treatments-xml/data/03/A3/87/03A387A7BC22FFCCFF655F1CFE638D4D.xml
2024-06-21 12:22:17 +02:00

334 lines
44 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="A2B58A77685D9D55F3613313BA066658" ID-DOI="10.5281/zenodo.197916" ID-GBIF-Dataset="5d407501-96a0-44cd-8a9c-47734417e6f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="197916" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460282467730" checkinUser="plazi" docAuthor="Zapata-Guardiola, Rebeca &amp; López-González, Pablo J." docDate="2010" docId="03A387A7BC22FFCCFF655F1CFE638D4D" docLanguage="en" docName="zt02616p068.pdf" docOrigin="Zootaxa 2616" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tokoprymno anatis Zapata-Guardiola &amp; López-González, 2010, sp. nov." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FF9AFFDFBC2DFFDEFFF25C1BFFB58C21" masterDocTitle="Redescription of Thouarella brucei Thomson and Ritchie, 1906 (Cnidaria: Octocorallia: Primnoidae) and description of two new Antarctic primnoid species" masterLastPageNumber="68" masterPageNumber="48" pageNumber="63" updateTime="1698249708632" updateUser="plazi">
<mods:mods id="B6CEBE91016E23F1FC59BE4B05D37D79" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9280AD781B271631E0DEF67EEA83F156">
<mods:title id="48805D83595A1CDC14F876D32E3FECF1">Redescription of Thouarella brucei Thomson and Ritchie, 1906 (Cnidaria: Octocorallia: Primnoidae) and description of two new Antarctic primnoid species</mods:title>
</mods:titleInfo>
<mods:name id="CB6BF59E79BAAB2034023D67478EFBD1" type="personal">
<mods:role id="C6E8EE3624C9F698F5919DB8FF787741">
<mods:roleTerm id="F4C2960369FC213860AD09FA6C32EAEC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14217FCD8BBA6D946B33477649CD2D4A">Zapata-Guardiola, Rebeca</mods:namePart>
</mods:name>
<mods:name id="92253E6CEB244011703F3679554436B6" type="personal">
<mods:role id="C4946CAA36CEA1F28DED327395D8981D">
<mods:roleTerm id="50695EC17AD8F55D07DD33466FAED089">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F75E03A354068BB3413944EEF201A716">López-González, Pablo J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="90D16C3F52D0D9EDBE409EF612711D08">text</mods:typeOfResource>
<mods:relatedItem id="639727778E8018CCE277A1716E3A8208" type="host">
<mods:titleInfo id="6F99217583E13913CE1F4160B958B2C9">
<mods:title id="7760FD8F9E16361BD280FAC43AED9099">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C2D2F2003384FB6F722B32BA95624E9D">
<mods:date id="A5D404FD54064447C74D106BA02047BD">2010</mods:date>
<mods:detail id="CE982B1180C81617779F37AF47DBAE7F" type="volume">
<mods:number id="40BD2EBD497BB97912AE7C9E41C78E56">2616</mods:number>
</mods:detail>
<mods:extent id="AB9579B3D5C45F2DD4639B82DD304425" unit="page">
<mods:start id="7B2BD0459518F1034B1C3CD06652C9F8">48</mods:start>
<mods:end id="9CE2CFD2AA6EE8D24DC211315C70F037">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="71D389492C6B69E503979DB1346DE4D2">journal article</mods:classification>
<mods:identifier id="D22FE10DFDF7437C7CBB649D80873729" type="DOI">10.5281/zenodo.197916</mods:identifier>
<mods:identifier id="564E75D3D663F53A7F9D73B81CEADDB8" type="GBIF-Dataset">5d407501-96a0-44cd-8a9c-47734417e6f9</mods:identifier>
<mods:identifier id="5349B0470B8D48B8BCDA04B4702AA0B9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="1A224592D3A995859036C930E5F3B355" type="Zenodo-Dep">197916</mods:identifier>
</mods:mods>
<treatment id="03A387A7BC22FFCCFF655F1CFE638D4D" ID-DOI="http://doi.org/10.5281/zenodo.6494511" ID-GBIF-Taxon="119396911" ID-Zenodo-Dep="6494511" LSID="urn:lsid:plazi:treatment:03A387A7BC22FFCCFF655F1CFE638D4D" httpUri="http://treatment.plazi.org/id/03A387A7BC22FFCCFF655F1CFE638D4D" lastPageId="18" lastPageNumber="66" pageId="15" pageNumber="63">
<subSubSection id="C310653ABC22FFD1FF655F1CFEF58F66" pageId="15" pageNumber="63" type="nomenclature">
<paragraph id="8BB536B1BC22FFD1FF655F1CFE518F00" blockId="15.[151,484,775,839]" box="[151,484,775,801]" pageId="15" pageNumber="63">
<heading id="D0FD81DDBC22FFD1FF655F1CFE518F00" bold="true" box="[151,484,775,801]" fontSize="11" level="1" pageId="15" pageNumber="63" reason="1">
<emphasis id="B97EEAA3BC22FFD1FF655F1CFE518F00" bold="true" box="[151,484,775,801]" pageId="15" pageNumber="63">
<taxonomicName id="4C0A4D32BC22FFD1FF655F1CFECA8F00" ID-CoL="866WH" box="[151,383,775,801]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="15" pageNumber="63" phylum="Cnidaria" rank="species" species="anatis" status="sp. nov.">
<emphasis id="B97EEAA3BC22FFD1FF655F1CFECA8F00" bold="true" box="[151,383,775,801]" italics="true" pageId="15" pageNumber="63">Tokoprymno anatis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57D8BC22FFD1FE755F1CFE518F00" box="[391,484,775,801]" pageId="15" pageNumber="63" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF655F36FEF58F66" blockId="15.[151,484,775,839]" box="[151,320,813,839]" pageId="15" pageNumber="63">
<figureCitation id="13312A34BC22FFD1FF655F36FEF58F66" box="[151,320,813,839]" captionStart-0="FIGURE 13" captionStart-1="FIGURE 14" captionStart-2="FIGURE 15" captionStart-3="FIGURE 16" captionStartId-0="16.[151,255,1504,1528]" captionStartId-1="17.[151,255,1915,1939]" captionStartId-2="18.[151,255,1928,1952]" captionStartId-3="19.[151,256,1314,1338]" captionTargetBox-0="[151,1436,194,1483]" captionTargetBox-1="[151,1436,306,1893]" captionTargetBox-2="[151,1436,421,1907]" captionTargetBox-3="[151,1436,194,1292]" captionTargetId-0="figure@16.[151,1436,194,1483]" captionTargetId-1="figure@17.[151,1436,306,1893]" captionTargetId-2="figure@18.[151,1436,421,1907]" captionTargetId-3="figure@19.[151,1436,194,1292]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionTargetPageId-2="18" captionTargetPageId-3="19" captionText-0="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." captionText-1="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." captionText-2="FIGURE 15. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, accessory opercular scale; B, opercular scales; C, marginal scales. inner surface view." captionText-3="FIGURE 16. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, body scales; B, coenenchymal scales. inner surface view." httpUri-0="https://zenodo.org/record/197928/files/figure.png" httpUri-1="https://zenodo.org/record/197929/files/figure.png" httpUri-2="https://zenodo.org/record/197930/files/figure.png" httpUri-3="https://zenodo.org/record/197931/files/figure.png" pageId="15" pageNumber="63">Figures 1316</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C310653ABC22FFD1FF655F6CFC508FFD" pageId="15" pageNumber="63" type="materials_examined">
<paragraph id="8BB536B1BC22FFD1FF655F6CFC508FFD" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
<emphasis id="B97EEAA3BC22FFD1FF655F6CFDBB8FB0" bold="true" box="[151,526,887,913]" pageId="15" pageNumber="63">
Material examined.
<typeStatus id="54B18813BC22FFD1FE605F6CFDBC8FB0" box="[402,521,887,913]" pageId="15" pageNumber="63" type="holotype">Holotype</typeStatus>
:
</emphasis>
<collectionCode id="ED1BAE74BC22FFD1FDE75F63FDE38FB3" box="[533,598,888,914]" httpUri="http://grbio.org/cool/kdnf-0379" name="Zoologisches Museum Hamburg" pageId="15" pageNumber="63">ZMH</collectionCode>
C11749, ANT XIX/3, stn PS61/046-08,
<geoCoordinate id="EE3E5076BC22FFD1FBC55F63FB0F8FB3" box="[1079,1210,888,914]" direction="south" orientation="latitude" pageId="15" pageNumber="63" precision="9" value="-60.6465">60°38.79'S</geoCoordinate>
,
<geoCoordinate id="EE3E5076BC22FFD1FB345F63FAE68FB3" box="[1222,1363,888,914]" direction="west" orientation="longitude" pageId="15" pageNumber="63" precision="9" value="-53.957">53°57.42'W</geoCoordinate>
, north east of Elephant Island,
<collectingCountry id="F31D7621BC22FFD1FE345F86FDFE8F96" box="[454,587,925,951]" name="Antarctica" pageId="15" pageNumber="63">Antarctica</collectingCountry>
,
<quantity id="4CF29B54BC22FFD1FDAA5F86FC868F96" box="[600,819,925,951]" metricMagnitude="3" metricUnit="m" metricValue="2.896" metricValueMax="2.8964" metricValueMin="2.8956" pageId="15" pageNumber="63" unit="m" value="2896.0" valueMax="2896.4" valueMin="2895.6">2895.62896.4 m</quantity>
depth,
<date id="FFB41071BC22FFD1FC635F86FBD78F96" box="[913,1122,925,951]" pageId="15" pageNumber="63" value="2002-02-02">2 February 2002</date>
, one colony, fragmented. Fragments of the
<typeStatus id="54B18813BC22FFD1FE975FD9FE7E8FFD" box="[357,459,962,988]" pageId="15" pageNumber="63" type="holotype">holotype</typeStatus>
have also been deposited in
<collectionCode id="ED1BAE74BC22FFD1FCEE5FD9FCC48FFD" LSID="urn:lsid:biocol.org:col:34871" box="[796,881,962,988]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="15" pageNumber="63">USNM</collectionCode>
1145316.
</paragraph>
</subSubSection>
<subSubSection id="C310653ABC22FFD1FF375FFCFBEE8BD3" pageId="15" pageNumber="63" type="description">
<paragraph id="8BB536B1BC22FFD1FF375FFCFCE48853" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
<emphasis id="B97EEAA3BC22FFD1FF375FFCFDA08820" bold="true" box="[197,533,999,1025]" pageId="15" pageNumber="63">
Description of the
<typeStatus id="54B18813BC22FFD1FE585FFCFDA08820" box="[426,533,999,1025]" pageId="15" pageNumber="63" type="holotype">holotype</typeStatus>
</emphasis>
. Colony bottlebrush (
<figureCitation id="13312A34BC22FFD1FCE15FF3FC338823" box="[787,902,1000,1026]" captionStart="FIGURE 13" captionStartId="16.[151,255,1504,1528]" captionTargetBox="[151,1436,194,1483]" captionTargetId="figure@16.[151,1436,194,1483]" captionTargetPageId="16" captionText="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="15" pageNumber="63">Figure 13</figureCitation>
A), fragmented in three main parts of 5, 7 and
<quantity id="4CF29B54BC22FFD1FF655816FF5F8806" box="[151,234,1037,1063]" metricMagnitude="-2" metricUnit="m" metricValue="9.5" pageId="15" pageNumber="63" unit="cm" value="9.5">9.5 cm</quantity>
in length,
<quantity id="4CF29B54BC22FFD1FE9D5816FE0E8806" box="[367,443,1037,1063]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="63" unit="cm" value="22.0">22 cm</quantity>
in total length and
<quantity id="4CF29B54BC22FFD1FD585816FCB98806" box="[682,780,1037,1063]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" pageId="15" pageNumber="63" unit="cm" value="10.5">10.5 cm</quantity>
in width. Simple or scarcely ramified stiff branchlets (
<figureCitation id="13312A34BC22FFD1FF6D5829FEA3886D" box="[159,278,1074,1100]" captionStart="FIGURE 13" captionStartId="16.[151,255,1504,1528]" captionTargetBox="[151,1436,194,1483]" captionTargetId="figure@16.[151,1436,194,1483]" captionTargetPageId="16" captionText="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="15" pageNumber="63">Figure 13</figureCitation>
B) up to
<quantity id="4CF29B54BC22FFD1FE715829FE62886D" box="[387,471,1074,1100]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="63" unit="cm" value="6.5">6.5 cm</quantity>
in length, proximally almost perpendicular to stem, then curving upward. Axis bronze, stiff, broken proximally. Basal axis diameter
<quantity id="4CF29B54BC22FFD1FCF75843FCF88853" box="[773,845,1112,1138]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="15" pageNumber="63" unit="mm" value="3.0">3 mm</quantity>
.
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF375866FCB2890D" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
Polyps perpendicular to branchlets (
<figureCitation id="13312A34BC22FFD1FD8C5866FD4288B6" box="[638,759,1149,1175]" captionStart="FIGURE 13" captionStartId="16.[151,255,1504,1528]" captionTargetBox="[151,1436,194,1483]" captionTargetId="figure@16.[151,1436,194,1483]" captionTargetPageId="16" captionText="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="15" pageNumber="63">Figure 13</figureCitation>
B, 14A), absent on main stem, singly or biserial placed (
<figureCitation id="13312A34BC22FFD1FF6D58B9FEA0889D" box="[159,277,1186,1212]" captionStart="FIGURE 14" captionStartId="17.[151,255,1915,1939]" captionTargetBox="[151,1436,306,1893]" captionTargetId="figure@17.[151,1436,306,1893]" captionTargetPageId="17" captionText="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="15" pageNumber="63">Figure 14</figureCitation>
A), 611 polyps per cm. Polyps (
<figureCitation id="13312A34BC22FFD1FD5358B9FCAD889D" box="[673,792,1186,1212]" captionStart="FIGURE 14" captionStartId="17.[151,255,1915,1939]" captionTargetBox="[151,1436,306,1893]" captionTargetId="figure@17.[151,1436,306,1893]" captionTargetPageId="17" captionText="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="15" pageNumber="63">Figure 14</figureCitation>
) relatively elongate, slightly clavate, up to
<quantity id="4CF29B54BC22FFD1FAD358B9FAC8889D" box="[1313,1405,1186,1212]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="15" pageNumber="63" unit="mm" value="2.4">2.4 mm</quantity>
in height and
<quantity id="4CF29B54BC22FFD1FED258D3FE6488C3" box="[288,465,1224,1250]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" metricValueMax="9.7" metricValueMin="6.1" pageId="15" pageNumber="63" unit="mm" value="0.79" valueMax="0.97" valueMin="0.61">0.610.97 mm</quantity>
in diameter, with a conical operculum. Polyp body with 8 longitudinal rows of scales somewhat disorganized, adaxial body scales smaller (
<figureCitation id="13312A34BC22FFD1FCAB58F6FC7B8926" box="[857,974,1261,1287]" captionStart="FIGURE 14" captionStartId="17.[151,255,1915,1939]" captionTargetBox="[151,1436,306,1893]" captionTargetId="figure@17.[151,1436,306,1893]" captionTargetPageId="17" captionText="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="15" pageNumber="63">Figure 14</figureCitation>
B), 45 transverse rows of scales in the abaxial aspect overlapping one another (
<figureCitation id="13312A34BC22FFD1FD9D5909FD51890D" box="[623,740,1298,1324]" captionStart="FIGURE 14" captionStartId="17.[151,255,1915,1939]" captionTargetBox="[151,1436,306,1893]" captionTargetId="figure@17.[151,1436,306,1893]" captionTargetPageId="17" captionText="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="15" pageNumber="63">Figure 14</figureCitation>
C).
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF375923FBFF89BD" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
Accessory opercular scales (
<figureCitation id="13312A34BC22FFD1FDEE5923FD268973" box="[540,659,1336,1362]" captionStart="FIGURE 15" captionStartId="18.[151,255,1928,1952]" captionTargetBox="[151,1436,421,1907]" captionTargetId="figure@18.[151,1436,421,1907]" captionTargetPageId="18" captionText="FIGURE 15. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, accessory opercular scale; B, opercular scales; C, marginal scales. inner surface view." httpUri="https://zenodo.org/record/197930/files/figure.png" pageId="15" pageNumber="63">Figure 15</figureCitation>
A), blade-shaped, variable in number from absent to two, small, about
<quantity id="4CF29B54BC22FFD1FF135946FEFE8956" box="[225,331,1373,1399]" metricMagnitude="-4" metricUnit="m" metricValue="4.9" pageId="15" pageNumber="63" unit="mm" value="0.49">0.49 mm</quantity>
in height and
<quantity id="4CF29B54BC22FFD1FE0A5946FDD78956" box="[504,610,1373,1399]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="15" pageNumber="63" unit="mm" value="0.12">0.12 mm</quantity>
in width. Proximal half of inner surface tuberculate, smooth distally, without keel. Margin finely serrated. Outer surface almost smooth, few granules.
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF3759B3FC0B8A13" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
Opercular scales (
<figureCitation id="13312A34BC22FFD1FE5359B3FDAC89E3" box="[417,537,1448,1474]" captionStart="FIGURE 15" captionStartId="18.[151,255,1928,1952]" captionTargetBox="[151,1436,421,1907]" captionTargetId="figure@18.[151,1436,421,1907]" captionTargetPageId="18" captionText="FIGURE 15. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, accessory opercular scale; B, opercular scales; C, marginal scales. inner surface view." httpUri="https://zenodo.org/record/197930/files/figure.png" pageId="15" pageNumber="63">Figure 15</figureCitation>
B) eight in number,
<quantity id="4CF29B54BC22FFD1FCE059B3FC7689E3" box="[786,963,1448,1474]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" metricValueMax="8.4" metricValueMin="5.8" pageId="15" pageNumber="63" unit="mm" value="0.71" valueMax="0.84" valueMin="0.58">0.580.84 mm</quantity>
in height and
<quantity id="4CF29B54BC22FFD1FB8759B3FA9289E3" box="[1141,1319,1448,1474]" metricMagnitude="-4" metricUnit="m" metricValue="2.15" metricValueMax="2.8" metricValueMin="1.5" pageId="15" pageNumber="63" unit="mm" value="0.215" valueMax="0.28" valueMin="0.15">0.150.28 mm</quantity>
in width, duck-beak shaped with rounded tips and square or bilobed base. Proximal inner surface tuberculate covering up to half of their length; distal part convex, with granules forming ridges or with a small keel. Outer surface almost smooth with a few granules proximally. Free margin serrated.
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF375A26FECF8AE6" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
Marginal scales (
<figureCitation id="13312A34BC22FFD1FE7F5A26FE4A8A76" box="[397,511,1597,1623]" captionStart="FIGURE 15" captionStartId="18.[151,255,1928,1952]" captionTargetBox="[151,1436,421,1907]" captionTargetId="figure@18.[151,1436,421,1907]" captionTargetPageId="18" captionText="FIGURE 15. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, accessory opercular scale; B, opercular scales; C, marginal scales. inner surface view." httpUri="https://zenodo.org/record/197930/files/figure.png" pageId="15" pageNumber="63">Figure 15</figureCitation>
C) eight in number, more-or-less triangular,
<quantity id="4CF29B54BC22FFD1FBF55A26FB078A76" box="[1031,1202,1597,1623]" metricMagnitude="-4" metricUnit="m" metricValue="7.15" metricValueMax="9.1" metricValueMin="5.2" pageId="15" pageNumber="63" unit="mm" value="0.715" valueMax="0.91" valueMin="0.52">0.520.91 mm</quantity>
in height and
<quantity id="4CF29B54BC22FFD1FAA85A26FF4A8A5D" metricMagnitude="-4" metricUnit="m" metricValue="3.55" metricValueMax="4.4" metricValueMin="2.7" pageId="15" pageNumber="63" unit="mm" value="0.355" valueMax="0.44" valueMin="0.27">0.27 0.44 mm</quantity>
in width. Inner surface tuberculate covering at least 75% of the scale, with distal longitudinal ridges. Outer surface almost smooth, with a few granules on proximal portion. Free margin serrated, proximal margin granular and lobed.
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF375AC9FB9F8B16" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
Body wall scales (
<figureCitation id="13312A34BC22FFD1FE6C5AC9FDA78ACD" box="[414,530,1746,1772]" captionStart="FIGURE 16" captionStartId="19.[151,256,1314,1338]" captionTargetBox="[151,1436,194,1292]" captionTargetId="figure@19.[151,1436,194,1292]" captionTargetPageId="19" captionText="FIGURE 16. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, body scales; B, coenenchymal scales. inner surface view." httpUri="https://zenodo.org/record/197931/files/figure.png" pageId="15" pageNumber="63">Figure 16</figureCitation>
A) roughly square shaped,
<quantity id="4CF29B54BC22FFD1FCA65AC9FBB58ACD" box="[852,1024,1746,1772]" metricMagnitude="-4" metricUnit="m" metricValue="4.85" metricValueMax="6.1" metricValueMin="3.5999999999999996" pageId="15" pageNumber="63" unit="mm" value="0.485" valueMax="0.61" valueMin="0.36">0.360.61 mm</quantity>
in maximum length. Inner surface almost completely tuberculate, outer surface almost smooth and covered with granules proximally. Free margin serrated with tendency to be lobed, basal margin as in marginal scales.
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF375B59FF4E8B86" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
Coenenchyme scales (
<figureCitation id="13312A34BC22FFD1FE245B59FDE58B7D" box="[470,592,1858,1884]" captionStart="FIGURE 16" captionStartId="19.[151,256,1314,1338]" captionTargetBox="[151,1436,194,1292]" captionTargetId="figure@19.[151,1436,194,1292]" captionTargetPageId="19" captionText="FIGURE 16. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, body scales; B, coenenchymal scales. inner surface view." httpUri="https://zenodo.org/record/197931/files/figure.png" pageId="15" pageNumber="63">Figure 16</figureCitation>
B) round in shape,
<quantity id="4CF29B54BC22FFD1FCCC5B59FC458B7D" box="[830,1008,1858,1884]" metricMagnitude="-4" metricUnit="m" metricValue="2.6" metricValueMax="3.4" metricValueMin="1.7999999999999998" pageId="15" pageNumber="63" unit="mm" value="0.26" valueMax="0.34" valueMin="0.18">0.180.34 mm</quantity>
in maximum length. Inner surface completely tuberculate, outer surface smooth with a few granules. Margin quite smooth, granulate or finely serrated.
</paragraph>
<paragraph id="8BB536B1BC22FFD1FF375BA9FBEE8BD3" blockId="15.[151,1437,887,2034]" pageId="15" pageNumber="63">
<emphasis id="B97EEAA3BC22FFD1FF375BA9FD668BED" bold="true" box="[197,723,1970,1996]" pageId="15" pageNumber="63">Geographic and bathymetric distribution.</emphasis>
At present,
<taxonomicName id="4C0A4D32BC22FFD1FC965BA8FBF08BED" box="[868,1093,1971,1996]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="15" pageNumber="63" phylum="Cnidaria" rank="species" species="anatis" status="sp. nov.">
<emphasis id="B97EEAA3BC22FFD1FC965BA8FBF08BED" box="[868,1093,1971,1996]" italics="true" pageId="15" pageNumber="63">Tokoprymno anatis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24D57D8BC22FFD1FBBF5BA9FB1E8BED" box="[1101,1195,1970,1996]" pageId="15" pageNumber="63" rank="species">
<emphasis id="B97EEAA3BC22FFD1FBBF5BA9FB1E8BED" bold="true" box="[1101,1195,1970,1996]" pageId="15" pageNumber="63">sp. nov.</emphasis>
</taxonomicNameLabel>
is only known from off Elephant Island,
<collectingCountry id="F31D7621BC22FFD1FE775BC3FE4A8BD3" box="[389,511,2008,2034]" name="Antarctica" pageId="15" pageNumber="63">Antarctica</collectingCountry>
(
<figureCitation id="13312A34BC22FFD1FDFC5BC3FDC48BD3" box="[526,625,2008,2034]" captionStart="FIGURE 1" captionStartId="2.[151,255,1225,1249]" captionTargetBox="[163,1416,208,1182]" captionTargetId="figure@2.[151,1436,194,1206]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of primnoid species studied in this paper: types of Thouarella brucei Thomson and Ritchie, 1906; new record of T. brucei; Digitogorgia brochi sp. nov.; ˔ Tokoprymno anatis sp. nov." httpUri="https://zenodo.org/record/197917/files/figure.png" pageId="15" pageNumber="63">Figure 1</figureCitation>
), between 2895.6 and
<quantity id="4CF29B54BC22FFD1FC8C5BC3FC588BD3" box="[894,1005,2008,2034]" metricMagnitude="3" metricUnit="m" metricValue="2.8964" pageId="15" pageNumber="63" unit="m" value="2896.4">2896.4 m</quantity>
in depth.
</paragraph>
</subSubSection>
<caption id="DF756639BC3DFFCEFF6559FBFA9B89D9" box="[151,1326,1504,1528]" httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="16" pageNumber="64" targetBox="[151,1436,194,1483]" targetPageId="16">
<paragraph id="8BB536B1BC3DFFCEFF6559FBFA9B89D9" blockId="16.[151,1326,1504,1528]" box="[151,1326,1504,1528]" pageId="16" pageNumber="64">
<emphasis id="B97EEAA3BC3DFFCEFF6559FBFE9389D9" bold="true" box="[151,294,1504,1528]" pageId="16" pageNumber="64">FIGURE 13.</emphasis>
<taxonomicName id="4C0A4D32BC3DFFCEFEDF59FAFE4C89D9" box="[301,505,1505,1528]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="species" species="anatis" status="sp. nov.">
<emphasis id="B97EEAA3BC3DFFCEFEDF59FAFE4C89D9" box="[301,505,1505,1528]" italics="true" pageId="16" pageNumber="64">Tokoprymno anatis</emphasis>
</taxonomicName>
<emphasis id="B97EEAA3BC3DFFCEFDF259FBFDE089D9" bold="true" box="[512,597,1504,1528]" pageId="16" pageNumber="64">
<taxonomicNameLabel id="A24D57D8BC3DFFCEFDF259FBFDE089D9" box="[512,597,1504,1528]" pageId="16" pageNumber="64" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (ZMH C11749):
<emphasis id="B97EEAA3BC3DFFCEFC8E59FBFC3A89D9" bold="true" box="[892,911,1504,1528]" pageId="16" pageNumber="64">A</emphasis>
, whole colony;
<emphasis id="B97EEAA3BC3DFFCEFBC859FBFBF989D9" bold="true" box="[1082,1100,1504,1528]" pageId="16" pageNumber="64">B</emphasis>
, detail of branchlets.
</paragraph>
</caption>
<subSubSection id="C310653ABC3DFFCEFF375A3DFC7F8A44" pageId="16" pageNumber="64" type="etymology">
<paragraph id="8BB536B1BC3DFFCEFF375A3DFC7F8A44" blockId="16.[151,1437,1574,2011]" pageId="16" pageNumber="64">
<emphasis id="B97EEAA3BC3DFFCEFF375A3DFEFF8A61" bold="true" box="[197,330,1574,1600]" pageId="16" pageNumber="64">Etymology</emphasis>
. The species name
<taxonomicName id="4C0A4D32BC3DFFCEFDC35A3DFDCC8A1E" box="[561,633,1574,1599]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="species" species="anatis">
<emphasis id="B97EEAA3BC3DFFCEFDC35A3DFDCC8A1E" box="[561,633,1574,1599]" italics="true" pageId="16" pageNumber="64">anatis</emphasis>
</taxonomicName>
, derived from the Latin and meaning duck-like, refers to the distinct shape of the opercular scales for their similarity to the beak of a duck.
</paragraph>
</subSubSection>
<subSubSection id="C310653ABC3DFFCCFF375A6BFE638D4D" lastPageId="18" lastPageNumber="66" pageId="16" pageNumber="64" type="discussion">
<paragraph id="8BB536B1BC3DFFCEFF375A6BFA988BB1" blockId="16.[151,1437,1574,2011]" pageId="16" pageNumber="64">
<emphasis id="B97EEAA3BC3DFFCEFF375A6BFE8A8AAB" bold="true" box="[197,319,1648,1674]" pageId="16" pageNumber="64">Remarks.</emphasis>
According to Cairns and Bayer's (2009) generic review of the gorgonian family
<taxonomicName id="4C0A4D32BC3DFFCEFAFE5A6AFA2D8AAA" box="[1292,1432,1649,1675]" class="Anthozoa" family="Primnoidae" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="family">Primnoidae</taxonomicName>
, the new taxon is included in the genus
<taxonomicName id="4C0A4D32BC3DFFCEFDAD5A8DFC338A91" authority="Bayer, 1996" authorityName="Bayer" authorityYear="1996" box="[607,902,1686,1712]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3DFFCEFDAD5A8DFD448A8E" box="[607,753,1686,1711]" italics="true" pageId="16" pageNumber="64">Tokoprymno</emphasis>
<bibRefCitation id="EF9B4B40BC3DFFCEFD055A8DFC338A91" author="Bayer" box="[759,902,1686,1712]" pageId="16" pageNumber="64" refString="Bayer, F. M. (1996) New Primnoid Gorgonians (Coelenterata: Octocorallia) from Antarctic waters. Bulletin of Marine Science, 58, 511 - 530." type="journal article" year="1996">Bayer, 1996</bibRefCitation>
</taxonomicName>
due to: (1) the presence of marginal scales of polyps not forming a circumoperculum (
<figureCitation id="13312A34BC3DFFCEFD825AA0FD508AF4" box="[624,741,1723,1749]" captionStart="FIGURE 14" captionStartId="17.[151,255,1915,1939]" captionTargetBox="[151,1436,306,1893]" captionTargetId="figure@17.[151,1436,306,1893]" captionTargetPageId="17" captionText="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="16" pageNumber="64">Figure 14</figureCitation>
), (2) the presence of an abundantly branched colony giving a bottlebrush shape (
<figureCitation id="13312A34BC3DFFCEFE7B5AFAFE498ADA" box="[393,508,1761,1787]" captionStart="FIGURE 13" captionStartId="16.[151,255,1504,1528]" captionTargetBox="[151,1436,194,1483]" captionTargetId="figure@16.[151,1436,194,1483]" captionTargetPageId="16" captionText="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="16" pageNumber="64">Figure 13</figureCitation>
A), (3) the fact that the polyps are often arranged biserially (
<figureCitation id="13312A34BC3DFFCEFB305AFAFA838ADA" box="[1218,1334,1761,1787]" captionStart="FIGURE 13" captionStartId="16.[151,255,1504,1528]" captionTargetBox="[151,1436,194,1483]" captionTargetId="figure@16.[151,1436,194,1483]" captionTargetPageId="16" captionText="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="16" pageNumber="64">Figure 13</figureCitation>
B, 14A), and (4) the opercular scales having a small keel on their inner surface. However, some species in the genus
<taxonomicName id="4C0A4D32BC3DFFCEFF655B37FEAE8B64" box="[151,283,1836,1861]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3DFFCEFF655B37FEAE8B64" box="[151,283,1836,1861]" italics="true" pageId="16" pageNumber="64">Thouarella</emphasis>
</taxonomicName>
also have a bottlebrush colony shape, polyps arranged singly, and the presence of a keel on the opercular scales. In
<emphasis id="B97EEAA3BC3DFFCEFE605B4AFD978B4B" box="[402,546,1873,1898]" italics="true" pageId="16" pageNumber="64">
<taxonomicName id="4C0A4D32BC3DFFCEFE605B4AFDA88B4B" box="[402,541,1873,1898]" class="Anthozoa" family="Primnoidae" genus="Thouarella" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="genus">Thouarella</taxonomicName>
,
</emphasis>
however, the marginals fold over the bases of the operculars forming a circumoperculum. In fact, this character is used to segregate both genera in Cairns and Bayers key.
</paragraph>
<paragraph id="8BB536B1BC3DFFCFFF375B80FDD18CDD" blockId="16.[151,1437,1574,2011]" lastBlockId="17.[151,1436,152,252]" lastPageId="17" lastPageNumber="65" pageId="16" pageNumber="64">
Also according to
<bibRefCitation id="EF9B4B40BC3DFFCEFE585B80FD558B94" author="Cairns" box="[426,736,1947,1973]" pageId="16" pageNumber="64" refString="Cairns, S. D. &amp; Bayer, F. M. (2009) A Generic Revision and Phylogenetic Analysis of the Primnoidae (Cnidaria: Octocorallia). Smithsonian Contributions to Zoology, 629, 1 - 79." type="journal article" year="2009">Cairns and Bayer (2009)</bibRefCitation>
, the genera
<taxonomicName id="4C0A4D32BC3DFFCEFC845B87FB248B94" authority="Gray, 1870" authorityName="Gray" authorityYear="1870" box="[886,1169,1947,1973]" class="Anthozoa" family="Primnoidae" genus="Plumarella" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3DFFCEFC845B87FC4A8B94" box="[886,1023,1948,1973]" italics="true" pageId="16" pageNumber="64">Plumarella</emphasis>
<bibRefCitation id="EF9B4B40BC3DFFCEFBFA5B80FB248B94" author="Gray" box="[1032,1169,1947,1973]" pageId="16" pageNumber="64" refString="Gray, J. E. (1870) Catalogue of the lithophytes or stony corals in the collection of the British Museum. British Museum, London, 51 pp." type="book" year="1870">Gray, 1870</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C0A4D32BC3DFFCEFB3C5B87FE7C8BFA" authority="Cairns and Bayer, 2004" authorityName="Cairns and Bayer" authorityYear="2004" class="Anthozoa" family="Primnoidae" genus="Acanthoprimnoa" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3DFFCEFB3C5B87FA298B94" box="[1230,1436,1948,1973]" italics="true" pageId="16" pageNumber="64">Acanthoprimnoa</emphasis>
<bibRefCitation id="EF9B4B40BC3DFFCEFF655BDAFE7C8BFA" author="Cairns" box="[151,457,1985,2011]" pageId="16" pageNumber="64" refString="Cairns, S. D. &amp; Bayer, F. M. (2004) Studies on western Atlantic Octocorallia (Coelenterata: Anthozoa). Part 5: The genera Plumarella Gray, 1870; Acanthoprimnoa, n. gen.; and Candidella Bayer, 1954. Proceedings of the Biological Society of Washington, 117, 447 - 487." type="journal article" year="2004">Cairns and Bayer, 2004</bibRefCitation>
</taxonomicName>
are morphologically closely related to the genus
<taxonomicName id="4C0A4D32BC3DFFCEFBBD5BDAFB588BFB" box="[1103,1261,1985,2010]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="16" pageNumber="64" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3DFFCEFBBD5BDAFB588BFB" box="[1103,1261,1985,2010]" italics="true" pageId="16" pageNumber="64">Tokoprymno</emphasis>
</taxonomicName>
. They can be distinguished, however, by the absence of a keel on the opercular scales, as well as their uniplanar colony shape (
<bibRefCitation id="EF9B4B40BC3CFFCFFF1B5CA6FEC58CF6" author="Gray" box="[233,368,189,215]" pageId="17" pageNumber="65" refString="Gray, J. E. (1870) Catalogue of the lithophytes or stony corals in the collection of the British Museum. British Museum, London, 51 pp." type="book" year="1870">Gray, 1870</bibRefCitation>
;
<bibRefCitation id="EF9B4B40BC3CFFCFFE895CA6FDCA8CF6" author="Cairns" box="[379,639,189,215]" pageId="17" pageNumber="65" refString="Cairns, S. D. &amp; Bayer, F. M. (2004) Studies on western Atlantic Octocorallia (Coelenterata: Anthozoa). Part 5: The genera Plumarella Gray, 1870; Acanthoprimnoa, n. gen.; and Candidella Bayer, 1954. Proceedings of the Biological Society of Washington, 117, 447 - 487." type="journal article" year="2004">Cairns &amp; Bayer, 2004</bibRefCitation>
), while
<taxonomicName id="4C0A4D32BC3CFFCFFD295CA6FCDC8CF7" box="[731,873,189,214]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="17" pageNumber="65" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3CFFCFFD295CA6FCDC8CF7" box="[731,873,189,214]" italics="true" pageId="17" pageNumber="65">Tokoprymno</emphasis>
</taxonomicName>
has a keel on the operculars (
<figureCitation id="13312A34BC3CFFCFFB345CA6FA8F8CF6" box="[1222,1338,189,215]" captionStart="FIGURE 14" captionStartId="17.[151,255,1915,1939]" captionTargetBox="[151,1436,306,1893]" captionTargetId="figure@17.[151,1436,306,1893]" captionTargetPageId="17" captionText="FIGURE 14. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, detail of a branchlet; B, polyp, adaxial view; C, polyp, abaxial view; D, polyp, lateral view." httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="17" pageNumber="65">Figure 14</figureCitation>
B) and a bottlebrush colony shape (
<figureCitation id="13312A34BC3CFFCFFE395CF9FD8B8CDD" box="[459,574,226,252]" captionStart="FIGURE 13" captionStartId="16.[151,255,1504,1528]" captionTargetBox="[151,1436,194,1483]" captionTargetId="figure@16.[151,1436,194,1483]" captionTargetPageId="16" captionText="FIGURE 13. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, whole colony; B, detail of branchlets." httpUri="https://zenodo.org/record/197928/files/figure.png" pageId="17" pageNumber="65">Figure 13</figureCitation>
A).
</paragraph>
<caption id="DF756639BC3CFFCFFF655B60FDD18B93" httpUri="https://zenodo.org/record/197929/files/figure.png" pageId="17" pageNumber="65" targetBox="[151,1436,306,1893]" targetPageId="17">
<paragraph id="8BB536B1BC3CFFCFFF655B60FDD18B93" blockId="17.[151,1436,1915,1971]" pageId="17" pageNumber="65">
<emphasis id="B97EEAA3BC3CFFCFFF655B60FE928BB2" bold="true" box="[151,295,1915,1939]" pageId="17" pageNumber="65">FIGURE 14.</emphasis>
<taxonomicName id="4C0A4D32BC3CFFCFFEDF5B60FE4F8BB3" box="[301,506,1915,1938]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="17" pageNumber="65" phylum="Cnidaria" rank="species" species="anatis" status="sp. nov.">
<emphasis id="B97EEAA3BC3CFFCFFEDF5B60FE4F8BB3" box="[301,506,1915,1938]" italics="true" pageId="17" pageNumber="65">Tokoprymno anatis</emphasis>
</taxonomicName>
<emphasis id="B97EEAA3BC3CFFCFFDF35B60FDE28BB2" bold="true" box="[513,599,1915,1939]" pageId="17" pageNumber="65">
<taxonomicNameLabel id="A24D57D8BC3CFFCFFDF35B60FDE28BB2" box="[513,599,1915,1939]" pageId="17" pageNumber="65" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype (ZMH C11749):
<emphasis id="B97EEAA3BC3CFFCFFC8C5B60FC248BB2" bold="true" box="[894,913,1915,1939]" pageId="17" pageNumber="65">A</emphasis>
, detail of a branchlet;
<emphasis id="B97EEAA3BC3CFFCFFB705B60FB218BB2" bold="true" box="[1154,1172,1915,1939]" pageId="17" pageNumber="65">B</emphasis>
, polyp, adaxial view;
<emphasis id="B97EEAA3BC3CFFCFFA705B60FA298BB2" bold="true" box="[1410,1436,1915,1939]" pageId="17" pageNumber="65">C,</emphasis>
polyp, abaxial view;
<emphasis id="B97EEAA3BC3CFFCFFE845B80FE3C8B92" bold="true" box="[374,393,1947,1971]" pageId="17" pageNumber="65">D</emphasis>
, polyp, lateral view.
</paragraph>
</caption>
<paragraph id="8BB536B1BC3FFFCCFF375C83FE638D4D" blockId="18.[151,1436,152,364]" pageId="18" pageNumber="66">
The genus
<taxonomicName id="4C0A4D32BC3FFFCCFEB85C83FE6A8C90" box="[330,479,152,177]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="66" phylum="Cnidaria" rank="genus">
<emphasis id="B97EEAA3BC3FFFCCFEB85C83FE6A8C90" box="[330,479,152,177]" italics="true" pageId="18" pageNumber="66">Tokoprymno</emphasis>
</taxonomicName>
has currently only one described species,
<taxonomicName id="4C0A4D32BC3FFFCCFC105C83FAEE8C93" authority="Bayer, 1996" authorityName="Bayer" authorityYear="1996" box="[994,1371,152,178]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="66" phylum="Cnidaria" rank="species" species="maia">
<emphasis id="B97EEAA3BC3FFFCCFC105C83FB098C90" box="[994,1212,152,177]" italics="true" pageId="18" pageNumber="66">Tokoprymno maia</emphasis>
<bibRefCitation id="EF9B4B40BC3FFFCCFB365C83FAEE8C93" author="Bayer" box="[1220,1371,152,178]" pageId="18" pageNumber="66" refString="Bayer, F. M. (1996) New Primnoid Gorgonians (Coelenterata: Octocorallia) from Antarctic waters. Bulletin of Marine Science, 58, 511 - 530." type="journal article" year="1996">Bayer, 1996</bibRefCitation>
</taxonomicName>
. This species is clearly differentiated from
<taxonomicName id="4C0A4D32BC3FFFCCFDBB5CA6FD188CF7" box="[585,685,189,214]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="66" phylum="Cnidaria" rank="species" species="anatis" status="sp. nov.">
<emphasis id="B97EEAA3BC3FFFCCFDBB5CA6FD188CF7" box="[585,685,189,214]" italics="true" pageId="18" pageNumber="66">T. anatis</emphasis>
</taxonomicName>
<emphasis id="B97EEAA3BC3FFFCCFD465CA6FCA48CF6" bold="true" box="[692,785,189,215]" pageId="18" pageNumber="66">
<taxonomicNameLabel id="A24D57D8BC3FFFCCFD465CA6FCA48CF6" box="[692,785,189,215]" pageId="18" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by the size of the polyps, from
<quantity id="4CF29B54BC3FFFCCFB775CA6FB4E8CF6" box="[1157,1275,189,215]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="18" pageNumber="66" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3 to 5 mm</quantity>
in the former (
<bibRefCitation id="EF9B4B40BC3FFFCCFF6D5CF9FED98CDD" author="Bayer" box="[159,364,226,252]" pageId="18" pageNumber="66" refString="Bayer, F. M. (1996) New Primnoid Gorgonians (Coelenterata: Octocorallia) from Antarctic waters. Bulletin of Marine Science, 58, 511 - 530." type="journal article" year="1996">Bayer, 1996: 514</bibRefCitation>
) depending on the kind of polyp (vegetative or brooder) and up to
<quantity id="4CF29B54BC3FFFCCFB895CF9FB608CDD" box="[1147,1237,226,252]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" pageId="18" pageNumber="66" unit="mm" value="2.4">2.4 mm</quantity>
in the latter; and by the distinct duck-beak shape of the opercular scales in
<taxonomicName id="4C0A4D32BC3FFFCCFCC95D13FC2A8D00" box="[827,927,264,289]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="66" phylum="Cnidaria" rank="species" species="anatis" status="sp. nov.">
<emphasis id="B97EEAA3BC3FFFCCFCC95D13FC2A8D00" box="[827,927,264,289]" italics="true" pageId="18" pageNumber="66">T. anatis</emphasis>
</taxonomicName>
<emphasis id="B97EEAA3BC3FFFCCFC545D1CFBB68D00" bold="true" box="[934,1027,263,289]" pageId="18" pageNumber="66">
<taxonomicNameLabel id="A24D57D8BC3FFFCCFC545D1CFBB68D00" box="[934,1027,263,289]" pageId="18" pageNumber="66" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="13312A34BC3FFFCCFBE15D13FB338D03" box="[1043,1158,264,290]" captionStart="FIGURE 15" captionStartId="18.[151,255,1928,1952]" captionTargetBox="[151,1436,421,1907]" captionTargetId="figure@18.[151,1436,421,1907]" captionTargetPageId="18" captionText="FIGURE 15. Tokoprymno anatis sp. nov., holotype (ZMH C 11749): A, accessory opercular scale; B, opercular scales; C, marginal scales. inner surface view." httpUri="https://zenodo.org/record/197930/files/figure.png" pageId="18" pageNumber="66">Figure 15</figureCitation>
A), while in
<taxonomicName id="4C0A4D32BC3FFFCCFAE85D13FADB8D00" box="[1306,1390,264,289]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="66" phylum="Cnidaria" rank="species" species="maia">
<emphasis id="B97EEAA3BC3FFFCCFAE85D13FADB8D00" box="[1306,1390,264,289]" italics="true" pageId="18" pageNumber="66">T. maia</emphasis>
</taxonomicName>
the operculars are isosceles triangle shaped (see
<figureCitation id="13312A34BC3FFFCCFD3C5D36FC8D8D66" box="[718,824,301,327]" captionStart="FIGURE 4" captionStartId="5.[151,255,750,774]" captionTargetBox="[151,1436,180,728]" captionTargetId="figure@5.[151,1436,180,728]" captionTargetPageId="5" captionText="FIGURE 4. Thouarella brucei Thomson and Ritchie, 1906, lectotype (NMS. Z. 1921.143.1298): polyp, oral view, stereo pair." httpUri="https://zenodo.org/record/197920/files/figure.png" pageId="18" pageNumber="66">
Figure
<quantity id="4CF29B54BC3FFFCCFCDA5D36FC8D8D66" box="[808,824,301,327]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="18" pageNumber="66" unit="in" value="4.0">4</quantity>
</figureCitation>
in
<bibRefCitation id="EF9B4B40BC3FFFCCFC905D36FC488D66" author="Bayer" box="[866,1021,301,327]" pageId="18" pageNumber="66" refString="Bayer, F. M. (1996) New Primnoid Gorgonians (Coelenterata: Octocorallia) from Antarctic waters. Bulletin of Marine Science, 58, 511 - 530." type="journal article" year="1996">Bayer, 1996</bibRefCitation>
) and for the absence of accesory opercular scales in
<taxonomicName id="4C0A4D32BC3FFFCCFE8A5D49FE7B8D4A" box="[376,462,338,363]" class="Anthozoa" family="Primnoidae" genus="Tokoprymno" kingdom="Animalia" order="Alcyonacea" pageId="18" pageNumber="66" phylum="Cnidaria" rank="species" species="maia">
<emphasis id="B97EEAA3BC3FFFCCFE8A5D49FE7B8D4A" box="[376,462,338,363]" italics="true" pageId="18" pageNumber="66">T. maia</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>