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<document id="EBEFE90970823F1632824AFDFAF3F13A" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF94CD78FF4CE939B02FEB29" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Georgiosoma bicornutum" docType="treatment" docVersion="8" lastPageNumber="71" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="68" updateTime="1698435542289" updateUser="ExternalLinkService">
<mods:mods id="3D8389A0C2110385AD26DEBC86376E8B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="84D3AA9088A6E7DFB2054E758D590C0E">
<mods:title id="B3A14CA6616D41CBAFB2C98EE7556EE6">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="6242E8C7CDD589BF143DC8D4178F8449" type="personal">
<mods:role id="A95513DFA52A31C647E1ECC5DE0EA329">
<mods:roleTerm id="E20315C9115A21013C2E645F14BFD202">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F6B53B183F264417A6B2DB25040C4898">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="7A60E11FF036CBDE0E7158308384C29B" type="personal">
<mods:role id="90F81AC6ABE0C4E5ABD10B886D621D66">
<mods:roleTerm id="CF9F53DF619CABA9BAB3C22D26170856">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9F928821EE64829F46DC9C6A3B4B4476">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="5B037A021C90B85EAAFFAD7567DC0D76">text</mods:typeOfResource>
<mods:relatedItem id="4F50D4F663DCDD8DF2EB5E709CD0CB25" type="host">
<mods:titleInfo id="0C494DF3E5AEC37411C8CA40095A17F5">
<mods:title id="EC7DA428DFCB1D31655BCAE47E4B7E2C">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8EB3DB48160F27124FB71C30F3970AD5">
<mods:date id="0D0E27E981E02882C8E1CB1E68D4FA97">2016</mods:date>
<mods:detail id="1A8CCDB9EC58DE8FAA3D2419D3CE51FA" type="volume">
<mods:number id="D3C79EA9F67BFDFA1DDC75E9529227CB">4211</mods:number>
</mods:detail>
<mods:detail id="FC39E14178E6054C0BEB4A3C6097C5A0" type="issue">
<mods:number id="956D23B412CA887498EF27D760E987AC">1</mods:number>
</mods:detail>
<mods:extent id="39CBE87D3626F8D8C78A8DF969689267" unit="page">
<mods:start id="68FC2888FD92ADE418E09B87DD9609C5">1</mods:start>
<mods:end id="00115E399E0B9B7896FDBFEF117E72E7">205</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6BDC2BDA5597B4DC0CB3E6EADFA84E95">journal article</mods:classification>
<mods:identifier id="C035B7314DAB73F24EE1CB94D72EC7E7" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="C31B835606AC5DED6314826D695F87E6" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="26D9E90887FEA7B8FFF5D54348905C1A" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
<mods:identifier id="A2B24D0EBE654DD45BFBBF21FBEF633B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="55B7DB7328C20C617F4331B68699E07F" type="Zenodo-Dep">272750</mods:identifier>
<mods:identifier id="E3C3DD9D3169A584559C50D5E229AB51" type="ZooBank">6B86C6BA-6AFE-4AAD-870D-04794C138D47</mods:identifier>
</mods:mods>
<treatment id="03B7878CFF94CD78FF4CE939B02FEB29" ID-DOI="http://doi.org/10.5281/zenodo.6074400" ID-GBIF-Taxon="127419149" ID-Zenodo-Dep="6074400" LSID="urn:lsid:plazi:treatment:03B7878CFF94CD78FF4CE939B02FEB29" httpUri="http://treatment.plazi.org/id/03B7878CFF94CD78FF4CE939B02FEB29" lastPageId="70" lastPageNumber="71" pageId="67" pageNumber="68">
<subSubSection id="C3046511FF94CD7DFF4CE939B066E8CE" pageId="67" pageNumber="68" type="nomenclature">
<paragraph id="8BA1369AFF94CD7DFF4CE939B3F9E82F" blockId="67.[151,650,332,391]" box="[151,650,332,358]" pageId="67" pageNumber="68">
<heading id="D0E981F6FF94CD7DFF4CE939B3F9E82F" bold="true" box="[151,650,332,358]" fontSize="11" level="1" pageId="67" pageNumber="68" reason="1">
<emphasis id="B96AEA88FF94CD7DFF4CE939B3F9E82F" bold="true" box="[151,650,332,358]" pageId="67" pageNumber="68">
<taxonomicName id="4C1E4D19FF94CD7DFF4CE939B0BFE82F" ID-CoL="848WC" box="[151,460,332,358]" class="Diplopoda" family="Anthroleucosomatidae" genus="Georgiosoma" kingdom="Animalia" order="Chordeumatida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="bicornutum">
<emphasis id="B96AEA88FF94CD7DFF4CE939B0BFE82F" bold="true" box="[151,460,332,358]" italics="true" pageId="67" pageNumber="68">Georgiosoma bicornutum</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF94CD7DFF4CE91BB066E8CE" blockId="67.[151,650,332,391]" box="[151,277,366,391]" pageId="67" pageNumber="68">
<figureCitation id="13252A1FFF94CD7DFF4CE91BB066E8CE" box="[151,277,366,391]" captionStart-0="FIGURE 53" captionStart-1="FIGURE 54" captionStart-2="FIGURE 55" captionStartId-0="67.[151,250,1655,1677]" captionStartId-1="68.[151,250,1668,1690]" captionStartId-2="69.[151,250,1696,1718]" captionTargetBox-0="[188,1398,767,1633]" captionTargetBox-1="[194,1392,193,1646]" captionTargetBox-2="[219,1299,225,1650]" captionTargetId-0="figure@67.[188,1398,767,1633]" captionTargetId-1="figure@68.[194,1392,193,1646]" captionTargetId-2="figure@69.[188,1398,193,1675]" captionTargetPageId-0="67" captionTargetPageId-1="68" captionTargetPageId-2="69" captionText-0="FIGURE 53. Georgiosoma bicornutum gen. et sp. nov., holotype male habitus. Scale line = 1 mm." captionText-1="FIGURE 54. Georgiosoma bicornutum gen. et sp. nov., paratype male. A. Leg 3. B. Leg 10. C. Leg 4. D. Leg 7. E. Leg 11. Scale lines = 0.2 mm." captionText-2="FIGURE 55. Georgiosoma bicornutum gen. et sp. nov., paratype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; n = distal notch; t = telopodite; cp = coxal process; cv = coxal vesicle; lg = logitudinal groove; mp = medial part; pl = posterior lamella, rs = row of setae; sp = sternal plate; ss = sternal sac; sah = subtriangular, acuminate horns. Scale line = 0.3 mm." httpUri-0="https://zenodo.org/record/272803/files/figure.png" httpUri-1="https://zenodo.org/record/272804/files/figure.png" httpUri-2="https://zenodo.org/record/272805/files/figure.png" pageId="67" pageNumber="68">Figs 53–55</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF94CD7DFF4CE9C3B0ABE886" box="[151,472,438,463]" pageId="67" pageNumber="68" type="diagnosis">
<paragraph id="8BA1369AFF94CD7DFF4CE9C3B0ABE886" blockId="67.[151,1436,438,715]" box="[151,472,438,463]" pageId="67" pageNumber="68">
<emphasis id="B96AEA88FF94CD7DFF4CE9C3B062E886" bold="true" box="[151,273,438,463]" pageId="67" pageNumber="68">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF94CD7DFF1CE9AFB465E8BA" box="[199,1302,474,499]" pageId="67" pageNumber="68" type="etymology">
<paragraph id="8BA1369AFF94CD7DFF1CE9AFB465E8BA" blockId="67.[151,1436,438,715]" box="[199,1302,474,499]" pageId="67" pageNumber="68">
<emphasis id="B96AEA88FF94CD7DFF1CE9AFB03DE8BA" bold="true" box="[199,334,474,499]" pageId="67" pageNumber="68">Etymology.</emphasis>
An adjective; the new species is named after a pair of horns on the anterior gonopods.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF94CD7DFF1CE98BB2B1EBEE" pageId="67" pageNumber="68" type="materials_examined">
<paragraph id="8BA1369AFF94CD7DFF1CE98BB5CAEB72" blockId="67.[151,1436,438,715]" pageId="67" pageNumber="68">
<materialsCitation id="3B763CC7FF94CD7DFF1CE98BB5CAEB72" ID-GBIF-Occurrence="1424567077" collectingDate="1983-09-21" collectingDateMax="1983-10-14" collectingDateMin="1983-09-21" collectionCode="ZMUM" collectorName="A. Tirakh &amp; V. Y. Dolzhanskiy" country="Georgia" county="Okureshi" elevation="2000" location="Material" municipality="Grot Lagernaya" pageId="67" pageNumber="68" specimenCount="3" specimenCount-juvenile="1" specimenCount-male="2" typeStatus="holotype">
<emphasis id="B96AEA88FF94CD7DFF1CE98BB0E2EB5E" bold="true" box="[199,401,510,535]" pageId="67" pageNumber="68">
<location id="8EC16041FF94CD7DFF1CE98BB05CEB5E" LSID="urn:lsid:plazi:treatment:03B7878CFF94CD78FF4CE939B02FEB29:8EC16041FF94CD7DFF1CE98BB05CEB5E" box="[199,303,510,535]" country="Georgia" county="Okureshi" municipality="Grot Lagernaya" name="Material" pageId="67" pageNumber="68">Material</location>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FF94CD7DFE32E98AB335EB5E" box="[489,582,510,535]" pageId="67" pageNumber="68" type="male">2 males</specimenCount>
,
<specimenCount id="9D18FD13FF94CD7DFD8EE98AB3BEEB5E" box="[597,717,510,535]" pageId="67" pageNumber="68" type="juvenile">1 juvenile</specimenCount>
).
<emphasis id="B96AEA88FF94CD7DFD39E98BB225EB5E" bold="true" box="[738,854,510,535]" pageId="67" pageNumber="68">
<typeStatus id="54A58838FF94CD7DFD39E98BB222EB5E" box="[738,849,510,535]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFF94CD7DFCBAE98AB297EB5E" box="[865,996,511,535]" name="Georgia" pageId="67" pageNumber="68">GEORGIA</collectingCountry>
: male,
<location id="8EC16041FF94CD7DFBE2E98AB500EB5E" LSID="urn:lsid:plazi:treatment:03B7878CFF94CD78FF4CE939B02FEB29:8EC16041FF94CD7DFBE2E98AB500EB5E" box="[1081,1139,511,535]" country="Georgia" county="Okureshi" municipality="Grot Lagernaya" name="Cave" pageId="67" pageNumber="68">Cave</location>
&quot;
<collectingMunicipality id="6BC5ACE0FF94CD7DFB5DE98AB44FEB5E" box="[1158,1340,511,535]" pageId="67" pageNumber="68">Grot Lagernaya</collectingMunicipality>
1&quot; near
<collectingCounty id="62C04E16FF94CD7DFF4CEA57B072EB72" box="[151,257,546,571]" pageId="67" pageNumber="68">Okureshi</collectingCounty>
,
<elevation id="0033D1A9FF94CD7DFED7EA56B010EB73" box="[268,355,547,571]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="m" value="2000.0">
<quantity id="4CE69B7FFF94CD7DFED7EA56B010EB73" box="[268,355,547,571]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="m" value="2000.0">2000 m</quantity>
</elevation>
,
<collectingDate id="EFE4E9B2FF94CD7DFEB5EA56B316EB72" box="[366,613,547,571]" pageId="67" pageNumber="68" value="1983-09-21" valueMax="1983-10-14" valueMin="1983-09-21">
<date id="FFA0105AFF94CD7DFEB5EA56B316EB72" box="[366,613,547,571]" pageId="67" pageNumber="68" value="1983-09-21" valueMax="1983-10-14" valueMin="1983-09-21">21 Sep.–14 Oct. 1983</date>
</collectingDate>
,
<collectorName id="26EB534CFF94CD7DFDABEA56B3AFEB72" box="[624,732,546,571]" pageId="67" pageNumber="68">A. Tirakh</collectorName>
&amp;
<collectorName id="26EB534CFF94CD7DFCDBEA56B2B0EB72" box="[768,963,546,571]" pageId="67" pageNumber="68">V.Y. Dolzhanskiy</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF94CD7DFBD9EA56B528EB72" box="[1026,1115,547,571]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="67" pageNumber="68">ZMUM</collectionCode>
ρ3229).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF94CD7DFF1CEA33B026EBCA" blockId="67.[151,1436,438,715]" pageId="67" pageNumber="68">
<emphasis id="B96AEA88FF94CD7DFF1CEA33B04CEB16" bold="true" box="[199,319,582,607]" pageId="67" pageNumber="68">Paratypes</emphasis>
(total: 1 male, 1 juveniles).
<materialsCitation id="3B763CC7FF94CD7DFD4CEA33B026EBCA" ID-GBIF-Occurrence="1424567093" collectingDate="1983-09-21" collectingDateMax="1983-10-14" collectingDateMin="1983-09-21" collectionCode="ZMUM" collectorName="A. Tirakh &amp; V. Y. Dolzhanskiy" country="Georgia" county="Okureshi" elevation="2000" location="Both" municipality="Grot Lagernaya" pageId="67" pageNumber="68" specimenCount="2" specimenCount-juvenile="1" specimenCount-male="1">
<location id="8EC16041FF94CD7DFD4CEA33B3BEEB16" LSID="urn:lsid:plazi:treatment:03B7878CFF94CD78FF4CE939B02FEB29:8EC16041FF94CD7DFD4CEA33B3BEEB16" box="[663,717,582,607]" country="Georgia" name="Both" pageId="67" pageNumber="68">Both</location>
from
<collectingCountry id="F309760AFF94CD7DFCC7EA32B2ECEB16" box="[796,927,583,607]" name="Georgia" pageId="67" pageNumber="68">GEORGIA</collectingCountry>
:
<specimenCount id="9D18FD13FF94CD7DFC76EA32B573EB16" box="[941,1024,582,607]" pageId="67" pageNumber="68" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF94CD7DFBCBEA32B5F4EB16" box="[1040,1159,582,607]" pageId="67" pageNumber="68" type="juvenile">1 juvenile</specimenCount>
, same data as holotype (
<collectionCode id="ED0FAE5FFF94CD7DFF45EA1EB184EBCA" box="[158,247,619,643]" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="67" pageNumber="68">ZMUM</collectionCode>
ρ3230).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF94CD7DFF1CEAFBB2B1EBEE" blockId="67.[151,1436,438,715]" box="[199,962,654,679]" pageId="67" pageNumber="68">
<materialsCitation id="3B763CC7FF94CD7DFF1CEAFBB2B1EBEE" ID-GBIF-Occurrence="1424567078" box="[199,962,654,679]" country="Georgia" location="Cave" municipality="Grot Lagernaya" pageId="67" pageNumber="68" specimenCount="1" typeStatus="holotype">
<emphasis id="B96AEA88FF94CD7DFF1CEAFBB016EBEE" bold="true" box="[199,357,654,679]" pageId="67" pageNumber="68">
<typeStatus id="54A58838FF94CD7DFF1CEAFBB071EBEE" box="[199,258,654,679]" pageId="67" pageNumber="68">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFF94CD7DFEB7EAFAB09CEBEE" box="[364,495,655,679]" name="Georgia" pageId="67" pageNumber="68">GEORGIA</collectingCountry>
:
<location id="8EC16041FF94CD7DFE22EAFAB340EBEE" LSID="urn:lsid:plazi:treatment:03B7878CFF94CD78FF4CE939B02FEB29:8EC16041FF94CD7DFE22EAFAB340EBEE" box="[505,563,655,679]" country="Georgia" municipality="Grot Lagernaya" name="Cave" pageId="67" pageNumber="68">Cave</location>
&quot;
<collectingMunicipality id="6BC5ACE0FF94CD7DFD99EAFAB386EBEE" box="[578,757,655,679]" pageId="67" pageNumber="68">Grot Lagernaya</collectingMunicipality>
1&quot; near Okureshi.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF94CD78FF1CEAC7B02FEB29" lastPageId="70" lastPageNumber="71" pageId="67" pageNumber="68" type="description">
<paragraph id="8BA1369AFF94CD7DFF1CEAC7B2D1EB82" blockId="67.[151,1436,438,715]" box="[199,930,690,715]" pageId="67" pageNumber="68">
<emphasis id="B96AEA88FF94CD7DFF1CEAC7B02AEB82" bold="true" box="[199,345,690,715]" pageId="67" pageNumber="68">Description.</emphasis>
Body with 30 segments (including telson) in adults.
</paragraph>
<caption id="DF616612FF94CD7DFF4CEE02B5F5EFC4" box="[151,1158,1655,1677]" httpUri="https://zenodo.org/record/272803/files/figure.png" pageId="67" pageNumber="68" targetBox="[188,1398,767,1633]" targetPageId="67">
<paragraph id="8BA1369AFF94CD7DFF4CEE02B5F5EFC4" blockId="67.[151,1158,1655,1677]" box="[151,1158,1655,1677]" pageId="67" pageNumber="68">
<emphasis id="B96AEA88FF94CD7DFF4CEE02B053EFC5" bold="true" box="[151,288,1655,1677]" pageId="67" pageNumber="68">FIGURE 53.</emphasis>
<taxonomicName id="4C1E4D19FF94CD7DFEFEEE0DB359EFC4" box="[293,554,1655,1677]" class="Diplopoda" family="Anthroleucosomatidae" genus="Georgiosoma" kingdom="Animalia" order="Chordeumatida" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="bicornutum">
<emphasis id="B96AEA88FF94CD7DFEFEEE0DB359EFC4" box="[293,554,1655,1677]" italics="true" pageId="67" pageNumber="68">Georgiosoma bicornutum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF94CD7DFDEBEE02B3BEEFC5" bold="true" box="[560,717,1655,1676]" pageId="67" pageNumber="68">gen. et sp. nov.</emphasis>
, holotype male habitus. Scale line = 1 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF94CD7DFF1CEEC0B5B2EF85" blockId="67.[151,1437,1716,1993]" box="[199,1217,1716,1741]" pageId="67" pageNumber="68">
MEASUREMENTS. Males
<quantity id="4CE69B7FFF94CD7DFE07EEC0B358EF85" box="[476,555,1717,1741]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="67" pageNumber="68" unit="mm" value="20.0">20 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF94CD7DFBBFEEC1B5CEEF85" box="[1124,1213,1716,1741]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="67" pageNumber="68" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFF94CD7DFF1CEEACB3E3EFB8" blockId="67.[151,1437,1716,1993]" box="[199,656,1752,1777]" pageId="67" pageNumber="68">
COLORATION (
<figureCitation id="13252A1FFF94CD7DFEB7EEADB0B3EFB8" box="[364,448,1752,1777]" captionStart="FIGURE 53" captionStartId="67.[151,250,1655,1677]" captionTargetBox="[188,1398,767,1633]" captionTargetId="figure@67.[188,1398,767,1633]" captionTargetPageId="67" captionText="FIGURE 53. Georgiosoma bicornutum gen. et sp. nov., holotype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272803/files/figure.png" pageId="67" pageNumber="68">Fig. 53</figureCitation>
). Yellowish white.
</paragraph>
<paragraph id="8BA1369AFF94CD7DFF1CEE88B401EE80" blockId="67.[151,1437,1716,1993]" pageId="67" pageNumber="68">
HEAD. Slightly concave in males. Labrum with three medial teeth and 4+4 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 10+12 setae, on each plate arranged in 3 irregular rows. Stipites with
<emphasis id="B96AEA88FF94CD7DFEF5EF33B03AEE14" box="[302,329,1862,1885]" italics="true" pageId="67" pageNumber="68">ca</emphasis>
40+40 setae. Antennae elongated,
<quantity id="4CE69B7FFF94CD7DFD07EF30B247EE15" box="[732,820,1861,1885]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="67" pageNumber="68" unit="mm" value="3.4">3.4 mm</quantity>
long in
<typeStatus id="54A58838FF94CD7DFC4DEF31B28FEE14" box="[918,1020,1860,1885]" pageId="67" pageNumber="68" type="holotype">holotype</typeStatus>
. Length of antennomeres (in mm): I (0.13), II (0.25), III (1), IV (0.54), V (0.98), VI (0.26), VII (0.22) and VIII (0.02). Length/breadth ratios of antennomeres I–VII: I (1), II (1.3), III (6.7), IV (3.4), V (6), VI (1.6) and VII (2). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 6, arranged in 3 rows in males.
</paragraph>
<caption id="DF616612FF93CD7AFF4CEEF1B000EFF0" httpUri="https://zenodo.org/record/272804/files/figure.png" pageId="68" pageNumber="69" targetBox="[194,1392,193,1646]" targetPageId="68">
<paragraph id="8BA1369AFF93CD7AFF4CEEF1B000EFF0" blockId="68.[151,1436,1668,1721]" pageId="68" pageNumber="69">
<emphasis id="B96AEA88FF93CD7AFF4CEEF1B052EFD0" bold="true" box="[151,289,1668,1690]" pageId="68" pageNumber="69">FIGURE 54.</emphasis>
<taxonomicName id="4C1E4D19FF93CD7AFEF3EEF0B35FEFD3" box="[296,556,1668,1690]" class="Diplopoda" family="Anthroleucosomatidae" genus="Georgiosoma" kingdom="Animalia" order="Chordeumatida" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="bicornutum">
<emphasis id="B96AEA88FF93CD7AFEF3EEF0B35FEFD3" box="[296,556,1668,1690]" italics="true" pageId="68" pageNumber="69">Georgiosoma bicornutum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF93CD7AFDEFEEF1B3A7EFD0" bold="true" box="[564,724,1668,1689]" pageId="68" pageNumber="69">gen. et sp. nov.</emphasis>
, paratype male.
<emphasis id="B96AEA88FF93CD7AFC5BEEF1B2EAEFD0" bold="true" box="[896,921,1668,1689]" pageId="68" pageNumber="69">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FF93CD7AFC32EEF1B573EFD0" bold="true" box="[1001,1024,1668,1689]" pageId="68" pageNumber="69">B.</emphasis>
Leg 10.
<emphasis id="B96AEA88FF93CD7AFB86EEF1B505EFD3" bold="true" box="[1117,1142,1668,1690]" pageId="68" pageNumber="69">C.</emphasis>
Leg 4.
<emphasis id="B96AEA88FF93CD7AFB1CEEF1B5ACEFD0" bold="true" box="[1223,1247,1668,1689]" pageId="68" pageNumber="69">D.</emphasis>
Leg 7.
<emphasis id="B96AEA88FF93CD7AFAEBEEF1B434EFD0" bold="true" box="[1328,1351,1668,1689]" pageId="68" pageNumber="69">E.</emphasis>
Leg 11. Scale lines = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF93CD7AFF1CEE95B189EE55" blockId="68.[151,1436,1760,2001]" pageId="68" pageNumber="69">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BA1369AFF93CD7AFF1CEF5CB36BEEC0" blockId="68.[151,1436,1760,2001]" pageId="68" pageNumber="69">
BODY SEGMENTS (
<figureCitation id="13252A1FFF93CD7AFE7AEF5DB088EE08" box="[417,507,1832,1857]" captionStart="FIGURE 53" captionStartId="67.[151,250,1655,1677]" captionTargetBox="[188,1398,767,1633]" captionTargetId="figure@67.[188,1398,767,1633]" captionTargetPageId="67" captionText="FIGURE 53. Georgiosoma bicornutum gen. et sp. nov., holotype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272803/files/figure.png" pageId="68" pageNumber="69">Fig. 53</figureCitation>
). Lateral keels well-developed. Macrochaetae medium-sized, outer longer than medial and inner, all trichoid. CIX (pleurotergite 15) ~ 0.75; MIX (pleurotergite 15) ~ 1.5; PIX (pleurotergite 15) ~ 0.6; MA (pleurotergite 15) ~ 135˚.
</paragraph>
<paragraph id="8BA1369AFF93CD7AFF1CEFE0B39FEE99" blockId="68.[151,1436,1760,2001]" pageId="68" pageNumber="69">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<caption id="DF616612FF92CD7BFF4CEED5B428EE5B" httpUri="https://zenodo.org/record/272805/files/figure.png" pageId="69" pageNumber="70" targetBox="[219,1299,225,1650]" targetPageId="69">
<paragraph id="8BA1369AFF92CD7BFF4CEED5B428EE5B" blockId="69.[151,1436,1696,1810]" pageId="69" pageNumber="70">
<emphasis id="B96AEA88FF92CD7BFF4CEED5B051EFFF" bold="true" box="[151,290,1696,1718]" pageId="69" pageNumber="70">FIGURE 55.</emphasis>
<taxonomicName id="4C1E4D19FF92CD7BFEF0EED4B341EFFF" box="[299,562,1696,1718]" class="Diplopoda" family="Anthroleucosomatidae" genus="Georgiosoma" kingdom="Animalia" order="Chordeumatida" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="bicornutum">
<emphasis id="B96AEA88FF92CD7BFEF0EED4B341EFFF" box="[299,562,1696,1718]" italics="true" pageId="69" pageNumber="70">Georgiosoma bicornutum</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF92CD7BFDE0EED4B393EFFF" bold="true" box="[571,736,1697,1718]" pageId="69" pageNumber="70">gen. et sp. nov.</emphasis>
, paratype male gonopods.
<emphasis id="B96AEA88FF92CD7BFC27EED5B566EFFC" bold="true" box="[1020,1045,1696,1717]" pageId="69" pageNumber="70">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FF92CD7BFA5FEED4B4E8EFFF" bold="true" box="[1412,1435,1697,1718]" pageId="69" pageNumber="70">B.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FF92CD7BFE33EECAB373EF9C" bold="true" box="[488,512,1727,1749]" pageId="69" pageNumber="70">C.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FF92CD7BFCAFEECAB2FEEF9D" bold="true" box="[884,909,1727,1748]" pageId="69" pageNumber="70">D.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B96AEA88FF92CD7BFADEEEB5B463EF9C" box="[1285,1296,1728,1749]" italics="true" pageId="69" pageNumber="70">c</emphasis>
= coxite;
<emphasis id="B96AEA88FF92CD7BFAA1EEB5B4F4EF9C" box="[1402,1415,1728,1749]" italics="true" pageId="69" pageNumber="70">n</emphasis>
= distal notch;
<emphasis id="B96AEA88FF92CD7BFEC6EEAAB057EFBD" box="[285,292,1759,1780]" italics="true" pageId="69" pageNumber="70">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FF92CD7BFE6CEEAAB0A3EFBD" box="[439,464,1759,1780]" italics="true" pageId="69" pageNumber="70">cp</emphasis>
= coxal process;
<emphasis id="B96AEA88FF92CD7BFD53EEAAB3EDEFBD" box="[648,670,1759,1780]" italics="true" pageId="69" pageNumber="70">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FF92CD7BFC8AEEA8B216EFBA" box="[849,869,1757,1779]" italics="true" pageId="69" pageNumber="70">lg</emphasis>
= logitudinal groove;
<emphasis id="B96AEA88FF92CD7BFB94EEAAB51DEFBD" box="[1103,1134,1759,1780]" italics="true" pageId="69" pageNumber="70">mp</emphasis>
= medial part;
<emphasis id="B96AEA88FF92CD7BFACBEEA8B450EFBA" box="[1296,1315,1757,1779]" italics="true" pageId="69" pageNumber="70">pl</emphasis>
= posterior lamella,
<emphasis id="B96AEA88FF92CD7BFF35EE88B072EE5B" box="[238,257,1789,1810]" italics="true" pageId="69" pageNumber="70">rs</emphasis>
= row of setae;
<emphasis id="B96AEA88FF92CD7BFE7EEE88B0C8EE5B" box="[421,443,1789,1810]" italics="true" pageId="69" pageNumber="70">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF92CD7BFD84EE88B300EE5B" box="[607,627,1789,1810]" italics="true" pageId="69" pageNumber="70">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF92CD7BFCDEEE89B25AEE5B" box="[773,809,1788,1810]" italics="true" pageId="69" pageNumber="70">sah</emphasis>
= subtriangular, acuminate horns. Scale line = 0.3 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF92CD7BFF1CEF35B2ECEE32" blockId="69.[151,1436,1855,2024]" pageId="69" pageNumber="70">
WALKING LEGS (
<figureCitation id="13252A1FFF92CD7BFE5EEF4AB0A9EE11" box="[389,474,1855,1880]" captionStart="FIGURE 53" captionStartId="67.[151,250,1655,1677]" captionTargetBox="[188,1398,767,1633]" captionTargetId="figure@67.[188,1398,767,1633]" captionTargetPageId="67" captionText="FIGURE 53. Georgiosoma bicornutum gen. et sp. nov., holotype male habitus. Scale line = 1 mm." httpUri="https://zenodo.org/record/272803/files/figure.png" pageId="69" pageNumber="70">Fig. 53</figureCitation>
). Leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae.
</paragraph>
<paragraph id="8BA1369AFF92CD78FF1CEFFDB0CBE99D" blockId="69.[151,1436,1855,2024]" lastBlockId="70.[151,1437,151,608]" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFF92CD7BFDC0EFF2B302EEE9" box="[539,625,1927,1952]" captionStart="FIGURE 54" captionStartId="68.[151,250,1668,1690]" captionTargetBox="[194,1392,193,1646]" captionTargetId="figure@68.[194,1392,193,1646]" captionTargetPageId="68" captionText="FIGURE 54. Georgiosoma bicornutum gen. et sp. nov., paratype male. A. Leg 3. B. Leg 10. C. Leg 4. D. Leg 7. E. Leg 11. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272804/files/figure.png" pageId="69" pageNumber="70">Fig. 54</figureCitation>
). Leg-pairs 3–7 enlarged. Leg-pairs 3 and 4 each with a basal external protrusion on prefemur. Leg-pair 5 with a strongly developed protrusion on prefemur. Leg-pair 6 without peculiarities. Coxae of leg-pair 7 with a well-developed, mesally curved horn, anteriorly with a subtriangular protrusion. Leg-pair 10 with coxal glands and well-developed coxal horns. Leg-pair 11 with coxal glands and short posterior coxal processes.
</paragraph>
<paragraph id="8BA1369AFF91CD78FF1CE895B209E8E5" blockId="70.[151,1437,151,608]" pageId="70" pageNumber="71">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFF91CD78FE0BE8AAB350E9B1" box="[464,547,223,248]" captionStart="FIGURE 55" captionStartId="69.[151,250,1696,1718]" captionTargetBox="[219,1299,225,1650]" captionTargetId="figure@69.[188,1398,193,1675]" captionTargetPageId="69" captionText="FIGURE 55. Georgiosoma bicornutum gen. et sp. nov., paratype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; n = distal notch; t = telopodite; cp = coxal process; cv = coxal vesicle; lg = logitudinal groove; mp = medial part; pl = posterior lamella, rs = row of setae; sp = sternal plate; ss = sternal sac; sah = subtriangular, acuminate horns. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272805/files/figure.png" pageId="70" pageNumber="71">Fig. 55</figureCitation>
A–C). Sternal plate (
<emphasis id="B96AEA88FF91CD78FCCAE894B259E9B1" box="[785,810,225,248]" italics="true" pageId="70" pageNumber="71">sp</emphasis>
) anteriorly with a medial, triangular, acuminate, hairy, sternal sac (
<emphasis id="B96AEA88FF91CD78FEC5E970B047E855" box="[286,308,261,284]" italics="true" pageId="70" pageNumber="71">ss</emphasis>
). Coxal processes (
<emphasis id="B96AEA88FF91CD78FDC9E970B35EE855" box="[530,557,261,284]" italics="true" pageId="70" pageNumber="71">cp</emphasis>
) fused, only distally with a notch (
<emphasis id="B96AEA88FF91CD78FC62E970B2B4E855" box="[953,967,261,284]" italics="true" pageId="70" pageNumber="71">n</emphasis>
) and an anterior longitudinal groove (
<emphasis id="B96AEA88FF91CD78FAADE971B4FFE854" box="[1398,1420,260,285]" italics="true" pageId="70" pageNumber="71">lg</emphasis>
), forming a rounded shield with two well-developed, subtriangular and acuminate horns (
<emphasis id="B96AEA88FF91CD78FB6BE952B5A4E809" box="[1200,1239,295,320]" italics="true" pageId="70" pageNumber="71">sah</emphasis>
) in anterior and posterior views. These horns connected to posterior lamellae (
<emphasis id="B96AEA88FF91CD78FCB6E939B2F0E82C" box="[877,899,332,357]" italics="true" pageId="70" pageNumber="71">pl</emphasis>
) supporting denticles and setae in upper half. Mediodistal parts (
<emphasis id="B96AEA88FF91CD78FEB2E904B0F8E8C1" box="[361,395,369,392]" italics="true" pageId="70" pageNumber="71">mp</emphasis>
) with denticulate upper edges and mesal edges folded inside and forming a denticulate winglike structure. Posteriorly, two diagonal rows of setae (
<emphasis id="B96AEA88FF91CD78FD20E9E0B262E8E5" box="[763,785,405,428]" italics="true" pageId="70" pageNumber="71">rs</emphasis>
) present.
</paragraph>
<paragraph id="8BA1369AFF91CD78FF1CE9CDB4E2E8BC" blockId="70.[151,1437,151,608]" pageId="70" pageNumber="71">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFF91CD78FE0CE9C2B358E899" box="[471,555,439,464]" captionStart="FIGURE 55" captionStartId="69.[151,250,1696,1718]" captionTargetBox="[219,1299,225,1650]" captionTargetId="figure@69.[188,1398,193,1675]" captionTargetPageId="69" captionText="FIGURE 55. Georgiosoma bicornutum gen. et sp. nov., paratype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, lateral view. C. Anterior gonopods, posterior view. D. Posterior gonopods, posterior view. c = coxite; n = distal notch; t = telopodite; cp = coxal process; cv = coxal vesicle; lg = logitudinal groove; mp = medial part; pl = posterior lamella, rs = row of setae; sp = sternal plate; ss = sternal sac; sah = subtriangular, acuminate horns. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272805/files/figure.png" pageId="70" pageNumber="71">Fig. 55</figureCitation>
D). Coxites (
<emphasis id="B96AEA88FF91CD78FD65E9CCB3B9E899" box="[702,714,441,464]" italics="true" pageId="70" pageNumber="71">c</emphasis>
) divided. Telopodites (
<emphasis id="B96AEA88FF91CD78FC0FE9CCB2AFE899" box="[980,988,441,464]" italics="true" pageId="70" pageNumber="71">t</emphasis>
) present on posterolateral side, clothed with setae. Coxal processes (
<emphasis id="B96AEA88FF91CD78FE00E9A8B085E8BD" box="[475,502,477,500]" italics="true" pageId="70" pageNumber="71">cp</emphasis>
) horn-like, clothed with minute setae. Coxal vesicles (
<emphasis id="B96AEA88FF91CD78FB81E9A8B500E8BD" box="[1114,1139,477,500]" italics="true" pageId="70" pageNumber="71">cv</emphasis>
) present on anterior side.
</paragraph>
<paragraph id="8BA1369AFF91CD78FF1CE98AB02FEB29" blockId="70.[151,1437,151,608]" pageId="70" pageNumber="71">
<emphasis id="B96AEA88FF91CD78FF1CE98AB062EB51" bold="true" box="[199,273,511,536]" pageId="70" pageNumber="71">Notes.</emphasis>
This species is only known from a cave and shows some troglomorphic features such as reduced body pigmentation and number of ocelli, a larger number of setae on the gnathochilarium, as well as elongated antennae and walking legs.
</paragraph>
</subSubSection>
</treatment>
</document>
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