97N</geoCoordinate>
,
<geoCoordinate degrees="115.43" direction="east" orientation="longitude" precision="555" value="115.43">115.43E</geoCoordinate>
,
<collectingDate value="2001-06-15">15.vi.2001</collectingDate>
, leg.
<collectorName>J. Cooter</collectorName>
&amp;
<collectorName>
P.
<normalizedToken originalValue="Hlavá">Hlava</normalizedToken>
</collectorName>
, mixed woodland litter,
<specimenCount type="female">1 ♀</specimenCount>
in MHNG
</materialsCitation>
;
<materialsCitation collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="K. Rost" country="China" location="Heilongjiang Prov." specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Heilongjiang Prov.">
<emphasis bold="true" pageId="0" pageNumber="17">
<collectingRegion country="China" name="Heilongjiang">Heilongjiang Prov.</collectingRegion>
</emphasis>
:
<normalizedToken originalValue="“Süd-Mandshurei">&quot;Sued-Mandshurei</normalizedToken>
Chi-Kuan Shan, leg
<collectorName>K. Rost</collectorName>
&quot;,
<specimenCount type="male">2 ♂</specimenCount>
and
<specimenCount type="female">1 ♀</specimenCount>
in FMNH
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="distribution">
<paragraph pageId="0" pageNumber="17">Distribution and natural history.</paragraph>
<paragraph pageId="0" pageNumber="17">
<taxonomicName baseAuthorityName="Illiger" baseAuthorityYear="1794" class="Insecta" family="Staphylinidae" genus="Megarthrus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Megarthrus hemipterus" order="Coleoptera" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="hemipterus">
<emphasis italics="true" pageId="0" pageNumber="17">Megarthrus hemipterus</emphasis>
</taxonomicName>
is the most widespread member of the genus, with records ranging from the United Kingdom in the West to China, South Korea, Japan, and Far East Russia in the East (
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.79957" author="Cuccodoro, G" journalOrPublisher="Revue suisse de Zoologie" pageId="0" pageNumber="17" pagination="475 - 524" refId="B3" refString="Cuccodoro, G, 1996. Revision of the Japanese rove-beetles of the genus Megarthrus (Coleoptera, Staphylinidae, Proteininae). Revue suisse de Zoologie 103 (2): 475 - 524, DOI: https://doi.org/10.5962/bhl.part.79957" title="Revision of the Japanese rove-beetles of the genus Megarthrus (Coleoptera, Staphylinidae, Proteininae)." url="https://doi.org/10.5962/bhl.part.79957" volume="103" year="1996">Cuccodoro 1996</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1080/00222939700770761" author="Cuccodoro, G" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="17" pagination="1347 - 1415" refId="B6" refString="Cuccodoro, G, Loebl, I, 1997. Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae). Journal of Natural History 31: 1347 - 1415, DOI: https://doi.org/10.1080/00222939700770761" title="Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae)." url="https://doi.org/10.1080/00222939700770761" volume="31" year="1997">
Cuccodoro and
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
1997
</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.4039/n11-004" author="Cuccodoro, G" journalOrPublisher="The Canadian Entomologist" pageId="0" pageNumber="17" pagination="317 - 331" refId="B7" refString="Cuccodoro, G, Kim, HM, Ahn, KJ, 2011. Megarthrus of Korea, with description of a new species (Coleoptera: Staphylinidae: Proteininae). The Canadian Entomologist 143: 317 - 331, DOI: https://doi.org/10.4039/n11-004" title="Megarthrus of Korea, with description of a new species (Coleoptera: Staphylinidae: Proteininae)." url="https://doi.org/10.4039/n11-004" volume="143" year="2011">Cuccodoro et al. 2011</bibRefCitation>
). It occurs in a wide range of microhabitats (carrion, fungi, leaf litter, and various types of decaying organic matter); in South Korea it was abundantly collected from fresh and rotten fungi. The first record for China without precise locality data (
<bibRefCitation DOI="https://doi.org/10.1080/00222939700770761" author="Cuccodoro, G" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="17" pagination="1347 - 1415" refId="B6" refString="Cuccodoro, G, Loebl, I, 1997. Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae). Journal of Natural History 31: 1347 - 1415, DOI: https://doi.org/10.1080/00222939700770761" title="Revision of the Palaearctic rove-beetles of the genus Megarthrus Curtis (Coleoptera: Staphylinidae: Proteininae)." url="https://doi.org/10.1080/00222939700770761" volume="31" year="1997">
Cuccodoro and
<normalizedToken originalValue="Löbl">Loebl</normalizedToken>
1997
</bibRefCitation>
) refers to the three specimens cited here from Chi-Kuan Shan from the Max
<normalizedToken originalValue="Bernhauer’s">Bernhauer's</normalizedToken>
collection, housed in the FMNH. The species, which occurs on most land masses surrounding the Sea of Japan, seems to reach its southern limit of extension in China in the Western Hills (Fig.
<figureCitation captionStart="Figure 41" captionStartId="F9" captionText="Figure 41. Distribution map in China of Megarthrus dentipes Bernhauer, M. flavolimbatus Cameron, M. hemipterus (Illiger) and Megarthrus panda sp. nov. Scale bar: 300 km." figureDoi="10.3897/zookeys.1056.66553.figure41" httpUri="https://binary.pensoft.net/fig/578551" pageId="0" pageNumber="17">41</figureCitation>
), where it was collected in mid-April at 1400 m a. s. l. from mixed woodland litter on Mount Dongling (Beijing).
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</subSubSection>
</treatment>
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