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<document id="8E7ECAC836984DD0DBBDA849C9FD2A0D" ID-DOI="10.11646/zootaxa.3647.2.2" ID-GBIF-Dataset="c49f48ba-b464-4b93-8269-40d6ed2fd447" ID-ISSN="1175-5326" ID-Zenodo-Dep="224159" ID-ZooBank="E917959A-ED11-47DA-ADAB-6D59F333705E" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460604624028" checkinUser="plazi" docAuthor="Barták, Miroslav &amp; Kubík, Štěpán" docDate="2013" docId="03C42E68FFBB19402FBF39F695E2D0FD" docLanguage="en" docName="zt03647p278.pdf" docOrigin="Zootaxa 3647 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bicellaria" docType="key" docVersion="8" lastPageNumber="276" masterDocId="FFFD5610FFAD19592F283C51907ED241" masterDocTitle="Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species" masterLastPageNumber="278" masterPageNumber="251" pageNumber="273" updateTime="1698327825972" updateUser="plazi">
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<mods:title id="D67E8BA7CD84850263B0CC246A471A89">Species of Bicellaria Macquart (Diptera: Hybotidae) of Europe, with descriptions of four new species</mods:title>
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<mods:namePart id="9D72FA6D376A3BB7F0544DC5B4C1C6AA">Barták, Miroslav</mods:namePart>
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<mods:namePart id="7160C5752C36F0F4D0F1FC3BA987FB32">Kubík, Štěpán</mods:namePart>
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<mods:title id="B17CFD1F27BE1F73D69AE6626FAA160A">Zootaxa</mods:title>
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<mods:date id="C17EAC55E36514F3951157E9A4D5CB0A">2013</mods:date>
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<mods:number id="5F46E2DF9098BAE727A4097D675DE025">3647</mods:number>
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<mods:identifier id="916EC54872406C5015C3AABCBBC4AFC9" type="DOI">10.11646/zootaxa.3647.2.2</mods:identifier>
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<treatment id="03C42E68FFBB19402FBF39F695E2D0FD" ID-DOI="http://doi.org/10.5281/zenodo.6148018" ID-GBIF-Taxon="119567646" ID-Zenodo-Dep="6148018" LSID="urn:lsid:plazi:treatment:03C42E68FFBB19402FBF39F695E2D0FD" httpUri="http://treatment.plazi.org/id/03C42E68FFBB19402FBF39F695E2D0FD" lastPageId="25" lastPageNumber="276" pageId="22" pageNumber="273">
<subSubSection id="C377CCF5FFBB194F2FBF39F694EBD59B" pageId="22" pageNumber="273" type="nomenclature">
<paragraph id="8BD29F7EFFBB194F2FBF39F69219D780" blockId="22.[151,615,1447,1473]" box="[151,615,1447,1473]" pageId="22" pageNumber="273">
<heading id="D09A2812FFBB194F2FBF39F69219D780" bold="true" box="[151,615,1447,1473]" fontSize="11" level="1" pageId="22" pageNumber="273" reason="1">
<emphasis id="B919436CFFBB194F2FBF39F69219D780" bold="true" box="[151,615,1447,1473]" pageId="22" pageNumber="273">
Key to species of
<taxonomicName id="4C6DE4FDFFBB194F2E4539F6919AD780" ID-CoL="62GCM" ID-ENA="192473" box="[365,484,1447,1473]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="genus">
<emphasis id="B919436CFFBB194F2E4539F6919AD780" bold="true" box="[365,484,1447,1473]" italics="true" pageId="22" pageNumber="273">Bicellaria</emphasis>
</taxonomicName>
of Europe
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD29F7EFFBB194F2FBF39BE94EBD59B" blockId="22.[151,1436,1518,2010]" pageId="22" pageNumber="273">
<emphasis id="B919436CFFBB194F2FBF39BE9175D446" bold="true" box="[151,267,1519,1543]" pageId="22" pageNumber="273">Remarks.</emphasis>
Absolute measures given in the text (performed on dry mounted material) are only benchmarks: they may slightly differ in unusually small or large individuals. Only those setae on the palpus are counted which are longer than
<quantity id="4C95329BFFBB194F2E0C3A6791F2D40F" box="[292,396,1590,1614]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="22" pageNumber="273" unit="mm" value="0.05">0.05 mm</quantity>
(those occurring on the palpifer are not counted). Long setae on dorsal side of the third antennal segment may be broken or their presence/absence may be variable (we found a single specimen of
<taxonomicName id="4C6DE4FDFFBB194F2AAC3A0D917DD4D7" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="subpilosa">
<emphasis id="B919436CFFBB194F2AAC3A0D917DD4D7" italics="true" pageId="22" pageNumber="273">B. subpilosa</emphasis>
</taxonomicName>
with long fine seta on the third antennal segment). Those setae on intrahumeral and posthumeral areas are counted which occur outside the single row of dorsocentrals in the presutural area. Prescutellar depression in some species contains differently long and more erect microtrichia giving this area lighter appearance in dorsal view, this area is mostly sharply delimited and this character state is referred in the text as follows: “prescutellar depression lighter than other parts of mesoscutum in dorsal view”. Nearly all characters sometimes vary (including precise shape of male hypandrial processes) which makes identification extremely difficult. Females are sometimes indistinguishable (most notably:
<taxonomicName id="4C6DE4FDFFBB194F2D243B09920AD52F" box="[524,628,1878,1903]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="italica">
<emphasis id="B919436CFFBB194F2D243B09920AD52F" box="[524,628,1878,1903]" italics="true" pageId="22" pageNumber="273">B. italica</emphasis>
</taxonomicName>
-
<taxonomicName id="4C6DE4FDFFBB194F2DA63B099356D52F" box="[654,808,1878,1903]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="intermedia">
<emphasis id="B919436CFFBB194F2DA63B099356D52F" box="[654,808,1878,1903]" italics="true" pageId="22" pageNumber="273">B. intermedia</emphasis>
</taxonomicName>
), thus faunistic records should be based only on males and always be supported by dissection of the genitalia. Both
<taxonomicName id="4C6DE4FDFFBB194F2C5B3B2D9473D5D2" box="[883,1037,1915,1939]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="albopilosa">
<emphasis id="B919436CFFBB194F2C5B3B2D9473D5D2" box="[883,1037,1915,1939]" italics="true" pageId="22" pageNumber="273">B. albopilosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6DE4FDFFBB194F2B663B2D94C4D5D2" box="[1102,1210,1916,1939]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="273" phylum="Arthropoda" rank="species" species="spuria">
<emphasis id="B919436CFFBB194F2B663B2D94C4D5D2" box="[1102,1210,1916,1939]" italics="true" pageId="22" pageNumber="273">B. spuria</emphasis>
</taxonomicName>
probably represent complexes of sibling species. For this reason, some recently published records should be considered carefully due to possible misidentifications (e.g. Raffone 2006, 2008; Jonassen 1984; Parvu 2003, 2005).
</paragraph>
</subSubSection>
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<paragraph id="8BD29F7EFFBA194E2FBF3CC695E5D345" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
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1 Antennal stylus together with slender portion of third segment half as long as broad basal portion of third segment (
<figureCitation id="135683FBFFBA194E2A5D3CC6915ED28B" captionStart="FIGURES 41 48. 41 44" captionStartId="20.[151,264,1709,1732]" captionTargetBox="[161,1426,582,1672]" captionTargetId="figure@20.[151,1436,573,1685]" captionTargetPageId="20" captionText="FIGURES 41 48. 41 44. Hypandrium (ventral view) and phallus with postgonites (lateral view). 41 42. Bicellaria uvens Melander. 43 44. Bicellaria vana Collin. 45 46. Distal part of front tibia and proximal part of front basitarsus (posterior view). 45. Bicellaria spuria (Fallén). 46. Bicellaria vana Collin. 47 48. Antennae. 47. Bicellaria dispar Oldenberg. 48. Bicellaria alpina Bezzi (microchaetae mostly omitted). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224165/files/figure.png" pageId="23" pageNumber="274">Fig. 47</figureCitation>
); acrostichals and dorsocentrals (except last two pairs of dorsocentrals) very short (less than
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long); all tibiae without any setae obviously longer than setulae; male hypandrial processes long setose along the whole inner margin; very small species (wing length
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)....................................................
<taxonomicName id="4C6DE4FDFFBA194E2BE73CBE95E5D345" ID-CoL="LRNM" authority="Oldenberg" authorityName="Oldenberg" box="[1231,1435,239,260]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="dispar">
<emphasis id="B919436CFFBA194E2BE73CBE95E5D345" bold="true" box="[1231,1435,239,260]" pageId="23" pageNumber="274">
<emphasis id="B919436CFFBA194E2BE73CBE9556D345" bold="true" box="[1231,1320,239,260]" italics="true" pageId="23" pageNumber="274">B. dispar</emphasis>
Oldenberg
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD29F7EFFBA194E2FBF3D5C95E2D37E" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3D5C95E2D37E" pageId="23" pageNumber="274">
- Antennal stylus together with slender portion of third segment at least as long as broad basal portion of third segment (as in
<figureCitation id="135683FBFFBA194E2E2D3D7B9132D37E" box="[261,332,298,319]" captionStart="FIGURES 41 48. 41 44" captionStartId="20.[151,264,1709,1732]" captionTargetBox="[161,1426,582,1672]" captionTargetId="figure@20.[151,1436,573,1685]" captionTargetPageId="20" captionText="FIGURES 41 48. 41 44. Hypandrium (ventral view) and phallus with postgonites (lateral view). 41 42. Bicellaria uvens Melander. 43 44. Bicellaria vana Collin. 45 46. Distal part of front tibia and proximal part of front basitarsus (posterior view). 45. Bicellaria spuria (Fallén). 46. Bicellaria vana Collin. 47 48. Antennae. 47. Bicellaria dispar Oldenberg. 48. Bicellaria alpina Bezzi (microchaetae mostly omitted). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224165/files/figure.png" pageId="23" pageNumber="274">Fig. 48</figureCitation>
); acrostichals and dorsocentrals longer than
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; remaining characters disagree....................... 2
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</paragraph>
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<keyStep id="309987DBFFBA194E2FBF3D1695E2D3D6" pageId="23" pageNumber="274">
<paragraph id="8BD29F7EFFBA194E2FBF3D1695E2D31D" blockId="23.[151,1437,151,2020]" box="[151,1436,327,348]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3D1695E2D31D" box="[151,1436,327,348]" pageId="23" pageNumber="274">2 (1) Palpus with 4 or more black setae; third antennal segment with dorsal seta(e); all body setae black.................. 3</keyLead>
</paragraph>
<paragraph id="8BD29F7EFFBA194E2FBF3D3495E2D3D6" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3D3495E2D3D6" pageId="23" pageNumber="274">
- Palpus with 13 black setae (rarely with an additional pale seta), if palpus with more than 4 setae (
<taxonomicName id="4C6DE4FDFFBA194E2BE63D349546D33B" box="[1230,1336,357,378]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="andorra" status="sp. nov.">
<emphasis id="B919436CFFBA194E2BE63D349546D33B" box="[1230,1336,357,378]" italics="true" pageId="23" pageNumber="274">B. andorra</emphasis>
</taxonomicName>
<emphasis id="B919436CFFBA194E2A683D3495F0D33B" bold="true" box="[1344,1422,357,378]" pageId="23" pageNumber="274">
<taxonomicNameLabel id="A22AFE17FFBA194E2A683D3495F0D33B" box="[1344,1422,357,378]" pageId="23" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), then sides of abdomen with yellowish-white setae; third antennal segment with or without dorsal seta(e)............. 6
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</paragraph>
</keyStep>
<keyStep id="309987DBFFBA194E2FBF3DCE95E2D006" pageId="23" pageNumber="274">
<paragraph id="8BD29F7EFFBA194E2FBF3DCE95E2D04D" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3DCE95E2D04D" pageId="23" pageNumber="274">
3 (2) Mesoscutum with two pale (silvery) stripes between rows of acrostichals and dorsocentrals apparent from both dorsal and caudal views (in female covering most of mesoscutum); usually less than 9 setae outside rows of dorsocentrals at presutural area; abdomen light bluish grey in lateral view; male genitalia as in
<figureCitation id="135683FBFFBA194E2CE93D8B944DD3AE" box="[961,1075,474,495]" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1650,1673]" captionTargetBox="[158,1430,608,1619]" captionTargetId="figure@3.[151,1436,595,1626]" captionTargetPageId="3" captionText="FIGURES 1 8. Hypandrium (ventral view) and phallus with postgonites (lateral view). 1 2. Bicellaria albopilosa Chvála. 3 4. Bicellaria alpina Bezzi. 5 6. Bicellaria andorra sp. nov. 7 8. Bicellaria austriaca Tuomikoski. Abbreviations: eapejaculatory apodeme, hph—phallic hooks, mep—median extensions of postgonites, pgn—postgonites, vap—ventral apodeme. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224160/files/figure.png" pageId="23" pageNumber="274">Figures 34</figureCitation>
(
<taxonomicName id="4C6DE4FDFFBA194E2B6A3D8B94E9D3AE" ID-CoL="5WJQ8" box="[1090,1175,474,495]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B919436CFFBA194E2B6A3D8B94E9D3AE" box="[1090,1175,474,495]" italics="true" pageId="23" pageNumber="274">B. pilosa</emphasis>
</taxonomicName>
complex-part)..............................................................................................
<taxonomicName id="4C6DE4FDFFBA194E2A2C3DA695E2D04D" ID-CoL="5WJPP" authority="Bezzi" authorityName="Bezzi" box="[1284,1436,503,524]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="alpina">
<emphasis id="B919436CFFBA194E2A2C3DA695E2D04D" bold="true" box="[1284,1436,503,524]" pageId="23" pageNumber="274">
<emphasis id="B919436CFFBA194E2A2C3DA69521D04D" bold="true" box="[1284,1375,503,524]" italics="true" pageId="23" pageNumber="274">B. alpina</emphasis>
Bezzi
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD29F7EFFBA194E2FBF3E4495E2D006" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3E4495E2D006" pageId="23" pageNumber="274">- Mesoscutum uniformly dark velvety black without any stripes; usually 10 or more setae outside rows of dorsocentrals at presutural area; abdomen dark greyish to blackish brown in lateral view....................................... 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="309987DBFFBA194E2FBF3E1E95E2D10E" pageId="23" pageNumber="274">
<paragraph id="8BD29F7EFFBA194E2FBF3E1E95E2D09B" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3E1E95E2D09B" pageId="23" pageNumber="274">
4 (3) Third antennal segment usually with 4 or more dorsal setae, second segment with curiously long lateral setae pointing forwards; fore tibia with posteroventral setae shorter than diameter of tibia and with a row of 46 erect posterodorsal setae about
<date id="FFD3B9BEFFBA194E2E683EDB9120D0DE" box="[320,350,650,671]" pageId="23" pageNumber="274">4X</date>
longer than diameter of tibia (
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long); hind tibia broadly thickened on apical third in male and slender in female; male hypandrium with short and broad processes bearing long apical seta, two phallic hooks present, phallus strongly sclerotized, forming flat and long valve-like structure (
<figureCitation id="135683FBFFBA194E2C4E3E9493ADD09B" box="[870,979,709,730]" captionStart="FIGURES 17 24" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[165,1424,888,1908]" captionTargetId="figure@10.[151,1436,878,1916]" captionTargetPageId="10" captionText="FIGURES 17 24. Hypandrium (ventral view) and phallus with postgonites (lateral view). 17 18. Bicellaria kocoureki sp. nov. 19 20. Bicellaria longisetosa Chvála. 21 22. Bicellaria mera Collin. 23 24. Bicellaria nigra (Meigen). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224162/files/figure.png" pageId="23" pageNumber="274">Figs 1920</figureCitation>
)...................
<taxonomicName id="4C6DE4FDFFBA194E2BEA3E9495E2D09B" ID-CoL="LRP3" authority="Chvala" authorityName="Chvala" box="[1218,1436,709,730]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="longisetosa">
<emphasis id="B919436CFFBA194E2BEA3E9495E2D09B" bold="true" box="[1218,1436,709,730]" pageId="23" pageNumber="274">
<emphasis id="B919436CFFBA194E2BEA3E949533D09B" bold="true" box="[1218,1357,709,730]" italics="true" pageId="23" pageNumber="274">B. longisetosa</emphasis>
Chvála
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD29F7EFFBA194E2FBF3EB395E2D10E" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
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- Third antennal segment usually with 13 dorsal setae, second segment with short lateral setae (but with long ventral setae); fore tibia differently setose; hind tibia more evenly dilated in both sexes; male hypandrial processes without long apical seta, three phallic hooks (left hook bifurcate), phallus membranous, not forming valve-like structure. (
<taxonomicName id="4C6DE4FDFFBA194E2BCC3F4C9545D173" ID-CoL="5WJQ8" box="[1252,1339,797,818]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis id="B919436CFFBA194E2BCC3F4C9545D173" box="[1252,1339,797,818]" italics="true" pageId="23" pageNumber="274">B. pilosa</emphasis>
</taxonomicName>
complexpart)............................................................................................ 5
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5 (4) Face narrow (about 56 facets combined), almost parallel throughout (
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); about 10 setae on posthumeral and intahumeral areas; male hypandrial processes broadened near apex, left phallic hook bifurcate, its anterior arm bent, postgonites strongly asymmetrical, right one with ventral extension (
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)..................
<emphasis id="B919436CFFBA194E2BE33FC395E2D1E6" bold="true" box="[1227,1436,914,935]" pageId="23" pageNumber="274">
<taxonomicName id="4C6DE4FDFFBA194E2BE33FC39534D1E6" box="[1227,1354,914,935]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="setipalpus" status="sp. nov.">
<emphasis id="B919436CFFBA194E2BE33FC39534D1E6" bold="true" box="[1227,1354,914,935]" italics="true" pageId="23" pageNumber="274">B. setipalpus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22AFE17FFBA194E2A673FC395E2D1E6" box="[1359,1436,914,935]" pageId="23" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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- Face broad (about 89 facets combined), in lower half even broader (
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) than in upper half (
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); usually more than 10 setae on posthumeral and intrahumeral areas; male hypandrial processes apically narrow; left phallic hook bifurcate, its anterior arm straight (
<figureCitation id="135683FBFFBA194E2D0C3FBB92ECD1BE" box="[548,658,1002,1023]" captionStart="FIGURES 25 32" captionStartId="13.[151,264,1898,1921]" captionTargetBox="[164,1430,747,1867]" captionTargetId="figure@13.[151,1436,735,1874]" captionTargetPageId="13" captionText="FIGURES 25 32. Hypandrium (ventral view) and phallus with postgonites (lateral view). 25 26. Bicellaria nigrita Collin. 27 28. Bicellaria pilosa Lundbeck. 29 30. Bicellaria setipalpus sp. nov. 31 32. Bicellaria simplicipes (Zetterstedt). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224163/files/figure.png" pageId="23" pageNumber="274">Figs 2728</figureCitation>
)..............................................
<taxonomicName id="4C6DE4FDFFBA194E2BFF3FBB95E2D1BE" ID-CoL="5WJQ8" authority="Lundbeck" authorityName="Lundbeck" box="[1239,1436,1002,1023]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="pilosa">
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<emphasis id="B919436CFFBA194E2BFF3FBB9550D1BE" bold="true" box="[1239,1326,1002,1023]" italics="true" pageId="23" pageNumber="274">B. pilosa</emphasis>
Lundbeck
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6 (2)
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basitarsus (and usually also the hind tarsomeres 2) distinctly swollen (usually more than 2.3X broader than hind tarsomere 3 and usually broader than
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); additional characters: fore tibia with ventral pilosity on apical third at most slightly shorter than on fore basitarsus and posteroventral setae short, at most as long as pilosity and usually shorter than depth of tibia (as in
<figureCitation id="135683FBFFBA194E2EE1380E926ED635" box="[457,528,1119,1140]" captionStart="FIGURES 41 48. 41 44" captionStartId="20.[151,264,1709,1732]" captionTargetBox="[161,1426,582,1672]" captionTargetId="figure@20.[151,1436,573,1685]" captionTargetPageId="20" captionText="FIGURES 41 48. 41 44. Hypandrium (ventral view) and phallus with postgonites (lateral view). 41 42. Bicellaria uvens Melander. 43 44. Bicellaria vana Collin. 45 46. Distal part of front tibia and proximal part of front basitarsus (posterior view). 45. Bicellaria spuria (Fallén). 46. Bicellaria vana Collin. 47 48. Antennae. 47. Bicellaria dispar Oldenberg. 48. Bicellaria alpina Bezzi (microchaetae mostly omitted). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224165/files/figure.png" pageId="23" pageNumber="274">Fig. 46</figureCitation>
), face very narrow (less than 3 facets combined, less than
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at narrowest point), abdomen at least partly pale setose (exception: male
<taxonomicName id="4C6DE4FDFFBA194E2DFD382C934CD6D3" ID-CoL="5WJPW" box="[725,818,1149,1170]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B919436CFFBA194E2DFD382C934CD6D3" box="[725,818,1149,1170]" italics="true" pageId="23" pageNumber="274">B. nigrita</emphasis>
</taxonomicName>
), third antennal segment without dorsal seta(e), hypandrial processes never strikingly shortened (
<taxonomicName id="4C6DE4FDFFBA194E2D6738CB92E0D6EE" ID-CoL="LRP8" box="[591,670,1178,1199]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B919436CFFBA194E2D6738CB92E0D6EE" box="[591,670,1178,1199]" italics="true" pageId="23" pageNumber="274">B. nigra</emphasis>
</taxonomicName>
complex)...................................................... 7
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-
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basitarsus slender (usually less than 2.0X broader than hind tarsomere 3 and usually narrower than
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); additional characters in different combinations.............................................................. 9
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<paragraph id="8BD29F7EFFBA194E2FBF38A395E2D71E" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF38A395E2D71E" pageId="23" pageNumber="274">
7 (6) Third antennal segment arched dorsally; mid tibia usually without dorsal setae differentiated from short hairs; hind femur shorter than
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; hind tibia dilated in basal third; male: postgonites shortened (about
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long), both phallic hooks present, long and strong, longer than postgonites, right one slightly shorter and narrower than left one (
<figureCitation id="135683FBFFBA194E2BC5397C954BD703" box="[1261,1333,1325,1346]" captionStart="FIGURES 25 32" captionStartId="13.[151,264,1898,1921]" captionTargetBox="[164,1430,747,1867]" captionTargetId="figure@13.[151,1436,735,1874]" captionTargetPageId="13" captionText="FIGURES 25 32. Hypandrium (ventral view) and phallus with postgonites (lateral view). 25 26. Bicellaria nigrita Collin. 27 28. Bicellaria pilosa Lundbeck. 29 30. Bicellaria setipalpus sp. nov. 31 32. Bicellaria simplicipes (Zetterstedt). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224163/files/figure.png" pageId="23" pageNumber="274">Fig. 25</figureCitation>
), squamae mostly with dark fringes................................................................
<taxonomicName id="4C6DE4FDFFBA194E2BD1391B95E2D71E" ID-CoL="5WJPW" authority="Collin" authorityName="Collin" box="[1273,1436,1354,1375]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="nigrita">
<emphasis id="B919436CFFBA194E2BD1391B95E2D71E" bold="true" box="[1273,1436,1354,1375]" pageId="23" pageNumber="274">
<emphasis id="B919436CFFBA194E2BD1391B9529D71E" bold="true" box="[1273,1367,1354,1375]" italics="true" pageId="23" pageNumber="274">B. nigrita</emphasis>
Collin
</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD29F7EFFBA194E2FBF393695E2D7F6" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF393695E2D7F6" pageId="23" pageNumber="274">
-. Third antennal segment nearly straight dorsally; mid tibia with 13 outstanding dorsal setae, about twice as long as short hairs on limb; hind femur longer than
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; hind tibia equally slender in basal third; male: postgonites not reduced (slightly shortened in
<taxonomicName id="4C6DE4FDFFBA194E2EFE39F39237D7F6" ID-CoL="LRNV" box="[470,585,1442,1463]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="halterata">
<emphasis id="B919436CFFBA194E2EFE39F39237D7F6" box="[470,585,1442,1463]" italics="true" pageId="23" pageNumber="274">B. halterata</emphasis>
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), phallic hooks different, squamae mostly with pale fringes...................... 8
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<keyStep id="309987DBFFBA194E2FBF39EE95E2D4E3" pageId="23" pageNumber="274">
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8 (7) All large setae on mesoscutum black; male: halter usually dark, postgonites long (more than twice as long as broad), left phallic hook long, without lateral process, right one strongly reduced (
<figureCitation id="135683FBFFBA194E2C8B398C9395D7B3" box="[931,1003,1501,1522]" captionStart="FIGURES 17 24" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[165,1424,888,1908]" captionTargetId="figure@10.[151,1436,878,1916]" captionTargetPageId="10" captionText="FIGURES 17 24. Hypandrium (ventral view) and phallus with postgonites (lateral view). 17 18. Bicellaria kocoureki sp. nov. 19 20. Bicellaria longisetosa Chvála. 21 22. Bicellaria mera Collin. 23 24. Bicellaria nigra (Meigen). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224162/files/figure.png" pageId="23" pageNumber="274">Fig. 23</figureCitation>
); female: broad portion of third antennal segment usually longer than
<quantity id="4C95329BFFBA194E2ED039AB922ED44E" box="[504,592,1530,1551]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="23" pageNumber="274" unit="mm" value="0.15">0.15 mm</quantity>
, lustrous portions of abdominal tergites 67 comprise about basal third of these tergites...................................................................................
<taxonomicName id="4C6DE4FDFFBA194E2BC13A4695E2D46D" ID-CoL="LRP8" authority="Meigen" authorityName="Meigen" box="[1257,1436,1559,1580]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B919436CFFBA194E2BC13A4695E2D46D" bold="true" box="[1257,1436,1559,1580]" pageId="23" pageNumber="274">
<emphasis id="B919436CFFBA194E2BC13A469544D46D" bold="true" box="[1257,1338,1559,1580]" italics="true" pageId="23" pageNumber="274">B. nigra</emphasis>
(Meigen)
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</taxonomicName>
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<paragraph id="8BD29F7EFFBA194E2FBF3A6495E2D4E3" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
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- Large setae on mesoscutum mostly white to brownish-yellow (at least scutellars, last pair of dorsocentrals and notopleurals); male: halter yellow, postgonites shorter (about 1.5X as long as broad), both phallic hooks long, left one with lateral process (best visible from below,
<figureCitation id="135683FBFFBA194E2D633A3E92EED4C5" box="[587,656,1647,1668]" captionStart="FIGURES 9 16" captionStartId="7.[151,264,1544,1567]" captionTargetBox="[164,1423,529,1514]" captionTargetId="figure@7.[151,1436,519,1520]" captionTargetPageId="7" captionText="FIGURES 9 16. Hypandrium (ventral view) and phallus with postgonites (lateral view). 9 10. Bicellaria collini Tuomikoski. 11 12. Bicellaria halterata Collin. 13 14. Bicellaria intermedia Lundbeck. 15 16. Bicellaria italica sp. nov. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224161/files/figure.png" pageId="23" pageNumber="274">Fig. 11</figureCitation>
); female: broad portion of third antennal segment usually shorter than
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, lustrous portions of abdominal tergites 67 comprise more than basal half of these tergites.........
<taxonomicName id="4C6DE4FDFFBA194E2BCA3ADC95E2D4E3" ID-CoL="LRNV" authority="Collin" authorityName="Collin" box="[1250,1436,1677,1698]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="halterata">
<emphasis id="B919436CFFBA194E2BCA3ADC95E2D4E3" bold="true" box="[1250,1436,1677,1698]" pageId="23" pageNumber="274">
<emphasis id="B919436CFFBA194E2BCA3ADC9529D4E3" bold="true" box="[1250,1367,1677,1698]" italics="true" pageId="23" pageNumber="274">B. halterata</emphasis>
Collin
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9 (6) Third antennal segment with one to several long dorsal setae; male with two phallic hooks (both exceeding postgonites) (
<taxonomicName id="4C6DE4FDFFBA194E2AAF3AFB9121D49C" ID-CoL="LRNG" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="austriaca">
<emphasis id="B919436CFFBA194E2AAF3AFB9121D49C" italics="true" pageId="23" pageNumber="274">B. austriaca</emphasis>
</taxonomicName>
complex)................................................................................ 10
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</paragraph>
<paragraph id="8BD29F7EFFBA194E2FBF3AB495E2D4BB" blockId="23.[151,1437,151,2020]" box="[151,1436,1765,1786]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3AB495E2D4BB" box="[151,1436,1765,1786]" pageId="23" pageNumber="274">- Third antennal segment without long dorsal setae; male with 12 phallic hooks................................ 16</keyLead>
</paragraph>
</keyStep>
<keyStep id="309987DBFFBA194E2FBF3B5395E5D575" pageId="23" pageNumber="274">
<paragraph id="8BD29F7EFFBA194E2FBF3B5395E5D556" blockId="23.[151,1437,151,2020]" box="[151,1435,1794,1815]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3B5395E5D556" box="[151,1435,1794,1815]" pageId="23" pageNumber="274">10 (9) Male........................................................................................... 11</keyLead>
</paragraph>
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<keyLead id="309C3C4BFFBA194E2FBF3B4E95E5D575" box="[151,1435,1823,1844]" pageId="23" pageNumber="274">- Female......................................................................................... 14</keyLead>
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<keyStep id="309987DBFFBA194E2FBF3B6C95E2D586" pageId="23" pageNumber="274">
<paragraph id="8BD29F7EFFBA194E2FBF3B6C95E5D5CD" blockId="23.[151,1437,151,2020]" pageId="23" pageNumber="274">
<keyLead id="309C3C4BFFBA194E2FBF3B6C95E5D5CD" pageId="23" pageNumber="274">
11 (10) Prescutellar depression lighter than other parts of mesoscutum in dorsal view (lighter area reaches hind acrostichals); halter yellow; hypandrial processes swollen apically, bearing several rather long setae on apex, both phallic hooks almost equally long, right one more sharply bent (
<figureCitation id="135683FBFFBA194E2D793B2692C0D5CD" box="[593,702,1911,1932]" captionStart="FIGURES 9 16" captionStartId="7.[151,264,1544,1567]" captionTargetBox="[164,1423,529,1514]" captionTargetId="figure@7.[151,1436,519,1520]" captionTargetPageId="7" captionText="FIGURES 9 16. Hypandrium (ventral view) and phallus with postgonites (lateral view). 9 10. Bicellaria collini Tuomikoski. 11 12. Bicellaria halterata Collin. 13 14. Bicellaria intermedia Lundbeck. 15 16. Bicellaria italica sp. nov. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224161/files/figure.png" pageId="23" pageNumber="274">Figs 1516</figureCitation>
)..............................................
<emphasis id="B919436CFFBA194E2BC63B2695E5D5CD" bold="true" box="[1262,1435,1911,1932]" pageId="23" pageNumber="274">
<taxonomicName id="4C6DE4FDFFBA194E2BC63B269537D5CD" box="[1262,1353,1911,1932]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="274" phylum="Arthropoda" rank="species" species="italica" status="sp. nov.">
<emphasis id="B919436CFFBA194E2BC63B269537D5CD" bold="true" box="[1262,1353,1911,1932]" italics="true" pageId="23" pageNumber="274">B. italica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22AFE17FFBA194E2A673B2695E5D5CD" box="[1359,1435,1911,1932]" pageId="23" pageNumber="274" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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12 (11) Face parallel-sided or slightly broadened ventrally (about 5 facets combined,
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broad at narrowest point); hypan-
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drial processes short (
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) and broadly U-shaped (distance between their inner margins of processes about
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), both phallic hooks equally long and equally bent (
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)......................
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- Face narrowing ventrally (about 3 facets combined,
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broad at narrowest point); hypandrial processes longer and narrowly U-shaped (distance between inner margins of processes about
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), phallic hooks different...... 13
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13 (12) Usually more than 9 setae in posthumeral and intrahumeral areas; hypandrial processes broader, with several rather long setae near apex, phallic hooks unequal, shorter and broader (
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)....................
<taxonomicName id="4C6DE4FDFFB519412B8D3D7B95E2D37E" ID-CoL="LRNG" authority="Tuomikoski" authorityName="Tuomikoski" box="[1189,1436,298,319]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="austriaca">
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- Usually less than 7 setae in posthumeral and intrahumeral areas; hypandrial processes narrower, with at most a single longer seta on apex, other setae much shorter, long and narrow phallic hooks nearly equal in length (
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).......................................................................................
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14 (10) Face parallel-sided or slightly broadening ventrally (about 5 facets combined,
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broad at narrowest point); abdominal tergites 67 broadly lustrous on basal half or more....................................
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- Face narrowed ventrally (about 3 facets combined,
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broad at narrowest point); abdominal tergites 67 nearly completely microtrichose.............................................................................. 15
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15 (14) Halter brown; usually more than 9 setae in posthumeral and intrahumeral areas.............
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- Halter yellow; usually less than 9 setae in posthumeral and intrahumeral areas.................................................................................................
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Lundbeck,
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italica
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16 (9) Fore tibia with ventral pilosity on apical third at most slightly shorter than pilosity on fore basitarsus and/or posteroventral setae on apical third of tibia shorter than depth of tibia (as in
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, observe in exactly posterior view)............ 17
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- Fore tibia with ventral pilosity on apical third at most one-third as long as on fore basitarsus and posteroventral setae on apical third of tibia at least twice as long as pilosity, and usually longer than depth of tibia (as in
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)............ 20
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17 (16)
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tibia slender throughout; lower occipital setae mostly pale; male: hypandrial processes short and broad, right phallic hook long, simply curved and exceeding postgonites, left one slightly shorter, triangular in shape (
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)....................................................................................
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-
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tibia distinctly swollen apically (less so in female); lower occipital setae black; male: hypandrial processes not conspicuously short and broad, right phallic hook strongly reduced or absent, left one long and simply curved (
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complex)........................................................................................... 18
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18 (17) Abdomen with long white setae on sides; the anteromost acrostichals and dorsocentrals subequally long as those just before suture (first three about
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long or even longer in males); hind femur without posteroventral setae................................................................................................
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- Abdomen entirely black setose; the anteromost acrostichals and dorsocentrals distinctly shorter than those just before suture (first three about
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long); hind femur with posteroventral setae (if abdomen partly pale setose, compare
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)....................................................................................... 19
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19 (18) Face very narrow (less than 2 facets combined,
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), narrowed ventrally; male presutural area evenly dark in dorsal view (male) or with pale spot sharply triangular-shaped (female); hind tibia more equally swollen towards tip, at its broadest point subequally broad as hind femur even in male (both
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); male hypandrial processes slender; apical corner of postgonites broadly rounded (
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)..............................................
<taxonomicName id="4C6DE4FDFFB519412A2338CB95E2D6EE" ID-CoL="LRPM" authority="Collin" authorityName="Collin" box="[1291,1436,1178,1199]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="vana">
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<emphasis id="B919436CFFB519412A2338CB9529D6EE" bold="true" box="[1291,1367,1178,1199]" italics="true" pageId="24" pageNumber="275">B. vana</emphasis>
Collin
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</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="309C3C4BFFB519412FBF38E695E5D703" pageId="24" pageNumber="275">
- Face broader (about 3 facets combined,
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), parallel-sided; presutural area lighter in dorsal view than remaining parts of mesoscutum in both sexes, of rectangular to semicircular appearance; hind tibia strongly swollen towards tip (usually more than
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at broadest point in male and some
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in female), at its broadest point broader than hind femur (
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); male hypandrial processes broader; apical corner of postgonites narrower and more pronounced (
<figureCitation id="135683FBFFB519412A59395E913CD703" captionStart="FIGURES 33 40" captionStartId="17.[151,264,1945,1968]" captionTargetBox="[163,1428,876,1910]" captionTargetId="figure@17.[151,1436,865,1921]" captionTargetPageId="17" captionText="FIGURES 33 40. Hypandrium (ventral view) and phallus with postgonites (lateral view). 33 34. Bicellaria spuria (Fallén). 35 36. Bicellaria stackelbergi Tuomikoski. 37 38. Bicellaria subpilosa Collin. 39 40. Bicellaria sulcata (Zetterstedt). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224164/files/figure.png" pageId="24" pageNumber="275">Figs 3940</figureCitation>
)........................................................................
<taxonomicName id="4C6DE4FDFFB519412B9A397C95E5D703" ID-CoL="LRPK" authority="Zetterstedt" authorityName="Zetterstedt" box="[1202,1435,1325,1346]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="sulcata">
<emphasis id="B919436CFFB519412B9A397C95E5D703" bold="true" box="[1202,1435,1325,1346]" pageId="24" pageNumber="275">
<emphasis id="B919436CFFB519412B9A397C956AD703" bold="true" box="[1202,1300,1325,1346]" italics="true" pageId="24" pageNumber="275">B. sulcata</emphasis>
(Zetterstedt)
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</keyStep>
<keyStep id="309987DBFFB519412FBF391B95E5D73D" pageId="24" pageNumber="275">
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<keyLead id="309C3C4BFFB519412FBF391B95E5D71E" box="[151,1435,1354,1375]" pageId="24" pageNumber="275">20 (16) Abdomen partly pale setose (at least on sides of tergite 2)................................................. 21</keyLead>
</paragraph>
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<keyLead id="309C3C4BFFB519412FBF393695E5D73D" box="[151,1435,1383,1404]" pageId="24" pageNumber="275">- Abdomen entirely black setose....................................................................... 22</keyLead>
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</keyStep>
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<keyLead id="309C3C4BFFB519412FBF39D495E2D7B3" pageId="24" pageNumber="275">
21 (20) Legs microtrichose, with longer and slender tarsi (combined length of fore+mid+hind basitarsus usually more than
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); usually less than 7 setae in posthumeral and intrahumeral areas; male hind tibia gradually slightly swollen, lateral parts of abdomen brown; hypandrial processes shorter (
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), both phallic hooks narrow, connected ventrally (
<figureCitation id="135683FBFFB519412A7B39EE916DD7B3" captionStart="FIGURES 1 8" captionStartId="3.[151,264,1650,1673]" captionTargetBox="[158,1430,608,1619]" captionTargetId="figure@3.[151,1436,595,1626]" captionTargetPageId="3" captionText="FIGURES 1 8. Hypandrium (ventral view) and phallus with postgonites (lateral view). 1 2. Bicellaria albopilosa Chvála. 3 4. Bicellaria alpina Bezzi. 5 6. Bicellaria andorra sp. nov. 7 8. Bicellaria austriaca Tuomikoski. Abbreviations: eapejaculatory apodeme, hph—phallic hooks, mep—median extensions of postgonites, pgn—postgonites, vap—ventral apodeme. Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224160/files/figure.png" pageId="24" pageNumber="275">Figs 1 2</figureCitation>
); female tergites 68 at least partly microtrichose......................................
<taxonomicName id="4C6DE4FDFFB519412BE3398C95E2D7B3" ID-CoL="LRND" authority="Chvala" authorityName="Chvala" box="[1227,1436,1501,1522]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="albopilosa">
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<emphasis id="B919436CFFB519412BE3398C9532D7B3" bold="true" box="[1227,1356,1501,1522]" italics="true" pageId="24" pageNumber="275">B. albopilosa</emphasis>
Chvála
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</taxonomicName>
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</paragraph>
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<keyLead id="309C3C4BFFB519412FBF39AB95E2D4C5" pageId="24" pageNumber="275">
- Legs lustrous, with shorter tarsi (combined length of fore+mid+hind basitarsus usually less than
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); usually more than 8 setae in posthumeral and intrahumeral areas; male hind tibia strongly and abruptly swollen, lateral parts of abdomen pale and almost silvery grey; hypandrial processes long (
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), both phallic hooks present, left one hoe-like broadened laterally (best visible from behind) (
<figureCitation id="135683FBFFB519412D763A0392B0D426" box="[606,718,1618,1639]" captionStart="FIGURES 33 40" captionStartId="17.[151,264,1945,1968]" captionTargetBox="[163,1428,876,1910]" captionTargetId="figure@17.[151,1436,865,1921]" captionTargetPageId="17" captionText="FIGURES 33 40. Hypandrium (ventral view) and phallus with postgonites (lateral view). 33 34. Bicellaria spuria (Fallén). 35 36. Bicellaria stackelbergi Tuomikoski. 37 38. Bicellaria subpilosa Collin. 39 40. Bicellaria sulcata (Zetterstedt). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224164/files/figure.png" pageId="24" pageNumber="275">Figs 3536</figureCitation>
); female tergites 68 almost entirely lustrous..................................................................................................
<taxonomicName id="4C6DE4FDFFB519412BA33A3E95E2D4C5" ID-CoL="LRPH" authority="Tuomikoski" authorityName="Tuomikoski" box="[1163,1436,1647,1668]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="stackelbergi">
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<emphasis id="B919436CFFB519412BA33A3E9563D4C5" bold="true" box="[1163,1309,1647,1668]" italics="true" pageId="24" pageNumber="275">B. stackelbergi</emphasis>
Tuomikoski
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</keyStep>
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<keyLead id="309C3C4BFFB519412FBF3ADC95E5D4FE" pageId="24" pageNumber="275">
22 (20) Halter yellow; male hypandrium large with contrastingly short, narrow, broadly U-shaped and long setose processes (
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); female tergites 58 almost entirely lustrous.........................................
<emphasis id="B919436CFFB519412BE33AFB95E5D4FE" bold="true" box="[1227,1435,1706,1727]" pageId="24" pageNumber="275">
<taxonomicName id="4C6DE4FDFFB519412BE33AFB9537D4FE" box="[1227,1353,1706,1727]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="kocoureki" status="sp. nov.">
<emphasis id="B919436CFFB519412BE33AFB9537D4FE" bold="true" box="[1227,1353,1706,1727]" italics="true" pageId="24" pageNumber="275">B. kocoureki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22AFE17FFB519412A673AFB95E5D4FE" box="[1359,1435,1706,1727]" pageId="24" pageNumber="275" rank="species">sp. nov.</taxonomicNameLabel>
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</keyLead>
</paragraph>
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<keyLead id="309C3C4BFFB519412FBF3A9695E2D49D" box="[151,1436,1735,1756]" pageId="24" pageNumber="275">- Halter brownish yellow to black; male hypandrium different; female tergites 58 at least partly microtrichose........ 23</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BD29F7EFFB519412FBF3AB495E5D4BB" blockId="24.[151,1437,151,2020]" box="[151,1435,1765,1786]" pageId="24" pageNumber="275">
<keyLead id="309C3C4BFFB519412FBF3AB495E5D4BB" box="[151,1435,1765,1786]" pageId="24" pageNumber="275">23 (22) Male........................................................................................... 24</keyLead>
</paragraph>
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<keyLead id="309C3C4BFFB519412FBF3B5395E5D556" box="[151,1435,1794,1815]" pageId="24" pageNumber="275">- Female......................................................................................... 27</keyLead>
</paragraph>
</keyStep>
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<keyLead id="309C3C4BFFB519412FBF3B4E95E2D513" pageId="24" pageNumber="275">
24 (23) Postgonites slightly broadened laterally and narrow in lateral view, hypandrium with short (
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) and broad processes (3
<date id="FFD3B9BEFFB519412E4D3B6C910FD513" box="[357,369,1853,1874]" pageId="24" pageNumber="275">4</date>
X as long as broad), two equally shaped and long phallic hooks (
<figureCitation id="135683FBFFB519412CF63B6C9432D513" box="[990,1100,1853,1874]" captionStart="FIGURES 41 48. 41 44" captionStartId="20.[151,264,1709,1732]" captionTargetBox="[161,1426,582,1672]" captionTargetId="figure@20.[151,1436,573,1685]" captionTargetPageId="20" captionText="FIGURES 41 48. 41 44. Hypandrium (ventral view) and phallus with postgonites (lateral view). 41 42. Bicellaria uvens Melander. 43 44. Bicellaria vana Collin. 45 46. Distal part of front tibia and proximal part of front basitarsus (posterior view). 45. Bicellaria spuria (Fallén). 46. Bicellaria vana Collin. 47 48. Antennae. 47. Bicellaria dispar Oldenberg. 48. Bicellaria alpina Bezzi (microchaetae mostly omitted). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224165/files/figure.png" pageId="24" pageNumber="275">Figs 4142</figureCitation>
)...........
<taxonomicName id="4C6DE4FDFFB519412BF43B6C95E2D513" ID-CoL="LRPL" authority="Melander" authorityName="Melander" box="[1244,1436,1853,1874]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="24" pageNumber="275" phylum="Arthropoda" rank="species" species="uvens">
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<emphasis id="B919436CFFB519412BF43B6C954CD513" bold="true" box="[1244,1330,1853,1874]" italics="true" pageId="24" pageNumber="275">B. uvens</emphasis>
Melander
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</taxonomicName>
</keyLead>
</paragraph>
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<keyLead id="309C3C4BFFB519412FBF3B0B95E2D5CD" pageId="24" pageNumber="275">- Postgonites not broadened laterally and trapezoid in lateral view, hypandrium with narrower and longer processes, a single long phallic hook present (the other less than one-quarter as long)........................................... 25</keyLead>
</paragraph>
</keyStep>
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25 (24) Face broad in ventral portion (usually more than 5 facets combined,
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); mid femur with complete row of anteroventrals at least half as long as depth of femur at mid-length; hind tibia with ventral setae about twice as long as depth of tibia; prescutellar depression paler than remaining parts of mesoscutum in dorsal view; phallic hook bent in S-shaped (
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).
</keyStep>
</keyLead>
</paragraph>
<paragraph id="8BD29F7EFFB419402E2F3CC695E2D37E" blockId="25.[151,1436,151,700]" pageId="25" pageNumber="276">
..................................................................................
<taxonomicName id="4C6DE4FDFFB419402BF63CC695E2D2ED" ID-CoL="LRPJ" authority="Collin" authorityName="Collin" box="[1246,1436,151,172]" class="Insecta" family="Hybotidae" genus="Bicellaria" kingdom="Animalia" order="Diptera" pageId="25" pageNumber="276" phylum="Arthropoda" rank="species" species="subpilosa">
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<emphasis id="B919436CFFB419402BF63CC69529D2ED" bold="true" box="[1246,1367,151,172]" italics="true" pageId="25" pageNumber="276">B. subpilosa</emphasis>
Collin
</emphasis>
</taxonomicName>
- Face narrow in ventral portion (less than 4 facets combined,
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); mid femur with anteroventrals shorter than half of femur depths at mid-length; hind tibia with ventral setae at most slightly longer than tibia depth of tibia; prescutellar depression paler or evenly dark as remaining parts of mesoscutum in dorsal view; phallic hook simply bowed (
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,
<figureCitation id="135683FBFFB419402A5E3CBE95F1D345" box="[1398,1423,239,260]" captionStart="FIGURES 17 24" captionStartId="10.[151,264,1940,1963]" captionTargetBox="[165,1424,888,1908]" captionTargetId="figure@10.[151,1436,878,1916]" captionTargetPageId="10" captionText="FIGURES 17 24. Hypandrium (ventral view) and phallus with postgonites (lateral view). 17 18. Bicellaria kocoureki sp. nov. 19 20. Bicellaria longisetosa Chvála. 21 22. Bicellaria mera Collin. 23 24. Bicellaria nigra (Meigen). Scale bars = 0.10 mm." httpUri="https://zenodo.org/record/224162/files/figure.png" pageId="25" pageNumber="276">21</figureCitation>
)................................................................................................ 26
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26 (25) Hypandrium with rather long (
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), slender and straight processes (
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)..................
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<emphasis id="B919436CFFB419402BC33D7B9538D37E" bold="true" box="[1259,1350,298,319]" italics="true" pageId="25" pageNumber="276">B. spuria</emphasis>
(Fallén)
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- Hypandrium with rather short (
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), apically foot-like bent and broadened processes (
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)..
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<emphasis id="B919436CFFB419402A203D169529D31D" bold="true" box="[1288,1367,327,348]" italics="true" pageId="25" pageNumber="276">B. mera</emphasis>
Collin
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27 (23) Face broad in ventral part (usually broader than
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, more than 4 facets combined); mid femur with complete row of anteroventrals at least half as long as depth of femur at mid-length; hind tibia with ventral setae about twice as long as depth of tibia; prescutellar depression paler than remaining parts of mesoscutum in dorsal view.....
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<emphasis id="B919436CFFB419402BF63DCE9529D3F5" bold="true" box="[1246,1367,415,436]" italics="true" pageId="25" pageNumber="276">B. subpilosa</emphasis>
Collin
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- Face narrower in ventral part (usually less than
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, about 3 facets combined); mid femur with anteroventrals shorter than one-third of femur depth at mid-length; hind tibia with ventral setae at most slightly longer than depth of tibia; prescutellar depression paler than remaining parts of mesoscutum in dorsal view or evenly dark...................... 28
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28 (27) Usually two dorsal setae on fore trochanter; usually more than 9 setae in posthumeral and intrahumeral areas; hind tibia swollen (usually about
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at broadest point).........................................
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<emphasis id="B919436CFFB419402BF43E63954CD006" bold="true" box="[1244,1330,562,583]" italics="true" pageId="25" pageNumber="276">B. uvens</emphasis>
Melander
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- Usually a single seta on fore trochanter; usually less than 9 setae in posthumeral and intrahumeral areas; hind tibia less swollen (usually less than
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at broadest point).................................................... 29
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29 (28) Tergite 8 at least in basal half lustrous....................................................
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<emphasis id="B919436CFFB419402BC33EDB9538D0DE" bold="true" box="[1259,1350,650,671]" italics="true" pageId="25" pageNumber="276">B. spuria</emphasis>
(Fallén)
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- Tergite 8 almost entirely microtrichose......................................................
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<emphasis id="B919436CFFB419402A203EF69529D0FD" bold="true" box="[1288,1367,679,700]" italics="true" pageId="25" pageNumber="276">B. mera</emphasis>
Collin
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