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<document id="D7DC9682AEB75E38CFB255C252F234D0" ID-CLB-Dataset="24929" ID-DOI="10.5281/zenodo.3532762" ID-GBIF-Dataset="db23a775-d25c-4253-abda-b5a007110f6d" ID-ISSN="0374-1036" ID-Zenodo-Dep="3532762" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1573206962882" checkinUser="plazi" docAuthor="Irmler, Ulrich" docDate="2013" docId="03A3FB753F5A1E64FE20FA4844C70C1B" docLanguage="en" docName="ActaEntMusNatPra.53.1.155-176.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 53 (1)" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Holotrochus hamatus Irmler 2013, sp. nov." docType="treatment" docVersion="7" lastPageNumber="167" masterDocId="FF9A830D3F511E68FF93FFED46660A2F" masterDocTitle="New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)" masterLastPageNumber="176" masterPageNumber="155" pageNumber="166" updateTime="1698764282444" updateUser="ExternalLinkService">
<mods:mods id="FF83755548B7AAB0D4A3BAA802278601" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="85AA489B5A8734C85F3F3B6B94CE76C4">New species of the genus Mimogonia and Holotrochus from South America (Coleoptera: Staphylindae: Osoriinae)</mods:title>
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<mods:namePart id="740071088F3B84B8A4B8E13DA7D78C6A">Irmler, Ulrich</mods:namePart>
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<mods:title id="FB87034E35A408AFF27179F84635A50C">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="CE25F530E9AF3F72479FDD0B18A59397">2013</mods:date>
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<mods:number id="C76436E3711BD1450B5AEDF3AFDDCC03">2013-07-15</mods:number>
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<mods:number id="4D645ADC61A1E999EF9244A2BBB25E00">53</mods:number>
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<mods:classification id="DA6F01DD63DE84BB59FE5FA742ADBD66">journal article</mods:classification>
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<mods:identifier id="48785BE2AAE3E327BCCA58F43269BB45" type="DOI">10.5281/zenodo.3532762</mods:identifier>
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<treatment id="03A3FB753F5A1E64FE20FA4844C70C1B" ID-DOI="http://doi.org/10.5281/zenodo.4427165" ID-GBIF-Taxon="160218245" ID-Zenodo-Dep="4427165" LSID="urn:lsid:plazi:treatment:03A3FB753F5A1E64FE20FA4844C70C1B" httpUri="http://treatment.plazi.org/id/03A3FB753F5A1E64FE20FA4844C70C1B" lastPageId="12" lastPageNumber="167" pageId="11" pageNumber="166">
<subSubSection id="C31019E83F5A1E63FE20FA4844990F92" box="[435,767,1445,1469]" pageId="11" pageNumber="166" type="nomenclature">
<paragraph id="8BB54A633F5A1E63FE20FA4844990F92" blockId="11.[435,767,1445,1494]" box="[435,767,1445,1469]" pageId="11" pageNumber="166">
<heading id="D0FDFD0F3F5A1E63FE20FA4844990F92" bold="true" box="[435,767,1445,1469]" centered="true" fontSize="10" level="3" pageId="11" pageNumber="166" reason="4">
<emphasis id="B97E96713F5A1E63FE20FA4844990F92" bold="true" box="[435,767,1445,1469]" pageId="11" pageNumber="166">
<taxonomicName id="4C0A31E03F5A1E63FE20FA4844C50F92" ID-CoL="3MBQL" authority="Irmler, 2013" authorityName="Irmler" authorityYear="2013" box="[435,675,1445,1469]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="166" phylum="Arthropoda" rank="species" species="hamatus" status="sp. nov.">
<emphasis id="B97E96713F5A1E63FE20FA4844C50F92" bold="true" box="[435,675,1445,1469]" italics="true" pageId="11" pageNumber="166">Holotrochus hamatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D2B0A3F5A1E63FD3AFA4844990F92" box="[681,767,1445,1469]" pageId="11" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5A1E63FD88FA2E44F00FF9" box="[539,662,1475,1494]" pageId="11" pageNumber="166" type="description">
<paragraph id="8BB54A633F5A1E63FD88FA2E44F00FF9" blockId="11.[435,767,1445,1494]" box="[539,662,1475,1494]" pageId="11" pageNumber="166">
(
<figureCitation id="133156E63F5A1E63FDB2FA2E44050FF9" box="[545,611,1475,1494]" captionStart="Figs 8–11. 8" captionStartId="9.[113,150,1524,1543]" captionTargetBox="[150,1050,427,1501]" captionTargetId="figure@9.[143,1058,419,1513]" captionTargetPageId="9" captionText="Figs 8–11. 8– Holotrochus yasuniensis sp. nov.; 9 – H. mrazi sp. nov.; 10 – H. paraguensis sp. nov.; 11 – H. hamatus sp. nov. Body parts:A – aedeagus; B – antenna, C – tergite XII, D – last sternite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3532768" httpUri="https://zenodo.org/record/3532768/files/figure.png" pageId="11" pageNumber="166">Figs 11</figureCitation>
,
<figureCitation id="133156E63F5A1E63FDFFFA2E44F60FF9" box="[620,656,1475,1494]" captionStart="Fig" captionStartId="17.[113,144,1541,1560]" captionTargetBox="[121,1067,217,1507]" captionTargetId="figure@17.[113,1086,211,1515]" captionTargetPageId="17" captionText="Fig. 18. Head, pronotum, and elytra of Holotrochus Erichson,1839 showing shape, punctation, and microsculpture.A – H. paraguensis sp. nov.; B – H.tortilis sp. nov.; C – H.hamatus sp. nov.; D – H.yanuniensis sp. nov.; E – H. mrazi sp. nov.; F – H. lobatus sp. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3532776" httpUri="https://zenodo.org/record/3532776/files/figure.png" pageId="11" pageNumber="166">18C</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5A1E63FFE2FA0044680C1A" pageId="11" pageNumber="166" type="materials_examined">
<paragraph id="8BB54A633F5A1E63FFE2FA0044240C2F" blockId="11.[113,1090,1517,1590]" box="[113,578,1517,1536]" pageId="11" pageNumber="166">
<materialsCitation id="3B62403E3F5A1E63FFE2FA0044580C2F" ID-GBIF-Occurrence="2449305682" box="[113,574,1517,1536]" country="Paraguay" location="Puerto Presidente Stroessner" pageId="11" pageNumber="166" specimenCount="1" typeStatus="holotype">
<emphasis id="B97E96713F5A1E63FFE2FA00468C0C2F" bold="true" box="[113,234,1517,1536]" pageId="11" pageNumber="166">
<typeStatus id="54B1F4C13F5A1E63FFE2FA0046F80C2F" box="[113,158,1517,1536]" pageId="11" pageNumber="166">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F31D0AF33F5A1E63FF7CFA0047590C2F" box="[239,319,1517,1536]" name="Paraguay" pageId="11" pageNumber="166">Paraguay</collectingCountry>
,
<location id="8ED51CB83F5A1E63FEDBFA0044580C2F" LSID="urn:lsid:plazi:treatment:03A3FB753F5A1E64FE20FA4844C70C1B:8ED51CB83F5A1E63FEDBFA0044580C2F" box="[328,574,1517,1536]" country="Paraguay" name="Puerto Presidente Stroessner" pageId="11" pageNumber="166">Puerto Presidente Stroessner</location>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB54A633F5A1E63FFE2F9E544680C1A" blockId="11.[113,1090,1517,1590]" pageId="11" pageNumber="166">
<emphasis id="B97E96713F5A1E63FFE2F9E546900C35" bold="true" box="[113,246,1543,1563]" pageId="11" pageNumber="166">Type material.</emphasis>
<materialsCitation id="3B62403E3F5A1E63FF68F9E544680C1A" ID-GBIF-Occurrence="2449305662" collectingDate="1966-06-01" collectionCode="HNHM" collectorName="L. S. Mahunka'" country="Paraguay" latitude="-25.516666" location="Puerto Presidente Stroessner" longLatPrecision="1246" longitude="-54.6" pageId="11" pageNumber="166" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B1F4C13F5A1E63FF68F9E547280C36" box="[251,334,1544,1563]" pageId="11" pageNumber="166" type="holotype">HOLOTYPE</typeStatus>
:, ‘
<collectingCountry id="F31D0AF33F5A1E63FEE4F9E547AD0C34" box="[375,459,1544,1563]" name="Paraguay" pageId="11" pageNumber="166">Paraguay</collectingCountry>
,
<location id="8ED51CB83F5A1E63FE47F9E544AE0C34" LSID="urn:lsid:plazi:treatment:03A3FB753F5A1E64FE20FA4844C70C1B:8ED51CB83F5A1E63FE47F9E544AE0C34" box="[468,712,1544,1563]" country="Paraguay" latitude="-25.516666" longLatPrecision="1246" longitude="-54.6" name="Puerto Presidente Stroessner" pageId="11" pageNumber="166">Puerto Presidente Stroessner</location>
(
<geoCoordinate id="EE3E2CA43F5A1E63FD40F9E545400C34" box="[723,806,1544,1563]" degrees="54" direction="west" minutes="36" orientation="longitude" pageId="11" pageNumber="166" precision="925" value="-54.6">54°36’ W</geoCoordinate>
,
<geoCoordinate id="EE3E2CA43F5A1E63FCBCF9E5451C0C34" box="[815,890,1544,1563]" degrees="25" direction="south" minutes="31" orientation="latitude" pageId="11" pageNumber="166" precision="925" value="-25.516666">25°31’ S</geoCoordinate>
), Hungarian Soil-Zool. Exp.,
<date id="FFB46CA33F5A1E63FF37F9CF47600C1A" box="[164,262,1570,1589]" pageId="11" pageNumber="166" value="1966-06-01">
<collectingDate id="EFF0954B3F5A1E63FF37F9CF47600C1A" box="[164,262,1570,1589]" pageId="11" pageNumber="166" value="1966-06-01">06.01.1966</collectingDate>
</date>
, leg.
<collectorName id="26FF2FB53F5A1E63FEA0F9CF47D40C1A" box="[307,434,1570,1589]" pageId="11" pageNumber="166">L.S. Mahunka’</collectorName>
(
<collectionCode id="ED1BD2A63F5A1E63FE2EF9CF44650C1A" box="[445,515,1570,1589]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="11" pageNumber="166">HNHM</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5D1E64FF04FF3146B2083B" pageId="12" pageNumber="167" type="diagnosis">
<paragraph id="8BB54A633F5D1E64FF04FF3146B2083B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">
<emphasis id="B97E96713F5D1E64FF04FF3147690ADB" bold="true" box="[151,271,220,244]" pageId="12" pageNumber="167">Diagnosis.</emphasis>
This species certainly belongs to the
<taxonomicName id="4C0A31E03F5D1E64FD25FF30454B0ADB" authorityName="Sharp" authorityYear="1887" box="[694,813,221,244]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="simplex">
<emphasis id="B97E96713F5D1E64FD25FF30454B0ADB" box="[694,813,221,244]" italics="true" pageId="12" pageNumber="167">H. simplex</emphasis>
</taxonomicName>
group due to the size, existence of a margined front edge of pronotum, and the absence of abdominal pubescence.
<taxonomicName id="4C0A31E03F5D1E64FF04FEF047E50B1B" authorityName="Irmler" authorityYear="2013" box="[151,387,285,308]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B97E96713F5D1E64FF04FEF047E50B1B" box="[151,387,285,308]" italics="true" pageId="12" pageNumber="167">Holotrochus hamatus</emphasis>
</taxonomicName>
is characterised by its short antennae with the fourth antennomere wider than long and the pronotum widest in the middle. Similarly short antennae are found in
<taxonomicName id="4C0A31E03F5D1E64FF20FEB047960B5A" authority="Irmler, 1981" authorityName="Irmler" authorityYear="1981" box="[179,496,348,373]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="blackwelderi">
<emphasis id="B97E96713F5D1E64FF20FEB047070B5B" box="[179,353,349,372]" italics="true" pageId="12" pageNumber="167">H. blackwelderi</emphasis>
<bibRefCitation id="EF9B37923F5D1E64FEF4FEB147960B5A" author="IRMLER U." box="[359,496,348,373]" journalOrPublisher="Entomologische Blatter" pageId="12" pageNumber="167" pagination="143 - 152" part="77" refId="ref10117" refString="IRMLER U. 1981 a: Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae). Entomologische Blatter 77: 143 - 152." title="Neue Arten der Gattung Mimogonia Coiffait (1978) aus der Neotropis (Coleoptera, Staphylinidae)" type="journal article" year="1981">Irmler, 1981</bibRefCitation>
</taxonomicName>
, but total size of the body is distinctly smaller. Among the most other similarly sized species of the group, the pronotum is widest in the apical third. In this respect
<taxonomicName id="4C0A31E03F5D1E64FEAAFE7047D00B9B" authorityName="Irmler" authorityYear="2013" box="[313,438,413,436]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B97E96713F5D1E64FEAAFE7047D00B9B" box="[313,438,413,436]" italics="true" pageId="12" pageNumber="167">H. hamatus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C0A31E03F5D1E64FDA1FE7045320B9A" authority="Bernhauer, 1939" authorityName="Bernhauer" authorityYear="1939" box="[562,852,412,437]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="vianai">
<emphasis id="B97E96713F5D1E64FDA1FE7044FE0B9B" box="[562,664,413,436]" italics="true" pageId="12" pageNumber="167">H. vianai</emphasis>
Bernhauer, 1939
</taxonomicName>
within the species found from the same region, but the structure of the endophallus differs significantly. In contrast to
<taxonomicName id="4C0A31E03F5D1E64FF20FE3047560BDB" authorityName="Irmler" authorityYear="2013" box="[179,304,477,500]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B97E96713F5D1E64FF20FE3047560BDB" box="[179,304,477,500]" italics="true" pageId="12" pageNumber="167">H. hamatus</emphasis>
</taxonomicName>
, the endophallus of
<taxonomicName id="4C0A31E03F5D1E64FD9DFE3044150BDB" authorityName="Bernhauer" authorityYear="1939" box="[526,627,477,500]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="vianai">
<emphasis id="B97E96713F5D1E64FD9DFE3044150BDB" box="[526,627,477,500]" italics="true" pageId="12" pageNumber="167">H. vianai</emphasis>
</taxonomicName>
, is simply spiralled with a row of short apical teeth.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5D1E64FF04FDF145A50FDB" pageId="12" pageNumber="167" type="description">
<paragraph id="8BB54A633F5D1E64FF04FDF144D8087B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">
<emphasis id="B97E96713F5D1E64FF04FDF14744081B" bold="true" box="[151,290,540,564]" pageId="12" pageNumber="167">Description.</emphasis>
Length:
<quantity id="4CF2E7863F5D1E64FEEDFDF047B4081B" box="[382,466,541,565]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="12" pageNumber="167" unit="mm" value="4.2">4.2 mm</quantity>
. Colouration black, elytra slightly lighter, brown; fine posterior margin of pronotum reddish; legs and antennae red.
</paragraph>
<paragraph id="8BB54A633F5D1E64FF26FDB147DF097B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">
Head
<quantity id="4CF2E7863F5D1E64FF67FDB14735085B" box="[244,339,604,628]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="12" pageNumber="167" unit="mm" value="0.65">0.65 mm</quantity>
long,
<quantity id="4CF2E7863F5D1E64FE04FDB04793085B" box="[407,501,605,629]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="12" pageNumber="167" unit="mm" value="0.81">0.81 mm</quantity>
wide; eyes not prominent; temples nearly twice as long as eyes; with fine supraocular margin continuing to base of antennae; 4 setiferous punctures within the margin; front edge with several setiferous punctures; on a line between posterior edge of eyes with row of four setiferous punctures; interstices between inner punctures twice as wide as interstices between inner punctures and outer punctures; irregular punctation on vertex distinct; on average, distance between punctures distinctly wider than diameter of punctures; without microsculpture on vertex, but with sparse and fine micro-punctation; surface polished and shiny.
</paragraph>
<paragraph id="8BB54A633F5D1E64FF26FCB045E6099B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">Antennae short, only slightly longer than head; second antennomere globular and only half as long as conical third; fourth antennomere again wider than long; following antennomeres slightly increasing in width; tenth antennomere twice as wide as long.</paragraph>
<paragraph id="8BB54A633F5D1E64FF26FC50474C0E9B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">
Pronotum
<quantity id="4CF2E7863F5D1E64FEB1FC50471809FB" box="[290,382,957,981]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="12" pageNumber="167" unit="mm" value="0.38">0.38 mm</quantity>
long,
<quantity id="4CF2E7863F5D1E64FE2EFC50447F09FB" box="[445,537,957,981]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="12" pageNumber="167" unit="mm" value="0.84">0.84 mm</quantity>
wide; widest in the middle; in posterior half only slightly narrowed, nearly parallel; in anterior half more distinctly narrowed; sides margined; margin continuing to anterior edge; in dorsal aspect, lateral margin partly hidden in anterior half; punctation similar as on head, but punctures posteriad increasing in size; irregular punctation on anterior half as dense as on head, on posterior half slightly denser; on average, interstices between punctures slightly shorter than diameter of punctures; with impunctate midline in posterior half; with very weak remains of microsculpture, but with sparse micro-punctation; surface shiny.
</paragraph>
<paragraph id="8BB54A633F5D1E64FF26FB5144650F1B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">
Elytra
<quantity id="4CF2E7863F5D1E64FF6EFB50473D0EFB" box="[253,347,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="12" pageNumber="167" unit="mm" value="1.07">1.07 mm</quantity>
long,
<quantity id="4CF2E7863F5D1E64FE0DFB50479A0EFB" box="[414,508,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="12" pageNumber="167" unit="mm" value="1.14">1.14 mm</quantity>
wide; shoulders distinct, nearly rectangular and with very short teeth; punctation coarse and moderately dense; in size and density, punctation similar to punctation of posterior pronotum; ground-sculpture coarse and coriaceous; surface less shiny than on head and pronotum.
</paragraph>
<paragraph id="8BB54A633F5D1E64FF26FAD147C70F9B" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">Abdomen with very fine punctation; punctures almost half as large as punctures on head or pronotum; surface with distinct net-like microsculpture; surface less shiny than head and pronotum; last sternite widely rounded at apex, with numerous net-like striae and pair of groups of fine punctures.</paragraph>
<paragraph id="8BB54A633F5D1E64FF26FA5145A50FDB" blockId="12.[151,1127,220,1589]" pageId="12" pageNumber="167">Aedeagus with shortly curved apical part and obtuse apex; parameres shorter than apical part of central lobe; endophallus bilobed; dorsal lobe with row of four teeth.</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5D1E64FF04FA1045B00C3B" box="[151,982,1532,1557]" pageId="12" pageNumber="167" type="etymology">
<paragraph id="8BB54A633F5D1E64FF04FA1045B00C3B" blockId="12.[151,1127,220,1589]" box="[151,982,1532,1557]" pageId="12" pageNumber="167">
<emphasis id="B97E96713F5D1E64FF04FA10477E0C3A" bold="true" box="[151,280,1533,1557]" pageId="12" pageNumber="167">Etymology.</emphasis>
The specific name
<taxonomicName id="4C0A31E03F5D1E64FE74FA1044270C3B" authorityName="Irmler" authorityYear="2013" box="[487,577,1533,1556]" class="Insecta" family="Staphylinidae" genus="Holotrochus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="167" phylum="Arthropoda" rank="species" species="hamatus">
<emphasis id="B97E96713F5D1E64FE74FA1044270C3B" box="[487,577,1533,1556]" italics="true" pageId="12" pageNumber="167">hamatus</emphasis>
</taxonomicName>
(Latin, adjective) means “hook-like”.
</paragraph>
</subSubSection>
<subSubSection id="C31019E83F5D1E64FF04F9F144C70C1B" box="[151,673,1564,1589]" pageId="12" pageNumber="167" type="distribution">
<paragraph id="8BB54A633F5D1E64FF04F9F144C70C1B" blockId="12.[151,1127,220,1589]" box="[151,673,1564,1589]" pageId="12" pageNumber="167">
<emphasis id="B97E96713F5D1E64FF04F9F1474C0C1B" bold="true" box="[151,298,1564,1588]" pageId="12" pageNumber="167">Distribution.</emphasis>
Known only from the
<typeStatus id="54B1F4C13F5D1E64FD8EF9F0442D0C1A" box="[541,587,1565,1589]" pageId="12" pageNumber="167">type</typeStatus>
locality
</paragraph>
</subSubSection>
</treatment>
</document>
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