treatments-xml/data/03/DC/87/03DC87904360FFF7FEBD0F8A8FCFF81F.xml

<document id="BE9AF07F1FCB999ADCEB0AAE4E2E89D3" ID-DOI="10.5281/zenodo.5355323" ID-ISSN="2345-7600" ID-Zenodo-Dep="5355323" ID-ZooBank="A24D5EE6-A22B-4E66-8F9C-1207DF7E0C9D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630215298288" checkinUser="valdenar" docAuthor="Jaafar, Zeehan, Polgar, Gianluca &amp; Zamroni, Yuliadi" docDate="2016" docId="03DC87904360FFF7FEBD0F8A8FCFF81F" docLanguage="en" docName="64rbz278-283.pdf" docOrigin="Raffles Bulletin of Zoology 64" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Periophthalmus pusing Jaafar &amp; Polgar &amp; Zamroni 2016, sp. nov." docType="treatment" docVersion="3" lastPageNumber="283" masterDocId="FFE5FFE84361FFF2FFA8081D8E23FFEA" masterDocTitle="Description of a new species of Periophthalmus (Teleostei: Gobiidae) from the Lesser Sunda Islands" masterLastPageNumber="283" masterPageNumber="278" pageNumber="279" updateTime="1699189195016" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="F1A629680A2E3E64D9BAD3845EBBF6A3">Description of a new species of Periophthalmus (Teleostei: Gobiidae) from the Lesser Sunda Islands</mods:title>
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<mods:namePart id="1E34262FE4DF10CF4339EE69344E1504">Jaafar, Zeehan</mods:namePart>
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<mods:namePart id="0A9978330B4D9C26C020EB115481AF6A">Polgar, Gianluca</mods:namePart>
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<mods:namePart id="8B4B9973AB2FC14E7DDBE6746EBCED48">Zamroni, Yuliadi</mods:namePart>
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<mods:title id="E37A8D851EBE38E269DAF2D39270DB8D">Raffles Bulletin of Zoology</mods:title>
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<mods:part id="AD80DFF0FCBA3132537B17C01ACE0518">
<mods:date id="1AC5095A2F9DF899086B8CFCCD05B2DD">2016</mods:date>
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<mods:number id="EA8A728B0FBA7A71F4A785C838C24607">2016-08-26</mods:number>
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<subSubSection id="C36F650D4360FFF3FEBD0F8A8C5FF845" box="[277,636,1943,1967]" pageId="1" pageNumber="279" type="nomenclature">
<paragraph id="8BCA36864360FFF3FEBD0F8A8C5FF845" blockId="1.[277,636,1943,1999]" box="[277,636,1943,1967]" pageId="1" pageNumber="279">
<heading id="D08281EA4360FFF3FEBD0F8A8C5FF845" bold="true" box="[277,636,1943,1967]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="279" reason="2">
<emphasis id="B901EA944360FFF3FEBD0F8A8C5FF845" bold="true" box="[277,636,1943,1967]" pageId="1" pageNumber="279">
<taxonomicName id="4C754D054360FFF3FEBD0F8A8C3BF845" ID-CoL="77663" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[277,536,1943,1967]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="279" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944360FFF3FEBD0F8A8C3BF845" bold="true" box="[277,536,1943,1967]" italics="true" pageId="1" pageNumber="279">Periophthalmus pusing</emphasis>
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<taxonomicNameLabel id="A23257EF4360FFF3FD8B0F8A8C5FF845" box="[547,636,1943,1967]" pageId="1" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C36F650D4360FFF3FEE20FAA8C66F825" box="[330,581,1975,1999]" pageId="1" pageNumber="279" type="description">
<paragraph id="8BCA36864360FFF3FEE20FAA8C66F825" blockId="1.[277,636,1943,1999]" box="[330,581,1975,1999]" pageId="1" pageNumber="279">
(
<figureCitation id="134E2A034360FFF3FEF90FAA8FB5F825" box="[337,406,1975,1999]" captionStart="Fig" captionStartId="2.[113,148,1896,1918]" captionTargetBox="[113,1451,1507,1880]" captionTargetId="figure-485@2.[113,1451,1507,1880]" captionTargetPageId="2" captionText="Fig. 1. Holotype of Periophthalmus pusing sp. nov. MZB 23015, female, 35.3 mm SL, collected in Sumba Island, Indonesia (Photograph by: Sandra Raredon)." figureDoi="http://doi.org/10.5281/zenodo.5355325" httpUri="https://zenodo.org/record/5355325/files/figure.png" pageId="1" pageNumber="279">Figs 1</figureCitation>
, 
<figureCitation id="134E2A034360FFF3FE0E0FAA8F96F825" box="[422,437,1975,1999]" captionStart="Fig" captionStartId="3.[135,170,722,744]" captionTargetBox="[135,1473,189,707]" captionTargetId="figure-7@3.[135,1473,189,707]" captionTargetPageId="3" captionText="Fig. 2. Live specimen of Periophthalmus pusing sp. nov., approximately 35 mm SL, collected in Sumba Island, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.5355327" httpUri="https://zenodo.org/record/5355327/files/figure.png" pageId="1" pageNumber="279">2</figureCitation>
, 
<tableCitation id="C6F7033D4360FFF3FE6B0FAA8C3CF825" box="[451,543,1975,1999]" captionStart="Table 1" captionStartId="1.[135,190,189,211]" captionText="Table 1. Measurements and counts of the holotype of Periophthalmus pusing sp. nov. (MZB 23015). Measurements are presented as percentages of the standard length (% SL) and head length (% HL)." pageId="1" pageNumber="279">Tables 1</tableCitation>
, 
<tableCitation id="C6F7033D4360FFF3FD870FAA8C1EF825" box="[559,573,1975,1999]" captionStart="Table 2" captionStartId="2.[113,168,189,211]" captionText="Table 2. Frequency table showing variation in fin element and scale counts in Periophthalmus pusing sp. nov. (n=15) and P. gracilis (n=27, *=presence of damaged specimens so that total sample size is not 27)." pageId="1" pageNumber="279">2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36F650D4360FFF3FF200FEA8A8EF985" pageId="1" pageNumber="279" type="materials_examined">
<paragraph id="8BCA36864360FFF3FF200FEA8D24F7C5" blockId="1.[136,775,2039,2095]" pageId="1" pageNumber="279">
<emphasis id="B901EA944360FFF3FF200FEA8FA5F7E5" bold="true" box="[136,390,2039,2063]" pageId="1" pageNumber="279">Materials examined.</emphasis>
<materialsCitation id="3B1D3CDB4360FFF3FE390FEA8D24F7C5" accessionNumber="MZB 23015" collectionCode="MZB" country="Indonesia" county="Kawangu" location="Sumba" municipality="Waingapu" pageId="1" pageNumber="279" specimenCount="1" typeStatus="holotype">
<typeStatus id="54CE88244360FFF3FE390FEA8C23F7E5" box="[401,512,2039,2063]" pageId="1" pageNumber="279" type="holotype">Holotype</typeStatus>
: 
<collectionCode id="ED64AE434360FFF3FDB80FEA8C6FF7E4" box="[528,588,2039,2062]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="1" pageNumber="279" type="Museum">MZB</collectionCode>
23015, female, 
<quantity id="4C8D9B634360FFF3FF21000A8ED3F7C4" box="[137,240,2071,2095]" metricMagnitude="-2" metricUnit="m" metricValue="3.53" pageId="1" pageNumber="279" unit="mm" value="35.3">35.3 mm</quantity>
SL, 
<collectingCountry id="F36276164360FFF3FE86000A8FBCF7C5" box="[302,415,2071,2095]" name="Indonesia" pageId="1" pageNumber="279">Indonesia</collectingCountry>
, 
<location id="8EAA605D4360FFF3FE07000A8FDCF7C5" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4360FFF3FE07000A8FDCF7C5" box="[431,511,2071,2095]" country="Indonesia" county="Kawangu" municipality="Waingapu" name="Sumba" pageId="1" pageNumber="279">Sumba</location>
, 
<collectingMunicipality id="6BAEACFC4360FFF3FDA7000A8CA6F7C5" box="[527,645,2071,2095]" pageId="1" pageNumber="279">Waingapu</collectingMunicipality>
, 
<collectingCounty id="62AB4E0A4360FFF3FD3D000A8D21F7C5" box="[661,770,2071,2095]" pageId="1" pageNumber="279">Kawangu</collectingCounty>
,
</materialsCitation>
</paragraph>
<paragraph id="8BCA36864360FFF3FCEC0D4A8A8EF985" blockId="1.[835,1476,1367,1647]" pageId="1" pageNumber="279">
<materialsCitation id="3B1D3CDB4360FFF3FCEC0D4A8DE4FA25" accessionNumber="MZB 23015" collectingDate="2012-11-10" collectionCode="MZB" collectorName="Jaafar Z. &amp; Zamroni Y." country="Indonesia" county="Kawangu" location="Sumba" municipality="Waingapu" pageId="1" pageNumber="279" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
collected by 
<collectorName id="268053504360FFF3FC660D4A8A0FFA84" box="[974,1068,1367,1391]" pageId="1" pageNumber="279">Jaafar Z.</collectorName>
and 
<collectorName id="268053504360FFF3FBC90D4A8AC3FA84" box="[1121,1248,1367,1391]" pageId="1" pageNumber="279">Zamroni Y.</collectorName>
, 
<date id="FFCB10464360FFF3FB450D4A8BE3FA85" box="[1261,1472,1367,1391]" pageId="1" pageNumber="279" value="2012-11-10">
<collectingDate id="EF8FE9AE4360FFF3FB450D4A8BE3FA85" box="[1261,1472,1367,1391]" pageId="1" pageNumber="279" value="2012-11-10">10 November 2012</collectingDate>
</date>
. 
<typeStatus id="54CE88244360FFF3FCEC0D6A8D8FFA65" box="[836,940,1399,1423]" pageId="1" pageNumber="279" type="paratype">Paratypes</typeStatus>
(n=14): AMS I.46780-001, male (
<quantity id="4C8D9B634360FFF3FA850D6A8BB3FA64" box="[1325,1424,1399,1423]" metricMagnitude="-2" metricUnit="m" metricValue="3.28" pageId="1" pageNumber="279" unit="mm" value="32.8">32.8 mm</quantity>
SL) and 
<specimenCount id="9D73FD0F4360FFF3FCDD0D8A8DC1FA45" box="[885,994,1431,1455]" pageId="1" pageNumber="279" type="female">2 females</specimenCount>
(
<quantity id="4C8D9B634360FFF3FC5C0D8A8AB4FA44" box="[1012,1175,1431,1455]" metricMagnitude="-2" metricUnit="m" metricValue="3.49" metricValueMax="3.66" metricValueMin="3.32" pageId="1" pageNumber="279" unit="mm" value="34.9" valueMax="36.6" valueMin="33.2">33.2–36.6 mm</quantity>
SL), collection data same as holotype
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4360FFF3FC7E0DAA8B9CFA05" accessionNumber="MZB 23015" collectionCode="RMNH, PISC" country="Indonesia" county="Kawangu" location="Sumba" municipality="Waingapu" pageId="1" pageNumber="279" specimenCount="2" specimenCount-female="2" typeStatus="paratype">
<collectionCode id="ED64AE434360FFF3FC7E0DAA8A0FFA25" box="[982,1068,1463,1487]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="1" pageNumber="279" type="Museum">RMNH</collectionCode>
. 
<collectionCode id="ED64AE434360FFF3FB870DAA8A48FA25" box="[1071,1131,1463,1487]" pageId="1" pageNumber="279">PISC</collectionCode>
.38420, male (
<quantity id="4C8D9B634360FFF3FAA30DAA8B53FA24" box="[1291,1392,1463,1487]" metricMagnitude="-2" metricUnit="m" metricValue="2.9699999999999998" pageId="1" pageNumber="279" unit="mm" value="29.7">29.7 mm</quantity>
SL), 
<specimenCount id="9D73FD0F4360FFF3FA1D0DAA8DB6FA05" pageId="1" pageNumber="279" type="female">2 females</specimenCount>
(
<quantity id="4C8D9B634360FFF3FC090DCA8A1BFA04" box="[929,1080,1495,1519]" metricMagnitude="-2" metricUnit="m" metricValue="3.145" metricValueMax="3.28" metricValueMin="3.01" pageId="1" pageNumber="279" unit="mm" value="31.45" valueMax="32.8" valueMin="30.1">30.1–32.8 mm</quantity>
SL), collection data same as holotype
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4360FFF3FCEB0DEA8B3BF9C5" accessionNumber="MZB 23015" collectionCode="USNM" country="Indonesia" county="Kawangu" location="Sumba" municipality="Waingapu" pageId="1" pageNumber="279" specimenCode="USNM 432497" specimenCount="3" specimenCount-female="3" typeStatus="paratype">
<specimenCode id="DBD39EFD4360FFF3FCEB0DEA8DCFF9E5" box="[835,1004,1527,1551]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="279" type="Museum">USNM 432497</specimenCode>
, male (
<quantity id="4C8D9B634360FFF3FB930DEA8ABEF9E4" box="[1083,1181,1527,1551]" metricMagnitude="-2" metricUnit="m" metricValue="2.96" pageId="1" pageNumber="279" unit="mm" value="29.6">29.6 mm</quantity>
SL), 
<specimenCount id="9D73FD0F4360FFF3FB750DEA8B65F9E5" box="[1245,1350,1527,1551]" pageId="1" pageNumber="279" type="female">3 females</specimenCount>
(
<quantity id="4C8D9B634360FFF3FAFD0DEA8D4FF9C4" metricMagnitude="-2" metricUnit="m" metricValue="3.31" metricValueMax="3.66" metricValueMin="2.96" pageId="1" pageNumber="279" unit="mm" value="33.1" valueMax="36.6" valueMin="29.6">29.6–36.6 mm</quantity>
SL), collection data same as holotype
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4360FFF3FA8D0E0A8A89F985" accessionNumber="ZRC 54416" collectionCode="ZRC" country="Indonesia" county="Kawangu" location="Sumba" municipality="Waingapu" pageId="1" pageNumber="279" specimenCount="4" specimenCount-female="2" specimenCount-male="2" typeStatus="paratype">
<collectionCode id="ED64AE434360FFF3FA8D0E0A8B7AF9C5" box="[1317,1369,1559,1583]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="279">ZRC</collectionCode>
54416, 
<specimenCount id="9D73FD0F4360FFF3FA1D0E0A8DA2F9A5" pageId="1" pageNumber="279" type="male">2 males</specimenCount>
(33.0– 
<quantity id="4C8D9B634360FFF3FC670E2A8A13F9A4" box="[975,1072,1591,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.6799999999999997" pageId="1" pageNumber="279" unit="mm" value="36.8">36.8 mm</quantity>
SL), 
<specimenCount id="9D73FD0F4360FFF3FBDA0E2A8AF8F9A5" box="[1138,1243,1591,1615]" pageId="1" pageNumber="279" type="female">2 females</specimenCount>
(
<quantity id="4C8D9B634360FFF3FB430E2A8BA9F9A4" box="[1259,1418,1591,1615]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" metricValueMax="3.72" metricValueMin="3.38" pageId="1" pageNumber="279" unit="mm" value="35.5" valueMax="37.2" valueMin="33.8">33.8–37.2 mm</quantity>
SL), collection data same as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F650D4360FFF3FCEB0E8A8B89F825" pageId="1" pageNumber="279" type="diagnosis">
<paragraph id="8BCA36864360FFF3FCEB0E8A8B89F825" blockId="1.[835,1474,1687,1999]" pageId="1" pageNumber="279">
<emphasis id="B901EA944360FFF3FCEB0E8A8DE2F945" bold="true" box="[835,961,1687,1711]" pageId="1" pageNumber="279">Diagnosis.</emphasis>
Adult 
<taxonomicName id="4C754D054360FFF3FBB20E8A8B0AF945" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[1050,1321,1687,1711]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="279" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944360FFF3FBB20E8A8B0AF945" box="[1050,1321,1687,1711]" italics="true" pageId="1" pageNumber="279">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4360FFF3FA9D0E8A8BB7F944" box="[1333,1428,1687,1710]" pageId="1" pageNumber="279" rank="species">sp. nov.</taxonomicNameLabel>
are distinguished from all congeners with the following suite of characters: pelvic frenum absent; pelvic fins separate for the entire length; a black spot posteriorly on first dorsal fin, spot usually between ninth and the ultimate spine; first dorsal fin taller than second dorsal fin; interdorsal distance less than half the length of the first dorsal-fin spine. First dorsal-fin elements XI–XV; second dorsal-fin elements I, 10–I, 12; anal-fin elements I, 10–I, 12; pectoral-fin elements 10–12; longitudinal scale rows 42–54; predorsal midline 14–19.
</paragraph>
</subSubSection>
<subSubSection id="C36F650D4360FFF1FCEB0FEA8B6DF7FD" lastPageId="3" lastPageNumber="281" pageId="1" pageNumber="279" type="description">
<paragraph id="8BCA36864360FFF1FCEB0FEA8FD1FBDD" blockId="1.[835,1474,2039,2095]" lastBlockId="3.[135,776,799,1079]" lastPageId="3" lastPageNumber="281" pageId="1" pageNumber="279">
<emphasis id="B901EA944360FFF3FCEB0FEA8DEDF7E5" bold="true" box="[835,974,2039,2063]" pageId="1" pageNumber="279">Description.</emphasis>
<tableCitation id="C6F7033D4360FFF3FC7D0FEA8A05F7E5" box="[981,1062,2039,2063]" captionStart="Table 1" captionStartId="1.[135,190,189,211]" captionText="Table 1. Measurements and counts of the holotype of Periophthalmus pusing sp. nov. (MZB 23015). Measurements are presented as percentages of the standard length (% SL) and head length (% HL)." pageId="1" pageNumber="279">Table 1</tableCitation>
shows the morphometric and meristic data for the 
<typeStatus id="54CE88244360FFF3FC6D000A8A01F7C5" box="[965,1058,2071,2095]" pageId="1" pageNumber="279" type="holotype">holotype</typeStatus>
(MZB 23015). A frequency table with meristic counts for 
<taxonomicName id="4C754D054362FFF1FEF50B028C74FCDD" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[349,599,799,823]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="281" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944362FFF1FEF50B028C74FCDD" box="[349,599,799,823]" italics="true" pageId="3" pageNumber="281">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4362FFF1FDC80B3D8C94FCDD" box="[608,695,800,823]" pageId="3" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C754D054362FFF1FD470B3D8EFBFCBD" authorityName="Eggert" authorityYear="1935" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="281" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944362FFF1FD470B3D8EFBFCBD" italics="true" pageId="3" pageNumber="281">P. gracilis</emphasis>
</taxonomicName>
is presented in 
<tableCitation id="C6F7033D4362FFF1FE200B228FFEFCBD" box="[392,477,831,855]" captionStart="Table 2" captionStartId="2.[113,168,189,211]" captionText="Table 2. Frequency table showing variation in fin element and scale counts in Periophthalmus pusing sp. nov. (n=15) and P. gracilis (n=27, *=presence of damaged specimens so that total sample size is not 27)." pageId="3" pageNumber="281">Table 2</tableCitation>
. The 
<typeStatus id="54CE88244362FFF1FDB30B228C5BFCBD" box="[539,632,831,855]" pageId="3" pageNumber="281" type="holotype">holotype</typeStatus>
was fixed in formalin and maintained in ethanol (see 
<figureCitation id="134E2A034362FFF1FDE40B428CADFC9D" box="[588,654,863,887]" captionStart="Fig" captionStartId="2.[113,148,1896,1918]" captionTargetBox="[113,1451,1507,1880]" captionTargetId="figure-485@2.[113,1451,1507,1880]" captionTargetPageId="2" captionText="Fig. 1. Holotype of Periophthalmus pusing sp. nov. MZB 23015, female, 35.3 mm SL, collected in Sumba Island, Indonesia (Photograph by: Sandra Raredon)." figureDoi="http://doi.org/10.5281/zenodo.5355325" httpUri="https://zenodo.org/record/5355325/files/figure.png" pageId="3" pageNumber="281">Fig. 1</figureCitation>
). 
<figureCitation id="134E2A034362FFF1FD000B428CC9FC9D" box="[680,746,863,887]" captionStart="Fig" captionStartId="3.[135,170,722,744]" captionTargetBox="[135,1473,189,707]" captionTargetId="figure-7@3.[135,1473,189,707]" captionTargetPageId="3" captionText="Fig. 2. Live specimen of Periophthalmus pusing sp. nov., approximately 35 mm SL, collected in Sumba Island, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.5355327" httpUri="https://zenodo.org/record/5355327/files/figure.png" pageId="3" pageNumber="281">Fig. 2</figureCitation>
is a photograph of a live specimen of 
<taxonomicName id="4C754D054362FFF1FDA60B628D2BFC7D" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[526,776,895,919]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="281" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944362FFF1FDA60B628D2BFC7D" box="[526,776,895,919]" italics="true" pageId="3" pageNumber="281">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4362FFF1FF2F0BBD8EFEFC5D" box="[135,221,928,951]" pageId="3" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
while 
<figureCitation id="134E2A034362FFF1FE800B828F4BFC5D" box="[296,360,927,951]" captionStart="Fig" captionStartId="4.[113,148,618,640]" captionTargetBox="[113,1451,189,603]" captionTargetId="figure-11@4.[113,1451,189,603]" captionTargetPageId="4" captionText="Fig. 3. Live specimen of Periophthalmus gracilis, approximately 48 mm SL, collected in Vietnam (Photograph by: Tran Dac Dinh)." figureDoi="http://doi.org/10.5281/zenodo.5355329" httpUri="https://zenodo.org/record/5355329/files/figure.png" pageId="3" pageNumber="281">Fig. 3</figureCitation>
is a photograph of a live specimen of 
<taxonomicName id="4C754D054362FFF1FF2F0BDD8ED4FC3D" authorityName="Eggert" authorityYear="1935" box="[135,247,959,983]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="281" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944362FFF1FF2F0BDD8ED4FC3D" box="[135,247,959,983]" italics="true" pageId="3" pageNumber="281">P. gracilis</emphasis>
</taxonomicName>
. The descriptions below provide details of adult specimens over 
<quantity id="4C8D9B634362FFF1FE980BC28F5AFC1D" box="[304,377,991,1015]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="281" unit="mm" value="30.0">30 mm</quantity>
SL. As with other congeners, juvenile specimens, especially those equal or less than 
<quantity id="4C8D9B634362FFF1FD210BE28CF5FBFD" box="[649,726,1023,1047]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="281" unit="mm" value="25.0">25 mm</quantity>
SL, are difficult to discern to species.
</paragraph>
<caption id="DF0A660E4363FFF0FFD908A08D4EFF1A" pageId="2" pageNumber="280" startId="2.[113,168,189,211]">
<paragraph id="8BCA36864363FFF0FFD908A08D4EFF1A" blockId="2.[113,1452,189,240]" pageId="2" pageNumber="280">
Table 2. Frequency table showing variation in fin element and scale counts in 
<taxonomicName id="4C754D054363FFF0FC2108A08A4FFF39" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[905,1132,189,211]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FC2108A08A4FFF39" box="[905,1132,189,211]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FBDD08A38AE6FF39" box="[1141,1221,190,211]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
(n=15) and 
<taxonomicName id="4C754D054363FFF0FAEC08A38B8FFF38" authorityName="Eggert" authorityYear="1935" box="[1348,1452,189,211]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FAEC08A38B70FF39" box="[1348,1363,190,211]" italics="true" pageId="2" pageNumber="280">P</emphasis>
. 
<emphasis id="B901EA944363FFF0FACA08A08B8FFF38" box="[1378,1452,189,210]" italics="true" pageId="2" pageNumber="280">gracilis</emphasis>
</taxonomicName>
(n=27, *=presence of damaged specimens so that total sample size is not 27).
</paragraph>
</caption>
<paragraph id="8BCA36864363FFF0FDE9093C8D03FA77" pageId="2" pageNumber="280">
<table id="F975C4264363000DFF28093C8BA2FA77" box="[128,1409,289,1437]" gridcols="9" gridrows="25" pageId="2" pageNumber="280">
<tr id="354534C44363000DFF28093C8BA2FEDD" box="[128,1409,289,311]" gridrow="0" pageId="2" pageNumber="280" rowspan-0="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFDA5093C8A04FEDD" box="[525,1063,289,311]" colspan="5" colspanRight="4" gridcol="1" gridrow="0" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FDE9093C8DFFFEDD" bold="true" box="[577,988,289,311]" pageId="2" pageNumber="280">VARIATION IN MERISTIC COUNTS</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF28094D8BA2FE8C" box="[128,1409,336,358]" gridrow="1" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF28094D8BA2FE8C" box="[128,1409,336,358]" colspan="9" colspanRight="8" gridcol="0" gridrow="1" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FCE2094D8A72FE8F" bold="true" box="[842,1105,336,358]" pageId="2" pageNumber="280">First Dorsal-fin Elements</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF28099D8BA2FE7F" box="[128,1409,384,405]" gridrow="2" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF28099D8F9AFE7F" box="[128,441,384,405]" gridcol="0" gridrow="2" pageId="2" pageNumber="280">Range</th>
<td id="76945DB84363000DFDA5099D8C6AFE7F" box="[525,585,384,405]" gridcol="1" gridrow="2" pageId="2" pageNumber="280">VIII</td>
<td id="76945DB84363000DFD2C099D8CE2FE7F" box="[644,705,384,405]" gridcol="2" gridrow="2" pageId="2" pageNumber="280">IX</td>
<td id="76945DB84363000DFD54099D8D12FE7F" box="[764,817,384,405]" gridcol="3" gridrow="2" pageId="2" pageNumber="280">X</td>
<td id="76945DB84363000DFCD3099D8D93FE7F" box="[891,944,384,405]" gridcol="4" gridrow="2" pageId="2" pageNumber="280">XI</td>
<td id="76945DB84363000DFC43099D8AB6FE7F" box="[1003,1173,384,405]" colspan="2" colspanRight="1" gridcol="5" gridrow="2" pageId="2" pageNumber="280">XII XIII</td>
<td id="76945DB84363000DFB4B099D8B2DFE7F" box="[1251,1294,384,405]" gridcol="7" gridrow="2" pageId="2" pageNumber="280">XIV</td>
<td id="76945DB84363000DFAF6099D8BA2FE7F" box="[1374,1409,384,405]" gridcol="8" gridrow="2" pageId="2" pageNumber="280">XV</td>
</tr>
<tr id="354534C44363000DFF2809B38BA2FE2E" box="[128,1409,430,452]" gridrow="3" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF2809B38F9AFE2E" box="[128,441,430,452]" gridcol="0" gridrow="3" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF2809B38F42FE2E" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[128,353,430,452]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FF2809B38F42FE2E" box="[128,353,430,452]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FEC309B28F9AFE2E" box="[363,441,431,452]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76945DB84363000DFDA509B38C6AFE2E" box="[525,585,430,452]" gridcol="1" gridrow="3" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFD2C09B38CE2FE2E" box="[644,705,430,452]" gridcol="2" gridrow="3" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFD5409B38D12FE2E" box="[764,817,430,452]" gridcol="3" gridrow="3" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFCD309B38D93FE2E" box="[891,944,430,452]" gridcol="4" gridrow="3" pageId="2" pageNumber="280">3</td>
<td id="76945DB84363000DFC4309B38A04FE2E" box="[1003,1063,430,452]" gridcol="5" gridrow="3" pageId="2" pageNumber="280">4</td>
<td id="76945DB84363000DFBD309B38AB6FE2E" box="[1147,1173,430,452]" gridcol="6" gridrow="3" pageId="2" pageNumber="280">4</td>
<td id="76945DB84363000DFB4B09B38B2DFE2E" box="[1251,1294,430,452]" gridcol="7" gridrow="3" pageId="2" pageNumber="280">2</td>
<td id="76945DB84363000DFAF609B38BA2FE2E" box="[1374,1409,430,452]" gridcol="8" gridrow="3" pageId="2" pageNumber="280">2</td>
</tr>
<tr id="354534C44363000DFF2809C08BA2FE19" box="[128,1409,477,499]" gridrow="4" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF2809C08F9AFE19" box="[128,441,477,499]" gridcol="0" gridrow="4" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF2809C08F49FE18" authorityName="Eggert" authorityYear="1935" box="[128,362,477,498]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FF2809C08F49FE18" box="[128,362,477,498]" italics="true" pageId="2" pageNumber="280">Periophthalmus gracilis</emphasis>
</taxonomicName>
</th>
<td id="76945DB84363000DFDA509C08C6AFE19" box="[525,585,477,499]" gridcol="1" gridrow="4" pageId="2" pageNumber="280">4</td>
<td id="76945DB84363000DFD2C09C08CE2FE19" box="[644,705,477,499]" gridcol="2" gridrow="4" pageId="2" pageNumber="280">10</td>
<td id="76945DB84363000DFD5409C08D12FE19" box="[764,817,477,499]" gridcol="3" gridrow="4" pageId="2" pageNumber="280">8</td>
<td id="76945DB84363000DFCD309C08D93FE19" box="[891,944,477,499]" gridcol="4" gridrow="4" pageId="2" pageNumber="280">4</td>
<td id="76945DB84363000DFC4309C08A04FE19" box="[1003,1063,477,499]" gridcol="5" gridrow="4" pageId="2" pageNumber="280">1</td>
<td id="76945DB84363000DFBD309C08AB6FE19" box="[1147,1173,477,499]" gridcol="6" gridrow="4" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFB4B09C08B2DFE19" box="[1251,1294,477,499]" gridcol="7" gridrow="4" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFAF609C08BA2FE19" box="[1374,1409,477,499]" gridcol="8" gridrow="4" pageId="2" pageNumber="280">–</td>
</tr>
<tr id="354534C44363000DFF280A118BA2FDC8" box="[128,1409,524,546]" gridrow="5" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF280A118BA2FDC8" box="[128,1409,524,546]" colspan="9" colspanRight="8" gridcol="0" gridrow="5" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FC960A118A7EFDCB" bold="true" box="[830,1117,524,546]" pageId="2" pageNumber="280">Second Dorsal-fin Elements</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF280A268BA2FDBA" box="[128,1409,571,592]" gridrow="6" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280A268F9AFDBA" box="[128,441,571,592]" gridcol="0" gridrow="6" pageId="2" pageNumber="280">Range</th>
<td id="76945DB84363000DFDA50A268C6AFDBA" box="[525,585,571,592]" gridcol="1" gridrow="6" pageId="2" pageNumber="280">I, 10</td>
<td id="76945DB84363000DFD2C0A268CE2FDBA" box="[644,705,571,592]" gridcol="2" gridrow="6" pageId="2" pageNumber="280">I, 11</td>
<td id="76945DB84363000DFD540A268D12FDBA" box="[764,817,571,592]" gridcol="3" gridrow="6" pageId="2" pageNumber="280">I, 12</td>
</tr>
<tr id="354534C44363000DFF280A778BA2FD95" box="[128,1409,618,639]" gridrow="7" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280A778F9AFD95" box="[128,441,618,639]" gridcol="0" gridrow="7" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280A778F42FD95" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[128,353,618,639]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FF280A778F42FD95" box="[128,353,618,639]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FEC30A778F9AFD95" box="[363,441,618,639]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76945DB84363000DFDA50A778C6AFD95" box="[525,585,618,639]" gridcol="1" gridrow="7" pageId="2" pageNumber="280">1</td>
<td id="76945DB84363000DFD2C0A778CE2FD95" box="[644,705,618,639]" gridcol="2" gridrow="7" pageId="2" pageNumber="280">12</td>
<td id="76945DB84363000DFD540A778D12FD95" box="[764,817,618,639]" gridcol="3" gridrow="7" pageId="2" pageNumber="280">2</td>
</tr>
<tr id="354534C44363000DFF280A848BA2FD44" box="[128,1409,665,686]" gridrow="8" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280A848F9AFD44" box="[128,441,665,686]" gridcol="0" gridrow="8" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280A848F49FD44" authorityName="Eggert" authorityYear="1935" box="[128,362,665,686]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FF280A848F49FD44" box="[128,362,665,686]" italics="true" pageId="2" pageNumber="280">Periophthalmus gracilis</emphasis>
</taxonomicName>
</th>
<td id="76945DB84363000DFDA50A848C6AFD44" box="[525,585,665,686]" gridcol="1" gridrow="8" pageId="2" pageNumber="280">17</td>
<td id="76945DB84363000DFD2C0A848CE2FD44" box="[644,705,665,686]" gridcol="2" gridrow="8" pageId="2" pageNumber="280">10</td>
<td id="76945DB84363000DFD540A848D12FD44" box="[764,817,665,686]" gridcol="3" gridrow="8" pageId="2" pageNumber="280">–</td>
</tr>
<tr id="354534C44363000DFF280ADA8BA2FD37" box="[128,1409,711,733]" gridrow="9" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF280ADA8BA2FD37" box="[128,1409,711,733]" colspan="9" colspanRight="8" gridcol="0" gridrow="9" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FCD80ADA8A08FD37" bold="true" box="[880,1067,711,733]" pageId="2" pageNumber="280">Anal-fin Elements</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF280AEA8BA2FCE6" box="[128,1409,759,780]" gridrow="10" pageId="2" pageNumber="280" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280AEA8F9AFCE6" box="[128,441,759,780]" gridcol="0" gridrow="10" pageId="2" pageNumber="280">Range</th>
<td id="76945DB84363000DFDA50AEA8C6AFCE6" box="[525,585,759,780]" gridcol="1" gridrow="10" pageId="2" pageNumber="280">I, 9</td>
<td id="76945DB84363000DFD2C0AEA8CE2FCE6" box="[644,705,759,780]" gridcol="2" gridrow="10" pageId="2" pageNumber="280">I, 10</td>
<td id="76945DB84363000DFD540AEA8D12FCE6" box="[764,817,759,780]" gridcol="3" gridrow="10" pageId="2" pageNumber="280">I, 11</td>
<td id="76945DB84363000DFCD30AEA8D93FCE6" box="[891,944,759,780]" gridcol="4" gridrow="10" pageId="2" pageNumber="280">I, 12</td>
</tr>
<tr id="354534C44363000DFF280B388BA2FCD1" box="[128,1409,805,827]" gridrow="11" pageId="2" pageNumber="280" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280B388F9AFCD1" box="[128,441,805,827]" gridcol="0" gridrow="11" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280B388F42FCD1" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[128,353,805,827]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FF280B388F42FCD1" box="[128,353,805,827]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FEC30B3B8F9AFCD1" box="[363,441,806,827]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76945DB84363000DFDA50B388C6AFCD1" box="[525,585,805,827]" gridcol="1" gridrow="11" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFD2C0B388CE2FCD1" box="[644,705,805,827]" gridcol="2" gridrow="11" pageId="2" pageNumber="280">1</td>
<td id="76945DB84363000DFD540B388D12FCD1" box="[764,817,805,827]" gridcol="3" gridrow="11" pageId="2" pageNumber="280">12</td>
<td id="76945DB84363000DFCD30B388D93FCD1" box="[891,944,805,827]" gridcol="4" gridrow="11" pageId="2" pageNumber="280">2</td>
</tr>
<tr id="354534C44363000DFF280B498BA2FC80" box="[128,1409,852,874]" gridrow="12" pageId="2" pageNumber="280" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280B498F9AFC80" box="[128,441,852,874]" gridcol="0" gridrow="12" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280B498F49FC83" authorityName="Eggert" authorityYear="1935" box="[128,362,852,873]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FF280B498F49FC83" box="[128,362,852,873]" italics="true" pageId="2" pageNumber="280">Periophthalmus gracilis</emphasis>
</taxonomicName>
*
</th>
<td id="76945DB84363000DFDA50B498C6AFC80" box="[525,585,852,874]" gridcol="1" gridrow="12" pageId="2" pageNumber="280">1</td>
<td id="76945DB84363000DFD2C0B498CE2FC80" box="[644,705,852,874]" gridcol="2" gridrow="12" pageId="2" pageNumber="280">12</td>
<td id="76945DB84363000DFD540B498D12FC80" box="[764,817,852,874]" gridcol="3" gridrow="12" pageId="2" pageNumber="280">6</td>
<td id="76945DB84363000DFCD30B498D93FC80" box="[891,944,852,874]" gridcol="4" gridrow="12" pageId="2" pageNumber="280">7</td>
</tr>
<tr id="354534C44363000DFF280B9E8BA2FC73" box="[128,1409,899,921]" gridrow="13" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF280B9E8BA2FC73" box="[128,1409,899,921]" colspan="9" colspanRight="8" gridcol="0" gridrow="13" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FCED0B9E8A75FC72" bold="true" box="[837,1110,899,921]" pageId="2" pageNumber="280">Left Pectoral-fin Elements</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF280BAF8BA2FC2D" box="[128,1409,946,967]" gridrow="14" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280BAF8F9AFC2D" box="[128,441,946,967]" gridcol="0" gridrow="14" pageId="2" pageNumber="280">Range</th>
<td id="76945DB84363000DFDA50BAF8C6AFC2D" box="[525,585,946,967]" gridcol="1" gridrow="14" pageId="2" pageNumber="280">10</td>
<td id="76945DB84363000DFD2C0BAF8CE2FC2D" box="[644,705,946,967]" gridcol="2" gridrow="14" pageId="2" pageNumber="280">11</td>
<td id="76945DB84363000DFD540BAF8D12FC2D" box="[764,817,946,967]" gridcol="3" gridrow="14" pageId="2" pageNumber="280">12</td>
</tr>
<tr id="354534C44363000DFF280BFC8BA2FC1D" box="[128,1409,993,1015]" gridrow="15" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280BFC8F9AFC1D" box="[128,441,993,1015]" gridcol="0" gridrow="15" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280BFC8F42FC1D" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[128,353,993,1015]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FF280BFC8F42FC1D" box="[128,353,993,1015]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FEC30BFC8F9AFC1C" box="[363,441,993,1014]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76945DB84363000DFDA50BFC8C6AFC1D" box="[525,585,993,1015]" gridcol="1" gridrow="15" pageId="2" pageNumber="280">1</td>
<td id="76945DB84363000DFD2C0BFC8CE2FC1D" box="[644,705,993,1015]" gridcol="2" gridrow="15" pageId="2" pageNumber="280">6</td>
<td id="76945DB84363000DFD540BFC8D12FC1D" box="[764,817,993,1015]" gridcol="3" gridrow="15" pageId="2" pageNumber="280">8</td>
</tr>
<tr id="354534C44363000DFF280C0D8BA2FBCF" box="[128,1409,1040,1061]" gridrow="16" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280C0D8F9AFBCF" box="[128,441,1040,1061]" gridcol="0" gridrow="16" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280C0D8F49FBCF" authorityName="Eggert" authorityYear="1935" box="[128,362,1040,1061]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FF280C0D8F49FBCF" box="[128,362,1040,1061]" italics="true" pageId="2" pageNumber="280">Periophthalmus gracilis</emphasis>
</taxonomicName>
*
</th>
<td id="76945DB84363000DFDA50C0D8C6AFBCF" box="[525,585,1040,1061]" gridcol="1" gridrow="16" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFD2C0C0D8CE2FBCF" box="[644,705,1040,1061]" gridcol="2" gridrow="16" pageId="2" pageNumber="280">7</td>
<td id="76945DB84363000DFD540C0D8D12FBCF" box="[764,817,1040,1061]" gridcol="3" gridrow="16" pageId="2" pageNumber="280">18</td>
</tr>
<tr id="354534C44363000DFF280C238BA2FBBE" box="[128,1409,1086,1108]" gridrow="17" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF280C238BA2FBBE" box="[128,1409,1086,1108]" colspan="9" colspanRight="8" gridcol="0" gridrow="17" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FCE40C238A6DFBBE" bold="true" box="[844,1102,1086,1108]" pageId="2" pageNumber="280">Longitudinal Scale Rows</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF280C708BA2FB69" box="[128,1409,1133,1155]" gridrow="18" pageId="2" pageNumber="280" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280C708F9AFB69" box="[128,441,1133,1155]" gridcol="0" gridrow="18" pageId="2" pageNumber="280">Range</th>
<td id="76945DB84363000DFDA50C708C6AFB69" box="[525,585,1133,1155]" gridcol="1" gridrow="18" pageId="2" pageNumber="280">35–40</td>
<td id="76945DB84363000DFD2C0C708CE2FB69" box="[644,705,1133,1155]" gridcol="2" gridrow="18" pageId="2" pageNumber="280">41–45</td>
<td id="76945DB84363000DFD540C708D93FB69" box="[764,944,1133,1155]" colspan="2" colspanRight="1" gridcol="3" gridrow="18" pageId="2" pageNumber="280">46–50 51–55</td>
<td id="76945DB84363000DFC430C708A04FB69" box="[1003,1063,1133,1155]" gridcol="5" gridrow="18" pageId="2" pageNumber="280">56–60</td>
</tr>
<tr id="354534C44363000DFF280C818BA2FB58" box="[128,1409,1180,1202]" gridrow="19" pageId="2" pageNumber="280" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280C818F9AFB58" box="[128,441,1180,1202]" gridcol="0" gridrow="19" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280C818F42FB58" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[128,353,1180,1202]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FF280C818F42FB58" box="[128,353,1180,1202]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FEC30C808F9AFB58" box="[363,441,1181,1202]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76945DB84363000DFDA50C818C6AFB58" box="[525,585,1180,1202]" gridcol="1" gridrow="19" pageId="2" pageNumber="280">–</td>
<td id="76945DB84363000DFD2C0C818CE2FB58" box="[644,705,1180,1202]" gridcol="2" gridrow="19" pageId="2" pageNumber="280">8</td>
<td id="76945DB84363000DFD540C818D12FB58" box="[764,817,1180,1202]" gridcol="3" gridrow="19" pageId="2" pageNumber="280">4</td>
<td id="76945DB84363000DFCD30C818D93FB58" box="[891,944,1180,1202]" gridcol="4" gridrow="19" pageId="2" pageNumber="280">3</td>
<td id="76945DB84363000DFC430C818A04FB58" box="[1003,1063,1180,1202]" gridcol="5" gridrow="19" pageId="2" pageNumber="280">–</td>
</tr>
<tr id="354534C44363000DFF280CD68BA2FB0B" box="[128,1409,1227,1249]" gridrow="20" pageId="2" pageNumber="280" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280CD68F9AFB0B" box="[128,441,1227,1249]" gridcol="0" gridrow="20" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280CD68F49FB0A" authorityName="Eggert" authorityYear="1935" box="[128,362,1227,1248]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FF280CD68F49FB0A" box="[128,362,1227,1248]" italics="true" pageId="2" pageNumber="280">Periophthalmus gracilis</emphasis>
</taxonomicName>
</th>
<td id="76945DB84363000DFDA50CD68C6AFB0B" box="[525,585,1227,1249]" gridcol="1" gridrow="20" pageId="2" pageNumber="280">8</td>
<td id="76945DB84363000DFD2C0CD68CE2FB0B" box="[644,705,1227,1249]" gridcol="2" gridrow="20" pageId="2" pageNumber="280">6</td>
<td id="76945DB84363000DFD540CD68D12FB0B" box="[764,817,1227,1249]" gridcol="3" gridrow="20" pageId="2" pageNumber="280">3</td>
<td id="76945DB84363000DFCD30CD68D93FB0B" box="[891,944,1227,1249]" gridcol="4" gridrow="20" pageId="2" pageNumber="280">6</td>
<td id="76945DB84363000DFC430CD68A04FB0B" box="[1003,1063,1227,1249]" gridcol="5" gridrow="20" pageId="2" pageNumber="280">4</td>
</tr>
<tr id="354534C44363000DFF280CE78BA2FAFA" box="[128,1409,1274,1296]" gridrow="21" pageId="2" pageNumber="280">
<th id="76945DB84363000DFF280CE78BA2FAFA" box="[128,1409,1274,1296]" colspan="9" colspanRight="8" gridcol="0" gridrow="21" pageId="2" pageNumber="280">
<emphasis id="B901EA944363FFF0FCC70CE78A08FAFA" bold="true" box="[879,1067,1274,1296]" pageId="2" pageNumber="280">Predorsal Midline</emphasis>
</th>
</tr>
<tr id="354534C44363000DFF280D348BA2FAD5" box="[128,1409,1321,1343]" gridrow="22" pageId="2" pageNumber="280" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280D348F9AFAD5" box="[128,441,1321,1343]" gridcol="0" gridrow="22" pageId="2" pageNumber="280">Range</th>
<td id="76945DB84363000DFDA50D348C6AFAD5" box="[525,585,1321,1343]" gridcol="1" gridrow="22" pageId="2" pageNumber="280">10–15</td>
<td id="76945DB84363000DFD2C0D348CE2FAD5" box="[644,705,1321,1343]" gridcol="2" gridrow="22" pageId="2" pageNumber="280">16–20</td>
<td id="76945DB84363000DFD540D348D93FAD5" box="[764,944,1321,1343]" colspan="2" colspanRight="1" gridcol="3" gridrow="22" pageId="2" pageNumber="280">21–25</td>
</tr>
<tr id="354534C44363000DFF280D458BA2FA84" box="[128,1409,1368,1390]" gridrow="23" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280D458F9AFA84" box="[128,441,1368,1390]" gridcol="0" gridrow="23" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280D458F42FA84" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[128,353,1368,1390]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FF280D458F42FA84" box="[128,353,1368,1390]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FEC30D458F9AFA87" box="[363,441,1368,1389]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76945DB84363000DFDA50D458C6AFA84" box="[525,585,1368,1390]" gridcol="1" gridrow="23" pageId="2" pageNumber="280">3</td>
<td id="76945DB84363000DFD2C0D458CE2FA84" box="[644,705,1368,1390]" gridcol="2" gridrow="23" pageId="2" pageNumber="280">12</td>
<td id="76945DB84363000DFD540D458D12FA84" box="[764,817,1368,1390]" gridcol="3" gridrow="23" pageId="2" pageNumber="280">–</td>
</tr>
<tr id="354534C44363000DFF280D9A8BA2FA77" box="[128,1409,1415,1437]" gridrow="24" pageId="2" pageNumber="280" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76945DB84363000DFF280D9A8F9AFA77" box="[128,441,1415,1437]" gridcol="0" gridrow="24" pageId="2" pageNumber="280">
<taxonomicName id="4C754D054363FFF0FF280D9A8F49FA76" authorityName="Eggert" authorityYear="1935" box="[128,362,1415,1436]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944363FFF0FF280D9A8F49FA76" box="[128,362,1415,1436]" italics="true" pageId="2" pageNumber="280">Periophthalmus gracilis</emphasis>
</taxonomicName>
*
</th>
<td id="76945DB84363000DFDA50D9A8C6AFA77" box="[525,585,1415,1437]" gridcol="1" gridrow="24" pageId="2" pageNumber="280">1</td>
<td id="76945DB84363000DFD2C0D9A8CE2FA77" box="[644,705,1415,1437]" gridcol="2" gridrow="24" pageId="2" pageNumber="280">19</td>
<td id="76945DB84363000DFD540D9A8D12FA77" box="[764,817,1415,1437]" gridcol="3" gridrow="24" pageId="2" pageNumber="280">6</td>
</tr>
</table>
</paragraph>
<caption id="DF0A660E4363FFF0FFD90F758F61F871" ID-DOI="http://doi.org/10.5281/zenodo.5355325" ID-Zenodo-Dep="5355325" httpUri="https://zenodo.org/record/5355325/files/figure.png" pageId="2" pageNumber="280" startId="2.[113,148,1896,1918]" targetBox="[113,1451,1507,1880]" targetPageId="2">
<paragraph id="8BCA36864363FFF0FFD90F758F61F871" blockId="2.[113,1451,1896,1947]" pageId="2" pageNumber="280">
Fig. 1. Holotype of 
<taxonomicName id="4C754D054363FFF0FE9B0F758C30F897" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[307,531,1896,1917]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="280" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944363FFF0FE9B0F758C30F897" box="[307,531,1896,1917]" italics="true" pageId="2" pageNumber="280">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4363FFF0FDB20F758C4BF897" box="[538,616,1896,1917]" pageId="2" pageNumber="280" rank="species">sp. nov.</taxonomicNameLabel>
MZB 23015, female, 35.3 mm SL, collected in Sumba Island, Indonesia (Photograph by: Sandra Raredon).
</paragraph>
</caption>
<caption id="DF0A660E4362FFF1FF2F0ACF8B1DFD02" ID-DOI="http://doi.org/10.5281/zenodo.5355327" ID-Zenodo-Dep="5355327" box="[135,1342,722,744]" httpUri="https://zenodo.org/record/5355327/files/figure.png" pageId="3" pageNumber="281" startId="3.[135,170,722,744]" targetBox="[135,1473,189,707]" targetPageId="3">
<paragraph id="8BCA36864362FFF1FF2F0ACF8B1DFD02" blockId="3.[135,1342,722,744]" box="[135,1342,722,744]" pageId="3" pageNumber="281">
Fig. 2. Live specimen of 
<taxonomicName id="4C754D054362FFF1FE290ACF8C41FD02" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[385,610,722,744]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="281" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944362FFF1FE290ACF8C41FD02" box="[385,610,722,744]" italics="true" pageId="3" pageNumber="281">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4362FFF1FDC20ACE8C99FD02" box="[618,698,723,744]" pageId="3" pageNumber="281" rank="species">sp. nov.</taxonomicNameLabel>
, approximately 35 mm SL, collected in Sumba Island, Indonesia.
</paragraph>
</caption>
<paragraph id="8BCA36864362FFF1FF2F0C428C3AF99D" blockId="3.[135,775,1119,1655]" pageId="3" pageNumber="281">
General description: A slender-bodied 
<taxonomicName id="4C754D054362FFF1FD8E0C428CF3FB9D" authorityName="Bloch &amp; Schneider" authorityYear="1801" box="[550,720,1119,1143]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="281" phylum="Chordata" rank="genus">
<emphasis id="B901EA944362FFF1FD8E0C428CF3FB9D" box="[550,720,1119,1143]" italics="true" pageId="3" pageNumber="281">Periophthalmus</emphasis>
</taxonomicName>
with compressed trunk and large, cylindrical head. One row of teeth in both upper and lower jaws; teeth caninoid, evenly spaced; no recurved canine teeth internal to lower jaw symphysis; tongue adnate. Trunk fully covered with scales, scales increase in size posteriorly; opercle and dorsal portion of preopercle with small embedded scales; scales absent on interorbital, isthmus, and snout. First dorsal fin tall, and somewhat rounded; first dorsal fin taller than second dorsal fin, up to two times the height of the second dorsal fin; first dorsal-fin elements almost reaching the second dorsal fin when adpressed; first and second dorsal fins closely set, distance between fins less than half the first spine length of the first dorsal fin. Pelvic frenum absent, pelvic fins separate throughout their length, no basal membrane joining innermost pelvic-fin rays; pelvic fins opaque beige, with light brown scattered pigment on the dorsal area.
</paragraph>
<paragraph id="8BCA36864362FFF1FF2F0E828DC9FB7D" blockId="3.[135,775,1695,2104]" lastBlockId="3.[834,1475,799,1175]" pageId="3" pageNumber="281">
Live colouration: Ground colour of live specimens greenishbrown to tan-brown dorsally, silvery white to light beige ventrally. Multiple silvery-white vertical lines on the trunk from the pectoral-fin base to the caudal peduncle; lines more prominent ventrally, almost indiscernible dorsally. Seven to eight dark brown saddle-like bars crossing dorsum as described below. First dorsal fin with distal silvery-white to hyaline band; a broad, submarginal dark brown or black band, posterior section intense brown to black, and appears as a spot; black spot aligns with third saddle-like bar on trunk (
<figureCitation id="134E2A034362FFF1FF780FC28F37F81D" box="[208,276,2015,2039]" captionStart="Fig" captionStartId="3.[135,170,722,744]" captionTargetBox="[135,1473,189,707]" captionTargetId="figure-7@3.[135,1473,189,707]" captionTargetPageId="3" captionText="Fig. 2. Live specimen of Periophthalmus pusing sp. nov., approximately 35 mm SL, collected in Sumba Island, Indonesia." figureDoi="http://doi.org/10.5281/zenodo.5355327" httpUri="https://zenodo.org/record/5355327/files/figure.png" pageId="3" pageNumber="281">Fig. 2</figureCitation>
); proximal to submarginal band with variable colouration, from greenish yellow anteriorly and reddishorange posteriorly, with two or three rows of bright yellow elongate to circular spots, to entirely reddish orange with one to two rows of hyaline spots. Second dorsal fin with distal hyaline band, with a broad submarginal reddish-orange and, another hyaline band, and a basal dusky reddish-brown band proximally; basal band with one to two rows of small circular dull yellow to hyaline spots. Anal fin hyaline; pectoral-fin membranes hyaline and dusky light green to brown along fin elements; caudal fin hyaline with aligned series of dark brown spots on interradial membranes, appearing as irregular seven to nine dull yellow-brown thin vertical bars; pelvic fins beige, with light subtle light yellow pigmentation on the dorsal area.
</paragraph>
<paragraph id="8BCA36864362FFF1FCEB0CA28B6DF7FD" blockId="3.[834,1476,1215,2071]" pageId="3" pageNumber="281">Colour in preservative: Ground colour of body beige to dusky brown, creamy beige ventrally; colour most intense anteriorly, dark brown around eyes and on snout. Seven to eight dark brown irregular saddle-like bars crossing dorsum; bars course anteroventrally in the following anterior to posterior pattern: immediately anterior to first dorsal fin, at mid-base of first dorsal fin, just anterior to first dorsal-fin terminus, immediately posterior to origin of second dorsal fin, mid-base of second dorsal fin, immediately posterior to the second dorsal fin, mid-caudal peduncle, posteriormost reach of caudal peduncle. First dorsal fin with thin, distal hyaline band; a broad, submarginal dark brown or black band, posterior section of the band intense brown to black, especially between the ultimate 4-5 spines that appears as a spot in the posterior portion of the first dorsal fin; spot aligns with third saddle-like bar on trunk; proximal to submarginal band, light to dusky brown anteriorly and white to hyaline posteriorly; white elongate spots in two to three rows on spines, especially anteriorly. Second dorsal fin with thin, distal hyaline band, with a broad submarginal brown band and another hyaline band, and a basal dusky brown band proximally; basal band with one to two rows of small circular spots especially on fin elements. Anal fin hyaline; pectoral-fin membranes hyaline and dusky light brown along fin elements; caudal fin hyaline with aligned series of dark brown speckles on interradial membranes, appearing as seven to nine irregular dusky brown thin vertical bars.</paragraph>
</subSubSection>
<caption id="DF0A660E4365FFF6FFD90A778BA4FD6A" ID-DOI="http://doi.org/10.5281/zenodo.5355329" ID-Zenodo-Dep="5355329" box="[113,1415,618,640]" httpUri="https://zenodo.org/record/5355329/files/figure.png" pageId="4" pageNumber="282" startId="4.[113,148,618,640]" targetBox="[113,1451,189,603]" targetPageId="4">
<paragraph id="8BCA36864365FFF6FFD90A778BA4FD6A" blockId="4.[113,1415,618,640]" box="[113,1415,618,640]" pageId="4" pageNumber="282">
Fig. 3. Live specimen of 
<taxonomicName id="4C754D054365FFF6FEC30A778C76FD95" authorityName="Eggert" authorityYear="1935" box="[363,597,618,639]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="282" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944365FFF6FEC30A778C76FD95" box="[363,597,618,639]" italics="true" pageId="4" pageNumber="282">Periophthalmus gracilis</emphasis>
</taxonomicName>
, approximately 48 mm SL, collected in Vietnam (Photograph by: Tran Dac Dinh).
</paragraph>
</caption>
<caption id="DF0A660E4365FFF6FFD90EDC8E82F91E" ID-DOI="http://doi.org/10.5281/zenodo.5355331" ID-Zenodo-Dep="5355331" httpUri="https://zenodo.org/record/5355331/files/figure.png" pageId="4" pageNumber="282" startId="4.[113,148,1729,1751]" targetBox="[113,1451,696,1713]" targetPageId="4">
<paragraph id="8BCA36864365FFF6FFD90EDC8E82F91E" blockId="4.[113,1453,1729,1780]" pageId="4" pageNumber="282">
Fig. 4. Occurrence of 
<taxonomicName id="4C754D054365FFF6FEEC0EDC8C00F93D" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[324,547,1729,1751]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="282" phylum="Chordata" rank="species" species="pusing">
<emphasis id="B901EA944365FFF6FEEC0EDC8C00F93D" box="[324,547,1729,1751]" italics="true" pageId="4" pageNumber="282">Periophthalmus pusing</emphasis>
</taxonomicName>
represented by a black circle—Sumba Island, Indonesia. The base map is courtesy of d-maps. com.
</paragraph>
</caption>
<subSubSection id="C36F650D4364FFF7FF2008A08CC9FE9F" pageId="5" pageNumber="283" type="etymology">
<paragraph id="8BCA36864364FFF7FF2008A08CC9FE9F" blockId="5.[136,775,189,373]" pageId="5" pageNumber="283">
<emphasis id="B901EA944364FFF7FF2008A08F29FF3F" bold="true" box="[136,266,189,213]" pageId="5" pageNumber="283">Etymology.</emphasis>
The specific epithet ‘pusing’, meaning ‘giddy’ in Indonesian, is the common name used by the coastal people of the 
<collectingRegion id="49B1F8644364FFF7FEBC08E38FD3FEFF" box="[276,496,253,277]" country="Indonesia" name="Nusa Tenggara Timur" pageId="5" pageNumber="283">Lesser Sunda Islands</collectingRegion>
to refer to 
<taxonomicName id="4C754D054364FFF7FDF508E08D26FEFF" authorityName="Bloch &amp; Schneider" authorityYear="1801" box="[605,773,253,277]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="genus">
<emphasis id="B901EA944364FFF7FDF508E08D26FEFF" box="[605,773,253,277]" italics="true" pageId="5" pageNumber="283">Periophthalmus</emphasis>
</taxonomicName>
mudskippers. These fishes are known as ‘Ikan Pusing’ (Indonesian: ikan=fish, pusing=giddy), as it is believed that consuming these fishes causes headaches and giddiness.
</paragraph>
</subSubSection>
<subSubSection id="C36F650D4364FFF7FF2009808FE7FD1F" pageId="5" pageNumber="283" type="distribution">
<paragraph id="8BCA36864364FFF7FF2009808FEFFE3F" blockId="5.[136,775,413,469]" pageId="5" pageNumber="283">
<emphasis id="B901EA944364FFF7FF2009808F39FE5F" bold="true" box="[136,282,413,437]" pageId="5" pageNumber="283">Distribution.</emphasis>
Presently known to occur only on the island of Sumba, 
<collectingCountry id="F36276164364FFF7FEAA09A08F49FE3F" box="[258,362,445,469]" name="Indonesia" pageId="5" pageNumber="283">Indonesia</collectingCountry>
(
<figureCitation id="134E2A034364FFF7FED309A08F9CFE3F" box="[379,447,445,469]" captionStart="Fig" captionStartId="4.[113,148,1729,1751]" captionTargetBox="[113,1451,696,1713]" captionTargetId="figure-32@4.[113,1451,696,1713]" captionTargetPageId="4" captionText="Fig. 4. Occurrence of Periophthalmus pusing represented by a black circle—Sumba Island, Indonesia. The base map is courtesy of d-maps. com." figureDoi="http://doi.org/10.5281/zenodo.5355331" httpUri="https://zenodo.org/record/5355331/files/figure.png" pageId="5" pageNumber="283">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BCA36864364FFF7FF2009E08FE7FD1F" blockId="5.[134,777,509,757]" pageId="5" pageNumber="283">
Field notes: The general locality in which this mudskipper was found—Kawangu—is a tide-dominated coastal system with relatively low wave action. Within the mangrove forest, 
<taxonomicName id="4C754D054364FFF7FD590A238EECFD9F" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944364FFF7FD590A238EECFD9F" italics="true" pageId="5" pageNumber="283">P. pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4364FFF7FF7E0A438F09FD9F" box="[214,298,606,629]" pageId="5" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
was found in a variety of microhabitat 
<typeStatus id="54CE88244364FFF7FD660A438D25FD9C" box="[718,774,606,630]" pageId="5" pageNumber="283">types</typeStatus>
in seaward and higher mangrove areas as well as within, and on sloping banks of tidal creeks. This species was syntopic with two congeners: 
<taxonomicName id="4C754D054364FFF7FEDF0AA08CF5FD3F" authorityName="Valenciennes" authorityYear="1837" box="[375,726,701,725]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="argentilineatus">
<emphasis id="B901EA944364FFF7FEDF0AA08CF5FD3F" box="[375,726,701,725]" italics="true" pageId="5" pageNumber="283">Periophthalmus argentilineatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C754D054364FFF7FF200AC08F9EFD1F" authorityName="Eggert" authorityYear="1935" box="[136,445,733,757]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="malaccensis">
<emphasis id="B901EA944364FFF7FF200AC08F9EFD1F" box="[136,445,733,757]" italics="true" pageId="5" pageNumber="283">Periophthalmus malaccensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F650D4364FFF7FF200B008FCFF81F" pageId="5" pageNumber="283" type="discussion">
<paragraph id="8BCA36864364FFF7FF200B008C5BFA9F" blockId="5.[135,776,797,1397]" pageId="5" pageNumber="283">
<emphasis id="B901EA944364FFF7FF200B008ED4FCDF" bold="true" box="[136,247,797,821]" pageId="5" pageNumber="283">Remarks.</emphasis>
Adult 
<taxonomicName id="4C754D054364FFF7FE960B008C16FCDF" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[318,565,797,821]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944364FFF7FE960B008C16FCDF" box="[318,565,797,821]" italics="true" pageId="5" pageNumber="283">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4364FFF7FD920B038CAEFCDF" box="[570,653,798,821]" pageId="5" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
differ from all other congeners, except 
<taxonomicName id="4C754D054364FFF7FE680B238C17FCBF" authorityName="Eggert" authorityYear="1935" box="[448,564,829,853]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FE680B238C17FCBF" box="[448,564,829,853]" italics="true" pageId="5" pageNumber="283">P. gracilis</emphasis>
</taxonomicName>
, in the absence of the pelvic frenum, pelvic fin not unified, and the presence of a black spot on the posterior first dorsal fin. Adult 
<taxonomicName id="4C754D054364FFF7FF200B808FA1FC5F" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[136,386,925,949]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944364FFF7FF200B808FA1FC5F" box="[136,386,925,949]" italics="true" pageId="5" pageNumber="283">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4364FFF7FE230B838FC1FC5F" box="[395,482,926,949]" pageId="5" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
can be differentiated from adult 
<taxonomicName id="4C754D054364FFF7FF6D0BA38F15FC3F" authorityName="Eggert" authorityYear="1935" box="[197,310,957,981]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FF6D0BA38F15FC3F" box="[197,310,957,981]" italics="true" pageId="5" pageNumber="283">P. gracilis</emphasis>
</taxonomicName>
in having XI–XV spines in the first dorsal fin (vs. VI–XII in 
<taxonomicName id="4C754D054364FFF7FEF40BC38FF1FC1F" authorityName="Eggert" authorityYear="1935" box="[348,466,989,1013]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FEF40BC38F4FFC1F" box="[348,364,990,1013]" italics="true" pageId="5" pageNumber="283">P</emphasis>
. 
<emphasis id="B901EA944364FFF7FED60BC08FF1FC1F" box="[382,466,989,1013]" italics="true" pageId="5" pageNumber="283">gracilis</emphasis>
</taxonomicName>
), first dorsal fin taller than second dorsal fin (first dorsal fin shorter or equal in height to second dorsal fin in 
<taxonomicName id="4C754D054364FFF7FED70C038FCCFBDF" authorityName="Eggert" authorityYear="1935" box="[383,495,1053,1077]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FED70C038FACFBDF" box="[383,399,1054,1077]" italics="true" pageId="5" pageNumber="283">P</emphasis>
. 
<emphasis id="B901EA944364FFF7FE360C008FCCFBDF" box="[414,495,1053,1077]" italics="true" pageId="5" pageNumber="283">gracilis</emphasis>
</taxonomicName>
), interdorsal distance less than half the length of the first dorsal-fin spine (interdorsal distance approximates the length of the first dorsal-fin spine in 
<taxonomicName id="4C754D054364FFF7FF0C0C638F30FB7F" authorityName="Eggert" authorityYear="1935" box="[164,275,1149,1173]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FF0C0C638E97FB7F" box="[164,180,1150,1173]" italics="true" pageId="5" pageNumber="283">P</emphasis>
. 
<emphasis id="B901EA944364FFF7FF6A0C608F30FB7F" box="[194,275,1149,1173]" italics="true" pageId="5" pageNumber="283">gracilis</emphasis>
</taxonomicName>
). Whereas juvenile specimens (approx. 
<quantity id="4C8D9B634364FFF7FD6C0C608E92FB5F" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="5" pageNumber="283" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
SL) of 
<taxonomicName id="4C754D054364FFF7FEA00C808C22FB5F" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[264,513,1181,1205]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944364FFF7FEA00C808C22FB5F" box="[264,513,1181,1205]" italics="true" pageId="5" pageNumber="283">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4364FFF7FDAF0C838C7EFB5F" box="[519,605,1182,1205]" pageId="5" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
and those of 
<taxonomicName id="4C754D054364FFF7FD590C838EF8FB3F" authorityName="Eggert" authorityYear="1935" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FD590C838EF8FB3F" italics="true" pageId="5" pageNumber="283">P. gracilis</emphasis>
</taxonomicName>
are superficially similar, differences are apparent upon scrutiny. For specimens of the same size, the first dorsal fin in 
<taxonomicName id="4C754D054364FFF7FE8B0CE08C08FAFF" authority="Jaafar &amp; Polgar &amp; Zamroni, 2016" authorityName="Jaafar &amp; Polgar &amp; Zamroni" authorityYear="2016" box="[291,555,1277,1301]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="pusing" status="sp. nov.">
<emphasis id="B901EA944364FFF7FE8B0CE08C08FAFF" box="[291,555,1277,1301]" italics="true" pageId="5" pageNumber="283">Periophthalmus pusing</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23257EF4364FFF7FD9F0CE38CB0FAFF" box="[567,659,1278,1301]" pageId="5" pageNumber="283" rank="species">sp. nov.</taxonomicNameLabel>
has more spines, is taller, and has a shorter interdorsal-fin distance. It is extremely difficult to identify the species of most 
<taxonomicName id="4C754D054364FFF7FF200D408F11FA9F" authorityName="Bloch &amp; Schneider" authorityYear="1801" box="[136,306,1373,1397]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="genus">
<emphasis id="B901EA944364FFF7FF200D408F11FA9F" box="[136,306,1373,1397]" italics="true" pageId="5" pageNumber="283">Periophthalmus</emphasis>
</taxonomicName>
specimens below 
<quantity id="4C8D9B634364FFF7FE550D408C6AFA9F" box="[509,585,1373,1397]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="283" unit="mm" value="25.0">25 mm</quantity>
SL.
</paragraph>
<paragraph id="8BCA36864364FFF7FF200D808FCFF81F" blockId="5.[136,776,1437,2037]" pageId="5" pageNumber="283">
<emphasis id="B901EA944364FFF7FF200D808FB0FA5F" bold="true" box="[136,403,1437,1461]" pageId="5" pageNumber="283">Comparative material.</emphasis>
<taxonomicName id="4C754D054364FFF7FE350D808D25FA5F" authority=": USNM" authorityName="USNM" box="[413,774,1437,1461]" class="Actinopterygii" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="283" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B901EA944364FFF7FE350D808C84FA5F" box="[413,679,1437,1461]" italics="true" pageId="5" pageNumber="283">Periophthalmus gracilis</emphasis>
: USNM
</taxonomicName>
278295, Vanikoro, 
<collectingCountry id="F36276164364FFF7FEF00DA08C2DFA3F" box="[344,526,1469,1493]" name="Solomon Islands" pageId="5" pageNumber="283">Solomon Islands</collectingCountry>
, female (
<quantity id="4C8D9B634364FFF7FDD80DA08CF3FA3F" box="[624,720,1469,1493]" metricMagnitude="-2" metricUnit="m" metricValue="4.21" pageId="5" pageNumber="283" unit="mm" value="42.1">42.1 mm</quantity>
SL); 
<materialsCitation id="3B1D3CDB4364FFF7FF200DC08D22FA1F" box="[136,769,1501,1525]" collectionCode="USNM" country="Solomon Islands" location="Solomon Islands" pageId="5" pageNumber="283" specimenCode="USNM 341322" specimenCount="1">
<specimenCode id="DBD39EFD4364FFF7FF200DC08F1BFA1F" box="[136,312,1501,1525]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="283" type="Museum">USNM 341322</specimenCode>
, 
<collectingCountry id="F36276164364FFF7FEE00DC08C2AFA1F" box="[328,521,1501,1525]" name="Solomon Islands" pageId="5" pageNumber="283">Solomon Islands</collectingCountry>
, male (
<quantity id="4C8D9B634364FFF7FDCA0DC08CE9FA1F" box="[610,714,1501,1525]" metricMagnitude="-2" metricUnit="m" metricValue="3.63" pageId="5" pageNumber="283" unit="mm" value="36.3">36.3 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FF210DE08F5EF93F" collectionCode="USNM" collectorName="Balikpapan" country="Philippines" location="Marieveles" municipality="Daintree River" pageId="5" pageNumber="283" specimenCode="USNM 278473" specimenCount="7" specimenCount-female="5" specimenCount-male="2" stateProvince="Kalimantan Timur">
<specimenCode id="DBD39EFD4364FFF7FF210DE08F16F9FF" box="[137,309,1533,1557]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="283" type="Museum">USNM 278473</specimenCode>
, 
<location id="8EAA605D4364FFF7FEEC0DE08FE3F9FF" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FEEC0DE08FE3F9FF" box="[324,448,1533,1557]" country="Philippines" municipality="Daintree River" name="Marieveles" pageId="5" pageNumber="283" stateProvince="Kalimantan Timur">Marieveles</location>
, 
<collectingCountry id="F36276164364FFF7FE670DE08C6EF9FF" box="[463,589,1533,1557]" name="Philippines" pageId="5" pageNumber="283">Philippines</collectingCountry>
, male (
<quantity id="4C8D9B634364FFF7FD0A0DE08D24F9FF" box="[674,775,1533,1557]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="5" pageNumber="283" unit="mm" value="35.8">35.8 mm</quantity>
SL), female (
<quantity id="4C8D9B634364FFF7FEB60E008FA7F9DF" box="[286,388,1565,1589]" metricMagnitude="-2" metricUnit="m" metricValue="3.55" pageId="5" pageNumber="283" unit="mm" value="35.5">35.5 mm</quantity>
SL); AMS I.22700–014, 
<collectingMunicipality id="6BAEACFC4364FFF7FD010E008EE0F9BF" pageId="5" pageNumber="283">Daintree River</collectingMunicipality>
mouth, 
<collectingCountry id="F36276164364FFF7FEB10E208F5EF9BF" box="[281,381,1597,1621]" name="Australia" pageId="5" pageNumber="283">Australia</collectingCountry>
, 
<specimenCount id="9D73FD0F4364FFF7FE2E0E208FCFF9BF" box="[390,492,1597,1621]" pageId="5" pageNumber="283" type="female">2 females</specimenCount>
(
<quantity id="4C8D9B634364FFF7FE500E208CB1F9BF" box="[504,658,1597,1621]" metricMagnitude="-2" metricUnit="m" metricValue="3.9450000000000003" metricValueMax="4.0600000000000005" metricValueMin="3.8299999999999996" pageId="5" pageNumber="283" unit="mm" value="39.45" valueMax="40.6" valueMin="38.3">38.3–40.6 mm</quantity>
SL); AMS I.26333-003, Queensland, 
<collectingCountry id="F36276164364FFF7FE0E0E408C28F99F" box="[422,523,1629,1653]" name="Australia" pageId="5" pageNumber="283">Australia</collectingCountry>
, male (
<quantity id="4C8D9B634364FFF7FDF20E408C99F99F" box="[602,698,1629,1653]" metricMagnitude="-2" metricUnit="m" metricValue="3.9299999999999997" pageId="5" pageNumber="283" unit="mm" value="39.3">39.3 mm</quantity>
SL), 
<specimenCount id="9D73FD0F4364FFF7FD510E408EF9F97F" pageId="5" pageNumber="283" type="female">3 females</specimenCount>
(
<quantity id="4C8D9B634364FFF7FF4E0E608F5CF97F" box="[230,383,1661,1685]" metricMagnitude="-2" metricUnit="m" metricValue="3.635" metricValueMax="3.96" metricValueMin="3.31" pageId="5" pageNumber="283" unit="mm" value="36.35" valueMax="39.6" valueMin="33.1">33.1–39.6 mm</quantity>
SL); AMS I.19355–022, 
<collectorName id="268053504364FFF7FD200E608D20F97F" box="[648,771,1661,1685]" pageId="5" pageNumber="283">Balikpapan</collectorName>
, 
<collectingRegion id="49B1F8644364FFF7FF200E838F63F95F" box="[136,320,1693,1717]" country="Indonesia" name="Kalimantan Timur" pageId="5" pageNumber="283">East Kalimantan</collectingRegion>
, 
<collectingCountry id="F36276164364FFF7FEE60E808F9AF95F" box="[334,441,1693,1717]" name="Indonesia" pageId="5" pageNumber="283">Indonesia</collectingCountry>
, 
<specimenCount id="9D73FD0F4364FFF7FE6E0E808C3FF95F" box="[454,540,1693,1717]" pageId="5" pageNumber="283" type="male">2 males</specimenCount>
(20.4–33.0 mm SL), female (
<quantity id="4C8D9B634364FFF7FF4B0EA08F6BF93F" box="[227,328,1725,1749]" metricMagnitude="-2" metricUnit="m" metricValue="3.31" pageId="5" pageNumber="283" unit="mm" value="33.1">33.1 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FE240EA08C6FF91F" accessionNumber="ZRC 50771" collectionCode="ZRC" country="Indonesia" location="Tritih" municipality="Cilacap" pageId="5" pageNumber="283" specimenCount="3" specimenCount-female="3" stateProvince="Jawa Timur">
<collectionCode id="ED64AE434364FFF7FE240EA08FE3F93F" box="[396,448,1725,1749]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="283">ZRC</collectionCode>
50771, 
<location id="8EAA605D4364FFF7FDB70EA08C7CF93F" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FDB70EA08C7CF93F" box="[543,607,1725,1749]" country="Indonesia" municipality="Cilacap" name="Tritih" pageId="5" pageNumber="283" stateProvince="Jawa Timur">Tritih</location>
, 
<collectingMunicipality id="6BAEACFC4364FFF7FDC50EA08CE1F93F" box="[621,706,1725,1749]" pageId="5" pageNumber="283">Cilacap</collectingMunicipality>
, 
<collectingRegion id="49B1F8644364FFF7FD780EA38D21F93F" box="[720,770,1726,1749]" country="Indonesia" name="Jawa Timur" pageId="5" pageNumber="283">Java</collectingRegion>
, 
<collectingCountry id="F36276164364FFF7FF200EC08ED1F91F" box="[136,242,1757,1781]" name="Indonesia" pageId="5" pageNumber="283">Indonesia</collectingCountry>
, 
<specimenCount id="9D73FD0F4364FFF7FEA80EC08F4AF91F" box="[256,361,1757,1781]" pageId="5" pageNumber="283" type="female">3 females</specimenCount>
(39.4–46.0 mm SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FDF20EC08C54F8DF" collectionCode="USNM" country="Brunei" location="River" pageId="5" pageNumber="283" specimenCode="USNM 366711" specimenCount="3" specimenCount-male="3" stateProvince="Temburong">
<specimenCode id="DBD39EFD4364FFF7FDF20EC08D20F91F" box="[602,771,1757,1781]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="5" pageNumber="283" type="Museum">USNM 366711</specimenCode>
, 
<collectingRegion id="49B1F8644364FFF7FF200EE08F2BF8FF" box="[136,264,1789,1813]" country="Brunei" name="Temburong" pageId="5" pageNumber="283">Temburong</collectingRegion>
<location id="8EAA605D4364FFF7FEBA0EE08F71F8FF" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FEBA0EE08F71F8FF" box="[274,338,1789,1813]" country="Brunei" name="River" pageId="5" pageNumber="283" stateProvince="Temburong">River</location>
, 
<collectingCountry id="F36276164364FFF7FEC90EE08F8CF8FF" box="[353,431,1789,1813]" name="Brunei" pageId="5" pageNumber="283">Brunei</collectingCountry>
, 
<specimenCount id="9D73FD0F4364FFF7FE160EE08C35F8FF" box="[446,534,1789,1813]" pageId="5" pageNumber="283" type="male">3 males</specimenCount>
(
<quantity id="4C8D9B634364FFF7FD800EE08CEFF8FF" box="[552,716,1789,1813]" metricMagnitude="-2" metricUnit="m" metricValue="3.34" metricValueMax="3.44" metricValueMin="3.2399999999999998" pageId="5" pageNumber="283" unit="mm" value="33.4" valueMax="34.4" valueMin="32.4">32.4–34.4 mm</quantity>
SL), female (35.0 mm SL), juvenile (
<quantity id="4C8D9B634364FFF7FE4B0F008C67F8DF" box="[483,580,1821,1845]" metricMagnitude="-2" metricUnit="m" metricValue="2.67" pageId="5" pageNumber="283" unit="mm" value="26.7">26.7 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FD2B0F008D21F8BF" accessionNumber="ZRC 27057" collectionCode="ZRC" country="Singapore" location="St. John's Island" pageId="5" pageNumber="283" specimenCount="2" specimenCount-male="2">
<collectionCode id="ED64AE434364FFF7FD2B0F008C94F8DF" box="[643,695,1821,1845]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="283">ZRC</collectionCode>
27057, 
<location id="8EAA605D4364FFF7FF200F208F62F8BF" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FF200F208F62F8BF" box="[136,321,1853,1877]" country="Singapore" name="St. John's Island" pageId="5" pageNumber="283">St. John’s Island</location>
, 
<collectingCountry id="F36276164364FFF7FEE60F208F9EF8BF" box="[334,445,1853,1877]" name="Singapore" pageId="5" pageNumber="283">Singapore</collectingCountry>
, 
<specimenCount id="9D73FD0F4364FFF7FE620F208C3CF8BF" box="[458,543,1853,1877]" pageId="5" pageNumber="283" type="male">2 males</specimenCount>
(
<quantity id="4C8D9B634364FFF7FD870F208CEDF8BF" box="[559,718,1853,1877]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" metricValueMax="3.4299999999999997" metricValueMin="3.41" pageId="5" pageNumber="283" unit="mm" value="34.2" valueMax="34.3" valueMin="34.1">34.1–34.3 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FF200F408C2AF87F" accessionNumber="ZRC 53201" collectionCode="ZRC" country="Singapore" location="Pandan Mangroves" pageId="5" pageNumber="283" specimenCount="2" specimenCount-female="2">
<collectionCode id="ED64AE434364FFF7FF200F408E9FF89F" box="[136,188,1885,1909]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="283">ZRC</collectionCode>
53201, 
<location id="8EAA605D4364FFF7FEBA0F408FC1F89C" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FEBA0F408FC1F89C" box="[274,482,1885,1910]" country="Singapore" name="Pandan Mangroves" pageId="5" pageNumber="283">Pandan Mangroves</location>
, 
<collectingCountry id="F36276164364FFF7FE440F408C78F89F" box="[492,603,1885,1909]" name="Singapore" pageId="5" pageNumber="283">Singapore</collectingCountry>
, male (
<quantity id="4C8D9B634364FFF7FD0F0F408D25F89F" box="[679,774,1885,1909]" metricMagnitude="-2" metricUnit="m" metricValue="3.41" pageId="5" pageNumber="283" unit="mm" value="34.1">34.1 mm</quantity>
SL), 
<specimenCount id="9D73FD0F4364FFF7FF170F608F0BF87F" box="[191,296,1917,1941]" pageId="5" pageNumber="283" type="female">2 females</specimenCount>
(
<quantity id="4C8D9B634364FFF7FE900F608FF6F87F" box="[312,469,1917,1941]" metricMagnitude="-2" metricUnit="m" metricValue="3.4349999999999996" metricValueMax="3.5799999999999996" metricValueMin="3.29" pageId="5" pageNumber="283" unit="mm" value="34.349999999999994" valueMax="35.8" valueMin="32.9">32.9–35.8 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FDBE0F608EC2F83F" accessionNumber="ZRC 37792" collectionCode="ZRC" country="Malaysia" location="East" pageId="5" pageNumber="283" specimenCount="6" specimenCount-female="6">
<collectionCode id="ED64AE434364FFF7FDBE0F608C69F87F" box="[534,586,1917,1941]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="283">ZRC</collectionCode>
37792, Sarawak, 
<location id="8EAA605D4364FFF7FF200F838E95F85F" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FF200F838E95F85F" box="[136,182,1950,1973]" country="Malaysia" name="East" pageId="5" pageNumber="283">East</location>
<collectingCountry id="F36276164364FFF7FF680F808F06F85F" box="[192,293,1949,1973]" name="Malaysia" pageId="5" pageNumber="283">Malaysia</collectingCountry>
, male (
<quantity id="4C8D9B634364FFF7FED00F808FFFF85F" box="[376,476,1949,1973]" metricMagnitude="-2" metricUnit="m" metricValue="3.45" pageId="5" pageNumber="283" unit="mm" value="34.5">34.5 mm</quantity>
SL), 
<specimenCount id="9D73FD0F4364FFF7FDB70F808CAAF85F" box="[543,649,1949,1973]" pageId="5" pageNumber="283" type="female">6 females</specimenCount>
(
<quantity id="4C8D9B634364FFF7FD330F808E92F83F" metricMagnitude="-2" metricUnit="m" metricValue="3.575" metricValueMax="3.8299999999999996" metricValueMin="3.32" pageId="5" pageNumber="283" unit="mm" value="35.75" valueMax="38.3" valueMin="33.2">33.2–38.3 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="3B1D3CDB4364FFF7FF430FA08FCBF81F" accessionNumber="ZRC 47563" collectionCode="ZRC" country="Malaysia" location="Pantai Batu Hitam" municipality="Peninsular" pageId="5" pageNumber="283" specimenCount="1" stateProvince="Pahang">
<collectionCode id="ED64AE434364FFF7FF430FA08F3CF83F" box="[235,287,1981,2005]" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="5" pageNumber="283">ZRC</collectionCode>
47563, 
<location id="8EAA605D4364FFF7FED90FA08C16F83F" LSID="urn:lsid:plazi:treatment:03DC87904360FFF7FEBD0F8A8FCFF81F:8EAA605D4364FFF7FED90FA08C16F83F" box="[369,565,1981,2005]" country="Malaysia" municipality="Peninsular" name="Pantai Batu Hitam" pageId="5" pageNumber="283" stateProvince="Pahang">Pantai Batu Hitam</location>
, 
<collectingRegion id="49B1F8644364FFF7FD960FA08CACF83F" box="[574,655,1981,2005]" country="Malaysia" name="Pahang" pageId="5" pageNumber="283">Pahang</collectingRegion>
, 
<collectingMunicipality id="6BAEACFC4364FFF7FD3F0FA08D24F83F" box="[663,775,1981,2005]" pageId="5" pageNumber="283">Peninsular</collectingMunicipality>
<collectingCountry id="F36276164364FFF7FF200FC08ECFF81F" box="[136,236,2013,2037]" name="Malaysia" pageId="5" pageNumber="283">Malaysia</collectingCountry>
, female (
<quantity id="4C8D9B634364FFF7FEFA0FC08F90F81F" box="[338,435,2013,2037]" metricMagnitude="-2" metricUnit="m" metricValue="3.9799999999999995" pageId="5" pageNumber="283" unit="mm" value="39.8">39.8 mm</quantity>
SL)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>