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<mods:title id="B2CC8ABDF0AF1B56B17A7CA7CA02617B">Palaemonella shirakawai, a new species of shrimp (Crustacea: Decapoda: Palaemonidae), associated with burrow dwelling opistognathid fish from the Ryukyu Islands, Japan</mods:title>
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<taxonomicName id="4C0A4D2A8845FFEFFF4AFF49A6F643CD" ID-CoL="b2ab8157-d77f-40f1-a9b1-b0a99ebe7e95" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[151,457,151,177]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8845FFEFFF4AFF49A6F643CD" bold="true" box="[151,457,151,177]" italics="true" pageId="1" pageNumber="522">Palaemonella shirakawai</emphasis>
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<figureCitation id="13312A2C8845FFEFFF4AFF64A63F43AF" box="[151,256,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1822,1844]" captionStartId-1="2.[151,250,1908,1930]" captionStartId-2="3.[151,250,1860,1882]" captionStartId-3="4.[151,250,1873,1895]" captionTargetBox-0="[160,1426,1152,1788]" captionTargetBox-1="[171,1420,442,1865]" captionTargetBox-2="[151,1400,198,1814]" captionTargetBox-3="[176,1370,165,1829]" captionTargetId-0="figure@1.[151,1436,1142,1800]" captionTargetId-1="figure@2.[151,1436,409,1886]" captionTargetId-2="figure@3.[151,1436,193,1838]" captionTargetId-3="figure@4.[175,1389,163,1842]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). Entire animal in lateral view." captionText-1="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." captionText-2="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." captionText-3="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri-0="https://zenodo.org/record/837067/files/figure.png" httpUri-1="https://zenodo.org/record/837069/files/figure.png" httpUri-2="https://zenodo.org/record/837071/files/figure.png" httpUri-3="https://zenodo.org/record/837073/files/figure.png" pageId="1" pageNumber="522">Figs. 14</figureCitation>
</paragraph>
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<paragraph id="8BB536A98845FFEFFF4AFEDDA45E422A" blockId="1.[151,1436,258,372]" pageId="1" pageNumber="522">
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<taxonomicName id="4C0A4D2A8845FFEFFF4AFEDDA66E4264" ID-CoL="f378d761-d93b-42a4-bece-669e10a5b69f" authorityName="Kemp" authorityYear="1922" box="[151,337,259,280]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B97EEABB8845FFEFFF4AFEDDA66E4264" box="[151,337,259,280]" italics="true" pageId="1" pageNumber="522">Palaemonella lata</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AAB10B88845FFEFFEB2FEDCA5004264" author="Debelius" box="[367,575,258,280]" page="192" pageId="1" pageNumber="522" year="1999">
<bibRefCitation id="EF9B4B588845FFEFFEB2FEDCA5324264" author="Debelius" box="[367,525,258,280]" pageId="1" pageNumber="522" refString="Debelius, H. (1999) Crustacean guide of the World: shrimps, crabs, lobsters, mantis shrimps, amphipods. Ikan- Unterwasserarchiv, Frankfurt, 321 pp." type="book" year="1999">Debelius, 1999</bibRefCitation>
: 192
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, unnumbered fig. in colour. Not
<taxonomicName id="4C0A4D2A8845FFEFFC57FEDDA3F54264" ID-CoL="f378d761-d93b-42a4-bece-669e10a5b69f" authority="Kemp, 1922" authorityName="Kemp" authorityYear="1922" box="[906,1226,258,280]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B97EEABB8845FFEFFC57FEDDA37B4264" box="[906,1092,259,280]" italics="true" pageId="1" pageNumber="522">Palaemonella lata</emphasis>
<bibRefCitation id="EF9B4B588845FFEFFB96FEDDA3F54264" author="Kemp" box="[1099,1226,258,280]" pageId="1" pageNumber="522" refString="Kemp, S. (1922) Notes on the Crustacea Decapoda in the Indian Museum. Records of the Indian Museum, 24, 113 - 288, pls. 3 - 9." type="journal article" year="1922">Kemp, 1922</bibRefCitation>
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.
<taxonomicName id="4C0A4D2A8845FFEFFF4AFEFFA651424A" ID-CoL="e2da6f38-e1ae-43d3-8fe8-050cbafaa0f9" authorityName="Marin" authorityYear="2008" box="[151,366,289,310]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8845FFEFFF4AFEFFA651424A" box="[151,366,289,310]" italics="true" pageId="1" pageNumber="522">Palaemonella aliska</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AAB10B88845FFEFFE56FEFFA5E2424B" author="Kuiter" box="[395,733,289,311]" page="101" pageId="1" pageNumber="522" year="2009">
<bibRefCitation id="EF9B4B588845FFEFFE56FEFFA59C424B" author="Kuiter" box="[395,675,289,311]" pageId="1" pageNumber="522" refString="Kuiter, R. H. &amp; Debelius, H. (2009) World atlas of marine fauna. Ikan-Unterwasserarchiv, Frankfult, 725 pp." type="book" year="2009">Kuiter &amp; Debelius, 2009</bibRefCitation>
: 101
</treatmentCitation>
, unnumbered figs. in colour;
<treatmentCitation id="0AAB10B88845FFEFFBF2FEFCA2A8424B" author="Humann" box="[1071,1431,289,311]" page="130" pageId="1" pageNumber="522" year="2010">
<bibRefCitation id="EF9B4B588845FFEFFBF2FEFCA261424B" author="Humann" box="[1071,1374,289,311]" pageId="1" pageNumber="522" refString="Humann, P. &amp; DeLoach, N. (2010) Reef creature identification: tropical Pacific. New World Publications, Jacksonville, 497 pp." type="book" year="2010">Humann &amp; DeLoach, 2010</bibRefCitation>
: 130
</treatmentCitation>
, unnumbered fig. in colour. Not
<taxonomicName id="4C0A4D2A8845FFEFFDDAFE9EA462422A" ID-CoL="e2da6f38-e1ae-43d3-8fe8-050cbafaa0f9" authority="Marin, 2008" authorityName="Marin" authorityYear="2008" box="[519,861,320,342]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8845FFEFFDDAFE9EA5E84229" box="[519,727,320,341]" italics="true" pageId="1" pageNumber="522">Palaemonella aliska</emphasis>
<bibRefCitation id="EF9B4B588845FFEFFD01FE9EA462422A" author="Marin" box="[732,861,320,342]" pageId="1" pageNumber="522" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin, 2008</bibRefCitation>
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<emphasis id="B97EEABB8845FFEFFF4AFE81A61B4208" box="[151,292,351,372]" italics="true" pageId="1" pageNumber="522">Palaemonella</emphasis>
aff.
<emphasis id="B97EEABB8845FFEFFE8FFE81A6B14208" box="[338,398,351,372]" italics="true" pageId="1" pageNumber="522">aliska</emphasis>
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.—
<treatmentCitation id="0AAB10B88845FFEFFE71FE80A5404208" author="Minemizu" box="[428,639,350,372]" page="42" pageId="1" pageNumber="522" year="2013">
<bibRefCitation id="EF9B4B588845FFEFFE71FE80A5664208" author="Minemizu" box="[428,601,350,372]" pageId="1" pageNumber="522" refString="Minemizu, R. (2013) Coral reef shrimps of Indo-West Pacific. Bun-ichi Sogo Shuppan, Co. Ltd., Tokyo, 144 pp. [in Japanese]" type="book" year="2013">Minemizu, 2013</bibRefCitation>
: 42
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, unnumbered figs. in colour. Not
<taxonomicName id="4C0A4D2A8845FFEFFC0EFE81A2164208" ID-CoL="e2da6f38-e1ae-43d3-8fe8-050cbafaa0f9" authority="Marin, 2008" authorityName="Marin" authorityYear="2008" box="[979,1321,350,372]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8845FFEFFC0EFE81A39E4208" box="[979,1185,351,372]" italics="true" pageId="1" pageNumber="522">Palaemonella aliska</emphasis>
<bibRefCitation id="EF9B4B588845FFEFFB75FE80A2164208" author="Marin" box="[1192,1321,350,372]" pageId="1" pageNumber="522" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin, 2008</bibRefCitation>
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0 2525, ovigerous female, CL
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</paragraph>
</subSubSection>
<subSubSection id="C31065228845FFEFFF1AFE37A476417E" box="[199,841,489,514]" pageId="1" pageNumber="522" type="biology_ecology">
<paragraph id="8BB536A98845FFEFFF1AFE37A476417E" blockId="1.[151,1437,417,1090]" box="[199,841,489,514]" pageId="1" pageNumber="522">
<emphasis id="B97EEABB8845FFEFFF1AFE37A639417E" bold="true" box="[199,262,489,514]" pageId="1" pageNumber="522">Host.</emphasis>
An unidentified fish belonging to
<taxonomicName id="4C0A4D2A8845FFEFFD54FE37A47A417E" authorityName="Bonaparte" authorityYear="1835" box="[649,837,489,514]" class="Actinopterygii" family="Opistognathidae" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="522" phylum="Chordata" rank="family">Opistognathidae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C31065228845FFEFFF1AFDD3A56C4182" pageId="1" pageNumber="522" type="diagnosis">
<paragraph id="8BB536A98845FFEFFF1AFDD3A56C4182" blockId="1.[151,1437,417,1090]" pageId="1" pageNumber="522">
<emphasis id="B97EEABB8845FFEFFF1AFDD3A67E415A" bold="true" box="[199,321,525,550]" pageId="1" pageNumber="522">Diagnosis.</emphasis>
A medium sized palaemonid shrimp with subcylindrical body. Carapace without supraorbital tooth, postorbital ridge obsolete, hepatic tooth located on normal for genus. Rostrum slender, straight, distinctly shorter than postorbital carapace length, dorsally with 3 postrostral teeth, with 2 of them posterior to level of hepatic tooth, ventrally with a single subapical tooth. Antennal scaphocerite elongate, bearing distolateral tooth reaching level of truncate distal margin of lamella. Second pereiopod with ischium and merus unarmed, carpus distinctly longer than chela, distomesial margin of carpus unarmed, fingers distinctly shorter than palm. Ambulatory pereiopods with propodi not subdivided, dactyli slender.
</paragraph>
</subSubSection>
<subSubSection id="C31065228845FFE8FF1AFCD7A6CD4629" lastPageId="6" lastPageNumber="527" pageId="1" pageNumber="522" type="description">
<paragraph id="8BB536A98845FFEFFF1AFCD7A336473E" blockId="1.[151,1437,417,1090]" pageId="1" pageNumber="522">
<emphasis id="B97EEABB8845FFEFFF1AFCD7A666405E" bold="true" box="[199,345,777,802]" pageId="1" pageNumber="522">Description.</emphasis>
Carapace (
<figureCitation id="13312A2C8845FFEFFE3EFCD7A513405E" box="[483,556,777,802]" captionStart="FIGURE 1" captionStartId="1.[151,250,1822,1844]" captionTargetBox="[160,1426,1152,1788]" captionTargetId="figure@1.[151,1436,1142,1800]" captionTargetPageId="1" captionText="FIGURE 1. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). Entire animal in lateral view." httpUri="https://zenodo.org/record/837067/files/figure.png" pageId="1" pageNumber="522">Fig. 1</figureCitation>
,
<figureCitation id="13312A2C8845FFEFFD9DFCD4A571405E" box="[576,590,778,802]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">2</figureCitation>
A) smooth, glabrous, supraorbital tooth absent; orbit feebly developed, postorbital ridge obsolete, inferior orbital angle strongly produced, angular; antennal tooth well developed, slender, submarginal, arising slightly ventral to orbital angle, overreaching distal margin of inferior orbital angle; hepatic tooth small, situated on normal position, arising ventral to level of antennal tooth; pterygostomial angle obsolete. Rostrum (
<figureCitation id="13312A2C8845FFEFFEDAFC47A67440CE" box="[263,331,921,946]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">Fig. 2</figureCitation>
B) straight, horizontal, reaching distal margin of intermediate segment of antennular peduncle, 0.6 times as long as carapace length; dorsal carina well developed, reaching midlength of carapace, armed with 7 low, acute teeth, including 4 on rostrum proper, anterior-most tooth minute, subequal to apex, thus, tip of rostrum visible as bifid (
<figureCitation id="13312A2C8845FFEFFF23FBDBA67B4762" box="[254,324,1029,1054]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">Fig. 2</figureCitation>
C), 3 postrostral teeth present, 2 of them posterior to level of hepatic tooth (posterior-most tooth damaged); ventral carina obsolete, armed with a single minute subapical tooth.
</paragraph>
<caption id="DF7566218845FFEFFF4AF8C0A2AE4448" box="[151,1425,1822,1844]" httpUri="https://zenodo.org/record/837067/files/figure.png" pageId="1" pageNumber="522" targetBox="[160,1426,1152,1788]" targetPageId="1">
<paragraph id="8BB536A98845FFEFFF4AF8C0A2AE4448" blockId="1.[151,1425,1822,1844]" box="[151,1425,1822,1844]" pageId="1" pageNumber="522">
<emphasis id="B97EEABB8845FFEFFF4AF8C0A62C444F" bold="true" box="[151,275,1822,1844]" pageId="1" pageNumber="522">FIGURE 1.</emphasis>
<taxonomicName id="4C0A4D2A8845FFEFFEC4F8C0A523444F" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[281,540,1822,1843]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="522" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8845FFEFFEC4F8C0A523444F" box="[281,540,1822,1843]" italics="true" pageId="1" pageNumber="522">Palaemonella shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08845FFEFFDFEF8C0A54C444F" box="[547,627,1822,1843]" pageId="1" pageNumber="522" rank="species">
<emphasis id="B97EEABB8845FFEFFDFEF8C0A54C444F" bold="true" box="[547,627,1822,1843]" pageId="1" pageNumber="522">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype ovigerous female (CMNH-ZC 02525). Entire animal in lateral view.
</paragraph>
</caption>
<paragraph id="8BB536A98845FFEFFF1AF885A44C44E4" blockId="1.[151,1436,1883,2016]" pageId="1" pageNumber="522">
Fourth thoracic sternite (
<figureCitation id="13312A2C8845FFEFFE2DF885A5064408" box="[496,569,1883,1908]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">Fig. 2</figureCitation>
D) with finger-like median process; fifth sternite (
<figureCitation id="13312A2C8845FFEFFB4AF885A3DF4408" box="[1175,1248,1883,1908]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">Fig. 2</figureCitation>
D) with pair of semiquadrate lobes posteriorly; sixth to eighth sternites unarmed.
</paragraph>
<paragraph id="8BB536A98845FFECFF1AF87DA61B4261" blockId="1.[151,1436,1883,2016]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="523" pageId="1" pageNumber="522">
Pleonites (
<figureCitation id="13312A2C8845FFEFFEE2F87DA6BD44C0" box="[319,386,1955,1980]" captionStart="FIGURE 1" captionStartId="1.[151,250,1822,1844]" captionTargetBox="[160,1426,1152,1788]" captionTargetId="figure@1.[151,1436,1142,1800]" captionTargetPageId="1" captionText="FIGURE 1. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). Entire animal in lateral view." httpUri="https://zenodo.org/record/837067/files/figure.png" pageId="1" pageNumber="522">Fig. 1</figureCitation>
) smooth, covers glabrous; first to fourth pleura broad, with margins rounded, fifth pleuron posteriorly produced, angular (
<figureCitation id="13312A2C8845FFEFFDDDF819A576449C" box="[512,585,1991,2016]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">Fig. 2</figureCitation>
E); sixth pleonite (
<figureCitation id="13312A2C8845FFEFFCF5F819A44F449C" box="[808,880,1991,2016]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="1" pageNumber="522">Fig. 2</figureCitation>
E) about half length of carapace, posterolateral process acute, posteroventral margin produced, angular. Telson (
<figureCitation id="13312A2C8846FFECFC5CFF49A4F843CC" box="[897,967,151,176]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="2" pageNumber="523">Fig. 2</figureCitation>
F) slightly tapering posteriorly, posterior margin (
<figureCitation id="13312A2C8846FFECFF24FF62A60043A8" box="[249,319,188,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="2" pageNumber="523">Fig. 2</figureCitation>
G) convex, rounded, with 3 pairs of spines (lateral and intermediate spines simple, intermediate spines longest, submedian spines plumose); 2 pairs of subequal dorsolateral spines at 0.4 and 0.7 of telson length respectively.
</paragraph>
<paragraph id="8BB536A98846FFECFF1AFEF9A4DE4219" blockId="2.[151,1437,151,357]" pageId="2" pageNumber="523">
Ophthalmic somite without interocular process. Eye (
<figureCitation id="13312A2C8846FFECFCFEFEF9A458423C" box="[803,871,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="2" pageNumber="523">Fig. 2</figureCitation>
A) with large, globular cornea, accessory pigment spot obscure; stalk short, subequal to maximum corneal diameter in length.
</paragraph>
<caption id="DF7566218846FFECFF4AF8AAA376449A" httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="2" pageNumber="523" targetBox="[171,1420,442,1865]" targetPageId="2">
<paragraph id="8BB536A98846FFECFF4AF8AAA376449A" blockId="2.[151,1436,1908,2022]" pageId="2" pageNumber="523">
<emphasis id="B97EEABB8846FFECFF4AF8AAA62944F5" bold="true" box="[151,278,1908,1930]" pageId="2" pageNumber="523">FIGURE 2.</emphasis>
<taxonomicName id="4C0A4D2A8846FFECFEFDF8AAA51844F5" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[288,551,1908,1929]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="523" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8846FFECFEFDF8AAA51844F5" box="[288,551,1908,1929]" italics="true" pageId="2" pageNumber="523">Palaemonella shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08846FFECFDECF8AAA5BA44F5" box="[561,645,1908,1929]" pageId="2" pageNumber="523" rank="species">
<emphasis id="B97EEABB8846FFECFDECF8AAA5BA44F5" bold="true" box="[561,645,1908,1929]" pageId="2" pageNumber="523">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view.
</paragraph>
</caption>
<caption id="DF7566218847FFEDFF4AF89AA4F744C9" httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="3" pageNumber="524" targetBox="[151,1400,198,1814]" targetPageId="3">
<paragraph id="8BB536A98847FFEDFF4AF89AA4F744C9" blockId="3.[151,1436,1860,1973]" pageId="3" pageNumber="524">
<emphasis id="B97EEABB8847FFEDFF4AF89AA6294425" bold="true" box="[151,278,1860,1882]" pageId="3" pageNumber="524">FIGURE 3.</emphasis>
<taxonomicName id="4C0A4D2A8847FFEDFEC2F89AA5194425" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[287,550,1860,1881]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="524" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8847FFEDFEC2F89AA5194425" box="[287,550,1860,1881]" italics="true" pageId="3" pageNumber="524">Palaemonella shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08847FFEDFDEDF89AA5BC4425" box="[560,643,1860,1881]" pageId="3" pageNumber="524" rank="species">
<emphasis id="B97EEABB8847FFEDFDEDF89AA5BC4425" bold="true" box="[560,643,1860,1881]" pageId="3" pageNumber="524">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view.
</paragraph>
</caption>
<caption id="DF7566218840FFEAFF4AF88FA5F5449E" httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="4" pageNumber="525" targetBox="[176,1370,165,1829]" targetPageId="4">
<paragraph id="8BB536A98840FFEAFF4AF88FA5F5449E" blockId="4.[151,1435,1873,2018]" pageId="4" pageNumber="525">
<emphasis id="B97EEABB8840FFEAFF4AF88FA62D441A" bold="true" box="[151,274,1873,1895]" pageId="4" pageNumber="525">FIGURE 4.</emphasis>
<taxonomicName id="4C0A4D2A8840FFEAFEC5F88CA524441B" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[280,539,1874,1895]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="525" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8840FFEAFEC5F88CA524441B" box="[280,539,1874,1895]" italics="true" pageId="4" pageNumber="525">Palaemonella shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08840FFEAFDFCF88CA54E441B" box="[545,625,1874,1895]" pageId="4" pageNumber="525" rank="species">
<emphasis id="B97EEABB8840FFEAFDFCF88CA54E441B" bold="true" box="[545,625,1874,1895]" pageId="4" pageNumber="525">sp. nov.</emphasis>
</taxonomicNameLabel>
Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view.
</paragraph>
</caption>
<paragraph id="8BB536A98841FFEBFF1AFF49A6B74289" blockId="5.[151,1437,151,1516]" pageId="5" pageNumber="526">
Antennular peduncle (
<figureCitation id="13312A2C8841FFEBFE1EFF49A53A43CC" box="[451,517,151,176]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
A, B) with proximal segment slightly longer than distal two segments combined; distolateral margin roundly produced, reaching level of proximal third of intermediate segment, with row of long setae, lateral margin almost straight, terminating distally in acute tooth falling slightly short of level of midlength of intermediate segment; ventromesial margin armed with small acute tooth; stylocerite slender, acute, reaching level of proximal third of proximal segment, with sparse setae laterally; statocyst well developed, oblong; intermediate segment slender, feebly lobed laterally, distal and mesial margin furnished with sparse long setae; distal segment subequal to intermediate segment in length, slender, non-setose. Upper flagellum (
<figureCitation id="13312A2C8841FFEBFB25FEB1A20442F4" box="[1272,1339,367,392]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
A) long, slender, somewhat compressed dorsoventrally, biramous, proximal 16 segments fused, shorter free ramus consisting of 5 segments; about 10 groups of long aesthetascs present; lower flagellum considerably more slender than upper flagellum.
</paragraph>
<paragraph id="8BB536A98841FFEBFF1AFDDEA4A241F9" blockId="5.[151,1437,151,1516]" pageId="5" pageNumber="526">
Antenna (
<figureCitation id="13312A2C8841FFEBFEE4FE21A6404164" box="[313,383,511,536]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
C) with stout basicerite armed ventrolaterally with acute tooth; carpocerite stout, reaching proximal two fifths of scaphocerite; scaphocerite elongate, overreaching distal margin of antennular peduncle, lateral and mesial margins subparallel, 4.4 times as long as maximum width, lateral margin concave, terminating in strong acute tooth reaching level of truncate distal margin of lamella.
</paragraph>
<paragraph id="8BB536A98841FFEBFF1AFD51A41E41D4" blockId="5.[151,1437,151,1516]" box="[199,801,655,680]" pageId="5" pageNumber="526">Epistome with low, rounded submedian prominences.</paragraph>
<paragraph id="8BB536A98841FFEBFF1AFD6AA6BB46BC" blockId="5.[151,1437,151,1516]" pageId="5" pageNumber="526">
Mandible (
<figureCitation id="13312A2C8841FFEBFE9CFD6AA6BC41B0" box="[321,387,692,717]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
D, E) robust, with slender, 2-segmented palp, these segments subequal in length, distally with sparse long setae; molar process obliquely truncated distally, with large, blunt teeth; incisor process stout, tapering distally, with 3 acute distal teeth. Maxillule (
<figureCitation id="13312A2C8841FFEBFD7BFD22A5D44068" box="[678,747,764,789]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
F) with strongly bilobed palp, lower lobe with small distal protuberance; upper lacinia narrow, curved, distal margin truncated, with 7 stout spines and sparse spiniform setae; lower lacinia tapering distally, with some serrulate setae distally. Maxilla (
<figureCitation id="13312A2C8841FFEBFC3CFC9AA31C4020" box="[993,1059,836,861]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
G) with short, stout palp tapering distally; distal endite well developed, narrow, deeply bilobed, distal lobe slightly broader than proximal lobe, with simple setae distally; proximal endite obsolete, mesial margin feebly sinuous; scaphognathite well developed, posterior lobe short, distal half of anterior lobe narrow. First maxilliped (
<figureCitation id="13312A2C8841FFEBFC05FC71A32440B4" box="[984,1051,943,968]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
H) with short simple palp distally narrow, without setae; distal endite subquadrate distally, with mesial margin bearing 2 rows of dense setae, subquadrate distally; proximal endite small, rounded, separated from distal endite by distinct notch; caridean lobe small; epipod large, feebly bilobed, anteriorly pointed, falling shightly short of distal margin of caridean lobe; exopod with well-developed flagellum distally with numerous long setae. Second maxilliped (
<figureCitation id="13312A2C8841FFEBFB65FBE1A3C64724" box="[1208,1273,1087,1112]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">
Fig.
<date id="FFB410698841FFEBFB31FB9EA3C64724" box="[1260,1273,1088,1112]" pageId="5" pageNumber="526">3</date>
</figureCitation>
I) with normal endopod; coxa inflated mesially, epipod angular, with unilamellar podobranch, shaft stout; ischium and basis completely fused; merus stout, obliquely articulated with ischiobasis; carpus with distinct ventromesial process; propodus with anterior margin broadly rounded, falling short of level of mesial dactylar margin, with slender spines; dactylus broad, mesial margin with numerous setae; exopod with well developed flagellum, distally with numerous slender setae. Third maxilliped (
<figureCitation id="13312A2C8841FFEBFD4AFB2AA5E14670" box="[663,734,1268,1293]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="5" pageNumber="526">Fig. 3</figureCitation>
J) with endopod slender, reaching level of distal fourth of scaphocerite; coxal plate oval, multilamellar arthrobranch present; ischiomerus and basis demarcated, external margin armed with row of small spines distally, mesial margin furnished with long sparse setae; penultimate segment 1.3 times as long as ultimate segment, mesially with long dense setae; ultimate segment tapering distally, external surface with 7 transverse rows of short dense setae; exopod with well developed flagellum, distally with numerous long setae.
</paragraph>
<paragraph id="8BB536A98841FFEBFF1AFA0DA55A4690" blockId="5.[151,1437,151,1516]" box="[199,613,1491,1516]" pageId="5" pageNumber="526">Branchial formula as in Table below.</paragraph>
<paragraph id="8BB536A98841FFEBFDF3F9D9A3AF4561" blockId="5.[159,1369,1543,1813]" box="[558,1168,1543,1565]" pageId="5" pageNumber="526">Maxillipeds Pereiopods</paragraph>
<paragraph id="8BB536A98841FFEBFDD3F9EFA2664469" blockId="5.[159,1369,1543,1813]" pageId="5" pageNumber="526">I II III I II III IV V Pleurobranchs — — — 1 1 1 1 1 Arthrobranchs — — 1 — — — — — Podobranchs — 1 — — — — — — Epipods 1 1 — — — — — — Exopods 1 1 1 — — — — —</paragraph>
<paragraph id="8BB536A98841FFE8FF1AF8E6A5084260" blockId="5.[151,1436,1848,2017]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="527" pageId="5" pageNumber="526">
First pereiopod (
<figureCitation id="13312A2C8841FFEBFE5FF8E6A6FA442C" box="[386,453,1848,1873]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="5" pageNumber="526">Fig. 4</figureCitation>
A) moderately slender, overreaching distal margin of scaphocerite by lengths of carpus and chela. Coxa with ventral margin sparsely setose, without medial process. Ischium and merus furnished with long sparse setae ventrally, merus 1.7 times as long as ischium. Carpus considerably elongate, 1.8 times as long as ischium, 1.2 times as long as merus, 1.8 times as long as chela, with longitudinal row of serrulate grooming setae distally. Chela (
<figureCitation id="13312A2C8841FFEBFE97F816A6B0449C" box="[330,399,1992,2017]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="5" pageNumber="526">Fig. 4</figureCitation>
B) about half of carapace length; palm subcylindrical, slightly compressed, slightly shorter than fingers, with 7 transverse rows of short serrulate grooming setae proximoventrally; dactylus slender, terminating in small, hooked tip, cutting edge situated laterally, edentate, with sparse short setae; fixed finger terminating in small, hooked tip, cutting edge situated laterally, edentate, with sparse short setae, ventral surface with 5 transverse rows of short setae.
</paragraph>
<paragraph id="8BB536A98842FFE8FF1AFEF9A26C411C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="527">
Second pereiopods (
<figureCitation id="13312A2C8842FFE8FE6DFEF9A6C8423C" box="[432,503,295,320]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
C) well developed, similar in size and form, considerably elongate, overreaching distal margin of scaphocerite by lengths of distal two fifths of merus, carpus and chela. Ischium and merus slender, unarmed, furnished with long sparse setae ventrally, merus 1.3 times as long as ischium. Carpus long, without distal lobe (
<figureCitation id="13312A2C8842FFE8FEC7FE4AA66242D0" box="[282,349,404,429]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
D), furnished uniformly with short sparse setae, 2.3 times as long as ischium, 1.7 times as long as merus, slightly longer than length of chela. Chela slightly bowed, 1.7 times as long as carapace length; palm 1.7 times as long as dactylus, slightly compressed; dactylus (
<figureCitation id="13312A2C8842FFE8FCFEFE02A4584288" box="[803,871,476,501]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
E) terminating in hooked, acutely pointed unguis, cutting edge situated laterally, armed proximally with 3 large subtriangular teeth, remaining part unarmed, sharply edged, sparsely with equidistant short setae; fixed finger (
<figureCitation id="13312A2C8842FFE8FCC7FDFAA4634140" box="[794,860,548,573]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
E) generally similar to dactylus, cutting edge armed proximally with 2 large subtriangular teeth opposite to notch among proximal teeth of dactylar cutting edge.
</paragraph>
<paragraph id="8BB536A98842FFE8FF1AFDB2A55340FC" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="527">
Third pereiopod (
<figureCitation id="13312A2C8842FFE8FE51FDB2A6F141F8" box="[396,462,620,645]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
F) slender, overreaching distal margin of scaphocerite by lengths of distal three fourths of carpus, propodus and dactylus. Merus 1.3 times as long as carpus, unarmed. Carpus elongate, unarmed. Propodus (
<figureCitation id="13312A2C8842FFE8FED2FD6AA66E41B0" box="[271,337,692,717]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
G) 1.2 times as long as carpus, with 2 distoventral spines and 8 single spines along ventral surface, dorsal surface with few short setae. Dactylus (
<figureCitation id="13312A2C8842FFE8FD11FD09A42A418C" box="[716,789,727,752]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
H) simple, compressed, feebly sinuous, unguis clearly demarcated. Fourth pereiopod similar to third, overreaching distal margin of scaphocerite by lengths of distal fourth of carpus, propodus and dactylus. Fifth pereiopod overreaching distal margin of scaphocerite by lengths of propodus and dactylus, with propodus (
<figureCitation id="13312A2C8842FFE8FD92FC9AA5AD4020" box="[591,658,836,861]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">
Fig.
<date id="FFB410698842FFE8FD5EFC9AA5AD4020" box="[643,658,836,860]" pageId="6" pageNumber="527">4</date>
</figureCitation>
I) bearing 1 spine and tufts of setae distoventrally (
<figureCitation id="13312A2C8842FFE8FB1BFC9AA2354020" box="[1222,1290,836,861]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="6" pageNumber="527">Fig. 4</figureCitation>
J), spaced set of 7 spines posterior to distoventral spine.
</paragraph>
<paragraph id="8BB536A98842FFE8FF1AFC52A2174091" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="527">
Uropod (
<figureCitation id="13312A2C8842FFE8FEF1FC52A64F40D8" box="[300,368,908,933]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="6" pageNumber="527">Fig. 2</figureCitation>
F) with protopodite posterolaterally produced; exopod broad, overreaching posterior margin of telson, broadly rounded distally, lateral margin nearly straight, terminating in small acute tooth, with larger mobile spine just mesial to distolateral tooth; endopod oval, falling slightly short of posterior margin of exopod.
</paragraph>
<paragraph id="8BB536A98842FFE8FF1AFC29A6CD4629" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="527">
<emphasis id="B97EEABB8842FFE8FF1AFC29A64D476C" bold="true" box="[199,370,1015,1040]" pageId="6" pageNumber="527">Colour in life.</emphasis>
Carapace and rostrum whitish transparent, lateral surface of carapace covered sparsely with some yellow spots. Pleonites with some yellow spots sparsely, midline of sternites brownish purpule, visible as longitudinal stripe in dorsal view; telson proximally brownish purple, distally colorless. Eyestalk with longitudinal brownish purple stripe, cornea silver. Antennular peduncle and upper flagellum brownish purple. Scaphocerite and antennal flagellum colorless. Ultimate segment of third maxilliped brownish purple, penultimate and antepenultimate segments colorless, both with yellow distal patches. First pereiopod brownish purple, distal part of merus colorless. Second pereiopod uniformly brownish purple. Ambulatory pereiopods with ischial-meral joint yellow; meri generally brownish purple, distal part white; proximal two thirds of carpi brownish purple, distal third white; propodi white, with scattered brownish purple chromatophores. (From the underwater photograph taken before capturing the
<typeStatus id="54B1880B8842FFE8FEA2FAE2A6D94629" box="[383,486,1340,1365]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
).
</paragraph>
</subSubSection>
<subSubSection id="C31065228842FFE8FF1AFA81A4AF46E0" pageId="6" pageNumber="527" type="distribution">
<paragraph id="8BB536A98842FFE8FF1AFA81A4AF46E0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="527">
<emphasis id="B97EEABB8842FFE8FF1AFA81A65F4604" bold="true" box="[199,352,1375,1400]" pageId="6" pageNumber="527">Distribution.</emphasis>
Known from the Ryukyu Islands, southern
<collectingCountry id="F31D76398842FFE8FC90FABFA4B24604" box="[845,909,1377,1400]" name="Japan" pageId="6" pageNumber="527">Japan</collectingCountry>
(present study),
<collectingCountry id="F31D76398842FFE8FB97FA81A3F74604" box="[1098,1224,1375,1400]" name="Philippines" pageId="6" pageNumber="527">Philippines</collectingCountry>
(
<bibRefCitation id="EF9B4B588842FFE8FB0AFA81A2AF4604" author="Minemizu" box="[1239,1424,1375,1400]" pageId="6" pageNumber="527" refString="Minemizu, R. (2013) Coral reef shrimps of Indo-West Pacific. Bun-ichi Sogo Shuppan, Co. Ltd., Tokyo, 144 pp. [in Japanese]" type="book" year="2013">Minemizu 2013</bibRefCitation>
),
<collectingCountry id="F31D76398842FFE8FF4AFA5AA7C446E1" box="[151,251,1412,1437]" name="Thailand" pageId="6" pageNumber="527">Thailand</collectingCountry>
(
<bibRefCitation id="EF9B4B588842FFE8FED4FA5AA68E46E0" author="Debelius" box="[265,433,1412,1437]" pageId="6" pageNumber="527" refString="Debelius, H. (1999) Crustacean guide of the World: shrimps, crabs, lobsters, mantis shrimps, amphipods. Ikan- Unterwasserarchiv, Frankfurt, 321 pp." type="book" year="1999">Debelius 1999</bibRefCitation>
), and
<collectingCountry id="F31D76398842FFE8FE29FA5AA55E46E1" box="[500,609,1412,1437]" name="Indonesia" pageId="6" pageNumber="527">Indonesia</collectingCountry>
(
<bibRefCitation id="EF9B4B588842FFE8FDB2FA5AA4BB46E0" author="Kuiter" box="[623,900,1412,1437]" pageId="6" pageNumber="527" refString="Kuiter, R. H. &amp; Debelius, H. (2009) World atlas of marine fauna. Ikan-Unterwasserarchiv, Frankfult, 725 pp." type="book" year="2009">Kuiter &amp; Debelius 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C31065228842FFE8FF1AFA79A48F4698" pageId="6" pageNumber="527" type="etymology">
<paragraph id="8BB536A98842FFE8FF1AFA79A32946BC" blockId="6.[151,1437,151,2013]" box="[199,1046,1447,1472]" pageId="6" pageNumber="527">
<emphasis id="B97EEABB8842FFE8FF1AFA79A67146BC" bold="true" box="[199,334,1447,1472]" pageId="6" pageNumber="527">Etymology.</emphasis>
Named for Mr. Naoki Shirakawa, the collector of the
<typeStatus id="54B1880B8842FFE8FC71FA79A32D46BC" box="[940,1042,1447,1472]" pageId="6" pageNumber="527" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="8BB536A98842FFE8FF1AFA12A48F4698" blockId="6.[151,1437,151,2013]" box="[199,944,1484,1509]" pageId="6" pageNumber="527">
<emphasis id="B97EEABB8842FFE8FF1AFA12A6BA4699" bold="true" box="[199,389,1484,1509]" pageId="6" pageNumber="527">Common name.</emphasis>
Agito-kakure-ebi (new standard Japanese name).
</paragraph>
</subSubSection>
<subSubSection id="C31065228842FFE8FF1AFA31A2A244A0" pageId="6" pageNumber="527" type="discussion">
<paragraph id="8BB536A98842FFE8FF1AFA31A2A244A0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="527">
<emphasis id="B97EEABB8842FFE8FF1AFA31A6044574" bold="true" box="[199,315,1519,1544]" pageId="6" pageNumber="527">Remarks.</emphasis>
Among the known species of
<taxonomicName id="4C0A4D2A8842FFE8FD7DFA2EA4034574" authorityName="Dana" authorityYear="1852" box="[672,828,1520,1544]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEABB8842FFE8FD7DFA2EA4034574" box="[672,828,1520,1544]" italics="true" pageId="6" pageNumber="527">Palaemonella</emphasis>
</taxonomicName>
, the present new species appears closest to
<taxonomicName id="4C0A4D2A8842FFE8FAE6FA2FA2A34574" authorityName="Marin" authorityYear="2008" box="[1339,1436,1520,1544]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8842FFE8FAE6FA2FA2A34574" box="[1339,1436,1520,1544]" italics="true" pageId="6" pageNumber="527">P. aliska</emphasis>
</taxonomicName>
living inside of the holes burrowed by snapping shrimp
<taxonomicName id="4C0A4D2A8842FFE8FCD9F9CAA4614550" box="[772,862,1556,1580]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEABB8842FFE8FCD9F9CAA4614550" box="[772,862,1556,1580]" italics="true" pageId="6" pageNumber="527">Alpheus</emphasis>
</taxonomicName>
cf.
<emphasis id="B97EEABB8842FFE8FC55F9CAA4DE4550" box="[904,993,1556,1580]" italics="true" pageId="6" pageNumber="527">bellulus</emphasis>
Miya &amp; Miyake,
<quantity id="4CF29B4C8842FFE8FB75F9CAA3C24551" box="[1192,1277,1556,1581]" metricMagnitude="1" metricUnit="m" metricValue="5.00126" pageId="6" pageNumber="527" unit="in" value="1969.0">1969 in</quantity>
the mangrove littoral zone of Nhatrang Bay,
<collectingCountry id="F31D76398842FFE8FE37F9E9A576452C" box="[490,585,1591,1616]" name="Vietnam" pageId="6" pageNumber="527">Vietnam</collectingCountry>
(
<bibRefCitation id="EF9B4B588842FFE8FD8AF9E9A5DF452C" author="Marin" box="[599,736,1591,1616]" pageId="6" pageNumber="527" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
), but presumably more widespread. In the original description,
<bibRefCitation id="EF9B4B588842FFE8FF4AF982A6114508" author="Marin" box="[151,302,1628,1653]" pageId="6" pageNumber="527" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin (2008)</bibRefCitation>
indicated that the opistognathid-associated shrimp reported as
<taxonomicName id="4C0A4D2A8842FFE8FC29F983A3034508" authorityName="Kemp" authorityYear="1922" box="[1012,1084,1628,1652]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="lata">
<emphasis id="B97EEABB8842FFE8FC29F983A3034508" box="[1012,1084,1628,1652]" italics="true" pageId="6" pageNumber="527">P. lata</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF9B4B588842FFE8FBBAF982A2224508" author="Debelius" box="[1127,1309,1628,1653]" pageId="6" pageNumber="527" refString="Debelius, H. (1999) Crustacean guide of the World: shrimps, crabs, lobsters, mantis shrimps, amphipods. Ikan- Unterwasserarchiv, Frankfurt, 321 pp." type="book" year="1999">Debelius (1999)</bibRefCitation>
is possibly also with
<taxonomicName id="4C0A4D2A8842FFE8FEDBF95FA65C45E4" authorityName="Marin" authorityYear="2008" box="[262,355,1664,1688]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8842FFE8FEDBF95FA65C45E4" box="[262,355,1664,1688]" italics="true" pageId="6" pageNumber="527">P. aliska</emphasis>
</taxonomicName>
, but herein referred to the new species. Morphologically, the present new species and
<taxonomicName id="4C0A4D2A8842FFE8FAE1F95FA2A345E4" authorityName="Marin" authorityYear="2008" box="[1340,1436,1664,1688]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8842FFE8FAE1F95FA2A345E4" box="[1340,1436,1664,1688]" italics="true" pageId="6" pageNumber="527">P. aliska</emphasis>
</taxonomicName>
share the post-rostral carina extending to the midpoint of the carapace and armed with teeth posterior to level of the hepatic tooth (
<figureCitation id="13312A2C8842FFE8FE99F919A6B2459C" box="[324,397,1735,1760]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="6" pageNumber="527">Fig. 2</figureCitation>
A), and the unarmed ishium, merus and carpus of the second pereiopod.
<taxonomicName id="4C0A4D2A8842FFE8FADDF916A62B4478" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8842FFE8FADDF916A62B4478" italics="true" pageId="6" pageNumber="527">Palaemonella shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08842FFE8FEC6F932A64A4479" box="[283,373,1772,1797]" pageId="6" pageNumber="527" rank="species">
<emphasis id="B97EEABB8842FFE8FEC6F932A64A4479" bold="true" box="[283,373,1772,1797]" pageId="6" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
is, however, readily distinguishable from
<taxonomicName id="4C0A4D2A8842FFE8FC8CF933A4914478" authorityName="Marin" authorityYear="2008" box="[849,942,1772,1796]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8842FFE8FC8CF933A4914478" box="[849,942,1772,1796]" italics="true" pageId="6" pageNumber="527">P. aliska</emphasis>
</taxonomicName>
on account of the following morphological characters: (1) a single, minute subterminal tooth is present on the ventral margin of the rostrum in
<taxonomicName id="4C0A4D2A8842FFE8FADEF8CFA2A34454" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[1283,1436,1808,1832]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8842FFE8FADEF8CFA2A34454" box="[1283,1436,1808,1832]" italics="true" pageId="6" pageNumber="527">P. shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08842FFE8FF4AF8EAA7CD4431" box="[151,242,1844,1869]" pageId="6" pageNumber="527" rank="species">
<emphasis id="B97EEABB8842FFE8FF4AF8EAA7CD4431" bold="true" box="[151,242,1844,1869]" pageId="6" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13312A2C8842FFE8FEDEF8EAA6774430" box="[259,328,1844,1869]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="6" pageNumber="527">Fig. 2</figureCitation>
B, C), instead of 23 acuminate rostral ventral teeth in
<taxonomicName id="4C0A4D2A8842FFE8FC1CF8EBA3224430" authorityName="Marin" authorityYear="2008" box="[961,1053,1844,1868]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8842FFE8FC1CF8EBA3224430" box="[961,1053,1844,1868]" italics="true" pageId="6" pageNumber="527">P. aliska</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9B4B588842FFE8FBECF8EAA3824430" author="Marin" box="[1073,1213,1844,1869]" pageId="6" pageNumber="527" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008">Marin 2008</bibRefCitation>
:
<figureCitation id="13312A2C8842FFE8FB14F8EAA2324430" box="[1225,1293,1844,1869]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="6" pageNumber="527">Fig. 2</figureCitation>
a, b); (2) the posterior margin of the telson is rounded in
<taxonomicName id="4C0A4D2A8842FFE8FD45F887A40C440C" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[664,819,1880,1904]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8842FFE8FD45F887A40C440C" box="[664,819,1880,1904]" italics="true" pageId="6" pageNumber="527">P. shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08842FFE8FCE1F889A4A6440C" box="[828,921,1879,1904]" pageId="6" pageNumber="527" rank="species">
<emphasis id="B97EEABB8842FFE8FCE1F889A4A6440C" bold="true" box="[828,921,1879,1904]" pageId="6" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13312A2C8842FFE8FC76F889A4CE440C" box="[939,1009,1879,1904]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="6" pageNumber="527">Fig. 2</figureCitation>
G), whereas its midpoint is distinctly protruding in
<taxonomicName id="4C0A4D2A8842FFE8FEE7F8A3A6A244E8" authorityName="Marin" authorityYear="2008" box="[314,413,1916,1940]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8842FFE8FEE7F8A3A6A244E8" box="[314,413,1916,1940]" italics="true" pageId="6" pageNumber="527">P. aliska</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9B4B588842FFE8FE72F8A2A5A044E9" author="Marin" box="[431,671,1916,1941]" pageId="6" pageNumber="527" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008" yearSuffix="f">Marin 2008: Fig. 5f</bibRefCitation>
); (3)
<taxonomicName id="4C0A4D2A8842FFE8FD39F8A2A33744E8" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[740,1032,1916,1940]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="527" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8842FFE8FD39F8A2A33744E8" box="[740,1032,1916,1940]" italics="true" pageId="6" pageNumber="527">Palaemonella shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08842FFE8FBCEF8A2A34E44E9" box="[1043,1137,1916,1941]" pageId="6" pageNumber="527" rank="species">
<emphasis id="B97EEABB8842FFE8FBCEF8A2A34E44E9" bold="true" box="[1043,1137,1916,1941]" pageId="6" pageNumber="527">sp. nov.</emphasis>
</taxonomicNameLabel>
has an elongate antennal scaphocerite with the subparallel lateral and mesial margins, its length is 4.4 times as long as the maximum width, and the distolateral tooth reaching the level of the truncate distal margin of lamella (
<figureCitation id="13312A2C8842FFE8FBAFF81AA38844A0" box="[1138,1207,1988,2013]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="6" pageNumber="527">Fig. 3</figureCitation>
C), in contrast in
<emphasis id="B97EEABB8842FFE8FA5AF81BA2A244A0" box="[1415,1437,1989,2012]" italics="true" pageId="6" pageNumber="527">P.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31065228843FFE9FF4AFF46A5E34289" pageId="7" pageNumber="528" type="reference_group">
<paragraph id="8BB536A98843FFE9FF4AFF46A2644218" blockId="7.[151,1437,151,501]" pageId="7" pageNumber="528">
<taxonomicName id="4C0A4D2A8843FFE9FF4AFF46A7E543CC" ID-CoL="e2da6f38-e1ae-43d3-8fe8-050cbafaa0f9" authorityName="Marin" authorityYear="2008" box="[151,218,152,176]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8843FFE9FF4AFF46A7E543CC" box="[151,218,152,176]" italics="true" pageId="7" pageNumber="528">aliska</emphasis>
</taxonomicName>
, the scaphocerite is relatively shorter, being about 3 times as long as the maximum width, and the distolateral tooth overreaches the rounded distal margin of lamella (
<bibRefCitation id="EF9B4B588843FFE9FCD6FF62A4D143A9" author="Marin" box="[779,1006,188,213]" pageId="7" pageNumber="528" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008" yearSuffix="f">
Marin 2008:
<figureCitation id="13312A2C8843FFE9FC7DFF62A4DD43A9" box="[928,994,188,213]" captionStart="FIGURE 2" captionStartId="2.[151,250,1908,1930]" captionTargetBox="[171,1420,442,1865]" captionTargetId="figure@2.[151,1436,409,1886]" captionTargetPageId="2" captionText="FIGURE 2. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, carapace, rostrum and cephalic appendages, dorsal view; B, carapace and rostrum, lateral view; C, tip of rostrum, lateral view; D, fourth to sixth thoracic sternites, ventral view; E, posterior part of fifth pleuron, sixth pleonite, proximal part of telson and uropodal basipodite, lateral view; F, telson and right uropods, dorsal view; G, tip of telson, dorsal view." httpUri="https://zenodo.org/record/837069/files/figure.png" pageId="7" pageNumber="528">Fig. 2</figureCitation>
f
</bibRefCitation>
); (4) the epipod of the first maxilliped of
<taxonomicName id="4C0A4D2A8843FFE9FF68FF3FA6714384" ID-CoL="b2ab8157-d77f-40f1-a9b1-b0a99ebe7e95" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[181,334,224,248]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8843FFE9FF68FF3FA6714384" box="[181,334,224,248]" italics="true" pageId="7" pageNumber="528">P. shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08843FFE9FE88FF01A6904384" box="[341,431,223,248]" pageId="7" pageNumber="528" rank="species">
<emphasis id="B97EEABB8843FFE9FE88FF01A6904384" bold="true" box="[341,431,223,248]" pageId="7" pageNumber="528">sp. nov.</emphasis>
</taxonomicNameLabel>
is long, with the anterior margin falling slightly short of the distal margin of the caridean lobe (
<figureCitation id="13312A2C8843FFE9FF04FEDAA6234260" box="[217,284,260,285]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="7" pageNumber="528">Fig. 3</figureCitation>
H), whereas the epipod in
<taxonomicName id="4C0A4D2A8843FFE9FD8DFEDBA5914260" ID-CoL="e2da6f38-e1ae-43d3-8fe8-050cbafaa0f9" authorityName="Marin" authorityYear="2008" box="[592,686,260,284]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8843FFE9FD8DFEDBA5914260" box="[592,686,260,284]" italics="true" pageId="7" pageNumber="528">P. aliska</emphasis>
</taxonomicName>
is short, and its anterior margin slightly overreaches level of the distal margin of the proximal endite (
<bibRefCitation id="EF9B4B588843FFE9FDE1FEF9A41C423C" author="Marin" box="[572,803,295,320]" pageId="7" pageNumber="528" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008" yearSuffix="e">
Marin 2008:
<figureCitation id="13312A2C8843FFE9FD0CFEF9A42B423C" box="[721,788,295,320]" captionStart="FIGURE 3" captionStartId="3.[151,250,1860,1882]" captionTargetBox="[151,1400,198,1814]" captionTargetId="figure@3.[151,1436,193,1838]" captionTargetPageId="3" captionText="FIGURE 3. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right antennule, dorsal view; B, right antennular peduncle, dorsal view; C, right antenna, dorsal view; D, right mandible, external view; E, same, distal part, mesial view; F, right first maxillule, external view; G, right maxilla, external view; H, right first maxilliped, external view; I, right second maxilliped, external view; J, right third maxilliped, external view." httpUri="https://zenodo.org/record/837071/files/figure.png" pageId="7" pageNumber="528">Fig. 3</figureCitation>
e
</bibRefCitation>
); and (5) the carpus of second pereiopod furnished with dense setae in
<taxonomicName id="4C0A4D2A8843FFE9FEE4FE93A6EE4218" ID-CoL="b2ab8157-d77f-40f1-a9b1-b0a99ebe7e95" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[313,465,332,356]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="shirakawai" status="sp. nov.">
<emphasis id="B97EEABB8843FFE9FEE4FE93A6EE4218" box="[313,465,332,356]" italics="true" pageId="7" pageNumber="528">P. shirakawai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08843FFE9FE05FE92A50E4219" box="[472,561,332,357]" pageId="7" pageNumber="528" rank="species">
<emphasis id="B97EEABB8843FFE9FE05FE92A50E4219" bold="true" box="[472,561,332,357]" pageId="7" pageNumber="528">sp. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="13312A2C8843FFE9FD9CFE92A5BB4218" box="[577,644,332,357]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="7" pageNumber="528">Fig. 4</figureCitation>
C), while the carpus is smooth in
<taxonomicName id="4C0A4D2A8843FFE9FC21FE93A3654218" ID-CoL="e2da6f38-e1ae-43d3-8fe8-050cbafaa0f9" authorityName="Marin" authorityYear="2008" box="[1020,1114,332,356]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="aliska">
<emphasis id="B97EEABB8843FFE9FC21FE93A3654218" box="[1020,1114,332,356]" italics="true" pageId="7" pageNumber="528">P. aliska</emphasis>
</taxonomicName>
(
<bibRefCitation id="EF9B4B588843FFE9FBB5FE92A2714218" author="Marin" box="[1128,1358,332,357]" pageId="7" pageNumber="528" refString="Marin, I. N. (2008) Description of two new species from the genera Palaemonella Dana, 1852 and Uir Holthuis, 1952 (Crustacea: Caridea: Palaemonidae: Pontoniinae). Zoologische Mededelingen, 82, 375 - 390." type="journal article" year="2008" yearSuffix="e">
Marin 2008:
<figureCitation id="13312A2C8843FFE9FB21FE92A27F4218" box="[1276,1344,332,357]" captionStart="FIGURE 4" captionStartId="4.[151,250,1873,1895]" captionTargetBox="[176,1370,165,1829]" captionTargetId="figure@4.[175,1389,163,1842]" captionTargetPageId="4" captionText="FIGURE 4. Palaemonella shirakawai sp. nov. Holotype ovigerous female (CMNH-ZC 02525). A, right first pereiopod, lateral view; B, same, chela, mesial view; C, right second pereiopod, lateral view; D, same, carpo-palm junction, ventral view; E, same, dactylus and fixed finger, mesial view; F, right third pereiopod, lateral view; G, same, propodus and dactylus, lateral view; H, same, distal part of propodus and dactylus, lateral view; I, right fifth pereiopod, propodus and dactylus, lateral view; J, same, distal part of propodus and dactylus, lateral view." httpUri="https://zenodo.org/record/837073/files/figure.png" pageId="7" pageNumber="528">Fig. 4</figureCitation>
e
</bibRefCitation>
).
</paragraph>
<paragraph id="8BB536A98843FFE9FF1AFEAEA5E34289" blockId="7.[151,1437,151,501]" pageId="7" pageNumber="528">
As an associate in holes constructed by partner fishes (see
<bibRefCitation id="EF9B4B588843FFE9FCB9FEB1A4D942F4" author="Colin" box="[868,998,367,392]" pageId="7" pageNumber="528" refString="Colin, P. L. (1973) Burrowing behavior of the yellowhead jawfish, Opistognathus aurifrons. Copeia, 1973, 84 - 90. https: // doi. org / 10.2307 / 1442361" type="journal article" year="1973">Colin 1973</bibRefCitation>
), ecology of both species is somewhat similar. However, the present study and previous records in the popular guidebooks (
<bibRefCitation id="EF9B4B588843FFE9FBA4FE4AA21B42D0" author="Debelius" box="[1145,1316,404,429]" pageId="7" pageNumber="528" refString="Debelius, H. (1999) Crustacean guide of the World: shrimps, crabs, lobsters, mantis shrimps, amphipods. Ikan- Unterwasserarchiv, Frankfurt, 321 pp." type="book" year="1999">Debelius 1999</bibRefCitation>
;
<bibRefCitation id="EF9B4B588843FFE9FAE9FE4AA67F42AC" author="Kuiter" pageId="7" pageNumber="528" refString="Kuiter, R. H. &amp; Debelius, H. (2009) World atlas of marine fauna. Ikan-Unterwasserarchiv, Frankfult, 725 pp." type="book" year="2009">Kuiter &amp; Debelius 2009</bibRefCitation>
;
<bibRefCitation id="EF9B4B588843FFE9FE92FE66A5B742AC" author="Humann" box="[335,648,439,464]" pageId="7" pageNumber="528" refString="Humann, P. &amp; DeLoach, N. (2010) Reef creature identification: tropical Pacific. New World Publications, Jacksonville, 497 pp." type="book" year="2010">Humann &amp; DeLoach 2010</bibRefCitation>
;
<bibRefCitation id="EF9B4B588843FFE9FD4AFE69A47042AC" author="Minemizu" box="[663,847,439,464]" pageId="7" pageNumber="528" refString="Minemizu, R. (2013) Coral reef shrimps of Indo-West Pacific. Bun-ichi Sogo Shuppan, Co. Ltd., Tokyo, 144 pp. [in Japanese]" type="book" year="2013">Minemizu 2013</bibRefCitation>
) suggest that
<taxonomicName id="4C0A4D2A8843FFE9FC29FE67A3AF42AC" authority="Okuno, 2017" authorityName="Okuno" authorityYear="2017" box="[1012,1168,440,464]" class="Malacostraca" family="Palaemonidae" genus="Palaemonella" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="528" phylum="Arthropoda" rank="species" species="shirawakai" status="sp. nov.">
<emphasis id="B97EEABB8843FFE9FC29FE67A3AF42AC" box="[1012,1168,440,464]" italics="true" pageId="7" pageNumber="528">P. shirawakai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57C08843FFE9FB47FE69A3C842AC" box="[1178,1271,439,464]" pageId="7" pageNumber="528" rank="species">
<emphasis id="B97EEABB8843FFE9FB47FE69A3C842AC" bold="true" box="[1178,1271,439,464]" pageId="7" pageNumber="528">sp. nov.</emphasis>
</taxonomicNameLabel>
is perhaps an obligatory associate of jawfishes rather than gobies.
</paragraph>
</subSubSection>
</treatment>
</document>