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<mods:title id="9D82553364EE7FDF42E8C96BDA05344D">The genus Drymeia Meigen, 1826 (Diptera: Muscidae) in Kyrgyzstan: new records and new species</mods:title>
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<mods:namePart id="67A306E8062576FB1B829607CD339E74">Sorokina, Vera S.</mods:namePart>
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<mods:affiliation id="6626F346E96C1330FE90BFDCA9BC6BD8">Institute of Systematics and Ecology of Animals, Russian Academy of Sciences, Siberian Branch, Frunze Street 11, Novosibirsk 630091, Russia. &amp; Tomsk State University, Lenin Avenue 36, Tomsk 634050, Russia.</mods:affiliation>
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<mods:namePart id="A78C37214FEB9581EC0ABD9A9767788C">Pont, Adrian C.</mods:namePart>
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<heading id="D08281A8C330FFE0FD1F9594FC14C7BD" box="[644,944,1486,1513]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="164" reason="2">
<taxonomicName id="4C754D47C330FFE0FD1F9594FCEAC7BD" ID-CoL="9J2XF" authority="Sorokina &amp; Pont, 2022" authorityName="Sorokina &amp; Pont" authorityYear="2022" box="[644,846,1486,1513]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="164" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
<emphasis id="B901EAD6C330FFE0FD1F9594FCEAC7BD" bold="true" box="[644,846,1486,1513]" italics="true" pageId="6" pageNumber="164">Drymeia ziegleri</emphasis>
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<taxonomicNameLabel id="A23257ADC330FFE0FCCE9595FC14C7BD" box="[853,944,1487,1513]" pageId="6" pageNumber="164" rank="species">sp. nov.</taxonomicNameLabel>
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<figureCitation id="134E2A41C330FFE0FD7C964EFCE8C47B" box="[743,844,1556,1583]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="2.[189,232,1962,1988]" captionStartId-1="3.[189,232,1998,2024]" captionStartId-2="7.[189,232,1963,1989]" captionStartId-3="8.[190,233,1962,1988]" captionTargetBox-0="[189,1398,869,1924]" captionTargetBox-1="[189,1398,1354,1959]" captionTargetBox-2="[189,1398,597,1924]" captionTargetBox-3="[189,1398,1116,1922]" captionTargetId-0="figure-128@2.[189,1398,869,1927]" captionTargetId-1="figure-147@3.[189,1398,1354,1959]" captionTargetId-2="figure-14@7.[189,1398,594,1924]" captionTargetId-3="figure-225@8.[189,1398,1116,1922]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="7" captionTargetPageId-3="8" captionText-0="Fig. 2. Angelica brevicaulis (Rupr.) covered with muscoid flies at Kalmak-Anshun Pass, Kyrgyzstan. Photo: J. Ziegler (Berlin, Germany)." captionText-1="Fig. 3. The distribution of the species of Drymeia Meigen, 1826 in Kyrgyzstan." captionText-2="Fig. 4. Drymeia ziegleri sp. nov., ♂, holotype (ZMHU), lateral view. Photo: K. Child (UK, formerly OUMNH)." captionText-3="Fig. 5. The type-locality of Drymeia ziegleri sp. nov. (Kyrgyzstan, Kalmak-Anshun Pass). Photo: J. Ziegler (Berlin, Germany)." figureDoi-0="http://doi.org/10.5281/zenodo.6658723" figureDoi-1="http://doi.org/10.5281/zenodo.6658731" figureDoi-2="http://doi.org/10.5281/zenodo.6658739" figureDoi-3="http://doi.org/10.5281/zenodo.6658747" httpUri-0="https://zenodo.org/record/6658723/files/figure.png" httpUri-1="https://zenodo.org/record/6658731/files/figure.png" httpUri-2="https://zenodo.org/record/6658739/files/figure.png" httpUri-3="https://zenodo.org/record/6658747/files/figure.png" pageId="6" pageNumber="164">Figs 25</figureCitation>
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<emphasis id="B901EAD6C330FFE0FF26960CFE93C424" bold="true" box="[189,311,1622,1648]" pageId="6" pageNumber="164">Diagnosis</emphasis>
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The new species is close to
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<emphasis id="B901EAD6C330FFE0FD9696DFFD69C4CA" box="[525,717,1669,1694]" italics="true" pageId="6" pageNumber="164">Drymeia vicana</emphasis>
(Harris, 1780)
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, which also has 4 postsutural dorsocentral setae, prealar absent and densely long-haired eyes. However,
<taxonomicName id="4C754D47C330FFE0FC0396F2FBB3C496" authorityName="Sorokina &amp; Pont" authorityYear="2022" box="[920,1047,1704,1730]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="164" phylum="Arthropoda" rank="species" species="ziegleri">
<emphasis id="B901EAD6C330FFE0FC0396F2FBB3C496" box="[920,1047,1704,1730]" italics="true" pageId="6" pageNumber="164">D. ziegleri</emphasis>
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is a larger black species with thin, elongate proboscis, mid femur slightly curved ventrally and with 3 short ventral spines closer to the middle, a row of long anteroventral setae on the apical ⅔ and a row of very long posteroventral setae; the male has the anepimeron and katepimeron bare; hind femur without long and strong posteroventrals; hind tibia with a small ventral apical prong, without apical posteroventral seta.
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<emphasis id="B901EAD6C330FFE0FF26972CFEE1C5C4" bold="true" box="[189,325,1910,1936]" pageId="6" pageNumber="164">Etymology</emphasis>
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The species epithet is a patronym in honour of the dipterist Dr Joachim Ziegler (
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) who collected the
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.
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<emphasis id="B901EAD6C331FFE1FF2697F1FEA9C591" bold="true" box="[189,269,1963,1989]" pageId="7" pageNumber="165">Fig. 4.</emphasis>
<taxonomicName id="4C754D47C331FFE1FE8297F6FE40C591" authority="Sorokina &amp; Pont, 2022" authorityName="Sorokina &amp; Pont" authorityYear="2022" box="[281,484,1963,1989]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="165" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
<emphasis id="B901EAD6C331FFE1FE8297F6FE40C591" box="[281,484,1963,1989]" italics="true" pageId="7" pageNumber="165">Drymeia ziegleri</emphasis>
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<taxonomicNameLabel id="A23257ADC331FFE1FE7097F6FDEEC592" box="[491,586,1964,1990]" pageId="7" pageNumber="165" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, holotype (ZMHU), lateral view. Photo: K. Child (UK, formerly OUMNH).
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<emphasis id="B901EAD6C33EFFEEFF269153FEC9C376" bold="true" box="[189,365,264,291]" pageId="8" pageNumber="166">Type material</emphasis>
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<typeStatus id="54CE8866C33EFFEEFF71916DFEF8C305" box="[234,348,311,337]" pageId="8" pageNumber="166" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3627654C33EFFEEFF269100FE25C320" box="[189,385,346,372]" name="Kyrgyzstan" pageId="8" pageNumber="166">KYRGYZSTAN</collectingCountry>
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;
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,
<collectingMunicipality id="6BAEACBEC33EFFEEFDEF9101FC91C320" box="[628,821,346,373]" pageId="8" pageNumber="166">Inner Tien Shan</collectingMunicipality>
,
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,
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;
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,
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;
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a.s.l.;
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<collectingDate id="EF8FE9ECC33EFFEEFBF49124FB58C3CC" box="[1135,1276,381,408]" pageId="8" pageNumber="166" value="2009-07-31">31 Jul. 2009</collectingDate>
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;
<collectorName id="26805312C33EFFEEFA929124FAD3C3C3" box="[1289,1399,381,408]" pageId="8" pageNumber="166">J. Ziegler</collectorName>
leg.;
<collectionCode id="ED64AE01C33EFFEEFF6391FBFEF6C3EF" box="[248,338,417,443]" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="166">ZMHU</collectionCode>
.
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</paragraph>
<paragraph id="8BCA36C4C33EFFEEFF7191B8FDB2C032" blockId="8.[189,1399,482,614]" pageId="8" pageNumber="166">
<materialsCitation id="3B1D3C99C33EFFEEFF7191B8FC5DC074" ID-GBIF-Occurrence="3816439317" collectingDate="2009-07-31" collectionCode="ZMHU" collectorName="J. Ziegler" country="Kyrgyzstan" county="Baidui Mtn Range" elevation="3420" latitude="41.914722" location="KYRGYZSTAN" longLatPrecision="19" longitude="75.42777" municipality="Inner Tien Shan" pageId="8" pageNumber="166" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B901EAD6C33EFFEEFF7191B8FEC3C3A8" bold="true" box="[234,359,482,508]" pageId="8" pageNumber="166">
<typeStatus id="54CE8866C33EFFEEFF7191B8FEC3C3A8" box="[234,359,482,508]" pageId="8" pageNumber="166" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="F3627654C33EFFEEFF26925FFE25C04B" box="[189,385,517,543]" name="Kyrgyzstan" pageId="8" pageNumber="166">KYRGYZSTAN</collectingCountry>
<specimenCount id="9D73FD4DC33EFFEEFE04925FFE74C074" box="[415,464,517,544]" pageId="8" pageNumber="166" type="male">1 ♂</specimenCount>
; same collection data as for holotype;
<collectionCode id="ED64AE01C33EFFEEFC38925CFC5DC074" box="[931,1017,518,544]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/99a5-d0z2" name="Zoologisches Museum der Humboldt Universitaet" pageId="8" pageNumber="166">ZMHU</collectionCode>
</materialsCitation>
<materialsCitation id="3B1D3C99C33EFFEEFB8C925FFDB6C032" ID-GBIF-Occurrence="3816439330" collectingDate="1964-08-27" collectionCode="ZMUM" collectorName="R. Zlotin" country="Kyrgyzstan" elevation="3500" latitude="41.733334" location="Tien-Shan" longLatPrecision="1154" longitude="77.88333" pageId="8" pageNumber="166" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D73FD4DC33EFFEEFB8C925FFBEDC074" box="[1047,1097,517,544]" pageId="8" pageNumber="166" type="male">1 ♂</specimenCount>
; “
<location id="8EAA601FC33EFFEEFBFA925FFB44C04B" LSID="urn:lsid:plazi:treatment:03DC87D2C330FFEFFD1F9594FE78C7ED:8EAA601FC33EFFEEFBFA925FFB44C04B" box="[1121,1248,517,543]" country="Kyrgyzstan" latitude="41.733334" longLatPrecision="1154" longitude="77.88333" name="Tien-Shan" pageId="8" pageNumber="166">Tien-Shan</location>
,
<location id="8EAA601FC33EFFEEFB74925FFEADC017" LSID="urn:lsid:plazi:treatment:03DC87D2C330FFEFFD1F9594FE78C7ED:8EAA601FC33EFFEEFB74925FFEADC017" country="Kyrgyzstan" latitude="41.733334" longLatPrecision="1154" longitude="77.88333" name="verkh. r. B. Naryn" pageId="8" pageNumber="166">verkh. r. B. Naryn</location>
” [Tien-Shan, upper reaches of B. Naryn River, ~
<geoCoordinate id="EE415003C33EFFEEFCFC927DFC72C017" box="[871,982,551,579]" degrees="41" direction="north" minutes="44" orientation="latitude" pageId="8" pageNumber="166" precision="925" value="41.733334">41°44 N</geoCoordinate>
,
<geoCoordinate id="EE415003C33EFFEEFC7F927DFBEBC017" box="[996,1103,551,579]" degrees="77" direction="east" minutes="53" orientation="longitude" pageId="8" pageNumber="166" precision="925" value="77.88333">77°53 E</geoCoordinate>
,
<elevation id="0058D1F7C33EFFEEFBC49272FB59C017" box="[1119,1277,552,579]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="166" unit="m" value="3500.0">
<quantity id="4C8D9B21C33EFFEEFBC49272FB1EC017" box="[1119,1210,552,579]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="8" pageNumber="166" unit="m" value="3500.0">3500 m</quantity>
a.s.l.
</elevation>
];
<date id="FFCB1004C33EFFEEFA8C9273FF5FC032" pageId="8" pageNumber="166" value="1964-08-27">
<collectingDate id="EF8FE9ECC33EFFEEFA8C9273FF5FC032" pageId="8" pageNumber="166" value="1964-08-27">27 Aug. 1964</collectingDate>
</date>
;
<collectorName id="26805312C33EFFEEFE9C9216FED5C032" box="[263,369,588,614]" pageId="8" pageNumber="166">R. Zlotin</collectorName>
leg.;
<collectionCode id="ED64AE01C33EFFEEFE289216FDB6C032" box="[435,530,588,614]" collectionName="Russia, Moscow, Moscow State University" country="China" httpUri="http://grbio.org/cool/49rk-3jpq" name="Zoological Museum, University of Amoy" pageId="8" pageNumber="166">ZMUM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F654FC33EFFEFFF2692D7FE9DC71C" lastPageId="9" lastPageNumber="167" pageId="8" pageNumber="166" type="description">
<paragraph id="8BCA36C4C33EFFEEFF2692D7FEEBC0F3" blockId="8.[189,335,653,679]" box="[189,335,653,679]" pageId="8" pageNumber="166">
<heading id="D08281A8C33EFFEEFF2692D7FEEBC0F3" bold="true" box="[189,335,653,679]" fontSize="11" level="3" pageId="8" pageNumber="166" reason="3">
<emphasis id="B901EAD6C33EFFEEFF2692D7FEEBC0F3" bold="true" box="[189,335,653,679]" pageId="8" pageNumber="166">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36C4C33EFFEEFF7192E6FE8EC082" blockId="8.[189,984,700,762]" box="[234,298,700,726]" pageId="8" pageNumber="166">
<emphasis id="B901EAD6C33EFFEEFF7192E6FE8EC082" bold="true" box="[234,298,700,726]" pageId="8" pageNumber="166">Male</emphasis>
</paragraph>
<paragraph id="8BCA36C4C33EFFEEFF269285FC7CC0AD" blockId="8.[189,984,700,762]" box="[189,984,735,762]" pageId="8" pageNumber="166">
MEASUREMENTS. Body length
<quantity id="4C8D9B21C33EFFEEFD8E9285FD01C0AD" box="[533,677,735,761]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="8.2" metricValueMin="6.8" pageId="8" pageNumber="166" unit="mm" value="7.5" valueMax="8.2" valueMin="6.8">6.88.2 mm</quantity>
; wing length
<quantity id="4C8D9B21C33EFFEEFCDF9285FC70C0AD" box="[836,980,735,762]" metricMagnitude="-3" metricUnit="m" metricValue="5.95" metricValueMax="6.5" metricValueMin="5.4" pageId="8" pageNumber="166" unit="mm" value="5.95" valueMax="6.5" valueMin="5.4">5.46.5 mm</quantity>
.
</paragraph>
<paragraph id="8BCA36C4C33EFFEFFF26937BFD60C33D" blockId="8.[188,1400,800,1038]" lastBlockId="9.[189,1400,265,362]" lastPageId="9" lastPageNumber="167" pageId="8" pageNumber="166">HEAD. Ground-colour black. Eye densely long-haired. Fronto-orbital plate and parafacial, face, gena and lower occiput brownish-grey pruinose. Fronto-orbital plates not touching, separated by a narrow frontal vitta. Frons at narrowest point 3 × as wide as diameter of anterior ocellus. 1618 pairs of frontal setae, including interstitials, reaching almost to anterior ocellus. Antenna black, postpedicel 1.5× as long as wide. Arista appearing almost bare, longest hairs much shorter than its basal diameter. Parafacial at level of insertion of arista slightly wider than or equal to width of postpedicel, hardly narrowing below. In lateral view, lower facial margin not projecting forward beyond level of antennal base. Gena broad, depth below lowest eye-margin equal to or slightly broader than length of postpedicel, densely setose and with a group of upcurved setae on anterior part of genal dilation. Palpus black. Proboscis thin, elongate, prementum dusted but subshining.</paragraph>
<caption id="DF0A664CC33EFFEEFF2597F0FE5EC5B3" ID-DOI="http://doi.org/10.5281/zenodo.6658747" ID-Zenodo-Dep="6658747" httpUri="https://zenodo.org/record/6658747/files/figure.png" pageId="8" pageNumber="166" startId="8.[190,233,1962,1988]" targetBox="[189,1398,1116,1922]" targetPageId="8">
<paragraph id="8BCA36C4C33EFFEEFF2597F0FE5EC5B3" blockId="8.[189,1398,1962,2023]" pageId="8" pageNumber="166">
<emphasis id="B901EAD6C33EFFEEFF2597F0FEABC590" bold="true" box="[190,271,1962,1988]" pageId="8" pageNumber="166">Fig. 5.</emphasis>
The type-locality of
<taxonomicName id="4C754D47C33EFFEEFD8997F1FD79C590" authority="Sorokina &amp; Pont, 2022" authorityName="Sorokina &amp; Pont" authorityYear="2022" box="[530,733,1962,1988]" class="Insecta" family="Muscidae" genus="Drymeia" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="166" phylum="Arthropoda" rank="species" species="ziegleri" status="sp. nov.">
<emphasis id="B901EAD6C33EFFEEFD8997F1FD79C590" box="[530,733,1962,1988]" italics="true" pageId="8" pageNumber="166">Drymeia ziegleri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257ADC33EFFEEFD7397F0FCE1C590" box="[744,837,1962,1988]" pageId="8" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
(Kyrgyzstan, Kalmak-Anshun Pass). Photo: J. Ziegler (Berlin, Germany).
</paragraph>
</caption>
<paragraph id="8BCA36C4C33FFFEFFF2691CBFC70C040" blockId="9.[189,1399,401,532]" pageId="9" pageNumber="167">THORAX. Ground-colour black. Scutum matt, black, brownish-grey dusted. Ground-setulae long and dense. Pleura black, subshining. Acrostichals indistinct, the presutural setulae in 45 irregular rows. Dorsocentrals 2+4. Prealar seta absent. Notopleuron densely setulose. Prosternum bare. Anepimeron and katepimeron bare. Scutellum black and dusted in anterior view.</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF269266FC12C18D" blockId="9.[188,1401,572,986]" pageId="9" pageNumber="167">LEGS. Black. Fore tibia with 01 posterodorsal and 23 posteroventral setae in middle. Fore tarsomere 1 with a row of posteroventral setulae on apical third. Mid femur slightly curved ventrally, with a row of long anteroventral setae on apical ⅔, the setae in middle part of femur directed upwards and slightly shifted on to anterior surface, apical setae directed downwards; with 3 short ventral spines closer to middle; with a row of very long posteroventral setae, 2.32.5 × as long as diameter of femur, except in apical and basal quarters where long setae are absent. Mid tibia without long strong anterior setae; 67 posterodorsals; 45 posteroventrals. Hind femur without posteroventrals but with a tuft of fine long setae on basal half of posterior surface; with a row of anteroventral setae that become longer and stronger on apical third. Hind tibia with a row of long, strong posterodorsals on basal half; with a row of anterodorsals, strong and long on basal half, short and erect on apical half; 46 anteroventrals and 45 posteroventrals in middle; without preapical setae, except 1 short anteroventral seta on small ventral apical prong. Hind tarsomere 1 with 2 short ventral setae at base.</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF26945BFC1FC66A" blockId="9.[189,1398,1024,1086]" pageId="9" pageNumber="167">WING. Brown, blackish at base. Basicosta and tegula black. Costa with weak spinules, without costal spine. Calypters and margins light yellow. Knob of haltere black.</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF26943FFE1FC6F6" blockId="9.[189,1398,1125,1187]" pageId="9" pageNumber="167">ABDOMEN. Ground-colour black. When viewed from behind, tergites 35 with brown dusted lateral spots. Sternite 1 bare.</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF269490FE5DC6B0" blockId="9.[189,505,1226,1252]" box="[189,505,1226,1252]" pageId="9" pageNumber="167">TERMINALIA. Not dissected.</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF719551FEE1C771" blockId="9.[189,325,1291,1352]" box="[234,325,1291,1317]" pageId="9" pageNumber="167">
<emphasis id="B901EAD6C33FFFEFFF719551FEE1C771" bold="true" box="[234,325,1291,1317]" pageId="9" pageNumber="167">Female</emphasis>
</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF269574FE9DC71C" blockId="9.[189,325,1291,1352]" box="[189,313,1326,1352]" pageId="9" pageNumber="167">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C36F654FC33FFFEFFF26952AFE78C7ED" box="[189,476,1392,1465]" pageId="9" pageNumber="167" type="distribution">
<paragraph id="8BCA36C4C33FFFEFFF26952AFEF2C7DE" blockId="9.[189,342,1392,1418]" box="[189,342,1392,1418]" pageId="9" pageNumber="167">
<heading id="D08281A8C33FFFEFFF26952AFEF2C7DE" bold="true" box="[189,342,1392,1418]" fontSize="11" level="3" pageId="9" pageNumber="167" reason="3">
<emphasis id="B901EAD6C33FFFEFFF26952AFEF2C7DE" bold="true" box="[189,342,1392,1418]" pageId="9" pageNumber="167">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BCA36C4C33FFFEFFF2695C4FE78C7ED" blockId="9.[189,476,1438,1465]" box="[189,476,1438,1465]" pageId="9" pageNumber="167">
Palaearctic:
<collectingCountry id="F3627654C33FFFEFFED695C5FE7CC7ED" box="[333,472,1439,1465]" name="Kyrgyzstan" pageId="9" pageNumber="167">Kyrgyzstan</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>