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<mods:title id="FB6E2D2C8FA92781F54B9A6A6B659C20">The Inocellia crassicornis species group (Raphidioptera: Inocelliidae) in mainland China, with description of two new species</mods:title>
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<mods:namePart id="B2579D5B28206267720378C4AA0DF1A8">Yang, Ding</mods:namePart>
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<mods:namePart id="5FE7BCAE30B126551FCE995515BEF9D5">Aspöck, Ulrike</mods:namePart>
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<treatment id="03DC875AFF9879031DA8FBFF77C966A4" ID-DOI="http://doi.org/10.5281/zenodo.6199297" ID-GBIF-Taxon="119390366" ID-Zenodo-Dep="6199297" LSID="urn:lsid:plazi:treatment:03DC875AFF9879031DA8FBFF77C966A4" httpUri="http://treatment.plazi.org/id/03DC875AFF9879031DA8FBFF77C966A4" lastPageId="12" lastPageNumber="52" pageId="6" pageNumber="46">
<subSubSection id="C36F65C7FF9879091DA8FBFF73086663" pageId="6" pageNumber="46" type="nomenclature">
<paragraph id="8BCA364CFF9879091DA8FBFF73AA664E" blockId="6.[151,497,1085,1146]" box="[151,497,1085,1111]" pageId="6" pageNumber="46">
<heading id="D0828120FF9879091DA8FBFF73AA664E" bold="true" box="[151,497,1085,1111]" fontSize="11" level="1" pageId="6" pageNumber="46" reason="1">
<taxonomicName id="4C754DCFFF9879091DA8FBFF73AA664E" ID-CoL="3PQGH" authority="Yang, 1999" authorityName="Yang" authorityYear="1999" box="[151,497,1085,1111]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9879091DA8FBFF73AA664E" bold="true" box="[151,497,1085,1111]" pageId="6" pageNumber="46">
<emphasis id="B901EA5EFF9879091DA8FBFF7339664E" bold="true" box="[151,354,1085,1111]" italics="true" pageId="6" pageNumber="46">Inocellia fujiana</emphasis>
<bibRefCitation id="EFE44BBDFF9879091C56FBFF73AA664E" author="Yang" box="[361,497,1085,1111]" pageId="6" pageNumber="54" refString="Yang, C. K. (1999) Raphidioptera: Family Inocelliidae. In: Huang, B. K. (ed). Fauna of insects in Fujian Province of China, Volume 3. Fujian Science Press, Fuzhou, pp. 177 - 180." type="book" year="1999">Yang, 1999</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BCA364CFF9879091DA8FBA273086663" blockId="6.[151,497,1085,1146]" box="[151,339,1120,1146]" pageId="6" pageNumber="46">
(
<figureCitation id="134E2AC9FF9879091DA0FBA272A86663" box="[159,243,1120,1146]" captionStart="FIGURES 3 4" captionStartId="5.[151,271,1672,1696]" captionTargetBox="[152,1435,208,1647]" captionTargetId="figure@5.[240,1386,185,997]" captionTargetPageId="5" captionText="FIGURES 3 4. Habitus images of the species of the Inocellia crassicornis group. 3. I. digitiformis sp. nov., holotype male; 4. I. fujiana Yang, holotype male. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/196508/files/figure.png" pageId="6" pageNumber="46">Figs. 4</figureCitation>
,
<figureCitation id="134E2AC9FF9879091DC0FBA273116663" box="[255,330,1120,1146]" captionStart-0="FIGURES 29 37" captionStart-1="FIGURES 38 43" captionStartId-0="10.[151,274,1734,1758]" captionStartId-1="11.[151,270,1217,1241]" captionTargetBox-0="[154,1429,546,1701]" captionTargetBox-1="[162,1421,275,1190]" captionTargetId-0="figure@10.[151,1436,530,1713]" captionTargetId-1="figure@11.[155,1433,269,1196]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." captionText-1="FIGURES 38 43. Geographic variation of the male genitalia of I. fujiana. 38. Genital segments, lateral view; 39. Male genital segments, caudal view; 40. Gonarcus (gonocoxites 11), dorsal view; 41. Genital segments, lateral view; 42. Male genital segments, caudal view; 43. Gonarcus (gonocoxites 11), dorsal view. 38 40: male from Beijing; 41 43: male from Shaanxi Prov. Scale bars = 0.5 mm." httpUri-0="https://zenodo.org/record/196512/files/figure.png" httpUri-1="https://zenodo.org/record/196513/files/figure.png" pageId="6" pageNumber="46">2943</figureCitation>
)
</paragraph>
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<subSubSection id="C36F65C7FF9879091DA8FB68711066F8" pageId="6" pageNumber="46" type="reference_group">
<paragraph id="8BCA364CFF9879091DA8FB68711066F8" blockId="6.[151,863,1194,1249]" pageId="6" pageNumber="46">
<treatmentCitationGroup id="AB651162FF9879091DA8FB68711066F8" pageId="6" pageNumber="46">
<taxonomicName id="4C754DCFFF9879091DA8FB68705A66D8" ID-CoL="3PQGH" authority="Yang, 1999: 178" authorityName="Yang" authorityPageNumber="178" authorityYear="1999" box="[151,513,1194,1217]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9879091DA8FB68731366D8" box="[151,328,1194,1217]" italics="true" pageId="6" pageNumber="46">Inocellia fujiana</emphasis>
<treatmentCitation id="0AD4105DFF9879091C71FB68705A66D8" author="Yang" box="[334,513,1194,1217]" page="178" pageId="6" pageNumber="46" year="1999">
<bibRefCitation id="EFE44BBDFF9879091C71FB68739766D8" author="Yang" box="[334,460,1194,1217]" pageId="6" pageNumber="54" refString="Yang, C. K. (1999) Raphidioptera: Family Inocelliidae. In: Huang, B. K. (ed). Fauna of insects in Fujian Province of China, Volume 3. Fujian Science Press, Fuzhou, pp. 177 - 180." type="book" year="1999">Yang, 1999</bibRefCitation>
: 178
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</taxonomicName>
.
<typeStatus id="54CE88EEFF9879091F34FB68701B66D8" box="[523,576,1194,1217]" pageId="6" pageNumber="46">Type</typeStatus>
locality: Fujian (Nanjing).
<taxonomicName id="4C754DCFFF9879091DA8FB08739766F8" ID-CoL="3PQGH" authority="Yang, 1999" authorityName="Yang" authorityYear="1999" box="[151,460,1226,1249]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="6" pageNumber="46" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9879091DA8FB08731366F8" box="[151,328,1226,1249]" italics="true" pageId="6" pageNumber="46">Inocellia fujiana</emphasis>
<bibRefCitation id="EFE44BBDFF9879091C71FB08739766F8" author="Yang" box="[334,460,1226,1249]" pageId="6" pageNumber="54" refString="Yang, C. K. (1999) Raphidioptera: Family Inocelliidae. In: Huang, B. K. (ed). Fauna of insects in Fujian Province of China, Volume 3. Fujian Science Press, Fuzhou, pp. 177 - 180." type="book" year="1999">Yang, 1999</bibRefCitation>
</taxonomicName>
: U. Aspöck
<emphasis id="B901EA5EFF9879091F70FB0870DE66F8" box="[591,645,1226,1249]" italics="true" pageId="6" pageNumber="46">et al.</emphasis>
2009 (taxonomy).
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</paragraph>
</subSubSection>
<subSubSection id="C36F65C7FF9879091DA8FACD71626756" pageId="6" pageNumber="46" type="diagnosis">
<paragraph id="8BCA364CFF9879091DA8FACD71626756" blockId="6.[151,1437,1295,2031]" pageId="6" pageNumber="46">
<emphasis id="B901EA5EFF9879091DA8FACD734C6730" bold="true" box="[151,279,1295,1321]" pageId="6" pageNumber="46">Diagnosis.</emphasis>
The species is characterized by the feebly prominent styli on the male ninth gonocoxite and by the gonarcus (fused gonocoxites 11) with a bifurcate process.
</paragraph>
</subSubSection>
<subSubSection id="C36F65C7FF9879051DF9FA9872AB63C5" lastPageId="10" lastPageNumber="50" pageId="6" pageNumber="46" type="description">
<paragraph id="8BCA364CFF9879091DF9FA987156676D" blockId="6.[151,1437,1295,2031]" box="[198,781,1370,1396]" pageId="6" pageNumber="46">
<emphasis id="B901EA5EFF9879091DF9FA987304676D" bold="true" box="[198,351,1370,1396]" pageId="6" pageNumber="46">Description.</emphasis>
Male. Forewing length 7.58.0 mm.
</paragraph>
<paragraph id="8BCA364CFF9879091DFAFA42710167FD" blockId="6.[151,1437,1295,2031]" pageId="6" pageNumber="46">Head subquadrate, black, clypeus yellowish brown, vertex laterally with two pairs of indistinct brownish cuneate markings and medially with a pair of brownish vittae. Antennal sclerite (torulus), scape, and pedicel yellow; flagellum pale brown. Mouthparts yellowish brown.</paragraph>
<paragraph id="8BCA364CFF9879091DFAFA3272AA6486" blockId="6.[151,1437,1295,2031]" pageId="6" pageNumber="46">Thorax blackish brown; anterior half of pronotum medially with a pair of narrow, hook-like, brownish markings; meso- and metanota each with a yellowish marking anteromedially, and with a yellowish transverse marking on posterior margin of scutellum. Legs yellow with yellowish setae. Wings hyaline, pterostigma brown, veins pale brown. Anterior branch of Rs with one forked vein and one simple vein running to wing margin.</paragraph>
<paragraph id="8BCA364CFF9879091DFAF96877C665F6" blockId="6.[151,1437,1295,2031]" pageId="6" pageNumber="46">
Abdomen blackish brown, venter much paler; each pregenital segment with a yellowish transverse marking on posterior margins of tergite and sternite; ninth segment with yellow posterior margin gonocoxite brown, ectoproct yellow. Ninth gonocoxite (
<figureCitation id="134E2AC9FF9879091F96F93771636516" box="[681,824,1781,1807]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="6" pageNumber="46">Figs. 2931</figureCitation>
) shell-like, nearly semicircular, slightly wider than long; subdistal portion of inner side dorsally with a blunt stylus bearing a bristle tuft, an additional bristle tuft lies anteriorly to the stylus, and a few bristles are present ventrally on posterior margin. Pseudostyli (basal parts of gonapophyses 9) (
<figureCitation id="134E2AC9FF9879091CDDF8A770666566" box="[482,573,1893,1919]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="6" pageNumber="46">Fig. 30</figureCitation>
) paired and foliate. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10) (
<figureCitation id="134E2AC9FF9879091F18F84870E865BD" box="[551,691,1930,1956]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="6" pageNumber="46">Figs. 31, 33</figureCitation>
) small, proximal portion flattened and roundly expanded, distal projection slightly curved dorsad with concave tip. Gonarcus (fused gonocoxites 11) (
<figureCitation id="134E2AC9FF9879091941F872775065D3" box="[1150,1291,1968,1994]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="6" pageNumber="46">Figs. 3032</figureCitation>
) shield-like, dorsal margin produced posteriad, in caudal view subtriangular, medially with a bifurcate projection.
</paragraph>
<paragraph id="8BCA364CFF9979081DA8FF5A70A462E5" blockId="7.[151,1437,152,290]" pageId="7" pageNumber="47">
Endophallus (
<figureCitation id="134E2AC9FF9979081C7CFF5A738262AB" box="[323,473,152,178]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="7" pageNumber="47">Figs. 3031</figureCitation>
) short, distal portion medially with a pair of bristle tufts directed dorsad and additionally two or four bristles directed ventrad. Ectoproct (
<figureCitation id="134E2AC9FF9979081E63FF7F71EA62CE" box="[860,945,189,215]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="7" pageNumber="47">Fig. 29</figureCitation>
) subquadrate in lateral view. Hypandrium internum (
<figureCitation id="134E2AC9FF9979081C2EFF20733D62E5" box="[273,358,226,252]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="7" pageNumber="47">Fig. 34</figureCitation>
) small, lobes considerably narrow.
</paragraph>
<paragraph id="8BCA364CFF9979081DFAFECA70CE633B" blockId="7.[151,1437,152,290]" box="[197,661,264,290]" pageId="7" pageNumber="47">Female. Forewing length 9.012.0 mm.</paragraph>
<caption id="DF0A66C4FF9979081DA8F91B704465E8" httpUri="https://zenodo.org/record/196509/files/figure.png" pageId="7" pageNumber="47" targetBox="[168,1421,346,1715]" targetPageId="7">
<paragraph id="8BCA364CFF9979081DA8F91B704465E8" blockId="7.[151,1437,1753,2033]" pageId="7" pageNumber="47">
<emphasis id="B901EA5EFF9979081DA8F91B730B64E8" bold="true" box="[151,336,1753,1777]" pageId="7" pageNumber="47">FIGURES 513</emphasis>
.
<taxonomicName id="4C754DCFFF9979081C61F918701064E8" box="[350,587,1754,1777]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="7" pageNumber="47" phylum="Arthropoda" rank="species" species="biprocessus" status="sp. nov.">
<emphasis id="B901EA5EFF9979081C61F918701064E8" box="[350,587,1754,1777]" italics="true" pageId="7" pageNumber="47">Inocellia biprocessus</emphasis>
</taxonomicName>
<emphasis id="B901EA5EFF9979081F6CF91B70F664E8" bold="true" box="[595,685,1753,1777]" pageId="7" pageNumber="47">
<taxonomicNameLabel id="A2325725FF9979081F6CF91B70F664E8" box="[595,685,1753,1777]" pageId="7" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(paratypes) 5. Male genital segments, lateral view; 6. Male genital segments, caudal view; 7. Male genital segments showing internal structures, lateral view; 8. Gonarcus (fused gonocoxites 11), dorsal view; 9. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10), dorsal view; 10. Hypandrium internum, ventral view; 11. Female genital segments, lateral view; 12. Female genital segments, ventral view; 13. Bursa copulatrix, lateral view. ab: atrium bursae; e: ectoproct; ep: endophallus; g: gonarcus (fused gonocoxites 11); gr: glandula receptaculi; gx9: gonocoxite 9; o: ovipositor; p: fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10); ps: pseudostylus (basal parts of gonapophysis 9); r: receptaculum seminis; s: stylus or stylus-like process (gonostylus 9); sb: sacculus bursae; sg: subgenitale; S7, 9: sternite 7 and 9; T79: tergite 7 and 9. Scale bars = 0.5 mm.
</paragraph>
</caption>
<caption id="DF0A66C4FF9679071DA8F90D77D0657E" httpUri="https://zenodo.org/record/196510/files/figure.png" pageId="8" pageNumber="48" targetBox="[161,1417,211,1706]" targetPageId="8">
<paragraph id="8BCA364CFF9679071DA8F90D77D0657E" blockId="8.[151,1437,1743,1895]" pageId="8" pageNumber="48">
<emphasis id="B901EA5EFF9679071DA8F90D730364FE" bold="true" box="[151,344,1743,1767]" pageId="8" pageNumber="48">FIGURES 1422</emphasis>
.
<taxonomicName id="4C754DCFFF9679071C59F912708664FE" authority="Schummel" authorityName="Schummel" box="[358,733,1744,1767]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="8" pageNumber="48" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B901EA5EFF9679071C59F912701464FE" box="[358,591,1744,1767]" italics="true" pageId="8" pageNumber="48">Inocellia crassicornis</emphasis>
(Schummel)
</taxonomicName>
. (male and female from Jilin Prov., northeastern China) 14. Male genital segments, lateral view; 15. Male genital segments, caudal view; 16. Male genital segments showing internal structures, lateral view; 17. Gonarcus (fused gonocoxites 11), dorsal view; 18. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10), dorsal view; 19. Hypandrium internum, ventral view; 20. Female genital segments, lateral view; 21. Female genital segments, ventral view; 22. Bursa copulatrix, lateral view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCA364CFF9679061DFAF857712D62E5" blockId="8.[151,1436,1941,2004]" lastBlockId="9.[151,1437,152,252]" lastPageId="9" lastPageNumber="49" pageId="8" pageNumber="48">
Seventh sternite (
<figureCitation id="134E2AC9FF9679071C9FF857706C65B6" box="[416,567,1941,1967]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="8" pageNumber="48">Figs. 3536</figureCitation>
) subtrapezoidal in lateral view, posteriorly produced; in ventral view posterior margin arcuately incised medially, forming a pair of small subtriangular processes. Subgenitale (
<figureCitation id="134E2AC9FF9779061D9FFF5A736F62AB" box="[160,308,152,178]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="9" pageNumber="49">Figs. 3536</figureCitation>
) small, suboval. Ectoproct (
<figureCitation id="134E2AC9FF9779061FB0FF5A70B162AB" box="[655,746,152,178]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="9" pageNumber="49">Fig. 35</figureCitation>
) subquadrate in lateral view, slightly wider than long in lateral view. Atrium bursae (
<figureCitation id="134E2AC9FF9779061CD3FF7F701862CE" box="[492,579,189,215]" captionStart="FIGURES 29 37" captionStartId="10.[151,274,1734,1758]" captionTargetBox="[154,1429,546,1701]" captionTargetId="figure@10.[151,1436,530,1713]" captionTargetPageId="10" captionText="FIGURES 29 37. Inocellia fujiana Yang. 29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="9" pageNumber="49">Fig. 36</figureCitation>
) slightly pointed dorsad; sacculus bursae tubular; receptaculum seminis long, with a pair of slender tubular glandula receptaculi distad.
</paragraph>
<caption id="DF0A66C4FF9779061DA8F96671EE6505" httpUri="https://zenodo.org/record/196511/files/figure.png" pageId="9" pageNumber="49" targetBox="[159,1430,326,1667]" targetPageId="9">
<paragraph id="8BCA364CFF9779061DA8F96671EE6505" blockId="9.[151,1436,1700,1820]" pageId="9" pageNumber="49">
<emphasis id="B901EA5EFF9779061DA8F966730164A5" bold="true" box="[151,346,1700,1724]" pageId="9" pageNumber="49">FIGURES 2328</emphasis>
.
<taxonomicName id="4C754DCFFF9779061C57F967701464A5" box="[360,591,1701,1724]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="9" pageNumber="49" phylum="Arthropoda" rank="species" species="digitiformis" status="sp. nov.">
<emphasis id="B901EA5EFF9779061C57F967701464A5" box="[360,591,1701,1724]" italics="true" pageId="9" pageNumber="49">Inocellia digitiformis</emphasis>
</taxonomicName>
<emphasis id="B901EA5EFF9779061F69F96670F564A5" bold="true" box="[598,686,1700,1724]" pageId="9" pageNumber="49">
<taxonomicNameLabel id="A2325725FF9779061F69F96670F564A5" box="[598,686,1700,1724]" pageId="9" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(holotype) 23. Male genital segments, lateral view; 24. Male genital segments, caudal view; 25. Male genital segments showing internal structures, lateral view; 26. Gonarcus (fused gonocoxites 11), dorsal view; 27. Fused parameres (complex of fused gonocoxites, gonapophyses, and gonostyli 10), dorsal view; 28. Hypandrium internum, ventral view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BCA364CFF9779051DFAF88872AB63C5" blockId="9.[151,1436,1866,2004]" lastBlockId="10.[151,1437,150,476]" lastPageId="10" lastPageNumber="50" pageId="9" pageNumber="49">
<emphasis id="B901EA5EFF9779061DFAF8887380657D" bold="true" box="[197,475,1866,1892]" pageId="9" pageNumber="49">Specimens examined.</emphasis>
<typeStatus id="54CE88EEFF9779061CDBF888700D657D" box="[484,598,1866,1892]" pageId="9" pageNumber="49" type="holotype">Holotype</typeStatus>
3 [pinned, genitalia missing],
<collectingCountry id="F36276DCFF9779061EE8F8887660657D" box="[983,1083,1866,1892]" name="China" pageId="9" pageNumber="49">CHINA</collectingCountry>
: Fujian Prov., Nanjing, Hexi [24°30΄N, 117°21΄E],
<date id="FFCB108CFF9779061C9DF8AD70776590" box="[418,556,1903,1929]" pageId="9" pageNumber="49" value="1965-04-20">20.IV.1965</date>
, S. Liu (CAU).
<collectingCountry id="F36276DCFF9779061FCDF8AD710D6590" box="[754,854,1903,1929]" name="China" pageId="9" pageNumber="49">CHINA</collectingCountry>
: 13 [preserved in alcohol], Beijing, Xiangshan [39°59΄N, 116°10΄E],
<date id="FFCB108CFF9779061C97F857706365B6" box="[424,568,1941,1967]" pageId="9" pageNumber="49" value="1961-06-24">24.VI.1961</date>
, F. Li (CAU); 131Ƥ [preserved in alcohol], Beijing, Xiangshan,
<date id="FFCB108CFF9779061851F857734565CD" pageId="9" pageNumber="49" value="1964-06-24" valueMax="1964-06-25" valueMin="1964-06-24">24 25.VI.1964</date>
, F. Li (CAU); 1Ƥ [preserved in alcohol], Beijing, Xiangshan,
<date id="FFCB108CFF9779061EC8F878762465CD" box="[1015,1151,1978,2004]" pageId="9" pageNumber="49" value="1964-06-24">24.VI.1964</date>
, C. Yang (CAU); 136Ƥ [pinned], Beijing, Xiangshan,
<date id="FFCB108CFF9479051F33FF5A70C062AB" box="[524,667,152,178]" pageId="10" pageNumber="50" value="1974-06-24">24.VI.1974</date>
, F. Li (CAU); 13 [preserved in alcohol], Beijing, Mentougou, Xiaolongmen [39°58΄N, 115°26΄E],
<quantity id="4C8D9BA9FF9479051F6AFF7F715A62CE" box="[597,769,189,215]" metricMagnitude="3" metricUnit="m" metricValue="1.3035" metricValueMax="1.43" metricValueMin="1.177" pageId="10" pageNumber="50" unit="m" value="1303.5" valueMax="1430.0" valueMin="1177.0">11771430 m</quantity>
,
<date id="FFCB108CFF9479051E31FF7F71CB62CE" box="[782,912,189,215]" pageId="10" pageNumber="50" value="2009-05-25">25.V.2009</date>
, H. Yu (CAU); 2Ƥ [preserved in alcohol], Beijing, Mentougou, Xiaolongmen,
<quantity id="4C8D9BA9FF9479051F5EFF20714962E5" box="[609,786,226,252]" metricMagnitude="3" metricUnit="m" metricValue="1.331" metricValueMax="1.43" metricValueMin="1.232" pageId="10" pageNumber="50" unit="m" value="1331.0" valueMax="1430.0" valueMin="1232.0">12321430 m</quantity>
,
<date id="FFCB108CFF9479051E1FFF2071F862E5" box="[800,931,226,252]" pageId="10" pageNumber="50" value="2009-05-24">24.V.2009</date>
, L. Liang (CAU); 7322Ƥ [preserved in alcohol], Shanxi Prov., Taigu [37°25΄N, 112°32΄E] (CAU); 13 [pinned], Shanxi Prov., Qingxu [37°36΄N, 112°21΄E],
<date id="FFCB108CFF9479051C22FEEF7325635E" box="[285,382,301,327]" pageId="10" pageNumber="50" value="1959-06">VI.1959</date>
(CAU); 23 [pinned], Shanxi Prov., Nanwutai [38°58΄N, 113°36΄E],
<date id="FFCB108CFF9479051994FEEF7756635E" box="[1195,1293,301,327]" pageId="10" pageNumber="50" value="1980-06">VI.1980</date>
(CAU); 1Ƥ [pinned], Shanxi Prov., Taiyuan [37°52΄N, 112°33΄E] (CAU); 132Ƥ [preserved in alcohol], Shaanxi Prov., Huashan [34°29΄N, 110°05΄E],
<date id="FFCB108CFF9479051F31FEBA703D638B" box="[526,614,376,402]" pageId="10" pageNumber="50" value="1951-05">V.1951</date>
, Y. Zhou (CAU); 536Ƥ [pinned], Shaanxi Prov., Huashan, 1963, Y. Zhou (CAU); 1Ƥ [pinned], Henan Prov., Lushi, Dakuaidi [33°54΄N, 110°43΄E],
<quantity id="4C8D9BA9FF9479051905FE5F76CE63AE" box="[1082,1173,413,439]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="10" pageNumber="50" unit="m" value="1800.0">1800 m</quantity>
,
<date id="FFCB108CFF947905199FFE5F776963AE" box="[1184,1330,413,439]" pageId="10" pageNumber="50" value="2001-07-20">20.VII.2001</date>
, X. Shen (CAU).
</paragraph>
</subSubSection>
<caption id="DF0A66C4FF9479051DA8F90477196544" httpUri="https://zenodo.org/record/196512/files/figure.png" pageId="10" pageNumber="50" targetBox="[154,1429,546,1701]" targetPageId="10">
<paragraph id="8BCA364CFF9479051DA8F90477196544" blockId="10.[151,1436,1734,1885]" pageId="10" pageNumber="50">
<emphasis id="B901EA5EFF9479051DA8F904733B64C7" bold="true" box="[151,352,1734,1758]" pageId="10" pageNumber="50">FIGURES 2937</emphasis>
.
<taxonomicName id="4C754DCFFF9479051C50F905702E64C4" authority="Yang." authorityName="Yang." box="[367,629,1734,1758]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9479051C50F905707064C7" box="[367,555,1735,1758]" italics="true" pageId="10" pageNumber="50">Inocellia fujiana</emphasis>
Yang.
</taxonomicName>
29. (male and female from Shanxi Prov., northern China) Male genital segments, lateral view; 30. Male genital segments, caudal view; 31. Male genital segments showing internal structures, lateral view; 32. Gonarcus fused (gonocoxites 11), dorsal view; 33. Fused parameres (complex of fused gonocoxites, gonapophyse, and gonostyli 10), dorsal view; 34. Hypandrium internum, ventral view; 35. Female genital segments, lateral view; 36. Female genital segments, ventral view; 37. Bursa copulatrix, lateral view. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C36F65C7FF9479041DFAF84E736E62CE" lastPageId="11" lastPageNumber="51" pageId="10" pageNumber="50" type="distribution">
<paragraph id="8BCA364CFF9479041DFAF84E736E62CE" blockId="10.[151,1436,1932,2033]" lastBlockId="11.[151,1436,152,215]" lastPageId="11" lastPageNumber="51" pageId="10" pageNumber="50">
<emphasis id="B901EA5EFF9479051DFAF84E733E65BF" bold="true" box="[197,357,1932,1958]" pageId="10" pageNumber="50">Distribution.</emphasis>
<collectingCountry id="F36276DCFF9479051C53F84E73E965BF" box="[364,434,1932,1958]" name="China" pageId="10" pageNumber="50">China</collectingCountry>
(Beijing, Fujian Prov., Henan Prov., Shanxi Prov., Shaanxi Prov.). Biogeographically,
<taxonomicName id="4C754DCFFF9479051DA8F873735865D3" box="[151,259,1969,1994]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="10" pageNumber="50" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9479051DA8F873735865D3" box="[151,259,1969,1994]" italics="true" pageId="10" pageNumber="50">I. fujiana</emphasis>
</taxonomicName>
probably represents a Manchurian faunal element of the Ussurian secondary refugial center with a remarkable expansion. However, it is possible that the species will turn out as a polycentric Manchurian- Sinopacific faunal element, if the gap between the northern records and the
<typeStatus id="54CE88EEFF9579041920FF5A760962AB" box="[1055,1106,152,178]" pageId="11" pageNumber="51">type</typeStatus>
locality in the south can be corroborated.
</paragraph>
</subSubSection>
<caption id="DF0A66C4FF9579041DA8FB0370486721" httpUri="https://zenodo.org/record/196513/files/figure.png" pageId="11" pageNumber="51" targetBox="[162,1421,275,1190]" targetPageId="11">
<paragraph id="8BCA364CFF9579041DA8FB0370486721" blockId="11.[151,1436,1217,1336]" pageId="11" pageNumber="51">
<emphasis id="B901EA5EFF9579041DA8FB03730C66C0" bold="true" box="[151,343,1217,1241]" pageId="11" pageNumber="51">FIGURES 3843</emphasis>
. Geographic variation of the male genitalia of
<taxonomicName id="4C754DCFFF9579041E6FFB0071E966C0" box="[848,946,1218,1241]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9579041E6FFB0071E966C0" box="[848,946,1218,1241]" italics="true" pageId="11" pageNumber="51">I. fujiana</emphasis>
</taxonomicName>
. 38. Genital segments, lateral view; 39. Male genital segments, caudal view; 40. Gonarcus (gonocoxites 11), dorsal view; 41. Genital segments, lateral view; 42. Male genital segments, caudal view; 43. Gonarcus (gonocoxites 11), dorsal view. 3840: male from Beijing; 4143: male from Shaanxi Prov. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C36F65C7FF9579031DFAFAA577C966A4" lastPageId="12" lastPageNumber="52" pageId="11" pageNumber="51" type="discussion">
<paragraph id="8BCA364CFF9579041DFAFAA5764B64C8" blockId="11.[151,1437,1383,1932]" pageId="11" pageNumber="51">
<emphasis id="B901EA5EFF9579041DFAFAA5731E6798" bold="true" box="[197,325,1383,1409]" pageId="11" pageNumber="51">Remarks.</emphasis>
The species was originally described by
<bibRefCitation id="EFE44BBDFF9579041E69FAA571A86798" author="Yang" box="[854,1011,1383,1409]" pageId="11" pageNumber="54" refString="Yang, C. K. (1999) Raphidioptera: Family Inocelliidae. In: Huang, B. K. (ed). Fauna of insects in Fujian Province of China, Volume 3. Fujian Science Press, Fuzhou, pp. 177 - 180." type="book" year="1999">Yang (1999)</bibRefCitation>
based on a single male specimen collected from Fujian Province in the southeastern part of
<collectingCountry id="F36276DCFF9579041E76FA4E71C867BF" box="[841,915,1420,1446]" name="China" pageId="11" pageNumber="51">China</collectingCountry>
. Unfortunately, the genitalia of the
<typeStatus id="54CE88EEFF957904180AFA4E77C767BF" box="[1333,1436,1420,1446]" pageId="11" pageNumber="51" type="holotype">holotype</typeStatus>
are missing, at present. However, based on the original illustration of the male genitalia, the identity of this species can be validly determined by the remarkable gonarcus having a bifurcate process. In the collection of CAU, we found a great number of
<taxonomicName id="4C754DCFFF9579041F02FA3E70F3640C" box="[573,680,1532,1557]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B901EA5EFF9579041F02FA3E70F3640C" box="[573,680,1532,1557]" italics="true" pageId="11" pageNumber="51">Inocellia</emphasis>
</taxonomicName>
specimens collected from several localities of northern
<collectingCountry id="F36276DCFF9579041872FA3E77C3640F" box="[1357,1432,1532,1558]" name="China" pageId="11" pageNumber="51">China</collectingCountry>
, which possess male genitalia quite similar to that of the
<typeStatus id="54CE88EEFF9579041E11F9E071C36425" box="[814,920,1570,1596]" pageId="11" pageNumber="51" type="holotype">holotype</typeStatus>
, so that we identified these specimens to be
<taxonomicName id="4C754DCFFF9579041DA8F98573596479" box="[151,258,1607,1632]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9579041DA8F98573596479" box="[151,258,1607,1632]" italics="true" pageId="11" pageNumber="51">I. fujiana</emphasis>
</taxonomicName>
and provide a redescription of the male, as well as the first description of the female. The geographic variation of the male genitalia is also documented (
<figureCitation id="134E2AC9FF9579041FC2F9AE71CA649F" box="[765,913,1644,1670]" captionStart="FIGURES 38 43" captionStartId="11.[151,270,1217,1241]" captionTargetBox="[162,1421,275,1190]" captionTargetId="figure@11.[155,1433,269,1196]" captionTargetPageId="11" captionText="FIGURES 38 43. Geographic variation of the male genitalia of I. fujiana. 38. Genital segments, lateral view; 39. Male genital segments, caudal view; 40. Gonarcus (gonocoxites 11), dorsal view; 41. Genital segments, lateral view; 42. Male genital segments, caudal view; 43. Gonarcus (gonocoxites 11), dorsal view. 38 40: male from Beijing; 41 43: male from Shaanxi Prov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196513/files/figure.png" pageId="11" pageNumber="51">Figs. 3842</figureCitation>
). Thus,
<taxonomicName id="4C754DCFFF9579041ECEF9AE7604649C" box="[1009,1119,1644,1669]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9579041ECEF9AE7604649C" box="[1009,1119,1644,1669]" italics="true" pageId="11" pageNumber="51">I. fujiana</emphasis>
</taxonomicName>
is herein recognized to be widely distributed from southeastern
<collectingCountry id="F36276DCFF9579041F63F95070FF64B5" box="[604,676,1682,1708]" name="China" pageId="11" pageNumber="51">China</collectingCountry>
to northern
<collectingCountry id="F36276DCFF9579041E05F95071DE64B5" box="[826,901,1682,1708]" name="China" pageId="11" pageNumber="51">China</collectingCountry>
, while it might be quite rare in southeastern
<collectingCountry id="F36276DCFF9579041DA8F975728664C8" box="[151,221,1719,1745]" name="China" pageId="11" pageNumber="51">China</collectingCountry>
because no specimens other than the
<typeStatus id="54CE88EEFF9579041FA8F97570A664C8" box="[663,765,1719,1745]" pageId="11" pageNumber="51" type="holotype">holotype</typeStatus>
have been found there.
</paragraph>
<paragraph id="8BCA364CFF9579041DF2F91E73536595" blockId="11.[151,1437,1383,1932]" pageId="11" pageNumber="51">
Regarding the male genitalic structures,
<taxonomicName id="4C754DCFFF9579041F8AF91E717F64EC" box="[693,804,1756,1781]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9579041F8AF91E717F64EC" box="[693,804,1756,1781]" italics="true" pageId="11" pageNumber="51">I. fujiana</emphasis>
</taxonomicName>
appears to be closely related to a recently described species,
<emphasis id="B901EA5EFF9579041DC5F8C073CB6502" box="[250,400,1794,1819]" italics="true" pageId="11" pageNumber="51">
<taxonomicName id="4C754DCFFF9579041DC5F8C073D76502" box="[250,396,1794,1819]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="shinohara">I. shinohara</taxonomicName>
,
</emphasis>
due to the similarity of the gonarcus having a bifurcate process.
<taxonomicName id="4C754DCFFF95790419B8F8C077126502" box="[1159,1353,1794,1819]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF95790419B8F8C077126502" box="[1159,1353,1794,1819]" italics="true" pageId="11" pageNumber="51">Inocellia fujiana</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4C754DCFFF9579041C40F8E5704B6559" box="[383,528,1831,1856]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="shinohara">
<emphasis id="B901EA5EFF9579041C40F8E5704B6559" box="[383,528,1831,1856]" italics="true" pageId="11" pageNumber="51">I. shinohara</emphasis>
</taxonomicName>
by the feebly prominent styli of the male ninth gonocoxite. In
<taxonomicName id="4C754DCFFF957904183CF8E577CE6559" box="[1283,1429,1831,1856]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="11" pageNumber="51" phylum="Arthropoda" rank="species" species="shinohara">
<emphasis id="B901EA5EFF957904183CF8E577CE6559" box="[1283,1429,1831,1856]" italics="true" pageId="11" pageNumber="51">I. shinohara</emphasis>
</taxonomicName>
, the male ninth gonocoxite possesses a pair of long and acutely pointed styli on its inner portions (U. Aspöck
<emphasis id="B901EA5EFF95790418B8F88E72F56592" italics="true" pageId="11" pageNumber="51">et al</emphasis>
. 2009).
</paragraph>
<paragraph id="8BCA364CFF9279031DA8FBA677C966A4" blockId="12.[151,1437,1122,1213]" pageId="12" pageNumber="52">
<figureCitation id="134E2AC9FF9279031DA8FBA6737D6665" box="[151,294,1124,1148]" captionStart="FIGURES 38 43" captionStartId="11.[151,270,1217,1241]" captionTargetBox="[162,1421,275,1190]" captionTargetId="figure@11.[155,1433,269,1196]" captionTargetPageId="11" captionText="FIGURES 38 43. Geographic variation of the male genitalia of I. fujiana. 38. Genital segments, lateral view; 39. Male genital segments, caudal view; 40. Gonarcus (gonocoxites 11), dorsal view; 41. Genital segments, lateral view; 42. Male genital segments, caudal view; 43. Gonarcus (gonocoxites 11), dorsal view. 38 40: male from Beijing; 41 43: male from Shaanxi Prov. Scale bars = 0.5 mm." httpUri="https://zenodo.org/record/196513/files/figure.png" pageId="12" pageNumber="52">
<emphasis id="B901EA5EFF9279031DA8FBA6737D6665" bold="true" box="[151,294,1124,1148]" pageId="12" pageNumber="52">FIGURE 43</emphasis>
</figureCitation>
. Distribution map of the species of
<taxonomicName id="4C754DCFFF9279031F83FBA771066665" box="[700,861,1125,1148]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B901EA5EFF9279031F83FBA771066665" box="[700,861,1125,1148]" italics="true" pageId="12" pageNumber="52">I. crassicornis</emphasis>
</taxonomicName>
group from
<collectingCountry id="F36276DCFF9279031ED3FBA776696665" box="[1004,1074,1125,1148]" name="China" pageId="12" pageNumber="52">China</collectingCountry>
. Ǻ:
<taxonomicName id="4C754DCFFF9279031959FBA7775A6665" box="[1126,1281,1125,1148]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="biprocessus" status="sp. nov.">
<emphasis id="B901EA5EFF9279031959FBA7775A6665" box="[1126,1281,1125,1148]" italics="true" pageId="12" pageNumber="52">I. biprocessus</emphasis>
</taxonomicName>
<emphasis id="B901EA5EFF9279031836FBA677386665" bold="true" box="[1289,1379,1124,1148]" pageId="12" pageNumber="52">
<taxonomicNameLabel id="A2325725FF9279031836FBA677386665" box="[1289,1379,1124,1148]" pageId="12" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;●:
<taxonomicName id="4C754DCFFF92790318B3FBA773F06684" authority="Schummel" authorityName="Schummel" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B901EA5EFF92790318B3FBA773476684" italics="true" pageId="12" pageNumber="52">I. crassicornis</emphasis>
(Schummel)
</taxonomicName>
; ˔:
<taxonomicName id="4C754DCFFF9279031CDEFB44702D6684" box="[481,630,1158,1181]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="digitiformis" status="sp. nov.">
<emphasis id="B901EA5EFF9279031CDEFB44702D6684" box="[481,630,1158,1181]" italics="true" pageId="12" pageNumber="52">I. digitiformis</emphasis>
</taxonomicName>
<emphasis id="B901EA5EFF9279031F41FB47708E6684" bold="true" box="[638,725,1157,1181]" pageId="12" pageNumber="52">
<taxonomicNameLabel id="A2325725FF9279031F41FB47708E6684" box="[638,725,1157,1181]" pageId="12" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
;
<emphasis id="B901EA5EFF9279031FDBFB4770A86684" bold="true" box="[740,755,1157,1181]" pageId="12" pageNumber="52">+</emphasis>
:
<taxonomicName id="4C754DCFFF9279031E3CFB4471F26684" authority="Yang" authorityName="Yang" box="[771,937,1158,1181]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="fujiana">
<emphasis id="B901EA5EFF9279031E3CFB44713D6684" box="[771,870,1158,1181]" italics="true" pageId="12" pageNumber="52">I. fujiana</emphasis>
Yang
</taxonomicName>
; Ο:
<taxonomicName id="4C754DCFFF9279031EEBFB44773A6684" authority="Aspock, Liu &amp; Aspock" authorityName="Aspock, Liu &amp; Aspock" box="[980,1377,1158,1181]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="shinohara">
<emphasis id="B901EA5EFF9279031EEBFB4476016684" box="[980,1114,1158,1181]" italics="true" pageId="12" pageNumber="52">I. shinohara</emphasis>
Aspöck, Liu &amp; Aspöck
</taxonomicName>
; :
<taxonomicName id="4C754DCFFF92790318B3FB44704766A4" authority="Aspock &amp; Aspock. Inserted" authorityName="Aspock &amp; Aspock. Inserted" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="taiwana">
<emphasis id="B901EA5EFF92790318B3FB4472B666A4" italics="true" pageId="12" pageNumber="52">I. taiwana</emphasis>
Aspöck &amp; Aspöck. Inserted
</taxonomicName>
map: Distribution of
<taxonomicName id="4C754DCFFF9279031E3DFB6471C166A4" box="[770,922,1190,1213]" class="Insecta" family="Inocelliidae" genus="Inocellia" kingdom="Animalia" order="Raphidioptera" pageId="12" pageNumber="52" phylum="Arthropoda" rank="species" species="crassicornis">
<emphasis id="B901EA5EFF9279031E3DFB6471C166A4" box="[770,922,1190,1213]" italics="true" pageId="12" pageNumber="52">I. crassicornis</emphasis>
</taxonomicName>
modified after H.
<bibRefCitation id="EFE44BBDFF927903195EFB64777566A4" author="Aspock" box="[1121,1326,1190,1213]" pageId="12" pageNumber="53" refString="Aspock, H., Aspock, U. &amp; Rausch, H. (1991) Die Raphidiopteren der Erde. Eine monographische Darstellung der Systematik, Taxonomie, Biologie, Okologie und Chorologie der rezenten Raphidiopteren der Erde, mit einer zusammenfassenden Ubersicht der fossilen Raphidiopteren (Insecta: Neuropteroidea). Goecke &amp; Evers, Krefeld, 2 volumes, 730 pp, 550 pp." type="book" year="1991">
Aspöck
<emphasis id="B901EA5EFF9279031986FB6476B366A4" box="[1209,1256,1190,1213]" italics="true" pageId="12" pageNumber="52">et al</emphasis>
. 1991
</bibRefCitation>
: fig. 184.
</paragraph>
</subSubSection>
</treatment>
</document>