treatments-xml/data/03/E1/AF/03E1AF0CFFABFFECFF76FB725050C503.xml

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<mods:title id="48BBDB1A57B5586AD2CA097ED47C02C6">Two new species of Domene Fauvel, 1873 (Coleoptera: Staphylinidae: Paederinae) from Portugal and faunistic notes</mods:title>
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<mods:namePart id="4B8CC0A23BCB1D46185A958A3B9282AF">Serrano, Artur R. M.</mods:namePart>
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<mods:namePart id="6FE9F3B76DDD9F1012D0BC2337BEA04C">Aguiar, Carlos A. S.</mods:namePart>
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<treatment id="03E1AF0CFFABFFECFF76FB725050C503" ID-DOI="http://doi.org/10.5281/zenodo.14963690" ID-GBIF-Taxon="254987536" ID-Zenodo-Dep="14963690" LSID="urn:lsid:plazi:treatment:03E1AF0CFFABFFECFF76FB725050C503" httpUri="http://treatment.plazi.org/id/03E1AF0CFFABFFECFF76FB725050C503" lastPageId="14" lastPageNumber="451" pageId="10" pageNumber="447" scope_family="Staphylinidae" scope_order="Coleoptera" scope_subFamily="Paederinae">
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<paragraph id="8BF71E1AFFABFFE8FF76FB7250B0C4DA" blockId="10.[151,904,1230,1257]" box="[151,904,1230,1257]" pageId="10" pageNumber="447">
<heading id="D0BFA976FFABFFE8FF76FB7250B0C4DA" bold="true" box="[151,904,1230,1257]" fontSize="11" level="1" pageId="10" pageNumber="447" reason="1">
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<taxonomicName id="4C486599FFABFFE8FF76FB725156C4DA" authority="Serrano" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[151,622,1230,1257]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="447" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov." subGenus="Lathromene">
<emphasis id="B93CC208FFABFFE8FF76FB7253C4C4DB" bold="true" box="[151,252,1231,1257]" italics="true" pageId="10" pageNumber="447">Domene</emphasis>
(
<emphasis id="B93CC208FFABFFE8FEECFB735299C4DB" bold="true" box="[269,417,1230,1257]" italics="true" pageId="10" pageNumber="447">Lathromene</emphasis>
) 
<emphasis id="B93CC208FFABFFE8FE52FB735138C4DB" bold="true" box="[435,512,1230,1257]" italics="true" pageId="10" pageNumber="447">pedroi</emphasis>
Serrano
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, 
<taxonomicNameLabel id="A20F7F73FFABFFE8FD9BFB7251E0C4DB" box="[634,728,1231,1257]" pageId="10" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1373029FFFABFFE8FD06FB73507CC4DA" box="[743,836,1230,1256]" captionStart="FIGURE 1" captionStartId="4.[152,255,1905,1929]" captionTargetBox="[151,1436,306,1862]" captionTargetId="figure-73@4.[151,1436,306,1879]" captionTargetPageId="4" captionText="FIGURE 1. (a), habitat of Domene boavidae sp. nov. near S. Pedro (Vila Viçosa, Viseu district); (b), D. boavidae sp. nov. male adult; (c), habitus of D. boavidae sp. nov. (dorsal view), holotype male; (d), habitus of Domene pedroi sp. nov. (dorsal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.14896598" httpUri="https://zenodo.org/record/14896598/files/figure.png" pageId="10" pageNumber="447">Figs 1d</figureCitation>
, 
<figureCitation id="1373029FFFABFFE8FCB3FB72505AC4DB" box="[850,866,1231,1257]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="10" pageNumber="447">5</figureCitation>
, 
<figureCitation id="1373029FFFABFFE8FC91FB735047C4DA" box="[880,895,1230,1256]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="10" pageNumber="447">6</figureCitation>
)
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BF71E1AFFABFFE8FF76FAAA5711C54B" blockId="10.[151,1437,1302,1401]" pageId="10" pageNumber="447">
<emphasis id="B93CC208FFABFFE8FF76FAAA527CC502" bold="true" box="[151,324,1302,1329]" pageId="10" pageNumber="447">Type material.</emphasis>
<materialsCitation id="3B201447FFABFFE8FEA8FAAA571DC54B" ID-GBIF-Occurrence="5074337302" collectingDate="2008-04-03" collectorName="A. Serrano &amp; C. Aguiar" country="Portugal" determinerName="A. Serrano" elevation="635" latitude="40.027225" location="Aldeia Cimeira" longLatPrecision="1" longitude="-7.972947" municipality="Pampilhosa da Serra" pageId="10" pageNumber="447" specimenCount="1" specimenCount-male="1" stateProvince="Distrito Coimbra" typeStatus="holotype">
<typeStatus id="54F3A0B8FFABFFE8FEA8FAAA5289C503" box="[329,433,1303,1329]" pageId="10" pageNumber="447">Holotype</typeStatus>
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: “ 
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(
<collectingMunicipality id="6B938460FFABFFE8FD5EFAAA509DC503" box="[703,933,1303,1329]" pageId="10" pageNumber="447">Pampilhosa da Serra</collectingMunicipality>
)\ 
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, 
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\ 
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<collectingDate id="EFB2C132FFABFFE8FB10FAAA5666C503" box="[1265,1374,1303,1329]" pageId="10" pageNumber="447" value="2008-04-03">3.IV.2008</collectingDate>
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\ Col. A. Serrano”\\ “ 
<collectorName id="26BD7BCCFFABFFE8FEA1FA865283C567" box="[320,443,1339,1365]" pageId="10" pageNumber="447">A. Serrano</collectorName>
&amp;\ 
<collectorName id="26BD7BCCFFABFFE8FE09FA86516EC567" box="[488,598,1339,1365]" pageId="10" pageNumber="447">C. Aguiar</collectorName>
leg.”\\ “ 
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\ 
<geoCoordinate id="EE7C78DDFFABFFE8FC9FFA865704C567" box="[894,1084,1338,1365]" degrees="7" direction="west" minutes="58" orientation="longitude" pageId="10" pageNumber="447" precision="1" seconds="22.61" value="-7.972947">7º 58´22.61´´ W</geoCoordinate>
\ 
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alt.”\\ “Holotype male\ 
<taxonomicName id="4C486599FFABFFE8FF76FAE253CCC54A" authorityName="Fauvel" authorityYear="1873" box="[151,244,1375,1400]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="447" phylum="Arthropoda" rank="genus">
<emphasis id="B93CC208FFABFFE8FF76FAE253CCC54A" box="[151,244,1375,1400]" italics="true" pageId="10" pageNumber="447">Domene</emphasis>
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\ (
<taxonomicName id="4C486599FFABFFE8FEF5FAE252A2C54B" authorityName="Koch" authorityYear="1938" box="[276,410,1375,1401]" pageId="10" pageNumber="454" rank="subGenus" subGenus="Lathromene">
<emphasis id="B93CC208FFABFFE8FEF5FAE252A2C54B" box="[276,410,1375,1401]" italics="true" pageId="10" pageNumber="447">Lathromene</emphasis>
</taxonomicName>
) \ 
<taxonomicName id="4C486599FFABFFE8FE58FAE25139C54B" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[441,513,1375,1401]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="447" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFABFFE8FE58FAE25139C54B" box="[441,513,1375,1401]" italics="true" pageId="10" pageNumber="447">pedroi</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="A20F7F73FFABFFE8FDE9FAE2515AC54B" box="[520,610,1375,1401]" pageId="10" pageNumber="447" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
\ 
<determinerName id="7E2BF2DEFFABFFE8FD8EFAE251D0C54B" box="[623,744,1375,1401]" pageId="10" pageNumber="447">A. Serrano</determinerName>
det. 2024” [t] [h] [red card]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3524D91FFABFFE8FF76FA76510CC647" pageId="10" pageNumber="447" type="etymology">
<paragraph id="8BF71E1AFFABFFE8FF76FA765226C5D7" blockId="10.[151,286,1483,1509]" box="[151,286,1483,1509]" pageId="10" pageNumber="447">
<emphasis id="B93CC208FFABFFE8FF76FA765226C5D7" bold="true" box="[151,286,1483,1509]" pageId="10" pageNumber="447">Etymology</emphasis>
</paragraph>
<paragraph id="8BF71E1AFFABFFE8FF76F9AE510CC647" blockId="10.[151,1437,1555,2013]" pageId="10" pageNumber="447">This new species is dedicated to Pedro Martins da Silva, a master’s and doctoral student of the first author and a renowned ecologist, who would like to have been a staphylinid taxonomist, but whose scientific career path led him to study insects encoded in numbers.</paragraph>
</subSubSection>
<subSubSection id="C3524D91FFABFFE9FF26F9C35038C53F" lastPageId="11" lastPageNumber="448" pageId="10" pageNumber="447" type="description">
<paragraph id="8BF71E1AFFABFFE8FF26F9C351B4C68F" blockId="10.[151,1437,1555,2013]" pageId="10" pageNumber="447">
<emphasis id="B93CC208FFABFFE8FF26F9C35261C6AA" bold="true" box="[199,345,1662,1688]" pageId="10" pageNumber="447">Description.</emphasis>
Microphthalmous, wingless and elongate body (
<figureCitation id="1373029FFFABFFE8FC91F9C250F8C6AB" box="[880,960,1663,1689]" captionStart="FIGURE 1" captionStartId="4.[152,255,1905,1929]" captionTargetBox="[151,1436,306,1862]" captionTargetId="figure-73@4.[151,1436,306,1879]" captionTargetPageId="4" captionText="FIGURE 1. (a), habitat of Domene boavidae sp. nov. near S. Pedro (Vila Viçosa, Viseu district); (b), D. boavidae sp. nov. male adult; (c), habitus of D. boavidae sp. nov. (dorsal view), holotype male; (d), habitus of Domene pedroi sp. nov. (dorsal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.14896598" httpUri="https://zenodo.org/record/14896598/files/figure.png" pageId="10" pageNumber="447">Fig. 1d</figureCitation>
). Coloration: body yellowish and abdomen slightly darker. Body length: 
<quantity id="4CB0B3FFFFABFFE8FE3EF91F510DC68F" box="[479,565,1698,1725]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="447" unit="mm" value="4.5">4.5 mm</quantity>
(male).
</paragraph>
<paragraph id="8BF71E1AFFABFFE9FF26F97A5744C1BB" blockId="10.[151,1437,1555,2013]" lastBlockId="11.[151,1437,150,1293]" lastPageId="11" lastPageNumber="448" pageId="10" pageNumber="447">
<emphasis id="B93CC208FFABFFE8FF26F97A523FC6D3" bold="true" box="[199,263,1735,1761]" pageId="10" pageNumber="447">Head</emphasis>
(
<figureCitation id="1373029FFFABFFE8FEF5F97A52BCC6D3" box="[276,388,1734,1761]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="10" pageNumber="447">Figs 5a–c</figureCitation>
): subquadrate/orbicular, 1.1 times longer than wide (length: 
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, width: 
<quantity id="4CB0B3FFFFABFFE8FB30F97A560FC6D3" box="[1233,1335,1735,1761]" metricMagnitude="-4" metricUnit="m" metricValue="6.8" pageId="10" pageNumber="447" unit="mm" value="0.68">0.68 mm</quantity>
), slightly wider than pronotum (1.2 times); eyes strongly reduced, with 7 ommatidia (
<figureCitation id="1373029FFFABFFE8FC30F9565718C737" box="[977,1056,1770,1797]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="10" pageNumber="447">Fig. 5c</figureCitation>
); long and thin seta in supraocular, small circular depression (supraocular trichobothrium); absence of patch of minor pubescence above depression and behind eyes (
<figureCitation id="1373029FFFABFFE8FECFF88E52B8C77F" box="[302,384,1842,1869]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="10" pageNumber="447">Fig. 5c</figureCitation>
); two dark spots in vertex marking insertions of dorsal arms of tentorium; gular sutures well defined, diverging slightly towards neck (
<figureCitation id="1373029FFFABFFE8FD8CF8EA51FAC743" box="[621,706,1878,1905]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="10" pageNumber="447">Fig. 5b</figureCitation>
); dorsal surface without any isodiametric microreticulation, with well-defined coarse and dense punctuation, similar to that of pronotum, on lateral sides and vertex; frons with sparse punctuation, almost smooth (
<figureCitation id="1373029FFFABFFE8FE3DF8225116C78B" box="[476,558,1950,1977]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="10" pageNumber="447">Fig. 5a</figureCitation>
); antennae almost moniliform (
<figureCitation id="1373029FFFABFFE8FC6FF82250E7C78B" box="[910,991,1951,1977]" captionStart="FIGURE 1" captionStartId="4.[152,255,1905,1929]" captionTargetBox="[151,1436,306,1862]" captionTargetId="figure-73@4.[151,1436,306,1879]" captionTargetPageId="4" captionText="FIGURE 1. (a), habitat of Domene boavidae sp. nov. near S. Pedro (Vila Viçosa, Viseu district); (b), D. boavidae sp. nov. male adult; (c), habitus of D. boavidae sp. nov. (dorsal view), holotype male; (d), habitus of Domene pedroi sp. nov. (dorsal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.14896598" httpUri="https://zenodo.org/record/14896598/files/figure.png" pageId="10" pageNumber="447">Fig. 1d</figureCitation>
), 
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long, not reaching the base of pronotum when directed backwards; all antennomeres slightly longer than wide, the 1 
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longest of all (length: 
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); 2 
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antennomere slightly longer than the 3 
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to 10 
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antennomeres (1.2–1.3 times); 3 
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to 10 
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antennomeres subequal and the 11 
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is 1.3‒1.4 times larger than the size of each of the previous eight antennomeres; labrum deeply emarginated with 17 large setae spread over the disk and the anterior margin (
<figureCitation id="1373029FFFAAFFE9FC18FF625773C0CB" box="[1017,1099,222,249]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">Fig. 5a</figureCitation>
); mandibles symmetrical with four distinct teeth in the inner edge, two of them are more pronounced; maxillary palpus with four palpomeres, second and third subequal, more than 3 times as long as broad, apical palpomere reduced and conical; internal lobe of maxilla with clusters of setae; labium with bilobed glossae, circumscribed by hairy paraglossae; labial palpus with three palpomeres, the 2 
<superScript id="7C3DB352FFAAFFE9FE34FED352DDC14E" attach="left" box="[469,485,366,380]" fontSize="6" pageId="11" pageNumber="448">nd</superScript>
being the longest and more robust, and the 3 
<superScript id="7C3DB352FFAAFFE9FC3CFED350D3C14E" attach="left" box="[989,1003,366,380]" fontSize="6" pageId="11" pageNumber="448">rd</superScript>
the thinnest.
</paragraph>
<paragraph id="8BF71E1AFFAAFFE9FF26FE2E527FC20F" blockId="11.[151,1437,150,1293]" pageId="11" pageNumber="448">
<emphasis id="B93CC208FFAAFFE9FF26FE2E5278C19F" bold="true" box="[199,320,403,429]" pageId="11" pageNumber="448">Pronotum</emphasis>
(
<figureCitation id="1373029FFFAAFFE9FEB3FE2E5290C19F" box="[338,424,402,429]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">Fig. 5d</figureCitation>
): 1.2 times longer than wide (length: 
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, width: 
<quantity id="4CB0B3FFFFAAFFE9FBC0FE2F57B3C19F" box="[1057,1163,402,429]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="11" pageNumber="448" unit="mm" value="0.59">0.59 mm</quantity>
), slightly narrower than head, widest anteriorly and distinctly tapering posteriad; anterior angles weakened, posterior angles marked but largely obtuse; basal margin well defined; punctuation similar to that of head, evenly distributed except for a smooth midline ranging from the anterior to the posterior parts; absence of any middle sulcus, interstices without microsculpture.
</paragraph>
<paragraph id="8BF71E1AFFAAFFE9FF26FDFA5715C2FF" blockId="11.[151,1437,150,1293]" pageId="11" pageNumber="448">
<emphasis id="B93CC208FFAAFFE9FF26FDFA522BC253" bold="true" box="[199,275,583,609]" pageId="11" pageNumber="448">Elytra</emphasis>
(
<figureCitation id="1373029FFFAAFFE9FEC3FDFA524BC253" box="[290,371,582,609]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">Fig. 5e</figureCitation>
): 1.4 times wider than long (length: 
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, width: 
<quantity id="4CB0B3FFFFAAFFE9FC23FDFB5711C253" box="[962,1065,582,609]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="11" pageNumber="448" unit="mm" value="0.59">0.59 mm</quantity>
), approximately 0.6 times shorter than head or pronotum [elytra length/head length: 0.57, elytra length/pronotum length: 0.60); only slightly narrower than head (0.9 times) and as wide as pronotum (
<quantity id="4CB0B3FFFFAAFFE9FD28FD33500DC29B" box="[713,821,654,681]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="11" pageNumber="448" unit="mm" value="0.59">0.59 mm</quantity>
); flat in dorsal view; humeral angles obsolete; lateral margins parallel; surface with coarse, rugose and wrinkled punctuation. Wingless.
</paragraph>
<paragraph id="8BF71E1AFFAAFFE9FF26FD6B5706C423" blockId="11.[151,1437,150,1293]" pageId="11" pageNumber="448">
<emphasis id="B93CC208FFAAFFE9FF26FD6B5204C2C2" bold="true" box="[199,316,726,752]" pageId="11" pageNumber="448">Abdomen</emphasis>
(
<figureCitation id="1373029FFFAAFFE9FEAAFD6A5283C2C3" box="[331,443,726,753]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">Figs 5f–h</figureCitation>
): Maximum width at segment VI (
<quantity id="4CB0B3FFFFAAFFE9FCDDFD6A509EC2C3" box="[828,934,727,753]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="11" pageNumber="448" unit="mm" value="0.69">0.69 mm</quantity>
), 1.2 times wider than elytra; tergites III‒VII with fine and dense punctuation within a polygonal microsculpture of transverse meshes, these more pronounced in the intersegmental membranes; sternite III with a middle carina (intercoxal carina) in the first half (
<figureCitation id="1373029FFFAAFFE9FAA5FCA256ACC30B" box="[1348,1428,798,825]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">Fig. 5f</figureCitation>
) surrounded in both sides by a slight depression to accommodate the resting metacoxae; sternites III‒VI with fine and dense punctuation within a microsculpture of transverse meshes, substituted by irregular rows of scale-shaped microtrichia anteriad near the intersegmental membranes and by regular posteriad quadrate meshes. Sternite VII without any median impression posteriorly; sternite VIII slightly transverse, distinctly and circularly excised at posterior margin surrounded by a small glabrous area, surface with a microsculpture of polygonal meshes (
<figureCitation id="1373029FFFAAFFE9FADAFC6E56B6C3DF" box="[1339,1422,978,1005]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">
Fig. 
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</figureCitation>
); sternite IX slightly arcuate apically; genital segment in ventral view as in figure 5h.
</paragraph>
<paragraph id="8BF71E1AFFAAFFE9FF26FBA65204C46B" blockId="11.[151,1437,150,1293]" pageId="11" pageNumber="448">
<emphasis id="B93CC208FFAAFFE9FF26FBA653C7C407" bold="true" box="[199,255,1051,1077]" pageId="11" pageNumber="448">Legs</emphasis>
: Long and slender; forelegs with antennal cleaning organ following the general pattern of the genus (e.g. see 
<figureCitation id="1373029FFFAAFFE9FF23FB825208C46B" box="[194,304,1087,1113]" captionStart="FIGURE 2" captionStartId="6.[152,255,1909,1933]" captionTargetBox="[195,1392,179,1883]" captionTargetId="figure-19@6.[195,1392,179,1883]" captionTargetPageId="6" captionText="FIGURE 2. SEM photographs of structural features of Domene boavidae sp. nov.: (a), head (dorsal view); (b), detail of the eye and chaetotaxy (lateral view); (c), labrum, clypeus and frons (part) (dorsal view); (d), antennae (dorsal view); (e), part of the cleaning organ in forefemur (latero‒ventral view); (f), part of the cleaning organ in foretibia (latero‒ventral view); (g), pronotum (dorsal view); (h), left elytron (dorsal view)." figureDoi="http://doi.org/10.5281/zenodo.14896600" httpUri="https://zenodo.org/record/14896600/files/figure.png" pageId="11" pageNumber="448">Figs 2e–f</figureCitation>
).
</paragraph>
<paragraph id="8BF71E1AFFAAFFE9FF26FBDF5038C53F" blockId="11.[151,1437,150,1293]" pageId="11" pageNumber="448">
<emphasis id="B93CC208FFAAFFE9FF26FBDF5202C44E" bold="true" box="[199,314,1122,1148]" pageId="11" pageNumber="448">Aedeagus</emphasis>
(
<figureCitation id="1373029FFFAAFFE9FEABFBDE52A8C44F" box="[330,400,1123,1149]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Fig. 6</figureCitation>
): Length 
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; sclerotized ventral process asymmetric, exceeding largely the edge of the dorsal plate (
<figureCitation id="1373029FFFAAFFE9FECBFB3A52A5C493" box="[298,413,1159,1185]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Figs 6a–c</figureCitation>
), with a twisted leaf shape (
<figureCitation id="1373029FFFAAFFE9FD3AFB3A5040C493" box="[731,888,1159,1185]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Figs 6a, 6c–d</figureCitation>
); the base of the ventral process horn-shaped on each side directed inwards and upwards (
<figureCitation id="1373029FFFAAFFE9FD80FB1651D8C4F7" box="[609,736,1195,1221]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Figs 6a, 6c</figureCitation>
); dorsal plate cylindrical‒shaped in dorsal view (
<figureCitation id="1373029FFFAAFFE9FAE2FB16566EC4F7" box="[1283,1366,1195,1221]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Fig. 6b</figureCitation>
), with fused lateral lobes (
<figureCitation id="1373029FFFAAFFE9FE8FFB725287C4DB" box="[366,447,1231,1257]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Fig. 6c</figureCitation>
), an inverted V‒shaped sclerotized area towards the ventral process, supporting the upper part and the apical edge of the dorsal plate (
<figureCitation id="1373029FFFAAFFE9FD61FB4E51CCC53F" box="[640,756,1267,1293]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Figs 6b–c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3524D91FFAAFFECFF76FAE35050C503" lastPageId="14" lastPageNumber="451" pageId="11" pageNumber="448" type="discussion">
<paragraph id="8BF71E1AFFAAFFE9FF76FAE351DEC54B" blockId="11.[151,742,1374,1401]" box="[151,742,1374,1401]" pageId="11" pageNumber="448">
<heading id="D0BFA976FFAAFFE9FF76FAE351DEC54B" bold="true" box="[151,742,1374,1401]" fontSize="11" level="1" pageId="11" pageNumber="448" reason="1">
<emphasis id="B93CC208FFAAFFE9FF76FAE351DEC54B" bold="true" box="[151,742,1374,1401]" pageId="11" pageNumber="448">Taxonomic, distribution and bionomic remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BF71E1AFFAAFFE9FF76FA1A509CC78B" blockId="11.[151,1437,1447,1977]" pageId="11" pageNumber="448">
According to 
<bibRefCitation id="EFD963EBFFAAFFE9FED6FA1A52DBC5F3" author="Coiffait, H." box="[311,483,1447,1473]" pageId="11" pageNumber="448" pagination="1 - 440" refId="ref11438" refString="Coiffait, H. (1982) Coleopteres Staphylinidae de la region palearctique occidentale. IV. Sous famille Paederinae. Tribu Paederini 1 (Paederi, Lathrobii). Supplement a la Nouvelle Revue d'Entomologie, 12 (4), 1-440." type="journal article" year="1982">Coiffait (1982)</bibRefCitation>
taxonomic criteria (e.g. aedeagus with fused lateral lobes and the development of the lateral and ventral process) 
<taxonomicName id="4C486599FFAAFFE9FDEBFA765183C5D7" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[522,699,1483,1509]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFAAFFE9FDEBFA765183C5D7" box="[522,699,1483,1509]" italics="true" pageId="11" pageNumber="448">Domene pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAAFFE9FD27FA76501BC5D7" bold="true" box="[710,803,1483,1509]" pageId="11" pageNumber="448">
<taxonomicNameLabel id="A20F7F73FFAAFFE9FD27FA76501BC5D7" box="[710,803,1483,1509]" pageId="11" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
should be included also in the subgenus 
<taxonomicName id="4C486599FFAAFFE9FAEEFA7656ADC5D7" authorityName="Koch" authorityYear="1938" box="[1295,1429,1483,1509]" pageId="11" pageNumber="448" rank="subGenus" subGenus="Lathromene">
<emphasis id="B93CC208FFAAFFE9FAEEFA7656ADC5D7" box="[1295,1429,1483,1509]" italics="true" pageId="11" pageNumber="448">Lathromene</emphasis>
</taxonomicName>
. Among other morphological characteristics the species shows a distinctive shape of the aedeagus. Taking into account the pieces of this structure, the new species can be easily segregated from all known 
<taxonomicName id="4C486599FFAAFFE9FB5CF9AE567BC61F" authorityName="Koch" authorityYear="1938" box="[1213,1347,1555,1581]" pageId="11" pageNumber="454" rank="subGenus" subGenus="Lathromene">
<emphasis id="B93CC208FFAAFFE9FB5CF9AE567BC61F" box="[1213,1347,1555,1581]" italics="true" pageId="11" pageNumber="448">Lathromene</emphasis>
</taxonomicName>
species with symmetric ventral process (
<taxonomicName id="4C486599FFAAFFE9FDEDF98A51B8C663" box="[524,640,1591,1617]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="bergidi">
<emphasis id="B93CC208FFAAFFE9FDEDF98A51B8C663" box="[524,640,1591,1617]" italics="true" pageId="11" pageNumber="448">D. bergidi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FD6EF98A5022C663" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[655,794,1591,1617]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="boavidae" status="sp. nov.">
<emphasis id="B93CC208FFAAFFE9FD6EF98A5022C663" box="[655,794,1591,1617]" italics="true" pageId="11" pageNumber="448">D. boavidae</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAAFFE9FCC3F98A5045C663" bold="true" box="[802,893,1591,1617]" pageId="11" pageNumber="448">
<taxonomicNameLabel id="A20F7F73FFAAFFE9FCC3F98A5045C663" box="[802,893,1591,1617]" pageId="11" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="4C486599FFAAFFE9FC6DF98A5711C663" authorityName="Coiffait" authorityYear="1973" box="[908,1065,1591,1617]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="cantabrica">
<emphasis id="B93CC208FFAAFFE9FC6DF98A5711C663" box="[908,1065,1591,1617]" italics="true" pageId="11" pageNumber="448">D. cantabrica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FBD8F98A57E3C663" authorityName="Outerelo, Gamarra &amp; Salgado" authorityYear="2000" box="[1081,1243,1591,1617]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="caurelensis">
<emphasis id="B93CC208FFAAFFE9FBD8F98A57E3C663" box="[1081,1243,1591,1617]" italics="true" pageId="11" pageNumber="448">D. caurelensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FB0BF98A564AC663" authorityName="Magrini &amp; Carotti" authorityYear="2019" box="[1258,1394,1591,1617]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="darinkae">
<emphasis id="B93CC208FFAAFFE9FB0BF98A564AC663" box="[1258,1394,1591,1617]" italics="true" pageId="11" pageNumber="448">D. darinkae</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FA60F98A53C8C647" authorityName="Struyve" authorityYear="2018" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="eufemia">
<emphasis id="B93CC208FFAAFFE9FA60F98A53C8C647" italics="true" pageId="11" pageNumber="448">D. eufemia</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FF1FF9E652A4C647" authorityName="Fagel" authorityYear="1967" box="[254,412,1627,1653]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="gridelliana">
<emphasis id="B93CC208FFAAFFE9FF1FF9E652A4C647" box="[254,412,1627,1653]" italics="true" pageId="11" pageNumber="448">D. gridelliana</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FE4AF9E651B0C647" authorityName="Struyve" authorityYear="2018" box="[427,648,1627,1653]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="invernadeirensis">
<emphasis id="B93CC208FFAAFFE9FE4AF9E651B0C647" box="[427,648,1627,1653]" italics="true" pageId="11" pageNumber="448">D. invernadeirensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FD74F9E6501EC647" authorityName="Reboleira &amp; Oromi" authorityYear="2011" box="[661,806,1627,1653]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="B93CC208FFAAFFE9FD74F9E6501EC647" box="[661,806,1627,1653]" italics="true" pageId="11" pageNumber="448">D. lusitanica</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FCD5F9E6509FC646" authorityName="Struyve" authorityYear="2018" box="[820,935,1627,1652]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="orensis">
<emphasis id="B93CC208FFAAFFE9FCD5F9E6509FC646" box="[820,935,1627,1652]" italics="true" pageId="11" pageNumber="448">D. orensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FC54F9E65771C647" authorityName="Struyve" authorityYear="2018" box="[949,1097,1627,1653]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="sistelensis">
<emphasis id="B93CC208FFAAFFE9FC54F9E65771C647" box="[949,1097,1627,1653]" italics="true" pageId="11" pageNumber="448">D. sistelensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAAFFE9FBB6F9E657D8C647" authorityName="Struyve" authorityYear="2018" box="[1111,1248,1627,1653]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="suidensis">
<emphasis id="B93CC208FFAAFFE9FBB6F9E657D8C647" box="[1111,1248,1627,1653]" italics="true" pageId="11" pageNumber="448">D. suidensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C486599FFAAFFE9FAF6F9E656B3C646" authorityName="Serrano &amp; Boieiro" authorityYear="2015" box="[1303,1419,1627,1652]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="viriatoi">
<emphasis id="B93CC208FFAAFFE9FAF6F9E656B3C646" box="[1303,1419,1627,1652]" italics="true" pageId="11" pageNumber="448">D. viriatoi</emphasis>
</taxonomicName>
). Among the species with asymmetric ventral process, the closest species to 
<taxonomicName id="4C486599FFAAFFE9FC34F9C25707C6AB" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[981,1087,1663,1689]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFAAFFE9FC34F9C25707C6AB" box="[981,1087,1663,1689]" italics="true" pageId="11" pageNumber="448">D. pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAAFFE9FBA4F9C257A6C6AB" bold="true" box="[1093,1182,1663,1689]" pageId="11" pageNumber="448">
<taxonomicNameLabel id="A20F7F73FFAAFFE9FBA4F9C257A6C6AB" box="[1093,1182,1663,1689]" pageId="11" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
seems to be 
<taxonomicName id="4C486599FFAAFFE9FACAF9C2526DC68F" authority="(Outerelo, 1977)" baseAuthorityName="Outerelo" baseAuthorityYear="1977" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="subiasi">
<emphasis id="B93CC208FFAAFFE9FACAF9C256A4C6AB" box="[1323,1436,1663,1689]" italics="true" pageId="11" pageNumber="448">D. subiasi</emphasis>
(
<bibRefCitation id="EFD963EBFFAAFFE9FF7EF91E5275C68F" author="Outerelo, R." box="[159,333,1699,1725]" pageId="11" pageNumber="448" pagination="23 - 31" refId="ref12055" refString="Outerelo, R. (1977) Seis nuevos estafilinoides del norte de la Provincia de Caceres y sur de la de Salamanca (Espana) (Col. Staphylinoidea). Nouvelle Revue d`Entomologie, 7 (1), 23-31." type="journal article" year="1977">Outerelo, 1977</bibRefCitation>
)
</taxonomicName>
. However, unlike the new species, the ventral process of 
<taxonomicName id="4C486599FFAAFFE9FC3BF91E5774C68F" baseAuthorityName="Outerelo" baseAuthorityYear="1977" box="[986,1100,1699,1725]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="subiasi">
<emphasis id="B93CC208FFAAFFE9FC3BF91E5774C68F" box="[986,1100,1699,1725]" italics="true" pageId="11" pageNumber="448">D. subiasi</emphasis>
</taxonomicName>
presents a dilated axe-shaped apical tip in lateral view (Figs 44, 
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<bibRefCitation id="EFD963EBFFAAFFE9FDB8F97A51C3C6D3" author="Outerelo, R." box="[601,763,1735,1761]" pageId="11" pageNumber="448" pagination="23 - 31" refId="ref12055" refString="Outerelo, R. (1977) Seis nuevos estafilinoides del norte de la Provincia de Caceres y sur de la de Salamanca (Espana) (Col. Staphylinoidea). Nouvelle Revue d`Entomologie, 7 (1), 23-31." type="journal article" year="1977">Outerelo 1977</bibRefCitation>
) (leaf-shaped in the new species) and a small tooth directed laterally and obliquely backwards in the middle region on the left side (absent in the new species) (cf. 
<figureCitation id="1373029FFFAAFFE9FAFFF9565654C737" box="[1310,1388,1771,1797]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Fig. 6c</figureCitation>
and fig. 
<quantity id="4CB0B3FFFFAAFFE9FF22F8B353C3C71B" box="[195,251,1806,1833]" metricMagnitude="0" metricUnit="m" metricValue="1.143" pageId="11" pageNumber="448" unit="in" value="45.0">45 in</quantity>
<bibRefCitation id="EFD963EBFFAAFFE9FEE0F8B2529CC71B" author="Outerelo, R." box="[257,420,1807,1833]" pageId="11" pageNumber="448" pagination="23 - 31" refId="ref12055" refString="Outerelo, R. (1977) Seis nuevos estafilinoides del norte de la Provincia de Caceres y sur de la de Salamanca (Espana) (Col. Staphylinoidea). Nouvelle Revue d`Entomologie, 7 (1), 23-31." type="journal article" year="1977">Outerelo 1977</bibRefCitation>
). Furthermore, the apical tip is directed outward and in the new species it is directed slightly inward (cf. 
<figureCitation id="1373029FFFAAFFE9FEFAF88E5252C77F" box="[283,362,1843,1869]" captionStart="FIGURE 6" captionStartId="13.[152,255,1962,1986]" captionTargetBox="[265,1324,210,1925]" captionTargetId="figure-17@13.[252,1336,198,1936]" captionTargetPageId="13" captionText="FIGURE 6. Aedeagus of Domene pedroi sp. nov.: (a), lateral view; (b), dorsal view; (c), ventral view; (d), apex of ventral blade (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896608" httpUri="https://zenodo.org/record/14896608/files/figure.png" pageId="11" pageNumber="448">Fig. 6a</figureCitation>
and fig. 
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<bibRefCitation id="EFD963EBFFAAFFE9FDF2F88E5181C77F" author="Outerelo, R." box="[531,697,1843,1869]" pageId="11" pageNumber="448" pagination="23 - 31" refId="ref12055" refString="Outerelo, R. (1977) Seis nuevos estafilinoides del norte de la Provincia de Caceres y sur de la de Salamanca (Espana) (Col. Staphylinoidea). Nouvelle Revue d`Entomologie, 7 (1), 23-31." type="journal article" year="1977">Outerelo 1977</bibRefCitation>
). Regarding morphological characteristics other than those of the aedeagus 
<taxonomicName id="4C486599FFAAFFE9FEE6F8EA5243C743" baseAuthorityName="Outerelo" baseAuthorityYear="1977" box="[263,379,1879,1905]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="448" phylum="Arthropoda" rank="species" species="subiasi">
<emphasis id="B93CC208FFAAFFE9FEE6F8EA5243C743" box="[263,379,1879,1905]" italics="true" pageId="11" pageNumber="448">D. subiasi</emphasis>
</taxonomicName>
has a narrow and glabrous median stripe on the upper part of the head and a slight median longitudinal depression without setae in the VII sternite (see figs 40, 
<quantity id="4CB0B3FFFFAAFFE9FC28F8C6573EC7A7" box="[969,1030,1915,1941]" metricMagnitude="0" metricUnit="m" metricValue="1.0668" pageId="11" pageNumber="448" unit="in" value="42.0">42 in</quantity>
<bibRefCitation id="EFD963EBFFAAFFE9FBF0F8C65781C7A7" author="Outerelo, R." box="[1041,1209,1915,1941]" pageId="11" pageNumber="448" pagination="23 - 31" refId="ref12055" refString="Outerelo, R. (1977) Seis nuevos estafilinoides del norte de la Provincia de Caceres y sur de la de Salamanca (Espana) (Col. Staphylinoidea). Nouvelle Revue d`Entomologie, 7 (1), 23-31." type="journal article" year="1977">Outerelo 1977</bibRefCitation>
, respectively), both characteristics are absent in the new species (e.g. see head in 
<figureCitation id="1373029FFFAAFFE9FCA7F82250AFC78B" box="[838,919,1950,1977]" captionStart="FIGURE 5" captionStartId="12.[152,255,1943,1967]" captionTargetBox="[202,1384,179,1910]" captionTargetId="figure-19@12.[202,1386,179,1917]" captionTargetPageId="12" captionText="FIGURE 5. Structural features of Domene pedroi sp. nov., holotype male: (a), (b) and (c), head (dorsal, ventral and lateral views, respectively); (d), pronotum (dorsal view); (e), elytra (dorsal view); (f), III abdominal sternite with intercoxal carina (arrow) (ventral view); (g), detail of VIII abdominal sternite (ventral view); (h), genital segment (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="11" pageNumber="448">Fig. 5a</figureCitation>
).
</paragraph>
<caption id="DF374E92FFADFFEEFF79F82A5639C7C5" ID-DOI="http://doi.org/10.5281/zenodo.14896606" ID-Zenodo-Dep="14896606" httpUri="https://zenodo.org/record/14896606/files/figure.png" pageId="12" pageNumber="449" startId="12.[152,255,1943,1967]" targetBox="[202,1384,179,1910]" targetPageId="12" targetType="figure">
<paragraph id="8BF71E1AFFADFFEEFF79F82A5639C7C5" blockId="12.[151,1436,1943,2040]" pageId="12" pageNumber="449">
<emphasis id="B93CC208FFADFFEEFF79F82A5222C79D" bold="true" box="[152,282,1943,1967]" pageId="12" pageNumber="449">FIGURE 5.</emphasis>
Structural features of 
<taxonomicName id="4C486599FFADFFEEFDE2F8255199C79D" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[515,673,1943,1967]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="449" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFADFFEEFDE2F8255199C79D" box="[515,673,1943,1967]" italics="true" pageId="12" pageNumber="449">Domene pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFADFFEEFD48F82A51C3C79D" bold="true" box="[681,763,1943,1967]" pageId="12" pageNumber="449">
<taxonomicNameLabel id="A20F7F73FFADFFEEFD48F82A51C3C79D" box="[681,763,1943,1967]" pageId="12" pageNumber="449" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male: 
<emphasis id="B93CC208FFADFFEEFC4AF82A50F1C79D" bold="true" box="[939,969,1943,1967]" pageId="12" pageNumber="449">(a)</emphasis>
, 
<emphasis id="B93CC208FFADFFEEFC36F82A50CEC79D" bold="true" box="[983,1014,1943,1967]" pageId="12" pageNumber="449">(b)</emphasis>
and 
<emphasis id="B93CC208FFADFFEEFBCAF82A577FC79D" bold="true" box="[1067,1095,1943,1967]" pageId="12" pageNumber="449">(c)</emphasis>
, head (dorsal, ventral and lateral views, respectively); 
<emphasis id="B93CC208FFADFFEEFE95F80652ABC7E1" bold="true" box="[372,403,1979,2003]" pageId="12" pageNumber="449">(d)</emphasis>
, pronotum (dorsal view); 
<emphasis id="B93CC208FFADFFEEFD44F80651F9C7E1" bold="true" box="[677,705,1979,2003]" pageId="12" pageNumber="449">(e)</emphasis>
, elytra (dorsal view); 
<emphasis id="B93CC208FFADFFEEFC4BF80650FBC7E1" bold="true" box="[938,963,1979,2003]" pageId="12" pageNumber="449">(f)</emphasis>
, III abdominal sternite with intercoxal carina (arrow) (ventral view); 
<emphasis id="B93CC208FFADFFEEFE67F862529BC7C5" bold="true" box="[390,419,2015,2039]" pageId="12" pageNumber="449">(g)</emphasis>
, detail of VIII abdominal sternite (ventral view); 
<emphasis id="B93CC208FFADFFEEFC7BF8625081C7C5" bold="true" box="[922,953,2015,2039]" pageId="12" pageNumber="449">(h)</emphasis>
, genital segment (ventral view).
</paragraph>
</caption>
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<paragraph id="8BF71E1AFFACFFEFFF79F8175252C7D4" blockId="13.[151,1436,1962,2022]" pageId="13" pageNumber="450">
<emphasis id="B93CC208FFACFFEFFF79F8175222C7F0" bold="true" box="[152,282,1962,1986]" pageId="13" pageNumber="450">FIGURE 6.</emphasis>
Aedeagus of 
<taxonomicName id="4C486599FFACFFEFFE4BF8165170C7F0" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[426,584,1962,1986]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="450" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFACFFEFFE4BF8165170C7F0" box="[426,584,1962,1986]" italics="true" pageId="13" pageNumber="450">Domene pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFACFFEFFDB1F817519AC7F0" bold="true" box="[592,674,1962,1986]" pageId="13" pageNumber="450">
<taxonomicNameLabel id="A20F7F73FFACFFEFFDB1F817519AC7F0" box="[592,674,1962,1986]" pageId="13" pageNumber="450" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: 
<emphasis id="B93CC208FFACFFEFFD53F81751E8C7F0" bold="true" box="[690,720,1962,1986]" pageId="13" pageNumber="450">(a)</emphasis>
, lateral view; 
<emphasis id="B93CC208FFACFFEFFC87F81750BDC7F0" bold="true" box="[870,901,1962,1986]" pageId="13" pageNumber="450">(b)</emphasis>
, dorsal view; 
<emphasis id="B93CC208FFACFFEFFBFBF817570EC7F0" bold="true" box="[1050,1078,1962,1986]" pageId="13" pageNumber="450">(c)</emphasis>
, ventral view; 
<emphasis id="B93CC208FFACFFEFFB32F81757CAC7F0" bold="true" box="[1235,1266,1962,1986]" pageId="13" pageNumber="450">(d)</emphasis>
, apex of ventral blade (ventral view).
</paragraph>
</caption>
<paragraph id="8BF71E1AFFAFFFECFF26FF2A51E0C327" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="451">
The remaining three species with asymmetric ventral blade are 
<taxonomicName id="4C486599FFAFFFECFC35FF2A574FC083" authorityName="Fauvel" authorityYear="1873" box="[980,1143,151,177]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="scopaeella">
<emphasis id="B93CC208FFAFFFECFC35FF2A574FC083" box="[980,1143,151,177]" italics="true" pageId="14" pageNumber="451">D. scopaeella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAFFFECFB6DFF2A5602C083" authorityName="Feldmann &amp; Hernando" authorityYear="2005" box="[1164,1338,151,177]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="gallaeciana">
<emphasis id="B93CC208FFAFFFECFB6DFF2A5602C083" box="[1164,1338,151,177]" italics="true" pageId="14" pageNumber="451">D. gallaeciana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C486599FFAFFFECFA60FF2A5201C0E7" authorityName="Outerelo &amp; Gamarra" authorityYear="2012" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="barraganensis">
<emphasis id="B93CC208FFAFFFECFA60FF2A5201C0E7" italics="true" pageId="14" pageNumber="451">D. barraganensis</emphasis>
</taxonomicName>
. All can be segregated from the new species by some features of the aedeagus. In the 
<taxonomicName id="4C486599FFAFFFECFAE3FF0656A4C0E7" authorityName="Fauvel" authorityYear="1873" box="[1282,1436,187,213]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="scopaeella">
<emphasis id="B93CC208FFAFFFECFAE3FF06562EC0E6" box="[1282,1302,187,212]" italics="true" pageId="14" pageNumber="451">D</emphasis>
. 
<emphasis id="B93CC208FFAFFFECFAC5FF0656A4C0E7" box="[1316,1436,187,213]" italics="true" pageId="14" pageNumber="451">scopaeella</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C486599FFAFFFECFF24FF6252B0C0CB" authorityName="Outerelo &amp; Gamarra" authorityYear="2012" box="[197,392,223,249]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="barraganensis">
<emphasis id="B93CC208FFAFFFECFF24FF6252B0C0CB" box="[197,392,223,249]" italics="true" pageId="14" pageNumber="451">D. barraganensis</emphasis>
</taxonomicName>
the ventral process has an apical dilated axe-shaped tip that bends inwards at a right angle [see figs 100G–H in 
<bibRefCitation id="EFD963EBFFAFFFECFEAFFEBE52CEC12F" author="Coiffait, H." box="[334,502,259,285]" pageId="14" pageNumber="451" pagination="1 - 440" refId="ref11438" refString="Coiffait, H. (1982) Coleopteres Staphylinidae de la region palearctique occidentale. IV. Sous famille Paederinae. Tribu Paederini 1 (Paederi, Lathrobii). Supplement a la Nouvelle Revue d'Entomologie, 12 (4), 1-440." type="journal article" year="1982">Coiffait (1982)</bibRefCitation>
and figs 10, 
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<bibRefCitation id="EFD963EBFFAFFFECFD28FEBE573EC12F" author="Outerelo, R. &amp; Gamarra, P." box="[713,1030,259,285]" pageId="14" pageNumber="451" pagination="123 - 128" refId="ref12141" refString="Outerelo, R. &amp; Gamarra, P. (2012) Domene (Lathromene) barraganensis, nueva especie de estafilinido de la Peninsula Iberica (Coleoptera: Staphylinidae, Paederinae). Arquivos Entomoloxicos, 7, 123-128." type="journal article" year="2012">Outerelo &amp; Gamarra (2012)</bibRefCitation>
, respectively]. Moreover, unlike the new species, the upper part and the apical edge of the dorsal plate in 
<taxonomicName id="4C486599FFAFFFECFC44FE9A5752C173" authorityName="Outerelo &amp; Gamarra" authorityYear="2012" box="[933,1130,295,321]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="barraganensis">
<emphasis id="B93CC208FFAFFFECFC44FE9A5752C173" box="[933,1130,295,321]" italics="true" pageId="14" pageNumber="451">D. barraganensis</emphasis>
</taxonomicName>
is not directed towards the ventral process (see fig. 
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<bibRefCitation id="EFD963EBFFAFFFECFE0BFEF6502FC157" author="Outerelo, R. &amp; Gamarra, P." box="[490,791,331,357]" pageId="14" pageNumber="451" pagination="123 - 128" refId="ref12141" refString="Outerelo, R. &amp; Gamarra, P. (2012) Domene (Lathromene) barraganensis, nueva especie de estafilinido de la Peninsula Iberica (Coleoptera: Staphylinidae, Paederinae). Arquivos Entomoloxicos, 7, 123-128." type="journal article" year="2012">Outerelo &amp; Gamarra 2012</bibRefCitation>
). Concerning 
<taxonomicName id="4C486599FFAFFFECFC55FEF65763C157" authorityName="Feldmann &amp; Hernando" authorityYear="2005" box="[948,1115,331,357]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="gallaeciana">
<emphasis id="B93CC208FFAFFFECFC55FEF65763C157" box="[948,1115,331,357]" italics="true" pageId="14" pageNumber="451">D. gallaeciana</emphasis>
</taxonomicName>
, the ventral process exceeds largely the edge of the aedeagus and from the middle to apex, gradually narrows, becoming filiform towards the tip (see figs 
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<bibRefCitation id="EFD963EBFFAFFFECFE80FE2E519DC19F" author="Feldmann, B. &amp; Hernando, C." box="[353,677,402,429]" pageId="14" pageNumber="451" pagination="399 - 406" refId="ref11702" refString="Feldmann, B. &amp; Hernando, C. (2005) Two new species of Domene Fauvel 1873 from Spain, with a new combination and a catalogue of the Iberian species of the genus (Coleoptera: Staphylinidae, Paederinae). Linzer Biologische Beitrage, 37 (1), 399-406." type="journal article" year="2005">Feldmann &amp; Hernando 2005</bibRefCitation>
); median region presents also two contiguous lateral expansions that are absent in the new species (ventral view) (fig. 
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<bibRefCitation id="EFD963EBFFAFFFECFCEDFE0A576EC1E3" author="Feldmann, B. &amp; Hernando, C." box="[780,1110,438,465]" pageId="14" pageNumber="451" pagination="399 - 406" refId="ref11702" refString="Feldmann, B. &amp; Hernando, C. (2005) Two new species of Domene Fauvel 1873 from Spain, with a new combination and a catalogue of the Iberian species of the genus (Coleoptera: Staphylinidae, Paederinae). Linzer Biologische Beitrage, 37 (1), 399-406." type="journal article" year="2005">Feldmann &amp; Hernando 2005</bibRefCitation>
). Furthermore, in these three species there are no horn-shaped process on each side of the base of the ventral process (present in the new species and in 
<taxonomicName id="4C486599FFAFFFECFF05FE42526EC22B" baseAuthorityName="Outerelo" baseAuthorityYear="1977" box="[228,342,511,537]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="subiasi">
<emphasis id="B93CC208FFAFFFECFF05FE42526EC22B" box="[228,342,511,537]" italics="true" pageId="14" pageNumber="451">D. subiasi</emphasis>
</taxonomicName>
). Some other morphological characteristics, like the ommatidia number, as well as male sternites VII and VIII features, allow to segregate the new species from 
<taxonomicName id="4C486599FFAFFFECFCB4FD9E50D7C20F" authorityName="Fauvel" authorityYear="1873" box="[853,1007,547,573]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="scopaeella">
<emphasis id="B93CC208FFAFFFECFCB4FD9E50D7C20F" box="[853,1007,547,573]" italics="true" pageId="14" pageNumber="451">D. scopaeella</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C486599FFAFFFECFC1CFD9E579AC20F" authorityName="Feldmann &amp; Hernando" authorityYear="2005" box="[1021,1186,547,573]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="gallaeciana">
<emphasis id="B93CC208FFAFFFECFC1CFD9E579AC20F" box="[1021,1186,547,573]" italics="true" pageId="14" pageNumber="451">D. gallaeciana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C486599FFAFFFECFB38FD9E56A4C20F" authorityName="Outerelo &amp; Gamarra" authorityYear="2012" box="[1241,1436,547,573]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="barraganensis">
<emphasis id="B93CC208FFAFFFECFB38FD9E56A4C20F" box="[1241,1436,547,573]" italics="true" pageId="14" pageNumber="451">D. barraganensis</emphasis>
</taxonomicName>
too. For instance, 
<taxonomicName id="4C486599FFAFFFECFEBEFDFA513DC253" authorityName="Feldmann &amp; Hernando" authorityYear="2005" box="[351,517,583,609]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="gallaeciana">
<emphasis id="B93CC208FFAFFFECFEBEFDFA513DC253" box="[351,517,583,609]" italics="true" pageId="14" pageNumber="451">D. gallaeciana</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C486599FFAFFFECFDDAFDFA51C7C253" authorityName="Outerelo &amp; Gamarra" authorityYear="2012" box="[571,767,583,609]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="barraganensis">
<emphasis id="B93CC208FFAFFFECFDDAFDFA51C7C253" box="[571,767,583,609]" italics="true" pageId="14" pageNumber="451">D. barraganensis</emphasis>
</taxonomicName>
are totally anophtalmous (7 ommatidae in the new species), the latter presents a middle depression on VII sternite and both have black modified setae in the sides of the posteriad excision of sternite VIII (not present in the new species). The congeners with populations closer to the new species are 
<taxonomicName id="4C486599FFAFFFECFF20FD0E526AC2FF" authorityName="Reboleira &amp; Oromi" authorityYear="2011" box="[193,338,691,717]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="lusitanica">
<emphasis id="B93CC208FFAFFFECFF20FD0E526AC2FF" box="[193,338,691,717]" italics="true" pageId="14" pageNumber="451">D. lusitanica</emphasis>
</taxonomicName>
(from Gruta da Cerâmica, Serra do Sicó, 
<collectingCountry id="F35F5E8AFFAFFFECFCCFFD0E50A9C2FF" box="[814,913,691,717]" name="Portugal" pageId="14" pageNumber="451">Portugal</collectingCountry>
), 
<taxonomicName id="4C486599FFAFFFECFC44FD0E5722C2FE" authorityName="Serrano &amp; Boieiro" authorityYear="2015" box="[933,1050,691,716]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="viriatoi">
<emphasis id="B93CC208FFAFFFECFC44FD0E5722C2FE" box="[933,1050,691,716]" italics="true" pageId="14" pageNumber="451">D. viriatoi</emphasis>
</taxonomicName>
(from Buraco da Moura, Serra da Estrela, 
<collectingCountry id="F35F5E8AFFAFFFECFF14FD6A5260C2C3" box="[245,344,727,753]" name="Portugal" pageId="14" pageNumber="451">Portugal</collectingCountry>
), 
<taxonomicName id="4C486599FFAFFFECFE8FFD6A52C7C2C3" authorityName="Outerelo" authorityYear="1985" box="[366,511,727,753]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="B93CC208FFAFFFECFE8FFD6A52C7C2C3" box="[366,511,727,753]" italics="true" pageId="14" pageNumber="451">D. hispanica</emphasis>
</taxonomicName>
(from El Cabaco, Salamanca, 
<collectingCountry id="F35F5E8AFFAFFFECFC83FD6A509BC2C3" box="[866,931,727,753]" name="Spain" pageId="14" pageNumber="451">Spain</collectingCountry>
) and 
<taxonomicName id="4C486599FFAFFFECFC07FD6A5762C2C3" baseAuthorityName="Outerelo" baseAuthorityYear="1977" box="[998,1114,727,753]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="subiasi">
<emphasis id="B93CC208FFAFFFECFC07FD6A5762C2C3" box="[998,1114,727,753]" italics="true" pageId="14" pageNumber="451">D. subiasi</emphasis>
</taxonomicName>
(from Nuñomoral, Cáceres, 
<collectingCountry id="F35F5E8AFFAFFFECFF76FD4653E0C327" box="[151,216,763,789]" name="Spain" pageId="14" pageNumber="451">Spain</collectingCountry>
) (see also fig. 
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<bibRefCitation id="EFD963EBFFAFFFECFE5DFD4651F4C327" author="Magrini, P. &amp; Carotti, G." box="[444,716,763,789]" pageId="14" pageNumber="451" pagination="495 - 502" refId="ref11935" refString="Magrini, P. &amp; Carotti, G. (2019) Una nuova specie appartenente al genere Domene Fauvel, 1873 (subg. Lathromene Koch, 1938) del Portogallo (Coleoptera, Staphylinidae, Paederinae). Giornale Italiano di Entomologia, 15 (64), 495-502. [http://giornaleitalianodientomologia.blogspot.com]" type="journal article" year="2019">Magrini &amp; Carotti 2019</bibRefCitation>
).
</paragraph>
<paragraph id="8BF71E1AFFAFFFECFF26FCA25766C3B3" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="451">
<taxonomicName id="4C486599FFAFFFECFF26FCA25249C30B" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[199,369,799,825]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFAFFFECFF26FCA25249C30B" box="[199,369,799,825]" italics="true" pageId="14" pageNumber="451">Domene pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAFFFECFE97FCA252F5C30B" bold="true" box="[374,461,799,825]" pageId="14" pageNumber="451">
<taxonomicNameLabel id="A20F7F73FFAFFFECFE97FCA252F5C30B" box="[374,461,799,825]" pageId="14" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can easily be separated from 
<taxonomicName id="4C486599FFAFFFECFCECFCA2577CC30B" authority="Outerelo, 1985" authorityName="Outerelo" authorityYear="1985" box="[781,1092,798,825]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="hispanica">
<emphasis id="B93CC208FFAFFFECFCECFCA250A2C30B" box="[781,922,799,825]" italics="true" pageId="14" pageNumber="451">D. hispanica</emphasis>
<bibRefCitation id="EFD963EBFFAFFFECFC7FFCA2577CC30B" author="Outerelo, R." box="[926,1092,798,825]" pageId="14" pageNumber="451" pagination="103 - 106" refId="ref12100" refString="Outerelo, R. (1985) Domene (Lathromene) hispanicum n. sp. de Espana, y una homonimia-sinonimia del subgenero Spelaeomene Espanol. Anales de Biologia, Seccion Biologia Animal, 3, 103-106." type="journal article" year="1985">Outerelo, 1985</bibRefCitation>
</taxonomicName>
also through several characters of the external morphology (e.g. body size, shape and punctuation of the head and pronotum, absence of a median sulcus on the pronotum, femoral tooth of front leg) (see description in 
<bibRefCitation id="EFD963EBFFAFFFECFC4CFCDA576AC3B3" author="Outerelo, R." box="[941,1106,870,897]" pageId="14" pageNumber="451" pagination="103 - 106" refId="ref12100" refString="Outerelo, R. (1985) Domene (Lathromene) hispanicum n. sp. de Espana, y una homonimia-sinonimia del subgenero Spelaeomene Espanol. Anales de Biologia, Seccion Biologia Animal, 3, 103-106." type="journal article" year="1985">Outerelo 1985</bibRefCitation>
).
</paragraph>
<paragraph id="8BF71E1AFFAFFFECFF26FC36567BC423" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="451">
The 
<typeStatus id="54F3A0B8FFAFFFECFF16FC365261C397" box="[247,345,907,933]" pageId="14" pageNumber="451" type="holotype">holotype</typeStatus>
specimen of 
<taxonomicName id="4C486599FFAFFFECFE08FC365169C397" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[489,593,907,933]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFAFFFECFE08FC365169C397" box="[489,593,907,933]" italics="true" pageId="14" pageNumber="451">D. pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAFFFECFDB4FC365195C397" bold="true" box="[597,685,907,933]" pageId="14" pageNumber="451">
<taxonomicNameLabel id="A20F7F73FFAFFFECFDB4FC365195C397" box="[597,685,907,933]" pageId="14" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was sampled by hand directly among plates of schistose rocks below thin layer of clayey brown-reddish soil on the side slopes of a country road. The locality habitat was dominated by 
<taxonomicName id="4C486599FFAFFFECFF76FC6E53EFC3DE" authorityName="Perry" authorityYear="1811" box="[151,215,979,1004]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="genus">
<emphasis id="B93CC208FFAFFFECFF76FC6E53EFC3DE" box="[151,215,979,1004]" italics="true" pageId="14" pageNumber="451">Pinus</emphasis>
</taxonomicName>
trees with rock-roses, brooms, lavender and rosemary shrubs [
<taxonomicName id="4C486599FFAFFFECFC77FC6E5772C3DF" box="[918,1098,979,1005]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="species" species="ladanifer">
<emphasis id="B93CC208FFAFFFECFC77FC6E5772C3DF" box="[918,1098,979,1005]" italics="true" pageId="14" pageNumber="451">Cistus ladanifer</emphasis>
</taxonomicName>
(
<taxonomicName id="4C486599FFAFFFECFBB9FC6E57F1C3DF" box="[1112,1225,979,1005]" class="Magnoliopsida" family="Cistaceae" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="family">Cistaceae</taxonomicName>
), 
<taxonomicName id="4C486599FFAFFFECFB3DFC6E56A4C3DF" box="[1244,1436,979,1005]" class="Magnoliopsida" family="Cistaceae" genus="Cistus" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="species" species="salvifolius">
<emphasis id="B93CC208FFAFFFECFB3DFC6E56A4C3DF" box="[1244,1436,979,1005]" italics="true" pageId="14" pageNumber="451">Cistus salvifolius</emphasis>
</taxonomicName>
(
<taxonomicName id="4C486599FFAFFFECFF7EFC4A5228C423" box="[159,272,1015,1041]" class="Magnoliopsida" family="Cistaceae" kingdom="Plantae" order="Malvales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="family">Cistaceae</taxonomicName>
), 
<taxonomicName id="4C486599FFAFFFECFEC2FC4A5293C423" box="[291,427,1015,1041]" class="Magnoliopsida" family="Fabaceae" genus="Cytisus" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93CC208FFAFFFECFEC2FC4A524DC423" box="[291,373,1015,1041]" italics="true" pageId="14" pageNumber="451">Cytisus</emphasis>
spp.
</taxonomicName>
(
<taxonomicName id="4C486599FFAFFFECFE58FC4A511FC423" box="[441,551,1015,1041]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
), 
<taxonomicName id="4C486599FFAFFFECFDDBFC4A51E1C423" box="[570,729,1015,1041]" class="Magnoliopsida" family="Lamiaceae" genus="Lavandula" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B93CC208FFAFFFECFDDBFC4A518AC423" box="[570,690,1015,1041]" italics="true" pageId="14" pageNumber="451">Lavandula</emphasis>
sp.
</taxonomicName>
(
<taxonomicName id="4C486599FFAFFFECFD06FC4A505CC423" box="[743,868,1015,1041]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
) and 
<taxonomicName id="4C486599FFAFFFECFC43FC4A57A7C423" box="[930,1183,1015,1041]" class="Magnoliopsida" family="Lamiaceae" genus="Rosmarinus" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="species" species="officinalis">
<emphasis id="B93CC208FFAFFFECFC43FC4A57A7C423" box="[930,1183,1015,1041]" italics="true" pageId="14" pageNumber="451">Rosmarinus officinalis</emphasis>
</taxonomicName>
(
<taxonomicName id="4C486599FFAFFFECFB4FFC4A5609C423" box="[1198,1329,1015,1041]" class="Magnoliopsida" family="Lamiaceae" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="family">Lamiaceae</taxonomicName>
)].
</paragraph>
<paragraph id="8BF71E1AFFAFFFECFF26FBA652CBC493" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="451">
With no more data on the distribution of the new species than that of the 
<typeStatus id="54F3A0B8FFAFFFECFBF5FBA6574EC407" box="[1044,1142,1051,1077]" pageId="14" pageNumber="451" type="holotype">holotype</typeStatus>
it is provisional accepted that 
<taxonomicName id="4C486599FFAFFFECFF29FB82520BC46B" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[200,307,1087,1113]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFAFFFECFF29FB82520BC46B" box="[200,307,1087,1113]" italics="true" pageId="14" pageNumber="451">D. pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAFFFECFEDBFB8252ACC46B" bold="true" box="[314,404,1087,1113]" pageId="14" pageNumber="451">
<taxonomicNameLabel id="A20F7F73FFAFFFECFEDBFB8252ACC46B" box="[314,404,1087,1113]" pageId="14" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is endemic of 
<collectingCountry id="F35F5E8AFFAFFFECFDD8FB8251A4C46B" box="[569,668,1087,1113]" name="Portugal" pageId="14" pageNumber="451">Portugal</collectingCountry>
. In the same region and date two endogean ground beetles were also found: 
<taxonomicName id="4C486599FFAFFFECFF09FBDE51C5C44F" authority="Serrano &amp; Aguiar, 2014" authorityName="Serrano &amp; Aguiar" authorityYear="2014" box="[232,765,1123,1149]" class="Insecta" family="Carabidae" genus="Typhlocharis" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="zaballozi">
<emphasis id="B93CC208FFAFFFECFF09FBDE52DFC44F" box="[232,487,1123,1149]" italics="true" pageId="14" pageNumber="451">Typhlocharis zaballozi</emphasis>
<bibRefCitation id="EFD963EBFFAFFFECFE0FFBDE51C5C44F" author="Serrano, A. R. M. &amp; Aguiar, C. A. S." box="[494,765,1123,1149]" pageId="14" pageNumber="451" pagination="202 - 212" refId="ref12675" refString="Serrano, A. R. M. &amp; Aguiar, C. A. S. (2014) The genus Typhlocharis Dieck 1869 (Coleoptera: Carabidae): description of a new species from Portugal and remarks on the external microsculpture of the last abdominal sternum &quot; belt &quot;. Annales de la Societe entomologique de France, New Series, 50 (2), 202-212. https://doi.org/10.1080/00379271.2014.937106" type="journal article" year="2014">Serrano &amp; Aguiar, 2014</bibRefCitation>
</taxonomicName>
and 
<taxonomicName id="4C486599FFAFFFECFCD4FBDE565FC44F" authority="Serrano &amp; Aguiar, 2004" authorityName="Serrano &amp; Aguiar" authorityYear="2004" box="[821,1383,1123,1149]" class="Magnoliopsida" family="Lamiaceae" genus="Hypothyphlus" kingdom="Plantae" order="Lamiales" pageId="14" pageNumber="451" phylum="Tracheophyta" rank="species" species="lusitanicus">
<emphasis id="B93CC208FFAFFFECFCD4FBDE5769C44F" box="[821,1105,1123,1149]" italics="true" pageId="14" pageNumber="451">Hypothyphlus lusitanicus</emphasis>
Serrano &amp; Aguiar, 2004
</taxonomicName>
(see 
<bibRefCitation id="EFD963EBFFAFFFECFF76FB3A52A7C493" author="Serrano, A. R. M. &amp; Aguiar, C. A. S." box="[151,415,1159,1185]" pageId="14" pageNumber="451" pagination="33 - 46" refId="ref12608" refString="Serrano, A. R. M. &amp; Aguiar, C. A. S. (2011) Two new species of the genus Geocharis Ehlers, 1883 and new data on Anillina species from Portugal (Coleoptera: Carabidae). Zootaxa, 3116 (1), 33-46. https://doi.org/10.11646/zootaxa.3116.1.2" type="journal article" year="2011">Serrano &amp; Aguiar 2011</bibRefCitation>
, 
<bibRefCitation id="EFD963EBFFAFFFECFE4BFB3A52DFC493" author="Serrano, A. R. M. &amp; Aguiar, C. A. S." box="[426,487,1159,1185]" pageId="14" pageNumber="451" pagination="202 - 212" refId="ref12675" refString="Serrano, A. R. M. &amp; Aguiar, C. A. S. (2014) The genus Typhlocharis Dieck 1869 (Coleoptera: Carabidae): description of a new species from Portugal and remarks on the external microsculpture of the last abdominal sternum &quot; belt &quot;. Annales de la Societe entomologique de France, New Series, 50 (2), 202-212. https://doi.org/10.1080/00379271.2014.937106" type="journal article" year="2014">2014</bibRefCitation>
).
</paragraph>
<paragraph id="8BF71E1AFFAFFFECFF26FB165050C503" blockId="14.[151,1437,151,1329]" pageId="14" pageNumber="451">
According to the assumption of 
<bibRefCitation id="EFD963EBFFAFFFECFDA1FB16506FC4F7" author="Reboleira A. S. P. S. &amp; Goncalves, F. &amp; Oromi, P." box="[576,855,1195,1221]" pageId="14" pageNumber="451" pagination="48 - 56" refId="ref12367" refString="Reboleira A. S. P. S., Goncalves, F. &amp; Oromi, P. (2011 b) On the Iberian endemic subgenus Lathromene Koch (Coleoptera: Staphylinidae: Paederinae): description of the first hypogean Domene Fauvel, 1872 from Portugal. Zootaxa, 2780 (1), 48-56. https://doi.org/10.11646/zootaxa.2780.1.5" type="journal article" year="2011">
Reboleira 
<emphasis id="B93CC208FFAFFFECFD59FB1651D4C4F7" box="[696,748,1195,1221]" italics="true" pageId="14" pageNumber="451">et al</emphasis>
. (2011b)
</bibRefCitation>
, this small size species would be more connected with the endogean environment. Indeed, 
<taxonomicName id="4C486599FFAFFFECFD89FB7251ECC4DB" authorityName="Serrano &amp; Aguiar" authorityYear="2025" box="[616,724,1231,1257]" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="pedroi" status="sp. nov.">
<emphasis id="B93CC208FFAFFFECFD89FB7251ECC4DB" box="[616,724,1231,1257]" italics="true" pageId="14" pageNumber="451">D. pedroi</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAFFFECFD3DFB72500FC4DB" bold="true" box="[732,823,1231,1257]" pageId="14" pageNumber="451">
<taxonomicNameLabel id="A20F7F73FFAFFFECFD3DFB72500FC4DB" box="[732,823,1231,1257]" pageId="14" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was found in a typical endogean environment like 
<taxonomicName id="4C486599FFAFFFECFA60FB7253C6C53F" authorityName="Serrano &amp; Aguiar" authorityYear="2025" class="Insecta" family="Staphylinidae" genus="Domene" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="451" phylum="Arthropoda" rank="species" species="boavidae" status="sp. nov.">
<emphasis id="B93CC208FFAFFFECFA60FB7253C6C53F" italics="true" pageId="14" pageNumber="451">D. boavidae</emphasis>
</taxonomicName>
<emphasis id="B93CC208FFAFFFECFEE5FB4E5264C53F" bold="true" box="[260,348,1267,1293]" pageId="14" pageNumber="451">
<taxonomicNameLabel id="A20F7F73FFAFFFECFEE5FB4E5264C53F" box="[260,348,1267,1293]" pageId="14" pageNumber="451" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(e.g. 
<figureCitation id="1373029FFFAFFFECFE7BFB4E52D0C53F" box="[410,488,1267,1293]" captionStart="FIGURE 1" captionStartId="4.[152,255,1905,1929]" captionTargetBox="[151,1436,306,1862]" captionTargetId="figure-73@4.[151,1436,306,1879]" captionTargetPageId="4" captionText="FIGURE 1. (a), habitat of Domene boavidae sp. nov. near S. Pedro (Vila Viçosa, Viseu district); (b), D. boavidae sp. nov. male adult; (c), habitus of D. boavidae sp. nov. (dorsal view), holotype male; (d), habitus of Domene pedroi sp. nov. (dorsal view), holotype male." figureDoi="http://doi.org/10.5281/zenodo.14896598" httpUri="https://zenodo.org/record/14896598/files/figure.png" pageId="14" pageNumber="451">Fig. 1a</figureCitation>
). Taking into account the reduced eyes, lack of hind wings, elongated body shape and long legs it is assumed that it is a subterranean dwelling species.
</paragraph>
</subSubSection>
</treatment>
</document>