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<document id="2B22F10FB8F8BC0D4C74DDD1AFBC65A7" ID-DOI="10.5281/zenodo.12612354" ID-ISSN="2064-2474" ID-Zenodo-Dep="12612354" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719865439135" checkinUser="felipe" docAuthor="Papp, L &amp; Sevcik, J" docDate="2011" docId="03C38795BE72FFE2263444C5EAAB067E" docLanguage="en" docName="ActaZoolAcadSciH.57.2.139-159.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 57 (2)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Leptomorphus utarensis Papp &amp; Sevcik, 2011, sp. n." docType="treatment" docVersion="2" lastPageNumber="156" masterDocId="FFFAFFEDBE7FFFF324694111EB490318" masterDocTitle="Eight New Oriental And Australasian Species Of Leptomorphus (Diptera: Mycetophilidae)" masterLastPageNumber="159" masterPageNumber="139" pageNumber="152" updateTime="1719866131965" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="820AC1C47986C224B32AF01729FF663C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="30A260BEE2DD7DA71001B3667DF8DCE2">
<mods:title id="77183642D195C410AD0875E7CE11E28F">Eight New Oriental And Australasian Species Of Leptomorphus (Diptera: Mycetophilidae)</mods:title>
</mods:titleInfo>
<mods:name id="EBA925F70DC19ADFA269D1D42B6215BB" type="personal">
<mods:role id="F9B0DEA51A2291F7F7694772E90B813A">
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<mods:namePart id="0A1B3B99292C85743AA82846CE946D3F">Papp, L</mods:namePart>
</mods:name>
<mods:name id="B3D6F7DCA0827BB624A9A9B500FC369C" type="personal">
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<mods:namePart id="43FF466B42314DB4372CC6684B9DBF96">Sevcik, J</mods:namePart>
</mods:name>
<mods:typeOfResource id="318CD6855613A1755FB10F70540B3FCD">text</mods:typeOfResource>
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<mods:title id="5BDD20C5FE6B4B4FD0D6E84312D1CBDD">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
</mods:titleInfo>
<mods:part id="748B4318299EBDD7A26C2896BB489CC4">
<mods:date id="8E4BB6691541B15E02C7243C8320C51B">2011</mods:date>
<mods:detail id="A3DEEDDFDF459482B0C73F2FFF467A88" type="pubDate">
<mods:number id="0C69119DA87C9605468ECC03A4CC4392">2011-06-10</mods:number>
</mods:detail>
<mods:detail id="27B1B09F8FF51AC4421CBCEC3F98D963" type="volume">
<mods:number id="9BB279AC856C7A772F916178185B9613">57</mods:number>
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<mods:start id="49032CB1E63F8B0D35C7511928B1121B">139</mods:start>
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<mods:classification id="5B86DFCBEC862D6773F0EA3177C50F6D">journal article</mods:classification>
<mods:identifier id="645EB6320FB8A5A5CF80D8179AD334D2" type="DOI">10.5281/zenodo.12612354</mods:identifier>
<mods:identifier id="F3277BA9F783983E9720240C7988CC1C" type="ISSN">2064-2474</mods:identifier>
<mods:identifier id="42346C23745F917C02A98B216E5043FE" type="Zenodo-Dep">12612354</mods:identifier>
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<treatment id="03C38795BE72FFE2263444C5EAAB067E" LSID="urn:lsid:plazi:treatment:03C38795BE72FFE2263444C5EAAB067E" httpUri="http://treatment.plazi.org/id/03C38795BE72FFE2263444C5EAAB067E" lastPageId="17" lastPageNumber="156" pageId="13" pageNumber="152">
<subSubSection id="C3706508BE72FFFE263444C5E89F06F6" box="[605,982,1492,1518]" pageId="13" pageNumber="152" type="nomenclature">
<paragraph id="8BD53683BE72FFFE263444C5E89F06F6" blockId="13.[605,982,1492,1518]" box="[605,982,1492,1518]" pageId="13" pageNumber="152">
<heading id="D09D81EFBE72FFFE263444C5E89F06F6" box="[605,982,1492,1518]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="152" reason="2">
<taxonomicName id="4C6A4D00BE72FFFE263444C5E8D906F6" ID-CoL="8CJ4L" authorityName="Papp &amp; Sevcik" authorityYear="2011" box="[605,912,1492,1518]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="152" phylum="Arthropoda" rank="species" species="utarensis" status="sp. nov.">
<emphasis id="B91EEA91BE72FFFE263444C5E8D906F6" bold="true" box="[605,912,1492,1518]" pageId="13" pageNumber="152">Leptomorphus utarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57EABE72FFFE27FE44C5E89F06F6" box="[919,982,1492,1518]" pageId="13" pageNumber="152" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3706508BE72FFE226D144E6EAAB067E" lastPageId="17" lastPageNumber="156" pageId="13" pageNumber="152" type="description">
<paragraph id="8BD53683BE72FFFE26D144E6E8330509" blockId="13.[696,890,1527,1553]" box="[696,890,1527,1553]" pageId="13" pageNumber="152">
<heading id="D09D81EFBE72FFFE26D144E6E8330509" box="[696,890,1527,1553]" centered="true" fontSize="11" level="3" pageId="13" pageNumber="152" reason="3">
(
<figureCitation id="13512A06BE72FFFE26A944E6E8010509" box="[704,840,1527,1553]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="13" pageNumber="152">Figs 50–57</figureCitation>
,
<figureCitation id="13512A06BE72FFFE273D44E6E83A0509" box="[852,883,1527,1553]" captionStart="Fig" captionStartId="17.[584,619,878,899]" captionTargetBox="[396,1191,426,843]" captionTargetId="figure-297@17.[396,1191,424,849]" captionTargetPageId="17" captionText="Fig. 60. Leptomorphus utarensis sp. n., egg" figureDoi="http://doi.org/10.5281/zenodo.12612408" httpUri="https://zenodo.org/record/12612408/files/figure.png" pageId="13" pageNumber="152">60</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BD53683BE72FFFE2517472AE88A0591" blockId="13.[317,1270,1595,1817]" pageId="13" pageNumber="152">
<materialsCitation id="3B023CDEBE72FFFE2517472AE88A0591" collectingDate="1985-07-25" collectionCode="BMNH" collectorName="Bone N. P." country="Indonesia" location="Sulawesi" pageId="13" pageNumber="152" specimenCount="1" stateProvince="Sulawesi Utara" typeStatus="holotype">
<typeStatus id="54D18821BE72FFFE2517472AEA9E0548" box="[382,471,1595,1616]" pageId="13" pageNumber="152" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED7BAE46BE72FFFE2674472AE9220548" box="[541,619,1595,1616]" pageId="13" pageNumber="152">BMNH</collectionCode>
):
<collectingCountry id="F37D7613BE72FFFE2617472AE9960548" box="[638,735,1595,1616]" name="Indonesia" pageId="13" pageNumber="152">Indonesia</collectingCountry>
:
<collectingRegion id="49AEF861BE72FFFE2683472AE80D0548" box="[746,836,1595,1616]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="152">Sulawesi</collectingRegion>
: Utara, Dumoga–
<collectorName id="269F5355BE72FFFE279B472AEF280548" box="[1010,1121,1595,1616]" pageId="13" pageNumber="152">Bone N. P.</collectorName>
, “1440 camp”, 30.6.–
<date id="FFD41043BE72FFFE25114749EA950575" box="[376,476,1624,1645]" pageId="13" pageNumber="152" value="1985-07-25">
<collectingDate id="EF90E9ABBE72FFFE25114749EA950575" box="[376,476,1624,1645]" pageId="13" pageNumber="152" value="1985-07-25">25.7.1985</collectingDate>
</date>
,
<collectingMethod id="522B4E94BE72FFFE258C4749E9160575" box="[485,607,1624,1645]" pageId="13" pageNumber="152">Malaise trap</collectingMethod>
. (left apical 4 flagellomeres lost, some tarsomeres, incl. all right hind tarsus, as well as all but fore left 5th tarsomeres with claws, lost).
</materialsCitation>
</paragraph>
<paragraph id="8BD53683BE72FFFE25174780EFE40401" blockId="13.[317,1270,1595,1817]" pageId="13" pageNumber="152">
<materialsCitation id="3B023CDEBE72FFFE25174780E8F105DB" collectingDate="1985-07-25" collectionCode="HNMH" collectorName="Bone N. P." country="Indonesia" location="Sulawesi" pageId="13" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<typeStatus id="54D18821BE72FFFE25174780EA9705BE" box="[382,478,1681,1702]" pageId="13" pageNumber="152" type="paratype">Paratypes</typeStatus>
.
<specimenCount id="9D6CFD0ABE72FFFE258F4780E96F05BE" box="[486,550,1681,1702]" count="1" pageId="13" pageNumber="152" type="male">1 male</specimenCount>
(
<collectionCode id="ED7BAE46BE72FFFE26584780E9C805BE" box="[561,641,1681,1702]" pageId="13" pageNumber="152">HNMH</collectionCode>
): data same as for the holotype (postabdomen from segment 7 with genitalia are prepared and kept in a plastic microvial with glycerol)
</materialsCitation>
.
<materialsCitation id="3B023CDEBE72FFFE27A947BFE98605F8" collectingDate="1985-06-20" collectionCode="BMNH" collectorName="Bone N. P." country="Indonesia" location="Toraut" pageId="13" pageNumber="152" specimenCount="1" specimenCount-female="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D6CFD0ABE72FFFE27A947BFEF5805DB" box="[960,1041,1710,1731]" count="1" pageId="13" pageNumber="152" type="female">1 female</specimenCount>
(
<collectionCode id="ED7BAE46BE72FFFE207447BFEF2205DB" box="[1053,1131,1710,1731]" pageId="13" pageNumber="152">BMNH</collectionCode>
): ibid.,
<location id="8EB56058BE72FFFE20D947BFEFBB05DB" LSID="urn:lsid:plazi:treatment:03C38795BE72FFE2263444C5EAAB067E:8EB56058BE72FFFE20D947BFEFBB05DB" box="[1200,1266,1710,1731]" name="Toraut" pageId="13" pageNumber="152">Toraut</location>
, forest edge, 15.6.–
<date id="FFD41043BE72FFFE258047DAE90705F8" box="[489,590,1739,1760]" pageId="13" pageNumber="152" value="1985-06-20">
<collectingDate id="EF90E9ABBE72FFFE258047DAE90705F8" box="[489,590,1739,1760]" pageId="13" pageNumber="152" value="1985-06-20">20.6.1985</collectingDate>
</date>
,
<collectingMethod id="522B4E94BE72FFFE263F47DAE98605F8" box="[598,719,1739,1760]" pageId="13" pageNumber="152">Malaise trap</collectingMethod>
</materialsCitation>
.
<materialsCitation id="3B023CDEBE72FFFE26BE47DAEFE00401" collectingDate="1985-08-10" collectingDateMax="1985-08-19" collectingDateMin="1985-08-10" collectionCode="MNHN, NMW" collectorName="A. H. Kirk-Spriggs. &amp; Citandui" country="Indonesia" county="Expedition" location="Toraut" municipality="Dumoga Irrigation Project" pageId="13" pageNumber="152" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Utara" typeStatus="paratype">
<specimenCount id="9D6CFD0ABE72FFFE26BE47DAE85F05F8" box="[727,790,1739,1760]" count="1" pageId="13" pageNumber="152" type="male">1 male</specimenCount>
(
<collectionCode id="ED7BAE46BE72FFFE274847DAE83805F8" box="[801,881,1739,1760]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="13" pageNumber="152">MNHN</collectionCode>
):
<collectionCode id="ED7BAE46BE72FFFE271747DAE8F205F8" box="[894,955,1739,1760]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="152">NMW</collectionCode>
<collectingCountry id="F37D7613BE72FFFE27A947DAEF5405F8" box="[960,1053,1739,1760]" name="Indonesia" pageId="13" pageNumber="152">Indonesia</collectingCountry>
<collectingCounty id="62B44E0FBE72FFFE204B47DAEFC205F8" box="[1058,1163,1739,1760]" pageId="13" pageNumber="152">Expedition</collectingCounty>
1985 (Project Wallace),
<collectionCode id="ED7BAE46BE72FFFE25B047F9E95005E5" box="[473,537,1768,1789]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="13" pageNumber="152">NMW</collectionCode>
. Z. 1985. 078 –
<collectingRegion id="49AEF861BE72FFFE26B747F9E83505E5" box="[734,892,1768,1789]" country="Indonesia" name="Sulawesi Utara" pageId="13" pageNumber="152">Sulawesi Utara</collectingRegion>
:
<collectingMunicipality id="6BB1ACF9BE72FFFE27E447F9EFD705E5" box="[909,1182,1768,1789]" pageId="13" pageNumber="152">Dumoga Irrigation Project</collectingMunicipality>
,
<location id="8EB56058BE72FFFE20D947F9EFBB05E5" LSID="urn:lsid:plazi:treatment:03C38795BE72FFE2263444C5EAAB067E:8EB56058BE72FFFE20D947F9EFBB05E5" box="[1200,1266,1768,1789]" country="Indonesia" county="Expedition" municipality="Dumoga Irrigation Project" name="Toraut" pageId="13" pageNumber="152" stateProvince="Sulawesi Utara">Toraut</location>
,
<date id="FFD41043BE72FFFE25544615EA9E0401" box="[317,471,1796,1817]" pageId="13" pageNumber="152" value="1985-08-10" valueMax="1985-08-19" valueMin="1985-08-10">
<collectingDate id="EF90E9ABBE72FFFE25544615EA9E0401" box="[317,471,1796,1817]" pageId="13" pageNumber="152" value="1985-08-10" valueMax="1985-08-19" valueMin="1985-08-10">10–19.viii.1985</collectingDate>
</date>
,
<collectorName id="269F5355BE72FFFE258A4615E9EC0401" box="[483,677,1796,1817]" pageId="13" pageNumber="152">A. H. Kirk-Spriggs.</collectorName>
–
<collectingMethod id="522B4E94BE72FFFE26D64615E8730401" box="[703,826,1796,1817]" pageId="13" pageNumber="152">Malaise trap</collectingMethod>
, irrigated rice vars.
<collectingRegion id="49AEF861BE72FFFE27964615EF7B0401" box="[1023,1074,1796,1817]" country="Indonesia" name="Aceh" pageId="13" pageNumber="152">Aceh</collectingRegion>
&amp;
<collectorName id="269F5355BE72FFFE203A4615EFE00401" box="[1107,1193,1796,1817]" pageId="13" pageNumber="152">Citandui</collectorName>
</materialsCitation>
.
</paragraph>
<caption id="DF15660BBE71FFFD25544795EF090431" ID-DOI="http://doi.org/10.5281/zenodo.12612393" ID-Zenodo-Dep="12612393" httpUri="https://zenodo.org/record/12612393/files/figure.png" pageId="14" pageNumber="153" startId="14.[317,360,1668,1689]" targetBox="[325,1264,441,1629]" targetPageId="14" targetType="figure">
<paragraph id="8BD53683BE71FFFD25544795EFBE0415" blockId="14.[317,1271,1668,1833]" pageId="14" pageNumber="153">
<emphasis id="B91EEA91BE71FFFD25544795EAE60581" bold="true" box="[317,431,1668,1689]" pageId="14" pageNumber="153">Figs 43–49.</emphasis>
<taxonomicName id="4C6A4D00BE71FFFD25DD4794E9C90582" authorityName="Papp &amp; Sevcik" authorityYear="2011" box="[436,640,1669,1690]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="14" pageNumber="153" phylum="Arthropoda" rank="species" species="papua" status="sp. nov.">
<emphasis id="B91EEA91BE71FFFD25DD4794E9C90582" box="[436,640,1669,1690]" italics="true" pageId="14" pageNumber="153">Leptomorphus papua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57EABE71FFFD26EC4795E9F00581" box="[645,697,1668,1689]" pageId="14" pageNumber="153" rank="species">sp. n.</taxonomicNameLabel>
, holotype, male genitalia: 43 = pregenital (8th) sternite, perpendicular view, 44 = pregenital (8th) tergite, perpendicular view, 45 = contours of tergo-gonocoxal complex, ventral view, 46 = caudal end of tergo-gonocoxal complex with gonostylus in view of the broadest extension of gonostylus (dorsal-sublateral view), 47 = tergo-gonocoxal complex with gonostyli, dorsal view, 48 = same, caudal view, 49 = aedeagus and ejaculatory apodeme, dorsal view.
</paragraph>
<paragraph id="8BD53683BE71FFFD259B4605EF090431" blockId="14.[317,1271,1668,1833]" box="[498,1088,1812,1833]" pageId="14" pageNumber="153">Scale bar: 0.2 mm for Figs 43–45, 49, 0.1 mm for Figs 46–48</paragraph>
</caption>
<caption id="DF15660BBE70FFFC25544795EFBF0414" ID-DOI="http://doi.org/10.5281/zenodo.12612397" ID-Zenodo-Dep="12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="15" pageNumber="154" startId="15.[317,360,1668,1689]" targetBox="[329,1253,449,1636]" targetPageId="15" targetType="figure">
<paragraph id="8BD53683BE70FFFC25544795EFBF0414" blockId="15.[317,1270,1668,1833]" pageId="15" pageNumber="154">
<emphasis id="B91EEA91BE70FFFC25544795EAF90581" bold="true" box="[317,432,1668,1689]" pageId="15" pageNumber="154">Figs 50–57.</emphasis>
<taxonomicName id="4C6A4D00BE70FFFC25DE4795E9E90581" authorityName="Papp &amp; Sevcik" authorityYear="2011" box="[439,672,1668,1689]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="154" phylum="Arthropoda" rank="species" species="utarensis" status="sp. nov.">
<emphasis id="B91EEA91BE70FFFC25DE4795E9E90581" box="[439,672,1668,1689]" italics="true" pageId="15" pageNumber="154">Leptomorphus utarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57EABE70FFFC26CF4795E9950581" box="[678,732,1668,1689]" pageId="15" pageNumber="154" rank="species">sp. n.</taxonomicNameLabel>
, paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51,
</paragraph>
</caption>
<paragraph id="8BD53683BE70FFFC26EE4605E8E20431" blockId="15.[317,1270,1668,1833]" box="[647,939,1812,1833]" pageId="15" pageNumber="154">
56–57,
<quantity id="4C929B66BE70FFFC26A64605E8510431" box="[719,792,1812,1833]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="15" pageNumber="154" unit="mm" value="0.1">0.1 mm</quantity>
for
<figureCitation id="13512A06BE70FFFC27294605E8E20431" box="[832,939,1812,1833]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="15" pageNumber="154">Figs 52–55</figureCitation>
</paragraph>
<caption id="DF15660BBE6FFFE32666454DEF6A0769" ID-DOI="http://doi.org/10.5281/zenodo.12612400" ID-Zenodo-Dep="12612400" box="[527,1059,1116,1138]" httpUri="https://zenodo.org/record/12612400/files/figure.png" pageId="16" pageNumber="155" startId="16.[527,562,1116,1137]" targetBox="[394,1192,424,1088]" targetPageId="16" targetType="figure">
<paragraph id="8BD53683BE6FFFE32666454DEF6A0769" blockId="16.[527,1059,1116,1138]" box="[527,1059,1116,1138]" pageId="16" pageNumber="155">
<emphasis id="B91EEA91BE6FFFE32666454DE9120769" bold="true" box="[527,603,1116,1137]" pageId="16" pageNumber="155">Fig. 58.</emphasis>
<taxonomicName id="4C6A4D00BE6FFFE32608454CE82D076A" authorityName="Papp &amp; Sevcik" authorityYear="2011" box="[609,868,1117,1138]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="155" phylum="Arthropoda" rank="species" species="ascutellatus" status="sp. nov.">
<emphasis id="B91EEA91BE6FFFE32608454CE82D076A" box="[609,868,1117,1138]" italics="true" pageId="16" pageNumber="155">Leptomorphus ascutellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57EABE6FFFE32703454DE8D60769" box="[874,927,1116,1137]" pageId="16" pageNumber="155" rank="species">sp. n.</taxonomicNameLabel>
, male habitus
</paragraph>
</caption>
<caption id="DF15660BBE6FFFE325A647D8EF2A05C6" ID-DOI="http://doi.org/10.5281/zenodo.12612404" ID-Zenodo-Dep="12612404" box="[463,1123,1737,1758]" httpUri="https://zenodo.org/record/12612404/files/figure.png" pageId="16" pageNumber="155" startId="16.[463,498,1737,1758]" targetBox="[402,1185,1179,1708]" targetPageId="16" targetType="figure">
<paragraph id="8BD53683BE6FFFE325A647D8EF2A05C6" blockId="16.[463,1123,1737,1758]" box="[463,1123,1737,1758]" pageId="16" pageNumber="155">
<emphasis id="B91EEA91BE6FFFE325A647D8E95205C6" bold="true" box="[463,539,1737,1758]" pageId="16" pageNumber="155">Fig. 59.</emphasis>
<taxonomicName id="4C6A4D00BE6FFFE3264847D8E86D05C6" authorityName="Papp &amp; Sevcik" authorityYear="2011" box="[545,804,1737,1758]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="16" pageNumber="155" phylum="Arthropoda" rank="species" species="ascutellatus" status="sp. nov.">
<emphasis id="B91EEA91BE6FFFE3264847D8E86D05C6" box="[545,804,1737,1758]" italics="true" pageId="16" pageNumber="155">Leptomorphus ascutellatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57EABE6FFFE3274347D8E81605C6" box="[810,863,1737,1758]" pageId="16" pageNumber="155" rank="species">sp. n.</taxonomicNameLabel>
, detail of female scutellum
</paragraph>
</caption>
<caption id="DF15660BBE6EFFE22621427FE8A2009B" ID-DOI="http://doi.org/10.5281/zenodo.12612408" ID-Zenodo-Dep="12612408" box="[584,1003,878,899]" httpUri="https://zenodo.org/record/12612408/files/figure.png" pageId="17" pageNumber="156" startId="17.[584,619,878,899]" targetBox="[396,1191,426,843]" targetPageId="17" targetType="figure">
<paragraph id="8BD53683BE6EFFE22621427FE8A2009B" blockId="17.[584,1003,878,899]" box="[584,1003,878,899]" pageId="17" pageNumber="156">
<emphasis id="B91EEA91BE6EFFE22621427FE9DD009B" bold="true" box="[584,660,878,899]" pageId="17" pageNumber="156">Fig. 60.</emphasis>
<taxonomicName id="4C6A4D00BE6EFFE226F3427FE8CB009B" authorityName="Papp &amp; Sevcik" authorityYear="2011" box="[666,898,878,899]" class="Insecta" family="Mycetophilidae" genus="Leptomorphus" kingdom="Animalia" order="Diptera" pageId="17" pageNumber="156" phylum="Arthropoda" rank="species" species="utarensis" status="sp. nov.">
<emphasis id="B91EEA91BE6EFFE226F3427FE8CB009B" box="[666,898,878,899]" italics="true" pageId="17" pageNumber="156">Leptomorphus utarensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22D57EABE6EFFE227E1427FE8F4009B" box="[904,957,878,899]" pageId="17" pageNumber="156" rank="species">sp. n.</taxonomicNameLabel>
, egg
</paragraph>
</caption>
<paragraph id="8BD53683BE6EFFE2251742AFEFAA00CB" blockId="17.[317,1271,958,1296]" box="[382,1251,958,979]" pageId="17" pageNumber="156">
Measurements (
<typeStatus id="54D18821BE6EFFE2267C42AFE92600CB" box="[533,623,958,979]" pageId="17" pageNumber="156" type="holotype">holotype</typeStatus>
): wing length:
<quantity id="4C929B66BE6EFFE2269142AFE80700CB" box="[760,846,958,979]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" pageId="17" pageNumber="156" unit="mm" value="5.75">5.75 mm</quantity>
, head + thorax
<quantity id="4C929B66BE6EFFE227B242AFEF7800CB" box="[987,1073,958,979]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" pageId="17" pageNumber="156" unit="mm" value="1.75">1.75 mm</quantity>
, abdomen
<quantity id="4C929B66BE6EFFE220FC42AFEF9600CB" box="[1173,1247,958,979]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="17" pageNumber="156" unit="mm" value="5.1">5.1 mm</quantity>
.
</paragraph>
<paragraph id="8BD53683BE6EFFE2251742CAEF8407EB" blockId="17.[317,1271,958,1296]" pageId="17" pageNumber="156">
Male terminalia. Setae on sternite 8 almost symmetrical (
<figureCitation id="13512A06BE6EFFE227DB42CAE8B200E8" box="[946,1019,987,1008]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 50</figureCitation>
), setae long thick and arranged in one row; sternite 8 submedially with a darker transverse thickening. Tergite 8 about as long as broad, dark setae evenly emerge on its apical 2/3 (
<figureCitation id="13512A06BE6EFFE227594505E83E0731" box="[816,887,1044,1065]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 51</figureCitation>
). Tergo-gonocoxal complex longer than broad, caudal lobes (
<figureCitation id="13512A06BE6EFFE2266C4520E905075E" box="[517,588,1073,1094]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 52</figureCitation>
) comparatively short, broadly bifid, i.e. the complex with deep medial emargination (
<figureCitation id="13512A06BE6EFFE225A3455FE95B077B" box="[458,530,1102,1123]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 52</figureCitation>
). Medial processes partly overlapping in dorsal view, short and blunt, better seen in caudal view (
<figureCitation id="13512A06BE6EFFE22662457AE9340798" box="[523,637,1131,1152]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Figs 52, 55</figureCitation>
). Gonostyli broad basally (
<figureCitation id="13512A06BE6EFFE227EC457AE8870798" box="[901,974,1131,1152]" captionStart="Figs 18–23" captionStartId="8.[317,360,1699,1720]" captionTargetBox="[322,1261,767,1663]" captionTargetId="figure-86@8.[317,1269,763,1669]" captionTargetPageId="8" captionText="Figs 18–23. Leptomorphus baramensis sp. n., holotype, male genitalia:18 = pregenital (8th) sternite, perpendicular view, 19 = pregenital (8th) tergite, perpendicular view, 20 = caudal end of tergo-gonocoxal complex, dorsal view, 21 = medial processes of gonocoxite, caudal view, 22 = gonostylus, in broadest extension (sublateral view), 23 = aedeagus and ejaculatory apodeme, dorsal view. Scale bar:" figureDoi="http://doi.org/10.5281/zenodo.12612379" httpUri="https://zenodo.org/record/12612379/files/figure.png" pageId="17" pageNumber="156">Fig. 22</figureCitation>
), but apical 2/3 rather narrow. Gonostylus not much different in dorsal view and in its broadest extension (
<figureCitation id="13512A06BE6EFFE220614599EF070785" box="[1032,1102,1160,1181]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 54</figureCitation>
, vs
<figureCitation id="13512A06BE6EFFE220184599EFFE0785" box="[1137,1207,1160,1181]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 53</figureCitation>
). Caudal view (
<figureCitation id="13512A06BE6EFFE225F445B5EAAA07A1" box="[413,483,1188,1209]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 55</figureCitation>
) reveals the large, widely rounded extension of the complex subventrally. Phallic organ medium large, aedeagus short but high (
<figureCitation id="13512A06BE6EFFE2268645D0E81507CE" box="[751,860,1217,1238]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Figs 56–57</figureCitation>
); ejaculatory apodeme comparatively short, rather high (
<figureCitation id="13512A06BE6EFFE225C645CFEABC07EB" box="[431,501,1246,1267]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 56</figureCitation>
), but rather narrow in dorsal view, without lateral (horizontal) lobes (
<figureCitation id="13512A06BE6EFFE2201545CFEF8B07EB" box="[1148,1218,1246,1267]" captionStart="Figs 50–57" captionStartId="15.[317,360,1668,1689]" captionTargetBox="[329,1253,449,1636]" captionTargetId="figure-13@15.[329,1257,444,1639]" captionTargetPageId="15" captionText="Figs 50–57. Leptomorphus utarensis sp. n., paratype, male genitalia: 50 = pregenital (8th) sternite, perpendicular view, 51 = pregenital (8th) tergite, perpendicular view, 52 = caudal end of tergo-gonocoxal complex without gonostylus, dorsal view, 53 = gonostylus, broadest (subdorsal) view, 54 = gonostylus, dorsal view, 55 = tergo-gonocoxal complex with gonostylus, caudal view, 56 = aedeagus and ejaculatory apodeme, lateral view, 57 = same, dorsal view. Scale bar: 0.2 mm for Figs 50–51," figureDoi="http://doi.org/10.5281/zenodo.12612397" httpUri="https://zenodo.org/record/12612397/files/figure.png" pageId="17" pageNumber="156">Fig. 57</figureCitation>
).
</paragraph>
<paragraph id="8BD53683BE6EFFE2251745EAEF920608" blockId="17.[317,1271,958,1296]" box="[382,1243,1275,1296]" pageId="17" pageNumber="156">
Egg (
<figureCitation id="13512A06BE6EFFE225D945EAEABF0608" box="[432,502,1275,1296]" captionStart="Fig" captionStartId="17.[584,619,878,899]" captionTargetBox="[396,1191,426,843]" captionTargetId="figure-297@17.[396,1191,424,849]" captionTargetPageId="17" captionText="Fig. 60. Leptomorphus utarensis sp. n., egg" figureDoi="http://doi.org/10.5281/zenodo.12612408" httpUri="https://zenodo.org/record/12612408/files/figure.png" pageId="17" pageNumber="156">Fig. 60</figureCitation>
). This is a formerly unknown morphological
<typeStatus id="54D18821BE6EFFE227F445EAE88F0608" box="[925,966,1275,1296]" pageId="17" pageNumber="156">type</typeStatus>
of egg (see
<bibRefCitation id="EFFB4B72BE6EFFE2205C45EAEF860608" author="PLACHTER, H." box="[1077,1231,1275,1296]" pageId="17" pageNumber="156" pagination="43 - 63" refId="ref6580" refString="PLACHTER, H. (1981) Chorionic structures of the eggshells of 15 fungus-and root-gnat species (Diptera: Mycetophiliodea). International Journal of Insect Morphology &amp; Embryology 10 (1): 43 - 63." type="journal article" year="1981">
<smallCapsWord id="8D33A05FBE6EFFE2205C45EAEFD10616" baselines="1291,1290" box="[1077,1176,1275,1296]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Plachter" pageId="17" pageNumber="156">PLACHTER</smallCapsWord>
1981
</bibRefCitation>
).
</paragraph>
<paragraph id="8BD53683BE6EFFE225174425EFBC0651" blockId="17.[317,1269,1332,1382]" box="[382,1269,1332,1353]" pageId="17" pageNumber="156">
Etymology. The specific epithet of the new species refers to its
<typeStatus id="54D18821BE6EFFE227B84425E8B30651" box="[977,1018,1332,1353]" pageId="17" pageNumber="156">type</typeStatus>
locality
<collectingRegion id="49AEF861BE6EFFE220224425EF920651" box="[1099,1243,1332,1353]" country="Indonesia" name="Sulawesi Utara" pageId="17" pageNumber="156">Sulawesi Utara</collectingRegion>
(=
</paragraph>
<paragraph id="8BD53683BE6EFFE225544440EAAB067E" blockId="17.[317,1269,1332,1382]" box="[317,482,1361,1382]" pageId="17" pageNumber="156">
<collectingRegion id="49AEF861BE6EFFE225544440EA91067E" box="[317,472,1361,1382]" country="Indonesia" name="Sulawesi Utara" pageId="17" pageNumber="156">North Sulawesi</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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