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<document id="24992469725D9237E5107BF18812B9E2" ID-DOI="10.5281/zenodo.184583" ID-GBIF-Dataset="f71e3c5c-bb32-48db-be4a-6c7c4b24307e" ID-ISSN="1175-5326" ID-Zenodo-Dep="184583" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459989436065" checkinUser="plazi" docAuthor="Hong, Yong" docDate="2008" docId="03B7878EFFD3FFACFF02498166692A8C" docLanguage="en" docName="zt01914p061.pdf" docOrigin="Zootaxa 1914" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Amynthas namphouinensis Hong, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FF8EFFF6FFD2FFAEFF954916646D2C19" masterDocTitle="Two new bithecal earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Laos" masterLastPageNumber="61" masterPageNumber="57" pageNumber="58" updateTime="1698232534732" updateUser="plazi">
<mods:mods id="5ED0DA72EA2295C1E30F3CD47EE17BCB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6C6B4D478956EF32B664C7D6DA27EAD0">
<mods:title id="BFF7C093C408FAFCBC5F413F974F30BB">Two new bithecal earthworms of the genus Amynthas (Oligochaeta: Megascolecidae) from Laos</mods:title>
</mods:titleInfo>
<mods:name id="63B1C8858EB416272E1F473B6267A747" type="personal">
<mods:role id="1947E0177905DDA9CC70C8CA2FCB6DC8">
<mods:roleTerm id="001298566F0AE68F0401323AA52AD284">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BED680AFF480CA4EFEADBE4376287F46">Hong, Yong</mods:namePart>
</mods:name>
<mods:typeOfResource id="F7711898B914ACD533D1AF44583806D6">text</mods:typeOfResource>
<mods:relatedItem id="7C07133197DBB4618EE470515863A776" type="host">
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<mods:title id="34B1A126C2A9348E25E7E3399528602B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="33D5AAADC02394F826E73A42F4E7EAB7">
<mods:date id="591678981BDD26E3A23E5F2228809692">2008</mods:date>
<mods:detail id="3BDD01752941FEADD600CBD9F340DA1C" type="volume">
<mods:number id="89B2AD26C0E969DBB9F763ED89BAB227">1914</mods:number>
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<mods:classification id="D02D3B5E814B106D2B9A23900F3919ED">journal article</mods:classification>
<mods:identifier id="F6B9128B255365926850F2A7A3F8B698" type="DOI">10.5281/zenodo.184583</mods:identifier>
<mods:identifier id="7BEB7FAA078E9531FA5D081B0C640702" type="GBIF-Dataset">f71e3c5c-bb32-48db-be4a-6c7c4b24307e</mods:identifier>
<mods:identifier id="DB0A363F0D8890722A24F32777DAF27F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="45FFDD59276FA144D15687258D7846E8" type="Zenodo-Dep">184583</mods:identifier>
</mods:mods>
<treatment id="03B7878EFFD3FFACFF02498166692A8C" ID-DOI="http://doi.org/10.5281/zenodo.6235536" ID-GBIF-Taxon="119379813" ID-Zenodo-Dep="6235536" LSID="urn:lsid:plazi:treatment:03B7878EFFD3FFACFF02498166692A8C" httpUri="http://treatment.plazi.org/id/03B7878EFFD3FFACFF02498166692A8C" lastPageId="2" lastPageNumber="58" pageId="1" pageNumber="58">
<subSubSection id="C3046513FFD3FFAFFF024981654B2CCD" pageId="1" pageNumber="58" type="nomenclature">
<paragraph id="8BA13698FFD3FFAFFF02498166282CA8" blockId="1.[151,581,151,212]" box="[151,581,151,177]" pageId="1" pageNumber="58">
<heading id="D0E981F4FFD3FFAFFF02498166282CA8" bold="true" box="[151,581,151,177]" fontSize="11" level="1" pageId="1" pageNumber="58" reason="1">
<emphasis id="B96AEA8AFFD3FFAFFF02498166282CA8" bold="true" box="[151,581,151,177]" pageId="1" pageNumber="58">
<taxonomicName id="4C1E4D1BFFD3FFAFFF024981658D2CA8" box="[151,480,151,177]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="namphouinensis" status="sp. nov.">
<emphasis id="B96AEA8AFFD3FFAFFF024981658D2CA8" bold="true" box="[151,480,151,177]" italics="true" pageId="1" pageNumber="58">Amynthas namphouinensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25957F1FFD3FFAFFE7D498166282CA8" box="[488,581,151,177]" pageId="1" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA13698FFD3FFAFFF0249AC654B2CCD" blockId="1.[151,581,151,212]" box="[151,294,186,212]" pageId="1" pageNumber="58">
(
<figureCitation id="13252A1DFFD3FFAFFF0A49AC648D2CCD" box="[159,224,186,212]" captionStart="FIGURE 1" captionStartId="2.[151,255,1328,1352]" captionTargetBox="[584,1003,189,1307]" captionTargetId="figure@2.[576,1014,184,1313]" captionTargetPageId="2" captionText="FIGURE 1. Amynthas namphouinensis sp. nov. A: ventral view; B: spermathecae. Scale bars = 2.5 mm (A), 1 mm (B)." httpUri="https://zenodo.org/record/184584/files/figure.png" pageId="1" pageNumber="58">Fig. 1</figureCitation>
A–B)
</paragraph>
</subSubSection>
<subSubSection id="C3046513FFD3FFAFFF02481C60D72D86" pageId="1" pageNumber="58" type="materials_examined">
<paragraph id="8BA13698FFD3FFAFFF02481C60D72D86" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">
<emphasis id="B96AEA8AFFD3FFAFFF02481C65662D3D" bold="true" box="[151,267,266,292]" pageId="1" pageNumber="58">Material.</emphasis>
<typeStatus id="54A5883AFFD3FFAFFE80481C65EA2D3D" box="[277,391,266,292]" pageId="1" pageNumber="58" type="holotype">Holotype</typeStatus>
: clitellate (
<collectionCode id="ED0FAE5DFFD3FFAFFD9E481C661F2D3D" box="[523,626,266,292]" pageId="1" pageNumber="58">BDNUL</collectionCode>
0001):
<collectingCountry id="F3097608FFD3FFAFFD47481C67632D3D" box="[722,782,266,292]" name="Laos" pageId="1" pageNumber="58">Laos</collectingCountry>
, Xayabouli Province, Nam Phouin
<collectionCode id="ED0FAE5DFFD3FFAFFB2F481C61632D3D" box="[1210,1294,266,292]" pageId="1" pageNumber="58">NBCA</collectionCode>
, Phou vang Phan (
<geoCoordinate id="EE2A505FFFD3FFAFFF74482365142D56" box="[225,377,306,335]" direction="north" orientation="latitude" pageId="1" pageNumber="58" precision="1" value="18.497967">18o 29.878’N</geoCoordinate>
,
<geoCoordinate id="EE2A505FFFD3FFAFFE114823664A2D56" box="[388,551,306,335]" direction="east" orientation="longitude" pageId="1" pageNumber="58" precision="1" value="101.48116">101o 28.870’E</geoCoordinate>
),
<quantity id="4CE69B7DFFD3FFAFFDAE482366EA2D56" box="[571,647,309,335]" metricMagnitude="2" metricUnit="m" metricValue="4.95" pageId="1" pageNumber="58" unit="m" value="495.0">495 m</quantity>
,
<quantity id="4CE69B7DFFD3FFAFFD06482366A32D56" box="[659,718,309,335]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="58" unit="km" value="5.0">5 km</quantity>
from village in old forest, black soil with sand, in soil and litter layers,
<date id="FFA01058FFD3FFAFFE84484B65A32D6E" box="[273,462,349,375]" pageId="1" pageNumber="58" value="2005-08-30">30 August 2005</date>
, K. Inkhavilay coll. 4
<typeStatus id="54A5883AFFD3FFAFFD45484B67282D6E" box="[720,837,349,375]" pageId="1" pageNumber="58" type="paratype">paratypes</typeStatus>
: 2 clitellates (
<collectionCode id="ED0FAE5DFFD3FFAFFC76484B60272D6E" box="[995,1098,349,375]" pageId="1" pageNumber="58">BDNUL</collectionCode>
0 0 0 3, 0004) and 2 clitellates (
<collectionCode id="ED0FAE5DFFD3FFAFFF0A489364862D86" box="[159,235,389,415]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="1" pageNumber="58">NIBR</collectionCode>
), same data as for
<typeStatus id="54A5883AFFD3FFAFFE51489366432D86" box="[452,558,389,415]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
. Other material: 2 clitellates, same data as for
<typeStatus id="54A5883AFFD3FFAFFBD9489360DB2D86" box="[1100,1206,389,415]" pageId="1" pageNumber="58" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C3046513FFD3FFAFFF5048BB67532DDE" box="[197,830,429,455]" pageId="1" pageNumber="58" type="etymology">
<paragraph id="8BA13698FFD3FFAFFF5048BB67532DDE" blockId="1.[151,1437,266,2024]" box="[197,830,429,455]" pageId="1" pageNumber="58">
<emphasis id="B96AEA8AFFD3FFAFFF5048BB653F2DDE" bold="true" box="[197,338,429,455]" pageId="1" pageNumber="58">Etymology.</emphasis>
The species is named for its
<typeStatus id="54A5883AFFD3FFAFFD3248BB66B72DDE" box="[679,730,429,455]" pageId="1" pageNumber="58">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C3046513FFD3FFAFFF5048CE65CE2E73" pageId="1" pageNumber="58" type="diagnosis">
<paragraph id="8BA13698FFD3FFAFFF5048CE65CE2E73" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">
<emphasis id="B96AEA8AFFD3FFAFFF5048CE65282DEB" bold="true" box="[197,325,472,498]" pageId="1" pageNumber="58">Diagnosis.</emphasis>
Spermathecal pores paired in 7/8 at 5th seta lines; male pores superficial in XVIII at 4th seta lines on small round porophores, 0.10–0.14 circumference apart ventrally; genital papillae equatorial in XVII, presetal widely paired in XIX; mid-ventral in 17/18; presetal, closely paired in mid-ventral in XIX; holandric, first dorsal pore in 5/6.
</paragraph>
</subSubSection>
<subSubSection id="C3046513FFD3FFAFFF504B6E67D42A11" pageId="1" pageNumber="58" type="description">
<paragraph id="8BA13698FFD3FFAFFF504B6E67CE2F43" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">
<emphasis id="B96AEA8AFFD3FFAFFF504B6E65302E8B" bold="true" box="[197,349,632,658]" pageId="1" pageNumber="58">Description.</emphasis>
Dimensions
<quantity id="4CE69B7DFFD3FFAFFE6D4B6E661B2E8B" box="[504,630,632,658]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="9.2" metricValueMin="6.3" pageId="1" pageNumber="58" unit="mm" value="77.5" valueMax="92.0" valueMin="63.0">63–92 mm</quantity>
by 3.5–4.0 mm at segment X, 4.0–
<quantity id="4CE69B7DFFD3FFAFFB874B6E60072E8B" box="[1042,1130,632,658]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="58" unit="mm" value="4.1">4.1 mm</quantity>
at segment XXX, 3.0–
<quantity id="4CE69B7DFFD3FFAFFAED4B6E64A82EA3" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="58" unit="mm" value="3.5">3.5 mm</quantity>
at clitellum; body cylindrical, segments 92–94. Setae regularly distributed around segmental equators, numbering 52–61 at VII, 53–58 at XX, 0–7 between male pores, setal formula AA:AB:ZZ:YZ= 1:1:5:3 at XIII. Female pore single in XIV, on an oval area
<quantity id="4CE69B7DFFD3FFAFFD5B4BE667362F13" box="[718,859,752,778]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="1" pageNumber="58" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
in width. Prostomium epilobic with tongue open, clitellum coffee color, dorsal brown pigment fading posteriorly, formalin preservation. First dorsal pore at 4/5, 5/6, or 6/7. Clitellum annular XIV–XVI; setae invisible externally.
</paragraph>
<paragraph id="8BA13698FFD3FFAFFF504A7D61152861" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">
Male pores superficial on small round porophores in XVIII at 4th seta lines, 0.10–0.14 circumference apart ventrally; distance between male pores
<quantity id="4CE69B7DFFD3FFAFFDEF4A8567632FB4" box="[634,782,915,941]" metricMagnitude="-3" metricUnit="m" metricValue="1.45" metricValueMax="1.5" metricValueMin="1.4" pageId="1" pageNumber="58" unit="mm" value="1.45" valueMax="1.5" valueMin="1.4">1.4–1.5 mm</quantity>
; porophores at medial edges of reniform dark genital papillae; small grooves around male pores and longitudinal grooves between male pores. Locations of genital papillae irregular, but always paired, equatorial in XVII, and in some of the following positions: presetal, widely paired in XIX; mid-ventral in 17/18; presetal, closely paired in mid-ventral in XIX; in total usually 5– 7 papillae. Spermathecal pores paired in 7/8 at 5th seta lines, conspicuous in intersegmental furrow, 0.15–0.28 circumference apart ventrally; distance between spermathecal pores 1.4–2.0 mm. Genital markings absent.
</paragraph>
<paragraph id="8BA13698FFD3FFAFFF504D90651F2989" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">
<collectingCountry id="F3097608FFD3FFAFFF504D90656B28B9" box="[197,262,1158,1184]" name="Spain" pageId="1" pageNumber="58">Septa</collectingCountry>
5/6–7/8 thin, 8/9, 9/10 absent,
<date id="FFA01058FFD3FFAFFDE34D90676728B9" box="[630,778,1158,1184]" pageId="1" pageNumber="58" value="1914-10-11" valueMax="1914-10-13" valueMin="1914-10-11">10/11–13/14</date>
thin. Gizzard large between VIII–X. Intestinal origin in XV, lymph glands not found. Typhlosole small from XXVII. Intestinal caeca originating in XXVII, extending anteriorly about to XXIV, simple finger-shaped. Hearts esophageal in X–XIII. Holandric; testes and funnels in ventrally joined sacs in X–XI. Seminal vesicles paired in XI–XII with round dorsal lobe in anterior dorsal notch. Prostates in XVIII; extending between XVII–XIX; glands consisting of 2 main lobes, each divided again into 2–3 small lobes; prostatic ducts slender, muscular. Genital marking glands sessile on body wall within XVII–XIX.
</paragraph>
<paragraph id="8BA13698FFD3FFAFFF504C8867D42A11" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">Ovaries in XIII. Spermathecae one pair in VIII; ampulla thumb-shaped, stout ducts shorter than ampulla, diverticulum chamber spherical, iridescent, with slender stalk shorter than ampulla, stalk attached to duct near body wall, no nephridia on ducts. Genital marking glands not found.</paragraph>
</subSubSection>
<subSubSection id="C3046513FFD3FFACFF504F0066692A8C" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58" type="discussion">
<paragraph id="8BA13698FFD3FFAFFF504F0061592B81" blockId="1.[151,1437,266,2024]" pageId="1" pageNumber="58">
<emphasis id="B96AEA8AFFD3FFAFFF504F0065532A29" bold="true" box="[197,318,1558,1584]" pageId="1" pageNumber="58">Remarks.</emphasis>
<taxonomicName id="4C1E4D1BFFD3FFAFFED24F0066EC2A36" box="[327,641,1558,1583]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="namphouinensis" status="sp. nov.">
<emphasis id="B96AEA8AFFD3FFAFFED24F0066EC2A36" box="[327,641,1558,1583]" italics="true" pageId="1" pageNumber="58">Amynthas namphouinensis</emphasis>
</taxonomicName>
<emphasis id="B96AEA8AFFD3FFAFFD1F4F0066842A29" bold="true" box="[650,745,1558,1584]" pageId="1" pageNumber="58">
<taxonomicNameLabel id="A25957F1FFD3FFAFFD1F4F0066842A29" box="[650,745,1558,1584]" pageId="1" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
belongs to the
<taxonomicName id="4C1E4D1BFFD3FFAFFC344F0067802A36" box="[929,1005,1558,1583]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="zebrus">
<emphasis id="B96AEA8AFFD3FFAFFC344F0067802A36" box="[929,1005,1558,1583]" italics="true" pageId="1" pageNumber="58">zebrus</emphasis>
</taxonomicName>
group (
<bibRefCitation id="EF8F4B69FFD3FFAFFBDF4F0061342A29" author="Sims" box="[1098,1369,1558,1584]" pageId="1" pageNumber="61" refString="Sims, R. W. &amp; Easton, E. G. (1972) A numerical revision of the earthworm genus Pheretima auct. (Megascolecidae: Oligochaeta) with the recognition of new genera and an appendix on the earthworms collected by the Royal Society North Borneo Expedition. The Biological Journal of the Linnean Society, London, 4, 169 - 268." type="journal article" year="1972">Sims and Easton 1972</bibRefCitation>
). The other Laotian species of the
<taxonomicName id="4C1E4D1BFFD3FFAFFE724F2866592A4E" box="[487,564,1598,1623]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="zebrus">
<emphasis id="B96AEA8AFFD3FFAFFE724F2866592A4E" box="[487,564,1598,1623]" italics="true" pageId="1" pageNumber="58">zebrus</emphasis>
</taxonomicName>
group is
<taxonomicName id="4C1E4D1BFFD3FFAFFD314F2860CC2A41" authority="Thai and Samphon, 1990" authorityName="Thai and Samphon" authorityYear="1990" box="[676,1185,1598,1624]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="xuongmontis">
<emphasis id="B96AEA8AFFD3FFAFFD314F2866DB2A4E" box="[676,694,1598,1623]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFD534F2867312A4E" box="[710,860,1598,1623]" italics="true" pageId="1" pageNumber="58">xuongmontis</emphasis>
(
<collectingCountry id="F3097608FFD3FFAFFCF94F2867CE2A41" box="[876,931,1598,1624]" name="Thailand" pageId="1" pageNumber="58">Thai</collectingCountry>
and Samphon, 1990)
</taxonomicName>
from Luangprabang. Among the
<taxonomicName id="4C1E4D1BFFD3FFAFFEBF4F70651B2A66" box="[298,374,1638,1663]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="zebrus">
<emphasis id="B96AEA8AFFD3FFAFFEBF4F70651B2A66" box="[298,374,1638,1663]" italics="true" pageId="1" pageNumber="58">zebrus</emphasis>
</taxonomicName>
group,
<taxonomicName id="4C1E4D1BFFD3FFAFFE4C4F7066122A66" box="[473,639,1638,1664]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="hilgendorfi">
<emphasis id="B96AEA8AFFD3FFAFFE4C4F7065862A66" box="[473,491,1638,1663]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFE684F7066122A66" box="[509,639,1638,1663]" italics="true" pageId="1" pageNumber="58">hilgendorfi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D1BFFD3FFAFFD064F70674B2A66" box="[659,806,1638,1664]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="palmosus">
<emphasis id="B96AEA8AFFD3FFAFFD064F7066C82A66" box="[659,677,1638,1663]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFD224F70674B2A66" box="[695,806,1638,1663]" italics="true" pageId="1" pageNumber="58">palmosus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1E4D1BFFD3FFAFFCAC4F7067D22A66" box="[825,959,1638,1664]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="eleganis">
<emphasis id="B96AEA8AFFD3FFAFFCAC4F7067262A66" box="[825,843,1638,1663]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFCC84F7067D22A66" box="[861,959,1638,1663]" italics="true" pageId="1" pageNumber="58">eleganis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D1BFFD3FFAFFB944F70608E2A66" box="[1025,1251,1638,1664]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="magnipapillatus">
<emphasis id="B96AEA8AFFD3FFAFFB944F70607E2A66" box="[1025,1043,1638,1663]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFBB04F70608E2A66" box="[1061,1251,1638,1663]" italics="true" pageId="1" pageNumber="58">magnipapillatus</emphasis>
</taxonomicName>
have manicate intestinal caeca.
<taxonomicName id="4C1E4D1BFFD3FFAFFEF54F98658D2ABE" box="[352,480,1678,1703]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="heaneyi">
<emphasis id="B96AEA8AFFD3FFAFFEF54F98658D2ABE" box="[352,480,1678,1703]" italics="true" pageId="1" pageNumber="58">A. heaneyi</emphasis>
</taxonomicName>
from Kitanglad, Mindanao Island,
<collectingCountry id="F3097608FFD3FFAFFC044F9860782AB1" box="[913,1045,1678,1704]" name="Philippines" pageId="1" pageNumber="58">Philippines</collectingCountry>
is easily distinguished from the other members of the group because it is proandric (
<bibRefCitation id="EF8F4B69FFD3FFAFFC8F4FA067C32AC9" author="James" box="[794,942,1718,1744]" pageId="1" pageNumber="61" refString="James, S. W. (2004) New species of Amynthas, Pheretima and Pleionogaster (Oligochaeta: Megascolecidae) of the Mt. Kitanglad Range, Mindanao, Philippines. Raffles Bulletin of Zoology, 52 (2), 289 - 313." type="journal article" year="2004">James 2004</bibRefCitation>
).
<taxonomicName id="4C1E4D1BFFD3FFAFFC504FA060332AD6" box="[965,1118,1718,1744]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="fasciculus">
<emphasis id="B96AEA8AFFD3FFAFFC504FA067BA2AD6" box="[965,983,1718,1743]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFC7C4FA060332AD6" box="[1001,1118,1718,1743]" italics="true" pageId="1" pageNumber="58">fasciculus</emphasis>
</taxonomicName>
from Guizhou,
<collectingCountry id="F3097608FFD3FFAFFAB14FA061072AC9" box="[1316,1386,1718,1744]" name="China" pageId="1" pageNumber="58">China</collectingCountry>
has coiled and kinked spermathecae, whereas
<taxonomicName id="4C1E4D1BFFD3FFAFFD1E4FC867062AEE" box="[651,875,1758,1783]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="namphouinensis">
<emphasis id="B96AEA8AFFD3FFAFFD1E4FC867062AEE" box="[651,875,1758,1783]" italics="true" pageId="1" pageNumber="58">A. namphouinensis</emphasis>
</taxonomicName>
has a straight simple diverticulum. First dorsal pore location is unusually far forward (5/6) in
<emphasis id="B96AEA8AFFD3FFAFFD504E1067C02B06" box="[709,941,1798,1823]" italics="true" pageId="1" pageNumber="58">
<taxonomicName id="4C1E4D1BFFD3FFAFFD504E1067C42B06" box="[709,937,1798,1823]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="namphouinensis">A. namphouinensis</taxonomicName>
,
</emphasis>
distinguishable from other species of the group. Two species from Sulawesi,
<taxonomicName id="4C1E4D1BFFD3FFAFFDA24E3866D72B5E" box="[567,698,1838,1863]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="culminis">
<emphasis id="B96AEA8AFFD3FFAFFDA24E3866D72B5E" box="[567,698,1838,1863]" italics="true" pageId="1" pageNumber="58">A. culminis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1E4D1BFFD3FFAFFD664E3867322B5E" box="[755,863,1838,1863]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="zebrus">
<emphasis id="B96AEA8AFFD3FFAFFD664E3867322B5E" box="[755,863,1838,1863]" italics="true" pageId="1" pageNumber="58">A. zebrus</emphasis>
</taxonomicName>
, have male fields and spermathecal anatomy different from
<emphasis id="B96AEA8AFFD3FFAFFEBF4E4067712B76" box="[298,796,1878,1903]" italics="true" pageId="1" pageNumber="58">
<taxonomicName id="4C1E4D1BFFD3FFAFFEBF4E40667D2B76" box="[298,528,1878,1903]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="namphouinensis">A. namphouinensis</taxonomicName>
.
<taxonomicName id="4C1E4D1BFFD3FFAFFD8B4E4067712B76" box="[542,796,1878,1903]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="principalis">Amynthas principalis</taxonomicName>
</emphasis>
, a large (
<quantity id="4CE69B7DFFD3FFAFFC064E4067962B69" box="[915,1019,1878,1904]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="58" unit="mm" value="300.0">300 mm</quantity>
) species from Bali, has secondary male pores with a very small penis inside each pore, which makes it unique among the
<taxonomicName id="4C1E4D1BFFD3FFAFFB014E68608D2B8E" box="[1172,1248,1918,1943]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="zebrus">
<emphasis id="B96AEA8AFFD3FFAFFB014E68608D2B8E" box="[1172,1248,1918,1943]" italics="true" pageId="1" pageNumber="58">zebrus</emphasis>
</taxonomicName>
group.
</paragraph>
<paragraph id="8BA13698FFD3FFACFF504EB066692A8C" blockId="1.[151,1437,266,2024]" lastBlockId="2.[151,1437,1398,1685]" lastPageId="2" lastPageNumber="59" pageId="1" pageNumber="58">
<taxonomicName id="4C1E4D1BFFD3FFAFFF504EB065932BA6" box="[197,510,1958,1983]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="namphouinensis" status="sp. nov.">
<emphasis id="B96AEA8AFFD3FFAFFF504EB065932BA6" box="[197,510,1958,1983]" italics="true" pageId="1" pageNumber="58">Amynthas namphouinensis</emphasis>
</taxonomicName>
<emphasis id="B96AEA8AFFD3FFAFFD924EB066092BD9" bold="true" box="[519,612,1958,1984]" pageId="1" pageNumber="58">
<taxonomicNameLabel id="A25957F1FFD3FFAFFD924EB066092BD9" box="[519,612,1958,1984]" pageId="1" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
shares some characteristics of the size (
<quantity id="4CE69B7DFFD3FFAFFBAC4EB060E02BD9" box="[1081,1165,1958,1984]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="58" unit="mm" value="90.0">90 mm</quantity>
x
<quantity id="4CE69B7DFFD3FFAFFB3E4EB061642BD9" box="[1195,1289,1958,1984]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="58" unit="mm" value="3.5">3.5 mm</quantity>
), first dorsal pore (5/6), segments (74), and setae (48 on VIII, 55 on XIX) with
<taxonomicName id="4C1E4D1BFFD3FFAFFC0A4ED860382BFE" box="[927,1109,1998,2024]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="1" pageNumber="58" phylum="Annelida" rank="species" species="xuongmontis">
<emphasis id="B96AEA8AFFD3FFAFFC0A4ED867DC2BFE" box="[927,945,1998,2023]" italics="true" pageId="1" pageNumber="58">A</emphasis>
.
<emphasis id="B96AEA8AFFD3FFAFFC2A4ED860382BFE" box="[959,1109,1998,2023]" italics="true" pageId="1" pageNumber="58">xuongmontis</emphasis>
</taxonomicName>
. It has no genital markings near the spermathecal pores and thus, differs from
<taxonomicName id="4C1E4D1BFFD0FFACFD734C6067A92996" box="[742,964,1398,1423]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="59" phylum="Annelida" rank="species" species="namphouinensis">
<emphasis id="B96AEA8AFFD0FFACFD734C6067A92996" box="[742,964,1398,1423]" italics="true" pageId="2" pageNumber="59">A. namphouinensis</emphasis>
</taxonomicName>
which has genital markings just front of 7/
<quantity id="4CE69B7DFFD0FFACFF3A4C8D64B629AC" box="[175,219,1435,1461]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="2" pageNumber="59" unit="in" value="8.0">8 in</quantity>
VII. It seems unlikely that this difference is due to individual variation when all
<specimenCount id="9D18FD11FFD0FFACFB074C8D614F29AC" box="[1170,1314,1435,1461]" pageId="2" pageNumber="59" type="generic">7 specimens</specimenCount>
examined lacked genital markings in the spermathecal segments, and all of the specimens examined by
<collectingCountry id="F3097608FFD0FFACFB784CD7614E29C2" box="[1261,1315,1473,1499]" name="Thailand" pageId="2" pageNumber="59">Thai</collectingCountry>
and Samphon (1990) had them.
<taxonomicName id="4C1E4D1BFFD0FFACFE2D4CF0669929E6" box="[440,756,1510,1535]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="59" phylum="Annelida" rank="species" species="namphouinensis" status="sp. nov.">
<emphasis id="B96AEA8AFFD0FFACFE2D4CF0669929E6" box="[440,756,1510,1535]" italics="true" pageId="2" pageNumber="59">Amynthas namphouinensis</emphasis>
</taxonomicName>
<emphasis id="B96AEA8AFFD0FFACFC954CF0670C2A19" bold="true" box="[768,865,1510,1536]" pageId="2" pageNumber="59">
<taxonomicNameLabel id="A25957F1FFD0FFACFC954CF0670C2A19" box="[768,865,1510,1536]" pageId="2" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
has four positions of genital papillae in XVII, XVIII, and XIX: equatorial in XVII, widely paired in XIX, mid-ventral in 17/18, and closely paired in midventral XIX. In contrast,
<taxonomicName id="4C1E4D1BFFD0FFACFE284F27661E2A53" box="[445,627,1585,1611]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="59" phylum="Annelida" rank="species" species="xuongmontis">
<emphasis id="B96AEA8AFFD0FFACFE284F2765A22A53" box="[445,463,1585,1610]" italics="true" pageId="2" pageNumber="59">A</emphasis>
.
<emphasis id="B96AEA8AFFD0FFACFE484F27661E2A53" box="[477,627,1585,1610]" italics="true" pageId="2" pageNumber="59">xuongmontis</emphasis>
</taxonomicName>
only has genital papillae only in XVIII: two pairs medial to the male porophores, and 2–5 presetal on XVIII. Also,
<taxonomicName id="4C1E4D1BFFD0FFACFD3B4F4067E02A76" box="[686,909,1622,1647]" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="59" phylum="Annelida" rank="species" species="namphouinensis">
<emphasis id="B96AEA8AFFD0FFACFD3B4F4067E02A76" box="[686,909,1622,1647]" italics="true" pageId="2" pageNumber="59">A. namphouinensis</emphasis>
</taxonomicName>
has 0–7 setae between the male pores, but
<taxonomicName id="4C1E4D1BFFD0FFACFA164F4065402A8D" class="Clitellata" family="Megascolecidae" genus="Amynthas" kingdom="Animalia" order="Opisthopora" pageId="2" pageNumber="59" phylum="Annelida" rank="species" species="xuongmontis">
<emphasis id="B96AEA8AFFD0FFACFA164F4061F82A76" box="[1411,1429,1622,1647]" italics="true" pageId="2" pageNumber="59">A</emphasis>
.
<emphasis id="B96AEA8AFFD0FFACFF024F6D65402A8D" box="[151,301,1659,1684]" italics="true" pageId="2" pageNumber="59">xuongmontis</emphasis>
</taxonomicName>
has more than 15.
</paragraph>
</subSubSection>
</treatment>
</document>
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