treatments-xml/data/03/D1/CF/03D1CF44F7158253A8B4DA91D06AF88E.xml
2024-06-21 12:22:17 +02:00

446 lines
49 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="ABEB02706F590D2980C87977CEB2AC00" ID-DOI="10.5281/zenodo.180677" ID-GBIF-Dataset="6b8ec8f2-f45d-4239-968e-d63cee165d73" ID-ISSN="1175-5326" ID-Zenodo-Dep="180677" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459812935578" checkinUser="plazi" docAuthor="Gerasimova, Elena, Erpenbeck, Dirk &amp; Plotkin, Alexander" docDate="2008" docId="03D1CF44F7158253A8B4DA91D06AF88E" docLanguage="en" docName="zt01694p037.pdf" docOrigin="Zootaxa 1694" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Johannesia reticulosa Thiele 1905" docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="FFE8B73CF7078249A823DB3ED42CFFC6" masterDocTitle="Vosmaeria Fristedt, 1885 (Porifera, Demospongiae, Halichondriidae): revision of species, phylogenetic reconstruction and evidence for split" masterLastPageNumber="37" masterPageNumber="1" pageNumber="19" updateTime="1698221197441" updateUser="plazi">
<mods:mods id="3EED5AD4CB8FA651746610FEC6A3C4B4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0F16F89432DA96D5F1ACD7B8D4EA2374">
<mods:title id="9C56C64707B266C8E5DAD2B280713094">Vosmaeria Fristedt, 1885 (Porifera, Demospongiae, Halichondriidae): revision of species, phylogenetic reconstruction and evidence for split</mods:title>
</mods:titleInfo>
<mods:name id="6887E944F983C015F9B1756E6C5F4FC7" type="personal">
<mods:role id="331C16708F5E30A2895B0EB6886E9211">
<mods:roleTerm id="79C6CFF8CEE83B6E33B4F1521AF34BFE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="842BC0FAC41F30F54EAA8CDBC9C1ECC5">Gerasimova, Elena</mods:namePart>
</mods:name>
<mods:name id="2A153C6CD9ED3195CBB93F076692C0FC" type="personal">
<mods:role id="549E7CE746A7AEBBBCAA82348A9D21F3">
<mods:roleTerm id="F44EA30E4AEEBA12B6294223FC84F0E5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C72998BDA48E7405C2CC8BFF72CA2399">Erpenbeck, Dirk</mods:namePart>
</mods:name>
<mods:name id="D31EEF376259F53F76A88344846B961A" type="personal">
<mods:role id="EA5F8702390A2109F88D8117805B4BA2">
<mods:roleTerm id="E88D502694C0E8A9837C881E73F5C6E2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98C1B9F8E595AA1EF805C2EDDFA3283F">Plotkin, Alexander</mods:namePart>
</mods:name>
<mods:typeOfResource id="5DDD79BB115A7EE192879797784A4EB3">text</mods:typeOfResource>
<mods:relatedItem id="C7AB279F878C8192C091F6FEDB4F4E63" type="host">
<mods:titleInfo id="38BED2012FFA4BA25CDAF2E14A232E21">
<mods:title id="A2110EF54FC02E6E4E8B51D359C40363">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="22D7446C8B7EF565A5D7CC96019D326F">
<mods:date id="0C0BD78560FE5116D516DC7F9CA5CEBB">2008</mods:date>
<mods:detail id="6099B3E9CDAA655692E786E13EBDD5C8" type="volume">
<mods:number id="6C3F242F5A8356D147045DAA379344F5">1694</mods:number>
</mods:detail>
<mods:extent id="747844AC717D1EAFA0CC01A4BBA6D914" unit="page">
<mods:start id="D8197238B36D5D331BC5CF15731C9CB5">1</mods:start>
<mods:end id="2B67F6BF00F85E6FC37801A11D89487A">37</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="FFD4DDA40C1D066F7D7A88AD90E6DEE8">journal article</mods:classification>
<mods:identifier id="0FAF4A5D935BE4837D89F9AD1944DB81" type="DOI">10.5281/zenodo.180677</mods:identifier>
<mods:identifier id="7760E3C7DA883F854D0A8A17A608C31D" type="GBIF-Dataset">6b8ec8f2-f45d-4239-968e-d63cee165d73</mods:identifier>
<mods:identifier id="7F1DE70B90D18743FC4BBEE140B6E1D6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="39133D2916853960A9598BCA009788F6" type="Zenodo-Dep">180677</mods:identifier>
</mods:mods>
<treatment id="03D1CF44F7158253A8B4DA91D06AF88E" ID-DOI="http://doi.org/10.5281/zenodo.6234169" ID-GBIF-Taxon="119354146" ID-Zenodo-Dep="6234169" LSID="urn:lsid:plazi:treatment:03D1CF44F7158253A8B4DA91D06AF88E" httpUri="http://treatment.plazi.org/id/03D1CF44F7158253A8B4DA91D06AF88E" lastPageId="26" lastPageNumber="27" pageId="18" pageNumber="19">
<subSubSection id="C3622DD9F715825BA8B4DA91D533FDD2" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BC77E52F715825BA8B4DA91D67FFE0F" blockId="18.[151,595,431,457]" box="[151,595,431,457]" pageId="18" pageNumber="19">
<heading id="D08FC93EF715825BA8B4DA91D67FFE0F" bold="true" box="[151,595,431,457]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<taxonomicName id="4C7805D1F715825BA8B4DA91D67FFE0F" authority="Thiele, 1905" authorityName="Thiele" authorityYear="1905" box="[151,595,431,457]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="18" pageNumber="19" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F715825BA8B4DA91D67FFE0F" bold="true" box="[151,595,431,457]" pageId="18" pageNumber="19">
<emphasis id="B90CA240F715825BA8B4DA91D58CFE0F" bold="true" box="[151,416,431,457]" italics="true" pageId="18" pageNumber="19">Johannesia reticulosa</emphasis>
(
<bibRefCitation id="EFE903A3F715825BA993DA91D667FE0F" author="Thiele" box="[432,587,431,457]" pageId="18" pageNumber="37" refString="Thiele, J. (1905) Die Kiesel- und Hornschwamme der Sammlung Plate. Zoologische Jahrbucher Supplement 6 (Fauna Chiliensis III), 407 - 496." type="book chapter" year="1905">Thiele, 1905</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC77E52F715825BA8B4DAC4D533FDD2" blockId="18.[151,1114,506,569]" box="[151,287,506,532]" pageId="18" pageNumber="19">Synonymy:</paragraph>
</subSubSection>
<subSubSection id="C3622DD9F7158253A8B4D91CD06AF88E" lastPageId="26" lastPageNumber="27" pageId="18" pageNumber="19" type="description">
<paragraph id="8BC77E52F715825BA8B4D91CD076FDFF" blockId="18.[151,1114,506,569]" box="[151,1114,546,569]" pageId="18" pageNumber="19">
<treatmentCitationGroup id="AB68597CF715825BA8B4D91CD076FDFF" box="[151,1114,546,569]" pageId="18" pageNumber="19">
<emphasis id="B90CA240F715825BA8B4D91CD5A7FDFF" box="[151,395,546,569]" italics="true" pageId="18" pageNumber="19">
<taxonomicName id="4C7805D1F715825BA8B4D91CD554FDFF" box="[151,376,546,569]" class="Demospongiae" family="Halichondriidae" genus="Vosmaeria" kingdom="Animalia" order="Halichondrida" pageId="18" pageNumber="19" phylum="Porifera" rank="species" species="reticulosa">Vosmaeria reticulosa</taxonomicName>
</emphasis>
<treatmentCitation id="0AD95843F715825BA9B1D91CD67DFDFF" author="Thiele" box="[402,593,546,569]" page="423" pageId="18" pageNumber="19" year="1905">
<bibRefCitation id="EFE903A3F715825BA9B1D91CD631FDFF" author="Thiele" box="[402,541,546,569]" pageId="18" pageNumber="37" refString="Thiele, J. (1905) Die Kiesel- und Hornschwamme der Sammlung Plate. Zoologische Jahrbucher Supplement 6 (Fauna Chiliensis III), 407 - 496." type="book chapter" year="1905">Thiele, 1905</bibRefCitation>
: 423
</treatmentCitation>
, figs. 45ac; Erpenbeck &amp; van Soest, 2002: 815.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC77E52F715825BA8B4D954D54BFD12" pageId="18" pageNumber="19">
<table id="F9788CF2F7157DB6A8B4D954D1B0FD12" box="[151,1436,618,724]" colsContinueIn="27.[151,1436,445,551]" gridcols="14" gridrows="3" pageId="18" pageNumber="19">
<tr id="35487C10F7157DB6A8B4D954D1B0FD42" box="[151,1436,618,644]" gridrow="0" pageId="18" pageNumber="19" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="7699156CF7157DB6A8B4D954D5AFFD42" box="[151,387,618,644]" gridcol="0" gridrow="0" pageId="18" pageNumber="19">Type material:</th>
</tr>
<tr id="35487C10F7157DB6A8B4D9ACD1B0FD6A" box="[151,1436,658,684]" gridrow="1" pageId="18" pageNumber="19">
<th id="7699156CF7157DB6A8B4D9ACD5AFFD6A" box="[151,387,658,684]" gridcol="0" gridrow="1" pageId="18" pageNumber="19">Holotype: ZMB</th>
<td id="7699156CF7157DB6A9A8D9ACD598FD6A" box="[395,436,658,684]" gridcol="1" gridrow="1" pageId="18" pageNumber="19">Por</td>
<td id="7699156CF7157DB6A998D9ACD5DAFD6A" box="[443,502,658,684]" gridcol="2" gridrow="1" pageId="18" pageNumber="19">3279</td>
<td id="7699156CF7157DB6A9DDD9ACD65AFD6A" box="[510,630,658,684]" gridcol="3" gridrow="1" pageId="18" pageNumber="19">(specimen</td>
<td id="7699156CF7157DB6AA5DD9ACD6B9FD6A" box="[638,661,658,684]" gridcol="4" gridrow="1" pageId="18" pageNumber="19">in</td>
<td id="7699156CF7157DB6AABFD9ACD729FD6A" box="[668,773,658,684]" gridcol="5" gridrow="1" pageId="18" pageNumber="19">alcohol):</td>
<td id="7699156CF7157DB6AB2FD9ACD741FD6A" box="[780,877,658,684]" gridcol="6" gridrow="1" pageId="18" pageNumber="19">Iquique,</td>
<td id="7699156CF7157DB6AB56D9ACD7F3FD6A" box="[885,991,658,684]" gridcol="7" gridrow="1" pageId="18" pageNumber="19">Northern</td>
<td id="7699156CF7157DB6ABC5D9ACD06DFD6A" box="[998,1089,658,684]" gridcol="8" gridrow="1" pageId="18" pageNumber="19">Chilean</td>
<td id="7699156CF7157DB6AC6AD9ACD0B8FD6A" box="[1097,1172,658,684]" gridcol="9" gridrow="1" pageId="18" pageNumber="19">Coast;</td>
<td id="7699156CF7157DB6ACBFD9ACD095FD6A" box="[1180,1209,658,684]" gridcol="10" gridrow="1" pageId="18" pageNumber="19">30</td>
<td id="7699156CF7157DB6ACE2D9ACD0CCFD6A" box="[1217,1248,658,684]" gridcol="11" gridrow="1" pageId="18" pageNumber="19">m;</td>
<td id="7699156CF7157DB6ACCBD9ACD17EFD6A" box="[1256,1362,658,684]" gridcol="12" gridrow="1" pageId="18" pageNumber="19">February</td>
<td id="7699156CF7157DB6AD7AD9ACD1B0FD6A" box="[1369,1436,658,684]" gridcol="13" gridrow="1" pageId="18" pageNumber="19">1895;</td>
</tr>
<tr id="35487C10F7157DB6A8B4D984D1B0FD12" box="[151,1436,698,724]" gridrow="2" pageId="18" pageNumber="19" rowspan-1="1" rowspan-10="1" rowspan-11="1" rowspan-12="1" rowspan-13="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1" rowspan-9="1">
<th id="7699156CF7157DB6A8B4D984D5AFFD12" box="[151,387,698,724]" gridcol="0" gridrow="2" pageId="18" pageNumber="19">collector L. Plate.</th>
</tr>
</table>
</paragraph>
<paragraph id="8BC77E52F715825BA8B4D834D7CCFCE2" blockId="18.[151,1437,778,924]" box="[151,992,778,804]" pageId="18" pageNumber="19">Additional material (collected by Helmut Lehnert and fixed in alcohol):</paragraph>
<paragraph id="8BC77E52F715825BA8E5D80CD6A8FC5A" blockId="18.[151,1437,778,924]" pageId="18" pageNumber="19">
Region of Paracas, Peruvian Coast: HL 2352 (
<specimenCount id="9D7EB5DBF715825BAACDD80CD758FC8A" box="[750,884,818,844]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
) and HL 2354 (
<specimenCount id="9D7EB5DBF715825BAC10D80CD09BFC8A" box="[1075,1207,818,844]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
used for both morphological studies and molecular analysis):
<collectingCountry id="F36F3EC2F715825BAA85D864D6FCFCB2" box="[678,720,858,884]" name="Iceland" pageId="18" pageNumber="19">Isla</collectingCountry>
San Gallan;
<quantity id="4C80D3B7F715825BAB52D864D7F2FCB2" box="[881,990,858,884]" metricMagnitude="1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="18" pageNumber="19" unit="m" value="13.5" valueMax="15.0" valueMin="12.0">1215 m</quantity>
;
<date id="FFC65892F715825BABCED864D059FCB2" box="[1005,1141,858,884]" pageId="18" pageNumber="19" value="2002-08-13">13.08.2002</date>
. HL 2425 (
<specimenCount id="9D7EB5DBF715825BAD26D864D1BCFCB2" box="[1285,1424,858,884]" pageId="18" pageNumber="19" type="generic">1 specimen</specimenCount>
): Islotes Dos Marias;
<quantity id="4C80D3B7F715825BA9A1D8BCD5C1FC5A" box="[386,493,898,924]" metricMagnitude="1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="18" pageNumber="19" unit="m" value="13.5" valueMax="15.0" valueMin="12.0">1215 m</quantity>
;
<date id="FFC65892F715825BA9DAD8BCD6ADFC5A" box="[505,641,898,924]" pageId="18" pageNumber="19" value="2002-08-18">18.08.2002</date>
.
</paragraph>
<paragraph id="8BC77E52F715825BA8B4D8ECD504FC2A" blockId="18.[151,1436,978,1164]" box="[151,296,978,1004]" pageId="18" pageNumber="19">Description:</paragraph>
<paragraph id="8BC77E52F715825BA8E5D8C4D59CFBD5" blockId="18.[151,1436,978,1164]" box="[198,432,1018,1043]" pageId="18" pageNumber="19">
<emphasis id="B90CA240F715825BA8E5D8C4D59CFBD5" box="[198,432,1018,1043]" italics="true" pageId="18" pageNumber="19">External characters</emphasis>
</paragraph>
<paragraph id="8BC77E52F715825BA8E5DF1CD73AFB4A" blockId="18.[151,1436,978,1164]" pageId="18" pageNumber="19">
<typeStatus id="54C3C0F0F715825BA8E5DF1CD51FFBFA" box="[198,307,1058,1084]" pageId="18" pageNumber="19" type="holotype">Holotype</typeStatus>
a massive fist-shaped sponge, measuring approx.
<date id="FFC65892F715825BABA2DF1CD782FBFA" box="[897,942,1058,1084]" pageId="18" pageNumber="19" value="1950-10">50x</date>
<date id="FFC65892F715825BAB8DDF1CD7F6FBFA" box="[942,986,1058,1084]" pageId="18" pageNumber="19" value="1948-10">48x</date>
<quantity id="4C80D3B7F715825BABFADF1CD000FBFA" box="[985,1068,1058,1084]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="18" pageNumber="19" unit="mm" value="34.0">34 mm</quantity>
(fig. 10). Colour beige in alcohol. Surface smooth and ridged. Several oscula of
<quantity id="4C80D3B7F715825BAAC9DF74D760FBA2" box="[746,844,1098,1124]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="4.0" metricValueMin="1.0" pageId="18" pageNumber="19" unit="mm" value="2.5" valueMax="4.0" valueMin="1.0">14 mm</quantity>
in diameter visible on the surface, ostia cannot be seen macroscopically. Ectosome loose and detachable.
</paragraph>
<caption id="DF072EDAF715825BA8B4DDE0D00AF930" box="[151,1062,1758,1782]" httpUri="https://zenodo.org/record/180687/files/figure.png" pageId="18" pageNumber="19" targetBox="[264,1323,1221,1735]" targetPageId="18">
<paragraph id="8BC77E52F715825BA8B4DDE0D00AF930" blockId="18.[151,1062,1758,1782]" box="[151,1062,1758,1782]" pageId="18" pageNumber="19">
<emphasis id="B90CA240F715825BA8B4DDE0D50AF930" bold="true" box="[151,294,1758,1782]" pageId="18" pageNumber="19">FIGURE 10.</emphasis>
<taxonomicName id="4C7805D1F715825BA90EDDE1D63BF930" box="[301,535,1759,1782]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="18" pageNumber="19" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F715825BA90EDDE1D63BF930" box="[301,535,1759,1782]" italics="true" pageId="18" pageNumber="19">Johannesia reticulosa</emphasis>
</taxonomicName>
, holotype: external morphology. Scale bar: 1 cm.
</paragraph>
</caption>
<paragraph id="8BC77E52F715825AA8E5DC6CD598FF1C" blockId="18.[151,1436,1874,2020]" lastBlockId="19.[151,1436,152,538]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
According to the description of
<bibRefCitation id="EFE903A3F715825BAA69DC6CD6DCF8AA" author="Thiele" box="[586,752,1874,1900]" pageId="18" pageNumber="37" refString="Thiele, J. (1905) Die Kiesel- und Hornschwamme der Sammlung Plate. Zoologische Jahrbucher Supplement 6 (Fauna Chiliensis III), 407 - 496." type="book chapter" year="1905">Thiele (1905)</bibRefCitation>
, the specimen grew together with
<taxonomicName id="4C7805D1F715825BACB1DC6CD1B0F8AD" box="[1170,1436,1874,1899]" class="Demospongiae" family="Suberitidae" genus="Plicatellopsis" kingdom="Animalia" order="Hadromerida" pageId="18" pageNumber="19" phylum="Porifera" rank="species" species="expansa">
<emphasis id="B90CA240F715825BACB1DC6CD1B0F8AD" box="[1170,1436,1874,1899]" italics="true" pageId="18" pageNumber="19">Plicatellopsis expansa</emphasis>
</taxonomicName>
(
<taxonomicName id="4C7805D1F715825BA882DC44D550F855" box="[161,380,1914,1939]" class="Demospongiae" family="Suberitidae" genus="Plicatella" higherTaxonomySource="CoL" kingdom="Animalia" order="Suberitida" pageId="18" pageNumber="19" phylum="Porifera" rank="species" species="expansa">
<emphasis id="B90CA240F715825BA882DC44D550F855" box="[161,380,1914,1939]" italics="true" pageId="18" pageNumber="19">Plicatella expansa</emphasis>
</taxonomicName>
in the original description), another sponge described by him. At the time of our study these two sponges had been separated from each other already, and stored in different jars.
<typeStatus id="54C3C0F0F715825BAC9DDC9CD0D4F87A" box="[1214,1272,1954,1980]" pageId="18" pageNumber="19">Type</typeStatus>
material of
<taxonomicName id="4C7805D1F715825BADA5DC9CD4DBF825" class="Demospongiae" family="Suberitidae" genus="Plicatella" higherTaxonomySource="CoL" kingdom="Animalia" order="Suberitida" pageId="18" pageNumber="19" phylum="Porifera" rank="species" species="expansa">
<emphasis id="B90CA240F715825BADA5DC9CD4DBF825" italics="true" pageId="18" pageNumber="19">P. expansa</emphasis>
</taxonomicName>
was represented by two fragments which macroscopically could not be distinguished from the specimen of
<taxonomicName id="4C7805D1F714825AA8D0DBA6D5C6FF77" box="[243,490,152,177]" class="Demospongiae" family="Halichondriidae" genus="Vosmaeria" kingdom="Animalia" order="Halichondrida" pageId="19" pageNumber="20" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F714825AA8D0DBA6D5C6FF77" box="[243,490,152,177]" italics="true" pageId="19" pageNumber="20">Vosmaeria reticulosa</emphasis>
</taxonomicName>
but differed greatly from it by skeleton structure and spicules (only styles were observed in
<taxonomicName id="4C7805D1F714825AA905DBFED58FFF1F" box="[294,419,192,217]" class="Demospongiae" family="Suberitidae" genus="Plicatella" higherTaxonomySource="CoL" kingdom="Animalia" order="Suberitida" pageId="19" pageNumber="20" phylum="Porifera" rank="species" species="expansa">
<emphasis id="B90CA240F714825AA905DBFED58FFF1F" box="[294,419,192,217]" italics="true" pageId="19" pageNumber="20">P. expansa</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BC77E52F714825AA8E5DBD6D6FFFEEC" blockId="19.[151,1436,152,538]" pageId="19" pageNumber="20">
Peruvian samples are represented by small sponge fragments each of few cubic cm. Their surface and inner texture are similar to those of the
<typeStatus id="54C3C0F0F714825AAA46DA2ED6E3FEEC" box="[613,719,272,298]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BC77E52F714825AA8E5DA06D506FE97" blockId="19.[151,1436,152,538]" box="[198,298,312,337]" pageId="19" pageNumber="20">
<emphasis id="B90CA240F714825AA8E5DA06D506FE97" box="[198,298,312,337]" italics="true" pageId="19" pageNumber="20">Skeleton</emphasis>
</paragraph>
<paragraph id="8BC77E52F714825AA8E5DA5ED6F3FDDC" blockId="19.[151,1436,152,538]" pageId="19" pageNumber="20">Choanosomal skeleton made by tracts of (sub)tylostyles running from sponge base to the surface (fig. 11A). The tracts branch and anastomose. Their thickness 1050 µm. Free tylostyles and oxeas scattered between the principal tracts. Ectosomal skeleton measuring 300500 µm in thickness constituted by the diverging principal tracts intercrossed with tylostyles and oxeas (fig. 11B). In tangential surface view this intercrossing forms an irregular network (fig. 12).</paragraph>
<caption id="DF072EDAF714825AA8B4DC80D713F833" httpUri="https://zenodo.org/record/180688/files/figure.png" pageId="19" pageNumber="20" targetBox="[264,1322,591,1967]" targetPageId="19">
<paragraph id="8BC77E52F714825AA8B4DC80D713F833" blockId="19.[151,1435,1982,2037]" pageId="19" pageNumber="20">
<emphasis id="B90CA240F714825AA8B4DC80D50AF810" bold="true" box="[151,294,1982,2006]" pageId="19" pageNumber="20">FIGURE 11.</emphasis>
<taxonomicName id="4C7805D1F714825AA90DDC80D636F813" box="[302,538,1982,2005]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="19" pageNumber="20" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F714825AA90DDC80D636F813" box="[302,538,1982,2005]" italics="true" pageId="19" pageNumber="20">Johannesia reticulosa</emphasis>
</taxonomicName>
, holotype: longitudinal section of the body with skeleton. Scale bars for A: 1 mm, scale bar for B: 0.5 mm. Ageneral view. Bdetail of ectosome.
</paragraph>
</caption>
<caption id="DF072EDAF713825DA8B4D8A3D0F7FC72" box="[151,1243,925,949]" httpUri="https://zenodo.org/record/180689/files/figure.png" pageId="20" pageNumber="21" targetBox="[265,1324,188,910]" targetPageId="20">
<paragraph id="8BC77E52F713825DA8B4D8A3D0F7FC72" blockId="20.[151,1243,925,949]" box="[151,1243,925,949]" pageId="20" pageNumber="21">
<emphasis id="B90CA240F713825DA8B4D8A3D50AFC73" bold="true" box="[151,294,925,949]" pageId="20" pageNumber="21">FIGURE 12.</emphasis>
<taxonomicName id="4C7805D1F713825DA90ED8A3D63BFC72" box="[301,535,925,948]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="20" pageNumber="21" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F713825DA90ED8A3D63BFC72" box="[301,535,925,948]" italics="true" pageId="20" pageNumber="21">Johannesia reticulosa</emphasis>
</taxonomicName>
, holotype: surface view of detached ectosome. Scale bar: 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BC77E52F713825DA8E5D8D2D5B7FBC3" blockId="20.[151,1437,1004,1909]" box="[198,411,1004,1029]" pageId="20" pageNumber="21">
<emphasis id="B90CA240F713825DA8E5D8D2D5B7FBC3" box="[198,411,1004,1029]" italics="true" pageId="20" pageNumber="21">Aquiferous system</emphasis>
</paragraph>
<paragraph id="8BC77E52F713825DA8E5DF2DD678FB63" blockId="20.[151,1437,1004,1909]" pageId="20" pageNumber="21">Oscula and ostia located in the holes of the ectosomal network. Ostia connected with large subectosomal cavities (300650 µm in diameter — fig. 11B). Inhalant canals run from the subectosomal cavities to the choanosomal chambers. Exhalant canals come from the choanocyte chambers to the choanosomal atriums, which are connected to oscula at the surface.</paragraph>
<paragraph id="8BC77E52F713825DA8E5DF8AD612FB0B" blockId="20.[151,1437,1004,1909]" box="[198,574,1204,1229]" pageId="20" pageNumber="21">
<emphasis id="B90CA240F713825DA8E5DF8AD612FB0B" box="[198,574,1204,1229]" italics="true" pageId="20" pageNumber="21">Spicules (means in parentheses)</emphasis>
</paragraph>
<paragraph id="8BC77E52F713825DA8E5DFE5D4E3FA7B" blockId="20.[151,1437,1004,1909]" pageId="20" pageNumber="21">
-(Sub)tylostyles (figs. 13CF): Altogether, 240 tylostyles observed (30 ectosomal and 30 choanosomal spicules in each specimen). Tylostyles always with acerate distal tips and usually slightly curved. In the
<typeStatus id="54C3C0F0F713825DAD7DDE3DD4E6FA83" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
most tylostyles with well developed tyles, which are located terminally in choanosomal spicules and slightly displaced in many ectosomal ones (table 5). Measurements (table 6): length 104369 µm (255 µm), proximal diameter 4.211.7 µm (6.7 µm), diameter of tyle 5.011.7 (8.0 µm), central diameter 4.211.7 (7.3 µm).
</paragraph>
<paragraph id="8BC77E52F713825DA8E5DEF5D67DF96B" blockId="20.[151,1437,1004,1909]" pageId="20" pageNumber="21">In the Peruvian specimen HL 2352 most tylostyles with weakly developed tyles, which are terminally located in choanosomal spicules but slightly displaced in many ectosomal ones (table 5). In the other two Peruvian specimens most tylostyles with weakly developed tyles, terminally located both in choanosomal as well as ectosomal spicules. Altogether, the tylostyles of the Peruvian specimens measure 158400 µm (276 µm) in length, 3.512.5 µm (5.9 µm) in proximal diameter with tyle diameter of 3.515 (7.3 µm) and 3.515 (6.5 µm) in central diameter (table 6).</paragraph>
<paragraph id="8BC77E52F713825DA8E5DD85D75DF8B3" blockId="20.[151,1437,1004,1909]" pageId="20" pageNumber="21">
-Oxeas (figs. 13AB): Altogether, 240 oxeas observed (30 ectosomal and 30 choanosomal spicules in each specimen). All spicules with acerate tips. In the
<typeStatus id="54C3C0F0F713825DAB27DDDDD742F93B" box="[772,878,1763,1789]" pageId="20" pageNumber="21" type="holotype">holotype</typeStatus>
, oxeas usually moderately or even considerably curved (table 5), measuring 292550 µm (447 µm) in length and 4.213.3 µm (10.8 µm) in central diameter (table 6). In the Peruvian specimens, oxeas mainly slightly curved (table 5), measuring 215710 µm (503 µm) in length and 520 µm (11.8 µm) in central diameter (table 6).
</paragraph>
<caption id="DF072EDAF712825CA8B4DBA9D765FF69" box="[151,841,151,176]" pageId="21" pageNumber="22">
<paragraph id="8BC77E52F712825CA8B4DBA9D765FF69" blockId="21.[151,1436,151,313]" box="[151,841,151,176]" pageId="21" pageNumber="22">
TABLE 5.
<taxonomicName id="4C7805D1F712825CA933DBA6D62EFF76" box="[272,514,152,176]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="21" pageNumber="22" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F712825CA933DBA6D62EFF76" bold="true" box="[272,514,152,176]" italics="true" pageId="21" pageNumber="22">Johannesia reticulosa</emphasis>
</taxonomicName>
: variability of spicule shape.
</paragraph>
</caption>
<paragraph id="8BC77E52F712825CA8B4DB84D6C1FEFF" blockId="21.[151,1436,151,313]" pageId="21" pageNumber="22">Number of spicules characterized by the respective feature is given in every cell. Abbreviations: Curvature: Sstraight, SCslightly curved, MCmoderately curved, CCconsiderably curved; Slenderness: Ffusiform, SLslender; Tyle development: Aabsent, WKweakly developed, WLwell developed; Tyle location: Tterminal, SDslightly displaced, MDmoderately displaced, CDconsiderably displaced.</paragraph>
<paragraph id="8BC77E52F712825CAB07DA61D57CFC7B" blockId="21.[804,1371,351,372]" lastBlockId="21.[269,1420,905,957]" pageId="21" pageNumber="22">
(Sub)tylostyles Oxeas Islotes Dos Ectosome
<date id="FFC65892F712825CAA39D8B7D6A7FC58" box="[538,651,905,926]" pageId="21" pageNumber="22" value="1928-01-01">1 28 1</date>
0 0
<date id="FFC65892F712825CAB39D8B7D788FC58" box="[794,932,905,926]" pageId="21" pageNumber="22" value="1923-07-30">30 7 23</date>
0 21 2 0 0 2 28 0 0 Marias
</paragraph>
<paragraph id="8BC77E52F712825CAA18DAB1D1A0FCB5" pageId="21" pageNumber="22">
<table id="F9788CF2F7127DB6A8BFDAB1D1B4FCB5" box="[156,1432,399,883]" gridcols="6" gridrows="7" pageId="21" pageNumber="22">
<tr id="35487C10F7127DB6A8BFDAB1D1B4FE02" box="[156,1432,399,452]" gridrow="0" pageId="21" pageNumber="22" rowspan-0="1">
<th id="7699156CF7127DB6A9A0DAB1D6E4FE02" box="[387,712,399,452]" gridcol="1" gridrow="0" pageId="21" pageNumber="22">Curvature</th>
<th id="7699156CF7127DB6AAC7DAB1D719FE02" box="[740,821,399,452]" gridcol="2" gridrow="0" pageId="21" pageNumber="22">Slender- ness</th>
<th id="7699156CF7127DB6AB79DAB1D7C8FE02" box="[858,996,399,452]" gridcol="3" gridrow="0" pageId="21" pageNumber="22">Tyle develop- ment</th>
<th id="7699156CF7127DB6AC21DAB1D0EAFE02" box="[1026,1222,399,452]" gridcol="4" gridrow="0" pageId="21" pageNumber="22">Tyle location</th>
<th id="7699156CF7127DB6ACC6DAB1D1B4FE02" box="[1253,1432,399,452]" gridcol="5" gridrow="0" pageId="21" pageNumber="22">Curvature</th>
</tr>
<tr id="35487C10F7127DB6A8BFDAE4D1B4FE29" box="[156,1432,474,495]" gridrow="1" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFDAE4D571FE29" box="[156,349,474,495]" gridcol="0" gridrow="1" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CA8BFDAE4D579FE29" bold="true" box="[156,341,474,495]" pageId="21" pageNumber="22">Specimen Region</emphasis>
</th>
<td id="7699156CF7127DB6A9A0DAE4D6E4FE29" box="[387,712,474,495]" gridcol="1" gridrow="1" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CA9A0DAE4D6E4FE29" bold="true" box="[387,712,474,495]" pageId="21" pageNumber="22">Body part S SC MC CC</emphasis>
</td>
<td id="7699156CF7127DB6AAC7DAE4D719FE29" box="[740,821,474,495]" gridcol="2" gridrow="1" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CAAC9DAE4D719FE29" bold="true" box="[746,821,474,495]" pageId="21" pageNumber="22">F SL</emphasis>
</td>
<td id="7699156CF7127DB6AB79DAE4D7C8FE29" box="[858,996,474,495]" gridcol="3" gridrow="1" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CAB7FDAE4D7C8FE29" bold="true" box="[860,996,474,495]" pageId="21" pageNumber="22">A WK WL</emphasis>
</td>
<td id="7699156CF7127DB6AC21DAE4D0EAFE29" box="[1026,1222,474,495]" gridcol="4" gridrow="1" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CAC25DAE4D0EAFE29" bold="true" box="[1030,1222,474,495]" pageId="21" pageNumber="22">T SD MD CD</emphasis>
</td>
<td id="7699156CF7127DB6ACC6DAE4D1B4FE29" box="[1253,1432,474,495]" gridcol="5" gridrow="1" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CACC9DAE4D1B4FE29" bold="true" box="[1258,1432,474,495]" pageId="21" pageNumber="22">S SC MC CC</emphasis>
</td>
</tr>
<tr id="35487C10F7127DB6A8BFD93AD1B4FD82" box="[156,1432,516,580]" gridrow="2" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFD93AD571FD82" box="[156,349,516,580]" gridcol="0" gridrow="2" pageId="21" pageNumber="22">ZMB Por Iquique 3279</th>
<td id="7699156CF7127DB6A9A0D93AD6E4FD82" box="[387,712,516,580]" gridcol="1" gridrow="2" pageId="21" pageNumber="22">Choanosome 1 15 12 2 Ectosome 5 17 8 0</td>
<td id="7699156CF7127DB6AAC7D93AD719FD82" box="[740,821,516,580]" gridcol="2" gridrow="2" pageId="21" pageNumber="22">3 27 10 20</td>
<td id="7699156CF7127DB6AB79D93AD7C8FD82" box="[858,996,516,580]" gridcol="3" gridrow="2" pageId="21" pageNumber="22">3 8 19 0 9 21</td>
<td id="7699156CF7127DB6AC21D93AD0EAFD82" box="[1026,1222,516,580]" gridcol="4" gridrow="2" pageId="21" pageNumber="22">24 2 1 0 3 26 1 0</td>
<td id="7699156CF7127DB6ACC6D93AD1B4FD82" box="[1253,1432,516,580]" gridcol="5" gridrow="2" pageId="21" pageNumber="22">2 2 16 10 0 4 15 11</td>
</tr>
<tr id="35487C10F7127DB6A8BFD947D1B4FD17" box="[156,1432,633,721]" gridrow="3" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFD947D571FD17" box="[156,349,633,721]" gridcol="0" gridrow="3" pageId="21" pageNumber="22">HL 2352 Paracas, Isla San Gallan</th>
<td id="7699156CF7127DB6A9A0D947D6E4FD17" box="[387,712,633,721]" gridcol="1" gridrow="3" pageId="21" pageNumber="22">Choanosome 5 25 0 0 Ectosome 7 23 0 0</td>
<td id="7699156CF7127DB6AAC7D947D719FD17" box="[740,821,633,721]" gridcol="2" gridrow="3" pageId="21" pageNumber="22">9 21 15 15</td>
<td id="7699156CF7127DB6AB79D947D7C8FD17" box="[858,996,633,721]" gridcol="3" gridrow="3" pageId="21" pageNumber="22">1 28 1 0 30 0</td>
<td id="7699156CF7127DB6AC21D947D0EAFD17" box="[1026,1222,633,721]" gridcol="4" gridrow="3" pageId="21" pageNumber="22">20 8 1 0 13 17 0 0</td>
<td id="7699156CF7127DB6ACC6D947D1B4FD17" box="[1253,1432,633,721]" gridcol="5" gridrow="3" pageId="21" pageNumber="22">4 26 0 0 10 20 0 0</td>
</tr>
<tr id="35487C10F7127DB6A8BFD9D7D1B4FD38" box="[156,1432,745,766]" gridrow="4" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFD9D7D571FD38" box="[156,349,745,766]" gridcol="0" gridrow="4" pageId="21" pageNumber="22">HL 2354 Paracas,</th>
<td id="7699156CF7127DB6A9A0D9D7D6E4FD38" box="[387,712,745,766]" gridcol="1" gridrow="4" pageId="21" pageNumber="22">Choanosome 8 22 0 0</td>
<td id="7699156CF7127DB6AAC7D9D7D719FD38" box="[740,821,745,766]" gridcol="2" gridrow="4" pageId="21" pageNumber="22">17 13</td>
<td id="7699156CF7127DB6AB79D9D7D7C8FD38" box="[858,996,745,766]" gridcol="3" gridrow="4" pageId="21" pageNumber="22">4 25 1</td>
<td id="7699156CF7127DB6AC21D9D7D0EAFD38" box="[1026,1222,745,766]" gridcol="4" gridrow="4" pageId="21" pageNumber="22">19 6 1 0</td>
<td id="7699156CF7127DB6ACC6D9D7D1B4FD38" box="[1253,1432,745,766]" gridcol="5" gridrow="4" pageId="21" pageNumber="22">6 22 2 0</td>
</tr>
<tr id="35487C10F7127DB6A8BFD82AD1B4FC8F" box="[156,1432,788,841]" gridrow="5" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFD82AD571FC8F" box="[156,349,788,841]" gridcol="0" gridrow="5" pageId="21" pageNumber="22">Isla San Gallan</th>
<td id="7699156CF7127DB6A9A0D82AD6E4FC8F" box="[387,712,788,841]" gridcol="1" gridrow="5" pageId="21" pageNumber="22">Ectosome 15 15 0 0</td>
<td id="7699156CF7127DB6AAC7D82AD719FC8F" box="[740,821,788,841]" gridcol="2" gridrow="5" pageId="21" pageNumber="22">13 17</td>
<td id="7699156CF7127DB6AB79D82AD7C8FC8F" box="[858,996,788,841]" gridcol="3" gridrow="5" pageId="21" pageNumber="22">3 26 1</td>
<td id="7699156CF7127DB6AC21D82AD0EAFC8F" box="[1026,1222,788,841]" gridcol="4" gridrow="5" pageId="21" pageNumber="22">25 1 1 0</td>
<td id="7699156CF7127DB6ACC6D82AD1B4FC8F" box="[1253,1432,788,841]" gridcol="5" gridrow="5" pageId="21" pageNumber="22">6 24 0 0</td>
</tr>
<tr id="35487C10F7127DB6A8BFD860D1B4FCB5" box="[156,1432,862,883]" gridrow="6" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFD860D571FCB5" box="[156,349,862,883]" gridcol="0" gridrow="6" pageId="21" pageNumber="22">HL 2425 Paracas,</th>
<td id="7699156CF7127DB6A9A0D860D6E4FCB5" box="[387,712,862,883]" gridcol="1" gridrow="6" pageId="21" pageNumber="22">Choanosome 3 26 1 0</td>
<td id="7699156CF7127DB6AAC7D860D719FCB5" box="[740,821,862,883]" gridcol="2" gridrow="6" pageId="21" pageNumber="22">0 30</td>
<td id="7699156CF7127DB6AB79D860D7C8FCB5" box="[858,996,862,883]" gridcol="3" gridrow="6" pageId="21" pageNumber="22">8 22 0</td>
<td id="7699156CF7127DB6AC21D860D0EAFCB5" box="[1026,1222,862,883]" gridcol="4" gridrow="6" pageId="21" pageNumber="22">18 3 1 0</td>
<td id="7699156CF7127DB6ACC6D860D1B4FCB5" box="[1253,1432,862,883]" gridcol="5" gridrow="6" pageId="21" pageNumber="22">3 27 0 0</td>
</tr>
</table>
</paragraph>
<caption id="DF072EDAF712825CA8B4D8C3D7ADFBD3" ID-Table-UUID="DF072EDAF712825CA8B4D8C3D7ADFBD3" box="[151,897,1021,1046]" httpUri="http://table.plazi.org/id/DF072EDAF712825CA8B4D8C3D7ADFBD3" pageId="21" pageNumber="22" targetBox="[156,1397,1202,1542]" targetIsTable="true" targetPageId="21">
<paragraph id="8BC77E52F712825CA8B4D8C3D7ADFBD3" blockId="21.[151,1436,1021,1108]" box="[151,897,1021,1046]" pageId="21" pageNumber="22">
TABLE 6.
<taxonomicName id="4C7805D1F712825CA933D8C0D62EFBD0" box="[272,514,1022,1046]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="21" pageNumber="22" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F712825CA933D8C0D62EFBD0" bold="true" box="[272,514,1022,1046]" italics="true" pageId="21" pageNumber="22">Johannesia reticulosa</emphasis>
</taxonomicName>
: results of spicule measurements.
</paragraph>
</caption>
<paragraph id="8BC77E52F712825CA8B4DF25D1ADFB92" blockId="21.[151,1436,1021,1108]" pageId="21" pageNumber="22">All measurements are given in µm. Each measurement is given as minimum-mean-maximum. Length is given in the upper row; diameter is given in the lower row. Diameter of tylostyles is given as proximal diameter / central diameter.</paragraph>
<paragraph id="8BC77E52F712825CA8BFDF43D7D5F93A" blockId="21.[156,1293,1149,1170]" lastBlockId="21.[151,1436,1602,1788]" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CA8BFDF43D121FB54" bold="true" box="[156,1293,1149,1170]" pageId="21" pageNumber="22">Specimen Region (Sub)tylostyles Oxeas</emphasis>
-Variability of spicule size: Variability of spicule size within each specimen insignificant, with no differences observed between ectosomal and choanosomal spicules (figs. 1416; table 7). Conversely, among-specimens variability of spicule length significant. Ectosomal (sub)tylostyles of the Peruvian specimen HL 2354 longer than those of other specimens (fig. 16A). Both choanosomal and ectosomal oxeas of the Peruvian specimens HL
<date id="FFC65892F712825CA930DDDCD5EDF93A" box="[275,449,1762,1788]" pageId="21" pageNumber="22" value="2352" valueMax="2354">2352 and 2354</date>
longer than those of other specimens (fig. 16C).
</paragraph>
<paragraph id="8BC77E52F712825CAA31DF8CD17BF9C0" pageId="21" pageNumber="22">
<table id="F9788CF2F7127DB6A8BFDF8CD159F9C0" box="[156,1397,1202,1542]" gridcols="5" gridrows="5" pageId="21" pageNumber="22">
<tr id="35487C10F7127DB6A8BFDF8CD159FB01" box="[156,1397,1202,1223]" gridrow="0" pageId="21" pageNumber="22" rowspan-0="1">
<th id="7699156CF7127DB6AA31DF8CD6D1FB01" box="[530,765,1202,1223]" gridcol="1" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CAA31DF8CD6BAFB01" bold="true" box="[530,662,1202,1223]" pageId="21" pageNumber="22">Choanosome</emphasis>
</th>
<th id="7699156CF7127DB6AB13DF8CD03AFB01" box="[816,1046,1202,1223]" gridcol="2" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CAB13DF8CD7BEFB01" bold="true" box="[816,914,1202,1223]" pageId="21" pageNumber="22">Ectosome</emphasis>
</th>
<th id="7699156CF7127DB6AC6EDF8CD0FDFB01" box="[1101,1233,1202,1223]" gridcol="3" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CAC6EDF8CD0FDFB01" bold="true" box="[1101,1233,1202,1223]" pageId="21" pageNumber="22">Choanosome</emphasis>
</th>
<th id="7699156CF7127DB6ACD7DF8CD159FB01" box="[1268,1397,1202,1223]" gridcol="4" gridrow="0" pageId="21" pageNumber="22">
<emphasis id="B90CA240F712825CACD7DF8CD17AFB01" bold="true" box="[1268,1366,1202,1223]" pageId="21" pageNumber="22">Ectosome</emphasis>
</th>
</tr>
<tr id="35487C10F7127DB6A8BFDFDCD159FAD1" box="[156,1397,1250,1303]" gridrow="1" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFDFDCD5C7FAD1" box="[156,491,1250,1303]" gridcol="0" gridrow="1" pageId="21" pageNumber="22">ZMB Por 3279 Iquique</th>
<td id="7699156CF7127DB6AA31DFDCD6D1FAD1" box="[530,765,1250,1303]" gridcol="1" gridrow="1" pageId="21" pageNumber="22">104-270-369 4.2-7.1-11.7 / 5-7.4-11.7</td>
<td id="7699156CF7127DB6AB13DFDCD03AFAD1" box="[816,1046,1250,1303]" gridcol="2" gridrow="1" pageId="21" pageNumber="22">133-240-333 4.2-6.4-9.2 / 4.2-7.2-10</td>
<td id="7699156CF7127DB6AC6EDFDCD0FDFAD1" box="[1101,1233,1250,1303]" gridcol="3" gridrow="1" pageId="21" pageNumber="22">358-446-525 4.2-10.4-13.3</td>
<td id="7699156CF7127DB6ACD7DFDCD159FAD1" box="[1268,1397,1250,1303]" gridcol="4" gridrow="1" pageId="21" pageNumber="22">292-447-550 8.3-11.2-13.3</td>
</tr>
<tr id="35487C10F7127DB6A8BFDE0CD159FAA1" box="[156,1397,1330,1383]" gridrow="2" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFDE0CD5C7FAA1" box="[156,491,1330,1383]" gridcol="0" gridrow="2" pageId="21" pageNumber="22">HL 2352 Paracas, Isla San Gallan</th>
<td id="7699156CF7127DB6AA31DE0CD6D1FAA1" box="[530,765,1330,1383]" gridcol="1" gridrow="2" pageId="21" pageNumber="22">165-249-400 3.5-5.4-10 / 3.5-5.8-10</td>
<td id="7699156CF7127DB6AB13DE0CD03AFAA1" box="[816,1046,1330,1383]" gridcol="2" gridrow="2" pageId="21" pageNumber="22">165-259-330 4-5.6-10 / 4.5-6.7-10</td>
<td id="7699156CF7127DB6AC6EDE0CD0FDFAA1" box="[1101,1233,1330,1383]" gridcol="3" gridrow="2" pageId="21" pageNumber="22">385-533-650 6-13.4-20</td>
<td id="7699156CF7127DB6ACD7DE0CD159FAA1" box="[1268,1397,1330,1383]" gridcol="4" gridrow="2" pageId="21" pageNumber="22">340-526-690 5-13.1-20</td>
</tr>
<tr id="35487C10F7127DB6A8BFDEBCD159FA71" box="[156,1397,1410,1463]" gridrow="3" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFDEBCD5C7FA71" box="[156,491,1410,1463]" gridcol="0" gridrow="3" pageId="21" pageNumber="22">HL 2354 Paracas, Isla San Gallan</th>
<td id="7699156CF7127DB6AA31DEBCD6D1FA71" box="[530,765,1410,1463]" gridcol="1" gridrow="3" pageId="21" pageNumber="22">180-282-375 4-6-9 / 4-6.7-10</td>
<td id="7699156CF7127DB6AB13DEBCD03AFA71" box="[816,1046,1410,1463]" gridcol="2" gridrow="3" pageId="21" pageNumber="22">185-318-395 5-7.1-12.5 / 5-8.1-15</td>
<td id="7699156CF7127DB6AC6EDEBCD0FDFA71" box="[1101,1233,1410,1463]" gridcol="3" gridrow="3" pageId="21" pageNumber="22">355-577-710 5-13.3-19</td>
<td id="7699156CF7127DB6ACD7DEBCD159FA71" box="[1268,1397,1410,1463]" gridcol="4" gridrow="3" pageId="21" pageNumber="22">360-536-680 10-13.8-17.5</td>
</tr>
<tr id="35487C10F7127DB6A8BFDEECD159F9C0" box="[156,1397,1490,1542]" gridrow="4" pageId="21" pageNumber="22">
<th id="7699156CF7127DB6A8BFDEECD5C7F9C0" box="[156,491,1490,1542]" gridcol="0" gridrow="4" pageId="21" pageNumber="22">HL 2425 Paracas, Islotes Dos Marias</th>
<td id="7699156CF7127DB6AA31DEECD6D1F9C0" box="[530,765,1490,1542]" gridcol="1" gridrow="4" pageId="21" pageNumber="22">158-234-320 4-5.1-9 / 4-5.1-9</td>
<td id="7699156CF7127DB6AB13DEECD03AF9C0" box="[816,1046,1490,1542]" gridcol="2" gridrow="4" pageId="21" pageNumber="22">175-254-345 3.5-5.1-7.5 / 3.5-5.1-7.5</td>
<td id="7699156CF7127DB6AC6EDEECD0FDF9C0" box="[1101,1233,1490,1542]" gridcol="3" gridrow="4" pageId="21" pageNumber="22">215-408-520 5-8-10.5</td>
<td id="7699156CF7127DB6ACD7DEECD159F9C0" box="[1268,1397,1490,1542]" gridcol="4" gridrow="4" pageId="21" pageNumber="22">320-437-590 5-9.1-12.5</td>
</tr>
</table>
</paragraph>
<caption id="DF072EDAF711825FA8B4DBA3D66AFF11" pageId="22" pageNumber="23">
<paragraph id="8BC77E52F711825FA8B4DBA3D66AFF11" blockId="22.[151,1413,157,215]" pageId="22" pageNumber="23">
<emphasis id="B90CA240F711825FA8B4DBA3D1A9FF73" bold="true" box="[151,1413,157,181]" pageId="22" pageNumber="23">
TABLE 7.
<taxonomicName id="4C7805D1F711825FA933DBA3D62EFF73" box="[272,514,157,181]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F711825FA933DBA3D62EFF73" bold="true" box="[272,514,157,181]" italics="true" pageId="22" pageNumber="23">Johannesia reticulosa</emphasis>
</taxonomicName>
: ANOVA results for comparison of spicule length between different specimens.
</emphasis>
Significant values are marked with grey.
</paragraph>
</caption>
<paragraph id="8BC77E52F711825FA8BFDBC4D17EFEC9" blockId="22.[156,1389,250,612]" box="[156,1362,250,271]" pageId="22" pageNumber="23">
Spicule
<typeStatus id="54C3C0F0F711825FA8C9DBC4D53FFEC9" box="[234,275,250,271]" pageId="22" pageNumber="23">type</typeStatus>
Factors SS dF MS F P
</paragraph>
<paragraph id="8BC77E52F711825FA8BFDA1BD065FE7C" blockId="22.[156,1389,250,612]" pageId="22" pageNumber="23">
(Sub)tylostyles Specimen
<date id="FFC65892F711825FAB26DA1BD767FEFC" box="[773,843,293,314]" pageId="22" pageNumber="23" value="69-11-15">111569</date>
3 37190 12.30 2*10-7 Part of body 5099 1 5099 1.69 0.195 Specimen+Part of body 36010 3 12003 3.97 0.009 Error 701644 232 3024
</paragraph>
<paragraph id="8BC77E52F711825FA8BFDAF1D065FDA2" blockId="22.[156,1389,250,612]" pageId="22" pageNumber="23">Oxeas Specimen 744742 3 248247 47.29 7*10-24 Part of body 1246 1 1246 0.24 0.627 Specimen+Part of body 37732 3 12577 2.40 0.069 Error 1217818 232 5249</paragraph>
<caption id="DF072EDAF711825FA8B4DCA9D533F828" httpUri="https://zenodo.org/record/180690/files/figure.png" pageId="22" pageNumber="23" targetBox="[277,1336,692,1942]" targetPageId="22">
<paragraph id="8BC77E52F711825FA8B4DCA9D533F828" blockId="22.[151,1436,1943,2030]" pageId="22" pageNumber="23">
<emphasis id="B90CA240F711825FA8B4DCA9D504F869" bold="true" box="[151,296,1943,1967]" pageId="22" pageNumber="23">FIGURE 13.</emphasis>
<taxonomicName id="4C7805D1F711825FA913DCA9D630F868" box="[304,540,1943,1966]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="22" pageNumber="23" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F711825FA913DCA9D630F868" box="[304,540,1943,1966]" italics="true" pageId="22" pageNumber="23">Johannesia reticulosa</emphasis>
</taxonomicName>
, holotype: spicules at SEM. Scale bars for AD: 0.1 mm, scale bars for EF: 0.01 mm. Astraight oxea. Bcurved oxea. Cstraight subtylostyle. Dcurved subtylostyle. Edetail of tyle. Fdistal tip of subtylostyle.
</paragraph>
</caption>
<caption id="DF072EDAF710825EA8B4DD15D6F9F9A2" httpUri="https://zenodo.org/record/180691/files/figure.png" pageId="23" pageNumber="24" targetBox="[278,1309,208,1541]" targetPageId="23">
<paragraph id="8BC77E52F710825EA8B4DD15D6F9F9A2" blockId="23.[151,1436,1579,1636]" pageId="23" pageNumber="24">
<emphasis id="B90CA240F710825EA8B4DD15D505F985" bold="true" box="[151,297,1579,1603]" pageId="23" pageNumber="24">FIGURE 14.</emphasis>
<taxonomicName id="4C7805D1F710825EA910DD15D633F984" box="[307,543,1579,1602]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="23" pageNumber="24" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F710825EA910DD15D633F984" box="[307,543,1579,1602]" italics="true" pageId="23" pageNumber="24">Johannesia reticulosa</emphasis>
</taxonomicName>
: frequency distribution of (sub)tylostyles length. Dotted barschoanosome, black barsectosome. AHolotype. B Peruvian specimens.
</paragraph>
</caption>
<caption id="DF072EDAF71F8251A8B4DD15D643F9A2" httpUri="https://zenodo.org/record/180692/files/figure.png" pageId="24" pageNumber="25" targetBox="[274,1313,205,1544]" targetPageId="24">
<paragraph id="8BC77E52F71F8251A8B4DD15D643F9A2" blockId="24.[151,1436,1579,1636]" pageId="24" pageNumber="25">
<emphasis id="B90CA240F71F8251A8B4DD15D50BF985" bold="true" box="[151,295,1579,1603]" pageId="24" pageNumber="25">FIGURE 15.</emphasis>
<taxonomicName id="4C7805D1F71F8251A90DDD15D635F984" box="[302,537,1579,1602]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="24" pageNumber="25" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F71F8251A90DDD15D635F984" box="[302,537,1579,1602]" italics="true" pageId="24" pageNumber="25">Johannesia reticulosa</emphasis>
</taxonomicName>
: frequency distribution of oxeas length. Dotted barschoanosome, black barsectosome. AHolotype. B Peruvian specimens.
</paragraph>
</caption>
<caption id="DF072EDAF71E8250A8B4DE8CD12BF9C9" httpUri="https://zenodo.org/record/180693/files/figure.png" pageId="25" pageNumber="26" targetBox="[293,1294,209,1420]" targetPageId="25">
<paragraph id="8BC77E52F71E8250A8B4DE8CD12BF9C9" blockId="25.[151,1436,1458,1551]" pageId="25" pageNumber="26">
<emphasis id="B90CA240F71E8250A8B4DE8CD50AFA0C" bold="true" box="[151,294,1458,1482]" pageId="25" pageNumber="26">FIGURE 16.</emphasis>
<taxonomicName id="4C7805D1F71E8250A90FDE8CD63AFA0F" box="[300,534,1458,1481]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="25" pageNumber="26" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F71E8250A90FDE8CD63AFA0F" box="[300,534,1458,1481]" italics="true" pageId="25" pageNumber="26">Johannesia reticulosa</emphasis>
</taxonomicName>
: variability of spicule size. Means and 95% confidence intervals are indicated. Specimens: 1holotype, 2HL 2352, 3HL 2354, 4HL 2425. Alength of (sub)tylostyles. Bcentral diameter of (sub)tylostyles. Clength of oxeas. Dcentral diameter of oxeas. Blank bars choanosome, dotted bars ectosome.
</paragraph>
</caption>
<caption id="DF072EDAF71D8253A8B4DD77D503F945" httpUri="https://zenodo.org/record/180694/files/figure.png" pageId="26" pageNumber="27" targetBox="[275,1308,205,1571]" targetPageId="26">
<paragraph id="8BC77E52F71D8253A8B4DD77D503F945" blockId="26.[151,1435,1609,1667]" pageId="26" pageNumber="27">
<emphasis id="B90CA240F71D8253A8B4DD77D50BF9A7" bold="true" box="[151,295,1609,1633]" pageId="26" pageNumber="27">FIGURE 17.</emphasis>
<taxonomicName id="4C7805D1F71D8253A90EDD77D634F9A6" box="[301,536,1609,1632]" class="Demospongiae" family="Halichondriidae" genus="Johannesia" kingdom="Animalia" order="Halichondrida" pageId="26" pageNumber="27" phylum="Porifera" rank="species" species="reticulosa">
<emphasis id="B90CA240F71D8253A90EDD77D634F9A6" box="[301,536,1609,1632]" italics="true" pageId="26" pageNumber="27">Johannesia reticulosa</emphasis>
</taxonomicName>
: geographical distribution. 1holotype from ZMB. 2specimens from the collection of H. Lehnert.
</paragraph>
</caption>
<paragraph id="8BC77E52F71D8253A8B4DC38D501F8E6" blockId="26.[151,1094,1798,1864]" box="[151,301,1798,1824]" pageId="26" pageNumber="27">Distribution:</paragraph>
<paragraph id="8BC77E52F71D8253A8D6DC10D06AF88E" blockId="26.[151,1094,1798,1864]" box="[245,1094,1838,1864]" pageId="26" pageNumber="27">
Chilean and Peruvian Coast, south-east Pacific (fig. 17), depth
<quantity id="4C80D3B7F71D8253ABF4DC10D06EF88E" box="[983,1090,1838,1864]" metricMagnitude="1" metricUnit="m" metricValue="2.1" metricValueMax="3.0" metricValueMin="1.2" pageId="26" pageNumber="27" unit="m" value="21.0" valueMax="30.0" valueMin="12.0">
12
<specimenCount id="9D7EB5DBF71D8253AC20DC10D06EF88E" box="[1027,1090,1838,1864]" pageId="26" pageNumber="27" type="generic">30 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>