274 lines
34 KiB
XML
274 lines
34 KiB
XML
<document id="1C22F7A9D29BD1026F425C3A3EA400ED" ID-CLB-Dataset="29085" ID-DOI="10.11646/zootaxa.4455.3.7" ID-GBIF-Dataset="e805d477-ca64-4ea3-8990-285677f540b1" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457495" ID-ZooBank="6B5ED1A9-FC34-4142-85AD-B35C73A427DE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539263023203" checkinUser="plazi" docAuthor="Fanti, Fabrizio & Pankowski, Mary K." docDate="2018" docId="03DC87CAFF8DFFD2FF2F13F8FF12FC85" docLanguage="en" docName="zootaxa.4455.3.7.pdf" docOrigin="Zootaxa 4455 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhagonycha (Rhagonycha) maryae Fanti & Pankowski 2018" docType="treatment" docVersion="7" lastPageNumber="516" masterDocId="FFE5FFB2FF8CFFD1FFB8106EFFD6FFBD" masterDocTitle="Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae)" masterLastPageNumber="524" masterPageNumber="513" pageNumber="514" updateTime="1698576187060" updateUser="ExternalLinkService">
|
||
<mods:mods id="17BDA0928F321E0066D54CB4A2671595" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="8E8F9D16EDF1B4859D3C517F9F25F077">
|
||
<mods:title id="4864D5A369B37B27ADEA1F8B222EAFC5">Three new species of soldier beetles from Baltic amber (Coleoptera, Cantharidae)</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="8459E80BF43F5BD25AB16ADD514C7C38" type="personal">
|
||
<mods:role id="BE8CCACF7D8B25D05536EB76854A0164">
|
||
<mods:roleTerm id="A3F606CE6EDA2B085D1CD8B3511C0B51">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="2D5AEE1807EC52E3CECE60E717F48E70">Fanti, Fabrizio</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="60381595D03A8653E682DADE49D6827C" type="personal">
|
||
<mods:role id="67E1A952D37B94A52225A560F78AB91F">
|
||
<mods:roleTerm id="142B8DA24DE9351DFC5366E7EFFF616A">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4E59A16DC1F9957BB191A6140285396E">Pankowski, Mary K.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="C02CE0582E93F9F4E063819E52618791">text</mods:typeOfResource>
|
||
<mods:relatedItem id="BC4E959C41F04F349BFBA4BDE1BAD56D" type="host">
|
||
<mods:titleInfo id="726B4ABBF2A33378F89473599ACF0E42">
|
||
<mods:title id="9CBF9F67DD84E7D82FC2CF8027F1135E">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="1FE9C24F2478CF493A38C7D4DBC91100">
|
||
<mods:date id="8938D49D4F4D9CF823DCF50CC4CEDE8B">2018</mods:date>
|
||
<mods:detail id="F2E61C5EDD2A17FC6E264AB8C7F67D1F" type="pubDate">
|
||
<mods:number id="98D58C2ACA4866868C1EA5CD481357E4">2018-08-03</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="53C8D0A44AEB270FE27E71C5706241A1" type="volume">
|
||
<mods:number id="4442C6D461E13A2ABDC15346B22F4B64">4455</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="316DF855C5F303EB5F2CE5A066A6785E" type="issue">
|
||
<mods:number id="83B17CDABF0A182599193E96CF2A0335">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4DF96A964103B8DC72131A45DFD1616B" unit="page">
|
||
<mods:start id="8BB6D1768F674AAEED80471958D9719E">513</mods:start>
|
||
<mods:end id="9518174A2281E71A67F4281EB53149C7">524</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="CE296D425B9480DCEC6EEA305D73EAAF">journal article</mods:classification>
|
||
<mods:identifier id="C0AB0155D320A2CF870556BC5FE9B64B" type="CLB-Dataset">29085</mods:identifier>
|
||
<mods:identifier id="23ED8089B178AE0AE0D6052FA770E6B2" type="DOI">10.11646/zootaxa.4455.3.7</mods:identifier>
|
||
<mods:identifier id="D0F4792E8F21A021603E0A354C606BDE" type="GBIF-Dataset">e805d477-ca64-4ea3-8990-285677f540b1</mods:identifier>
|
||
<mods:identifier id="44B52E70CFAFD6C99FA22A6B23C38667" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="1D3278E94727FA454C59042722832DF0" type="Zenodo-Dep">1457495</mods:identifier>
|
||
<mods:identifier id="F1BB620C3F0A345266749A708361D1F8" type="ZooBank">6B5ED1A9-FC34-4142-85AD-B35C73A427DE</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03DC87CAFF8DFFD2FF2F13F8FF12FC85" ID-DOI="http://doi.org/10.5281/zenodo.5961866" ID-GBIF-Taxon="148691444" ID-Zenodo-Dep="5961866" LSID="urn:lsid:plazi:treatment:03DC87CAFF8DFFD2FF2F13F8FF12FC85" httpUri="http://treatment.plazi.org/id/03DC87CAFF8DFFD2FF2F13F8FF12FC85" lastPageId="3" lastPageNumber="516" pageId="1" pageNumber="514">
|
||
<subSubSection id="C36F6557FF8DFFD0FF2F13F8FF3DFC6F" pageId="1" pageNumber="514" type="nomenclature">
|
||
<paragraph id="8BCA36DCFF8DFFD0FF2F13F8FBEEFC0D" blockId="1.[151,1080,918,978]" box="[151,1080,918,944]" pageId="1" pageNumber="514">
|
||
<heading id="D08281B0FF8DFFD0FF2F13F8FBEEFC0D" bold="true" box="[151,1080,918,944]" fontSize="11" level="1" pageId="1" pageNumber="514" reason="1">
|
||
<emphasis id="B901EACEFF8DFFD0FF2F13F8FBEEFC0D" bold="true" box="[151,1080,918,944]" pageId="1" pageNumber="514">
|
||
<taxonomicName id="4C754D5FFF8DFFD0FF2F13F8FD92FC0D" authorityName="Fanti & Pankowski" authorityYear="2018" box="[151,580,918,944]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="514" phylum="Arthropoda" rank="species" species="maryae" subGenus="Rhagonycha">
|
||
<emphasis id="B901EACEFF8DFFD0FF2F13F8FEE6FC0D" bold="true" box="[151,304,918,944]" italics="true" pageId="1" pageNumber="514">Rhagonycha</emphasis>
|
||
(
|
||
<emphasis id="B901EACEFF8DFFD0FEF913F8FE0CFC0D" bold="true" box="[321,474,918,944]" italics="true" pageId="1" pageNumber="514">Rhagonycha</emphasis>
|
||
)
|
||
<emphasis id="B901EACEFF8DFFD0FE5313F8FD92FC0D" bold="true" box="[491,580,918,944]" italics="true" pageId="1" pageNumber="514">maryae</emphasis>
|
||
</taxonomicName>
|
||
FANTI & M. K. PANKOWSKI sp. nov.
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCA36DCFF8DFFD0FF2F13D7FF3DFC6F" blockId="1.[151,1080,918,978]" box="[151,235,953,978]" pageId="1" pageNumber="514">
|
||
(
|
||
<figureCitation id="134E2A59FF8DFFD0FF2713D7FF36FC6F" box="[159,224,953,978]" captionStart="FIGURE 1" captionStartId="2.[151,250,1899,1921]" captionTargetBox="[153,1436,267,1874]" captionTargetId="figure@2.[150,1436,267,1874]" captionTargetPageId="2" captionText="FIGURE 1. Rhagonycha maryae sp. nov. in Baltic amber. A: Holotype, dorsal view, scale bar = 1.0 mm. B: Holotype, detail of head, pronotum and left antenna, scale bar = 0.5 mm. C: Holotype, ventral view, scale bar = 0.5 mm. D: Holotype, detail of leg and claws, scale bar = 0.2 mm." httpUri="https://zenodo.org/record/1457497/files/figure.png" pageId="1" pageNumber="514">Fig. 1</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6557FF8DFFD0FF2F1394FE38F8BA" pageId="1" pageNumber="514" type="description">
|
||
<paragraph id="8BCA36DCFF8DFFD0FF2F1394FE38F8BA" blockId="1.[151,1437,1018,2015]" pageId="1" pageNumber="514">
|
||
<emphasis id="B901EACEFF8DFFD0FF2F1394FEFFFBAE" bold="true" box="[151,297,1018,1043]" pageId="1" pageNumber="514">Description.</emphasis>
|
||
Adult, winged, slender and elongated. Female, defined on the basis of the large last sternite as wide as last tergite and apically sinuous and concave in the middle. Body length:
|
||
<quantity id="4C8D9B39FF8DFFD0FBBA1471FBB6FB8B" box="[1026,1120,1055,1079]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="1" pageNumber="514" unit="mm" value="3.7">3.7 mm</quantity>
|
||
; elytra:
|
||
<quantity id="4C8D9B39FF8DFFD0FB7B1471FAF7FB8B" box="[1219,1313,1055,1079]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="1" pageNumber="514" unit="mm" value="3.1">3.1 mm</quantity>
|
||
; antennae: approximately
|
||
<quantity id="4C8D9B39FF8DFFD0FEFE142DFE73FBE7" box="[326,421,1091,1115]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="514" unit="mm" value="2.8">2.8 mm</quantity>
|
||
. Entirely dark brown including legs and antennae. Head almost completely exposed, rounded, with thin and shallow punctation. Eyes rounded, convex, inserted laterally to the head, inter-ocular dorsal distance about 2.4 times greater than eye diameter. Mandibles not visible. Maxillary palps 4-segmented, unequal in length, with the last palpomere securiform. Labial palps 3-segmented, last segment securiform. Antennae filiform, 11-segmented and with all segments pubescent, relative short, surpassing half of elytra; antennomere I (scape) robust, elongated, club–shaped; antennomere II robust, short, 1.7 times shorter than scape; antennomeres III–V elongated, robust (particularly the third), slightly shorter than first article; antennomeres VI–IX filiform, subequal, thinner and around 0.1–0.2 times shorter than previous three articles; antennomere X filiform, approximately 0.3 times shorter than previous one; antennomere XI filiform as long as antennomeres VI–IX. Pronotum larger basally, trapezoidal, larger than head, posterior corners rounded, posterior margin strongly bordered, anterior margin and sides slightly bordered, sides concave at half length, surface irregular and equipped with short, thick pubescence. Scutellum subquadrate with apex truncate and straight. Elytra slender, parallel-sided, very elongated and completely covering and slightly surpassing the last abdominal segments, wider than pronotum, apex rounded, surface rugose and equipped with many short setae. Posterior wings present, almost completely covered by elytra. Metasternum dark brown, rounded posteriorly, equipped with setae; sternites short, transverse, wrinkled; last sternite wide, rounded and sinuous at apex with central concavity. Legs short and pubescent; coxae massive; trochanters rounded; femora slightly enlarged; tibiae thin, cylindrical, longer than femora. All tarsi 5-segmented; first segment elongated; second tarsomere elongated and longer than first tarsomere; third about 2.3 times shorter than second, straight at apex; fourth tarsomere bilobed at sides; fifth flat and elongated; claws apparently bifid at apex without denticles at base.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6557FF8DFFD0FF7F177CFE22F8CE" pageId="1" pageNumber="514" type="etymology">
|
||
<paragraph id="8BCA36DCFF8DFFD0FF7F177CFE22F8CE" blockId="1.[151,1437,1018,2015]" pageId="1" pageNumber="514">
|
||
<emphasis id="B901EACEFF8DFFD0FF7F177CFE98F896" bold="true" box="[199,334,1810,1835]" pageId="1" pageNumber="514">Etymology.</emphasis>
|
||
Species named after Mary L. Pankowski, paternal grandmother of the second author, who provides immense love to her family, supports her community in numerous ways and inspires all with her indomitable spirit of optimism.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6557FF8DFFD3FF7F1710FDF7FF68" lastPageId="2" lastPageNumber="515" pageId="1" pageNumber="514" type="materials_examined">
|
||
<paragraph id="8BCA36DCFF8DFFD0FF7F1710FA4BF82A" blockId="1.[151,1437,1018,2015]" box="[199,1437,1918,1943]" pageId="1" pageNumber="514">
|
||
<emphasis id="B901EACEFF8DFFD0FF7F1710FEEDF82A" bold="true" box="[199,315,1918,1943]" pageId="1" pageNumber="514">
|
||
<typeStatus id="54CE887EFF8DFFD0FF7F1710FEE0F82A" box="[199,310,1918,1943]" pageId="1" pageNumber="514" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Female, adult specimen included in Baltic amber, accession No.
|
||
<collectionCode id="ED64AE19FF8DFFD0FBAB1711FBB2F82A" box="[1043,1124,1919,1943]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="514">USNM</collectionCode>
|
||
<collectionCode id="ED64AE19FF8DFFD0FBD31711FB4BF82A" box="[1131,1181,1919,1943]" country="Italy" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15292" name="Herbarium Mediterraneum Panormitanum" pageId="1" pageNumber="514">PAL</collectionCode>
|
||
<quantity id="4C8D9B39FF8DFFD0FB1C1710FAC3F82A" box="[1188,1301,1918,1943]" metricMagnitude="4" metricUnit="m" metricValue="1.8098363599999998" pageId="1" pageNumber="514" unit="in" value="712534.0">712534 in</quantity>
|
||
the
|
||
<collectionCode id="ED64AE19FF8DFFD0FAFC1711FA4FF82A" box="[1348,1433,1919,1943]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="514">USNM</collectionCode>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BCA36DCFF8DFFD0FF7F17CCFE9EF862" blockId="1.[151,1437,1018,2015]" pageId="1" pageNumber="514">
|
||
<materialsCitation id="3B1D3C81FF8DFFD0FF7F17CCFE9EF862" ID-GBIF-Occurrence="1917075957" country="Russia" county="Peninsula" location="Yantarny" municipality="Samland" pageId="1" pageNumber="514" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
|
||
<emphasis id="B901EACEFF8DFFD0FF7F17CCFEB9F806" bold="true" box="[199,367,1954,1979]" pageId="1" pageNumber="514">
|
||
<typeStatus id="54CE887EFF8DFFD0FF7F17CCFED4F806" box="[199,258,1954,1979]" pageId="1" pageNumber="514">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EAA6007FF8DFFD0FEC717CDFE32F806" LSID="urn:lsid:plazi:treatment:03DC87CAFF8DFFD2FF2F13F8FF12FC85:8EAA6007FF8DFFD0FEC717CDFE32F806" box="[383,484,1955,1979]" country="Russia" county="Peninsula" municipality="Samland" name="Yantarny" pageId="1" pageNumber="514" stateProvince="Kaliningrad">Yantarny</location>
|
||
settlement (formerly
|
||
<location id="8EAA6007FF8DFFD0FD4D17CCFCA8F806" LSID="urn:lsid:plazi:treatment:03DC87CAFF8DFFD2FF2F13F8FF12FC85:8EAA6007FF8DFFD0FD4D17CCFCA8F806" box="[757,894,1954,1979]" country="Russia" county="Peninsula" municipality="Samland" name="Palmnicken" pageId="1" pageNumber="514" stateProvince="Kaliningrad">Palmnicken</location>
|
||
),
|
||
<location id="8EAA6007FF8DFFD0FC2217CCFC2AF806" LSID="urn:lsid:plazi:treatment:03DC87CAFF8DFFD2FF2F13F8FF12FC85:8EAA6007FF8DFFD0FC2217CCFC2AF806" box="[922,1020,1954,1979]" country="Russia" county="Peninsula" municipality="Samland" name="Sambian" pageId="1" pageNumber="514" stateProvince="Kaliningrad">Sambian</location>
|
||
(
|
||
<collectingMunicipality id="6BAEACA6FF8DFFD0FBAC17CCFBAFF806" box="[1044,1145,1954,1979]" pageId="1" pageNumber="514">Samland</collectingMunicipality>
|
||
)
|
||
<collectingCounty id="62AB4E50FF8DFFD0FB2A17CCFAD4F806" box="[1170,1282,1954,1979]" pageId="1" pageNumber="514">Peninsula</collectingCounty>
|
||
,
|
||
<collectingRegion id="49B1F83EFF8DFFD0FAAE17CCFA4AF806" box="[1302,1436,1954,1979]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="514">Kaliningrad</collectingRegion>
|
||
Region,
|
||
<collectingCountry id="F362764CFF8DFFD0FF4E17A8FE92F862" box="[246,324,1990,2015]" name="Russia" pageId="1" pageNumber="514">Russia</collectingCountry>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BCA36DCFF8EFFD3FF7F10F9FDF7FF68" blockId="2.[199,1409,151,213]" pageId="2" pageNumber="515">
|
||
<emphasis id="B901EACEFF8EFFD3FF7F10F9FE83FF0D" bold="true" box="[199,341,151,176]" pageId="2" pageNumber="515">
|
||
<typeStatus id="54CE887EFF8EFFD3FF7F10F9FED4FF0D" box="[199,258,151,176]" pageId="2" pageNumber="515">Type</typeStatus>
|
||
strata.
|
||
</emphasis>
|
||
Baltic amber, Upper Eocene, Prussian Formation (Priabonian). Estimated age: 37.8–33.9 Mya.
|
||
<emphasis id="B901EACEFF8EFFD3FF7F10D2FEA7FF68" bold="true" box="[199,369,188,213]" pageId="2" pageNumber="515">Syninclusions.</emphasis>
|
||
Wood remains.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0A6654FF8EFFD3FF2F1705FE07F803" httpUri="https://zenodo.org/record/1457497/files/figure.png" pageId="2" pageNumber="515" startId="2.[151,250,1899,1921]" targetBox="[153,1436,267,1874]" targetPageId="2">
|
||
<paragraph id="8BCA36DCFF8EFFD3FF2F1705FE07F803" blockId="2.[151,1435,1899,1982]" pageId="2" pageNumber="515">
|
||
<emphasis id="B901EACEFF8EFFD3FF2F1705FEC4F83D" bold="true" box="[151,274,1899,1921]" pageId="2" pageNumber="515">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C754D5FFF8EFFD3FEA01705FE3EF83C" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[280,488,1899,1921]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="515" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
|
||
<emphasis id="B901EACEFF8EFFD3FEA01705FE3EF83C" box="[280,488,1899,1921]" italics="true" pageId="2" pageNumber="515">Rhagonycha maryae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257B5FF8EFFD3FE561705FDE9F83D" box="[494,575,1899,1920]" pageId="2" pageNumber="515" rank="species">
|
||
<emphasis id="B901EACEFF8EFFD3FE561705FDE9F83D" bold="true" box="[494,575,1899,1920]" pageId="2" pageNumber="515">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
in Baltic amber. A: Holotype, dorsal view, scale bar = 1.0 mm. B: Holotype, detail of head, pronotum and left antenna, scale bar = 0.5 mm. C: Holotype, ventral view, scale bar = 0.5 mm. D: Holotype, detail of leg and claws, scale bar = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36F6557FF8FFFD2FF7F10F9FD9BFD4D" pageId="3" pageNumber="516" type="diagnosis">
|
||
<paragraph id="8BCA36DCFF8FFFD2FF7F10F9FD9BFD4D" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="516">
|
||
<emphasis id="B901EACEFF8FFFD2FF7F10F9FE1BFF0D" bold="true" box="[199,461,151,176]" pageId="3" pageNumber="516">Differential diagnosis.</emphasis>
|
||
The new species belongs to the genus
|
||
<taxonomicName id="4C754D5FFF8FFFD2FCC210F9FB01FF0D" authority="Eschscholtz, 1830" authorityName="Eschscholtz" authorityYear="1830" box="[890,1239,151,176]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF8FFFD2FCC210F9FBD0FF0D" box="[890,1030,151,176]" italics="true" pageId="3" pageNumber="516">Rhagonycha</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FBB410F9FB01FF0D" box="[1036,1239,151,176]" pageId="3" pageNumber="516" refId="ref5218" refString="Eschscholtz, J. F. G. von (1830) Nova genera Coleopterorum Faunae Europaeae. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 2 (1), 63 - 66." type="journal article">Eschscholtz, 1830</bibRefCitation>
|
||
</taxonomicName>
|
||
based on its bifid claws and for its anteriorly narrowed pronotum. Two species of the genus
|
||
<taxonomicName id="4C754D5FFF8FFFD2FBB510D2FB4FFF68" authorityName="Eschscholtz" authorityYear="1830" box="[1037,1177,188,213]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF8FFFD2FBB510D2FB4FFF68" box="[1037,1177,188,213]" italics="true" pageId="3" pageNumber="516">Rhagonycha</emphasis>
|
||
</taxonomicName>
|
||
have previously been described from inclusions in Baltic amber:
|
||
<taxonomicName id="4C754D5FFF8FFFD2FD29108FFB73FF45" authority="(Yablokov-Khnzorian, 1960)" authorityName="Yablokov-Khnzorian" authorityYear="1960" baseAuthorityName="Yablokov-Khnzorian" baseAuthorityYear="1960" box="[657,1189,223,248]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="kryshtofovichi">
|
||
<emphasis id="B901EACEFF8FFFD2FD29108FFC85FF45" box="[657,851,223,248]" italics="true" pageId="3" pageNumber="516">R. kryshtofovichi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FCDC10B1FB4BFF45" box="[868,1181,223,248]" pageId="3" pageNumber="516" refId="ref7394" refString="Yablokov-Khnzorian, S. M. (1960) New beetles from the Baltic amber. Paleontologicheskii Zhurnal, 3, 90 - 101. [in Russian, title: Novye zhestkokrylye iz Baltiyskogo yantarya]" type="journal article">Yablokov-Khnzorian, 1960</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C754D5FFF8FFFD2FB5A108FFE4AFEA0" authority="(Poinar & Fanti, 2016)" authorityName="Poinar & Fanti" authorityYear="2016" baseAuthorityName="Poinar & Fanti" baseAuthorityYear="2016" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sucinobaltica">
|
||
<emphasis id="B901EACEFF8FFFD2FB5A108FFA4AFF45" box="[1250,1436,223,248]" italics="true" pageId="3" pageNumber="516">R. sucinobaltica</emphasis>
|
||
(
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FF26116AFE42FEA0" box="[158,404,260,285]" pageId="3" pageNumber="516" refId="ref6966" refString="Poinar, G. O. Jr. & Fanti, F. (2016) New fossil soldier beetles (Coleoptera: Cantharidae) in Burmese, Baltic and Dominican amber. Palaeodiversity, 9, 1 - 17. https: // doi. org / 10.18476 / pale. v 9. a 1" type="journal article">Poinar & Fanti, 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C754D5FFF8FFFD2FE1F116AFD0CFEA0" baseAuthorityName="Yablokov-Khnzorian" baseAuthorityYear="1960" box="[423,730,260,285]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="kryshtofovichi">
|
||
<emphasis id="B901EACEFF8FFFD2FE1F116AFD0CFEA0" box="[423,730,260,285]" italics="true" pageId="3" pageNumber="516">Rhagonycha kryshtofovichi</emphasis>
|
||
</taxonomicName>
|
||
initially attributed to the genus
|
||
<taxonomicName id="4C754D5FFF8FFFD2FBFC116AFA4AFEA0" authority="Kiesenwetter, 1863" authorityName="Kiesenwetter" authorityYear="1863" box="[1092,1436,260,285]" class="Insecta" family="Cantharidae" genus="Malchinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF8FFFD2FBFC116AFB6CFEA0" box="[1092,1210,260,285]" italics="true" pageId="3" pageNumber="516">Malchinus</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FB7B116AFA4AFEA0" box="[1219,1436,260,285]" pageId="3" pageNumber="516" refId="ref6290" refString="Kiesenwetter, E. A. H. von (1863) Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Vierter Band. Nicolaische Verlagsbuchhandlung, Berlin, vi + 745 + [1] pp. [issued in parts, pp. 1 - 178 in 1857; pp. 179 - 386 in 1858; pp. 387 - 570 in 1860; pp. 571 - 745 + [1] in 1863]." type="journal article">Kiesenwetter, 1863</bibRefCitation>
|
||
</taxonomicName>
|
||
was then transferred to
|
||
<taxonomicName id="4C754D5FFF8FFFD2FE1D1149FDE7FEFD" authorityName="Eschscholtz" authorityYear="1830" box="[421,561,295,320]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF8FFFD2FE1D1149FDE7FEFD" box="[421,561,295,320]" italics="true" pageId="3" pageNumber="516">Rhagonycha</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FDE61146FCF1FEFD" box="[606,807,296,320]" pageId="3" pageNumber="516" refId="ref6094" refString="Kazantsev, S. V. (2013) New taxa of Baltic amber soldier beetles (Insecta: Coleoptera: Cantharidae) with synonymic and taxonomic notes. Russian Entomological Journal, 22 (4), 283 - 291." type="journal article">Kazantsev (2013)</bibRefCitation>
|
||
based on the anteriorly narrowed pronotum, the fourth palpomere securiform and unmodified
|
||
<emphasis id="B901EACEFF8FFFD2FDEA1122FD11FED8" box="[594,711,332,357]" italics="true" pageId="3" pageNumber="516">terminalia</emphasis>
|
||
. In the genus
|
||
<taxonomicName id="4C754D5FFF8FFFD2FCD41122FC2EFED8" authorityName="Eschscholtz" authorityYear="1830" box="[876,1016,332,357]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF8FFFD2FCD41122FC2EFED8" box="[876,1016,332,357]" italics="true" pageId="3" pageNumber="516">Rhagonycha</emphasis>
|
||
</taxonomicName>
|
||
males and females possess the same pronotal shape. Thus, because the pronotal shape of
|
||
<taxonomicName id="4C754D5FFF8FFFD2FD53111FFC89FE35" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[747,863,369,392]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
|
||
<emphasis id="B901EACEFF8FFFD2FD53111FFC89FE35" box="[747,863,369,392]" italics="true" pageId="3" pageNumber="516">R. maryae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257B5FF8FFFD2FCDF1101FC14FE35" box="[871,962,367,392]" pageId="3" pageNumber="516" rank="species">
|
||
<emphasis id="B901EACEFF8FFFD2FCDF1101FC14FE35" bold="true" box="[871,962,367,392]" pageId="3" pageNumber="516">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is different from
|
||
<taxonomicName id="4C754D5FFF8FFFD2FB34111FFA9DFE35" baseAuthorityName="Yablokov-Khnzorian" baseAuthorityYear="1960" box="[1164,1355,367,392]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="kryshtofovichi">
|
||
<emphasis id="B901EACEFF8FFFD2FB34111FFA9DFE35" box="[1164,1355,367,392]" italics="true" pageId="3" pageNumber="516">R. kryshtofovichi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C754D5FFF8FFFD2FA3C111FFEF8FE10" baseAuthorityName="Poinar & Fanti" baseAuthorityYear="2016" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sucinobaltica">
|
||
<emphasis id="B901EACEFF8FFFD2FA3C111FFEF8FE10" italics="true" pageId="3" pageNumber="516">R. sucinobaltica</emphasis>
|
||
</taxonomicName>
|
||
, it can be distinguished as a new species. (It’s worth noting here that the specimen of
|
||
<taxonomicName id="4C754D5FFF8FFFD2FB4611FBFAA4FE11" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[1278,1394,405,428]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
|
||
<emphasis id="B901EACEFF8FFFD2FB4611FBFAA4FE11" box="[1278,1394,405,428]" italics="true" pageId="3" pageNumber="516">R. maryae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257B5FF8FFFD2FAC211FAFF1EFE6D" pageId="3" pageNumber="516" rank="species">
|
||
<emphasis id="B901EACEFF8FFFD2FAC211FAFF1EFE6D" bold="true" pageId="3" pageNumber="516">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is a female because its last sternite is as wide as its last tergite, and it has an apical margin sinuous and concave in the middle. In the males of
|
||
<taxonomicName id="4C754D5FFF8FFFD2FDB911B2FD5BFE48" authorityName="Eschscholtz" authorityYear="1830" box="[513,653,476,501]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B901EACEFF8FFFD2FDB911B2FD5BFE48" box="[513,653,476,501]" italics="true" pageId="3" pageNumber="516">Rhagonycha</emphasis>
|
||
</taxonomicName>
|
||
, the last tergite is triangular and narrower than the last tergite.)
|
||
<taxonomicName id="4C754D5FFF8FFFD2FA3C11B3FEE0FDA5" baseAuthorityName="Yablokov-Khnzorian" baseAuthorityYear="1960" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="kryshtofovichi">
|
||
<emphasis id="B901EACEFF8FFFD2FA3C11B3FEE0FDA5" italics="true" pageId="3" pageNumber="516">R. kryshtofovichi</emphasis>
|
||
</taxonomicName>
|
||
is easily distinguishable from
|
||
<taxonomicName id="4C754D5FFF8FFFD2FD2D126FFCDCFDA5" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[661,778,513,536]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
|
||
<emphasis id="B901EACEFF8FFFD2FD2D126FFCDCFDA5" box="[661,778,513,536]" italics="true" pageId="3" pageNumber="516">R. maryae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257B5FF8FFFD2FCAB1191FCB8FDA5" box="[787,878,511,536]" pageId="3" pageNumber="516" rank="species">
|
||
<emphasis id="B901EACEFF8FFFD2FCAB1191FCB8FDA5" bold="true" box="[787,878,511,536]" pageId="3" pageNumber="516">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
based on its larger size (with a body length of
|
||
<quantity id="4C8D9B39FF8FFFD2FA36126EFF15FD81" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="516" unit="mm" value="7.0">7 mm</quantity>
|
||
compared to
|
||
<quantity id="4C8D9B39FF8FFFD2FEE6124AFE62FD81" box="[350,436,548,572]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="3" pageNumber="516" unit="mm" value="3.7">3.7 mm</quantity>
|
||
for
|
||
<taxonomicName id="4C754D5FFF8FFFD2FE5C124BFD8EFD81" authorityName="Fanti & Pankowski" authorityYear="2018" box="[484,600,549,572]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="maryae">
|
||
<emphasis id="B901EACEFF8FFFD2FE5C124BFD8EFD81" box="[484,600,549,572]" italics="true" pageId="3" pageNumber="516">R. maryae</emphasis>
|
||
</taxonomicName>
|
||
), different pronotum and different length of the antennomeres (Yablokov-
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FF2F1229FE86FDDD" box="[151,336,583,608]" pageId="3" pageNumber="516" refId="ref7394" refString="Yablokov-Khnzorian, S. M. (1960) New beetles from the Baltic amber. Paleontologicheskii Zhurnal, 3, 90 - 101. [in Russian, title: Novye zhestkokrylye iz Baltiyskogo yantarya]" type="journal article">Khnzorian 1960</bibRefCitation>
|
||
). While the other known species
|
||
<taxonomicName id="4C754D5FFF8FFFD2FD701227FBA3FDDD" authority="Poinar & Fanti, 2016" authorityName="Poinar & Fanti" authorityYear="2016" box="[712,1141,583,608]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="sucinobaltica">
|
||
<emphasis id="B901EACEFF8FFFD2FD701227FCA9FDDD" box="[712,895,583,608]" italics="true" pageId="3" pageNumber="516">R. sucinobaltica</emphasis>
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FC3E1229FBA3FDDD" box="[902,1141,583,608]" pageId="3" pageNumber="516" refId="ref6966" refString="Poinar, G. O. Jr. & Fanti, F. (2016) New fossil soldier beetles (Coleoptera: Cantharidae) in Burmese, Baltic and Dominican amber. Palaeodiversity, 9, 1 - 17. https: // doi. org / 10.18476 / pale. v 9. a 1" type="journal article">Poinar & Fanti, 2016</bibRefCitation>
|
||
</taxonomicName>
|
||
has similar dimensions (a body length of
|
||
<quantity id="4C8D9B39FF8FFFD2FEFB1202FE4BFD39" box="[323,413,620,645]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="516" unit="mm" value="3.5">3.5 mm</quantity>
|
||
), it possesses a different pronotum that is more curved at the sides and less bordered at the posterior margin (
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FED912E1FD84FD15" box="[353,594,655,680]" pageId="3" pageNumber="516" refId="ref6966" refString="Poinar, G. O. Jr. & Fanti, F. (2016) New fossil soldier beetles (Coleoptera: Cantharidae) in Burmese, Baltic and Dominican amber. Palaeodiversity, 9, 1 - 17. https: // doi. org / 10.18476 / pale. v 9. a 1" type="journal article">Poinar & Fanti 2016</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C754D5FFF8FFFD2FDDD12E1FC98FD15" authority="Fanti & Pankowski, 2018" authorityName="Fanti & Pankowski" authorityYear="2018" box="[613,846,655,680]" class="Insecta" family="Cantharidae" genus="Rhagonycha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="516" phylum="Arthropoda" rank="species" species="maryae" status="sp. nov.">
|
||
<emphasis id="B901EACEFF8FFFD2FDDD12E1FC98FD15" box="[613,846,655,680]" italics="true" pageId="3" pageNumber="516">Rhagonycha maryae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23257B5FF8FFFD2FCEE12E1FC67FD15" box="[854,945,655,680]" pageId="3" pageNumber="516" rank="species">
|
||
<emphasis id="B901EACEFF8FFFD2FCEE12E1FC67FD15" bold="true" box="[854,945,655,680]" pageId="3" pageNumber="516">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
also differs from other inclusions in Baltic amber known at a generic level as well as specimens known from mold impression and compression fossils in various rocky sediments (
|
||
<bibRefCitation id="EFE44B2DFF8FFFD2FE0E12B9FD97FD4D" box="[438,577,727,752]" pageId="3" pageNumber="516" refId="ref5255" refString="Fanti, F. (2017 a) Catalogo Cantharidae fossili del mondo. Fossils & Minerals Review, 2, 1 - 18 [abbreviated Italian version. Available: 12 March 2017 / World catalog of fossil Cantharidae. Fossils & Minerals Review, 2 (Special Issue), 1 - 52, extended English version, available: 25 May 2017]." type="journal article">Fanti 2017a</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36F6557FF8FFFD2FF7F1292FF12FC85" pageId="3" pageNumber="516" type="discussion">
|
||
<paragraph id="8BCA36DCFF8FFFD2FF7F1292FF12FC85" blockId="3.[151,1437,151,824]" pageId="3" pageNumber="516">
|
||
<emphasis id="B901EACEFF8FFFD2FF7F1292FEEDFCA8" bold="true" box="[199,315,764,789]" pageId="3" pageNumber="516">Remarks.</emphasis>
|
||
The amber piece measures
|
||
<date id="FFCB101CFF8FFFD2FDC21292FD68FCA9" box="[634,702,764,788]" pageId="3" pageNumber="516" value="1919-10-19">19x19</date>
|
||
x
|
||
<quantity id="4C8D9B39FF8FFFD2FD751292FCC6FCA9" box="[717,784,764,788]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="516" unit="mm" value="4.0">4 mm</quantity>
|
||
and weighs
|
||
<quantity id="4C8D9B39FF8FFFD2FC191292FBC3FCA8" box="[929,1045,764,789]" metricMagnitude="-4" metricUnit="kg" metricValue="8.0" pageId="3" pageNumber="516" unit="g" value="0.8">0.8 grams</quantity>
|
||
. It has a round shape and is rather flat.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |