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<document id="735537C476324136590AAE1F7B3C7C99" ID-DOI="10.1111/j.1096-3642.2008.00503.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630455791949" checkinUser="felipe" docAuthor="Rotheray, Graham E., Marcos-García, Maria-Angeles, Hancock, Geoff, Pérez-Bañón, Celeste &amp; Maier, Chris T." docDate="2009" docId="03DC8791FFDDFFD0FCBAF97B16C5C6F2" docLanguage="en" docName="j.1096-3642.2008.00503.x.pdf" docOrigin="Zoological Journal of the Linnean Society 156 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00503.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Copestylum bequaerti Curran 1930" docType="treatment" docVersion="3" lastPageNumber="708" masterDocId="FFE5FFE9FFD7FFDBFFC8FF8E174BC358" masterDocTitle="Neotropical Copestylum (Diptera, Syrphidae) breeding in Agavaceae and Cactaceae including seven new species" masterLastPageNumber="749" masterPageNumber="697" pageNumber="707" updateTime="1699277160858" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="D9C16D8BB8066DA4C603BC459A4A0A03">Neotropical Copestylum (Diptera, Syrphidae) breeding in Agavaceae and Cactaceae including seven new species</mods:title>
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<mods:namePart id="D2A7FC20EE13146DDD7A0DF829739B9E">Rotheray, Graham E.</mods:namePart>
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<mods:namePart id="D355D3E188F508B267ABE7544F8A5BCE">Marcos-García, Maria-Angeles</mods:namePart>
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<mods:namePart id="1E4E6D3539C82253260A5D107ABFC47B">Hancock, Geoff</mods:namePart>
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<mods:namePart id="2679B053FBF1971AB15B7AF100228A10">Pérez-Bañón, Celeste</mods:namePart>
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<mods:namePart id="B764655A2C31F95DA4145B7C51D19AC6">Maier, Chris T.</mods:namePart>
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<mods:date id="D03E255627FC0CCF0403A627C2C399A0">2009</mods:date>
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<treatment id="03DC8791FFDDFFD0FCBAF97B16C5C6F2" LSID="urn:lsid:plazi:treatment:03DC8791FFDDFFD0FCBAF97B16C5C6F2" httpUri="http://treatment.plazi.org/id/03DC8791FFDDFFD0FCBAF97B16C5C6F2" lastPageId="11" lastPageNumber="708" pageId="10" pageNumber="707">
<subSubSection id="C36F650CFFDDFFD0FCBAF97B15AFC05A" lastPageId="11" lastPageNumber="708" pageId="10" pageNumber="707" type="nomenclature">
<paragraph id="8BCA3687FFDDFFD1FCBAF97B120AC452" blockId="10.[882,1345,1781,1802]" box="[882,1345,1781,1802]" pageId="10" pageNumber="707">
<heading id="D08281EBFFDDFFD1FCBAF97B120AC452" box="[882,1345,1781,1802]" centered="true" fontSize="9" level="2" pageId="10" pageNumber="707" reason="2">
<taxonomicName id="4C754D04FFDDFFD1FCBAF97B120AC452" ID-CoL="8BKCY" authority="CURRAN, 1930" authorityName="Curran" authorityYear="1930" box="[882,1345,1781,1802]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="707" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="B901EA95FFDDFFD1FCBAF97B13DBC451" box="[882,1168,1781,1802]" italics="true" pageId="10" pageNumber="707">COPESTYLUM BEQUAERTI</emphasis>
CURRAN, 1930
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCA3687FFDDFFD0FCEEF89215AFC05A" blockId="10.[806,1421,1820,1903]" lastBlockId="11.[164,779,198,771]" lastPageId="11" lastPageNumber="708" pageId="10" pageNumber="707">
<emphasis id="B901EA95FFDDFFD1FCEEF8921379C469" box="[806,1074,1820,1841]" italics="true" pageId="10" pageNumber="707">Larva and puparium:</emphasis>
Anterior fold with a wide spicule band. Anterior spiracles present, more than ten openings arranged in a crescent at the fan-like apex and the whole structure longer than surrounding spicules. Crochets of metathoracic prolegs forming a U-shaped band. Thoracic vestiture comprising fine setae, flattened against the integument. Anterodorsal and lateral margins of mesothorax and metathorax with spicules. Anteroventral margin of metathorax with a paired group of numerous spicules of varying sizes becoming shorter posteriorly. Abdominal vestiture comprising short and long setae, long setae confined to middle transverse folds on each segment. Vestiture between prolegs and on final section of anal segment, very short and inconspicuous. Abdominal prolegs with three, transverse rows of black crochets. Sensilla 911 on the ventral surface of the anal segment on short fleshy projections. Posterior breathing tube with TR deep and conspicuous. AP dark brown, faintly coriaceous, not strongly shining and 0.7¥ length of TR. Pupal spiracles long and spiracles almost reaching base, about 2.6¥ longer than AP.
</paragraph>
</subSubSection>
<subSubSection id="C36F650CFFDCFFD0FF6CFCA016C5C6F2" pageId="11" pageNumber="708" type="description">
<paragraph id="8BCA3687FFDCFFD0FF6CFCA0166CC7EA" blockId="11.[164,779,813,1202]" pageId="11" pageNumber="708">
<emphasis id="B901EA95FFDCFFD0FF6CFCA0162CC01B" box="[164,359,814,835]" italics="true" pageId="11" pageNumber="708">Taxonomic notes:</emphasis>
The early stages of
<taxonomicName id="4C754D04FFDCFFD0FD9AFCA31596C01A" authorityName="Curran" authorityYear="1930" box="[594,733,813,834]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="B901EA95FFDCFFD0FD9AFCA31596C01A" box="[594,733,813,834]" italics="true" pageId="11" pageNumber="708">C. bequaerti</emphasis>
</taxonomicName>
are very distinctive because of the large anterior spiracles with 10+ openings, spicules on the anterodorsal margin of the metathorax, long and short abdominal vestiture, and pupal spiracles with openings almost reaching the base. No other species considered here has such a combination of characters. On adult features such as the brush-like form of the arista,
<taxonomicName id="4C754D04FFDCFFD0FD3AFB8D1645C76F" authorityName="Curran" authorityYear="1930" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="B901EA95FFDCFFD0FD3AFB8D1645C76F" italics="true" pageId="11" pageNumber="708">C. bequaerti</emphasis>
</taxonomicName>
is part of the Marginatum group. On larval characters
<taxonomicName id="4C754D04FFDCFFD0FEEDFBCF16FAC70D" authorityName="Curran" authorityYear="1930" box="[293,433,1088,1110]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="bequaerti">
<emphasis id="B901EA95FFDCFFD0FEEDFBCF16FAC70D" box="[293,433,1088,1110]" italics="true" pageId="11" pageNumber="708">C. bequaerti</emphasis>
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is similar to the Satur group species in size, presence of anterior spiracles and sensilla 911 of the anal segment on short, fleshy projections.
</paragraph>
<paragraph id="8BCA3687FFDCFFD0FF6CFB5216C5C6F2" blockId="11.[164,779,1244,1450]" pageId="11" pageNumber="708">
<emphasis id="B901EA95FFDCFFD0FF6CFB5216CEC7A9" box="[164,389,1244,1265]" italics="true" pageId="11" pageNumber="708">Material examined:</emphasis>
<specimenCount id="9D73FD0EFFDCFFD0FE59FB53168BC7A9" box="[401,448,1245,1266]" pageId="11" pageNumber="708" type="female">2 ♀</specimenCount>
, two puparia
<collectingCountry id="F3627617FFDCFFD0FDA4FB531441C7AA" box="[620,778,1245,1266]" name="Costa Rica" pageId="11" pageNumber="708">COSTA RICA</collectingCountry>
<collectingRegion id="49B1F865FFDCFFD0FF6CFB751659C649" box="[164,274,1275,1297]" country="Costa Rica" name="San Jose" pageId="11" pageNumber="708">San José</collectingRegion>
, Santa Cruz, Central, Ext. Barra Honda
<date id="FFCB1047FFDCFFD0FF6CFA941648C677" box="[164,259,1306,1327]" pageId="11" pageNumber="708" value="1999-10-09">9.x.1999</date>
Y. Cárdenas ex decaying cactus (INBio);
<specimenCount id="9D73FD0EFFDCFFD0FD1EFA94144FC676" box="[726,772,1306,1327]" pageId="11" pageNumber="708" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D73FD0EFFDCFFD0FF6CFAB6179BC615" box="[164,208,1336,1357]" pageId="11" pageNumber="708" type="female">1 ♀</specimenCount>
, two puparia,
<specimenCount id="9D73FD0EFFDCFFD0FEB3FAB61542C616" box="[379,521,1336,1358]" pageId="11" pageNumber="708" type="larva">three larvae</specimenCount>
<collectingCountry id="F3627617FFDCFFD0FDD9FAB615E5C615" box="[529,686,1336,1357]" name="Costa Rica" pageId="11" pageNumber="708">COSTA RICA</collectingCountry>
<collectingRegion id="49B1F865FFDCFFD0FD7EFAB617ABC634" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="708">Guanacaste</collectingRegion>
, Santa Rosa, Playa Naranjo
<date id="FFCB1047FFDCFFD0FDE7FAD915EFC635" box="[559,676,1367,1389]" pageId="11" pageNumber="708" value="2000-06-28">28.vi.2000</date>
DBC, E. G. H., G. E. R. ex decaying
<taxonomicName id="4C754D04FFDCFFD0FE30FAF81796C6F1" authorityName="Britton and Rose" authorityYear="1909" baseAuthorityName="F. A. C. Weber" class="Magnoliopsida" family="Cactaceae" genus="Hylocereus" higherTaxonomySource="GBIF" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="708" phylum="Tracheophyta" rank="species" species="costaricensis">
<emphasis id="B901EA95FFDCFFD0FE30FAF815E6C6D3" box="[504,685,1398,1419]" italics="true" pageId="11" pageNumber="708">H. costaricensis</emphasis>
(Cactaceae)
</taxonomicName>
(INBio, NMS).
</paragraph>
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</treatment>
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