498 lines
58 KiB
XML
498 lines
58 KiB
XML
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<mods:title>The genus Pupa Röding, 1798 (Mollusca, Gastropoda, Acteonidae) in New Caledonia with notes on Recent species</mods:title>
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<mods:namePart>Valdés, Ángel</mods:namePart>
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<mods:affiliation>Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. & aavaldes @ cpp. edu; https: // orcid. org / 0000 - 0002 - 2347 - 4896</mods:affiliation>
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<mods:namePart>Feliciano, Kendall</mods:namePart>
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<mods:affiliation>Department of Biological Sciences, California State Polytechnic University, 3801 West Temple Avenue, Pomona, California 91768, USA. & https: // orcid. org / 0000 - 0003 - 0536 - 2628</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Malaquias, Manuel A. E.</mods:namePart>
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<mods:affiliation>Department of Natural History, University Museum of Bergen, University of Bergen, PB 7800, N- 5020 Bergen, Norway. https: // orcid. org / 0000 - 0002 - 9668 - 945 X</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<mods:number>2023-04-21</mods:number>
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<mods:number>5270</mods:number>
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<mods:number>3</mods:number>
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<paragraph blockId="23.[151,465,1778,1841]" box="[151,465,1778,1805]" pageId="23" pageNumber="494">
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<heading bold="true" box="[151,465,1778,1805]" fontSize="11" level="1" pageId="23" pageNumber="494" reason="1">
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<taxonomicName ID-CoL="7T6P4" authority="(Angas, 1871)" baseAuthorityName="Angas" baseAuthorityYear="1871" box="[151,465,1778,1805]" class="Gastropoda" family="Acteonidae" genus="Pupa" kingdom="Animalia" order="Cephalaspidea" pageId="23" pageNumber="494" phylum="Mollusca" rank="species" species="nivea">
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<emphasis bold="true" box="[151,465,1778,1805]" pageId="23" pageNumber="494">
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<emphasis bold="true" box="[151,287,1778,1805]" italics="true" pageId="23" pageNumber="494">Pupa nivea</emphasis>
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(
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<bibRefCitation author="Angas, G. F." box="[303,457,1778,1804]" pageId="23" pageNumber="494" pagination="13 - 21" refId="ref21498" refString="Angas, G. F. (1871) Description of thirty-four new species of shells from Australia. Proceedings of the Zoological Society of London, 1871, 13 - 21, pl. 1." type="journal article" year="1871">Angas, 1871</bibRefCitation>
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)
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</emphasis>
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</taxonomicName>
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||
</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,315,1814,1841]" pageId="23" pageNumber="494" type="description">
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<paragraph blockId="23.[151,465,1778,1841]" box="[151,315,1814,1841]" pageId="23" pageNumber="494">
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(
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<figureCitation box="[159,307,1814,1841]" captionStart="FIGURE 15" captionStartId="25.[152,255,1667,1691]" captionTargetBox="[151,1436,525,1644]" captionTargetId="figure-156@25.[151,1436,525,1644]" captionTargetPageId="25" captionText="FIGURE 15. Photographs of shell specimens of Pupa nivea (Angas, 1871) (A–G) and Pupa tragulata Iredale, 1936 (H–J), ventral view. A–B. Syntypes of Buccinulus niveus Angas, 1871 (NHMUK 1871.7.5.24), ©NHMUK, photos by K. Webb. C. Holotype of Buccinulus kirki Hutton, 1873 (NMNZ M.001814), ©NMNZ. D–F. Possible syntypes of Buccinulus intermedius Angas, 1879 (NHMUK 1879.1.31.4), ©NHMUK, photos by K. Webb. G. Holotype of Buccinulus gracilis Kirk, 1882 (NMNZ M.000050), ©NMNZ. H. Holotype of Pupa tragulata Iredale, 1936 (AM C.60701), ©AM. I. Holotype of Pupa strigosa sekii Habe, 1958 (NSMT Mo.39815), ©NSMT, photo by K. Hasegawa. J. Holotype of Pupa sinica Lin, 1989 (IOCAS Mo 26971), ©IOCAS, photo by H. Wang." figureDoi="http://doi.org/10.5281/zenodo.7860404" httpUri="https://zenodo.org/record/7860404/files/figure.png" pageId="23" pageNumber="494">Figs. 15A–G</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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<subSubSection pageId="23" pageNumber="494" type="reference_group">
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||
<paragraph blockId="23.[151,1302,1883,2030]" box="[151,1302,1883,1907]" pageId="23" pageNumber="494">
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<treatmentCitationGroup box="[151,1302,1883,1907]" pageId="23" pageNumber="494">
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<treatmentCitation author="Angas, G. F." box="[151,508,1883,1907]" page="19" pageId="23" pageNumber="494" year="1871">
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<taxonomicName ID-CoL="NL3H" authority="Angas, 1871: 19" authorityName="Angas" authorityPageNumber="19" authorityYear="1871" box="[151,508,1883,1907]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="23" pageNumber="494" phylum="Mollusca" rank="species" species="niveus">
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<emphasis box="[151,335,1883,1907]" italics="true" pageId="23" pageNumber="494">Buccinulus niveus</emphasis>
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<materialsCitation box="[340,508,1883,1907]" country="Australia" county="Port Jackson" location="Near Sow" municipality="Pigs Reef" pageId="23" pageNumber="494" specimenCount="1">
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<bibRefCitation author="Angas, G. F." box="[340,508,1883,1907]" pageId="23" pageNumber="494" pagination="13 - 21" refId="ref21498" refString="Angas, G. F. (1871) Description of thirty-four new species of shells from Australia. Proceedings of the Zoological Society of London, 1871, 13 - 21, pl. 1." type="journal article" year="1871">Angas, 1871: 19</bibRefCitation>
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</materialsCitation>
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||
</taxonomicName>
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</treatmentCitation>
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, pl. 1, fig. 27.
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||
<typeStatus box="[655,705,1883,1907]" pageId="23" pageNumber="494">Type</typeStatus>
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||
locality:
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<location LSID="urn:lsid:plazi:treatment:03C38798FF9A46778ED72AEF4471FA53:8EB56055FF9A46788D612B4742BCF8C5" box="[801,902,1883,1907]" country="Australia" county="Port Jackson" municipality="Pigs Reef" name="Near Sow" pageId="23" pageNumber="494">Near Sow</location>
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and
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<collectingMunicipality box="[951,1051,1883,1907]" pageId="23" pageNumber="494">Pigs Reef</collectingMunicipality>
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,
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<collectingCounty box="[1062,1193,1883,1907]" pageId="23" pageNumber="494">Port Jackson</collectingCounty>
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,
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<collectingCountry box="[1202,1298,1883,1907]" name="Australia" pageId="23" pageNumber="494">Australia</collectingCountry>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="23.[151,1302,1883,2030]" box="[151,882,1913,1938]" pageId="23" pageNumber="494">
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<treatmentCitationGroup box="[151,882,1913,1938]" pageId="23" pageNumber="494">
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<treatmentCitation author="Hutton, F. W." box="[151,498,1913,1937]" page="51" pageId="23" pageNumber="494" year="1873">
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<taxonomicName ID-CoL="NL3G" authority="Hutton, 1873: 51" authorityName="Hutton" authorityPageNumber="51" authorityYear="1873" box="[151,498,1913,1937]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="23" pageNumber="494" phylum="Mollusca" rank="species" species="kirki">
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<emphasis box="[151,316,1913,1937]" italics="true" pageId="23" pageNumber="494">Buccinulus kirki</emphasis>
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<materialsCitation box="[323,498,1913,1937]" country="New Zealand" location="Omaha" pageId="23" pageNumber="494" specimenCount="1">
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<bibRefCitation author="Hutton, F. W." box="[323,498,1913,1937]" pageId="23" pageNumber="494" refId="ref23963" refString="Hutton, F. W. (1873) Catalogue of the Marine Mollusca of New Zealand with Diagnoses of the Species. Didsbury, Wellington, New Zealand, xx pp., 1 pl., 116 pp." type="book" year="1873">Hutton, 1873: 51</bibRefCitation>
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</materialsCitation>
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.
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<typeStatus box="[507,557,1914,1938]" pageId="23" pageNumber="494">Type</typeStatus>
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locality:
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<location LSID="urn:lsid:plazi:treatment:03C38798FF9A46778ED72AEF4471FA53:8EB56055FF9A46788CCD2B6543E3F827" box="[653,729,1913,1937]" country="New Zealand" name="Omaha" pageId="23" pageNumber="494">Omaha</location>
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||
,
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||
<collectingCountry box="[739,878,1913,1937]" name="New Zealand" pageId="23" pageNumber="494">New Zealand</collectingCountry>
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.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="23.[151,1302,1883,2030]" box="[151,949,1944,1968]" pageId="23" pageNumber="494">
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<treatmentCitationGroup box="[151,949,1944,1968]" pageId="23" pageNumber="494">
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<treatmentCitation author="Hutton, F. W." box="[151,506,1944,1968]" page="51" pageId="23" pageNumber="494" year="1873">
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<taxonomicName ID-CoL="NL3B" authority="Hutton, 1873: 51" authorityName="Hutton" authorityPageNumber="51" authorityYear="1873" box="[151,506,1944,1968]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="23" pageNumber="494" phylum="Mollusca" rank="species" species="albus">
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||
<emphasis box="[151,325,1944,1968]" italics="true" pageId="23" pageNumber="494">Buccinulus albus</emphasis>
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||
<materialsCitation box="[331,506,1944,1968]" country="New Zealand" location="New Zealand" pageId="23" pageNumber="494" specimenCount="1">
|
||
<bibRefCitation author="Hutton, F. W." box="[331,506,1944,1968]" pageId="23" pageNumber="494" refId="ref23963" refString="Hutton, F. W. (1873) Catalogue of the Marine Mollusca of New Zealand with Diagnoses of the Species. Didsbury, Wellington, New Zealand, xx pp., 1 pl., 116 pp." type="book" year="1873">Hutton, 1873: 51</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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||
<typeStatus box="[516,566,1944,1968]" pageId="23" pageNumber="494">Type</typeStatus>
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||
locality: undermined [
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||
<collectingCountry box="[795,939,1944,1968]" name="New Zealand" pageId="23" pageNumber="494">New Zealand</collectingCountry>
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].
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||
</treatmentCitationGroup>
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||
</paragraph>
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<paragraph blockId="23.[151,1302,1883,2030]" box="[151,1183,1974,1999]" pageId="23" pageNumber="494">
|
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<treatmentCitationGroup box="[151,1183,1974,1999]" pageId="23" pageNumber="494">
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<treatmentCitation author="Angas, G. F." box="[151,576,1975,1999]" page="862" pageId="23" pageNumber="494" year="1879">
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||
<taxonomicName ID-CoL="NL3F" authority="Angas, 1879: 862" authorityName="Angas" authorityPageNumber="862" authorityYear="1879" box="[151,576,1975,1999]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="23" pageNumber="494" phylum="Mollusca" rank="species" species="intermedius">
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<emphasis box="[151,390,1975,1999]" italics="true" pageId="23" pageNumber="494">Buccinulus intermedius</emphasis>
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||
<materialsCitation box="[395,576,1975,1999]" location="Aldinga Bay" pageId="23" pageNumber="494" specimenCount="1" stateProvince="South Australia">
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<bibRefCitation author="Angas, G. F." box="[395,576,1975,1999]" pageId="23" pageNumber="494" pagination="861 - 864" refId="ref21535" refString="Angas, G. F. (1879) Descriptions of ten species of marine shells from the province of South Australia. Proceedings of the Zoological Society of London, 1878, 861 - 864, pl. 54. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08029. x" type="journal article" year="1879">Angas, 1879: 862</bibRefCitation>
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</materialsCitation>
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</taxonomicName>
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</treatmentCitation>
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||
, pl. 54, fig. 11.
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<typeStatus box="[735,785,1975,1999]" pageId="23" pageNumber="494">Type</typeStatus>
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locality:
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<location LSID="urn:lsid:plazi:treatment:03C38798FF9A46778ED72AEF4471FA53:8EB56055FF9A46788D2F2BAB42CBF879" box="[879,1009,1975,1999]" name="Aldinga Bay" pageId="23" pageNumber="494" stateProvince="South Australia">Aldinga Bay</location>
|
||
,
|
||
<collectingRegion box="[1019,1179,1975,1999]" country="Australia" name="South Australia" pageId="23" pageNumber="494">South Australia</collectingRegion>
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.
|
||
</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="23.[151,1302,1883,2030]" box="[151,940,2005,2030]" pageId="23" pageNumber="494">
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<treatmentCitationGroup box="[151,940,2005,2030]" pageId="23" pageNumber="494">
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<treatmentCitation author="Kirk, T. W." box="[151,518,2005,2029]" page="268" pageId="23" pageNumber="494" year="1882">
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<taxonomicName ID-CoL="NL3D" authority="Kirk, 1882: 268" authorityName="Kirk" authorityPageNumber="268" authorityYear="1882" box="[151,518,2005,2029]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="23" pageNumber="494" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis box="[151,347,2005,2029]" italics="true" pageId="23" pageNumber="494">Buccinulus gracilis</emphasis>
|
||
<materialsCitation box="[354,518,2005,2029]" country="New Zealand" location="Wellington" pageId="23" pageNumber="494" specimenCount="1" stateProvince="Wellington">
|
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<bibRefCitation author="Kirk, T. W." box="[354,518,2005,2029]" pageId="23" pageNumber="494" pagination="268 - 269" refId="ref24560" refString="Kirk, T. W. (1882) Additions to the list of New Zealand shells. Transactions and Proceedings of the New Zealand Institute, 14, 268 - 269." type="journal article" year="1882">Kirk, 1882: 268</bibRefCitation>
|
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</materialsCitation>
|
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</taxonomicName>
|
||
</treatmentCitation>
|
||
.
|
||
<typeStatus box="[527,577,2006,2030]" pageId="23" pageNumber="494">Type</typeStatus>
|
||
locality:
|
||
<collectingRegion box="[673,788,2005,2029]" country="New Zealand" name="Wellington" pageId="23" pageNumber="494">Wellington</collectingRegion>
|
||
,
|
||
<collectingCountry box="[797,936,2005,2029]" name="New Zealand" pageId="23" pageNumber="494">New Zealand</collectingCountry>
|
||
.
|
||
</treatmentCitationGroup>
|
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</paragraph>
|
||
</subSubSection>
|
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<subSubSection pageId="24" pageNumber="495" type="materials_examined">
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<paragraph blockId="24.[151,1437,150,1509]" pageId="24" pageNumber="495">
|
||
<emphasis bold="true" box="[151,321,150,177]" pageId="24" pageNumber="495">Type material</emphasis>
|
||
.
|
||
<emphasis box="[339,570,151,177]" italics="true" pageId="24" pageNumber="495">
|
||
<taxonomicName authorityName="Angas" authorityYear="1871" box="[339,542,151,177]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="niveus">Buccinulus niveus</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
<specimenCount box="[570,696,151,177]" pageId="24" pageNumber="495" type="generic" typeStatus="syntypes">2 syntypes</specimenCount>
|
||
, dry shells,
|
||
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|
||
long (
|
||
<collectionCode box="[1083,1191,151,177]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="24" pageNumber="495" type="Museum">NHMUK</collectionCode>
|
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1871.7.5.24), photos examined, one of them illustrated by
|
||
<bibRefCitation author="Beu, A. G." box="[586,711,187,213]" pageId="24" pageNumber="495" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu (2004</bibRefCitation>
|
||
: fig. 24D);
|
||
<emphasis box="[847,1057,187,213]" italics="true" pageId="24" pageNumber="495">
|
||
<taxonomicName authorityName="Hutton" authorityYear="1873" box="[847,1029,187,213]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="kirki">Buccinulus kirki</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
<typeStatus box="[1057,1164,187,213]" pageId="24" pageNumber="495">Holotype</typeStatus>
|
||
, dry shell,
|
||
<quantity box="[1295,1377,187,213]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="495" unit="mm" value="22.0">22 mm</quantity>
|
||
long (
|
||
<collectionCode box="[159,243,223,249]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="24" pageNumber="495" type="Museum">NMNZ</collectionCode>
|
||
M.001814), photo examined;
|
||
<taxonomicName authorityName="Hutton" authorityYear="1873" box="[600,796,223,249]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="albus">
|
||
<emphasis box="[600,796,223,249]" italics="true" pageId="24" pageNumber="495">Buccinulus albus</emphasis>
|
||
</taxonomicName>
|
||
—type material originally at
|
||
<collectionCode box="[1137,1222,223,249]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="24" pageNumber="495" type="Museum">NMNZ</collectionCode>
|
||
, currently missing (
|
||
<bibRefCitation author="Marshall, B. A." box="[159,333,259,285]" pageId="24" pageNumber="495" pagination="1 - 85" refId="ref25036" refString="Marshall, B. A. (1996) Molluscan name-bearing types in the Museum of New Zealand Te Papa Tongarewa. Records of the Museum of New Zealand Te Papa Tongarewa, 9, 1 - 85." type="journal article" year="1996">Marshall, 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Beu, A. G." box="[344,463,259,285]" pageId="24" pageNumber="495" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu, 2004</bibRefCitation>
|
||
);
|
||
<taxonomicName authorityName="Angas" authorityYear="1879" box="[482,745,259,285]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="intermedius">
|
||
<emphasis box="[482,745,259,285]" italics="true" pageId="24" pageNumber="495">Buccinulus intermedius</emphasis>
|
||
</taxonomicName>
|
||
—3 possible
|
||
<typeStatus box="[892,993,259,285]" pageId="24" pageNumber="495">syntypes</typeStatus>
|
||
, dry shells,
|
||
<quantity box="[1124,1259,258,285]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" metricValueMax="9.7" metricValueMin="7.5" pageId="24" pageNumber="495" unit="mm" value="8.6" valueMax="9.7" valueMin="7.5">7.5–9.7 mm</quantity>
|
||
long (
|
||
<collectionCode box="[1328,1436,259,285]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="24" pageNumber="495" type="Museum">NHMUK</collectionCode>
|
||
1879.1.31.4), photos examined;
|
||
<emphasis box="[513,755,295,321]" italics="true" pageId="24" pageNumber="495">
|
||
<taxonomicName authorityName="Kirk" authorityYear="1882" box="[513,725,295,321]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="gracilis">Buccinulus gracilis</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
<typeStatus box="[755,862,295,321]" pageId="24" pageNumber="495">Holotype</typeStatus>
|
||
, dry shell,
|
||
<quantity box="[984,1063,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="24" pageNumber="495" unit="mm" value="25.0">25 mm</quantity>
|
||
long (
|
||
<collectionCode box="[1134,1218,295,321]" country="New Zealand" httpUri="http://grbio.org/cool/fg7h-t7tk" name="Museum of New Zealand Te Papa Tongarewa" pageId="24" pageNumber="495" type="Museum">NMNZ</collectionCode>
|
||
M.000050).
|
||
</paragraph>
|
||
<paragraph blockId="24.[151,1437,150,1509]" pageId="24" pageNumber="495">
|
||
<emphasis bold="true" box="[199,457,331,357]" pageId="24" pageNumber="495">External morphology</emphasis>
|
||
. Live animal unknown. Shell solid, very narrow, elongate, widest at 1/3 from anterior end, with convex to parallel sides, rounded to elongate anterior end (
|
||
<figureCitation box="[909,1058,366,393]" captionStart="FIGURE 15" captionStartId="25.[152,255,1667,1691]" captionTargetBox="[151,1436,525,1644]" captionTargetId="figure-156@25.[151,1436,525,1644]" captionTargetPageId="25" captionText="FIGURE 15. Photographs of shell specimens of Pupa nivea (Angas, 1871) (A–G) and Pupa tragulata Iredale, 1936 (H–J), ventral view. A–B. Syntypes of Buccinulus niveus Angas, 1871 (NHMUK 1871.7.5.24), ©NHMUK, photos by K. Webb. C. Holotype of Buccinulus kirki Hutton, 1873 (NMNZ M.001814), ©NMNZ. D–F. Possible syntypes of Buccinulus intermedius Angas, 1879 (NHMUK 1879.1.31.4), ©NHMUK, photos by K. Webb. G. Holotype of Buccinulus gracilis Kirk, 1882 (NMNZ M.000050), ©NMNZ. H. Holotype of Pupa tragulata Iredale, 1936 (AM C.60701), ©AM. I. Holotype of Pupa strigosa sekii Habe, 1958 (NSMT Mo.39815), ©NSMT, photo by K. Hasegawa. J. Holotype of Pupa sinica Lin, 1989 (IOCAS Mo 26971), ©IOCAS, photo by H. Wang." figureDoi="http://doi.org/10.5281/zenodo.7860404" httpUri="https://zenodo.org/record/7860404/files/figure.png" pageId="24" pageNumber="495">Figs. 15A–G</figureCitation>
|
||
). Body whorl large, about 2/3–3/4 of total length. Spire conical, elongate, with 4–5 whorls. Suture slightly channeled. Aperture narrow, elongate, wider anteriorly, narrowing abruptly towards posterior end, ending at 1/4–1/6 of the posterior end of first whorl. Columellar margin thickened, slightly oblique, with large, channeled anterior fold starting at anterior end of aperture; small, simple posterior fold located close to aperture mid-length, separated from anterior fold by short gap. Protoconch not observed. Sculpture composed of numerous punctuated spiral grooves. Punctuations conspicuous, irregular, oval, situated next to each other, often fused together, within each groove. Grooves separated by gaps wider than grooves. Shell color uniformly white, with faint irregular orange pigment in some specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1253,654,681]" pageId="24" pageNumber="495" type="description">
|
||
<paragraph blockId="24.[151,1437,150,1509]" box="[199,1253,654,681]" pageId="24" pageNumber="495">
|
||
<emphasis bold="true" box="[199,414,654,681]" pageId="24" pageNumber="495">Geographic range</emphasis>
|
||
. Possibly endemic to
|
||
<collectingCountry box="[661,811,655,681]" name="New Zealand" pageId="24" pageNumber="495">New Zealand</collectingCountry>
|
||
and southern
|
||
<collectingCountry box="[967,1070,655,681]" name="Australia" pageId="24" pageNumber="495">Australia</collectingCountry>
|
||
(present paper).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="24" pageNumber="495" type="discussion">
|
||
<paragraph blockId="24.[151,1437,150,1509]" pageId="24" pageNumber="495">
|
||
<emphasis bold="true" box="[199,308,691,717]" pageId="24" pageNumber="495">Remarks</emphasis>
|
||
. The taxonomic history of the various names introduced for some
|
||
<collectingCountry box="[1118,1274,691,717]" name="New Zealand" pageId="24" pageNumber="495">New Zealand</collectingCountry>
|
||
and southern Australian species is complex and following
|
||
<bibRefCitation author="Beu, A. G." box="[663,791,727,753]" pageId="24" pageNumber="495" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu (2004)</bibRefCitation>
|
||
nearly all of them are currently considered synonyms of
|
||
<taxonomicName authority="(A. Adams, 1855)" baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[151,508,762,789]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[151,290,763,789]" italics="true" pageId="24" pageNumber="495">Pupa affinis</emphasis>
|
||
(A.
|
||
<bibRefCitation author="Adams, A." box="[344,499,762,789]" pageId="24" pageNumber="495" pagination="58 - 62" refId="ref21400" refString="Adams, A. (1855) Monographs of Actaeon and Solidula, two genera of gasteropodous Mollusca with descriptions of several new species from the Cumingian Collection. Proceedings of the Zoological Society of London, 1854, 58 - 62. https: // doi. org / 10.1111 / j. 1469 - 7998.1854. tb 07231. x" type="journal article" year="1855">Adams, 1855</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Several nominal species were described by
|
||
<bibRefCitation author="Angas, G. F." box="[1023,1171,763,789]" pageId="24" pageNumber="495" pagination="13 - 21" refId="ref21498" refString="Angas, G. F. (1871) Description of thirty-four new species of shells from Australia. Proceedings of the Zoological Society of London, 1871, 13 - 21, pl. 1." type="journal article" year="1871">Angas (1871</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Angas, G. F." box="[1186,1246,763,789]" pageId="24" pageNumber="495" pagination="861 - 864" refId="ref21535" refString="Angas, G. F. (1879) Descriptions of ten species of marine shells from the province of South Australia. Proceedings of the Zoological Society of London, 1878, 861 - 864, pl. 54. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08029. x" type="journal article" year="1879">1879</bibRefCitation>
|
||
),
|
||
<bibRefCitation author="Hutton, F. W." box="[1267,1431,763,789]" pageId="24" pageNumber="495" refId="ref23963" refString="Hutton, F. W. (1873) Catalogue of the Marine Mollusca of New Zealand with Diagnoses of the Species. Didsbury, Wellington, New Zealand, xx pp., 1 pl., 116 pp." type="book" year="1873">Hutton (1873)</bibRefCitation>
|
||
, and
|
||
<bibRefCitation author="Kirk, T. W." box="[198,333,799,825]" pageId="24" pageNumber="495" pagination="268 - 269" refId="ref24560" refString="Kirk, T. W. (1882) Additions to the list of New Zealand shells. Transactions and Proceedings of the New Zealand Institute, 14, 268 - 269." type="journal article" year="1882">Kirk (1882)</bibRefCitation>
|
||
, including
|
||
<taxonomicName authority="Angas, 1871" authorityName="Angas" authorityYear="1871" box="[457,809,799,825]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="niveus">
|
||
<emphasis box="[457,661,799,825]" italics="true" pageId="24" pageNumber="495">Buccinulus niveus</emphasis>
|
||
<bibRefCitation author="Angas, G. F." box="[665,809,799,825]" pageId="24" pageNumber="495" pagination="13 - 21" refId="ref21498" refString="Angas, G. F. (1871) Description of thirty-four new species of shells from Australia. Proceedings of the Zoological Society of London, 1871, 13 - 21, pl. 1." type="journal article" year="1871">Angas, 1871</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Angas, 1879" authorityName="Angas" authorityYear="1879" box="[821,1234,799,825]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="intermedius">
|
||
<emphasis box="[821,1084,799,825]" italics="true" pageId="24" pageNumber="495">Buccinulus intermedius</emphasis>
|
||
<bibRefCitation author="Angas, G. F." box="[1089,1234,799,825]" pageId="24" pageNumber="495" pagination="861 - 864" refId="ref21535" refString="Angas, G. F. (1879) Descriptions of ten species of marine shells from the province of South Australia. Proceedings of the Zoological Society of London, 1878, 861 - 864, pl. 54. https: // doi. org / 10.1111 / j. 1469 - 7998.1878. tb 08029. x" type="journal article" year="1879">Angas, 1879</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hutton, 1873" authorityName="Hutton" authorityYear="1873" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="albus">
|
||
<emphasis box="[1244,1437,799,825]" italics="true" pageId="24" pageNumber="495">Buccinulus albus</emphasis>
|
||
<bibRefCitation author="Hutton, F. W." box="[151,303,835,861]" pageId="24" pageNumber="495" refId="ref23963" refString="Hutton, F. W. (1873) Catalogue of the Marine Mollusca of New Zealand with Diagnoses of the Species. Didsbury, Wellington, New Zealand, xx pp., 1 pl., 116 pp." type="book" year="1873">Hutton, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Hutton, 1873" authorityName="Hutton" authorityYear="1873" box="[316,660,835,861]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="kirki">
|
||
<emphasis box="[316,499,835,861]" italics="true" pageId="24" pageNumber="495">Buccinulus kirki</emphasis>
|
||
<bibRefCitation author="Hutton, F. W." box="[508,660,835,861]" pageId="24" pageNumber="495" refId="ref23963" refString="Hutton, F. W. (1873) Catalogue of the Marine Mollusca of New Zealand with Diagnoses of the Species. Didsbury, Wellington, New Zealand, xx pp., 1 pl., 116 pp." type="book" year="1873">Hutton, 1873</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Kirk, 1882" authorityName="Kirk" authorityYear="1882" box="[673,1025,835,861]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis box="[673,891,835,861]" italics="true" pageId="24" pageNumber="495">Buccinulus gracilis</emphasis>
|
||
<bibRefCitation author="Kirk, T. W." box="[899,1025,835,861]" pageId="24" pageNumber="495" pagination="268 - 269" refId="ref24560" refString="Kirk, T. W. (1882) Additions to the list of New Zealand shells. Transactions and Proceedings of the New Zealand Institute, 14, 268 - 269." type="journal article" year="1882">Kirk, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Kirk, 1882" authorityName="Kirk" authorityYear="1882" box="[1086,1432,835,861]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="huttoni">
|
||
<emphasis box="[1086,1298,835,861]" italics="true" pageId="24" pageNumber="495">Buccinulus huttoni</emphasis>
|
||
<bibRefCitation author="Kirk, T. W." box="[1307,1432,835,861]" pageId="24" pageNumber="495" pagination="268 - 269" refId="ref24560" refString="Kirk, T. W. (1882) Additions to the list of New Zealand shells. Transactions and Proceedings of the New Zealand Institute, 14, 268 - 269." type="journal article" year="1882">Kirk, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Suter, H." box="[151,292,871,897]" pageId="24" pageNumber="495" refId="ref26877" refString="Suter, H. (1913) Manual of the New Zealand Mollusca. With an Atlas of Quarto Plates. John MacKay, Wellington, xxiii + 1120 pp. https: // doi. org / 10.5962 / bhl. title. 1716" type="book" year="1913">Suter (1913)</bibRefCitation>
|
||
abandoned the use of the genus name
|
||
<taxonomicName authorityName="Herrmannsen" authorityYear="1852" box="[723,847,871,897]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[723,847,871,897]" italics="true" pageId="24" pageNumber="495">Buccinulus</emphasis>
|
||
</taxonomicName>
|
||
in favor of
|
||
<taxonomicName authorityName="Draparnaud" authorityYear="1801" box="[979,1038,871,896]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[979,1038,871,896]" italics="true" pageId="24" pageNumber="495">Pupa</emphasis>
|
||
</taxonomicName>
|
||
and regarded
|
||
<taxonomicName authorityName="Hutton" authorityYear="1873" box="[1197,1275,871,897]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="kirki">
|
||
<emphasis box="[1197,1275,871,897]" italics="true" pageId="24" pageNumber="495">P. kirki</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Kirk" authorityYear="1882" box="[1330,1436,871,897]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="huttoni">
|
||
<emphasis box="[1330,1436,871,897]" italics="true" pageId="24" pageNumber="495">P. huttoni</emphasis>
|
||
</taxonomicName>
|
||
as synonyms of
|
||
<taxonomicName baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[341,441,907,933]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[341,441,907,933]" italics="true" pageId="24" pageNumber="495">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
(see remarks on
|
||
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[644,764,907,933]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="solidula">
|
||
<emphasis box="[644,764,907,933]" italics="true" pageId="24" pageNumber="495">P. solidula</emphasis>
|
||
</taxonomicName>
|
||
) but considered
|
||
<taxonomicName authorityName="Folin & Berillon" authorityYear="1877" box="[960,1040,907,933]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="alba">
|
||
<emphasis box="[960,1040,907,933]" italics="true" pageId="24" pageNumber="495">P. alba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Kirk" authorityYear="1882" box="[1100,1216,907,933]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis box="[1100,1216,907,933]" italics="true" pageId="24" pageNumber="495">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
as a valid, distinct species of
|
||
<taxonomicName authorityName="Draparnaud" authorityYear="1801" box="[270,329,943,968]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="genus">
|
||
<emphasis box="[270,329,943,968]" italics="true" pageId="24" pageNumber="495">Pupa</emphasis>
|
||
</taxonomicName>
|
||
. On the contrary,
|
||
<bibRefCitation author="Finlay, H. J." box="[532,685,943,969]" pageId="24" pageNumber="495" pagination="320 - 485" refId="ref22950" refString="Finlay, H. J. (1926) A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the Royal Society of New Zealand, 57, 320 - 485." type="journal article" year="1926">Finlay (1926)</bibRefCitation>
|
||
rejected the use of
|
||
<taxonomicName baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[905,1003,943,969]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[905,1003,943,969]" italics="true" pageId="24" pageNumber="495">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<collectingCountry box="[1050,1200,943,969]" name="New Zealand" pageId="24" pageNumber="495">New Zealand</collectingCountry>
|
||
specimens, regarded
|
||
<taxonomicName authorityName="Folin & Berillon" authorityYear="1877" box="[151,230,979,1005]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="alba">
|
||
<emphasis box="[151,230,979,1005]" italics="true" pageId="24" pageNumber="495">P. alba</emphasis>
|
||
</taxonomicName>
|
||
as valid name, and designated a
|
||
<typeStatus box="[605,694,979,1005]" pageId="24" pageNumber="495">neotype</typeStatus>
|
||
for this species. Moreover,
|
||
<bibRefCitation author="Finlay, H. J." box="[1010,1165,979,1005]" pageId="24" pageNumber="495" pagination="320 - 485" refId="ref22950" refString="Finlay, H. J. (1926) A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the Royal Society of New Zealand, 57, 320 - 485." type="journal article" year="1926">Finlay (1926)</bibRefCitation>
|
||
determined that
|
||
<taxonomicName authorityName="Hutton" authorityYear="1873" box="[1356,1436,979,1005]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="kirki">
|
||
<emphasis box="[1356,1436,979,1005]" italics="true" pageId="24" pageNumber="495">P. kirki</emphasis>
|
||
</taxonomicName>
|
||
is also valid and a senior synonym of
|
||
<taxonomicName authorityName="Kirk" authorityYear="1882" box="[603,720,1015,1041]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis box="[603,720,1015,1041]" italics="true" pageId="24" pageNumber="495">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
, but that
|
||
<taxonomicName authorityName="Kirk" authorityYear="1882" box="[836,946,1015,1041]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="huttoni">
|
||
<emphasis box="[836,946,1015,1041]" italics="true" pageId="24" pageNumber="495">P. huttoni</emphasis>
|
||
</taxonomicName>
|
||
was possibly different (see remarks on
|
||
<taxonomicName baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="solidula">
|
||
<emphasis italics="true" pageId="24" pageNumber="495">P. solidula</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<bibRefCitation author="Rudman, W. B." box="[266,444,1051,1077]" pageId="24" pageNumber="495" pagination="205 - 214" refId="ref26117" refString="Rudman, W. B. (1971) The family Acteonidae (Opisthobranchia, Gastropoda) in New Zealand, Journal of the Malacological Society of Australia, 2, 205 - 214. https: // doi. org / 10.1080 / 00852988.1971.10673853" type="journal article" year="1971">Rudman (1971)</bibRefCitation>
|
||
indicated that the specimens identified by
|
||
<bibRefCitation author="Suter, H." box="[935,1077,1051,1077]" pageId="24" pageNumber="495" refId="ref26877" refString="Suter, H. (1913) Manual of the New Zealand Mollusca. With an Atlas of Quarto Plates. John MacKay, Wellington, xxiii + 1120 pp. https: // doi. org / 10.5962 / bhl. title. 1716" type="book" year="1913">Suter (1913)</bibRefCitation>
|
||
as
|
||
<taxonomicName baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[1118,1217,1051,1077]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[1118,1217,1051,1077]" italics="true" pageId="24" pageNumber="495">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
are in fact
|
||
<taxonomicName authorityName="Hutton" authorityYear="1873" box="[1349,1429,1051,1077]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="kirki">
|
||
<emphasis box="[1349,1429,1051,1077]" italics="true" pageId="24" pageNumber="495">P. kirki</emphasis>
|
||
</taxonomicName>
|
||
, which according to him is a distinct and valid species, highly variable in color.
|
||
<bibRefCitation author="Rudman, W. B." box="[1064,1242,1087,1113]" pageId="24" pageNumber="495" pagination="205 - 214" refId="ref26117" refString="Rudman, W. B. (1971) The family Acteonidae (Opisthobranchia, Gastropoda) in New Zealand, Journal of the Malacological Society of Australia, 2, 205 - 214. https: // doi. org / 10.1080 / 00852988.1971.10673853" type="journal article" year="1971">Rudman (1971)</bibRefCitation>
|
||
also commented that the description of
|
||
<taxonomicName authorityName="Folin & Berillon" authorityYear="1877" box="[400,477,1123,1149]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="alba">
|
||
<emphasis box="[400,477,1123,1149]" italics="true" pageId="24" pageNumber="495">P. alba</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Hutton, F. W." box="[517,676,1123,1149]" pageId="24" pageNumber="495" refId="ref23963" refString="Hutton, F. W. (1873) Catalogue of the Marine Mollusca of New Zealand with Diagnoses of the Species. Didsbury, Wellington, New Zealand, xx pp., 1 pl., 116 pp." type="book" year="1873">Hutton (1873)</bibRefCitation>
|
||
was totally inadequate and that would fit several species. Moreover,
|
||
<bibRefCitation author="Rudman, W. B." box="[151,330,1159,1185]" pageId="24" pageNumber="495" pagination="205 - 214" refId="ref26117" refString="Rudman, W. B. (1971) The family Acteonidae (Opisthobranchia, Gastropoda) in New Zealand, Journal of the Malacological Society of Australia, 2, 205 - 214. https: // doi. org / 10.1080 / 00852988.1971.10673853" type="journal article" year="1971">Rudman (1971)</bibRefCitation>
|
||
regarded
|
||
<bibRefCitation author="Finlay, H. J." box="[446,620,1159,1185]" pageId="24" pageNumber="495" pagination="320 - 485" refId="ref22950" refString="Finlay, H. J. (1926) A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the Royal Society of New Zealand, 57, 320 - 485." type="journal article" year="1926">Finlay’s (1926)</bibRefCitation>
|
||
designation of a
|
||
<typeStatus box="[822,911,1159,1185]" pageId="24" pageNumber="495">neotype</typeStatus>
|
||
for
|
||
<taxonomicName authorityName="Folin & Berillon" authorityYear="1877" box="[961,1041,1159,1185]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="alba">
|
||
<emphasis box="[961,1041,1159,1185]" italics="true" pageId="24" pageNumber="495">P. alba</emphasis>
|
||
</taxonomicName>
|
||
invalid and suggested this species name should be rendered as a
|
||
<emphasis box="[494,652,1195,1221]" italics="true" pageId="24" pageNumber="495">nomen nudum</emphasis>
|
||
.
|
||
<bibRefCitation author="Beu, A. G." box="[667,795,1195,1221]" pageId="24" pageNumber="495" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu (2004)</bibRefCitation>
|
||
expanded the synonymy of
|
||
<taxonomicName baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[1115,1214,1195,1221]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[1115,1214,1195,1221]" italics="true" pageId="24" pageNumber="495">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
to include the
|
||
<collectingCountry name="New Zealand" pageId="24" pageNumber="495">New Zealand</collectingCountry>
|
||
species
|
||
<taxonomicName baseAuthorityName="Angas" baseAuthorityYear="1871" box="[341,432,1231,1256]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="nivea">
|
||
<emphasis box="[341,432,1231,1256]" italics="true" pageId="24" pageNumber="495">P. nivea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Morelet" authorityYear="1851" box="[492,643,1231,1257]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="intermedia">
|
||
<emphasis box="[492,643,1231,1257]" italics="true" pageId="24" pageNumber="495">P. intermedia</emphasis>
|
||
</taxonomicName>
|
||
, but also species names originally described from other Indo-Pacific locations such as
|
||
<taxonomicName authority="Gould, 1859" authorityName="Gould" authorityYear="1859" box="[344,723,1266,1293]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="strigosus">
|
||
<emphasis box="[344,575,1267,1293]" italics="true" pageId="24" pageNumber="495">Buccinulus strigosus</emphasis>
|
||
<bibRefCitation author="Gould, A. A." box="[581,723,1266,1293]" pageId="24" pageNumber="495" pagination="138 - 142" refId="ref23364" refString="Gould, A. A. (1859) Descriptions of new species of shells brought home by the North Pacific Exploring Expedition. Proceedings of the Boston Society of Natural History, 7, 138 - 142. https: // doi. org / 10.5962 / bhl. part. 4821" type="journal article" year="1859">Gould, 1859</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Reeve, 1865" authorityName="Reeve" authorityYear="1865" box="[732,1079,1266,1293]" class="Gastropoda" family="Acteonidae" genus="Tornatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="fumata">
|
||
<emphasis box="[732,932,1267,1293]" italics="true" pageId="24" pageNumber="495">Tornatella fumata</emphasis>
|
||
<bibRefCitation author="Reeve, L. A." box="[937,1079,1266,1293]" pageId="24" pageNumber="495" pagination="1 - 4" refId="ref25851" refString="Reeve, L. A. (1865) Monograph of the genus Tornatella. In: Conchologia Iconica: or, Illustrations of the Shells of Molluscous Animals. Vol. 15. Contains Monographs of the Genera Solarium. Sigaretus. Marginella. Ancillaria. Ovulum. Erato. Carinaria. Tornatella. Pyramidella. Cerithium. Eulima. Vertagus. Telescopium. Cerithidea. Pyrazus. Lampania. Tympanotonos. Leiostraca. Niso. Potamides. Lovell Reeve & Co., London, pp. [89 - 95], pls. 1 - 4" type="book chapter" year="1865">Reeve, 1865</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Dunker, 1882" authorityName="Dunker" authorityYear="1882" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="fraterculus">
|
||
<emphasis box="[1089,1341,1267,1293]" italics="true" pageId="24" pageNumber="495">Buccinulus fraterculus</emphasis>
|
||
<bibRefCitation author="Dunker, G." pageId="24" pageNumber="495" refId="ref22856" refString="Dunker, G. (1882) Index Molluscorum Maris Japonici, Conscriptus et Tabulis Iconum XVI Illustratus. Fischer, Cassellis Cattorum [Kassel], vii + 301 pp., 11 pls. https: // doi. org / 10.5962 / bhl. title. 10277" type="book" year="1882">Dunker, 1882</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Pilsbry, 1917" authorityName="Pilsbry" authorityYear="1917" box="[221,550,1303,1329]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="thaanumi">
|
||
<emphasis box="[221,393,1303,1329]" italics="true" pageId="24" pageNumber="495">Pupa thaanumi</emphasis>
|
||
<bibRefCitation author="Pilsbry, H. A." box="[399,550,1303,1329]" pageId="24" pageNumber="495" pagination="207 - 230" refId="ref25559" refString="Pilsbry, H. A. (1917) Marine Mollusks of Hawaii, I - III. Proceedings of the Academy of Natural Sciences of Philadelphia, 69, 207 - 230, pls. 14 - 15." type="journal article" year="1917">Pilsbry, 1917</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName authority="Iredale, 1936" authorityName="Iredale" authorityYear="1936" box="[561,888,1303,1329]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="tragulata">
|
||
<emphasis box="[561,730,1303,1329]" italics="true" pageId="24" pageNumber="495">Pupa tragulata</emphasis>
|
||
<bibRefCitation author="Iredale, T." box="[737,888,1303,1329]" pageId="24" pageNumber="495" pagination="267 - 340" refId="ref24124" refString="Iredale, T. (1936) Australian molluscan notes No. 2. Records of the Australian Museum, 19, 267 - 340, pls. 20 - 24. https: // doi. org / 10.3853 / j. 0067 - 1975.19.1936.704" type="journal article" year="1936">Iredale, 1936</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName authority="Habe, 1958" authorityName="Habe" authorityYear="1958" box="[946,1299,1302,1329]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="subSpecies" species="strigosa" subSpecies="sekii">
|
||
<emphasis box="[946,1160,1303,1329]" italics="true" pageId="24" pageNumber="495">Pupa strigosa sekii</emphasis>
|
||
<bibRefCitation author="Habe, T." box="[1167,1299,1302,1329]" pageId="24" pageNumber="495" pagination="117 - 120" refId="ref23499" refString="Habe, T. (1958) On the shell-bearing opisthobranchiate molluscan fauna from off Choshi, Chiba Pref., Japan. Annotationes Zoologicae Japonenses, 31, 117 - 120." type="journal article" year="1958">Habe, 1958</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Beu, A. G." box="[1310,1436,1303,1329]" pageId="24" pageNumber="495" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu (2004)</bibRefCitation>
|
||
also disagreed with Sutter (1913) and moved
|
||
<taxonomicName authorityName="Folin & Berillon" authorityYear="1877" box="[657,734,1339,1365]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="alba">
|
||
<emphasis box="[657,734,1339,1365]" italics="true" pageId="24" pageNumber="495">P. alba</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Kirk" authorityYear="1882" box="[788,901,1339,1365]" class="Gastropoda" family="Acteonidae" genus="Buccinulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cephalaspidea" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="gracilis">
|
||
<emphasis box="[788,901,1339,1365]" italics="true" pageId="24" pageNumber="495">P. gracilis</emphasis>
|
||
</taxonomicName>
|
||
back to the synonymy of
|
||
<taxonomicName baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[1188,1285,1339,1365]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[1188,1285,1339,1365]" italics="true" pageId="24" pageNumber="495">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation author="Burn, R." box="[1299,1436,1339,1365]" pageId="24" pageNumber="495" pagination="1 - 42" refId="ref22159" refString="Burn, R. (2006) A checklist and bibliography of the Opisthobranchia (Mollusca: Gastropoda) of Victoria and the Bass Strait area, south-eastern Australia. Museum Victoria Science Reports, 10, 1 - 42. https: // doi. org / 10.24199 / j. mvsr. 2006.10" type="journal article" year="2006">Burn (2006)</bibRefCitation>
|
||
agreed with
|
||
<bibRefCitation author="Beu, A. G." box="[287,413,1375,1401]" pageId="24" pageNumber="495" pagination="111 - 265" refId="ref21645" refString="Beu, A. G. (2004) Marine Mollusca of oxygen isotope stages of the last 2 million years in New Zealand. Part 1: Revised generic positions and recognition of warm-water and cool-water migrants. Journal of the Royal Society of New Zealand, 34, 111 - 265. [With an appendix: Age and correlation of Ototoka tephra by Alloway, B. V., Pillans, B. J., Naish, T. R. & Westgate, J. A.] https: // doi. org / 10.1080 / 03014223.2004.9517766" type="journal article" year="2004">Beu (2004)</bibRefCitation>
|
||
considering
|
||
<taxonomicName baseAuthorityName="Angas" baseAuthorityYear="1871" box="[555,643,1375,1400]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="nivea">
|
||
<emphasis box="[555,643,1375,1400]" italics="true" pageId="24" pageNumber="495">P. nivea</emphasis>
|
||
</taxonomicName>
|
||
as a synonym of
|
||
<taxonomicName baseAuthorityName="A. Adams" baseAuthorityYear="1855" box="[833,930,1375,1401]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="affinis">
|
||
<emphasis box="[833,930,1375,1401]" italics="true" pageId="24" pageNumber="495">P. affinis</emphasis>
|
||
</taxonomicName>
|
||
but because of slight differences in the radula supported the retention of
|
||
<taxonomicName authorityName="Iredale" authorityYear="1936" box="[456,590,1411,1437]" class="Gastropoda" family="Acteonidae" genus="Pupa" higherTaxonomySource="CoL" kingdom="Animalia" order="Architaenioglossa" pageId="24" pageNumber="495" phylum="Mollusca" rank="species" species="tragulata">
|
||
<emphasis box="[456,590,1411,1437]" italics="true" pageId="24" pageNumber="495">P. tragulata</emphasis>
|
||
</taxonomicName>
|
||
as a distinct deeper-water species found in
|
||
<collectingCountry box="[1097,1249,1411,1437]" name="New Zealand" pageId="24" pageNumber="495">New Zealand</collectingCountry>
|
||
and
|
||
<collectingRegion country="Australia" name="New South Wales" pageId="24" pageNumber="495">New South Wales</collectingRegion>
|
||
,
|
||
<collectingCountry box="[232,338,1447,1473]" name="Australia" pageId="24" pageNumber="495">Australia</collectingCountry>
|
||
. Further research on this species complex, including molecular techniques, is necessary to clarify the taxonomic status and validity of these available names but this is beyond the scope of the present paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |