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<document id="1DA1EDD94740DC08ED9912C1291A4928" ID-CLB-Dataset="37322" ID-DOI="10.11646/zootaxa.4211.1.1" ID-GBIF-Dataset="7af458af-d56d-44e3-a6f7-af97868175d9" ID-ISSN="1175-5326" ID-Zenodo-Dep="272750" ID-ZooBank="6B86C6BA-6AFE-4AAD-870D-04794C138D47" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1484386885843" checkinUser="plazi" docAuthor="Antić, Dragan Ž. &amp; Makarov, Slobodan E." docDate="2016" docId="03B7878CFF59CDACFF4CEB82B055ED11" docLanguage="en" docName="zootaxa.4211.1.1.pdf.imf" docOrigin="Zootaxa 4211 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Acanthophorella barjadzei" docType="treatment" docVersion="8" lastPageNumber="147" masterDocId="FF8EFFF4FFD7CD3EFFDBE875B173E949" masterDocTitle="The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)" masterLastPageNumber="205" masterPageNumber="1" pageNumber="143" updateTime="1698435542289" updateUser="ExternalLinkService">
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<mods:title id="7D4F7B74D5B200E4D85EB77C1996B8D7">The Caucasus as a major hotspot of biodiversity: Evidence from the millipede family Anthroleucosomatidae (Diplopoda, Chordeumatida)</mods:title>
</mods:titleInfo>
<mods:name id="F11DC22FDD438247885C3C0239190117" type="personal">
<mods:role id="873ED11F0EBA77FACDAA92A8FC81764E">
<mods:roleTerm id="9E025F8D5404AEE86BE8F79D234BD590">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F49ACA218DC0F9D5CB4FBEB498889AEE">Antić, Dragan Ž.</mods:namePart>
</mods:name>
<mods:name id="1FADDB4B1BEA934BB07B37F85317BFE2" type="personal">
<mods:role id="14668FC9FB14C906C7958902B355DD9C">
<mods:roleTerm id="81DF954D5C7E46EF07EBCF9F269810ED">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD2138856580410AC5550DC8C3223AB9">Makarov, Slobodan E.</mods:namePart>
</mods:name>
<mods:typeOfResource id="982C172611208CF0DCE9B2078168DD49">text</mods:typeOfResource>
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<mods:titleInfo id="2788E7E786EAADF99FB695CD5050172D">
<mods:title id="3DDC0293AEC96AA8BAC5EA12E174DF21">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C9D62B9B4E2DE705E91DE9ED96DB366E">
<mods:date id="80231DB7120E35E507370B4293276326">2016</mods:date>
<mods:detail id="147C637ED09C49EE8CAFC2EB47CC1B8F" type="volume">
<mods:number id="1F4FB53603CB65A5795D8DD811663C6E">4211</mods:number>
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<mods:number id="BF5BC77075448B4B854E24053EAEC97F">1</mods:number>
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<mods:classification id="5A921007394FE7FB6B09BBF4874B017A">journal article</mods:classification>
<mods:identifier id="C6E481B03A144A86A8A168698457AB81" type="CLB-Dataset">37322</mods:identifier>
<mods:identifier id="75C9A8F1C12CAE4D5E7FF1BB354D2012" type="DOI">10.11646/zootaxa.4211.1.1</mods:identifier>
<mods:identifier id="E95EAF8616EEDEB13FA66C5716327A61" type="GBIF-Dataset">7af458af-d56d-44e3-a6f7-af97868175d9</mods:identifier>
<mods:identifier id="0D88F45C384E79CAB07D4C3D23CD3FFB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03B7878CFF59CDACFF4CEB82B055ED11" ID-DOI="http://doi.org/10.5281/zenodo.6074466" ID-GBIF-Taxon="127419185" ID-Zenodo-Dep="6074466" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11" httpUri="http://treatment.plazi.org/id/03B7878CFF59CDACFF4CEB82B055ED11" lastPageId="146" lastPageNumber="147" pageId="142" pageNumber="143">
<subSubSection id="C3046511FF59CDB0FF4CEB82B05CED7A" pageId="142" pageNumber="143" type="nomenclature">
<paragraph id="8BA1369AFF59CDB0FF4CEB82B3E5ED5B" blockId="142.[151,662,1015,1075]" box="[151,662,1015,1042]" pageId="142" pageNumber="143">
<heading id="D0E981F6FF59CDB0FF4CEB82B3E5ED5B" bold="true" box="[151,662,1015,1042]" fontSize="11" level="1" pageId="142" pageNumber="143" reason="1">
<emphasis id="B96AEA88FF59CDB0FF4CEB82B3E5ED5B" bold="true" box="[151,662,1015,1042]" pageId="142" pageNumber="143">
<taxonomicName id="4C1E4D19FF59CDB0FF4CEB82B0ABED58" ID-CoL="8466W" box="[151,472,1015,1041]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="142" pageNumber="143" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B96AEA88FF59CDB0FF4CEB82B0ABED58" bold="true" box="[151,472,1015,1041]" italics="true" pageId="142" pageNumber="143">Acanthophorella barjadzei</emphasis>
</taxonomicName>
gen. et sp. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA1369AFF59CDB0FF4CEC6FB05CED7A" blockId="142.[151,662,1015,1075]" box="[151,303,1050,1075]" pageId="142" pageNumber="143">
<figureCitation id="13252A1FFF59CDB0FF4CEC6FB05CED7A" box="[151,303,1050,1075]" captionStart-0="FIGURE 118" captionStart-1="FIGURE 119" captionStart-2="FIGURE 120" captionStartId-0="143.[151,250,1932,1954]" captionStartId-1="144.[151,250,1949,1971]" captionStartId-2="145.[151,250,1875,1897]" captionTargetBox-0="[188,1398,337,1911]" captionTargetBox-1="[188,1398,193,1927]" captionTargetBox-2="[272,1245,214,1817]" captionTargetId-0="figure@143.[188,1398,337,1911]" captionTargetId-1="figure@144.[188,1398,193,1927]" captionTargetId-2="figure@145.[226,1351,193,1879]" captionTargetPageId-0="143" captionTargetPageId-1="144" captionTargetPageId-2="145" captionText-0="FIGURE 118. Acanthophorella barjadzei gen. et sp. nov., habitus. A. Paratype female. Scale line = 1 mm. B. Holotype male (photo S. Barjadze), without scale." captionText-1="FIGURE 119. Acanthophorella barjadzei gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 7. D. Leg 10. Scale lines = 0.2 mm." captionText-2="FIGURE 120. Acanthophorella barjadzei gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. c = coxite; t = telopodite; cp = coxal process; cv = coxal vesicle; hl = hairy lever; ll = lateral lamella; lo = anterior lobe; mp = medial shield; sp = sternal plate; ss = sternal sac; sw = widening; tp = thorn-like process; us = undefined structure; ppr = posterior projection; str = striae. Scale line = 0.3 mm." httpUri-0="https://zenodo.org/record/272867/files/figure.png" httpUri-1="https://zenodo.org/record/272868/files/figure.png" httpUri-2="https://zenodo.org/record/272869/files/figure.png" pageId="142" pageNumber="143">Figs 118120</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF59CDB0FF4CEC17B2F8EC43" pageId="142" pageNumber="143" type="diagnosis">
<paragraph id="8BA1369AFF59CDB0FF4CEC17B2F8EC43" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
<emphasis id="B96AEA88FF59CDB0FF4CEC17B062ED32" bold="true" box="[151,273,1122,1147]" pageId="142" pageNumber="143">Diagnosis.</emphasis>
The new species differs from the other two species of the genus by the structure of the anterior gonopods and body, such as the presence of very wide shields on the anterior gonopods with a pair of lobes at the base or the presence of narrower and more gracile processes on the anterior side of the anterior gonopods, as well as a twice larger, pigmentless, yellowish white body with less numerous ocelli, with elongated walking legs and antennae and a very characteristic coxal process on male leg-pair 7.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF59CDB0FF1CED63B05BEC1A" pageId="142" pageNumber="143" type="etymology">
<paragraph id="8BA1369AFF59CDB0FF1CED63B05BEC1A" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
<emphasis id="B96AEA88FF59CDB0FF1CED63B03DEC66" bold="true" box="[199,334,1302,1327]" pageId="142" pageNumber="143">Etymology.</emphasis>
The new species is named after Dr. Shalva Barjadze, a Georgian entomologist and biospeleologist, the collector.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF59CDB0FF1CED2BB013EF06" pageId="142" pageNumber="143" type="materials_examined">
<paragraph id="8BA1369AFF59CDB0FF1CED2BB00AECF6" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
<materialsCitation id="3B763CC7FF59CDB0FF1CED2BB00AECF6" ID-GBIF-Occurrence="1424566986" collectingDate="2014-10-13" collectionCode="IZB" collectorName="S. Barjadze" country="Georgia" county="Cave Dolabistavi" location="Material" municipality="Velevi" pageId="142" pageNumber="143" specimenCount="8" specimenCount-female="2" specimenCount-juvenile="5" specimenCount-male="1" typeStatus="holotype">
<emphasis id="B96AEA88FF59CDB0FF1CED2BB0E5EC3E" bold="true" box="[199,406,1374,1399]" pageId="142" pageNumber="143">
<location id="8EC16041FF59CDB0FF1CED2BB05CEC3E" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FF1CED2BB05CEC3E" box="[199,303,1374,1399]" country="Georgia" county="Cave Dolabistavi" municipality="Velevi" name="Material" pageId="142" pageNumber="143">Material</location>
studied
</emphasis>
(total:
<specimenCount id="9D18FD13FF59CDB0FE23ED2AB33CEC3E" box="[504,591,1374,1399]" pageId="142" pageNumber="143" type="male">1 male</specimenCount>
,
<specimenCount id="9D18FD13FF59CDB0FDB8ED2AB3A8EC3E" box="[611,731,1374,1399]" pageId="142" pageNumber="143" type="female">2 females</specimenCount>
,
<specimenCount id="9D18FD13FF59CDB0FD34ED2BB205EC3E" box="[751,886,1374,1399]" pageId="142" pageNumber="143" type="juvenile">5 juveniles</specimenCount>
).
<emphasis id="B96AEA88FF59CDB0FC4AED2BB575EC3E" bold="true" box="[913,1030,1374,1399]" pageId="142" pageNumber="143">
<typeStatus id="54A58838FF59CDB0FC4AED2BB573EC3E" box="[913,1024,1374,1399]" pageId="142" pageNumber="143" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectingCountry id="F309760AFF59CDB0FBCEED2AB5EBEC3E" box="[1045,1176,1375,1399]" name="Georgia" pageId="142" pageNumber="143">GEORGIA</collectingCountry>
: male,
<location id="8EC16041FF59CDB0FB23ED2BB4EBEC3E" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FB23ED2BB4EBEC3E" box="[1272,1432,1374,1399]" country="Georgia" county="Cave Dolabistavi" municipality="Velevi" name="Racha region" pageId="142" pageNumber="143">Racha region</location>
,
<location id="8EC16041FF59CDB0FF4CEDF7B001ECD2" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FF4CEDF7B001ECD2" box="[151,370,1410,1435]" country="Georgia" county="Cave Dolabistavi" municipality="Velevi" name="Ambrolauri district" pageId="142" pageNumber="143">Ambrolauri district</location>
,
<location id="8EC16041FF59CDB0FEA4EDF7B0B6ECD2" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FEA4EDF7B0B6ECD2" box="[383,453,1410,1435]" country="Georgia" county="Cave Dolabistavi" municipality="Velevi" name="Racha" pageId="142" pageNumber="143">Racha</location>
karst massif,
<quantity id="4CE69B7FFF59CDB0FDBEEDF6B3ECECD2" box="[613,671,1410,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="142" pageNumber="143" unit="km" value="1.0">1 km</quantity>
from village
<collectingMunicipality id="6BC5ACE0FF59CDB0FCE1EDF7B2F6ECD2" box="[826,901,1410,1435]" pageId="142" pageNumber="143">Velevi</collectingMunicipality>
,
<collectingCounty id="62C04E16FF59CDB0FC4AEDF6B524ECD2" box="[913,1111,1410,1435]" pageId="142" pageNumber="143">Cave Dolabistavi</collectingCounty>
, dark zone,
<collectingDate id="EFE4E9B2FF59CDB0FB39EDF6B40AECD2" box="[1250,1401,1411,1435]" pageId="142" pageNumber="143" value="2014-10-13">
<date id="FFA0105AFF59CDB0FB39EDF6B40AECD2" box="[1250,1401,1411,1435]" pageId="142" pageNumber="143" value="2014-10-13">13 Oct. 2014</date>
</collectingDate>
,
<collectorName id="26EB534CFF59CDB0FA5DEDF6B188ECF6" pageId="142" pageNumber="143">S. Barjadze</collectorName>
leg. (
<collectionCode id="ED0FAE5FFF59CDB0FEE1EDD2B01FECF6" box="[314,364,1447,1471]" pageId="142" pageNumber="143">IZB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF59CDB0FF1CEDBFB06AEF4E" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
<emphasis id="B96AEA88FF59CDB0FF1CEDBFB04CECAA" bold="true" box="[199,319,1482,1507]" pageId="142" pageNumber="143">Paratypes</emphasis>
(total: 2 females, 3 juveniles).
<materialsCitation id="3B763CC7FF59CDB0FD79EDBFB06AEF4E" ID-GBIF-Occurrence="1424566985" collectingDate="2014-10-13" collectionCode="ISU, IZB" collectorName="S. Barjadze" country="Georgia" county="Cave Dolabistavi" location="All" municipality="Velevi" pageId="142" pageNumber="143" specimenCount="7" specimenCount-female="2" specimenCount-juvenile="5">
<location id="8EC16041FF59CDB0FD79EDBFB3B5ECAA" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FD79EDBFB3B5ECAA" box="[674,710,1482,1507]" country="Georgia" name="All" pageId="142" pageNumber="143">All</location>
from
<collectingCountry id="F309760AFF59CDB0FCD6EDBEB2E3ECAA" box="[781,912,1483,1507]" name="Georgia" pageId="142" pageNumber="143">GEORGIA</collectingCountry>
:
<specimenCount id="9D18FD13FF59CDB0FC40EDBEB578ECAA" box="[923,1035,1482,1507]" pageId="142" pageNumber="143" type="female">2 females</specimenCount>
,
<specimenCount id="9D18FD13FF59CDB0FBC3EDBFB5E5ECAA" box="[1048,1174,1482,1507]" pageId="142" pageNumber="143" type="juvenile">5 juveniles</specimenCount>
, same data as holotype (
<collectionCode id="ED0FAE5FFF59CDB0FF44ED9AB1A3EF4E" box="[159,208,1519,1543]" country="USA" httpUri="http://grbio.org/cool/jgjv-qdg3" name="Indiana State University" pageId="142" pageNumber="143">ISU</collectionCode>
,
<collectionCode id="ED0FAE5FFF59CDB0FF00ED9AB07EEF4E" box="[219,269,1519,1543]" pageId="142" pageNumber="143">IZB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BA1369AFF59CDB0FF1CEE67B013EF06" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
<materialsCitation id="3B763CC7FF59CDB0FF1CEE67B013EF06" ID-GBIF-Occurrence="1424566989" country="Georgia" county="Velevi" location="Racha region" municipality="Racha" pageId="142" pageNumber="143" specimenCount="1" typeStatus="holotype">
<emphasis id="B96AEA88FF59CDB0FF1CEE67B014EF62" bold="true" box="[199,359,1554,1579]" pageId="142" pageNumber="143">
<typeStatus id="54A58838FF59CDB0FF1CEE67B071EF62" box="[199,258,1554,1579]" pageId="142" pageNumber="143">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F309760AFF59CDB0FEABEE66B080EF62" box="[368,499,1555,1579]" name="Georgia" pageId="142" pageNumber="143">GEORGIA</collectingCountry>
:
<location id="8EC16041FF59CDB0FE25EE67B3EBEF62" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FE25EE67B3EBEF62" box="[510,664,1554,1579]" country="Georgia" county="Velevi" municipality="Racha" name="Racha region" pageId="142" pageNumber="143">Racha region</location>
,
<location id="8EC16041FF59CDB0FD7EEE67B2F2EF62" LSID="urn:lsid:plazi:treatment:03B7878CFF59CDACFF4CEB82B055ED11:8EC16041FF59CDB0FD7EEE67B2F2EF62" box="[677,897,1554,1579]" country="Georgia" county="Velevi" municipality="Racha" name="Ambrolauri district" pageId="142" pageNumber="143">Ambrolauri district</location>
,
<collectingMunicipality id="6BC5ACE0FF59CDB0FC55EE67B2A7EF62" box="[910,980,1554,1579]" pageId="142" pageNumber="143">Racha</collectingMunicipality>
karst massif,
<quantity id="4CE69B7FFF59CDB0FBAEEE66B5C3EF62" box="[1141,1200,1554,1579]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="142" pageNumber="143" unit="km" value="1.0">1 km</quantity>
from village
<collectingCounty id="62C04E16FF59CDB0FA95EE67B4EBEF62" box="[1358,1432,1554,1579]" pageId="142" pageNumber="143">Velevi</collectingCounty>
, Dolabistavi Cave.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF59CDACFF1CEE2FB1B8EAC9" lastPageId="146" lastPageNumber="147" pageId="142" pageNumber="143" type="description">
<paragraph id="8BA1369AFF59CDB0FF1CEE2FB2D1EF3A" blockId="142.[151,1436,1122,2011]" box="[199,930,1626,1651]" pageId="142" pageNumber="143">
<emphasis id="B96AEA88FF59CDB0FF1CEE2FB02AEF3A" bold="true" box="[199,345,1626,1651]" pageId="142" pageNumber="143">Description.</emphasis>
Body with 31 segments (including telson) in adults.
</paragraph>
<paragraph id="8BA1369AFF59CDB0FF1CEE0AB2A4EFF3" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
MEASUREMENTS.
<typeStatus id="54A58838FF59CDB0FE49EE0BB089EFDE" box="[402,506,1662,1687]" pageId="142" pageNumber="143" type="holotype">Holotype</typeStatus>
male
<quantity id="4CE69B7FFF59CDB0FD99EE0AB3E0EFDF" box="[578,659,1663,1687]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" pageId="142" pageNumber="143" unit="mm" value="17.0">17 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF59CDB0FB03EE0AB440EFDF" box="[1240,1331,1663,1687]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="142" pageNumber="143" unit="mm" value="1.4">1.4 mm</quantity>
. Females
<quantity id="4CE69B7FFF59CDB0FF4CEED7B063EFF3" box="[151,272,1698,1723]" metricMagnitude="-2" metricUnit="m" metricValue="1.7" metricValueMax="1.8" metricValueMin="1.6" pageId="142" pageNumber="143" unit="mm" value="17.0" valueMax="18.0" valueMin="16.0">1618 mm</quantity>
long, vertical diameter of the largest pleurotergite
<quantity id="4CE69B7FFF59CDB0FC92EED7B2A1EFF3" box="[841,978,1698,1723]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="142" pageNumber="143" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.41.6 mm</quantity>
.
</paragraph>
<paragraph id="8BA1369AFF59CDB0FF1CEEB2B25CEF96" blockId="142.[151,1436,1122,2011]" box="[199,815,1734,1759]" pageId="142" pageNumber="143">
COLORATION (
<figureCitation id="13252A1FFF59CDB0FEB7EEB3B0BEEF96" box="[364,461,1734,1759]" captionStart="FIGURE 118" captionStartId="143.[151,250,1932,1954]" captionTargetBox="[188,1398,337,1911]" captionTargetId="figure@143.[188,1398,337,1911]" captionTargetPageId="143" captionText="FIGURE 118. Acanthophorella barjadzei gen. et sp. nov., habitus. A. Paratype female. Scale line = 1 mm. B. Holotype male (photo S. Barjadze), without scale." httpUri="https://zenodo.org/record/272867/files/figure.png" pageId="142" pageNumber="143">Fig. 118</figureCitation>
). Pigmentless, yellowish white.
</paragraph>
<paragraph id="8BA1369AFF59CDB0FF1CEE9EB517EE92" blockId="142.[151,1436,1122,2011]" pageId="142" pageNumber="143">
HEAD. With a longitudinal depression in male. Labrum with three medial teeth and 3+2 labral and 2+2 supralabral setae. Promentum triangular, without setae. Lingual plates with 10+9 setae, on each plate arranged in three irregular rows. Stipites with
<emphasis id="B96AEA88FF59CDB0FDC3EF41B340EE02" box="[536,563,1844,1867]" italics="true" pageId="142" pageNumber="143">ca</emphasis>
32+32 setae. Antennae elongated and slender in both sexes,
<quantity id="4CE69B7FFF59CDB0FB3AEF47B436EE03" box="[1249,1349,1842,1867]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" pageId="142" pageNumber="143" unit="mm" value="3.05">3.05 mm</quantity>
long in
<typeStatus id="54A58838FF59CDB0FF4CEF23B18EEE26" box="[151,253,1878,1903]" pageId="142" pageNumber="143" type="holotype">holotype</typeStatus>
. Length of antennomeres (in mm): I (0.15), II (0.34), III (0.83), IV (0.45), V (0.75), VI (0.27), VII (0.21) and VIII (0.05). Length/breadth ratios of antennomeres IVII: I (1), II (2), III (6), IV (3), V (5), VI (1.5) and VII (1.6). Antennomeres II, IV, V, VI and VII with one, three, one, four and one sensillum, respectively. Number of ocelli 6, arranged in 2 irregular rows in
<typeStatus id="54A58838FF59CDB0FD8FEFB7B3C9EE92" box="[596,698,1986,2011]" pageId="142" pageNumber="143" type="holotype">holotype</typeStatus>
;
<quantity id="4CE69B7FFF59CDB0FD1EEFB7B27FEE92" box="[709,780,1986,2011]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="142" pageNumber="143" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">57 in</quantity>
23 irregular rows in females.
</paragraph>
<paragraph id="8BA1369AFF58CDB1FF1CE8EDB189E99D" blockId="143.[151,1436,151,285]" pageId="143" pageNumber="144">COLLUM. Narrower than head, with six macrochaetae. Anterior edge semi-circular, posterior margin gently concave.</paragraph>
<paragraph id="8BA1369AFF58CDB1FF1CE895B4E3E854" blockId="143.[151,1436,151,285]" pageId="143" pageNumber="144">
BODY SEGMENTS (
<figureCitation id="13252A1FFF58CDB1FE72E8AAB367E9B1" box="[425,532,223,248]" captionStart="FIGURE 118" captionStartId="143.[151,250,1932,1954]" captionTargetBox="[188,1398,337,1911]" captionTargetId="figure@143.[188,1398,337,1911]" captionTargetPageId="143" captionText="FIGURE 118. Acanthophorella barjadzei gen. et sp. nov., habitus. A. Paratype female. Scale line = 1 mm. B. Holotype male (photo S. Barjadze), without scale." httpUri="https://zenodo.org/record/272867/files/figure.png" pageId="143" pageNumber="144">Fig. 118</figureCitation>
). Lateral keels moderate. Macrochaetae relatively long and trichoid. CIX (pleurotergite 15) ~ 0.6; MIX (pleurotergite 15) ~ 1.8; PIX (pleurotergite 15) ~ 0.5; MA (pleurotergite 15) ~ 115˚.
</paragraph>
<caption id="DF616612FF58CDB1FF4CEFF9B085EE88" httpUri="https://zenodo.org/record/272867/files/figure.png" pageId="143" pageNumber="144" targetBox="[188,1398,337,1911]" targetPageId="143">
<paragraph id="8BA1369AFF58CDB1FF4CEFF9B085EE88" blockId="143.[151,1436,1932,1985]" pageId="143" pageNumber="144">
<emphasis id="B96AEA88FF58CDB1FF4CEFF9B05AEEE8" bold="true" box="[151,297,1932,1954]" pageId="143" pageNumber="144">FIGURE 118.</emphasis>
<taxonomicName id="4C1E4D19FF58CDB1FEE9EFF9B330EEEB" box="[306,579,1932,1954]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="143" pageNumber="144" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B96AEA88FF58CDB1FEE9EFF9B330EEEB" box="[306,579,1932,1954]" italics="true" pageId="143" pageNumber="144">Acanthophorella barjadzei</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF58CDB1FD92EFF8B39BEEEB" bold="true" box="[585,744,1933,1954]" pageId="143" pageNumber="144">gen. et sp. nov.</emphasis>
, habitus.
<emphasis id="B96AEA88FF58CDB1FC96EFF9B215EEE8" bold="true" box="[845,870,1932,1953]" pageId="143" pageNumber="144">A.</emphasis>
Paratype female. Scale line = 1 mm.
<emphasis id="B96AEA88FF58CDB1FB3CEFF8B58DEEEB" bold="true" box="[1255,1278,1933,1954]" pageId="143" pageNumber="144">B.</emphasis>
Holotype male (photo S. Barjadze), without scale.
</paragraph>
</caption>
<caption id="DF616612FF47CDAEFF4CEFE8B073EE9B" httpUri="https://zenodo.org/record/272868/files/figure.png" pageId="144" pageNumber="145" targetBox="[188,1398,193,1927]" targetPageId="144">
<paragraph id="8BA1369AFF47CDAEFF4CEFE8B073EE9B" blockId="144.[151,1436,1949,2002]" pageId="144" pageNumber="145">
<emphasis id="B96AEA88FF47CDAEFF4CEFE8B05AEEFB" bold="true" box="[151,297,1949,1971]" pageId="144" pageNumber="145">FIGURE 119.</emphasis>
<taxonomicName id="4C1E4D19FF47CDAEFEEAEFE8B333EEFA" box="[305,576,1949,1971]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="144" pageNumber="145" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B96AEA88FF47CDAEFEEAEFE8B333EEFA" box="[305,576,1949,1971]" italics="true" pageId="144" pageNumber="145">Acanthophorella barjadzei</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF47CDAEFD9DEFE8B390EEFB" bold="true" box="[582,739,1949,1970]" pageId="144" pageNumber="145">gen. et sp. nov.</emphasis>
, holotype male.
<emphasis id="B96AEA88FF47CDAEFC57EFE8B2D6EEFB" bold="true" box="[908,933,1949,1970]" pageId="144" pageNumber="145">A.</emphasis>
Leg 3.
<emphasis id="B96AEA88FF47CDAEFC29EFE8B57AEEFB" bold="true" box="[1010,1033,1949,1970]" pageId="144" pageNumber="145">B.</emphasis>
Leg 4.
<emphasis id="B96AEA88FF47CDAEFB8CEFE8B51CEEFA" bold="true" box="[1111,1135,1949,1971]" pageId="144" pageNumber="145">C.</emphasis>
Leg 7.
<emphasis id="B96AEA88FF47CDAEFB67EFE8B5A6EEFB" bold="true" box="[1212,1237,1949,1970]" pageId="144" pageNumber="145">D.</emphasis>
Leg 10. Scale lines = 0.2 mm.
</paragraph>
</caption>
<caption id="DF616612FF46CDAFFF4CEF26B0B2EEAA" httpUri="https://zenodo.org/record/272869/files/figure.png" pageId="145" pageNumber="146" targetBox="[272,1245,214,1817]" targetPageId="145">
<paragraph id="8BA1369AFF46CDAFFF4CEF26B0B2EEAA" blockId="145.[151,1436,1875,2019]" pageId="145" pageNumber="146">
<emphasis id="B96AEA88FF46CDAFFF4CEF26B05EEE21" bold="true" box="[151,301,1875,1897]" pageId="145" pageNumber="146">FIGURE 120.</emphasis>
<taxonomicName id="4C1E4D19FF46CDAFFEEFEF26B336EE20" box="[308,581,1875,1897]" class="Diplopoda" family="Anthroleucosomatidae" genus="Acanthophorella" kingdom="Animalia" order="Chordeumatida" pageId="145" pageNumber="146" phylum="Arthropoda" rank="species" species="barjadzei">
<emphasis id="B96AEA88FF46CDAFFEEFEF26B336EE20" box="[308,581,1875,1897]" italics="true" pageId="145" pageNumber="146">Acanthophorella barjadzei</emphasis>
</taxonomicName>
<emphasis id="B96AEA88FF46CDAFFD97EF26B39EEE21" bold="true" box="[588,749,1875,1896]" pageId="145" pageNumber="146">gen. et sp. nov.</emphasis>
, holotype male gonopods.
<emphasis id="B96AEA88FF46CDAFFBDFEF26B56FEE21" bold="true" box="[1028,1052,1875,1896]" pageId="145" pageNumber="146">A.</emphasis>
Anterior gonopods, anterior view.
<emphasis id="B96AEA88FF46CDAFFA5FEF26B4E8EE21" bold="true" box="[1412,1435,1875,1896]" pageId="145" pageNumber="146">B.</emphasis>
Anterior gonopods, posterior view.
<emphasis id="B96AEA88FF46CDAFFDDFEF04B36FEECE" bold="true" box="[516,540,1905,1927]" pageId="145" pageNumber="146">C.</emphasis>
Anterior gonopods, lateral view.
<emphasis id="B96AEA88FF46CDAFFCADEF07B2FCEECE" bold="true" box="[886,911,1906,1927]" pageId="145" pageNumber="146">D.</emphasis>
Posterior gonopods, posterior view.
<emphasis id="B96AEA88FF46CDAFFAD2EF07B467EECE" box="[1289,1300,1906,1927]" italics="true" pageId="145" pageNumber="146">c</emphasis>
= coxite;
<emphasis id="B96AEA88FF46CDAFFAA4EF07B4F5EECE" box="[1407,1414,1906,1927]" italics="true" pageId="145" pageNumber="146">t</emphasis>
= telopodite;
<emphasis id="B96AEA88FF46CDAFFED2EFE4B052EEEF" box="[265,289,1937,1958]" italics="true" pageId="145" pageNumber="146">cp</emphasis>
= coxal process;
<emphasis id="B96AEA88FF46CDAFFE0AEFE4B094EEEF" box="[465,487,1937,1958]" italics="true" pageId="145" pageNumber="146">cv</emphasis>
= coxal vesicle;
<emphasis id="B96AEA88FF46CDAFFD4AEFE5B3D6EEEF" box="[657,677,1936,1958]" italics="true" pageId="145" pageNumber="146">hl</emphasis>
= hairy lever;
<emphasis id="B96AEA88FF46CDAFFCE3EFE5B235EEEF" box="[824,838,1936,1958]" italics="true" pageId="145" pageNumber="146">ll</emphasis>
= lateral lamella;
<emphasis id="B96AEA88FF46CDAFFC26EFE5B562EEEF" box="[1021,1041,1936,1958]" italics="true" pageId="145" pageNumber="146">lo</emphasis>
= anterior lobe;
<emphasis id="B96AEA88FF46CDAFFB63EFE4B5A4EEEF" box="[1208,1239,1937,1958]" italics="true" pageId="145" pageNumber="146">mp</emphasis>
= medial shield;
<emphasis id="B96AEA88FF46CDAFFA5EEFE4B4E8EEEF" box="[1413,1435,1937,1958]" italics="true" pageId="145" pageNumber="146">sp</emphasis>
= sternal plate;
<emphasis id="B96AEA88FF46CDAFFEEFEFC5B03BEE8C" box="[308,328,1968,1989]" italics="true" pageId="145" pageNumber="146">ss</emphasis>
= sternal sac;
<emphasis id="B96AEA88FF46CDAFFE02EFC5B080EE8C" box="[473,499,1968,1989]" italics="true" pageId="145" pageNumber="146">sw</emphasis>
= widening;
<emphasis id="B96AEA88FF46CDAFFDA0EFC5B3FCEE8C" box="[635,655,1968,1989]" italics="true" pageId="145" pageNumber="146">tp</emphasis>
= thorn-like process;
<emphasis id="B96AEA88FF46CDAFFCB7EFC5B2F1EE8C" box="[876,898,1968,1989]" italics="true" pageId="145" pageNumber="146">us</emphasis>
= undefined structure;
<emphasis id="B96AEA88FF46CDAFFBB4EFC5B5E1EE8C" box="[1135,1170,1968,1989]" italics="true" pageId="145" pageNumber="146">ppr</emphasis>
= posterior projection;
<emphasis id="B96AEA88FF46CDAFFA5AEFC5B4EFEE8C" box="[1409,1436,1968,1989]" italics="true" pageId="145" pageNumber="146">str</emphasis>
= striae. Scale line = 0.3 mm.
</paragraph>
</caption>
<paragraph id="8BA1369AFF45CDACFF1CE8EDB39FE99D" blockId="146.[151,1437,151,1112]" pageId="146" pageNumber="147">TELSON. Epiproct with a pair of spinnerets and 3+3 setae (1+1 paramedian, 2+2 marginal). Hypoproct with 1+1 apical setae. Paraprocts with 3+3 marginal setae.</paragraph>
<paragraph id="8BA1369AFF45CDACFF1CE895B581E855" blockId="146.[151,1437,151,1112]" pageId="146" pageNumber="147">WALKING LEGS. Elongated and slender. In both sexes, leg-pairs 1 and 2 with tarsal combs; prefemora with several long and robust setae; femora and postfemora with a group of several long and robust setae.</paragraph>
<paragraph id="8BA1369AFF45CDACFF1CE95DB0B6E8BC" blockId="146.[151,1437,151,1112]" pageId="146" pageNumber="147">
MALE SEXUAL CHARACTERS (
<figureCitation id="13252A1FFF45CDACFDCDE952B30BE809" box="[534,632,295,320]" captionStart="FIGURE 119" captionStartId="144.[151,250,1949,1971]" captionTargetBox="[188,1398,193,1927]" captionTargetId="figure@144.[188,1398,193,1927]" captionTargetPageId="144" captionText="FIGURE 119. Acanthophorella barjadzei gen. et sp. nov., holotype male. A. Leg 3. B. Leg 4. C. Leg 7. D. Leg 10. Scale lines = 0.2 mm." httpUri="https://zenodo.org/record/272868/files/figure.png" pageId="146" pageNumber="147">Fig. 119</figureCitation>
). Leg-pairs 37 enlarged, especially so leg-pairs 3, 4 and 7. Leg-pairs 3 and 4 each with a basal exterior protrusion on prefemur. Leg-pair 5 with a basal, subtriangular, oral protrusion on prefemur. Leg-pair 6 with a basal, exterior, small protrusion on prefemur. Leg-pair 7 with a characteristic, very wide, posteriorly orientated, lamellar, coxal process, acuminate mesally and rounded exteriorly. Leg-pair 10 with coxal glands and with a coxal subtriangular protrusion orientated posteriorly. Leg-pair 11 with coxal glands, without other peculiarities.
</paragraph>
<paragraph id="8BA1369AFF45CDACFF1CEA75B5DDEA5D" blockId="146.[151,1437,151,1112]" pageId="146" pageNumber="147">
ANTERIOR GONOPODS (
<figureCitation id="13252A1FFF45CDACFE0AE98AB340EB51" box="[465,563,511,536]" captionStart="FIGURE 120" captionStartId="145.[151,250,1875,1897]" captionTargetBox="[272,1245,214,1817]" captionTargetId="figure@145.[226,1351,193,1879]" captionTargetPageId="145" captionText="FIGURE 120. Acanthophorella barjadzei gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. c = coxite; t = telopodite; cp = coxal process; cv = coxal vesicle; hl = hairy lever; ll = lateral lamella; lo = anterior lobe; mp = medial shield; sp = sternal plate; ss = sternal sac; sw = widening; tp = thorn-like process; us = undefined structure; ppr = posterior projection; str = striae. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272869/files/figure.png" pageId="146" pageNumber="147">Fig. 120</figureCitation>
AC). Sternal plate (
<emphasis id="B96AEA88FF45CDACFCF9EA74B248EB51" box="[802,827,513,536]" italics="true" pageId="146" pageNumber="147">sp</emphasis>
) anteriorly with a small, medial, nippled, sternal sac (
<emphasis id="B96AEA88FF45CDACFF7BEA50B1C5EB75" box="[160,182,549,572]" italics="true" pageId="146" pageNumber="147">ss</emphasis>
). At base of gonopods, two anterior lobes (
<emphasis id="B96AEA88FF45CDACFD79EA51B3CBEB74" box="[674,696,548,573]" italics="true" pageId="146" pageNumber="147">lo</emphasis>
) can be seen. Medial parts characterized by the presence of welldeveloped and wide medial shields (
<emphasis id="B96AEA88FF45CDACFDE3EA3CB329EB29" box="[568,602,585,608]" italics="true" pageId="146" pageNumber="147">mp</emphasis>
) with mesal edges folded inside and forming posterior projections (
<emphasis id="B96AEA88FF45CDACFABFEA3CB4F9EB29" box="[1380,1418,585,608]" italics="true" pageId="146" pageNumber="147">ppr</emphasis>
). Shields with numerous and very thin striae (
<emphasis id="B96AEA88FF45CDACFD46EA18B3C9EBCD" box="[669,698,621,644]" italics="true" pageId="146" pageNumber="147">str</emphasis>
) anteriorly. Lateral sides of shields with a small widening (
<emphasis id="B96AEA88FF45CDACFAAEEA18B4E1EBCD" box="[1397,1426,621,644]" italics="true" pageId="146" pageNumber="147">sw</emphasis>
) which covers medial parts of slender, thorn-like processes (
<emphasis id="B96AEA88FF45CDACFC95EAE4B217EBE1" box="[846,868,657,680]" italics="true" pageId="146" pageNumber="147">tp</emphasis>
). The lateralmost parts of anterior gonopods are lateral lamellae (
<emphasis id="B96AEA88FF45CDACFE87EAC1B018EB84" box="[348,363,692,717]" italics="true" pageId="146" pageNumber="147">ll</emphasis>
) with small nipples. These lamellae almost as high as apex of lateral widening of shields. Posteriorly, anterior gonopods with a pair of mesally situated coxal vesicles (
<emphasis id="B96AEA88FF45CDACFBD8EAACB56FEBB9" box="[1027,1052,729,752]" italics="true" pageId="146" pageNumber="147">cv</emphasis>
), with well-developed and highly hairy levers (
<emphasis id="B96AEA88FF45CDACFEF0EA89B032EA5C" box="[299,321,764,789]" italics="true" pageId="146" pageNumber="147">hl</emphasis>
) situated laterally to coxal vesicles, as well as with two lateralmost lobes (
<emphasis id="B96AEA88FF45CDACFB5EEA88B5EDEA5D" box="[1157,1182,765,788]" italics="true" pageId="146" pageNumber="147">us</emphasis>
).
</paragraph>
<paragraph id="8BA1369AFF45CDACFF1CEB55B1B8EAC9" blockId="146.[151,1437,151,1112]" pageId="146" pageNumber="147">
POSTERIOR GONOPODS (
<figureCitation id="13252A1FFF45CDACFE0EEB6AB34BEA71" box="[469,568,799,824]" captionStart="FIGURE 120" captionStartId="145.[151,250,1875,1897]" captionTargetBox="[272,1245,214,1817]" captionTargetId="figure@145.[226,1351,193,1879]" captionTargetPageId="145" captionText="FIGURE 120. Acanthophorella barjadzei gen. et sp. nov., holotype male gonopods. A. Anterior gonopods, anterior view. B. Anterior gonopods, posterior view. C. Anterior gonopods, lateral view. D. Posterior gonopods, posterior view. c = coxite; t = telopodite; cp = coxal process; cv = coxal vesicle; hl = hairy lever; ll = lateral lamella; lo = anterior lobe; mp = medial shield; sp = sternal plate; ss = sternal sac; sw = widening; tp = thorn-like process; us = undefined structure; ppr = posterior projection; str = striae. Scale line = 0.3 mm." httpUri="https://zenodo.org/record/272869/files/figure.png" pageId="146" pageNumber="147">Fig. 120</figureCitation>
D). Coxites (
<emphasis id="B96AEA88FF45CDACFD13EB54B3A7EA71" box="[712,724,801,824]" italics="true" pageId="146" pageNumber="147">c</emphasis>
) divided. Telopodites (
<emphasis id="B96AEA88FF45CDACFC07EB54B297EA71" box="[988,996,801,824]" italics="true" pageId="146" pageNumber="147">t</emphasis>
) small, present on posterolateral sides. Coxal processes (
<emphasis id="B96AEA88FF45CDACFE86EB30B004EA15" box="[349,375,837,860]" italics="true" pageId="146" pageNumber="147">cp</emphasis>
) orientated laterally and their apices curved anteriorly. Coxal vesicles (
<emphasis id="B96AEA88FF45CDACFB44EB30B5CBEA15" box="[1183,1208,837,860]" italics="true" pageId="146" pageNumber="147">cv</emphasis>
) present on anterior side.
</paragraph>
</subSubSection>
<subSubSection id="C3046511FF45CDACFF1CEBF9B055ED11" pageId="146" pageNumber="147" type="distribution">
<paragraph id="8BA1369AFF45CDACFF1CEBF9B528EAEC" blockId="146.[151,1437,151,1112]" box="[199,1115,908,933]" pageId="146" pageNumber="147">
<emphasis id="B96AEA88FF45CDACFF1CEBF9B013EAEC" bold="true" box="[199,352,908,933]" pageId="146" pageNumber="147">Distribution.</emphasis>
<collectingRegion id="49DAF878FF45CDACFEBCEBF9B0B2EAEC" box="[359,449,908,933]" country="United States of America" name="Georgia" pageId="146" pageNumber="147">
<collectingCountry id="F309760AFF45CDACFEBCEBF9B0B2EAEC" box="[359,449,908,933]" name="Georgia" pageId="146" pageNumber="147">Georgia</collectingCountry>
</collectingRegion>
(known only from Cave Dolabistavi) (
<figureCitation id="13252A1FFF45CDACFCAEEBF9B2A6EAEC" box="[885,981,908,933]" captionStart="FIGURE 169" captionStartId="197.[151,250,1981,2003]" captionTargetBox="[207,1379,1136,1959]" captionTargetId="figure@197.[207,1379,1136,1959]" captionTargetPageId="197" captionText="FIGURE 169. Distribution of the Caucasian Anthroleucosomatidae: the Flagellophorella complex." httpUri="https://zenodo.org/record/272918/files/figure.png" pageId="146" pageNumber="147">Fig. 169</figureCitation>
, red circle).
</paragraph>
<paragraph id="8BA1369AFF45CDACFF1CEBDAB055ED11" blockId="146.[151,1437,151,1112]" pageId="146" pageNumber="147">
<emphasis id="B96AEA88FF45CDACFF1CEBDAB062EA81" bold="true" box="[199,273,943,968]" pageId="146" pageNumber="147">Notes.</emphasis>
This species is known only from a cave and shows some troglomorphic features such as a pigmentless, yellowish white body, a reduced number of ocelli, and elongated and slender antennae and walking legs. This species probably inhabits some other caves in the Racha region. One female and two juveniles probably of this species were collected in Cave Muradis, Racha region, but in the absence of an adult male, these specimens remain unidentified.
</paragraph>
</subSubSection>
</treatment>
</document>