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<document id="CED6464D122FD47F652290CC3540FC04" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629678646461" checkinUser="felipe" docAuthor="Melika, George, Pujade-Villar, Juli, Abe, Yoshihisa, Tang, Chang-Ti, Nicholls, James, Wachi, Nakatada, Ide, Tatsuya, Yang, Man-Miao, Pénzes, Zsolt, Csóka, György &amp; Stone, Graham N." docDate="2010" docId="03DC095CFFD0330ECAD8E0E683BAFA3B" docLanguage="en" docName="zt02470p079.pdf" docOrigin="Zootaxa 2470" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Trichagalma formosana , Melika &amp; Tang 2010, new species" docType="treatment" docVersion="2" lastPageNumber="13" masterDocId="FFE57124FFDA3302CA4FE53C8719FF89" masterDocTitle="2470" masterLastPageNumber="79" masterPageNumber="1" pageNumber="11" updateTime="1699094741932" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="BE5A2E67E197FF3BC2829202472DD52E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="BB816041B010AE1545DD224CE02AF30C">
<mods:title id="7BB89E7C763FBEC146318C613241D298">2470</mods:title>
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<mods:namePart id="EDC337D1FB1E770DF5C21F897D700988">Melika, George</mods:namePart>
</mods:name>
<mods:name id="CDC535C69B88AC922D838DF3DEDAEA9F" type="personal">
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<mods:namePart id="D7489F22944467F76D88E0FCB3DBE646">Pujade-Villar, Juli</mods:namePart>
</mods:name>
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</mods:role>
<mods:namePart id="26C2D6550E731FB6212BF04E0FCD1F6F">Abe, Yoshihisa</mods:namePart>
</mods:name>
<mods:name id="9B2973FFFC9205CFB11304EE84457C3D" type="personal">
<mods:role id="2F414A97447C13DB9355FF6593A722E8">
<mods:roleTerm id="E0455CFAECCBC84B089BB4AFE2BA82E8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2FD5603EECFFE3687BCDD38A7380AEB7">Tang, Chang-Ti</mods:namePart>
</mods:name>
<mods:name id="9031D3DED93C7643BC04B5185FFCA50A" type="personal">
<mods:role id="F31BE0261EEC35EFAE94DA57CAF96D24">
<mods:roleTerm id="FEE0F88529E3BBBB6B31EE38FE2265FA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="945C75E262E31B210FAE7E9899D2AC1D">Nicholls, James</mods:namePart>
</mods:name>
<mods:name id="1FFC1B1F798DBE901BF6005C10634694" type="personal">
<mods:role id="A70CBC60D527CC5AD2331D6563F1DF1E">
<mods:roleTerm id="A1D80FC858CC6D51EA40BD3F42C9268A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7A7404B9E0DD2EF9E2DBB39D2C04AA1B">Wachi, Nakatada</mods:namePart>
</mods:name>
<mods:name id="F71FDCCD00CC634C5E3A82193AFECB2A" type="personal">
<mods:role id="EEA3A81E91A6D07042F424C895B4BE71">
<mods:roleTerm id="6432B48692EA3AEBCD5F73903F2AF4B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8F81F5EFB889552458D63C5D3C6E12D8">Ide, Tatsuya</mods:namePart>
</mods:name>
<mods:name id="653F6BE3C23067506B88DC928FE1AB3C" type="personal">
<mods:role id="EB1F0EF1612F93087AC3CA688BBE3F5D">
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</mods:role>
<mods:namePart id="583C3EA95479D10239417C9CA640065C">Yang, Man-Miao</mods:namePart>
</mods:name>
<mods:name id="E464F1CCA2329B0FF9E97588A6A81FD1" type="personal">
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<mods:namePart id="4DBD39438552232F5385DD82D23439C0">Pénzes, Zsolt</mods:namePart>
</mods:name>
<mods:name id="0D3AEDE22CEDFCE73BA22BE3E078F5F3" type="personal">
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</mods:role>
<mods:namePart id="4782C79463E1AA973F8091FE60515762">Csóka, György</mods:namePart>
</mods:name>
<mods:name id="78639DF3E174AA704B1F864DCEF30326" type="personal">
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</mods:role>
<mods:namePart id="B27DBE31483B3687BB2D3422369E6014">Stone, Graham N.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0CBBCFC3322FE2D0D3DADADAA032BAD9">text</mods:typeOfResource>
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<mods:title id="151A18AD55D309C2254EBB8E74DD8D51">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3AA79B8FBA329A0F7846EFA70FC564D6">
<mods:date id="182D852A772D12987F283C3858A1B22B">2010</mods:date>
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<mods:number id="604D76DC1C95CE8D95164CC8CD48346C">2010-05-14</mods:number>
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<mods:number id="684BBE8E82FBAA4C5395D713BB3B3429">2470</mods:number>
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<mods:start id="8D986315CF00FBC70ED361A7C24A2AC7">1</mods:start>
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<mods:classification id="206AAD832BCEE2918CEC74EFBD1CB030">journal article</mods:classification>
<mods:identifier id="8DCD1D8DD1E95BDF17D0BF70840C9967" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="03DC095CFFD0330ECAD8E0E683BAFA3B" LSID="urn:lsid:plazi:treatment:03DC095CFFD0330ECAD8E0E683BAFA3B" httpUri="http://treatment.plazi.org/id/03DC095CFFD0330ECAD8E0E683BAFA3B" lastPageId="12" lastPageNumber="13" pageId="10" pageNumber="11">
<subSubSection id="C36FEBC1FFD03308CAD8E0E6843FFA7D" box="[151,806,1498,1524]" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BCAB84AFFD03308CAD8E0E6843FFA7D" blockId="10.[151,806,1498,1559]" box="[151,806,1498,1524]" pageId="10" pageNumber="11">
<heading id="D0820F26FFD03308CAD8E0E6843FFA7D" bold="true" box="[151,806,1498,1524]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B9016458FFD03308CAD8E0E6843FFA7D" bold="true" box="[151,806,1498,1524]" pageId="10" pageNumber="11">
<taxonomicName id="4C75C3C9FFD03308CAD8E0E68590FA7D" authority="Melika &amp; Tang" authorityName=", Melika &amp; Tang" authorityYear="2010" box="[151,649,1498,1524]" class="Insecta" family="Cynipidae" genus="Trichagalma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosana" status="sp. nov.">
<emphasis id="B9016458FFD03308CAD8E0E686A2FA7D" bold="true" box="[151,443,1498,1524]" italics="true" pageId="10" pageNumber="11">Trichagalma formosana</emphasis>
, Melika &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A232D923FFD03308C8D9E0E6843FFA7D" box="[662,806,1498,1524]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD03308CAD8E0C186C8F99E" box="[151,465,1533,1559]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BCAB84AFFD03308CAD8E0C186C8F99E" blockId="10.[151,806,1498,1559]" box="[151,465,1533,1559]" pageId="10" pageNumber="11">
<figureCitation id="134EA4CFFFD03308CAD8E0C18605F99E" box="[151,284,1533,1559]" captionStart="FIGURES 31–35" captionStartId="44.[151,269,1680,1704]" captionTargetBox="[151,1407,209,1619]" captionTargetId="figure-45@44.[151,1436,194,1656]" captionTargetPageId="44" captionText="FIGURES 31–35. Trichagalma formosana, new species, asexual female: 31–33, head: 31, anterior view, 32, posterior view, 33, dorsal view. 34, antenna, 35, forewing." pageId="10" pageNumber="11">Figs 31–35</figureCitation>
,
<figureCitation id="134EA4CFFFD03308CB66E0C1866CF99E" box="[297,373,1533,1559]" captionStart="FIGURES 36–42" captionStartId="45.[151,269,1888,1912]" captionTargetBox="[165,1418,213,1864]" captionTargetId="figure-60@45.[151,1436,194,1864]" captionTargetPageId="45" captionText="FIGURES 36–42. Trichagalma formosana, new species, asexual female: 36, mesosoma, dorsal view; 37, mesosoma, lateral view; 38, metascutellum and propodeum, posterodorsal view; 39, hind tarsus; 40, metasoma, lateral view; 41–42, ventral spine of hypopygium: 41, lateral view, 42, ventral view." pageId="10" pageNumber="11">36–42</figureCitation>
,
<figureCitation id="134EA4CFFFD03308CBCFE0C186D7F99E" box="[384,462,1533,1559]" captionStart="FIGURES 43–49" captionStartId="46.[151,269,1337,1361]" captionTargetBox="[151,1436,194,1313]" captionTargetId="figure-44@46.[151,1436,194,1313]" captionTargetPageId="46" captionText="FIGURES 43–49. Asexual galls of: 43–47, Trichagalma formosana, new species, 48–49, Trichagalma acutissimae (photos by Chang-Ti Tang and G. Csóka)." pageId="10" pageNumber="11">43–47</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD03308CAD8E37B86FFF8DB" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BCAB84AFFD03308CAD8E37B86FFF8DB" blockId="10.[151,1436,1607,1914]" pageId="10" pageNumber="11">
<emphasis id="B9016458FFD03308CAD8E37B8656F9E8" bold="true" box="[151,335,1607,1633]" pageId="10" pageNumber="11">Type material.</emphasis>
<materialsCitation id="3B1DB217FFD03308CB15E37484C5F903" collectingDate="2008-11-07" collectingDateMax="2008-12-07" collectingDateMin="2008-11-07" country="Taiwan" county="Cingjing Farm" location="Tang" municipality="Renai Township" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Nantou" typeStatus="holotype">
<typeStatus id="54CE06E8FFD03308CB15E37486ECF9EB" box="[346,501,1608,1634]" pageId="10" pageNumber="11" type="holotype">HOLOTYPE</typeStatus>
female:
<collectingCountry id="F362F8DAFFD03308C82EE37485CFF9EB" box="[609,726,1608,1634]" name="Taiwan" pageId="10" pageNumber="11">TAIWAN</collectingCountry>
,
<collectingRegion id="49B176A8FFD03308C8ABE3748423F9EB" box="[740,826,1608,1634]" country="Taiwan" name="Nantou" pageId="10" pageNumber="11">Nantou</collectingRegion>
Co.,
<collectingCounty id="62ABC0C6FFD03308C9CFE3748328F9EB" box="[896,1073,1608,1634]" pageId="10" pageNumber="11">Cingjing Farm</collectingCounty>
,
<collectingMunicipality id="6BAE2230FFD03308CE0FE374821DF9EB" box="[1088,1284,1608,1634]" pageId="10" pageNumber="11">Renai Township</collectingMunicipality>
, ex
<taxonomicName id="4C75C3C9FFD03308CF75E374861EF900" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B9016458FFD03308CF75E374861EF900" italics="true" pageId="10" pageNumber="11">Quercus variabilis</emphasis>
</taxonomicName>
, leg. Chang-Ti
<location id="8EAAEE91FFD03308CB8CE34C8519F903" LSID="urn:lsid:plazi:treatment:03DC095CFFD0330ECAD8E0E683BAFA3B:8EAAEE91FFD03308CB8CE34C8519F903" box="[451,512,1648,1674]" country="Taiwan" county="Cingjing Farm" municipality="Renai Township" name="Tang" pageId="10" pageNumber="11" stateProvince="Nantou">Tang</location>
,
<date id="FFCB9E8AFFD03308C841E34C8595F903" box="[526,652,1648,1674]" pageId="10" pageNumber="11" value="2008-11-07">
<collectingDate id="EF8F6762FFD03308C841E34C8595F903" box="[526,652,1648,1674]" pageId="10" pageNumber="11" value="2008-11-07">2008.XI.7.</collectingDate>
</date>
adult em.
<date id="FFCB9E8AFFD03308C943E34C848DF903" box="[780,916,1648,1674]" pageId="10" pageNumber="11" value="2008-12-07">
<collectingDate id="EF8F6762FFD03308C943E34C848DF903" box="[780,916,1648,1674]" pageId="10" pageNumber="11" value="2008-12-07">2008.XII.7.</collectingDate>
</date>
Eight
</materialsCitation>
<typeStatus id="54CE06E8FFD03308C9AAE34C836FF903" box="[997,1142,1648,1674]" pageId="10" pageNumber="11" type="paratype">PARATYPE</typeStatus>
females:
<materialsCitation id="3B1DB217FFD03308CEA3E34C8285F903" box="[1260,1436,1648,1674]" collectionCode="TAI-" country="Taiwan" pageId="10" pageNumber="11" specimenCount="1" specimenCount-female="1" stateProvince="Nantou" typeStatus="paratype">
<specimenCount id="9D7373C3FFD03308CEA3E34C824EF903" box="[1260,1367,1648,1674]" pageId="10" pageNumber="11" type="female">1 female</specimenCount>
:
<collectionCode id="ED64208FFFD03308CF2BE34C8285F903" box="[1380,1436,1648,1674]" pageId="10" pageNumber="11">TAI-</collectionCode>
</materialsCitation>
<materialsCitation id="3B1DB217FFD03308CAD8E3A48467F953" collectingDate="2001-03-01" collectorName="Chui-Mei Wang &amp; Seven" country="Taiwan" county="County" location="Kukuan" municipality="Central Cross Island Road" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Taichung" typeStatus="paratype">
WAN,
<collectingRegion id="49B176A8FFD03308CAA6E3A4864FF93B" box="[233,342,1688,1714]" country="Taiwan" name="Taichung" pageId="10" pageNumber="11">Taichung</collectingRegion>
<collectingCounty id="62ABC0C6FFD03308CB11E3A486ACF93B" box="[350,437,1688,1714]" pageId="10" pageNumber="11">County</collectingCounty>
, 36,5 K
<collectingMunicipality id="6BAE2230FFD03308C854E3A4844CF93B" box="[539,853,1688,1714]" pageId="10" pageNumber="11">Central Cross Island Road</collectingMunicipality>
,
<location id="8EAAEE91FFD03308C92DE3A484DBF93B" LSID="urn:lsid:plazi:treatment:03DC095CFFD0330ECAD8E0E683BAFA3B:8EAAEE91FFD03308C92DE3A484DBF93B" box="[866,962,1688,1714]" country="Taiwan" county="County" municipality="Central Cross Island Road" name="Kukuan" pageId="10" pageNumber="11" stateProvince="Taichung">Kukuan</location>
,
<location id="8EAAEE91FFD03308C981E3A483BBF93B" LSID="urn:lsid:plazi:treatment:03DC095CFFD0330ECAD8E0E683BAFA3B:8EAAEE91FFD03308C981E3A483BBF93B" box="[974,1186,1688,1714]" country="Taiwan" county="County" municipality="Central Cross Island Road" name="Heping Township" pageId="10" pageNumber="11" stateProvince="Taichung">Heping Township</location>
, ex
<taxonomicName id="4C75C3C9FFD03308CE9CE3A487AAF950" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B9016458FFD03308CE9CE3A487AAF950" italics="true" pageId="10" pageNumber="11">Quercus variabilis</emphasis>
</taxonomicName>
, leg.
<collectorName id="2680DD9CFFD03308CAB7E3FC86A0F953" box="[248,441,1728,1754]" pageId="10" pageNumber="11">Chui-Mei Wang</collectorName>
,
<date id="FFCB9E8AFFD03308CB88E3FC854AF953" box="[455,595,1728,1754]" pageId="10" pageNumber="11" value="2001-03-01">
<collectingDate id="EF8F6762FFD03308CB88E3FC854AF953" box="[455,595,1728,1754]" pageId="10" pageNumber="11" value="2001-03-01">1. III. 2001</collectingDate>
</date>
.
<collectorName id="2680DD9CFFD03308C82FE3FC85BEF953" box="[608,679,1728,1754]" pageId="10" pageNumber="11">Seven</collectorName>
female
<typeStatus id="54CE06E8FFD03308C945E3FC8467F953" box="[778,894,1728,1754]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
</materialsCitation>
:
<materialsCitation id="3B1DB217FFD03308C9C4E3FC84D9F8A3" collectingDate="2009-11-19" collectingDateMax="2009-12-22" collectingDateMin="2009-11-19" collectionCode="NMNS" collectorName="Tang &amp; The" country="Taiwan" county="Co." location="Heping Township" municipality="Central Cross Island Road" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Taichung" typeStatus="holotype">
<collectingCountry id="F362F8DAFFD03308C9C4E3FC8319F953" box="[907,1024,1728,1754]" name="Taiwan" pageId="10" pageNumber="11">TAIWAN</collectingCountry>
,
<collectingRegion id="49B176A8FFD03308CE42E3FC8360F953" box="[1037,1145,1728,1754]" country="Taiwan" name="Taichung" pageId="10" pageNumber="11">Taichung</collectingRegion>
<collectingCounty id="62ABC0C6FFD03308CECCE3FC83B6F953" box="[1155,1199,1728,1754]" pageId="10" pageNumber="11">Co.</collectingCounty>
, 29K
<collectingMunicipality id="6BAE2230FFD03308CEB5E3FC8631F88B" pageId="10" pageNumber="11">Central Cross Island Road</collectingMunicipality>
,
<location id="8EAAEE91FFD03308CB7BE3D48511F88B" LSID="urn:lsid:plazi:treatment:03DC095CFFD0330ECAD8E0E683BAFA3B:8EAAEE91FFD03308CB7BE3D48511F88B" box="[308,520,1768,1794]" country="Taiwan" county="Co." municipality="Central Cross Island Road" name="Heping Township" pageId="10" pageNumber="11" stateProvince="Taichung">Heping Township</location>
, ex
<taxonomicName id="4C75C3C9FFD03308C876E3D4840AF888" box="[569,787,1768,1793]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="10" pageNumber="11" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B9016458FFD03308C876E3D4840AF888" box="[569,787,1768,1793]" italics="true" pageId="10" pageNumber="11">Quercus variabilis</emphasis>
</taxonomicName>
, leg. Chang-Ti
<collectorName id="2680DD9CFFD03308C981E3D48312F88B" box="[974,1035,1768,1794]" pageId="10" pageNumber="11">Tang</collectorName>
,
<date id="FFCB9E8AFFD03308CE57E3D483BCF88B" box="[1048,1189,1768,1794]" pageId="10" pageNumber="11" value="2009-11-19">
<collectingDate id="EF8F6762FFD03308CE57E3D483BCF88B" box="[1048,1189,1768,1794]" pageId="10" pageNumber="11" value="2009-11-19">2009.XI.19.</collectingDate>
</date>
adult dissected from galls on
<date id="FFCB9E8AFFD03308CB4CE22C8680F8A3" box="[259,409,1808,1834]" pageId="10" pageNumber="11" value="2009-12-22">
<collectingDate id="EF8F6762FFD03308CB4CE22C8680F8A3" box="[259,409,1808,1834]" pageId="10" pageNumber="11" value="2009-12-22">2009.XII.22.</collectingDate>
</date>
<collectorName id="2680DD9CFFD03308CBEBE22C86CBF8A3" box="[420,466,1808,1834]" pageId="10" pageNumber="11">The</collectorName>
<typeStatus id="54CE06E8FFD03308CB91E22C855DF8A3" box="[478,580,1808,1834]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
female is deposited in
<collectionCode id="ED64208FFFD03308C927E22C84D9F8A3" box="[872,960,1808,1834]" country="Taiwan" httpUri="http://grbio.org/cool/2nry-s1yg" name="National Museum of Natural Science" pageId="10" pageNumber="11">NMNS</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B1DB217FFD03308C99FE22C83C6F8A3" box="[976,1247,1808,1834]" collectionCode="NCHU" country="Taiwan" pageId="10" pageNumber="11" specimenCount="3" stateProvince="Taichung" typeStatus="paratype">
<specimenCount id="9D7373C3FFD03308C99FE22C8342F8A3" box="[976,1115,1808,1834]" pageId="10" pageNumber="11" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
in
<collectionCode id="ED64208FFFD03308CEC7E22C83C6F8A3" box="[1160,1247,1808,1834]" country="Taiwan" httpUri="http://grbio.org/cool/e7yf-mhkb" name="National Chung Hsing University" pageId="10" pageNumber="11">NCHU</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B1DB217FFD03308CEA0E22C87F6F8DB" collectionCode="USNM" country="Taiwan" pageId="10" pageNumber="11" specimenCount="2" stateProvince="Taichung" typeStatus="paratype">
<specimenCount id="9D7373C3FFD03308CEA0E22C8263F8A3" box="[1263,1402,1808,1834]" pageId="10" pageNumber="11" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
in
<collectionCode id="ED64208FFFD03308CAD8E20487F6F8DB" box="[151,239,1848,1874]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">USNM</collectionCode>
</materialsCitation>
;
<specimenCount id="9D7373C3FFD03308CAB4E2048698F8DB" box="[251,385,1848,1874]" pageId="10" pageNumber="11" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
in PDL.
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD03308CA8AE26383EAF8F3" box="[197,1267,1887,1914]" pageId="10" pageNumber="11" type="etymology">
<paragraph id="8BCAB84AFFD03308CA8AE26383EAF8F3" blockId="10.[151,1436,1607,1914]" box="[197,1267,1887,1914]" pageId="10" pageNumber="11">
<emphasis id="B9016458FFD03308CA8AE2638648F8F0" bold="true" box="[197,337,1887,1913]" pageId="10" pageNumber="11">Etymology.</emphasis>
The species is named after Formosa, a previous name for the island of
<collectingCountry id="F362F8DAFFD03308CEDAE25C83F6F8F3" box="[1173,1263,1888,1914]" name="Taiwan" pageId="10" pageNumber="11">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD03309CA8AE2938450FE7B" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BCAB84AFFD03309CA8AE2938450FE7B" blockId="10.[151,1436,1967,2034]" lastBlockId="11.[151,1437,151,2018]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B9016458FFD03308CA8AE293865DF840" bold="true" box="[197,324,1967,1993]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
Closely related to
<taxonomicName id="4C75C3C9FFD03308C86FE28C8404F840" baseAuthorityName="Ashmead" baseAuthorityYear="1904" box="[544,797,1968,1993]" class="Insecta" family="Cynipidae" genus="Trichagalma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="serratae">
<emphasis id="B9016458FFD03308C86FE28C8404F840" box="[544,797,1968,1993]" italics="true" pageId="10" pageNumber="11">Trichagalma serratae</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C75C3C9FFD03308C904E28C84FDF840" authority="Melika &amp; Pujade-Villar &amp; Abe &amp; Tang &amp; Nicholls &amp; Wachi &amp; Ide &amp; Yang &amp; Pénzes &amp; Csóka &amp; Stone, 2010" authorityName="Melika &amp; Pujade-Villar &amp; Abe &amp; Tang &amp; Nicholls &amp; Wachi &amp; Ide &amp; Yang &amp; Pénzes &amp; Csóka &amp; Stone" authorityYear="2010" box="[843,996,1968,1993]" class="Insecta" family="Cynipidae" genus="Trichagalma" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="formosana" status="sp. nov.">
<emphasis id="B9016458FFD03308C904E28C84FDF840" box="[843,996,1968,1993]" italics="true" pageId="10" pageNumber="11">T. formosana</emphasis>
</taxonomicName>
,
<emphasis id="B9016458FFD03308C9BCE293839BF840" bold="true" box="[1011,1154,1967,1993]" pageId="10" pageNumber="11">
<taxonomicNameLabel id="A232D923FFD03308C9BCE293839BF840" box="[1011,1154,1967,1993]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
, the body is brown with darker brown marks; the clypeus is coriaceous, brown, with a median incision ventrally; the scape, pedicel and F1
<emphasis id="B9016458FFD13309CAA8E5AB87EFFF38" bold="true" box="[231,246,151,177]" pageId="11" pageNumber="12">–</emphasis>
F2 brown, remaining flagellomeres dark brown; the head in anterior view is ovate, the broadest part of genae behind eyes is level with the transfacial line; the mesosoma in lateral view much higher; the forewing margin with very short dense cilia; the radial cell of the forewing 3.5
<emphasis id="B9016458FFD13309C9F9E5DB84DCFE88" bold="true" box="[950,965,231,257]" pageId="11" pageNumber="12">–</emphasis>
3.9 times as long as broad, R1 on a short distance running along the wing margin, while in
<taxonomicName id="4C75C3C9FFD13309C8A3E42C8472FEA0" baseAuthorityName="Ashmead" baseAuthorityYear="1904" box="[748,875,272,298]" class="Insecta" family="Cynipidae" genus="Trichagalma" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="serratae">
<emphasis id="B9016458FFD13309C8A3E42C85E3FEA0" box="[748,762,272,297]" italics="true" pageId="11" pageNumber="12">T</emphasis>
.
<emphasis id="B9016458FFD13309C945E42C8472FEA0" box="[778,875,272,297]" italics="true" pageId="11" pageNumber="12">serratae</emphasis>
</taxonomicName>
the body is rusty brown with black marks; the clypeus is uniformly microreticulate, black, without median incision ventrally (
<figureCitation id="134EA4CFFFD13309CE0CE4048385FEDB" box="[1091,1180,312,338]" captionStart="FIGURES 24–30" captionStartId="43.[151,269,1493,1517]" captionTargetBox="[151,1426,212,1469]" captionTargetId="figure-52@43.[151,1436,194,1469]" captionTargetPageId="43" captionText="FIGURES 24–30. Trichagalma serratae, asexual female and galls: 24, head, anterior view; 25, antenna; 26, metascutellum and propodeum, posterodorsal view; 27, mesosoma, lateral view; 28, forewing; 29–30, galls (photos by G. Csóka)." pageId="11" pageNumber="12">Fig. 24</figureCitation>
); the antenna entirely and uniformly black (
<figureCitation id="134EA4CFFFD13309CBD8E45C86E9FEF3" box="[407,496,352,378]" captionStart="FIGURES 24–30" captionStartId="43.[151,269,1493,1517]" captionTargetBox="[151,1426,212,1469]" captionTargetId="figure-52@43.[151,1436,194,1469]" captionTargetPageId="43" captionText="FIGURES 24–30. Trichagalma serratae, asexual female and galls: 24, head, anterior view; 25, antenna; 26, metascutellum and propodeum, posterodorsal view; 27, mesosoma, lateral view; 28, forewing; 29–30, galls (photos by G. Csóka)." pageId="11" pageNumber="12">Fig. 25</figureCitation>
); the head in anterior view is more trapezoid, the broadest part of genae behind eyes is above the level of transfacial line (
<figureCitation id="134EA4CFFFD13309C8CAE4B485C7FE2B" box="[645,734,392,418]" captionStart="FIGURES 24–30" captionStartId="43.[151,269,1493,1517]" captionTargetBox="[151,1426,212,1469]" captionTargetId="figure-52@43.[151,1436,194,1469]" captionTargetPageId="43" captionText="FIGURES 24–30. Trichagalma serratae, asexual female and galls: 24, head, anterior view; 25, antenna; 26, metascutellum and propodeum, posterodorsal view; 27, mesosoma, lateral view; 28, forewing; 29–30, galls (photos by G. Csóka)." pageId="11" pageNumber="12">Fig. 24</figureCitation>
); the mesosoma in lateral view shorter (
<figureCitation id="134EA4CFFFD13309CEFBE4B48214FE2B" box="[1204,1293,392,418]" captionStart="FIGURES 24–30" captionStartId="43.[151,269,1493,1517]" captionTargetBox="[151,1426,212,1469]" captionTargetId="figure-52@43.[151,1436,194,1469]" captionTargetPageId="43" captionText="FIGURES 24–30. Trichagalma serratae, asexual female and galls: 24, head, anterior view; 25, antenna; 26, metascutellum and propodeum, posterodorsal view; 27, mesosoma, lateral view; 28, forewing; 29–30, galls (photos by G. Csóka)." pageId="11" pageNumber="12">Fig. 27</figureCitation>
); the smoky spots on forewing much darker and well-marked; the forewing margin with long cilia, the radial cell 4.3
<emphasis id="B9016458FFD13309CF26E493826EFE40" bold="true" box="[1385,1399,431,457]" pageId="11" pageNumber="12">–</emphasis>
4.5 times as long as broad, R1 reaching wing margin (
<figureCitation id="134EA4CFFFD13309C8ABE4E48425FE7B" box="[740,828,472,498]" captionStart="FIGURES 24–30" captionStartId="43.[151,269,1493,1517]" captionTargetBox="[151,1426,212,1469]" captionTargetId="figure-52@43.[151,1436,194,1469]" captionTargetPageId="43" captionText="FIGURES 24–30. Trichagalma serratae, asexual female and galls: 24, head, anterior view; 25, antenna; 26, metascutellum and propodeum, posterodorsal view; 27, mesosoma, lateral view; 28, forewing; 29–30, galls (photos by G. Csóka)." pageId="11" pageNumber="12">Fig. 28</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD1330ECA8AE4C387D3FBC3" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="description">
<paragraph id="8BCAB84AFFD13309CA8AE4C38300FD1B" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="12">
<emphasis id="B9016458FFD13309CA8AE4C38644FD90" bold="true" box="[197,349,511,537]" pageId="11" pageNumber="12">Description.</emphasis>
ASEXUAL FEMALE (
<typeStatus id="54CE06E8FFD13309C8CBE73C85E8FD93" box="[644,753,512,538]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
). Body, legs uniformly brown; head posteriorly, antenna (except scape, pedicel, F1
<emphasis id="B9016458FFD13309CB95E71B86F0FDC8" bold="true" box="[474,489,551,577]" pageId="11" pageNumber="12">–</emphasis>
F2), mandibles, stripes along anterior parallel and parapsidal lines, scutellar foveae, preaxilla, lateral axillar area, axillula, metascutellum, central propodeal area dark brown to black; body with dense white setae; wing veins dark brown, with dark brown spots.
</paragraph>
<paragraph id="8BCAB84AFFD13309CA8AE79C8471FA3B" blockId="11.[151,1437,151,2018]" pageId="11" pageNumber="12">
Head coriaceous, with uniformly very dense white setae, 2.5 times as broad as long from above; 1.4 times as broad as high in anterior view and equal to width of mesosoma. Gena coriaceous, strongly broadened behind eye, broader than cross diameter of eye. Malar space coriaceous, with dense setae, 0.4 times as long as height of eye, without striae and malar sulcus. POL 1.2 times as long as OOL; OOL 3.3 times as long as diameter of lateral ocellus, 2.0 times as long as LOL; lateral ocelli slightly ovate, larger than median rounded ocellus. Transfacial distance 1.4 times as long as height of eye and 1.8 times as long as height of lower face (distance between antennal rim and ventral margin of clypeus); diameter of antennal socket slightly shorter than distance between them, distance between socket and eye margin 1.5 times as long as diameter of socket. Lower face coriaceous, with strongly elevated median area and dense setae. Clypeus rectangular, flat, above 2.0 times as broad as high, coriaceous, with deep anterior tentorial pits, distinct epistomal sulcus and clypeopleurostomal line; ventrally emarginate and slightly incised medially. Frons coriaceous, with deep smooth and shiny impression below median ocellus; vertex and occiput uniformly dull coriaceous; interocellar area elevated. Postocciput around occipital foramen impressed, with numerous striae extending to level of gula; posterior tentorial pits large, deep, elongate; hypostomal bridge at least 2.0 times as high as broad, lower part narrowed down to emarginate hypostomal carina; occipital foramen slightly shorter than height of hypostomal bridge, around 1.5 times shorter than height of oral foramen. Antenna with 12 flagellomeres (or 13, indistinct suture between F13 and F12 visible in the
<typeStatus id="54CE06E8FFD13309C8DBE01C85E7FAB3" box="[660,766,1312,1338]" pageId="11" pageNumber="12" type="paratype">paratype</typeStatus>
), longer than head+mesosoma; pedicel nearly as long as broad, F1 4.1 times as long as pedicel, 1.3 times as long as F2, F2 1.1 times as long as F3, F4 slightly shorter than F3, F6
<emphasis id="B9016458FFD13309CB50E0538637FA00" bold="true" box="[287,302,1391,1417]" pageId="11" pageNumber="12">–</emphasis>
F11 shorter than F5 and nearly equal in length; fused F12+F13 1.7 times as long as F11; placodeal sensilla on F3–F12, in numerous rows, absent on F1–F2.
</paragraph>
<paragraph id="8BCAB84AFFD1330ECA8AE0FC84E0FD93" blockId="11.[151,1437,151,2018]" lastBlockId="12.[151,1437,152,1458]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">Mesosoma longer than high in lateral view, with uniform dense white setae. Pronotum uniformly coriaceous; with uniform dense white setae and irregular wrinkles, emarginate along lateral edge, followed by deep longitudinal invagination. Anterior rim of pronotum narrow, emarginate; propleuron with black stripe along lateral side, delicately coriaceous, with white setae, strongly concave in mediocentral part. Mesoscutum coriaceous, with distinct punctures in anterior 1/3; longer than broad (width measured across basis of tegulae); notauli complete, superficially impressed for full length; median mesoscutal line absent; anterior parallel and parapsidal lines not impressed, broad, smooth and shiny, extending to half of mesoscutum length, marked with black stripes. Transscutal articulation absent. Mesoscutellum slightly longer than broad, 1.5 times shorter than length of mesoscutum, uniformly coriaceous, overhanging metanotum; without distinct scutellar foveae, only transverse, slightly impressed area indicated by much darker colour present. Mesopleuron, including speculum, delicately coriaceous, shiny, with dense white setae, narrowly impressed along posterior edge; mesopleural triangle rugose, with dense white setae and numerous strong wrinkles. Metapleural sulcus reaching mesopleuron at half height; preaxilla coriaceous; lateral axillar area with parallel wrinkles, without setae; axillar carina broad, with longitudinal striae; axillula slightly ovate, smooth, with dense white setae and punctures; subaxillular bar narrow, smooth, shiny, in most posterior end narrower than height of metanotal trough. Metascutellum black, uniformly delicately punctate, slightly higher than height of smooth, shiny ventral impressed area; metanotal trough smooth, shiny, without setae. Lateral propodeal carinae absent, central propodeal area smooth, shiny, with central longitudinal delicate carina, delimited only by darker colour, without setae; lateral propodeal area uniformly coriaceous, with dense white setae; nucha very short, with few delicate longitudinal sulci dorsolaterally and laterally. Forewing longer than body, with dark brown veins and with dark irregular sclerotized pigmented spots, margin with very short dense cilia; radial cell 3.9 times as long as broad, R1 on a short distance running along wing margin, Rs nearly reaching wing margin; areolet large, triangular, well-delimited by distinct veins; projection of Rs+M reaching basalis in the lower third. Hind tarsomere I to V ratio as 1.0:0.6:0.3:0.1:0.5. Tarsal claws simple, without basal lobe.</paragraph>
<paragraph id="8BCAB84AFFD6330ECA8AE71483E8FD6B" blockId="12.[151,1437,152,1458]" pageId="12" pageNumber="13">Metasoma shorter than head+mesosoma (in fresh alive specimens metasoma longer than head+mesosoma), higher than long in lateral view, smooth, shiny, without setae; only 2nd and 3rd metasomal tergites posterolaterally with white setae; 2nd tergite extending to 1/3 length of metasoma; prominent part of ventral spine of hypopygium extremely short, as long as broad ventrally, with dense long white setae, extending far beyond apex of spine; setae located only on both lateral sides of ventral spine, absent ventrally.</paragraph>
<paragraph id="8BCAB84AFFD6330ECA8AE7CC83F6FC83" blockId="12.[151,1437,152,1458]" box="[197,1263,751,778]" pageId="12" pageNumber="13">
Body length
<quantity id="4C8D15AFFFD6330ECB13E7CC86F0FC83" box="[348,489,751,778]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="4.7" metricValueMin="4.5" pageId="12" pageNumber="13" unit="mm" value="4.6" valueMax="4.7" valueMin="4.5">
4.5
<emphasis id="B9016458FFD6330ECBCEE7D38689FC80" bold="true" box="[385,400,751,777]" pageId="12" pageNumber="13">–</emphasis>
4.7 mm
</quantity>
(n=4). In freshly killed specimens body length
<quantity id="4C8D15AFFFD6330ECE58E7CC8387FC83" box="[1047,1182,752,778]" metricMagnitude="-3" metricUnit="m" metricValue="5.199999999999999" metricValueMax="5.3" metricValueMin="5.1" pageId="12" pageNumber="13" unit="mm" value="5.199999999999999" valueMax="5.3" valueMin="5.1">5.1-5.3 mm</quantity>
(n=2).
</paragraph>
<paragraph id="8BCAB84AFFD6330ECA8AE62B87D3FBC3" blockId="12.[151,1437,152,1458]" pageId="12" pageNumber="13">
<emphasis id="B9016458FFD6330ECA8AE62B87E2FCB8" bold="true" box="[197,251,791,817]" pageId="12" pageNumber="13">Gall</emphasis>
. The gall when young is juicy, soft, covered with small raised tubercles, and green with purple spots on areas exposed to direct sunlight (
<figureCitation id="134EA4CFFFD6330EC805E67C85CEFCD3" box="[586,727,832,858]" captionStart="FIGURES 43–49" captionStartId="46.[151,269,1337,1361]" captionTargetBox="[151,1436,194,1313]" captionTargetId="figure-44@46.[151,1436,194,1313]" captionTargetPageId="46" captionText="FIGURES 43–49. Asexual galls of: 43–47, Trichagalma formosana, new species, 48–49, Trichagalma acutissimae (photos by Chang-Ti Tang and G. Csóka)." pageId="12" pageNumber="13">Figs 43, 45</figureCitation>
). The gall when mature reaches
<quantity id="4C8D15AFFFD6330ECE2FE67C83ADFCD3" box="[1120,1204,832,858]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="mm" value="15.0">15 mm</quantity>
in diameter, and is brown with a slightly irregular surface (
<figureCitation id="134EA4CFFFD6330EC827E65485E9FC0B" box="[616,752,872,898]" captionStart="FIGURES 43–49" captionStartId="46.[151,269,1337,1361]" captionTargetBox="[151,1436,194,1313]" captionTargetId="figure-44@46.[151,1436,194,1313]" captionTargetPageId="46" captionText="FIGURES 43–49. Asexual galls of: 43–47, Trichagalma formosana, new species, 48–49, Trichagalma acutissimae (photos by Chang-Ti Tang and G. Csóka)." pageId="12" pageNumber="13">Figs 44, 47</figureCitation>
). The mature gall is hollow, with a tough woody wall
<quantity id="4C8D15AFFFD6330ECF26E65487E9FC23" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="12" pageNumber="13" unit="mm" value="2.0" valueMax="2.5" valueMin="1.5">1.5– 2.5 mm</quantity>
thick. The interior space contains a single larval chamber, attached to the wall by a stalk, which, however, dry out when the gall is mature (
<figureCitation id="134EA4CFFFD6330EC815E68485AAFC5B" box="[602,691,952,978]" captionStart="FIGURES 43–49" captionStartId="46.[151,269,1337,1361]" captionTargetBox="[151,1436,194,1313]" captionTargetId="figure-44@46.[151,1436,194,1313]" captionTargetPageId="46" captionText="FIGURES 43–49. Asexual galls of: 43–47, Trichagalma formosana, new species, 48–49, Trichagalma acutissimae (photos by Chang-Ti Tang and G. Csóka)." pageId="12" pageNumber="13">Fig. 46</figureCitation>
). The larval chamber is ovoid, up to
<quantity id="4C8D15AFFFD6330ECE26E68483B4FC5B" box="[1129,1197,952,978]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="mm" value="6.0">6 mm</quantity>
in length, and has a tough but thin wall. Galls may be found singly or in groups, most commonly on lateral buds of young shoots. Solitary galls are almost spherical, while closely clustered galls may be deformed. Old galls persist on the host tree.
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD6330ECA8AE16B8486FA9B" pageId="12" pageNumber="13" type="biology_ecology">
<paragraph id="8BCAB84AFFD6330ECA8AE16B8486FA9B" blockId="12.[151,1437,152,1458]" pageId="12" pageNumber="13">
<emphasis id="B9016458FFD6330ECA8AE16B8630FBF8" bold="true" box="[197,297,1111,1137]" pageId="12" pageNumber="13">Biology.</emphasis>
Only the asexual generation is known from galls on
<taxonomicName id="4C75C3C9FFD6330EC9D6E164836AFBF8" box="[921,1139,1112,1137]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="variabilis">
<emphasis id="B9016458FFD6330EC9D6E164836AFBF8" box="[921,1139,1112,1137]" italics="true" pageId="12" pageNumber="13">Quercus variabilis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C75C3C9FFD6330ECEE2E1648256FBF8" box="[1197,1359,1112,1137]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="12" pageNumber="13" phylum="Tracheophyta" rank="species" species="acutissima">
<emphasis id="B9016458FFD6330ECEE2E1648256FBF8" box="[1197,1359,1112,1137]" italics="true" pageId="12" pageNumber="13">Q. acutissima</emphasis>
</taxonomicName>
. Galls appear on the tree from early August. Under the laboratory conditions, adults emerged in December, in the nature they might overwinter in the gall and emerge in spring of the following year. Galls mostly remains on the tree for one year. Common, however, the infestation by inquilines and parasitoids is quite high what is strongly influence the emerging of the cynipid gall inducer wasps.
</paragraph>
</subSubSection>
<subSubSection id="C36FEBC1FFD6330ECA8AE02383BAFA3B" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BCAB84AFFD6330ECA8AE02383BAFA3B" blockId="12.[151,1437,152,1458]" pageId="12" pageNumber="13">
<emphasis id="B9016458FFD6330ECA8AE0238647FAB0" bold="true" box="[197,350,1311,1337]" pageId="12" pageNumber="13">Distribution</emphasis>
.
<collectingCountry id="F362F8DAFFD6330ECB3EE01C86DEFAB3" box="[369,455,1312,1338]" name="Taiwan" pageId="12" pageNumber="13">Taiwan</collectingCountry>
(
<collectingRegion id="49B176A8FFD6330ECB95E01C85BDFAB3" box="[474,676,1312,1338]" country="Taiwan" name="Taoyuan" pageId="12" pageNumber="13">Taoyuan County</collectingRegion>
, Fuhsing Township;
<collectingRegion id="49B176A8FFD6330EC9EAE01C8373FAB3" box="[933,1130,1312,1338]" country="Taiwan" name="Hsinchu" pageId="12" pageNumber="13">Hsinchu County</collectingRegion>
, Hsinfong Township and Jiashih Township;
<collectingRegion id="49B176A8FFD6330ECB36E07486FCFAEB" box="[377,485,1352,1378]" country="Taiwan" name="Taichung" pageId="12" pageNumber="13">Taichung</collectingRegion>
County, Central Cross Island Road, Heping Township;
<collectingRegion id="49B176A8FFD6330ECEC0E0748251FAEB" box="[1167,1352,1352,1378]" country="Taiwan" name="Nantou" pageId="12" pageNumber="13">Nantou County</collectingRegion>
, Renai Township). Same
<typeStatus id="54CE06E8FFD6330ECB25E04C8685FA03" box="[362,412,1392,1418]" pageId="12" pageNumber="13">type</typeStatus>
of galls were found in
<collectingCountry id="F362F8DAFFD6330EC8E7E04C85F7FA03" box="[680,750,1392,1418]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
, and it is illustrated in
<bibRefCitation id="EFE4C5BBFFD6330EC9BAE04C8230FA03" author="Yukawa, J. &amp; Masuda, H." box="[1013,1321,1392,1418]" pageId="12" pageNumber="13" refId="ref35517" refString="Yukawa, J. &amp; Masuda, H. (eds) (1996) Insect and mite galls of Japan in colors. Zenkoku-Nouson-Kyoukai, Tokyo, 826 pp. (in Japanese)." type="book" year="1996">Yukawa &amp; Masuda (1996)</bibRefCitation>
under C
<emphasis id="B9016458FFD6330ECFC2E0538285FA00" bold="true" box="[1421,1436,1391,1417]" pageId="12" pageNumber="13">–</emphasis>
076. However, adults were never reared in
<collectingCountry id="F362F8DAFFD6330EC8C0E0A485CBFA3B" box="[655,722,1432,1458]" name="Japan" pageId="12" pageNumber="13">Japan</collectingCountry>
and thus the species was not described.
</paragraph>
</subSubSection>
</treatment>
</document>
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