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<document id="F391BB098B74FF2B01A75B3838C92CFA" ID-DOI="10.1111/j.1096-3642.2008.00503.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5443474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630455791949" checkinUser="felipe" docAuthor="Rotheray, Graham E., Marcos-García, Maria-Angeles, Hancock, Geoff, Pérez-Bañón, Celeste &amp; Maier, Chris T." docDate="2009" docId="03DC8791FFDCFFD7FCB8FD6F1584C288" docLanguage="en" docName="j.1096-3642.2008.00503.x.pdf" docOrigin="Zoological Journal of the Linnean Society 156 (4)" docSource="https://academic.oup.com/zoolinnean/article-lookup/doi/10.1111/j.1096-3642.2008.00503.x" docStyle="DocumentStyle:0DD8C314D74634CE09062A86991413F8.2:ZoolJLinnSoc.2002-2009.journal_article" docStyleId="0DD8C314D74634CE09062A86991413F8" docStyleName="ZoolJLinnSoc.2002-2009.journal_article" docStyleVersion="2" docTitle="Copestylum limbipenne Williston 1887" docType="treatment" docVersion="3" lastPageNumber="708" masterDocId="FFE5FFE9FFD7FFDBFFC8FF8E174BC358" masterDocTitle="Neotropical Copestylum (Diptera, Syrphidae) breeding in Agavaceae and Cactaceae including seven new species" masterLastPageNumber="749" masterPageNumber="697" pageNumber="708" updateTime="1699277160858" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="DC11E1C1167F77EC48E57337D26A0A68">Neotropical Copestylum (Diptera, Syrphidae) breeding in Agavaceae and Cactaceae including seven new species</mods:title>
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<mods:namePart id="5FA017257EF4A277D05444B1F077EAAB">Rotheray, Graham E.</mods:namePart>
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<mods:namePart id="2A4F56F910A19AA45299919EEAB7C37F">Marcos-García, Maria-Angeles</mods:namePart>
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<mods:namePart id="F08A51303F58D920680FA25B337B00E0">Hancock, Geoff</mods:namePart>
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<mods:namePart id="6870314789C138270BD3F50ECEE48B3F">Pérez-Bañón, Celeste</mods:namePart>
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<mods:namePart id="694CD49779B0C274F1653FA4C65E5B14">Maier, Chris T.</mods:namePart>
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<mods:date id="F9363E2E74D22B03A34BF572CB423FF0">2009</mods:date>
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<treatment id="03DC8791FFDCFFD7FCB8FD6F1584C288" LSID="urn:lsid:plazi:treatment:03DC8791FFDCFFD7FCB8FD6F1584C288" httpUri="http://treatment.plazi.org/id/03DC8791FFDCFFD7FCB8FD6F1584C288" lastPageId="12" lastPageNumber="708" pageId="11" pageNumber="708">
<subSubSection id="C36F650CFFDCFFD0FCB8FD6F1256C08C" pageId="11" pageNumber="708" type="nomenclature">
<paragraph id="8BCA3687FFDCFFD0FCB8FD6F1220C1AE" blockId="11.[880,1387,737,758]" box="[880,1387,737,758]" pageId="11" pageNumber="708">
<heading id="D08281EBFFDCFFD0FCB8FD6F1220C1AE" box="[880,1387,737,758]" centered="true" fontSize="9" level="2" pageId="11" pageNumber="708" reason="2">
<taxonomicName id="4C754D04FFDCFFD0FCB8FD6F1220C1AE" ID-CoL="8BKD9" authority="WILLISTON, 1887" authorityName="Williston" authorityYear="1887" box="[880,1387,737,758]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="limbipenne">
<emphasis id="B901EA95FFDCFFD0FCB8FD6F13D1C1AD" box="[880,1178,737,758]" italics="true" pageId="11" pageNumber="708">COPESTYLUM LIMBIPENNE</emphasis>
WILLISTON, 1887
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCA3687FFDCFFD0FCF2FC861256C08C" blockId="11.[826,1441,775,981]" pageId="11" pageNumber="708">
<emphasis id="B901EA95FFDCFFD0FCF2FC861375C045" box="[826,1086,775,797]" italics="true" pageId="11" pageNumber="708">Larva and puparium:</emphasis>
Anterior fold with a narrow spicule band. Crochets of metathoracic prolegs forming a U-shaped band. Metathorax with lateral folds bearing broad setae at sensilla 4 and 5. Anal lobe with spicules. Posterior breathing tube with AP 1.5¥ length of TR, smooth, shining and tapering. Pupal spiracles about 0.94¥ length of AP.
</paragraph>
</subSubSection>
<subSubSection id="C36F650CFFDCFFD7FCF2FC711584C288" lastPageId="12" lastPageNumber="709" pageId="11" pageNumber="708" type="description">
<paragraph id="8BCA3687FFDCFFD0FCF2FC711360C650" blockId="11.[826,1441,1022,1289]" pageId="11" pageNumber="708">
<emphasis id="B901EA95FFDCFFD0FCF2FC71127BC74B" box="[826,1328,1022,1044]" italics="true" pageId="11" pageNumber="708">
Taxonomic notes:
<taxonomicName id="4C754D04FFDCFFD0FBD3FC70127BC74B" authorityName="Williston" authorityYear="1887" box="[1051,1328,1022,1043]" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="limbipenne">Copestylum limbipenne</taxonomicName>
</emphasis>
is most similar to other members of the Marginatum group in having the AP longer than the width of the TR. It is distinguished from all these species except
<taxonomicName id="4C754D04FFDCFFD0FAE5FBD41423C7D6" authority="Rotheray &amp; Marcos-García &amp; Hancock &amp; Pérez-Bañón &amp; Maier, 2009" authorityName="Rotheray &amp; Marcos-García &amp; Hancock &amp; Pérez-Bañón &amp; Maier" authorityYear="2009" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="truncatum" status="sp. nov.">
<emphasis id="B901EA95FFDCFFD0FAE5FBD41423C7D6" italics="true" pageId="11" pageNumber="708">C. truncatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EEFFDCFFD0FCBBFBF71487C7D6" box="[883,972,1145,1166]" pageId="11" pageNumber="708" rank="species">sp. nov.</taxonomicNameLabel>
by the narrow band of spicules on the anterior fold and U-shaped arrangement of crochets on the mesothoracic prolegs. It is separated from
<taxonomicName id="4C754D04FFDCFFD0FA40FB3814F9C7B2" authorityName="Rotheray &amp; Marcos-García &amp; Hancock &amp; Pérez-Bañón &amp; Maier" authorityYear="2009" class="Insecta" family="Syrphidae" genus="Copestylum" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="708" phylum="Arthropoda" rank="species" species="truncatum">
<emphasis id="B901EA95FFDCFFD0FA40FB3814F9C7B2" italics="true" pageId="11" pageNumber="708">C. truncatum</emphasis>
</taxonomicName>
by the tapering AP, which is longer than the width of the TR.
</paragraph>
<paragraph id="8BCA3687FFDCFFD7FCF2FABC1584C288" blockId="11.[826,1441,1330,1903]" lastBlockId="12.[144,759,198,465]" lastPageId="12" lastPageNumber="709" pageId="11" pageNumber="708">
<emphasis id="B901EA95FFDCFFD0FCF2FABC124FC61F" box="[826,1284,1330,1351]" italics="true" pageId="11" pageNumber="708">Material examined (CIBIO, NMS):</emphasis>
<specimenCount id="9D73FD0EFFDCFFD0FAE9FABC12EAC610" box="[1313,1441,1330,1352]" pageId="11" pageNumber="708" type="larva">one larva</specimenCount>
<collectingCountry id="F3627617FFDCFFD0FCF2FADF14EDC63E" box="[826,934,1361,1382]" name="Mexico" pageId="11" pageNumber="708">MEXICO</collectingCountry>
, Tehuacán,
<collectingRegion id="49B1F865FFDCFFD0FB84FADF13D0C63F" box="[1100,1179,1361,1383]" country="Mexico" name="Puebla" pageId="11" pageNumber="708">Puebla</collectingRegion>
Zapotitlán, Salinas, Jardín Botánico
<date id="FFCB1047FFDCFFD0FC34FAE113DFC6DD" box="[1020,1172,1391,1413]" pageId="11" pageNumber="708" value="1999-07-06" valueMax="1999-07-10" valueMin="1999-07-06">610.vii.1999</date>
E. G. H., M. A. M., G. E. R. ex decaying platyclade of unidentified
<taxonomicName id="4C754D04FFDCFFD0FA8BFA00135CC69B" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="708" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B901EA95FFDCFFD0FA8BFA0012EAC6FB" box="[1347,1441,1422,1443]" italics="true" pageId="11" pageNumber="708">Opuntia</emphasis>
species (Cactaceae)
</taxonomicName>
;
<specimenCount id="9D73FD0EFFDCFFD0FBEAFA231316C699" box="[1058,1117,1453,1474]" pageId="11" pageNumber="708" type="male">23 ♂</specimenCount>
,
<specimenCount id="9D73FD0EFFDCFFD0FBA2FA2313EAC699" box="[1130,1185,1453,1474]" pageId="11" pageNumber="708" type="female">17 ♀</specimenCount>
, 40 puparia
<collectingCountry id="F3627617FFDCFFD0FAF9FA2312D6C69A" box="[1329,1437,1453,1474]" name="Mexico" pageId="11" pageNumber="708">MEXICO</collectingCountry>
,
<collectingRegion id="49B1F865FFDCFFD0FCF2FA4514DAC6B8" box="[826,913,1483,1504]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="708">Oaxaca</collectingRegion>
, Santiago Dominguillo
<date id="FFCB1047FFDCFFD0FB63FA451263C6B9" box="[1195,1320,1483,1505]" pageId="11" pageNumber="708" value="1999-07-14">14.vii.1999</date>
E. G. H., M. A. M., G. E. R. ex decaying stem of fallen
<taxonomicName id="4C754D04FFDCFFD0FA42FA641365C546" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="708" phylum="Tracheophyta" rank="species" species="stellatus">
<emphasis id="B901EA95FFDCFFD0FA42FA6414D1C545" italics="true" pageId="11" pageNumber="708">S. stellatus</emphasis>
(Cactaceae)
</taxonomicName>
;
<specimenCount id="9D73FD0EFFDCFFD0FB88F986133FC545" box="[1088,1140,1544,1565]" pageId="11" pageNumber="708" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D73FD0EFFDCFFD0FB41F98613F2C545" box="[1161,1209,1544,1565]" pageId="11" pageNumber="708" type="female">2 ♀</specimenCount>
, four puparia,
<specimenCount id="9D73FD0EFFDCFFD0FAB0F98714C9C565" pageId="11" pageNumber="708" type="larva">two larvae</specimenCount>
<collectingCountry id="F3627617FFDCFFD0FC43F9A914BCC564" box="[907,1015,1575,1596]" name="Mexico" pageId="11" pageNumber="708">MEXICO</collectingCountry>
,
<collectingRegion id="49B1F865FFDCFFD0FBCCF9A91310C564" box="[1028,1115,1575,1596]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="708">Oaxaca</collectingRegion>
, Valerio Trujano
<date id="FFCB1047FFDCFFD0FAECF9A912EAC565" box="[1316,1441,1575,1597]" pageId="11" pageNumber="708" value="1999-07-14">14.vii.1999</date>
E. G. H., M. A. M., G. E. R. ex decaying stem of
<taxonomicName id="4C754D04FFDCFFD0FCF2F9EB1270C522" authority="(F. A. C. Weber) Rose 1906" authorityName="Rose" authorityYear="1906" baseAuthorityName="F. A. C. Weber" box="[826,1339,1636,1658]" class="Magnoliopsida" family="Cactaceae" genus="Escontria" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="708" phylum="Tracheophyta" rank="species" species="chiotilla">
<emphasis id="B901EA95FFDCFFD0FCF2F9EB1346C521" box="[826,1037,1636,1658]" italics="true" pageId="11" pageNumber="708">Escontria chiotilla</emphasis>
(F.A.C. Weber) Rose 1906
</taxonomicName>
,
<specimenCount id="9D73FD0EFFDCFFD0FA80F9EA123CC521" box="[1352,1399,1636,1657]" pageId="11" pageNumber="708" type="male">8 ♂</specimenCount>
,
<specimenCount id="9D73FD0EFFDCFFD0FA4EF9EA1404C5CF" pageId="11" pageNumber="708" type="female">12 ♀</specimenCount>
, 20 puparia
<collectingCountry id="F3627617FFDCFFD0FC3DF90D132AC5C0" box="[1013,1121,1667,1688]" name="Mexico" pageId="11" pageNumber="708">MEXICO</collectingCountry>
,
<collectingRegion id="49B1F865FFDCFFD0FBBBF90D1381C5C0" box="[1139,1226,1667,1688]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="708">Oaxaca</collectingRegion>
, Cuicatlán, Cerro Colorado
<date id="FFCB1047FFDCFFD0FC61F92F1304C5EF" box="[937,1103,1697,1719]" pageId="11" pageNumber="708" value="1999-07-14" valueMax="1999-07-17" valueMin="1999-07-14">1417.vii.1999</date>
E. G. H., M. A. M., G. E. R. ex decaying stem of fallen
<taxonomicName id="4C754D04FFDCFFD0FB5BF94E1251C58D" box="[1171,1306,1728,1749]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="708" phylum="Tracheophyta" rank="species" species="stellatus">
<emphasis id="B901EA95FFDCFFD0FB5BF94E1251C58D" box="[1171,1306,1728,1749]" italics="true" pageId="11" pageNumber="708">S. stellatus</emphasis>
</taxonomicName>
;
<specimenCount id="9D73FD0EFFDCFFD0FAF9F94E1228C58D" box="[1329,1379,1728,1749]" pageId="11" pageNumber="708" type="female">1 ♀</specimenCount>
, one puparium,
<specimenCount id="9D73FD0EFFDCFFD0FC09F9511379C5AC" box="[961,1074,1758,1780]" pageId="11" pageNumber="708" type="larva">one larva</specimenCount>
<collectingCountry id="F3627617FFDCFFD0FB8AF95113E5C5AC" box="[1090,1198,1759,1780]" name="Mexico" pageId="11" pageNumber="708">MEXICO</collectingCountry>
,
<collectingRegion id="49B1F865FFDCFFD0FB09F9511253C5AC" box="[1217,1304,1759,1780]" country="Mexico" name="Oaxaca" pageId="11" pageNumber="708">Oaxaca</collectingRegion>
, Cuicatlán, Cerro Colorado
<date id="FFCB1047FFDCFFD0FC25F9731321C44B" box="[1005,1130,1789,1811]" pageId="11" pageNumber="708" value="1999-07-17">17.vii.1999</date>
E. G. H., M. A. M., G. E. R. ex decaying stem of fallen
<taxonomicName id="4C754D04FFDCFFD0FB52F8921257C408" authority="(F. A. C. Weber) Buxb. 1961" authorityName="Buxb." authorityYear="1961" baseAuthorityName="F. A. C. Weber" class="Magnoliopsida" family="Cactaceae" genus="Pachycereus" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="708" phylum="Tracheophyta" rank="species" species="hollianus">
<emphasis id="B901EA95FFDCFFD0FB52F89212EAC469" box="[1178,1441,1820,1841]" italics="true" pageId="11" pageNumber="708">Pachycereus hollianus</emphasis>
(F.A.C. Weber) Buxb. 1961 (Cactaceae)
</taxonomicName>
;
<specimenCount id="9D73FD0EFFDCFFD0FAF9F8B41222C417" box="[1329,1385,1850,1872]" pageId="11" pageNumber="708" type="male">6 ♂</specimenCount>
, six puparia
<collectingCountry id="F3627617FFDCFFD0FC53F8D7137EC436" box="[923,1077,1881,1902]" name="Costa Rica" pageId="11" pageNumber="708">COSTA RICA</collectingCountry>
<collectingRegion id="49B1F865FFDCFFD0FBF3F8D7138FC436" box="[1083,1220,1881,1902]" country="Costa Rica" name="Guanacaste" pageId="11" pageNumber="708">Guanacaste</collectingRegion>
, Santa Rosa, Playa Naranjo
<date id="FFCB1047FFDBFFD7FF32FF4816D3C384" box="[250,408,198,220]" pageId="12" pageNumber="709" value="2000-06-27" valueMax="2000-06-29" valueMin="2000-06-27">2729.vi.2000</date>
DBC, E. G. H., G. E. R. ex decaying platyclades of
<taxonomicName id="4C754D04FFDBFFD7FE6DFF6B15BDC3A3" authorityName="Britton and Rose" authorityYear="1919" box="[421,758,228,251]" class="Magnoliopsida" family="Cactaceae" genus="Opuntia" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="709" phylum="Tracheophyta" rank="species" species="guatemalensis">
<emphasis id="B901EA95FFDBFFD7FE6DFF6B1521C3A1" box="[421,618,228,250]" italics="true" pageId="12" pageNumber="709">O. guatemalensis</emphasis>
(Cactaceae)
</taxonomicName>
(INBio);
<specimenCount id="9D73FD0EFFDBFFD7FF3FFE8D1662C240" box="[247,297,259,280]" pageId="12" pageNumber="709" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D73FD0EFFDBFFD7FEF4FE8D1621C240" box="[316,362,259,280]" pageId="12" pageNumber="709" type="female">3 ♀</specimenCount>
, five puparia
<collectingCountry id="F3627617FFDBFFD7FDDEFE8D15C9C240" box="[534,642,259,280]" name="Mexico" pageId="12" pageNumber="709">MEXICO</collectingCountry>
,
<collectingRegion id="49B1F865FFDBFFD7FD5AFE8D15B9C241" box="[658,754,259,281]" country="Mexico" name="Chiapas" pageId="12" pageNumber="709">Chiapas</collectingRegion>
, Mazapa de Maderos,
<quantity id="4C8D9B62FFDBFFD7FE45FEAF169EC26F" box="[397,469,289,311]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="12" pageNumber="709" unit="m" value="850.0">850 m</quantity>
,
<date id="FFCB1047FFDBFFD7FE2DFEAF15C8C26F" box="[485,643,289,311]" pageId="12" pageNumber="709" value="2004-06-22" valueMax="2004-06-25" valueMin="2004-06-22">2225.vi.2004</date>
M. A. M. and C. P. B.;
<specimenCount id="9D73FD0EFFDBFFD7FEE5FECE1615C20D" box="[301,350,320,341]" pageId="12" pageNumber="709" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D73FD0EFFDBFFD7FEA6FECE16D1C20D" box="[366,410,320,341]" pageId="12" pageNumber="709" type="female">2 ♀</specimenCount>
, three puparia
<collectingCountry id="F3627617FFDBFFD7FD9BFECE15B9C20D" box="[595,754,320,341]" name="Costa Rica" pageId="12" pageNumber="709">COSTA RICA</collectingCountry>
,
<collectingRegion id="49B1F865FFDBFFD7FF58FED11652C22C" box="[144,281,351,372]" country="Costa Rica" name="Guanacaste" pageId="12" pageNumber="709">Guanacaste</collectingRegion>
, Santa Rosa,
<date id="FFCB1047FFDBFFD7FE7CFED1150EC22D" box="[436,581,351,373]" pageId="12" pageNumber="709" value="2005-01-18" valueMax="2005-01-19" valueMin="2005-01-18">1819.i.2005</date>
C. P. B. and M. Zumbado (INBio);
<specimenCount id="9D73FD0EFFDBFFD7FEB4FEF016FDC2CA" box="[380,438,382,403]" pageId="12" pageNumber="709" type="male">2 ♂</specimenCount>
, two puparia,
<specimenCount id="9D73FD0EFFDBFFD7FD48FEF015BCC2CB" box="[640,759,381,403]" pageId="12" pageNumber="709" type="larva">21 larvae</specimenCount>
<collectingCountry id="F3627617FFDBFFD7FF58FE1217B7C2E9" box="[144,252,412,433]" name="Mexico" pageId="12" pageNumber="709">MEXICO</collectingCountry>
,
<collectingRegion id="49B1F865FFDBFFD7FEDEFE121639C2EA" box="[278,370,412,434]" country="Mexico" name="Hidalgo" pageId="12" pageNumber="709">Hidalgo</collectingRegion>
, Metztitlán
<date id="FFCB1047FFDBFFD7FDD2FE12158CC2EA" box="[538,711,412,434]" pageId="12" pageNumber="709" value="2005-08-18" valueMax="2005-08-22" valueMin="2005-08-18">1822.viii.2005</date>
ex decaying stem of fallen
<taxonomicName id="4C754D04FFDBFFD7FE60FE35151CC288" box="[424,599,443,464]" class="Magnoliopsida" family="Cactaceae" genus="Stenocereus" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="709" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="B901EA95FFDBFFD7FE60FE351566C288" box="[424,557,443,464]" italics="true" pageId="12" pageNumber="709">Stenocereus</emphasis>
sp.
</taxonomicName>
M. A. M..
</paragraph>
</subSubSection>
</treatment>
</document>