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<document id="E8D432E9E1B19C57C7D9C92CF51D5DB8" ID-CLB-Dataset="46739" ID-DOI="10.5281/zenodo.277769" ID-GBIF-Dataset="59773ab9-2c22-4291-ab1d-ab419ec7b0b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="277769" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460364746379" checkinUser="plazi" docAuthor="Ziani, Stefano" docDate="2011" docId="03DC8232D207FFD749F6FF322FD532B2" docLanguage="en" docName="zt02907p050.pdf" docOrigin="Zootaxa 2907" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Onthophagus semicornis" docType="key" docVersion="9" lastPageNumber="50" masterDocId="FFE5FA4AD204FFD44961FFA52A493606" masterDocTitle="A new Iranian Onthophagus species belonging to the O. semicornis group (Coleoptera: Scarabaeidae: Onthophagini)" masterLastPageNumber="50" masterPageNumber="47" pageNumber="50" updateTime="1698295045633" updateUser="ExternalLinkService">
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<mods:title id="088B86CE5CF79521D0973F34242C9838">A new Iranian Onthophagus species belonging to the O. semicornis group (Coleoptera: Scarabaeidae: Onthophagini)</mods:title>
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<mods:namePart id="B4F90EF69127D48DDCABCCF01CA13E4B">Ziani, Stefano</mods:namePart>
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<mods:date id="5E2F2A0464CE15E9BE250ED72C9C4D2C">2011</mods:date>
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<treatment id="03DC8232D207FFD749F6FF322FD532B2" ID-DOI="http://doi.org/10.5281/zenodo.6193256" ID-GBIF-Taxon="119410537" ID-Zenodo-Dep="6193256" LSID="urn:lsid:plazi:treatment:03DC8232D207FFD749F6FF322FD532B2" httpUri="http://treatment.plazi.org/id/03DC8232D207FFD749F6FF322FD532B2" lastPageNumber="50" pageId="3" pageNumber="50">
<subSubSection id="C36F60AFD207FFD749F6FF322E0736A9" box="[151,1102,151,176]" pageId="3" pageNumber="50" type="nomenclature">
<paragraph id="8BCA3324D207FFD749F6FF322E0736A9" blockId="3.[151,1102,151,176]" box="[151,1102,151,176]" pageId="3" pageNumber="50">
<heading id="D0828448D207FFD749F6FF322E0736A9" bold="true" box="[151,1102,151,176]" fontSize="10" level="2" pageId="3" pageNumber="50" reason="2">
<emphasis id="B901EF36D207FFD749F6FF322E0736A9" bold="true" box="[151,1102,151,176]" pageId="3" pageNumber="50">
Key to the species in the
<taxonomicName id="4C7548A7D207FFD748CEFF3D280F36B6" ID-CoL="49MQ4" box="[431,582,152,176]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="semicornis">
<emphasis id="B901EF36D207FFD748CEFF3D280F36B6" bold="true" box="[431,582,152,176]" italics="true" pageId="3" pageNumber="50">O. semicornis</emphasis>
</taxonomicName>
group modified from
<bibRefCitation id="EFE44ED5D207FFD74A21FF322E0736A9" author="Ziani" box="[832,1102,151,175]" pageId="3" pageNumber="50" refString="Ziani, S. &amp; Gudenzi, I. (2007) Studies on Palaearctic Onthophagus associated with burrows of small mammals. II. O. semicornis and closely related species (Coleoptera: Scarabaeidae: Onthophagini). Folia Heyrovskyana, 15 (2), 89 - 114." type="journal article" year="2007">Ziani &amp; Gudenzi (2007)</bibRefCitation>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36F60AFD207FFD749F6FF782FD532B2" pageId="3" pageNumber="50" type="key">
<keyStep id="30812B81D207FFD749F6FF782FD53752" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FF782FD23712" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FF782FD23712" pageId="3" pageNumber="50">1. Clypeal margin more-or-less shallowly emarginate medially; pronotal punctures on disc separated by 0.52.0 diameters .............................................................................................................................................................................. 2</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FEB82FD53752" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FEB82FD53752" pageId="3" pageNumber="50">
-. Clypeal margin arcuately excised medially; pronotal punctures on disc separated by 24 diameters.
<collectingCountry id="F36273B4D207FFD74C78FEB82FD13732" box="[1305,1432,285,308]" name="Kazakhstan" pageId="3" pageNumber="50">Kazakhstan</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD749A6FE982B0B3752" box="[199,322,317,340]" name="Uzbekistan" pageId="3" pageNumber="50">Uzbekistan</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD7482DFE982B8F3752" box="[332,454,317,340]" name="Kyrgyzstan" pageId="3" pageNumber="50">Kyrgyzstan</collectingCountry>
<emphasis id="B901EF36D207FFD748AAFE982ED33752" box="[459,1178,317,340]" italics="true" pageId="3" pageNumber="50">............................................................................................................</emphasis>
<taxonomicName id="4C7548A7D207FFD74DFCFE982FD53752" ID-CoL="49MQ7" authority="Balthasar, 1960" authorityName="Balthasar" authorityYear="1960" box="[1181,1436,317,340]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="silus">
<emphasis id="B901EF36D207FFD74DFCFE982EA63752" box="[1181,1263,317,340]" italics="true" pageId="3" pageNumber="50">O. silus</emphasis>
Balthasar, 1960
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FEF82FD23792" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FEF82FD23772" blockId="3.[151,1436,221,1204]" box="[151,1435,349,372]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FEF82FD23772" box="[151,1435,349,372]" pageId="3" pageNumber="50">2. Pronotum with two anteromedian tubercles, tubercles distinctly separated................................................................ 3</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FED82FD23792" blockId="3.[151,1436,221,1204]" box="[151,1435,381,404]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FED82FD23792" box="[151,1435,381,404]" pageId="3" pageNumber="50">-. Pronotum with an anteromedian prominence, prominence sometimes appearing bilobate ......................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FE382FD237D2" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FE382FD537B2" blockId="3.[151,1436,221,1204]" box="[151,1436,413,436]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FE382FD537B2" box="[151,1436,413,436]" pageId="3" pageNumber="50">
3. Pronotal punctures simple, large, umbilicate on disc.
<collectingCountry id="F36273B4D207FFD74B9EFE3829CB37B2" box="[767,898,413,436]" name="Afghanistan" pageId="3" pageNumber="50">Afghanistan</collectingCountry>
.............................
<taxonomicName id="4C7548A7D207FFD74D2DFE382FD537B2" ID-CoL="49MPB" authority="Kabakov, 1982" authorityName="Kabakov" authorityYear="1982" box="[1100,1436,413,436]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="nurestanicus">
<emphasis id="B901EF36D207FFD74D2DFE382EBD37B2" box="[1100,1268,413,436]" italics="true" pageId="3" pageNumber="50">O. nurestanicus</emphasis>
Kabakov, 1982
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FE182FD237D2" blockId="3.[151,1436,221,1204]" box="[151,1435,445,468]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FE182FD237D2" box="[151,1435,445,468]" pageId="3" pageNumber="50">-. Pronotal punctures with a small granule at anterior margin ........................................................................................ 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FE782FD53472" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FE782FD53412" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FE782FD53412" pageId="3" pageNumber="50">
4. Superior edge of occipital carina straight in dorsal view, with no trace of lateral horns; occipital carina broader basally than the clypeofrontal carina.
<collectingCountry id="F36273B4D207FFD74B39FE5828A23412" box="[600,747,509,532]" name="Turkmenistan" pageId="3" pageNumber="50">Turkmenistan</collectingCountry>
<emphasis id="B901EF36D207FFD74B90FE582E233412" box="[753,1130,509,532]" italics="true" pageId="3" pageNumber="50">........................................................</emphasis>
<taxonomicName id="4C7548A7D207FFD74D08FE582FD53412" ID-CoL="49MPY" authority="Kabakov, 1982" authorityName="Kabakov" authorityYear="1982" box="[1129,1436,509,532]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="rufimanus">
<emphasis id="B901EF36D207FFD74D08FE582EBD3412" box="[1129,1268,509,532]" italics="true" pageId="3" pageNumber="50">O.rufimanus</emphasis>
Kabakov, 1982
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FDB82FD53472" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FDB82FD53472" pageId="3" pageNumber="50">
-. Superior edge of occipital carina sinuate in dorsal view, weakly dentate at middle and/or with a lateral horn more-orless erect on each side; occipital carina basally at least as broad as the clypeofrontal carina. Eastern
<collectingCountry id="F36273B4D207FFD74D8CFD982F7E3452" box="[1261,1335,573,596]" name="Russia" pageId="3" pageNumber="50">Russia</collectingCountry>
, northern and central
<collectingCountry id="F36273B4D207FFD74822FDF82BCC3472" box="[323,389,605,628]" name="China" pageId="3" pageNumber="50">China</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD748F1FDF82BB33472" box="[400,506,605,628]" name="Mongolia" pageId="3" pageNumber="50">Mongolia</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD74B64FDF828003472" box="[517,585,605,628]" name="South Korea" pageId="3" pageNumber="50">Korea</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD74B35FDF828D83472" box="[596,657,605,628]" name="Japan" pageId="3" pageNumber="50">Japan</collectingCountry>
<emphasis id="B901EF36D207FFD74BF2FDF82E0D3472" box="[659,1092,605,628]" italics="true" pageId="3" pageNumber="50">.................................................................</emphasis>
<taxonomicName id="4C7548A7D207FFD74D2AFDF82FD53472" ID-CoL="49MPD" authority="Boucomont, 1924" authorityName="Boucomont" authorityYear="1924" box="[1099,1436,605,628]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="olsoufieffi">
<emphasis id="B901EF36D207FFD74D2AFDF82E9F3472" box="[1099,1238,605,628]" italics="true" pageId="3" pageNumber="50">O. olsoufieffi</emphasis>
Boucomont, 1924
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FDD82FDC34D2" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FDD82FD534B2" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FDD82FD534B2" pageId="3" pageNumber="50">
5. Pronotum doubly sinuate laterally; occipital carina with a high, slender, bifurcate horn; anteromedian pronotal prominence slender, prolonged to top of occipital horn.
<collectingCountry id="F36273B4D207FFD74A78FD3829EA34B2" box="[793,931,669,692]" name="South Korea" pageId="3" pageNumber="50">South Korea</collectingCountry>
.
<emphasis id="B901EF36D207FFD74ACAFD382EC734B2" box="[939,1166,669,692]" italics="true" pageId="3" pageNumber="50">..................................</emphasis>
<taxonomicName id="4C7548A7D207FFD74DF0FD382FD534B2" ID-CoL="49MNR" authority="Kim, 1985" authorityName="Kim" authorityYear="1985" box="[1169,1436,669,692]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="koryoensis">
<emphasis id="B901EF36D207FFD74DF0FD382F6A34B2" box="[1169,1315,669,692]" italics="true" pageId="3" pageNumber="50">O. koryoensis</emphasis>
Kim, 1985
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FD182FDC34D2" blockId="3.[151,1436,221,1204]" box="[151,1429,701,724]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FD182FDC34D2" box="[151,1429,701,724]" pageId="3" pageNumber="50">-. Sides of pronotum only distinctly sinuate behind anterior angles ............................................................................... 6</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FD782FD23552" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FD782FD23512" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FD782FD23512" pageId="3" pageNumber="50">6. Occipital carina with an erect, terminally narrow crest; base of carina distinctly not extending from side to side of head ............................................................................................................................................................................... 7</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FCB82FD23552" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FCB82FD23552" pageId="3" pageNumber="50">-. Occipital carina with a simple, more-or-less erect lamina; carina base extended, or nearly extended, from side to side of head.......................................................................................................................................................................... 8</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FCF82FD235F2" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FCF82FD535B2" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FCF82FD535B2" pageId="3" pageNumber="50">
7. Anteromedian pronotal prominence broad, transverse, more than twice as wide as long, only slightly more projecting forward than the anterolateral tubercles (
<figureCitation id="134E2FA1D207FFD74BC6FCD828A13592" box="[679,744,893,916]" captionStart="FIGURES 7 10" captionStartId="2.[151,269,1005,1029]" captionTargetBox="[176,1419,219,967]" captionTargetId="figure@2.[152,1434,194,981]" captionTargetPageId="2" captionText="FIGURES 7 10. Onthophagus “ semicornis lineage ”, head and pronotum of male in dorsal view. 7: Onthophagus samai n. sp., holotype; 8: Onthophagus semicornis (Panzer, 1798) (Italy, Abruzzo Province); 9: Onthophagus eulaminicornis Pittino, 2006 (Greece, Thrace); 10: Onthophagus sacharovskii Olsoufieff, 1918 (Turkey, Kayseri Province). Photographs by A. Degiovanni." httpUri="https://zenodo.org/record/277770/files/figure.png" pageId="3" pageNumber="50">Fig. 8</figureCitation>
). Dorsal surface shiny. Central and southern Europe;
<collectingCountry id="F36273B4D207FFD74C4AFCD82ABD35B2" name="Turkmenistan" pageId="3" pageNumber="50">Turkmenistan</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD7499EFC382B3735B2" box="[255,382,925,948]" name="Kazakhstan" pageId="3" pageNumber="50">Kazakhstan</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD748E8FC38284D35B2" box="[393,516,925,948]" name="Uzbekistan" pageId="3" pageNumber="50">Uzbekistan</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD74B6FFC38283F35B2" box="[526,630,925,948]" name="Mongolia" pageId="3" pageNumber="50">Mongolia</collectingCountry>
........................................................................
<taxonomicName id="4C7548A7D207FFD74D04FC382FD535B2" ID-CoL="49MQ4" authority="Panzer, 1798" authorityName="Panzer" authorityYear="1798" box="[1125,1436,925,948]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="semicornis">
<emphasis id="B901EF36D207FFD74D04FC382EB035B2" box="[1125,1273,925,948]" italics="true" pageId="3" pageNumber="50">O. semicornis</emphasis>
(Panzer, 1798)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FC182FD235F2" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FC182FD235F2" pageId="3" pageNumber="50">
-. Anteromedian pronotal prominence less than twice wider than long, clearly more projecting forward than anterolateral tubercles (
<figureCitation id="134E2FA1D207FFD74805FC782BEC35F2" box="[356,421,989,1012]" captionStart="FIGURES 7 10" captionStartId="2.[151,269,1005,1029]" captionTargetBox="[176,1419,219,967]" captionTargetId="figure@2.[152,1434,194,981]" captionTargetPageId="2" captionText="FIGURES 7 10. Onthophagus “ semicornis lineage ”, head and pronotum of male in dorsal view. 7: Onthophagus samai n. sp., holotype; 8: Onthophagus semicornis (Panzer, 1798) (Italy, Abruzzo Province); 9: Onthophagus eulaminicornis Pittino, 2006 (Greece, Thrace); 10: Onthophagus sacharovskii Olsoufieff, 1918 (Turkey, Kayseri Province). Photographs by A. Degiovanni." httpUri="https://zenodo.org/record/277770/files/figure.png" pageId="3" pageNumber="50">Fig. 7</figureCitation>
). Dorsal surface dull. Northeastern
<collectingCountry id="F36273B4D207FFD74A43FC78290435F2" box="[802,845,989,1012]" name="Iran" pageId="3" pageNumber="50">Iran</collectingCountry>
<emphasis id="B901EF36D207FFD74A2FFC782F1335F2" box="[846,1370,989,1012]" italics="true" pageId="3" pageNumber="50">
................................................................
<taxonomicName id="4C7548A7D207FFD74D9FFC782F1335F2" ID-CoL="49MQ2" box="[1278,1370,989,1012]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="samai" status="sp. nov.">O. samai</taxonomicName>
</emphasis>
<taxonomicNameLabel id="A232524DD207FFD74C01FC792FD235F2" box="[1376,1435,988,1012]" pageId="3" pageNumber="50" rank="species">
<emphasis id="B901EF36D207FFD74C01FC792FD235F2" bold="true" box="[1376,1435,988,1012]" pageId="3" pageNumber="50">n. sp.</emphasis>
</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30812B81D207FFD749F6FC582FD532B2" pageId="3" pageNumber="50">
<paragraph id="8BCA3324D207FFD749F6FC582FD53232" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FC582FD53232" pageId="3" pageNumber="50">
8. Occipital carina erected in a rectangular lamina with vertical sides, straight in dorsal view, extending at base to inner edge of eyes (
<figureCitation id="134E2FA1D207FFD7483BFBB82BD53232" box="[346,412,1053,1076]" captionStart="FIGURES 7 10" captionStartId="2.[151,269,1005,1029]" captionTargetBox="[176,1419,219,967]" captionTargetId="figure@2.[152,1434,194,981]" captionTargetPageId="2" captionText="FIGURES 7 10. Onthophagus “ semicornis lineage ”, head and pronotum of male in dorsal view. 7: Onthophagus samai n. sp., holotype; 8: Onthophagus semicornis (Panzer, 1798) (Italy, Abruzzo Province); 9: Onthophagus eulaminicornis Pittino, 2006 (Greece, Thrace); 10: Onthophagus sacharovskii Olsoufieff, 1918 (Turkey, Kayseri Province). Photographs by A. Degiovanni." httpUri="https://zenodo.org/record/277770/files/figure.png" pageId="3" pageNumber="50">Fig. 9</figureCitation>
).
<collectingCountry id="F36273B4D207FFD748D1FBB82BB73232" box="[432,510,1053,1076]" name="Greece" pageId="3" pageNumber="50">Greece</collectingCountry>
,
<collectingCountry id="F36273B4D207FFD74B69FBB8282D3232" box="[520,612,1053,1076]" name="Bulgaria" pageId="3" pageNumber="50">Bulgaria</collectingCountry>
<emphasis id="B901EF36D207FFD74B05FBB82E0D3232" box="[612,1092,1053,1076]" italics="true" pageId="3" pageNumber="50">........................................................................</emphasis>
<taxonomicName id="4C7548A7D207FFD74D2AFBB82FD53232" ID-CoL="49MN3" authority="Pittino, 2006" authorityName="Pittino" authorityYear="2006" box="[1099,1436,1053,1076]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="eulaminicornis">
<emphasis id="B901EF36D207FFD74D2AFBB82F423232" box="[1099,1291,1053,1076]" italics="true" pageId="3" pageNumber="50">O. eulaminicornis</emphasis>
Pittino, 2006
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BCA3324D207FFD749F6FB982FD532B2" blockId="3.[151,1436,221,1204]" pageId="3" pageNumber="50">
<keyLead id="30849011D207FFD749F6FB982FD532B2" pageId="3" pageNumber="50">
-. Occipital carina weakly elevated, sometimes raised at edges, sinuate in dorsal view, distinctly bent back laterally, clearly twisted over each lateral extremity and visibly turning out and backwards, extending at base almost to outer edge of eyes (
<figureCitation id="134E2FA1D207FFD7483BFBD82BE23292" box="[346,427,1149,1172]" captionStart="FIGURES 7 10" captionStartId="2.[151,269,1005,1029]" captionTargetBox="[176,1419,219,967]" captionTargetId="figure@2.[152,1434,194,981]" captionTargetPageId="2" captionText="FIGURES 7 10. Onthophagus “ semicornis lineage ”, head and pronotum of male in dorsal view. 7: Onthophagus samai n. sp., holotype; 8: Onthophagus semicornis (Panzer, 1798) (Italy, Abruzzo Province); 9: Onthophagus eulaminicornis Pittino, 2006 (Greece, Thrace); 10: Onthophagus sacharovskii Olsoufieff, 1918 (Turkey, Kayseri Province). Photographs by A. Degiovanni." httpUri="https://zenodo.org/record/277770/files/figure.png" pageId="3" pageNumber="50">Fig. 10</figureCitation>
).
<collectingCountry id="F36273B4D207FFD748DCFBD828423292" box="[445,523,1149,1172]" name="Turkey" pageId="3" pageNumber="50">Turkey</collectingCountry>
, Caucasus, southern European
<collectingCountry id="F36273B4D207FFD74A37FBD829E83292" box="[854,929,1149,1172]" name="Russia" pageId="3" pageNumber="50">Russia</collectingCountry>
; northwestern
<collectingCountry id="F36273B4D207FFD74D5CFBD82E213292" box="[1085,1128,1149,1172]" name="Iran" pageId="3" pageNumber="50">Iran</collectingCountry>
............................................. ...................................................................................................................................
<taxonomicName id="4C7548A7D207FFD74D5EFB382FD532B2" ID-CoL="49MPZ" authority="Olsoufieff, 1918" authorityName="Olsoufieff" authorityYear="1918" box="[1087,1436,1181,1204]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="50" phylum="Arthropoda" rank="species" species="sacharovskii">
<emphasis id="B901EF36D207FFD74D5EFB382EAC32B2" box="[1087,1253,1181,1204]" italics="true" pageId="3" pageNumber="50">O. sacharovskii</emphasis>
Olsoufieff, 1918
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>