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<mods:title id="DBED1EBC510B88104A47F670320B77BC">Taxonomic revision of the genus Ectamenognonus Buysson (Coleoptera, Elateridae, Elaterinae) from Japan</mods:title>
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<treatment id="03D1A92AFFADFFCC3085105AFA51EE03" ID-DOI="http://doi.org/10.5281/zenodo.5824226" ID-GBIF-Taxon="191928805" ID-Zenodo-Dep="5824226" LSID="urn:lsid:plazi:treatment:03D1A92AFFADFFCC3085105AFA51EE03" httpUri="http://treatment.plazi.org/id/03D1A92AFFADFFCC3085105AFA51EE03" lastPageId="3" lastPageNumber="309" pageId="2" pageNumber="308">
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Key to species of the genus
<taxonomicName id="4C7863BFFFADFFCD31FE105AFD7DEB31" authorityName="Buysson" authorityYear="1893" box="[492,688,1883,1909]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="308" phylum="Arthropoda" rank="genus">
<emphasis id="B90CC42EFFADFFCD31FE105AFD7DEB31" bold="true" box="[492,688,1883,1909]" italics="true" pageId="2" pageNumber="308">Ectamenogonus</emphasis>
</taxonomicName>
from
<collectingCountry id="F36F58ACFFADFFCD32E9105AFC85EB31" box="[763,840,1883,1909]" name="Japan" pageId="2" pageNumber="308">Japan</collectingCountry>
</emphasis>
</heading>
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<subSubSection id="C3624BB7FFADFFCC308510A1FA51EE03" lastPageId="3" lastPageNumber="309" pageId="2" pageNumber="308" type="key">
<keyStep id="308C0099FFADFFCC308510A1FA51ECEA" lastPageId="3" lastPageNumber="309" pageId="2" pageNumber="308">
<paragraph id="8BC7183CFFADFFCD308510A1FA51EB96" blockId="2.[151,1436,1952,2032]" pageId="2" pageNumber="308">
<keyLead id="3089BB09FFADFFCD308510A1FA51EB96" pageId="2" pageNumber="308">
1. Body slender (EI:&gt;2.3, BI:&gt;1.9) (
<figureCitation id="134304B9FFADFFCD323F10A1FDB4EBF2" box="[557,633,1952,1974]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="2" pageNumber="308">Fig. 1A</figureCitation>
). Spiculum ventrale 3.64.2 x longer than sternite VIII. Ovipositor 1.21.3 x longer than length of midline of abdominal ventrites............................................................... 2
</keyLead>
</paragraph>
<paragraph id="8BC7183CFFADFFCC308510DAFA51ECEA" blockId="2.[151,1436,1952,2032]" lastBlockId="3.[151,1437,151,584]" lastPageId="3" lastPageNumber="309" pageId="2" pageNumber="308">
<keyLead id="3089BB09FFADFFCC308510DAFA51ECEA" lastPageId="3" lastPageNumber="309" pageId="2" pageNumber="308">
- Body robust (EI: &lt;2.3, BI: &lt;1.9) (
<figureCitation id="134304B9FFADFFCD323810DBFD58EBB4" box="[554,661,2010,2032]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="2" pageNumber="308">Fig. 1BG</figureCitation>
). Spiculum ventrale 2.5 x longer than sternite VIII. Ovipositor 0.9 x longer than length of midline of abdominal ventrites................................................
<taxonomicName id="4C7863BFFFACFFCC34B41799FA51ECEA" authority="(Kishii, 1973)" baseAuthorityName="Kishii" baseAuthorityYear="1973" box="[1190,1436,151,174]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="309" phylum="Arthropoda" rank="species" species="matobai">
<emphasis id="B90CC42EFFACFFCC34B41799FB78ECE9" box="[1190,1205,152,173]" italics="true" pageId="3" pageNumber="309">E</emphasis>
.
<emphasis id="B90CC42EFFACFFCC34D31796FADDECE8" box="[1217,1296,151,172]" italics="true" pageId="3" pageNumber="309">matobai</emphasis>
(
<bibRefCitation id="EFE965CDFFACFFCC350E1796FA58ECEA" author="Kishii, T." box="[1308,1429,151,174]" pageId="3" pageNumber="309" pagination="1 - 24" refId="ref17825" refString="Kishii, T. (1973) On the elaterid-beetles from Ryukyu Archipelago collected by Mr. Iso Matoba &quot; The snappers of Island (VI) &quot;. Bulletin of the Heian High School, 17, 1 - 24, 4 pls., tab. https: // doi. org / 10.5962 / bhl. part. 3500" type="journal article" year="1973">Kishii, 1973</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308C0099FFACFFCC308517B4FA51EDDC" pageId="3" pageNumber="309">
<paragraph id="8BC7183CFFACFFCC308517B4FA51ED66" blockId="3.[151,1437,151,584]" pageId="3" pageNumber="309">
<keyLead id="3089BB09FFACFFCC308517B4FA51ED66" pageId="3" pageNumber="309">
2. Antennomere IV 1.11.5 x longer than IIIII combined (
<figureCitation id="134304B9FFACFFCC330D17B4FCBAEC8F" box="[799,887,181,203]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="3" pageNumber="309">Figs. 5D</figureCitation>
,
<figureCitation id="134304B9FFACFFCC339717B4FC6EEC8F" box="[901,931,181,203]" captionStart="FIGURE 8" captionStartId="13.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure-30@13.[151,1436,181,1748]" captionTargetPageId="13" captionText="FIGURE 8. Ectamenogonus plebejus (Candèze, 1873), male. AF, HJ: [EPM01]; G: [KEP17]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum; H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edges of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824248" httpUri="https://zenodo.org/record/5824248/files/figure.png" pageId="3" pageNumber="309">8D</figureCitation>
). Prothorax somewhat slender (PI: 109121) (
<figureCitation id="134304B9FFACFFCC356717B4FED0ECAC" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="3" pageNumber="309">Fig. 1BE</figureCitation>
). Aedeagus slender (3.64.3 times longer than wide) (
<figureCitation id="134304B9FFACFFCC333817D3FC5BECAC" box="[810,918,210,232]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="3" pageNumber="309">Figs. 6E, F</figureCitation>
,
<figureCitation id="134304B9FFACFFCC33B217D3FC2FECAC" box="[928,994,210,232]" captionStart="FIGURE 11" captionStartId="16.[151,250,1453,1477]" captionTargetBox="[199,1430,489,1414]" captionTargetId="figure-172@16.[151,1436,489,1429]" captionTargetPageId="16" captionText="FIGURE 11. Ectamenogonus plebejus (Candèze, 1873), male [EPM03]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of lateral lobe of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824258" httpUri="https://zenodo.org/record/5824258/files/figure.png" pageId="3" pageNumber="309">11E, F</figureCitation>
). Median lobe exceeding apices of parameres by apical 1/4 to 1/3 (
<figureCitation id="134304B9FFACFFCC31BE17EEFDD4ED41" box="[428,537,239,261]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="3" pageNumber="309">Figs. 6E, F</figureCitation>
,
<figureCitation id="134304B9FFACFFCC323117F1FDAEED41" box="[547,611,240,261]" captionStart="FIGURE 11" captionStartId="16.[151,250,1453,1477]" captionTargetBox="[199,1430,489,1414]" captionTargetId="figure-172@16.[151,1436,489,1429]" captionTargetPageId="16" captionText="FIGURE 11. Ectamenogonus plebejus (Candèze, 1873), male [EPM03]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of lateral lobe of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824258" httpUri="https://zenodo.org/record/5824258/files/figure.png" pageId="3" pageNumber="309">11E, F</figureCitation>
). Paramere apex length 1.92.2 x width of parameres at expansions (
<figureCitation id="134304B9FFACFFCC351717EEFA96ED41" box="[1285,1371,239,261]" captionStart="FIGURE 6" captionStartId="9.[151,250,1145,1169]" captionTargetBox="[151,1436,181,1121]" captionTargetId="figure-30@9.[151,1436,181,1121]" captionTargetPageId="9" captionText="FIGURE 6. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of paramera of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824244" httpUri="https://zenodo.org/record/5824244/files/figure.png" pageId="3" pageNumber="309">Figs. 6G</figureCitation>
,
<figureCitation id="134304B9FFACFFCC357417F1FA42ED42" box="[1382,1423,240,262]" captionStart="FIGURE 11" captionStartId="16.[151,250,1453,1477]" captionTargetBox="[199,1430,489,1414]" captionTargetId="figure-172@16.[151,1436,489,1429]" captionTargetPageId="16" captionText="FIGURE 11. Ectamenogonus plebejus (Candèze, 1873), male [EPM03]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of lateral lobe of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824258" httpUri="https://zenodo.org/record/5824258/files/figure.png" pageId="3" pageNumber="309">11G</figureCitation>
).
<collectingRegion id="49BCD6DEFFACFFCC30F0160CFED7ED66" box="[226,282,269,290]" country="Turkey" name="Bursa" pageId="3" pageNumber="309">Bursa</collectingRegion>
copulatrix long (1.72.7 times length of uterus), with two clusters of sclerotized spines (
<figureCitation id="134304B9FFACFFCC347D160CFB09ED66" box="[1135,1220,269,291]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.AC, FH: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="3" pageNumber="309">Figs. 7A</figureCitation>
,
<figureCitation id="134304B9FFACFFCC34DC160CFB35ED67" box="[1230,1272,269,291]" captionStart="FIGURE 13" captionStartId="18.[151,250,1658,1682]" captionTargetBox="[151,1436,525,1634]" captionTargetId="figure-219@18.[151,1436,525,1634]" captionTargetPageId="18" captionText="FIGURE 13. Ectamenogonus plebejus (Candèze, 1873), female. AC, FH: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824262" httpUri="https://zenodo.org/record/5824262/files/figure.png" pageId="3" pageNumber="309">13A</figureCitation>
)........... 3
</keyLead>
</paragraph>
<paragraph id="8BC7183CFFACFFCC3085162AFA51EDDC" blockId="3.[151,1437,151,584]" pageId="3" pageNumber="309">
<keyLead id="3089BB09FFACFFCC3085162AFA51EDDC" pageId="3" pageNumber="309">
- Antennomere IV 0.91.1 x longer than IIIII combined (
<figureCitation id="134304B9FFACFFCC331C162BFCABED04" box="[782,870,298,320]" captionStart="FIGURE 14" captionStartId="21.[151,250,1863,1887]" captionTargetBox="[151,1436,273,1833]" captionTargetId="figure-78@21.[151,1436,273,1840]" captionTargetPageId="21" captionText="FIGURE 14. Ectamenogonus robustus (Kishii, 1966), male. A: [ERM07]; BF, J: [ERM01]; G: [KER14]; H, I: [ERM06]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum; H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edges of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824266" httpUri="https://zenodo.org/record/5824266/files/figure.png" pageId="3" pageNumber="309">Fig. 14D</figureCitation>
). Prothorax somewhat robust (PI: 101111) (
<figureCitation id="134304B9FFACFFCC3534162BFA40ED7B" box="[1318,1421,298,320]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="3" pageNumber="309">Fig. 1F, G</figureCitation>
). Aedeagus robust (3.3 times longer than wide) (
<figureCitation id="134304B9FFACFFCC32B91646FCD0ED19" box="[683,797,327,349]" captionStart="FIGURE 16" captionStartId="23.[151,250,1324,1348]" captionTargetBox="[211,1420,453,1286]" captionTargetId="figure-164@23.[161,1426,453,1301]" captionTargetPageId="23" captionText="FIGURE 16. Ectamenogonus robustus (Kishii, 1966), male [EPM03]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of lateral lobe of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824270" httpUri="https://zenodo.org/record/5824270/files/figure.png" pageId="3" pageNumber="309">Fig. 16E, F</figureCitation>
). Median lobe exceeding apices of parameres by apical 1/5 to 1/4 (
<figureCitation id="134304B9FFACFFCC30FB1664FE94ED3E" box="[233,345,357,379]" captionStart="FIGURE 16" captionStartId="23.[151,250,1324,1348]" captionTargetBox="[211,1420,453,1286]" captionTargetId="figure-164@23.[161,1426,453,1301]" captionTargetPageId="23" captionText="FIGURE 16. Ectamenogonus robustus (Kishii, 1966), male [EPM03]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of lateral lobe of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824270" httpUri="https://zenodo.org/record/5824270/files/figure.png" pageId="3" pageNumber="309">Fig. 16E, F</figureCitation>
). Paramere apex length 1.31.8 x width of parameres at expansions (
<figureCitation id="134304B9FFACFFCC33ED1664FB95ED3F" box="[1023,1112,357,379]" captionStart="FIGURE 16" captionStartId="23.[151,250,1324,1348]" captionTargetBox="[211,1420,453,1286]" captionTargetId="figure-164@23.[161,1426,453,1301]" captionTargetPageId="23" captionText="FIGURE 16. Ectamenogonus robustus (Kishii, 1966), male [EPM03]. A: tergite VIII; B: sternite VIII; C: tergites IXX; D: sternite IX; E: aedeagus, dorsal view; F: aedeagus, ventral view; G: apical part of lateral lobe of aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.5824270" httpUri="https://zenodo.org/record/5824270/files/figure.png" pageId="3" pageNumber="309">Fig. 16G</figureCitation>
).
<collectingRegion id="49BCD6DEFFACFFCC347E1664FB69ED3E" box="[1132,1188,357,378]" country="Turkey" name="Bursa" pageId="3" pageNumber="309">Bursa</collectingRegion>
copulatrix short (1.01.1 times length of uterus), with a cluster of sclerotized spines (
<figureCitation id="134304B9FFACFFCC330D1683FCBEEDDC" box="[799,883,386,408]" captionStart="FIGURE 18" captionStartId="24.[151,250,1314,1338]" captionTargetBox="[151,1436,181,1290]" captionTargetId="figure-16@24.[151,1436,181,1290]" captionTargetPageId="24" captionText="FIGURE 18. Ectamenogonus robustus (Kishii, 1966), female. A, CG: [ERF09]; B: [ERF15], H: [ERF01]. A: genitalia; B: genitalia; C: tergite VIII, dorsal view; D: sternite VIII and tergite VIII, ventral view; E: sclerotized spines cluster in bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824274" httpUri="https://zenodo.org/record/5824274/files/figure.png" pageId="3" pageNumber="309">Fig. 18E</figureCitation>
)........................
<taxonomicName id="4C7863BFFFACFFCC34B11682FA51EDDC" authority="(Kishii, 1966)" baseAuthorityName="Kishii" baseAuthorityYear="1966" box="[1187,1436,386,408]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="309" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B90CC42EFFACFFCC34B11682FB7FEDDC" box="[1187,1202,387,408]" italics="true" pageId="3" pageNumber="309">E</emphasis>
.
<emphasis id="B90CC42EFFACFFCC34AC1683FADDEDD3" box="[1214,1296,386,407]" italics="true" pageId="3" pageNumber="309">robustus</emphasis>
(
<bibRefCitation id="EFE965CDFFACFFCC350E1683FA58EDDC" author="Kishii, T." box="[1308,1429,386,408]" pageId="3" pageNumber="309" refId="ref17798" refString="Kishii, T. (1966) Elateridae of Kyoto and its adjacent regions. Biological Laboratory, Heian High School, Kyoto, 54 pp." type="book" year="1966">Kishii, 1966</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="308C0099FFACFFCC308516A1FA51EE03" pageId="3" pageNumber="309">
<paragraph id="8BC7183CFFACFFCC308516A1FA51EDB4" blockId="3.[151,1437,151,584]" pageId="3" pageNumber="309">
<keyLead id="3089BB09FFACFFCC308516A1FA51EDB4" pageId="3" pageNumber="309">
3. Antennae just reaching or extending beyond pronotum posterior lateral apices by apical antennomere in male (
<figureCitation id="134304B9FFACFFCC350B169EFA5FEDF2" box="[1305,1426,415,438]" captionStart="FIGURE 10" captionStartId="15.[151,250,753,777]" captionTargetBox="[151,1433,185,729]" captionTargetId="figure-30@15.[151,1436,181,729]" captionTargetPageId="15" captionText="FIGURE 10. Ectamenogonus spp., right antenna. A: E. miyakoensis, holotype, male [EMM01]; B: E. plebejus [GN. 2124]; C: E. plebejus [KEP20]; D: E. yakuensis, holotype, male (= E. plebejus)." figureDoi="http://doi.org/10.5281/zenodo.5824254" httpUri="https://zenodo.org/record/5824254/files/figure.png" pageId="3" pageNumber="309">Fig. 10BD</figureCitation>
). Posterior edge of scutellar shield rounded but truncate in some (
<figureCitation id="134304B9FFACFFCC332D16BCFC47ED97" box="[831,906,445,467]" captionStart="FIGURE 8" captionStartId="13.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1741]" captionTargetId="figure-30@13.[151,1436,181,1748]" captionTargetPageId="13" captionText="FIGURE 8. Ectamenogonus plebejus (Candèze, 1873), male. AF, HJ: [EPM01]; G: [KEP17]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum; H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edges of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824248" httpUri="https://zenodo.org/record/5824248/files/figure.png" pageId="3" pageNumber="309">Fig. 8G</figureCitation>
). Elytra 2.02.2 times longer than pronotum length (BI: 202229) (
<figureCitation id="134304B9FFACFFCC315916DBFE7EEDB4" box="[331,435,474,496]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="3" pageNumber="309">Fig. 1D, E</figureCitation>
).
<collectingRegion id="49BCD6DEFFACFFCC31D616DAFE31EDB4" box="[452,508,475,496]" country="Turkey" name="Bursa" pageId="3" pageNumber="309">Bursa</collectingRegion>
copulatrix shorter (1.72.3 times length of uterus) (
<figureCitation id="134304B9FFACFFCC33FD16DBFB8BEDB4" box="[1007,1094,474,496]" captionStart="FIGURE 13" captionStartId="18.[151,250,1658,1682]" captionTargetBox="[151,1436,525,1634]" captionTargetId="figure-219@18.[151,1436,525,1634]" captionTargetPageId="18" captionText="FIGURE 13. Ectamenogonus plebejus (Candèze, 1873), female. AC, FH: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824262" httpUri="https://zenodo.org/record/5824262/files/figure.png" pageId="3" pageNumber="309">Fig. 13A</figureCitation>
).....
<taxonomicName id="4C7863BFFFACFFCC349F16DAFA51EDB4" authority="(Candeze, 1873)" baseAuthorityName="Candeze" baseAuthorityYear="1873" box="[1165,1436,474,496]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="309" phylum="Arthropoda" rank="species" species="plebejus">
<emphasis id="B90CC42EFFACFFCC349F16DAFB51EDB4" box="[1165,1180,475,496]" italics="true" pageId="3" pageNumber="309">E</emphasis>
.
<emphasis id="B90CC42EFFACFFCC34BA16DBFB35EDAB" box="[1192,1272,474,495]" italics="true" pageId="3" pageNumber="309">plebejus</emphasis>
(
<bibRefCitation id="EFE965CDFFACFFCC351616DBFA58EDB4" author="Candeze, E. C. A." box="[1284,1429,474,496]" pageId="3" pageNumber="309" pagination="1 - 32" refId="ref17336" refString="Candeze, E. C. A. (1873) Insects recueillis au Japon par Mr. G. Lewis. Elaterides. Memoires de la Societe Royale des Sciences de Liege, Series 2, 5, 1 - 32." type="journal article" year="1873">Candèze, 1873</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC7183CFFACFFCC308516F9FA51EE03" blockId="3.[151,1437,151,584]" pageId="3" pageNumber="309">
<keyLead id="3089BB09FFACFFCC308516F9FA51EE03" pageId="3" pageNumber="309">
- Antennae not reaching pronotum posterior lateral apices by apical antennomere in male (
<figureCitation id="134304B9FFACFFCC342516F6FB5CEE4A" box="[1079,1169,503,526]" captionStart="FIGURE 10" captionStartId="15.[151,250,753,777]" captionTargetBox="[151,1433,185,729]" captionTargetId="figure-30@15.[151,1436,181,729]" captionTargetPageId="15" captionText="FIGURE 10. Ectamenogonus spp., right antenna. A: E. miyakoensis, holotype, male [EMM01]; B: E. plebejus [GN. 2124]; C: E. plebejus [KEP20]; D: E. yakuensis, holotype, male (= E. plebejus)." figureDoi="http://doi.org/10.5281/zenodo.5824254" httpUri="https://zenodo.org/record/5824254/files/figure.png" pageId="3" pageNumber="309">Fig. 10A</figureCitation>
). Posterior edge of scutellar shield emarginate medially (
<figureCitation id="134304B9FFACFFCC31E51514FD8BEE6F" box="[503,582,533,555]" captionStart="FIGURE 5" captionStartId="8.[151,250,1772,1796]" captionTargetBox="[151,1436,181,1740]" captionTargetId="figure-16@8.[151,1436,181,1748]" captionTargetPageId="8" captionText="FIGURE 5. Ectamenogonus miyakoensis, holotype, male [EMM01]. A: habitus, ventral view; B: head, anterior view; C: head, ventral view; D: basal parts of right antenna and head, laterodorsal view; E: apical parts of right antenna; F: head, prothorax and mesothorax, lateral view; G: scutellum (arrow: posterior edge of scutellum shallowly emarginate medially); H: mesothorax and metasternum, lateroventral view (white arrow: hypomeron slightly grooved next to impunctate ridge); I: border between mesosternum and metasternum, ventral view (arrow: posterior edge of mesosternum triangularly emarginate); J: posterior edge of elytra." figureDoi="http://doi.org/10.5281/zenodo.5824242" httpUri="https://zenodo.org/record/5824242/files/figure.png" pageId="3" pageNumber="309">Fig. 5G</figureCitation>
: arrow). Elytra 1.92.0 times longer than pronotum length (BI: 191199) (
<figureCitation id="134304B9FFACFFCC35341514FA5CEE6F" box="[1318,1425,533,555]" captionStart="FIGURE 1" captionStartId="4.[151,250,1852,1876]" captionTargetBox="[166,1431,357,1814]" captionTargetId="figure-97@4.[151,1436,345,1829]" captionTargetPageId="4" captionText="FIGURE 1. Ectamenogonus spp., habitus, dorsal view. A: E. matobai, holotype, female; B: E. miyakoensis, holotype, male [EMM01]; C: E. miyakoensis, paratype, female [EMF01]; D; E. plebejus, male [EPM01]; E: E. plebejus, female [EPF02]; F: E. robustus, male [ERM01]; G: E. robustus [ERF11]." figureDoi="http://doi.org/10.5281/zenodo.5824234" httpUri="https://zenodo.org/record/5824234/files/figure.png" pageId="3" pageNumber="309">Fig. 1B, C</figureCitation>
).
<collectingRegion id="49BCD6DEFFACFFCC30F01532FED7EE0C" box="[226,282,563,584]" country="Turkey" name="Bursa" pageId="3" pageNumber="309">Bursa</collectingRegion>
copulatrix longer (2.42.7 times length of uterus) (
<figureCitation id="134304B9FFACFFCC331A1533FC9EEE0C" box="[776,851,562,584]" captionStart="FIGURE 7" captionStartId="10.[151,250,1694,1718]" captionTargetBox="[151,1436,561,1670]" captionTargetId="figure-193@10.[151,1436,561,1670]" captionTargetPageId="10" captionText="FIGURE 7. Ectamenogonus miyakoensis, female, paratype.AC, FH: [EMF01]; D, E: [EMF02]. A: genitalia; B: tergite VIII, dorsal view; C: sternite VIII; and tergite VIII, ventral view; D: sclerotized spines cluster in first half of bursa copulatrix; E: sclerotized spines rows in latter half of bursa copulatrix; F, G: long thin sac extending from apex of bursa copulatrix; H: apical part of ovipositor, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5824246" httpUri="https://zenodo.org/record/5824246/files/figure.png" pageId="3" pageNumber="309">Fig. 7A</figureCitation>
)........................
<taxonomicName id="4C7863BFFFACFFCC34681532FAADEE0C" authority="Arimoto" authorityName="Arimoto" authorityYear="2022" box="[1146,1376,562,584]" class="Insecta" family="Elateridae" genus="Ectamenogonus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="309" phylum="Arthropoda" rank="species" species="miyakoensis" status="sp. nov.">
<emphasis id="B90CC42EFFACFFCC34681532FB44EE0C" box="[1146,1161,563,584]" italics="true" pageId="3" pageNumber="309">E</emphasis>
.
<emphasis id="B90CC42EFFACFFCC34871533FAC7EE03" box="[1173,1290,562,583]" italics="true" pageId="3" pageNumber="309">miyakoensis</emphasis>
Arimoto
</taxonomicName>
<emphasis id="B90CC42EFFACFFCC35741533FA51EE03" bold="true" box="[1382,1436,562,583]" pageId="3" pageNumber="309">
<taxonomicNameLabel id="A23F7955FFACFFCC35741533FA51EE03" box="[1382,1436,562,583]" pageId="3" pageNumber="309" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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</subSubSection>
</treatment>
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