614 lines
80 KiB
XML
614 lines
80 KiB
XML
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<mods:title id="44AD8B519209441B7149CBB41EB93F0A">A systematic revision of Draculoides (Schizomida: Hubbardiidae) of the Pilbara, Western Australia, Part I: the Western Pilbara</mods:title>
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<taxonomicName id="4C674D4CFF9AFFCFCC8B0487FBC2FA99" authority="(Harvey, Berry, Edward and Humphreys, 2008)" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,1058,1275,1302]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anachoretus">
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<emphasis id="B913EADDFF9AFFCFCC8B0487FE27FA9A" bold="true" box="[151,455,1275,1302]" italics="true" pageId="28" pageNumber="29">Draculoides anachoretus</emphasis>
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(
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<bibRefCitation id="EFF64B3EFF9AFFCFCDCA0487FBFAFA99" author="Harvey, M. S. & Berry, O. & Edward, K. L. & Humphreys, G." box="[470,1050,1275,1302]" pageId="28" pageNumber="29" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. & Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">Harvey, Berry, Edward and Humphreys, 2008</bibRefCitation>
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)
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<paragraph id="8BD836CFFF9AFFCFCC8B0563FE85FAB6" blockId="28.[151,1146,1275,1373]" box="[151,357,1311,1338]" pageId="28" pageNumber="29">
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(
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<figureCitation id="135C2A4AFF9AFFCFCC830563FEEBFAB6" box="[159,267,1311,1338]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStartId-0="16.[151,250,1868,1892]" captionStartId-1="17.[151,250,1910,1934]" captionStartId-2="18.[151,250,1547,1571]" captionStartId-3="19.[151,250,1882,1906]" captionStartId-4="20.[151,250,1216,1240]" captionStartId-5="21.[151,250,1709,1733]" captionStartId-6="22.[151,250,1755,1779]" captionStartId-7="23.[151,250,1919,1943]" captionTargetBox-0="[346,1243,188,1844]" captionTargetBox-1="[332,1254,181,1881]" captionTargetBox-2="[158,1420,181,1521]" captionTargetBox-3="[151,1425,182,1858]" captionTargetBox-4="[158,1424,183,1189]" captionTargetBox-5="[155,1423,181,1683]" captionTargetBox-6="[159,1425,181,1729]" captionTargetBox-7="[151,1430,181,1893]" captionTargetId-0="figure-20@16.[339,1249,181,1844]" captionTargetId-1="figure-17@17.[332,1254,181,1886]" captionTargetId-2="figure-20@18.[151,1436,181,1524]" captionTargetId-3="figure-17@19.[151,1436,181,1859]" captionTargetId-4="figure-20@20.[151,1436,181,1192]" captionTargetId-5="figure-17@21.[155,1432,181,1685]" captionTargetId-6="figure-20@22.[151,1436,181,1732]" captionTargetId-7="figure-17@23.[151,1436,181,1894]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionTargetPageId-4="20" captionTargetPageId-5="21" captionTargetPageId-6="22" captionTargetPageId-7="23" captionText-0="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." captionText-1="FIGURE 2. Maximum Likelihood tree, based on the reduced dataset. See Methods for details. Each terminal represents a single species or OTU, with named species in bold. Clades are identified using dashed boxes and shaded boxes encompass the species discussed in this study. Nodes with bootstrap support <80 are not shown, except for the common ancestor of the Draculoides radiation." captionText-2="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-3="FIGURE 4. Maximum Likelihood tree based on COI mini-barcodes (Fig. 3) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-4="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-5="FIGURE 6. Maximum Likelihood tree based on COI mini-barcodes (Fig. 5) showing similarity of mini-barcode sequences relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." captionText-6="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." captionText-7="FIGURE 8. Maximum Likelihood tree based on ITS2 mini-barcodes (Fig. 7) showing similarity of mini-barcode sequences, relative to species identity. This phylogeny does not recover an accurate tree topology, and evolutionary relationships should not be inferred from this figure." figureDoi-0="http://doi.org/10.5281/zenodo.4416851" figureDoi-1="http://doi.org/10.5281/zenodo.4416855" figureDoi-2="http://doi.org/10.5281/zenodo.4416857" figureDoi-3="http://doi.org/10.5281/zenodo.4416859" figureDoi-4="http://doi.org/10.5281/zenodo.4416863" figureDoi-5="http://doi.org/10.5281/zenodo.4416865" figureDoi-6="http://doi.org/10.5281/zenodo.4416867" figureDoi-7="http://doi.org/10.5281/zenodo.4416869" httpUri-0="https://zenodo.org/record/4416851/files/figure.png" httpUri-1="https://zenodo.org/record/4416855/files/figure.png" httpUri-2="https://zenodo.org/record/4416857/files/figure.png" httpUri-3="https://zenodo.org/record/4416859/files/figure.png" httpUri-4="https://zenodo.org/record/4416863/files/figure.png" httpUri-5="https://zenodo.org/record/4416865/files/figure.png" httpUri-6="https://zenodo.org/record/4416867/files/figure.png" httpUri-7="https://zenodo.org/record/4416869/files/figure.png" pageId="28" pageNumber="29">Figs. 1–8</figureCitation>
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,
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<figureCitation id="135C2A4AFF9AFFCFCD0B055CFEBDFAB6" box="[279,349,1312,1338]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="29.[151,250,1356,1380]" captionStartId-1="30.[151,250,1579,1603]" captionTargetBox-0="[152,1435,181,1331]" captionTargetBox-1="[149,1439,957,1550]" captionTargetId-0="figure-17@29.[151,1436,181,1332]" captionTargetId-1="figure-382@30.[149,1439,957,1556]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURE 11. Draculoides anachoretus male (WAM T140985): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 2 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." captionText-1="FIGURE 12. Draculoides anachoretus male (WAM T140985): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. The scale bar is 0.1 mm and applies to A–C." figureDoi-0="http://doi.org/10.5281/zenodo.4416877" figureDoi-1="http://doi.org/10.5281/zenodo.4416881" httpUri-0="https://zenodo.org/record/4416877/files/figure.png" httpUri-1="https://zenodo.org/record/4416881/files/figure.png" pageId="28" pageNumber="29">11–12</figureCitation>
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)
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</paragraph>
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http://zoobank.org/NomenclaturalActs/
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<subSubSection id="C37D6544FF9AFFCFCC8B05F4FB1CFA33" pageId="28" pageNumber="29" type="reference_group">
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<paragraph id="8BD836CFFF9AFFCFCC8B05F4FB99FA2C" blockId="28.[151,1276,1416,1471]" box="[151,1145,1416,1440]" pageId="28" pageNumber="29">
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<taxonomicName id="4C674D4CFF9AFFCFCC8B05F4FC16FA2C" ID-CoL="75P5M" authority="Harvey, Berry, Edward and Humphreys 2008: 185 - 187" authorityName="Harvey, Berry, Edward and Humphreys" authorityPageNumber="185 - 187" authorityYear="2008" box="[151,1014,1416,1440]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anachoretus">
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<emphasis id="B913EADDFF9AFFCFCC8B05F4FE24FA2C" box="[151,452,1416,1440]" italics="true" pageId="28" pageNumber="29">Paradraculoides anachoretus</emphasis>
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<bibRefCitation id="EFF64B3EFF9AFFCFCDD605F4FC16FA2C" author="Harvey, M. S. & Berry, O. & Edward, K. L. & Humphreys, G." box="[458,1014,1416,1440]" pageId="28" pageNumber="29" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. & Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">Harvey, Berry, Edward and Humphreys 2008:185–187</bibRefCitation>
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</taxonomicName>
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, figs. 32–35.
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</paragraph>
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<paragraph id="8BD836CFFF9AFFCFCC8B05DBFB1CFA33" blockId="28.[151,1276,1416,1471]" box="[151,1276,1446,1471]" pageId="28" pageNumber="29">
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<taxonomicName id="4C674D4CFF9AFFCFCC8B05DBFCA3FA33" authority="(Harvey, Berry, Edward and Humphreys)" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,835,1447,1471]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="anachoretus">
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<emphasis id="B913EADDFF9AFFCFCC8B05DBFE77FA33" box="[151,407,1447,1471]" italics="true" pageId="28" pageNumber="29">Draculoides anachoretus</emphasis>
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(Harvey, Berry, Edward and Humphreys)
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</taxonomicName>
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:
|
||
<bibRefCitation id="EFF64B3EFF9AFFCFCF5705DBFBEDFA33" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[843,1037,1447,1471]" pageId="28" pageNumber="29" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
|
||
Abrams
|
||
<emphasis id="B913EADDFF9AFFCFCFBD05DBFC2EFA33" box="[929,974,1447,1471]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
. 2019
|
||
</bibRefCitation>
|
||
MPE 106532: 8, fig. 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FF9AFFCFCC8B0590FA6FF81F" pageId="28" pageNumber="29" type="materials_examined">
|
||
<paragraph id="8BD836CFFF9AFFCFCC8B0590FA6AF9A7" blockId="28.[151,1437,1516,1940]" pageId="28" pageNumber="29">
|
||
<emphasis id="B913EADDFF9AFFCFCC8B0590FE61F98A" bold="true" box="[151,385,1516,1542]" pageId="28" pageNumber="29">Material examined.</emphasis>
|
||
<materialsCitation id="3B0F3C92FF9AFFCFCD950591FA6AF9A7" ID-GBIF-Occurrence="3013449326" accessionNumber="T63327" collectionCode="WAM, T, COI" country="Australia" latitude="-21.669722" location="Pannawonica" longLatPrecision="21" longitude="115.88694" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
|
||
<emphasis id="B913EADDFF9AFFCFCD950591FE10F98B" box="[393,496,1517,1543]" italics="true" pageId="28" pageNumber="29">
|
||
<typeStatus id="54DC886DFF9AFFCFCD950591FE10F98B" box="[393,496,1517,1543]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
female.
|
||
<collectingCountry id="F370765FFF9AFFCFCE4C0591FD13F98B" box="[592,755,1517,1543]" name="Australia" pageId="28" pageNumber="29">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFF9AFFCFCEE20592FC27F98B" box="[766,967,1517,1543]" italics="true" pageId="28" pageNumber="29">
|
||
<collectingRegion id="49A3F82DFF9AFFCFCEE20592FC27F98B" box="[766,967,1517,1543]" country="Australia" name="Western Australia" pageId="28" pageNumber="29">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectorName id="26925319FF9AFFCFCFCA0591FBCCF98B" box="[982,1068,1517,1543]" pageId="28" pageNumber="29">Mesa A</collectorName>
|
||
(Borehole 2497),
|
||
<quantity id="4C9F9B2AFF9AFFCFC8E70591FAB8F98B" box="[1275,1368,1517,1544]" metricMagnitude="4" metricUnit="m" metricValue="4.52" pageId="28" pageNumber="29" unit="km" value="45.2">45.2 km</quantity>
|
||
W. of
|
||
<location id="8EB86014FF9AFFCFCC8B066DFECFF9A7" LSID="urn:lsid:plazi:treatment:03CE87D9FF9AFFCDCC8B0487FDB6F97D:8EB86014FF9AFFCFCC8B066DFECFF9A7" box="[151,303,1553,1579]" country="Australia" latitude="-21.669722" longLatPrecision="21" longitude="115.88694" name="Pannawonica" pageId="28" pageNumber="29" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCD26066DFE20F9A7" box="[314,448,1553,1579]" degrees="21" direction="south" minutes="40" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="11" value="-21.669722">21°40’11”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCDD0066DFD81F9A0" box="[460,609,1553,1580]" degrees="115" direction="east" minutes="53" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="13" value="115.88694">115°53’13”E</geoCoordinate>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFCE71066DFCECF9A7" box="[621,780,1553,1579]" pageId="28" pageNumber="29">M. Greenham</collectorName>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFCF0A066DFC72F9A7" box="[790,914,1553,1579]" pageId="28" pageNumber="29">D. Kamien</collectorName>
|
||
and
|
||
<collectorName id="26925319FF9AFFCFCFD4066DFBD2F9A7" box="[968,1074,1553,1579]" pageId="28" pageNumber="29">L. Mould</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFF9AFFCFC85C066DFB67F9A7" box="[1088,1159,1553,1579]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<collectionCode id="ED76AE0AFF9AFFCFC891066DFB7EF9A0" box="[1165,1182,1553,1580]" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T</collectionCode>
|
||
63327) (DNA:
|
||
<collectionCode id="ED76AE0AFF9AFFCFC955066DFA9EF9A7" box="[1353,1406,1553,1579]" country="Portugal" lsid="urn:lsid:biocol.org:col:14498" name="University of Coimbra Botany Department" pageId="28" pageNumber="29" type="Herbarium">COI</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF9AFFCFCCDB064AFE04F96C" blockId="28.[151,1437,1516,1940]" pageId="28" pageNumber="29">
|
||
<materialsCitation id="3B0F3C92FF9AFFCFCCDB064AFC7FF9C3" ID-GBIF-Occurrence="3013449435" box="[199,927,1589,1615]" collectorName="All & Mesa A" country="Australia" location="Western Australia" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<emphasis id="B913EADDFF9AFFCFCCDB064AFED9F9C3" box="[199,313,1590,1615]" italics="true" pageId="28" pageNumber="29">
|
||
<typeStatus id="54DC886DFF9AFFCFCCDB064AFED9F9C3" box="[199,313,1590,1615]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectingCountry id="F370765FFF9AFFCFCD540649FE0BF9C3" box="[328,491,1589,1615]" name="Australia" pageId="28" pageNumber="29">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFF9AFFCFCDE4064AFD22F9C3" box="[504,706,1589,1615]" italics="true" pageId="28" pageNumber="29">
|
||
<collectingRegion id="49A3F82DFF9AFFCFCDE4064AFD22F9C3" box="[504,706,1589,1615]" country="Australia" name="Western Australia" pageId="28" pageNumber="29">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<collectorName id="26925319FF9AFFCFCEC90649FD19F9C3" box="[725,761,1589,1615]" pageId="28" pageNumber="29">All</collectorName>
|
||
from
|
||
<collectorName id="26925319FF9AFFCFCF5F0649FC7FF9C3" box="[835,927,1589,1615]" pageId="28" pageNumber="29">Mesa A</collectorName>
|
||
</materialsCitation>
|
||
:
|
||
<materialsCitation id="3B0F3C92FF9AFFCFCFB10649FA62F9F8" ID-GBIF-Occurrence="3013449342" accessionNumber="T63331" collectionCode="WAM, T" collectorName="M. Greenham & D. Kamien & Mould" country="Australia" latitude="-21.651388" location="Borehole" longLatPrecision="21" longitude="115.89333" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<specimenCount id="9D61FD46FF9AFFCFCFB10649FC39F9C3" box="[941,985,1589,1615]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EB86014FF9AFFCFCFF60649FBB1F9C3" LSID="urn:lsid:plazi:treatment:03CE87D9FF9AFFCDCC8B0487FDB6F97D:8EB86014FF9AFFCFCFF60649FBB1F9C3" box="[1002,1105,1589,1615]" country="Australia" latitude="-21.651388" longLatPrecision="21" longitude="115.89333" name="Borehole" pageId="28" pageNumber="29" stateProvince="Western Australia">Borehole</location>
|
||
2501,
|
||
<quantity id="4C9F9B2AFF9AFFCFC8BF0649FAE1F9C3" box="[1187,1281,1589,1616]" metricMagnitude="4" metricUnit="m" metricValue="4.45" pageId="28" pageNumber="29" unit="km" value="44.5">44.5 km</quantity>
|
||
W of Panna- wonica,
|
||
<geoCoordinate id="EE535008FF9AFFCFCCEB0625FE8FF9F8" box="[247,367,1625,1652]" degrees="21" direction="south" minutes="39" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="5" value="-21.651388">21°39’5”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCD600625FDF1F9F8" box="[380,529,1625,1652]" degrees="115" direction="east" minutes="53" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="36" value="115.89333">115°53’36”E</geoCoordinate>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFCE020625FD48F9FF" box="[542,680,1625,1651]" pageId="28" pageNumber="29">March–May</collectorName>
|
||
2005,
|
||
<collectorName id="26925319FF9AFFCFCEEB0625FC77F9FF" box="[759,919,1625,1651]" pageId="28" pageNumber="29">M. Greenham</collectorName>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFCFBF0625FBFFF9FF" box="[931,1055,1625,1651]" pageId="28" pageNumber="29">D. Kamien</collectorName>
|
||
and L.
|
||
<collectorName id="26925319FF9AFFCFC86A0625FB21F9FF" box="[1142,1217,1625,1651]" pageId="28" pageNumber="29">Mould</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFF9AFFCFC8CC0625FAF7F9FF" box="[1232,1303,1625,1651]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<collectionCode id="ED76AE0AFF9AFFCFC9020625FAD0F9F8" box="[1310,1328,1625,1652]" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T</collectionCode>
|
||
63331)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FF9AFFCFC9920625FE2DF930" ID-GBIF-Occurrence="3013449364" accessionNumber="T63311" collectingDate="2004-12-21" collectionCode="WAM, T" collectorName="G. Humphreys & M. Greenham" country="Australia" latitude="-21.665833" location="Borehole" longLatPrecision="21" longitude="115.89333" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<specimenCount id="9D61FD46FF9AFFCFC9920625FF4CF91B" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EB86014FF9AFFCFCCA60601FEC1F91B" LSID="urn:lsid:plazi:treatment:03CE87D9FF9AFFCDCC8B0487FDB6F97D:8EB86014FF9AFFCFCCA60601FEC1F91B" box="[186,289,1661,1687]" country="Australia" latitude="-21.665833" longLatPrecision="21" longitude="115.89333" name="Borehole" pageId="28" pageNumber="29" stateProvince="Western Australia">Borehole</location>
|
||
2501,
|
||
<locationDeviation id="9A6A2FEDFF9AFFCFCD720601FD48F91B" box="[366,680,1661,1688]" location="Pannawonica" pageId="28" pageNumber="29">
|
||
<quantity id="4C9F9B2AFF9AFFCFCD720601FE2AF91B" box="[366,458,1661,1688]" metricMagnitude="4" metricUnit="m" metricValue="4.45" pageId="28" pageNumber="29" unit="km" value="44.5">44.5 km</quantity>
|
||
W of Pannawonica
|
||
</locationDeviation>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCEAF0601FCDAF914" box="[691,826,1661,1688]" degrees="21" direction="south" minutes="39" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="57" value="-21.665833">21°39’57”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCF590601FC3AF914" box="[837,986,1661,1688]" degrees="115" direction="east" minutes="53" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="36" value="115.89333">115°53’36”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF9AFFCFCFFA0601FB5EF91B" box="[998,1214,1661,1687]" pageId="28" pageNumber="29" value="2004-12-21">
|
||
<collectingDate id="EF9DE9E7FF9AFFCFCFFA0601FB5EF91B" box="[998,1214,1661,1687]" pageId="28" pageNumber="29" value="2004-12-21">21 December 2004</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFC8D50601FA92F91B" box="[1225,1394,1661,1687]" pageId="28" pageNumber="29">G. Humphreys</collectorName>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFC9600601FEECF937" pageId="28" pageNumber="29">M. Greenham</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFF9AFFCFCD0706DDFE82F937" box="[283,354,1697,1723]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<collectionCode id="ED76AE0AFF9AFFCFCD7606DDFE9CF930" box="[362,380,1697,1724]" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T</collectionCode>
|
||
63311)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FF9AFFCFCDC506DDFE00F96C" ID-GBIF-Occurrence="3013449427" accessionNumber="T66235" collectingDate="2005-07-25" collectingDateMax="2005-09-08" collectingDateMin="2005-07-25" collectionCode="WAM, T" country="Australia" latitude="-21.67861" location="Borehole" longLatPrecision="21" longitude="115.873055" pageId="28" pageNumber="29" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia" typeStatus="paratype">
|
||
<specimenCount id="9D61FD46FF9AFFCFCDC506DDFDE5F937" box="[473,517,1697,1723]" pageId="28" pageNumber="29" type="female">1 ♀</specimenCount>
|
||
,
|
||
<location id="8EB86014FF9AFFCFCE0806DDFD9BF937" LSID="urn:lsid:plazi:treatment:03CE87D9FF9AFFCDCC8B0487FDB6F97D:8EB86014FF9AFFCFCE0806DDFD9BF937" box="[532,635,1697,1723]" country="Australia" latitude="-21.67861" longLatPrecision="21" longitude="115.873055" name="Borehole" pageId="28" pageNumber="29" stateProvince="Western Australia">Borehole</location>
|
||
3073,
|
||
<geoCoordinate id="EE535008FF9AFFCFCED606DEFCADF931" box="[714,845,1698,1725]" degrees="21" direction="south" minutes="40" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="43" value="-21.67861">21°40′43″S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCF4606DEFC0BF931" box="[858,1003,1698,1725]" degrees="115" direction="east" minutes="52" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="23" value="115.873055">115°52′23″E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF9AFFCFCFE506DDFACBF930" box="[1017,1323,1697,1724]" pageId="28" pageNumber="29" value="2005-07-25" valueMax="2005-09-08" valueMin="2005-07-25">
|
||
<collectingDate id="EF9DE9E7FF9AFFCFCFE506DDFACBF930" box="[1017,1323,1697,1724]" pageId="28" pageNumber="29" value="2005-07-25" valueMax="2005-09-08" valueMin="2005-07-25">25 July–8 September 2005</collectingDate>
|
||
</date>
|
||
, G. Hum- phreys
|
||
<emphasis id="B913EADDFF9AFFCFCCF506BAFEFAF953" box="[233,282,1733,1759]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
. (
|
||
<collectionCode id="ED76AE0AFF9AFFCFCD3306B9FE96F953" box="[303,374,1733,1759]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<collectionCode id="ED76AE0AFF9AFFCFCD6006B9FE6EF96C" box="[380,398,1733,1760]" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T</collectionCode>
|
||
66235)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF9AFFCFCCDB0695FA6FF81F" blockId="28.[151,1437,1516,1940]" pageId="28" pageNumber="29">
|
||
<emphasis id="B913EADDFF9AFFCFCCDB0695FE99F88F" box="[199,377,1769,1795]" italics="true" pageId="28" pageNumber="29">Other material.</emphasis>
|
||
<materialsCitation id="3B0F3C92FF9AFFCFCD9E0695FBC0F8A4" ID-GBIF-Occurrence="3013449411" collectingDate="2016-06-29" collectionCode="MWH, WAM" collectorName="S. Lange & Global" country="Australia" latitude="-21.662222" location="Pannawonica" longLatPrecision="21" longitude="115.888885" municipality="Mesa" pageId="28" pageNumber="29" specimenCode="LN5816, T140985" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<collectingCountry id="F370765FFF9AFFCFCD9E0695FDC5F88F" box="[386,549,1769,1795]" name="Australia" pageId="28" pageNumber="29">AUSTRALIA</collectingCountry>
|
||
:
|
||
<emphasis id="B913EADDFF9AFFCFCE2F0696FD1EF88F" box="[563,766,1769,1795]" italics="true" pageId="28" pageNumber="29">
|
||
<collectingRegion id="49A3F82DFF9AFFCFCE2F0696FD1EF88F" box="[563,766,1769,1795]" country="Australia" name="Western Australia" pageId="28" pageNumber="29">Western Australia</collectingRegion>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="9D61FD46FF9AFFCFCF0C0695FCA0F88F" box="[784,832,1769,1795]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FF9AFFCFCF530695FC6CF88F" box="[847,908,1769,1795]" pageId="28" pageNumber="29">Mesa</collectingMunicipality>
|
||
A,
|
||
<quantity id="4C9F9B2AFF9AFFCFCFA70695FBE5F88F" box="[955,1029,1769,1796]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="28" pageNumber="29" unit="km" value="15.0">15 km</quantity>
|
||
SW. of
|
||
<location id="8EB86014FF9AFFCFC8760695FAE2F88F" LSID="urn:lsid:plazi:treatment:03CE87D9FF9AFFCDCC8B0487FDB6F97D:8EB86014FF9AFFCFC8760695FAE2F88F" box="[1130,1282,1769,1795]" country="Australia" latitude="-21.662222" longLatPrecision="21" longitude="115.888885" municipality="Mesa" name="Pannawonica" pageId="28" pageNumber="29" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFC90D0695FA77F88F" box="[1297,1431,1769,1795]" degrees="21" direction="south" minutes="39" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="44" value="-21.662222">21°39’44”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCC8B0771FECCF8A4" box="[151,300,1805,1832]" degrees="115" direction="east" minutes="53" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="20" value="115.888885">115°53’20”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF9AFFCFCD240771FE2FF8AB" box="[312,463,1805,1831]" pageId="28" pageNumber="29" value="2016-06-29">
|
||
<collectingDate id="EF9DE9E7FF9AFFCFCD240771FE2FF8AB" box="[312,463,1805,1831]" pageId="28" pageNumber="29" value="2016-06-29">29 June 2016</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFCDC50771FDDCF8AB" box="[473,572,1805,1831]" pageId="28" pageNumber="29">S. Lange</collectorName>
|
||
(
|
||
<collectionCode id="ED76AE0AFF9AFFCFCE560771FD73F8AB" box="[586,659,1805,1831]" pageId="28" pageNumber="29">MWH</collectionCode>
|
||
<collectorName id="26925319FF9AFFCFCE860771FD06F8AB" box="[666,742,1805,1831]" pageId="28" pageNumber="29">Global</collectorName>
|
||
<specimenCode id="DBC19EB4FF9AFFCFCEF10771FCACF8A4" box="[749,844,1805,1832]" collectionCode="LN" pageId="28" pageNumber="29">LN5816</specimenCode>
|
||
) (
|
||
<collectionCode id="ED76AE0AFF9AFFCFCF7D0771FC48F8AB" box="[865,936,1805,1831]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FF9AFFCFCFB20771FBF8F8A4" box="[942,1048,1805,1832]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T140985</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FF9AFFCFC8360771FEBCF8E3" ID-GBIF-Occurrence="3013449413" collectingDate="2010-10-07" collectionCode="WAM" collectorName="Mesa A & S. Lange & Biota Environmental Sciences" country="Australia" latitude="-21.656666" longLatPrecision="21" longitude="115.89305" pageId="28" pageNumber="29" specimenCode="MEARC4284, T3, T110813" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D61FD46FF9AFFCFC8360771FBB6F8AB" box="[1066,1110,1805,1831]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFC87D0771FB5AF8AB" box="[1121,1210,1805,1831]" pageId="28" pageNumber="29">Mesa A</collectorName>
|
||
,
|
||
<quantity id="4C9F9B2AFF9AFFCFC8D90771FAC0F8AB" box="[1221,1312,1805,1831]" metricMagnitude="4" metricUnit="m" metricValue="4.47" pageId="28" pageNumber="29" unit="km" value="44.7">44.7 km</quantity>
|
||
W. of Pan- nawonica,
|
||
<geoCoordinate id="EE535008FF9AFFCFCD13074DFE75F8C7" box="[271,405,1841,1867]" degrees="21" direction="south" minutes="39" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="24" value="-21.656666">21°39’24”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCD83074DFDD5F8C0" box="[415,565,1841,1868]" degrees="115" direction="east" minutes="53" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="35" value="115.89305">115°53’35”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF9AFFCFCE23074DFD0CF8C7" box="[575,748,1841,1867]" pageId="28" pageNumber="29" value="2010-10-07">
|
||
<collectingDate id="EF9DE9E7FF9AFFCFCE23074DFD0CF8C7" box="[575,748,1841,1867]" pageId="28" pageNumber="29" value="2010-10-07">7 October 2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFCEEA074DFCB7F8C7" box="[758,855,1841,1867]" pageId="28" pageNumber="29">S. Lange</collectorName>
|
||
(
|
||
<collectorName id="26925319FF9AFFCFCF78074DFB55F8C7" box="[868,1205,1841,1867]" pageId="28" pageNumber="29">Biota Environmental Sciences</collectorName>
|
||
<specimenCode id="DBC19EB4FF9AFFCFC8A7074DFABFF8C7" box="[1211,1375,1841,1867]" collectionCode="MEARC" pageId="28" pageNumber="29">MEARC4284</specimenCode>
|
||
P1
|
||
<specimenCode id="DBC19EB4FF9AFFCFC96B074DFA75F8C7" box="[1399,1429,1841,1867]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T3</specimenCode>
|
||
) (
|
||
<collectionCode id="ED76AE0AFF9AFFCFCC820729FF05F8E3" box="[158,229,1877,1903]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FF9AFFCFCCF70729FEB4F8E3" box="[235,340,1877,1903]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T110813</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B0F3C92FF9AFFCFCD7B0729FB89F818" ID-GBIF-Occurrence="3013449336" collectingDate="2012-05-22" collectingDateMax="2012-07-19" collectingDateMin="2012-05-22" collectionCode="WAM" collectorName="J. Alexander & Biota Environmental Sciences" country="Australia" latitude="-21.656944" location="Pannawonica" longLatPrecision="21" longitude="115.90528" municipality="Mesa" pageId="28" pageNumber="29" specimenCode="MEARC4291-0712, T1-1, T127076" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
|
||
<specimenCount id="9D61FD46FF9AFFCFCD7B0729FE72F8E3" box="[359,402,1877,1903]" pageId="28" pageNumber="29" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingMunicipality id="6BBCACB5FF9AFFCFCD820729FE3BF8E3" box="[414,475,1877,1903]" pageId="28" pageNumber="29">Mesa</collectingMunicipality>
|
||
A,
|
||
<quantity id="4C9F9B2AFF9AFFCFCE1D0729FD8BF8E3" box="[513,619,1877,1904]" metricMagnitude="4" metricUnit="m" metricValue="4.345" pageId="28" pageNumber="29" unit="km" value="43.45">43.45 km</quantity>
|
||
W. of
|
||
<location id="8EB86014FF9AFFCFCEA80729FCACF8E3" LSID="urn:lsid:plazi:treatment:03CE87D9FF9AFFCDCC8B0487FDB6F97D:8EB86014FF9AFFCFCEA80729FCACF8E3" box="[692,844,1877,1903]" country="Australia" latitude="-21.656944" longLatPrecision="21" longitude="115.90528" municipality="Mesa" name="Pannawonica" pageId="28" pageNumber="29" stateProvince="Western Australia">Pannawonica</location>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCF4B0729FC3EF8FC" box="[855,990,1877,1904]" degrees="21" direction="south" minutes="39" orientation="latitude" pageId="28" pageNumber="29" precision="15" seconds="25" value="-21.656944">21°39’25”S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE535008FF9AFFCFCFF50729FB9EF8FC" box="[1001,1150,1877,1904]" degrees="115" direction="east" minutes="54" orientation="longitude" pageId="28" pageNumber="29" precision="15" seconds="19" value="115.90528">115°54’19”E</geoCoordinate>
|
||
,
|
||
<date id="FFD9100FFF9AFFCFC8950729FA60F8E3" box="[1161,1408,1877,1903]" pageId="28" pageNumber="29" value="2012-05-22" valueMax="2012-07-19" valueMin="2012-05-22">
|
||
<collectingDate id="EF9DE9E7FF9AFFCFC8950729FA60F8E3" box="[1161,1408,1877,1903]" pageId="28" pageNumber="29" value="2012-05-22" valueMax="2012-07-19" valueMin="2012-05-22">22 May–19 July 2012</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26925319FF9AFFCFC9960729FEECF81F" pageId="28" pageNumber="29">J. Alexander</collectorName>
|
||
(
|
||
<collectorName id="26925319FF9AFFCFCD060705FD8FF81F" box="[282,623,1913,1939]" pageId="28" pageNumber="29">Biota Environmental Sciences</collectorName>
|
||
<specimenCode id="DBC19EB4FF9AFFCFCE6A0705FC82F81F" box="[630,866,1913,1939]" collectionCode="MEARC" pageId="28" pageNumber="29">MEARC4291-0712</specimenCode>
|
||
-
|
||
<specimenCode id="DBC19EB4FF9AFFCFCF750705FC74F81F" box="[873,916,1913,1939]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T1-1</specimenCode>
|
||
) (
|
||
<collectionCode id="ED76AE0AFF9AFFCFCFB60705FC11F81F" box="[938,1009,1913,1939]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="28" pageNumber="29" type="Museum">WAM</collectionCode>
|
||
<specimenCode id="DBC19EB4FF9AFFCFCFEB0705FB81F818" box="[1015,1121,1913,1940]" collectionCode="T" name="Tavera, Department of Geology and Geophysics" pageId="28" pageNumber="29">T127076</specimenCode>
|
||
)
|
||
</materialsCitation>
|
||
; see
|
||
<bibRefCitation id="EFF64B3EFF9AFFCFC8820705FA6AF81F" author="Abrams, K. M. & Huey, J. A. & Hillyer, M. J. & Humphreys, W. F. & Didham, R. K. & Harvey, M. S." box="[1182,1418,1913,1939]" pageId="28" pageNumber="29" pagination="106532" refId="ref46122" refString="Abrams, K. M., Huey, J. A., Hillyer, M. J., Humphreys, W. F., Didham, R. K. & Harvey, M. S. (2019) Too hot to handle: Tertiary aridification drives multiple independent incursions of Schizomida (Hubbardiidae) into hypogean environments. Molecular Phylogenetics and Evolution, 139, 106532. https: // doi. org / 10.1016 / j. ympev. 2019.106532" type="journal article" year="2019">
|
||
Abrams
|
||
<emphasis id="B913EADDFF9AFFCFC8E10706FACFF81F" box="[1277,1327,1913,1939]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
|
||
. (2019)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF186647FF9BFFCECC8B0530FC69FA20" ID-DOI="http://doi.org/10.5281/zenodo.4416877" ID-Zenodo-Dep="4416877" httpUri="https://zenodo.org/record/4416877/files/figure.png" pageId="29" pageNumber="30" startId="29.[151,250,1356,1380]" targetBox="[152,1435,181,1331]" targetPageId="29">
|
||
<paragraph id="8BD836CFFF9BFFCECC8B0530FC69FA20" blockId="29.[151,1437,1356,1453]" pageId="29" pageNumber="30">
|
||
<emphasis id="B913EADDFF9BFFCECC8B0530FEFFFAE8" bold="true" box="[151,287,1356,1380]" pageId="29" pageNumber="30">FIGURE 11.</emphasis>
|
||
<taxonomicName id="4C674D4CFF9BFFCECD3A0530FDC6FAE8" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[294,550,1356,1380]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF9BFFCECD3A0530FDC6FAE8" box="[294,550,1356,1380]" italics="true" pageId="29" pageNumber="30">Draculoides anachoretus</emphasis>
|
||
</taxonomicName>
|
||
male (WAM T140985): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 2 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37D6544FF9BFFCECCDB05A6FB8AF844" pageId="29" pageNumber="30" type="diagnosis">
|
||
<paragraph id="8BD836CFFF9BFFCECCDB05A6FB8AF844" blockId="29.[151,1437,1498,2029]" pageId="29" pageNumber="30">
|
||
<emphasis id="B913EADDFF9BFFCECCDB05A6FEA2FA78" bold="true" box="[199,322,1498,1524]" pageId="29" pageNumber="30">Diagnosis.</emphasis>
|
||
The shape of the male flagellum of
|
||
<taxonomicName id="4C674D4CFF9BFFCECEC705A6FC16FA78" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[731,1014,1498,1524]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF9BFFCECEC705A6FC16FA78" box="[731,1014,1498,1524]" italics="true" pageId="29" pageNumber="30">Draculoides anachoretus</emphasis>
|
||
</taxonomicName>
|
||
most closely resembles
|
||
<taxonomicName id="4C674D4CFF9BFFCEC91605A7FA93FA78" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1290,1395,1498,1524]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B913EADDFF9BFFCEC91605A7FA93FA78" box="[1290,1395,1498,1524]" italics="true" pageId="29" pageNumber="30">D. affinis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCEC99D05A7FF08F994" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bythius">
|
||
<emphasis id="B913EADDFF9BFFCEC99D05A7FF08F994" italics="true" pageId="29" pageNumber="30">D. bythius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECCEA0583FE6DF994" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[246,397,1534,1560]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cochranus">
|
||
<emphasis id="B913EADDFF9BFFCECCEA0583FE6DF994" box="[246,397,1534,1560]" italics="true" pageId="29" pageNumber="30">D. cochranus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECD870583FDF5F994" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[411,533,1535,1560]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="eremius">
|
||
<emphasis id="B913EADDFF9BFFCECD870583FDF5F994" box="[411,533,1535,1560]" italics="true" pageId="29" pageNumber="30">D. eremius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECE3E0583FD23F994" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[546,707,1534,1560]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="gnophicola">
|
||
<emphasis id="B913EADDFF9BFFCECE3E0583FD23F994" box="[546,707,1534,1560]" italics="true" pageId="29" pageNumber="30">D. gnophicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECECD0583FCA5F994" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[721,837,1534,1560]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kryptus">
|
||
<emphasis id="B913EADDFF9BFFCECECD0583FCA5F994" box="[721,837,1534,1560]" italics="true" pageId="29" pageNumber="30">D. kryptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECF4F0583FBC6F994" authorityName="Abrams and Harvey" authorityYear="2020" box="[851,1062,1534,1560]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mckechnieorum">
|
||
<emphasis id="B913EADDFF9BFFCECF4F0583FBC6F994" box="[851,1062,1534,1560]" italics="true" pageId="29" pageNumber="30">D. mckechnieorum</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B913EADDFF9BFFCEC8410583FAE8F994" box="[1117,1288,1534,1560]" italics="true" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C674D4CFF9BFFCEC8410583FAE5F994" authorityName="Abrams and Harvey" authorityYear="2020" box="[1117,1285,1534,1560]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="warramboo">D. warramboo</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
especially in the presence of a broad base. It differs from
|
||
<emphasis id="B913EADDFF9BFFCECE67065FFAF1F9B0" box="[635,1297,1570,1596]" italics="true" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C674D4CFF9BFFCECE67065FFD05F9B0" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[635,741,1570,1596]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="affinis">D. affinis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECEF2065FFC83F9B0" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[750,867,1570,1596]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bythius">D. bythius</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECF71065FFC08F9B0" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[877,1000,1571,1596]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECFEE065FFB74F9B0" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1010,1172,1570,1596]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCEC882065FFAF1F9B0" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[1182,1297,1570,1596]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kryptus">D. kryptus</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C674D4CFF9BFFCEC959065FFEFBF9EC" authorityName="Abrams and Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mckechnieorum">
|
||
<emphasis id="B913EADDFF9BFFCEC959065FFEFBF9EC" italics="true" pageId="29" pageNumber="30">D. mckechnieorum</emphasis>
|
||
</taxonomicName>
|
||
in the anterior position of dm4, well set back from the posterior margin (in line with dl
|
||
<quantity id="4C9F9B2AFF9BFFCEC8ED063AFAFBF9EC" box="[1265,1307,1606,1632]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="29" pageNumber="30" unit="in" value="3.0">3 in</quantity>
|
||
<taxonomicName id="4C674D4CFF9BFFCEC93E063BFA75F9EC" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1314,1429,1606,1632]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bythius">
|
||
<emphasis id="B913EADDFF9BFFCEC93E063BFA75F9EC" box="[1314,1429,1606,1632]" italics="true" pageId="29" pageNumber="30">D. bythius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B913EADDFF9BFFCECC8B0617FE20F908" box="[151,448,1642,1668]" italics="true" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C674D4CFF9BFFCECC8B0617FEDAF908" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,314,1642,1668]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="gnophicola">D. gnophicola</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECD590617FE20F908" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[325,448,1643,1668]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="eremius">D. eremius</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="B913EADDFF9BFFCECDEA0617FD91F908" box="[502,625,1642,1668]" italics="true" pageId="29" pageNumber="30">
|
||
<taxonomicName id="4C674D4CFF9BFFCECDEA0617FD8DF908" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[502,621,1642,1668]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kryptus">D. kryptus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
slightly anterior to dl3 and close to posterior margin in
|
||
<taxonomicName id="4C674D4CFF9BFFCEC8FE0617FAABF908" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[1250,1355,1642,1668]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="affinis">
|
||
<emphasis id="B913EADDFF9BFFCEC8FE0617FAABF908" box="[1250,1355,1642,1668]" italics="true" pageId="29" pageNumber="30">D. affinis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C674D4CFF9BFFCEC99D0617FEA8F924" authorityName="Abrams and Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mckechnieorum">
|
||
<emphasis id="B913EADDFF9BFFCEC99D0617FEA8F924" italics="true" pageId="29" pageNumber="30">D. mckechnieorum</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName id="4C674D4CFF9BFFCECE1906F3FD7CF924" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[517,668,1678,1704]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cochranus">
|
||
<emphasis id="B913EADDFF9BFFCECE1906F3FD7CF924" box="[517,668,1678,1704]" italics="true" pageId="29" pageNumber="30">D. cochranus</emphasis>
|
||
</taxonomicName>
|
||
by the posterior position of dm
|
||
<quantity id="4C9F9B2AFF9BFFCECFE006F2FBC7F924" box="[1020,1063,1678,1704]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="29" pageNumber="30" unit="in" value="4.0">4 in</quantity>
|
||
relation to dl1 (dm4 is level with dl
|
||
<quantity id="4C9F9B2AFF9BFFCECCAD06CEFF39F940" box="[177,217,1714,1740]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="30" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C674D4CFF9BFFCECCFE06CFFE9AF940" baseAuthorityName="Framenau, Hamilton, Finston, Humphreys, Abrams, Huey and Harvey" baseAuthorityYear="2018" box="[226,378,1714,1740]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="cochranus">
|
||
<emphasis id="B913EADDFF9BFFCECCFE06CFFE9AF940" box="[226,378,1714,1740]" italics="true" pageId="29" pageNumber="30">D. cochranus</emphasis>
|
||
</taxonomicName>
|
||
) and from
|
||
<taxonomicName id="4C674D4CFF9BFFCECDE006CFFD42F940" authorityName="Abrams and Harvey" authorityYear="2020" box="[508,674,1714,1740]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="warramboo">
|
||
<emphasis id="B913EADDFF9BFFCECDE006CFFD42F940" box="[508,674,1714,1740]" italics="true" pageId="29" pageNumber="30">D. warramboo</emphasis>
|
||
</taxonomicName>
|
||
in the anterior position of vm
|
||
<quantity id="4C9F9B2AFF9BFFCECFE706CEFBC6F940" box="[1019,1062,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="29" pageNumber="30" unit="in" value="5.0">5 in</quantity>
|
||
relation to dl1 (vm
|
||
<quantity id="4C9F9B2AFF9BFFCEC91906CEFAD0F940" box="[1285,1328,1714,1741]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="29" pageNumber="30" unit="in" value="5.0">5 in</quantity>
|
||
line with dl
|
||
<quantity id="4C9F9B2AFF9BFFCECCAD06AAFF38F97C" box="[177,216,1750,1776]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="29" pageNumber="30" unit="in" value="1.0">1 in</quantity>
|
||
<taxonomicName id="4C674D4CFF9BFFCECCC206ABFE63F97C" authorityName="Abrams and Harvey" authorityYear="2020" box="[222,387,1750,1776]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="warramboo">
|
||
<emphasis id="B913EADDFF9BFFCECCC206ABFE63F97C" box="[222,387,1750,1776]" italics="true" pageId="29" pageNumber="30">D. warramboo</emphasis>
|
||
</taxonomicName>
|
||
). Females are most similar to
|
||
<taxonomicName id="4C674D4CFF9BFFCECEC906ABFC90F97C" authorityName="Abrams and Harvey" authorityYear="2020" box="[725,880,1750,1776]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFF9BFFCECEC906ABFC90F97C" box="[725,880,1750,1776]" italics="true" pageId="29" pageNumber="30">D. immortalis</emphasis>
|
||
</taxonomicName>
|
||
but differ in the position of dm1 of the flagellum, which is slightly anterior to the annulus separating flagellomeres II and III in
|
||
<taxonomicName id="4C674D4CFF9BFFCEC81C0687FB4CF898" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1024,1196,1786,1812]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF9BFFCEC81C0687FB4CF898" box="[1024,1196,1786,1812]" italics="true" pageId="29" pageNumber="30">D. anachoretus</emphasis>
|
||
</taxonomicName>
|
||
and very close to the annulus in
|
||
<taxonomicName id="4C674D4CFF9BFFCECD0F0763FE4EF8B4" authorityName="Abrams and Harvey" authorityYear="2020" box="[275,430,1822,1848]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFF9BFFCECD0F0763FE4EF8B4" box="[275,430,1822,1848]" italics="true" pageId="29" pageNumber="30">D. immortalis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C674D4CFF9BFFCECDA10762FD38F8B4" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[445,728,1822,1848]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF9BFFCECDA10762FD38F8B4" box="[445,728,1822,1848]" italics="true" pageId="29" pageNumber="30">Draculoides anachoretus</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFF9BFFCEC8560762FB34F8B4" authorityName="Harvey" authorityYear="1992" box="[1098,1236,1822,1848]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFF9BFFCEC8560762FB34F8B4" box="[1098,1236,1822,1848]" italics="true" pageId="29" pageNumber="30">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at COI and 12S by the 50bp mini-barcodes shown in
|
||
<figureCitation id="135C2A4AFF9BFFCECF76073EFC57F8D1" box="[874,951,1858,1885]" captionStart="FIGURE 3" captionStartId="18.[151,250,1547,1571]" captionTargetBox="[158,1420,181,1521]" captionTargetId="figure-20@18.[151,1436,181,1524]" captionTargetPageId="18" captionText="FIGURE 3. COI mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913085). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416857" httpUri="https://zenodo.org/record/4416857/files/figure.png" pageId="29" pageNumber="30">Figs. 3</figureCitation>
|
||
and
|
||
<figureCitation id="135C2A4AFF9BFFCECFF2073EFC1EF8D1" box="[1006,1022,1858,1885]" captionStart="FIGURE 5" captionStartId="20.[151,250,1216,1240]" captionTargetBox="[158,1424,183,1189]" captionTargetId="figure-20@20.[151,1436,181,1192]" captionTargetPageId="20" captionText="FIGURE 5. 12S mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913012). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416863" httpUri="https://zenodo.org/record/4416863/files/figure.png" pageId="29" pageNumber="30">5</figureCitation>
|
||
.
|
||
<taxonomicName id="4C674D4CFF9BFFCEC817073EFAC5F8D0" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1035,1317,1858,1884]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF9BFFCEC817073EFAC5F8D0" box="[1035,1317,1858,1884]" italics="true" pageId="29" pageNumber="30">Draculoides anachoretus</emphasis>
|
||
</taxonomicName>
|
||
can be diagnosed from all other
|
||
<taxonomicName id="4C674D4CFF9BFFCECD87071AFDC5F80C" authorityName="Harvey" authorityYear="1992" box="[411,549,1894,1920]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B913EADDFF9BFFCECD87071AFDC5F80C" box="[411,549,1894,1920]" italics="true" pageId="29" pageNumber="30">Draculoides</emphasis>
|
||
</taxonomicName>
|
||
species that were sequenced at ITS2 (
|
||
<figureCitation id="135C2A4AFF9BFFCECFCC071AFBF3F80D" box="[976,1043,1894,1921]" captionStart="FIGURE 7" captionStartId="22.[151,250,1755,1779]" captionTargetBox="[159,1425,181,1729]" captionTargetId="figure-20@22.[151,1436,181,1732]" captionTargetPageId="22" captionText="FIGURE 7. ITS2 mini-barcodes for all Draculoides species including the operational taxonomic units used in Abrams et al. (2019). This barcode is the maximally diagnostic 50 bp fragment of DNA. Genetic variation within species is shown using IUPAC Ambiguity Codes. See Methods for details. The position of the mini-barcode is reported relative to a specimen of Draculoides celatus (WAM T98698, GenBank number MG913105). The blue tree on the left groups minibarcodes which are most similar to each other but does not represent an accurate phylogeny. The coloured bases in the figure are those that differ from the consensus reference sequence." figureDoi="http://doi.org/10.5281/zenodo.4416867" httpUri="https://zenodo.org/record/4416867/files/figure.png" pageId="29" pageNumber="30">Fig. 7</figureCitation>
|
||
) except for
|
||
<taxonomicName id="4C674D4CFF9BFFCEC886071BFAEEF80C" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[1178,1294,1894,1920]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="bythius">
|
||
<emphasis id="B913EADDFF9BFFCEC886071BFAEEF80C" box="[1178,1294,1894,1920]" italics="true" pageId="29" pageNumber="30">D. bythius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCEC907071BFA76F80C" baseAuthorityName="Abrams and Harvey" baseAuthorityYear="2015" box="[1307,1430,1895,1920]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="eremius">
|
||
<emphasis id="B913EADDFF9BFFCEC907071BFA76F80C" box="[1307,1430,1895,1920]" italics="true" pageId="29" pageNumber="30">D. eremius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECC8B07F7FED7F828" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[151,311,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="gnophicola">
|
||
<emphasis id="B913EADDFF9BFFCECC8B07F7FED7F828" box="[151,311,1930,1956]" italics="true" pageId="29" pageNumber="30">D. gnophicola</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECD5807F7FE57F828" authorityName="Harvey, Berry, Edward and Humphreys" authorityYear="2008" box="[324,439,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Paradraculoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="kryptus">
|
||
<emphasis id="B913EADDFF9BFFCECD5807F7FE57F828" box="[324,439,1930,1956]" italics="true" pageId="29" pageNumber="30">D. kryptus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECDD907F7FD77F828" authorityName="Abrams and Harvey" authorityYear="2020" box="[453,663,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="mckechnieorum">
|
||
<emphasis id="B913EADDFF9BFFCECDD907F7FD77F828" box="[453,663,1930,1956]" italics="true" pageId="29" pageNumber="30">D. mckechnieorum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECEB907F7FCA9F828" authorityName="Abrams and Harvey" authorityYear="2020" box="[677,841,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="warramboo">
|
||
<emphasis id="B913EADDFF9BFFCECEB907F7FCA9F828" box="[677,841,1930,1956]" italics="true" pageId="29" pageNumber="30">D. warramboo</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECF4A07F7FC10F828" authorityName="Abrams and Harvey" authorityYear="2020" box="[854,1008,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="immortalis">
|
||
<emphasis id="B913EADDFF9BFFCECF4A07F7FC10F828" box="[854,1008,1930,1956]" italics="true" pageId="29" pageNumber="30">D. immortalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCECFE207F7FB7BF828" authorityName="Abrams and Harvey" authorityYear="2020" box="[1022,1179,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="belalugosii">
|
||
<emphasis id="B913EADDFF9BFFCECFE207F7FB7BF828" box="[1022,1179,1930,1956]" italics="true" pageId="29" pageNumber="30">D. belalugosii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCEC8B507F7FA94F828" authorityName="Abrams and Harvey" authorityYear="2020" box="[1193,1396,1930,1956]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="christopherleei">
|
||
<emphasis id="B913EADDFF9BFFCEC8B507F7FA94F828" box="[1193,1396,1930,1956]" italics="true" pageId="29" pageNumber="30">D. christopherleei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C674D4CFF9BFFCEC99D07F7FEEFF844" authorityName="Abrams and Harvey" authorityYear="2020" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="piscivultus">
|
||
<emphasis id="B913EADDFF9BFFCEC99D07F7FEEFF844" italics="true" pageId="29" pageNumber="30">D. piscivultus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C674D4CFF9BFFCECD5907D3FE24F844" authorityName="Abrams & Huey & Hillyer & Didham & Harvey" authorityYear="2020" box="[325,452,1966,1992]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="akashae">
|
||
<emphasis id="B913EADDFF9BFFCECD5907D3FE24F844" box="[325,452,1966,1992]" italics="true" pageId="29" pageNumber="30">D. akashae</emphasis>
|
||
</taxonomicName>
|
||
, which are not distinguishable using the ITS2 mini-barcode.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37D6544FF9BFFCDCCDB07AEFD13FC29" lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="description">
|
||
<paragraph id="8BD836CFFF9BFFCECCDB07AEFD4EF861" blockId="29.[151,1437,1498,2029]" box="[199,686,2002,2029]" pageId="29" pageNumber="30">
|
||
<emphasis id="B913EADDFF9BFFCECCDB07AEFEB9F860" bold="true" box="[199,345,2002,2028]" pageId="29" pageNumber="30">Description.</emphasis>
|
||
Adult male (WAM T140985).
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB00EBFC32FF3D" blockId="30.[151,1437,151,934]" box="[199,978,151,177]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB00EBFEF6FF3D" box="[199,278,151,177]" italics="true" pageId="30" pageNumber="31">Colour</emphasis>
|
||
. Yellow-brown; propeltidium and pedipalps somewhat darker.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB00C7FC60FE91" blockId="30.[151,1437,151,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB00C7FE8FFF59" box="[199,367,187,213]" italics="true" pageId="30" pageNumber="31">Cephalothorax</emphasis>
|
||
. Propeltidium with 2+1 apical setae in a triangular formation on anterior process and 2 + 1 + 2 setae; eye spots absent. Mesopeltidia separated. Metapeltidium divided. Anterior sternum with 15 setae (including 2 sternapophysial setae); posterior sternum triangular with 7 setae.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB015BFCBFFE79" blockId="30.[151,1437,151,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB015BFED5FECD" box="[199,309,295,321]" italics="true" pageId="30" pageNumber="31">Chelicera</emphasis>
|
||
. Fixed finger with 2 large teeth plus 5 smaller teeth between these, including 1 tiny lateral tooth on proximal tooth; membranous area between fixed and movable fingers with 3 large, lanceolate, terminally pilose setae (G1); G2 composed of 8 setae; G3 composed of 4 setae; internal face of chelicera with 5 short whip-like setae (G4); brush at base of fixed finger composed of 7 setae (G5A), each densely pilose in distal half and G5B composed of 10 setae; G6 with one seta; G7 composed of 3 setae. Movable finger serrula composed of 17 long lamellae, blunt guard tooth present subdistally; 1 large accessory tooth present.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB0183FA6FFDB1" blockId="30.[151,1437,151,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB0183FECCFD95" box="[199,300,511,537]" italics="true" pageId="30" pageNumber="31">Pedipalp</emphasis>
|
||
. Without apophyses; trochanter with sharply produced ventro-distal extension, ventral margin with ca. 8 stout setae, without mesal spur; tarsus and tibia without spines; tarsal spur present; claw 0.30 × length of tarsus.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB023BFB0EFDED" blockId="30.[151,1437,151,934]" box="[199,1262,583,609]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB023BFEE4FDEC" box="[199,260,583,608]" italics="true" pageId="30" pageNumber="31">Legs.</emphasis>
|
||
Tarsus I with 6 segments; baso-dorsal margin of femur IV produced at about a 90° angle.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB0217FA61FD26" blockId="30.[151,1437,151,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB0217FECFFD09" box="[199,303,619,645]" italics="true" pageId="30" pageNumber="31">Abdomen</emphasis>
|
||
. Chaetotaxy of tergites I: 2 macrosetae + 4 microsetae (microsetae diagonal), tergite II: 3 macrosetae + 6 microsetae (microsetae in column), tergites III–IX: 2: 2: 2: 2: 2: 3, segment XII with small dorsal process (
|
||
<specimenCount id="9D61FD46FF98FFCDC93802F3FAD9FD25" box="[1316,1337,655,681]" pageId="30" pageNumber="31" type="male">♂</specimenCount>
|
||
only).
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB02CFFAC3FCB5" blockId="30.[151,1437,151,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB02CFFEDAFD41" box="[199,314,691,717]" italics="true" pageId="30" pageNumber="31">Flagellum</emphasis>
|
||
. Male: Dorsoventrally compressed (
|
||
<figureCitation id="135C2A4AFF98FFCDCEC902CFFCCDFD41" box="[725,813,691,717]" captionStart="FIGURE 11" captionStartId="29.[151,250,1356,1380]" captionTargetBox="[152,1435,181,1331]" captionTargetId="figure-17@29.[151,1436,181,1332]" captionTargetPageId="29" captionText="FIGURE 11. Draculoides anachoretus male (WAM T140985): A. Body, dorsal; B. Body, ventral; C. Body, lateral; D. Flagellum, dorsal; E. Flagellum, ventral; F. Flagellum, lateral. The scale bar shown in image A is 2 mm and also applies to B and C and the scale bar shown in image D is 200 µm and also applies to E and F." figureDoi="http://doi.org/10.5281/zenodo.4416877" httpUri="https://zenodo.org/record/4416877/files/figure.png" pageId="30" pageNumber="31">Figs. 11</figureCitation>
|
||
D–F, 12A–C); 2.08 × longer than broad; seta dm1 situated dorso-medially, closer to anterior margin; seta dm4 set back from posterior margin, anterior to dl3; dl1posterior to vl1; dl3 close to posterior margin; vm2 uneven, one anterior to vm1 and one posterior to vm1; vm5 situated closer to vl2 than to vm3; left vm3 slightly anterior to right vm3; five pairs of microsetae between vl1 and dl3.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB033FFB17FC0D" blockId="30.[151,1437,151,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB033FFE6EFCD1" box="[199,398,835,861]" italics="true" pageId="30" pageNumber="31">Dimensions (mm)</emphasis>
|
||
. Male (WAM T140985): Body length 4.04. Propeltidium 1.19/0.65. Chelicera 0.73. Flagellum 0.45/0.22. Pedipalp: trochanter 0.38, femur 0.58, patella 0.60, tibia 0.42, tarsus 0.29, claw 0.09.
|
||
</paragraph>
|
||
<paragraph id="8BD836CFFF98FFCDCCDB03F7FD13FC29" blockId="30.[151,1437,151,934]" box="[199,755,907,934]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB03F7FECFFC28" box="[199,303,907,932]" italics="true" pageId="30" pageNumber="31">Variation</emphasis>
|
||
. Body length (males) 3.13–4.04 (n = 3).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF186647FF98FFCDCC8B0657FD6AF9EB" ID-DOI="http://doi.org/10.5281/zenodo.4416881" ID-Zenodo-Dep="4416881" httpUri="https://zenodo.org/record/4416881/files/figure.png" pageId="30" pageNumber="31" startId="30.[151,250,1579,1603]" targetBox="[149,1439,957,1550]" targetPageId="30">
|
||
<paragraph id="8BD836CFFF98FFCDCC8B0657FD6AF9EB" blockId="30.[151,1436,1579,1639]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCC8B0657FEC0F9CF" bold="true" box="[151,288,1579,1603]" pageId="30" pageNumber="31">FIGURE 12.</emphasis>
|
||
<taxonomicName id="4C674D4CFF98FFCDCD3B0657FDC7F9CF" baseAuthorityName="Harvey, Berry, Edward and Humphreys" baseAuthorityYear="2008" box="[295,551,1579,1603]" class="Arachnida" family="Hubbardiidae" genus="Draculoides" kingdom="Animalia" order="Schizomida" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="anachoretus">
|
||
<emphasis id="B913EADDFF98FFCDCD3B0657FDC7F9CF" box="[295,551,1579,1603]" italics="true" pageId="30" pageNumber="31">Draculoides anachoretus</emphasis>
|
||
</taxonomicName>
|
||
male (WAM T140985): A. Flagellum, dorsal; B. Flagellum, ventral; C. Flagellum, lateral. The scale bar is 0.1 mm and applies to A–C.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37D6544FF98FFCDCCDB06F3FDB6F97D" pageId="30" pageNumber="31" type="discussion">
|
||
<paragraph id="8BD836CFFF98FFCDCCDB06F3FDB6F97D" blockId="30.[151,1436,1679,1777]" pageId="30" pageNumber="31">
|
||
<emphasis id="B913EADDFF98FFCDCCDB06F3FEDBF925" bold="true" box="[199,315,1679,1705]" pageId="30" pageNumber="31">Remarks.</emphasis>
|
||
<bibRefCitation id="EFF64B3EFF98FFCDCD5D06F3FDC1F925" author="Harvey, M. S. & Berry, O. & Edward, K. L. & Humphreys, G." box="[321,545,1679,1705]" pageId="30" pageNumber="31" pagination="167 - 194" refId="ref46967" refString="Harvey, M. S., Berry, O., Edward, K. L. & Humphreys, G. (2008) Molecular and morphological systematics of hypogean schizomids (Schizomida: Hubbardiidae) in semi-arid Australia. Invertebrate Systematics, 22 (2), 167 - 194. https: // doi. org / 10.1071 / IS 07026" type="journal article" year="2008">
|
||
Harvey
|
||
<emphasis id="B913EADDFF98FFCDCD8506F3FE2AF925" box="[409,458,1679,1705]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
|
||
. (2008)
|
||
</bibRefCitation>
|
||
named this species from several females and juveniles collected from Mesa A in the Robe Valley,
|
||
<collectingRegion id="49A3F82DFF98FFCDCD4B06CFFDFFF941" box="[343,543,1715,1741]" country="Australia" name="Western Australia" pageId="30" pageNumber="31">Western Australia</collectingRegion>
|
||
(
|
||
<figureCitation id="135C2A4AFF98FFCDCE3206CFFD68F941" box="[558,648,1715,1741]" captionStart="FIGURE 1" captionStartId="16.[151,250,1868,1892]" captionTargetBox="[346,1243,188,1844]" captionTargetId="figure-20@16.[339,1249,181,1844]" captionTargetPageId="16" captionText="FIGURE 1. Map of an area of the Pilbara described as “West Pilbara” for the purposes of this study, showing the distribution of the named Draculoides species. Inset maps show species distributions in finer detail encompassing areas A: Warramboo, Mesas A, B and C and “BudgieBore”; B: Bungaroo, Mesas G, H, J, K and L; C: Cochrane and Jewell, Kens Bore, Cane and Upper Cane River, Trinity Bore and Catho Well, Mt. Stuart Station; D: Middle Robe, Robe Valley; E: Bungaroo South, West Pit, Buckland Hills and Dragon deposit." figureDoi="http://doi.org/10.5281/zenodo.4416851" httpUri="https://zenodo.org/record/4416851/files/figure.png" pageId="30" pageNumber="31">Fig. 1A</figureCitation>
|
||
). Additional specimens, including three males that are described here, have since been collected from Mesa A.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |