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<document id="8D5FA3C7527BCE12C56D8650552FAE09" ID-DOI="10.5852/ejt.2024.927.2475" ID-ISSN="2118-9773" ID-Zenodo-Dep="10853688" ID-ZooBank="B6E43365-FACA-49BA-8CCD-77E4BF8F0016" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="carolina" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1711115020933" checkinUser="plazi" docAuthor="Zallot, Enrico, Kamchev, Panche, Schilthuizen, Menno, Fehér, Zoltán, Mattia, Willy De &amp; Gittenberger, Edmund" docDate="2024" docId="03A387B9FFE1FFC0FDEC1EEEFE08A792" docLanguage="en" docName="EJT.2024.927.1-163.pdf" docOrigin="European Journal of Taxonomy 927" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2475/11013" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Cochlostoma (Turritus) pageti Klemm 1962" docType="treatment" docVersion="3" lastPageNumber="42" masterDocId="FF9AFFC1FFC4FFE9FFD91D5FFFB5A629" masterDocTitle="Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)" masterLastPageNumber="163" masterPageNumber="1" pageNumber="38" updateTime="1711568280370" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AE3951470E6CF2CAE370631D38E12637">Cochlostoma Jan, 1830 revised: an overview of the subgenus Turritus Westerlund, 1883 and its species (Caenogastropoda, Cochlostomatidae)</mods:title>
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<mods:namePart id="71788A61B8B79A712E4472382F57CC0C">Zallot, Enrico</mods:namePart>
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<mods:affiliation id="665EAAD9193406E6275C5FE9949AF471">Per Ligont 1, 33070 Budoia, Italy.</mods:affiliation>
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<mods:namePart id="7294E44DAE6FEA7F6120B066F3BAA1C7">Kamchev, Panche</mods:namePart>
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<mods:affiliation id="060A3EDB23E5E5D4866DB749D710518A">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands.</mods:affiliation>
<mods:nameIdentifier id="63E605B24FA7BABF5FB8494A28A3A0E9" type="email">panekamchev@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B1670231A0A626AAE8EFFB7268F09CE0">Schilthuizen, Menno</mods:namePart>
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<mods:affiliation id="033F78797353ACCBBA26A8C20BA3B1D6">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, the Netherlands. &amp; Taxon Expeditions, Rembrandtstraat 20, 2311 VW Leiden, the Netherlands.</mods:affiliation>
<mods:nameIdentifier id="F352C088047BE11A527A0042118A1E03" type="email">menno.schilthuizen@naturalis.nl</mods:nameIdentifier>
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<mods:namePart id="B1DC8FA951B7E9C9EA1B974734385864">Fehér, Zoltán</mods:namePart>
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<mods:affiliation id="44584FBE05434CFBED007CFCC6419781">WWF Hungary, Álmos vezér útja 69 / A, 1141 Budapest, Hungary.</mods:affiliation>
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<mods:namePart id="D3749EB8D589635CF6B3B25403B75A5B">Mattia, Willy De</mods:namePart>
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<mods:affiliation id="BFB2A60E59C3546B7176392A0023E659">Central Research Laboratories, Burgring 7, 1010, Vienna, Austria; Department of Evolutionary Biology, University of Vienna, Djerassiplatz 1, 1030, Vienna, Austria.</mods:affiliation>
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<mods:namePart id="1EB338751229F48CE970A8A3780D6B4D">Gittenberger, Edmund</mods:namePart>
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<mods:affiliation id="8E10E77CB81EFE28D246580861BA4DB7">Naturalis Biodiversity Center, P. O. Box 9517, 2300 RA Leiden, the Netherlands.</mods:affiliation>
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<mods:title id="2E4E266D185A537A3F5FBF893E2CA13A">European Journal of Taxonomy</mods:title>
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<mods:part id="3EDBE87FF30AC4C3978B608107725CCC">
<mods:date id="C6A1C6ED86544B0789846AB59BAA5746">2024</mods:date>
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<mods:number id="2F1F99EA0DDD4EFC172E92E0566CA9B8">927</mods:number>
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<mods:classification id="6EF81967E99DE5E70B227C5C40EF47C5">journal article</mods:classification>
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<treatment id="03A387B9FFE1FFC0FDEC1EEEFE08A792" ID-DOI="http://doi.org/10.5281/zenodo.10854662" ID-Zenodo-Dep="10854662" LSID="urn:lsid:plazi:treatment:03A387B9FFE1FFC0FDEC1EEEFE08A792" httpUri="http://treatment.plazi.org/id/03A387B9FFE1FFC0FDEC1EEEFE08A792" lastPageId="41" lastPageNumber="42" pageId="37" pageNumber="38">
<subSubSection id="C3106524FFE1FFCCFDEC1EEEFC4BA5E2" box="[565,1022,945,972]" pageId="37" pageNumber="38" type="nomenclature">
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<heading id="D0FD81C3FFE1FFCCFDEC1EEEFC4BA5E2" box="[565,1022,945,972]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="4C0A4D2CFFE1FFCCFDEC1EEEFC4BA5E2" ID-CoL="WNNW" authority="Klemm, 1962" authorityName="Klemm" authorityYear="1962" box="[565,1022,945,972]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE1FFCCFDEC1EEEFD66A5E5" bold="true" box="[565,723,945,972]" italics="true" pageId="37" pageNumber="38">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFFE1FFCCFD3D1EEDFD4EA5E5" bold="true" box="[740,763,946,972]" italics="true" pageId="37" pageNumber="38">T.</emphasis>
)
<emphasis id="B97EEABDFFE1FFCCFCD51EEEFCE0A5E2" bold="true" box="[780,853,945,971]" italics="true" pageId="37" pageNumber="38">pageti</emphasis>
<bibRefCitation id="EF9B4B5EFFE1FFCCFC841EEEFC4BA5E2" author="Klemm W." box="[861,1022,945,971]" pageId="37" pageNumber="38" pagination="203 - 258" refId="ref37223" refString="Klemm W. 1962. X. Teil. Die Gehauseschnecken. In: Beier M. (ed.) Zoologische Studien in West- Griechenland. Sitzungsberichte, Osterreichische Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 171 (6 / 7): 203 - 258." type="journal article" year="1962">Klemm, 1962</bibRefCitation>
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<paragraph id="8BB536AFFFE1FFCCFDBC1E8CFC7BA5C7" blockId="37.[565,1022,945,1006]" box="[613,974,979,1006]" pageId="37" pageNumber="38">
<figureCitation id="13312A2AFFE1FFCCFDBC1E8CFD09A5C4" box="[613,700,979,1005]" captionStart="Fig" captionStartId="31.[189,232,1925,1951]" captionTargetBox="[189,1398,1203,1883]" captionTargetId="figure-230@31.[189,1398,1203,1883]" captionTargetPageId="31" captionText="Fig. 26. Distribution of taxa part of clade A: cyan dots =Cochlostoma (T.) reitteri (Boettger, 1880); green dots =C. (T.) pallgergelyi sp. nov.; red dots =C. (T.) pageti Klemm, 1962. Black center indicates samples with amplified 16S." figureDoi="http://doi.org/10.5281/zenodo.10853746" httpUri="https://zenodo.org/record/10853746/files/figure.png" pageId="37" pageNumber="38">Figs 26</figureCitation>
(red dots), 27D, 32–33
</paragraph>
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<subSubSection id="C3106524FFE1FFCCFF64194BFC3BA278" pageId="37" pageNumber="38" type="reference_group">
<paragraph id="8BB536AFFFE1FFCCFF64194BFCAFA207" blockId="37.[189,910,1044,1105]" box="[189,794,1044,1070]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB1A1181FFE1FFCCFF64194BFCAFA207" box="[189,794,1044,1070]" pageId="37" pageNumber="38">
<treatmentCitation id="0AAB10BEFFE1FFCCFF64194BFD25A207" author="Klemm W." box="[189,656,1044,1070]" page="213" pageId="37" pageNumber="38" year="1962">
<taxonomicName id="4C0A4D2CFFE1FFCCFF64194BFD25A207" ID-CoL="WNNW" authority="Klemm, 1962: 213" authorityName="Klemm" authorityPageNumber="213" authorityYear="1962" box="[189,656,1044,1070]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="pageti">
<emphasis id="B97EEABDFFE1FFCCFF64194BFE1DA207" box="[189,424,1044,1070]" italics="true" pageId="37" pageNumber="38">Cochlostoma pageti</emphasis>
<bibRefCitation id="EF9B4B5EFFE1FFCCFE69194BFD25A207" author="Klemm W." box="[432,656,1044,1070]" pageId="37" pageNumber="38" pagination="203 - 258" refId="ref37223" refString="Klemm W. 1962. X. Teil. Die Gehauseschnecken. In: Beier M. (ed.) Zoologische Studien in West- Griechenland. Sitzungsberichte, Osterreichische Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 171 (6 / 7): 203 - 258." type="journal article" year="1962">Klemm, 1962: 213</bibRefCitation>
</taxonomicName>
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, pl. 1 fig. 1.
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</paragraph>
<paragraph id="8BB536AFFFE1FFCCFF641969FC3BA278" blockId="37.[189,910,1044,1105]" box="[189,910,1078,1105]" pageId="37" pageNumber="38">
<treatmentCitationGroup id="AB1A1181FFE1FFCCFF641969FC3BA278" box="[189,910,1078,1105]" pageId="37" pageNumber="38">
<treatmentCitation id="0AAB10BEFFE1FFCCFF641969FCB7A279" author="Schutt H." box="[189,770,1078,1104]" page="21" pageId="37" pageNumber="38" year="1977">
<taxonomicName id="4C0A4D2CFFE1FFCCFF641969FCB7A279" ID-CoL="5DW76" authority="Schutt, 1977: 21" authorityName="Schutt" authorityPageNumber="21" authorityYear="1977" box="[189,770,1078,1104]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="37" pageNumber="38" phylum="Mollusca" rank="subSpecies" species="gracile" subSpecies="subaiorum">
<emphasis id="B97EEABDFFE1FFCCFF641969FD82A279" box="[189,567,1078,1104]" italics="true" pageId="37" pageNumber="38">Cochlostoma gracile subaiorum</emphasis>
<bibRefCitation id="EF9B4B5EFFE1FFCCFDE71969FCB7A279" author="Schutt H." box="[574,770,1078,1104]" pageId="37" pageNumber="38" pagination="17 - 35" refId="ref38098" refString="Schutt H. 1977. Revision der griechischen Cochlostoma. Archiv fur Molluskenkunde 108 (1 / 3): 17 - 35." type="journal article" year="1977">Schütt, 1977: 21</bibRefCitation>
</taxonomicName>
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, pl. 1 fig. 7.
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</paragraph>
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<subSubSection id="C3106524FFE1FFCCFF641928FCD3A397" pageId="37" pageNumber="38" type="materials_examined">
<materialsCitation id="3B623CF2FFE1FFCCFF641928FAA9A296" box="[189,1308,1143,1215]" collectingDate="1933-05" collectionCode="NHMW" collectorName="Beier" country="Greece" elevation="1500" latitude="39.5682" location="Paraskevi" longLatPrecision="6" longitude="21.2607" pageId="37" pageNumber="38" specimenCode="NHMW74159" specimenCount="1" specimenCount-female="1" stateProvince="Epirus" typeStatus="holotype">
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</emphasis>
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</paragraph>
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<collectingCountry id="F31D763FFFE1FFCCFF6419FAFE9AA296" box="[189,303,1189,1215]" name="Greece" pageId="37" pageNumber="38">GREECE</collectingCountry>
• probably
<specimenCount id="9D0CFD26FFE1FFCCFE6119FAFE7AA296" box="[440,463,1189,1215]" count="1" pageId="37" pageNumber="38" type="female">♀</specimenCount>
; “
<location id="8ED56074FFE1FFCCFE3119FBFDD5A297" LSID="urn:lsid:plazi:treatment:03A387B9FFE1FFC0FDEC1EEEFE08A792:8ED56074FFE1FFCCFE3119FBFDD5A297" box="[488,608,1188,1214]" country="Greece" name="Paraskevi" pageId="37" pageNumber="38" stateProvince="Epirus">Paraskevi</location>
,
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, im
<collectingRegion id="49CEF84DFFE1FFCCFD2019FBFCFEA297" box="[761,843,1188,1214]" country="Greece" name="Ipeiros" pageId="37" pageNumber="38">Epirus</collectingRegion>
”;
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<collectingDate id="EFF0E987FFE1FFCCFCBB19FAFC68A296" box="[866,989,1189,1215]" pageId="37" pageNumber="38" value="1933-05">May 1933</collectingDate>
</date>
;
<collectorName id="26FF5379FFE1FFCCFC3019FBFB9CA297" box="[1001,1065,1188,1214]" pageId="37" pageNumber="38">Beier</collectorName>
leg.;
<specimenCode id="DBAC9ED4FFE1FFCCFBB219FBFAADA296" box="[1131,1304,1188,1215]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="37" pageNumber="38" type="Museum">NHMW74159</specimenCode>
.
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</materialsCitation>
<paragraph id="8BB536AFFFE1FFCCFF6419BAFE3BA2D6" blockId="37.[189,398,1253,1279]" box="[189,398,1253,1279]" pageId="37" pageNumber="38">
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<emphasis id="B97EEABDFFE1FFCCFF6419BAFE3BA2D6" bold="true" box="[189,398,1253,1279]" pageId="37" pageNumber="38">Other specimens</emphasis>
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<materialsCitation id="3B623CF2FFE1FFCCFF64184CFB70A304" box="[189,1221,1298,1325]" collectionCode="NHMWK" collectorName="Subai" country="Greece" latitude="39.4488" location="Terovo" longLatPrecision="6" longitude="20.8833" pageId="37" pageNumber="38" specimenCode="NHMWK69266" specimenCount="1">
<collectingCountry id="F31D763FFFE1FFCCFF64184CFE9AA304" box="[189,303,1299,1325]" name="Greece" pageId="37" pageNumber="38">GREECE</collectingCountry>
• 2-
<location id="8ED56074FFE1FFCCFEA3184CFE64A304" LSID="urn:lsid:plazi:treatment:03A387B9FFE1FFC0FDEC1EEEFE08A792:8ED56074FFE1FFCCFEA3184CFE64A304" box="[378,465,1299,1325]" country="Greece" latitude="39.4488" longLatPrecision="6" longitude="20.8833" name="Terovo" pageId="37" pageNumber="38">Terovo</location>
;
<geoCoordinate id="EE3E5068FFE1FFCCFE3B184CFDC6A304" box="[482,627,1299,1325]" degrees="39.4488" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="39.4488">39.4488° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FFE1FFCCFD5C184CFCA7A304" box="[645,786,1299,1325]" degrees="20.8833" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5" value="20.8833">20.8833° E</geoCoordinate>
; 1975;
<collectorName id="26FF5379FFE1FFCCFCAC184CFC0DA304" box="[885,952,1299,1325]" pageId="37" pageNumber="38">Subai</collectorName>
leg.;
<specimenCode id="DBAC9ED4FFE1FFCCFBDF184CFB70A304" box="[1030,1221,1299,1325]" collectionCode="NHMWK" pageId="37" pageNumber="38">NHMWK69266</specimenCode>
</materialsCitation>
•
<materialsCitation id="3B623CF2FFE1FFCCFB33184CFAA3A366" collectionCode="HNHM" collectorName="Kontschan &amp; Muranyi &amp; Szederjesi &amp; Ujvari" country="Greece" latitude="39.6333" location="Katafito" longLatPrecision="6" longitude="21.2242" pageId="37" pageNumber="38" specimenCode="HNHM99329" specimenCount="1">
3-
<location id="8ED56074FFE1FFCCFAC9184CFAC7A304" LSID="urn:lsid:plazi:treatment:03A387B9FFE1FFC0FDEC1EEEFE08A792:8ED56074FFE1FFCCFAC9184CFAC7A304" box="[1296,1394,1299,1325]" country="Greece" latitude="39.6333" longLatPrecision="6" longitude="21.2242" name="Katafito" pageId="37" pageNumber="38">Katafito</location>
;
<geoCoordinate id="EE3E5068FFE1FFCCFF64186AFEF2A366" box="[189,327,1333,1359]" degrees="39.6333" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="39.6333">39.6333° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FFE1FFCCFE8A186AFE6EA366" box="[339,475,1333,1359]" degrees="21.2242" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5" value="21.2242">21.2242° E</geoCoordinate>
; 2011;
<collectorName id="26FF5379FFE1FFCCFDE9186AFD04A366" box="[560,689,1333,1359]" pageId="37" pageNumber="38">Kontschán</collectorName>
,
<collectorName id="26FF5379FFE1FFCCFD65186AFC91A366" box="[700,804,1333,1359]" pageId="37" pageNumber="38">Murányi</collectorName>
,
<collectorName id="26FF5379FFE1FFCCFCE9186AFC1CA366" box="[816,937,1333,1359]" pageId="37" pageNumber="38">Szederjesi</collectorName>
and
<collectorName id="26FF5379FFE1FFCCFC38186AFB99A366" box="[993,1068,1333,1359]" pageId="37" pageNumber="38">Ujvári</collectorName>
leg.;
<specimenCode id="DBAC9ED4FFE1FFCCFBB7186AFAA3A366" box="[1134,1302,1333,1359]" collectionCode="HNHM" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="37" pageNumber="38">HNHM99329</specimenCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB536AFFFE1FFCCFF641829FEE8A3A6" blockId="37.[189,349,1397,1424]" box="[189,349,1397,1424]" pageId="37" pageNumber="38">
<heading id="D0FD81C3FFE1FFCCFF641829FEE8A3A6" bold="true" box="[189,349,1397,1424]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B97EEABDFFE1FFCCFF641829FEE8A3A6" bold="true" box="[189,349,1397,1424]" pageId="37" pageNumber="38">Type locality</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFFE1FFCCFF6418FCFCD3A397" blockId="37.[189,870,1443,1470]" box="[189,870,1443,1470]" pageId="37" pageNumber="38">
<collectingCountry id="F31D763FFFE1FFCCFF6418FCFE9AA394" box="[189,303,1443,1469]" name="Greece" pageId="37" pageNumber="38">GREECE</collectingCountry>
• 1-
<collectingRegion id="49CEF84DFFE1FFCCFEB118FCFE0DA394" box="[360,440,1443,1469]" country="Greece" name="Ipeiros" pageId="37" pageNumber="38">Epirus</collectingRegion>
,
<location id="8ED56074FFE1FFCCFE1A18FCFD8CA394" LSID="urn:lsid:plazi:treatment:03A387B9FFE1FFC0FDEC1EEEFE08A792:8ED56074FFE1FFCCFE1A18FCFD8CA394" box="[451,569,1443,1469]" country="Greece" latitude="39.5682" longLatPrecision="6" longitude="21.2607" name="Paraskevi" pageId="37" pageNumber="38" stateProvince="Epirus">Paraskevi</location>
;
<geoCoordinate id="EE3E5068FFE1FFCCFD9C18FCFD7AA397" box="[581,719,1443,1470]" degrees="39.5682" direction="north" orientation="latitude" pageId="37" pageNumber="38" precision="5" value="39.5682">39.5682° N</geoCoordinate>
,
<geoCoordinate id="EE3E5068FFE1FFCCFD0218FCFCD7A397" box="[731,866,1443,1470]" degrees="21.2607" direction="east" orientation="longitude" pageId="37" pageNumber="38" precision="5" value="21.2607">21.2607° E</geoCoordinate>
.
</paragraph>
</subSubSection>
<subSubSection id="C3106524FFE1FFCCFF6418BCFBE5A171" pageId="37" pageNumber="38" type="description">
<paragraph id="8BB536AFFFE1FFCCFF6418BCFEFBA3D4" blockId="37.[189,334,1507,1533]" box="[189,334,1507,1533]" pageId="37" pageNumber="38">
<heading id="D0FD81C3FFE1FFCCFF6418BCFEFBA3D4" bold="true" box="[189,334,1507,1533]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B97EEABDFFE1FFCCFF6418BCFEFBA3D4" bold="true" box="[189,334,1507,1533]" pageId="37" pageNumber="38">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFFE1FFCCFF641B4EFE8CA059" blockId="37.[189,1399,1553,1648]" pageId="37" pageNumber="38">
<smallCapsWord id="8D53A073FFE1FFCCFF641B4EFEB4A003" baselines="1572,1573" box="[189,257,1553,1579]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="37" pageNumber="38">SHELL</smallCapsWord>
. Spaced riblets on last part of protoconch. Teleoconch spotless, with prominent and rather thin ribs becoming weaker toward aperture. Moderately strong lip with indented columellar lobe covering umbilicus.
</paragraph>
<paragraph id="8BB536AFFFE1FFCCFF641BC8FB3CA0FA" blockId="37.[189,1398,1686,1748]" pageId="37" pageNumber="38">
<smallCapsWord id="8D53A073FFE1FFCCFF641BC8FED9A086" baselines="1706,1706" box="[189,364,1687,1713]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="37" pageNumber="38">MEASUREMENTS</smallCapsWord>
.
<specimenCount id="9D0CFD26FFE1FFCCFEA71BC9FE73A098" box="[382,454,1686,1713]" count="5" pageId="37" pageNumber="38" type="female">5 ♀♀</specimenCount>
: whorls=7.1–8, H =
<quantity id="4CF29B4AFFE1FFCCFD641BC9FCE5A098" box="[701,848,1686,1713]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" metricValueMax="7.4" metricValueMin="5.4" pageId="37" pageNumber="38" unit="mm" value="6.4" valueMax="7.4" valueMin="5.4">5.4–7.4 mm</quantity>
, H/W=2.43–2.62, roundness =0.13–0.16, ribs incl.= 56–66°, apert. incl.= 14–26°, ribs/mm 1
<superScript id="7C7F9BE7FFE1FFCCFD051BE7FD52A0EF" attach="left" box="[732,743,1720,1734]" fontSize="6" pageId="37" pageNumber="38">st</superScript>
wh.= 6–10, ribs/mm 4
<superScript id="7C7F9BE7FFE1FFCCFC2A1BE7FBB5A0EF" attach="left" box="[1011,1024,1720,1734]" fontSize="6" pageId="37" pageNumber="38">th</superScript>
wh.=7–10.
</paragraph>
<paragraph id="8BB536AFFFE1FFCCFF641BA5FBE5A171" blockId="37.[189,1399,1785,1880]" pageId="37" pageNumber="38">
<smallCapsWord id="8D53A073FFE1FFCCFF641BA5FEA1A13B" baselines="1805,1805" box="[189,276,1786,1812]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Female" pageId="37" pageNumber="38">FEMALE</smallCapsWord>
<smallCapsWord id="8D53A073FFE1FFCCFEC61BA0FEC3A13B" baselines="1805" box="[287,374,1791,1810]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="genital" pageId="37" pageNumber="38">GENITAL</smallCapsWord>
<smallCapsWord id="8D53A073FFE1FFCCFE5B1BA0FE63A13B" baselines="1805" box="[386,470,1791,1810]" lowerCaseFontSize="8" mainFontSize="11" normCase="lower" normString="organs" pageId="37" pageNumber="38">ORGANS</smallCapsWord>
. Ventral-posterior connection of pedunculus to bursa copulatrix. Short, oval seminal receptacle confined to ventral side of body with 2–3 loops close to apex. Junction of uterus gland moderately far from connection between pedunculus and distal oviduct.
</paragraph>
</subSubSection>
<subSubSection id="C3106524FFE1FFC0FF641A21FE08A792" lastPageId="41" lastPageNumber="42" pageId="37" pageNumber="38" type="discussion">
<paragraph id="8BB536AFFFE1FFCCFF641A21FE9AA1B1" blockId="37.[189,303,1918,1944]" box="[189,303,1918,1944]" pageId="37" pageNumber="38">
<heading id="D0FD81C3FFE1FFCCFF641A21FE9AA1B1" bold="true" box="[189,303,1918,1944]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B97EEABDFFE1FFCCFF641A21FE9AA1B1" bold="true" box="[189,303,1918,1944]" pageId="37" pageNumber="38">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFFE1FFCEFF641AF3FB9BA7A4" blockId="37.[189,1399,1964,2025]" lastBlockId="39.[189,1399,265,397]" lastPageId="39" lastPageNumber="40" pageId="37" pageNumber="38">
The analysis of the
<typeStatus id="54B1880DFFE1FFCCFE6A1AF2FE53A1EE" box="[435,486,1965,1991]" pageId="37" pageNumber="38">type</typeStatus>
series indicates that
<taxonomicName id="4C0A4D2CFFE1FFCCFD321AF3FC3BA1EF" authorityName="Klemm" authorityYear="1962" box="[747,910,1964,1990]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="37" pageNumber="38" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE1FFCCFD321AF3FCB0A1EF" box="[747,773,1964,1990]" italics="true" pageId="37" pageNumber="38">C.</emphasis>
(
<emphasis id="B97EEABDFFE1FFCCFCC21AF2FC84A1EF" box="[795,817,1965,1990]" italics="true" pageId="37" pageNumber="38">T.</emphasis>
)
<emphasis id="B97EEABDFFE1FFCCFC9C1AF2FC3BA1EF" box="[837,910,1965,1990]" italics="true" pageId="37" pageNumber="38">pageti</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D2CFFE1FFCCFC091AF3FAC2A1EF" authority="Schutt, 1977" authorityName="Schutt" authorityYear="1977" box="[976,1399,1964,1990]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="37" pageNumber="38" phylum="Mollusca" rank="subSpecies" species="gracile" subSpecies="subaiorum">
<emphasis id="B97EEABDFFE1FFCCFC091AF3FB67A1EF" box="[976,1234,1964,1990]" italics="true" pageId="37" pageNumber="38">C. gracile subaiorum</emphasis>
<bibRefCitation id="EF9B4B5EFFE1FFCCFB071AF3FAC2A1EF" author="Schutt H." box="[1246,1399,1964,1990]" pageId="37" pageNumber="38" pagination="17 - 35" refId="ref38098" refString="Schutt H. 1977. Revision der griechischen Cochlostoma. Archiv fur Molluskenkunde 108 (1 / 3): 17 - 35." type="journal article" year="1977">Schütt, 1977</bibRefCitation>
</taxonomicName>
are conspecific. The short distance between the
<typeStatus id="54B1880DFFE1FFCCFCD81A90FC81A1C0" box="[769,820,1999,2025]" pageId="37" pageNumber="38">type</typeStatus>
localities supports this conclusion. In Wagner’s collection (MIZPAS -W7085) there is a specimen (
<figureCitation id="13312A2AFFE3FFCEFCF61C56FC3EA70A" box="[815,907,265,291]" captionStart="Fig" captionStartId="40.[189,232,1642,1668]" captionTargetBox="[398,1173,265,1605]" captionTargetId="figure-19@40.[398,1173,265,1605]" captionTargetPageId="40" captionText="Fig. 34. Specimen in Wagner’s collection (MIZPAS-W7085) from the Acarnania region of Greece, named as Pomatias pindicus (nomen nudum)." figureDoi="http://doi.org/10.5281/zenodo.10853765" httpUri="https://zenodo.org/record/10853765/files/figure.png" pageId="39" pageNumber="40">Fig. 34</figureCitation>
), named
<emphasis id="B97EEABDFFE3FFCEFC241C56FB34A70A" box="[1021,1153,265,291]" italics="true" pageId="39" pageNumber="40">
<taxonomicName id="4C0A4D2CFFE3FFCEFC241C56FBC3A70A" authorityName="A.Letourneux &amp; Bourguignat" authorityYear="1887" box="[1021,1142,265,291]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pindicus">“pindicus</taxonomicName>
”
</emphasis>
from the Acarnania region, roughly
<quantity id="4CF29B4AFFE3FFCEFEA51C73FE63A76F" box="[380,470,300,326]" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="39" pageNumber="40" unit="km" value="100.0">100 km</quantity>
south of the range of
<taxonomicName id="4C0A4D2CFFE3FFCEFD391C72FCCBA76F" authorityName="Klemm" authorityYear="1962" box="[736,894,300,327]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE3FFCEFD391C72FD4EA76E" box="[736,763,301,327]" italics="true" pageId="39" pageNumber="40">C.</emphasis>
(
<emphasis id="B97EEABDFFE3FFCEFCD41C72FC96A76F" box="[781,803,301,326]" italics="true" pageId="39" pageNumber="40">T.</emphasis>
)
<emphasis id="B97EEABDFFE3FFCEFCEC1C72FCCBA76F" box="[821,894,301,326]" italics="true" pageId="39" pageNumber="40">pageti</emphasis>
</taxonomicName>
. Despite the low resolution of the picture, the deep indentation of the columellar lobe is clearly visible, like in
<taxonomicName id="4C0A4D2CFFE3FFCEFC291C0FFB3AA740" authorityName="Klemm" authorityYear="1962" box="[1008,1167,336,362]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE3FFCEFC291C0FFBBEA743" box="[1008,1035,336,362]" italics="true" pageId="39" pageNumber="40">C.</emphasis>
(
<emphasis id="B97EEABDFFE3FFCEFBC71C0FFB81A740" box="[1054,1076,336,361]" italics="true" pageId="39" pageNumber="40">T.</emphasis>
)
<emphasis id="B97EEABDFFE3FFCEFB9F1C0FFB3AA740" box="[1094,1167,336,361]" italics="true" pageId="39" pageNumber="40">pageti</emphasis>
</taxonomicName>
. It remains unclear whether this population is conspecific with or only related to
<taxonomicName id="4C0A4D2CFFE3FFCEFC551C2CFB92A7A5" authorityName="Klemm" authorityYear="1962" box="[908,1063,371,397]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE3FFCEFC551C2CFC13A7A4" box="[908,934,371,397]" italics="true" pageId="39" pageNumber="40">C.</emphasis>
(
<emphasis id="B97EEABDFFE3FFCEFC611C2CFC7BA7A5" box="[952,974,371,396]" italics="true" pageId="39" pageNumber="40">T.</emphasis>
)
<emphasis id="B97EEABDFFE3FFCEFC071C2CFB92A7A5" box="[990,1063,371,396]" italics="true" pageId="39" pageNumber="40">pageti</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="DF756627FFE2FFCFFF641AF9FAC5A1CD" ID-DOI="http://doi.org/10.5281/zenodo.10853759" ID-Zenodo-Dep="10853759" httpUri="https://zenodo.org/record/10853759/files/figure.png" pageId="38" pageNumber="39" startId="38.[189,232,1958,1984]" targetBox="[189,1398,265,1914]" targetPageId="38">
<paragraph id="8BB536AFFFE2FFCFFF641AF9FAC5A1CD" blockId="38.[189,1399,1958,2020]" pageId="38" pageNumber="39">
<emphasis id="B97EEABDFFE2FFCFFF641AF9FE83A1E9" bold="true" box="[189,310,1958,1984]" pageId="38" pageNumber="39">Fig. 32. A</emphasis>
. Holotype of
<taxonomicName id="4C0A4D2CFFE2FFCFFE031AF8FC2AA1E8" authority="Klemm, 1962" authorityName="Klemm" authorityYear="1962" box="[474,927,1959,1985]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE2FFCFFE031AF8FDC0A1E8" box="[474,629,1959,1985]" italics="true" pageId="38" pageNumber="39">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFFE2FFCFFD5F1AF8FD29A1E9" box="[646,668,1959,1984]" italics="true" pageId="38" pageNumber="39">T.</emphasis>
)
<emphasis id="B97EEABDFFE2FFCFFD751AF8FD40A1E9" box="[684,757,1959,1984]" italics="true" pageId="38" pageNumber="39">pageti</emphasis>
<bibRefCitation id="EF9B4B5EFFE2FFCFFD241AF8FC2AA1E8" author="Klemm W." box="[765,927,1959,1985]" pageId="38" pageNumber="39" pagination="203 - 258" refId="ref37223" refString="Klemm W. 1962. X. Teil. Die Gehauseschnecken. In: Beier M. (ed.) Zoologische Studien in West- Griechenland. Sitzungsberichte, Osterreichische Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 171 (6 / 7): 203 - 258." type="journal article" year="1962">Klemm, 1962</bibRefCitation>
</taxonomicName>
(NHMW-74159).
<emphasis id="B97EEABDFFE2FFCFFBA31AF8FB3BA1E8" bold="true" box="[1146,1166,1959,1985]" pageId="38" pageNumber="39">B</emphasis>
. Paratype of
<taxonomicName id="4C0A4D2CFFE2FFCFFAF21AF8FD84A1CD" authority="Schutt, 1977" authorityName="Schutt" authorityYear="1977" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="38" pageNumber="39" phylum="Mollusca" rank="subSpecies" species="gracile" subGenus="Turritus" subSpecies="subaiorum">
<emphasis id="B97EEABDFFE2FFCFFAF21AF8FAF3A1E8" box="[1323,1350,1959,1985]" italics="true" pageId="38" pageNumber="39">C.</emphasis>
(
<emphasis id="B97EEABDFFE2FFCFFA8E1AF8FAD8A1E9" box="[1367,1389,1959,1984]" italics="true" pageId="38" pageNumber="39">T.</emphasis>
)
<emphasis id="B97EEABDFFE2FFCFFF641A95FE21A1CD" box="[189,404,1994,2020]" italics="true" pageId="38" pageNumber="39">gracile subaiorum</emphasis>
<bibRefCitation id="EF9B4B5EFFE2FFCFFE451A95FD84A1CD" author="Schutt H." box="[412,561,1994,2020]" pageId="38" pageNumber="39" pagination="17 - 35" refId="ref38098" refString="Schutt H. 1977. Revision der griechischen Cochlostoma. Archiv fur Molluskenkunde 108 (1 / 3): 17 - 35." type="journal article" year="1977">Schütt, 1977</bibRefCitation>
</taxonomicName>
(NHMW-K69266). The latter is here synonymized with
<taxonomicName id="4C0A4D2CFFE2FFCFFB141A95FAD9A1CD" authorityName="Klemm" authorityYear="1962" box="[1229,1388,1994,2020]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="38" pageNumber="152" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE2FFCFFB141A95FB5DA1CD" box="[1229,1256,1994,2020]" italics="true" pageId="38" pageNumber="39">C.</emphasis>
(
<emphasis id="B97EEABDFFE2FFCFFB201A94FABAA1CD" box="[1273,1295,1995,2020]" italics="true" pageId="38" pageNumber="39">T.</emphasis>
) pageti
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF756627FFE3FFCEFF641A91FBCBA1C1" ID-DOI="http://doi.org/10.5281/zenodo.10853761" ID-Zenodo-Dep="10853761" box="[189,1150,1998,2024]" httpUri="https://zenodo.org/record/10853761/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1998,2024]" targetBox="[189,1398,547,1957]" targetPageId="39">
<paragraph id="8BB536AFFFE3FFCEFF641A91FBCBA1C1" blockId="39.[189,1150,1998,2024]" box="[189,1150,1998,2024]" pageId="39" pageNumber="40">
<emphasis id="B97EEABDFFE3FFCEFF641A91FEAFA1C1" bold="true" box="[189,282,1998,2024]" pageId="39" pageNumber="40">Fig. 33.</emphasis>
<taxonomicName id="4C0A4D2CFFE3FFCEFEF91A91FD52A1C1" authority="Klemm, 1962" authorityName="Klemm" authorityYear="1962" box="[288,743,1998,2024]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="39" pageNumber="40" phylum="Mollusca" rank="species" species="pageti" subGenus="Turritus">
<emphasis id="B97EEABDFFE3FFCEFEF91A91FE0EA1C1" box="[288,443,1998,2024]" italics="true" pageId="39" pageNumber="40">Cochlostoma</emphasis>
(
<emphasis id="B97EEABDFFE3FFCEFE151A90FE57A1C1" box="[460,482,1999,2024]" italics="true" pageId="39" pageNumber="40">T.</emphasis>
)
<emphasis id="B97EEABDFFE3FFCEFE2A1A90FD89A1C1" box="[499,572,1999,2024]" italics="true" pageId="39" pageNumber="40">pageti</emphasis>
<bibRefCitation id="EF9B4B5EFFE3FFCEFD9D1A91FD52A1C1" author="Klemm W." box="[580,743,1998,2024]" pageId="39" pageNumber="40" pagination="203 - 258" refId="ref37223" refString="Klemm W. 1962. X. Teil. Die Gehauseschnecken. In: Beier M. (ed.) Zoologische Studien in West- Griechenland. Sitzungsberichte, Osterreichische Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse, Abteilung I 171 (6 / 7): 203 - 258." type="journal article" year="1962">Klemm, 1962</bibRefCitation>
</taxonomicName>
, 3- Katafico, GR (HNHM-99329).
</paragraph>
</caption>
<caption id="DF756627FFECFFC1FF641B35FD62A081" ID-DOI="http://doi.org/10.5281/zenodo.10853765" ID-Zenodo-Dep="10853765" httpUri="https://zenodo.org/record/10853765/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,232,1642,1668]" targetBox="[398,1173,265,1605]" targetPageId="40">
<paragraph id="8BB536AFFFECFFC1FF641B35FD62A081" blockId="40.[189,1398,1642,1704]" pageId="40" pageNumber="41">
<emphasis id="B97EEABDFFECFFC1FF641B35FEA8A0AD" bold="true" box="[189,285,1642,1668]" pageId="40" pageNumber="41">Fig. 34.</emphasis>
Specimen in Wagner’s collection (MIZPAS-W7085) from the Acarnania region of Greece, named as
<taxonomicName id="4C0A4D2CFFECFFC1FEEB1BD1FDBEA081" authorityName="A.Letourneux &amp; Bourguignat" authorityYear="1887" box="[306,523,1678,1704]" class="Gastropoda" family="Pomatiidae" genus="Pomatias" kingdom="Animalia" order="Littorinimorpha" pageId="40" pageNumber="41" phylum="Mollusca" rank="species" species="pindicus">
<emphasis id="B97EEABDFFECFFC1FEEB1BD1FDBEA081" box="[306,523,1678,1704]" italics="true" pageId="40" pageNumber="41">Pomatias pindicus</emphasis>
</taxonomicName>
(nomen nudum).
</paragraph>
</caption>
<paragraph id="8BB536AFFFEDFFC0FF641C57FDB6A70B" blockId="41.[189,515,264,291]" box="[189,515,264,291]" pageId="41" pageNumber="42">
<heading id="D0FD81C3FFEDFFC0FF641C57FDB6A70B" bold="true" box="[189,515,264,291]" fontSize="11" level="3" pageId="41" pageNumber="42" reason="3">
<emphasis id="B97EEABDFFEDFFC0FF641C57FDB6A70B" bold="true" box="[189,515,264,291]" pageId="41" pageNumber="42">Additional note to clade A</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536AFFFEDFFC0FF641C68FE08A792" blockId="41.[189,1399,311,443]" pageId="41" pageNumber="42">
A terminal branch of clade A includes samples from the central and southern Apennines that can be referred to
<taxonomicName id="4C0A4D2CFFEDFFC0FEE61C04FD96A75D" baseAuthorityName="Paulucci" baseAuthorityYear="1878" box="[319,547,347,373]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="cassiniacum" subGenus="Turritus">
<emphasis id="B97EEABDFFEDFFC0FEE61C04FEEFA75C" box="[319,346,347,373]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(
<emphasis id="B97EEABDFFEDFFC0FEB21C04FE34A75D" box="[363,385,347,372]" italics="true" pageId="41" pageNumber="42">T.</emphasis>
)
<emphasis id="B97EEABDFFEDFFC0FE4B1C04FD96A75D" box="[402,547,347,372]" italics="true" pageId="41" pageNumber="42">cassiniacum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D2CFFEDFFC0FD851C04FCB6A75D" baseAuthorityName="Paulucci" baseAuthorityYear="1879" box="[604,771,346,373]" class="Gastropoda" family="Cochlostomatidae" genus="Cochlostoma" kingdom="Animalia" order="Architaenioglossa" pageId="41" pageNumber="42" phylum="Mollusca" rank="species" species="adamii" subGenus="Turritus">
<emphasis id="B97EEABDFFEDFFC0FD851C04FDC3A75C" box="[604,630,347,373]" italics="true" pageId="41" pageNumber="42">C.</emphasis>
(
<emphasis id="B97EEABDFFEDFFC0FD511C04FD2BA75D" box="[648,670,347,372]" italics="true" pageId="41" pageNumber="42">T.</emphasis>
)
<emphasis id="B97EEABDFFEDFFC0FD771C05FCB6A75D" box="[686,771,346,372]" italics="true" pageId="41" pageNumber="42">adamii</emphasis>
</taxonomicName>
<emphasis id="B97EEABDFFEDFFC0FCDA1C05FCB2A75D" box="[771,775,346,372]" italics="true" pageId="41" pageNumber="42">.</emphasis>
They share the female genital morphology and some shell features, like the umbilicus hidden by the curved columellar lobe and the pronounced ribbing, with rounded, whitish ribs.
</paragraph>
</subSubSection>
</treatment>
</document>
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