676 lines
80 KiB
XML
676 lines
80 KiB
XML
<document id="75092502C1BFA04D72560764A65C77E4" ID-CLB-Dataset="45386" ID-DOI="10.11646/zootaxa.4103.2.6" ID-GBIF-Dataset="7f2289fa-1b9a-48c0-bcbf-4d6e85770fa6" ID-ISSN="1175-5326" ID-Zenodo-Dep="262518" ID-ZooBank="6FE194B5-091E-4CBE-A105-7A72A7D445E5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460442941453" checkinUser="plazi" docAuthor="Campillo, Gustavo, Dávila-Galavíz, Luis Fernando, Flores-Villela, Oscar & Campbell, Jonathan A." docDate="2016" docId="03A387A43F4EFFBB2FB5FAB2FBD97117" docLanguage="en" docName="zootaxa.4103.2.6.pdf.imf" docOrigin="Zootaxa 4103 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhadinella dysmica Campillo, Dávila-Galavíz, Flores-Villela & Campbell, 2016, sp. nov." docType="treatment" docVersion="11" lastPageNumber="172" masterDocId="FF9AFFDC3F4FFFBC2F22FFB8FFEB7540" masterDocTitle="A new species of Rhadinella (Serpentes: Colubridae) from the Sierra Madre del Sur of Guerrero, Mexico" masterLastPageNumber="173" masterPageNumber="165" pageNumber="166" updateTime="1698333093312" updateUser="ExternalLinkService">
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<mods:mods id="16D27DAA2F1BEB373A541FE40134E95F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="08670E57E9D86CD8F8A1D0D6A4D7ECDC">
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<mods:title id="BBCDCD27A6E0CE49472AED3DEEEA1049">A new species of Rhadinella (Serpentes: Colubridae) from the Sierra Madre del Sur of Guerrero, Mexico</mods:title>
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</mods:titleInfo>
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<mods:name id="C84EBD08390734662969E81015C2C204" type="personal">
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<mods:role id="4FD6E0DC0F142CBF2384B3DC467621FC">
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<mods:roleTerm id="05C5AD26A9AFC691CF2EE84FC5228FAD">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="EF14FC12559A9CE869B55CA47863AA1B">Campillo, Gustavo</mods:namePart>
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</mods:name>
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<mods:name id="54E86FE7D9E259490C12920CA0541D71" type="personal">
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<mods:roleTerm id="6411E658368C2D978141FD992C3D7614">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="AD7FDE0A209838C8B810495FD17BB4AA">Dávila-Galavíz, Luis Fernando</mods:namePart>
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</mods:name>
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<mods:name id="40E4B7417A8B2B426D7AF9501AA5FC8C" type="personal">
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<mods:roleTerm id="2821595C66BBF45DAF9C7C3433CA1330">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C51A349B6DDE8EF8C85005D72B1BC461">Flores-Villela, Oscar</mods:namePart>
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</mods:name>
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<mods:name id="3D584AC4D7F831B2DA58562109272604" type="personal">
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<mods:role id="EDEBE89D6CD7DF097E6F1958933711BB">
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<mods:roleTerm id="C7634EB2A012191BC5EA8680266C292E">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="71492E844E19C4FC759E34B941F493F0">Campbell, Jonathan A.</mods:namePart>
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<mods:typeOfResource id="CEBFCA7AC6B96EB34CECE27AB5B1DC92">text</mods:typeOfResource>
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<mods:title id="2EDE12C77F5017369BEDD32AA1D126CE">Zootaxa</mods:title>
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<mods:part id="CA51FA1E20ECE7B7E5F93885D20C0FC2">
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<mods:date id="EC3466047C54662169953EFDF8879BD3">2016</mods:date>
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<mods:detail id="863D51B60CBCF214FB872EC5C1C2464A" type="volume">
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<mods:number id="A534DCBB0C89903E38B943FA55DD54BA">4103</mods:number>
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<mods:detail id="885D0D70F121272CAABED208277D15B1" type="issue">
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<mods:number id="934089488A6DEF699A55B7FDA70E4A63">2</mods:number>
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<mods:classification id="13EEB25D899933C2D14CC1A3D1B719BD">journal article</mods:classification>
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<mods:identifier id="512B7D4600CB13D94D04FDF9F5FA5E97" type="CLB-Dataset">45386</mods:identifier>
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<mods:identifier id="37DAD429F1246397FAAF6E6C80C32A3B" type="DOI">10.11646/zootaxa.4103.2.6</mods:identifier>
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<mods:identifier id="7C9102CF3538B6B7ACC07F723FB1AAC3" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A387A43F4EFFBB2FB5FAB2FBD97117" ID-DOI="http://doi.org/10.5281/zenodo.5682458" ID-GBIF-Taxon="119508879" ID-Zenodo-Dep="5682458" LSID="urn:lsid:plazi:treatment:03A387A43F4EFFBB2FB5FAB2FBD97117" httpUri="http://treatment.plazi.org/id/03A387A43F4EFFBB2FB5FAB2FBD97117" lastPageId="7" lastPageNumber="172" pageId="1" pageNumber="166">
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<subSubSection id="C31065393F4EFFBD2FB5FAB2FE8B7006" pageId="1" pageNumber="166" type="nomenclature">
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<paragraph id="8BB536B23F4EFFBD2FB5FAB2FE017065" blockId="1.[151,490,1290,1350]" box="[151,490,1290,1317]" pageId="1" pageNumber="166">
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<heading id="D0FD81DE3F4EFFBD2FB5FAB2FE017065" bold="true" box="[151,490,1290,1317]" fontSize="11" level="1" pageId="1" pageNumber="166" reason="1">
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<emphasis id="B97EEAA03F4EFFBD2FB5FAB2FE017065" bold="true" box="[151,490,1290,1317]" pageId="1" pageNumber="166">
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<taxonomicName id="4C0A4D313F4EFFBD2FB5FAB2FE6D7064" box="[151,390,1290,1316]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="dysmica" status="sp. nov.">
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<emphasis id="B97EEAA03F4EFFBD2FB5FAB2FE6D7064" bold="true" box="[151,390,1290,1316]" italics="true" pageId="1" pageNumber="166">Rhadinella dysmica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24D57DB3F4EFFBD2EAEFAB3FE017065" box="[396,490,1291,1317]" pageId="1" pageNumber="166" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="8BB536B23F4EFFBD2FB5FA95FE8B7006" blockId="1.[151,490,1290,1350]" box="[151,352,1325,1350]" pageId="1" pageNumber="166">
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<figureCitation id="13312A373F4EFFBD2FB5FA95FEEA7006" box="[151,257,1325,1350]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,866,888]" captionStartId-1="3.[151,250,1813,1835]" captionTargetBox-0="[176,1408,196,846]" captionTargetBox-1="[154,1348,269,1732]" captionTargetId-0="figure@2.[174,1410,194,849]" captionTargetId-1="figure@3.[151,1435,193,1810]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Dorsal view in life of the holotype of Rhadinella dysmica (ENCB 18951) from near Cueva de Tepozonales, Cuajilotla, 20 km S Mochitlán, Guerrero, Mexico. Snout-vent length 256 mm, total length 341 + N mm." captionText-1="FIGURE 2. Dorsal (A), lateral (B), and ventral (C) views of head of the holotype of Rhadinella dysmica (ENCB 18951). Head length 9.5 mm. Photos by N. V. López Valentín." httpUri-0="https://zenodo.org/record/262519/files/figure.png" httpUri-1="https://zenodo.org/record/262520/files/figure.png" pageId="1" pageNumber="166">Figs. 1–2</figureCitation>
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,
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<tableCitation id="C68803093F4EFFBD2E2FFA95FE8B7006" box="[269,352,1325,1350]" captionStart="TABLE 1" captionStartId="6.[151,239,656,679]" captionTargetBox="[151,1411,707,1381]" captionTargetPageId="6" captionText="TABLE 1. Comparison of selected features from Mexican species of Rhadinella with dark-colored dorsum." httpUri="http://table.plazi.org/id/DF75663A3F49FFBA2FB5FD28FAFE77E7" pageId="1" pageNumber="166" tableUuid="DF75663A3F49FFBA2FB5FD28FAFE77E7">Table 1</tableCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31065393F4EFFBD2FB5FACDFBF37096" pageId="1" pageNumber="166" type="materials_examined">
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<paragraph id="8BB536B23F4EFFBD2FB5FACDFBF37096" blockId="1.[151,1437,1397,2034]" pageId="1" pageNumber="166">
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<emphasis id="B97EEAA03F4EFFBD2FB5FACDFEE070CE" bold="true" box="[151,267,1397,1422]" pageId="1" pageNumber="166">
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<typeStatus id="54B188103F4EFFBD2FB5FACDFEED70CE" box="[151,262,1397,1422]" pageId="1" pageNumber="166" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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Adult female,
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<collectionCode id="ED1BAE773F4EFFBD2E97FACEFDEB70CE" box="[437,512,1398,1422]" httpUri="http://grbio.org/cool/1x45-fdpe" name="Universidad de Autonoma de Baja California" pageId="1" pageNumber="166">ENCB</collectionCode>
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18951, obtained
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<date id="FFB410723F4EFFBD2DE6FACEFCBD70CE" box="[708,854,1397,1422]" pageId="1" pageNumber="166" value="2014-07-17">17 July 2014</date>
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by L. F. Dávila Galavíz near cueva de Tepozonales (
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<geoCoordinate id="EE3E50753F4EFFBD2FBCFA21FEF170F1" box="[158,282,1433,1458]" direction="north" orientation="latitude" pageId="1" pageNumber="166" precision="5" value="17.2853">17.2853 N</geoCoordinate>
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, –
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||
<geoCoordinate id="EE3E50753F4EFFBD2E11FA21FE5970F1" box="[307,434,1433,1458]" direction="west" orientation="longitude" pageId="1" pageNumber="166" precision="5" value="-99.3662">99.3662 W</geoCoordinate>
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;
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<quantity id="4CF29B573F4EFFBD2E9CFA22FDEE70F1" box="[446,517,1434,1458]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="1" pageNumber="166" unit="m" value="432.0">432 m</quantity>
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above sea level, asl hereafter), Cuajilotla,
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<quantity id="4CF29B573F4EFFBD2CC7FA22FBC670F2" box="[997,1069,1433,1458]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="166" unit="km" value="20.0">20 km</quantity>
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S Mochitlán, Guerrero,
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||
<collectingCountry id="F31D76223F4EFFBD2A62FA21FA7270F2" box="[1344,1433,1433,1458]" name="Mexico" pageId="1" pageNumber="166">Mexico</collectingCountry>
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. The individual was found near the entrance of the cave, crawling on the ground.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C31065393F4EFFBE2FE5FA59FC4B70EA" lastPageId="2" lastPageNumber="167" pageId="1" pageNumber="166" type="diagnosis">
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||
<paragraph id="8BB536B23F4EFFBD2FE5FA59FEFE727E" blockId="1.[151,1437,1397,2034]" pageId="1" pageNumber="166">
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||
<emphasis id="B97EEAA03F4EFFBD2FE5FA59FED170BA" bold="true" box="[199,314,1505,1530]" pageId="1" pageNumber="166">Diagnosis</emphasis>
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. A member of
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<taxonomicName id="4C0A4D313F4EFFBD2ED0FA59FD8670B9" box="[498,621,1505,1529]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="genus">
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<emphasis id="B97EEAA03F4EFFBD2ED0FA59FD8670B9" box="[498,621,1505,1529]" italics="true" pageId="1" pageNumber="166">Rhadinella</emphasis>
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</taxonomicName>
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, genus characterized by gracile body, head only slightly distinct from neck; inverted dark U-shaped marking on rostral; anterior supralabials usually with dark edges and pale centers; conspicuous or at least evidence of pale collar on neck; with reduced to small supracloacal tubercles in adult females; no subpreocular scale; posterior (2–7) teeth larger and thicker than preceding teeth, all enlarged teeth on same plane, no broad diastema; with basal nude pocket on hemipenis.
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<taxonomicName id="4C0A4D313F4EFFBD2C90F9C9FB7A73C9" box="[946,1169,1649,1673]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="dysmica">
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<emphasis id="B97EEAA03F4EFFBD2C90F9C9FB7A73C9" box="[946,1169,1649,1673]" italics="true" pageId="1" pageNumber="166">Rhadinella dysmica</emphasis>
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</taxonomicName>
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is further characterized by the combination of a distinctive white collar, undivided at middorsum, extending 2.0–2.5 scale lengths at middorsum and 3–4 scale lengths on side of head; temporals 1 + 2; smooth dorsal scale rows 17, unreduced throughout body; ventrals 172; cloacal scute divided; divided subcaudals incomplete 63+N; dorsal coloration dark brown to blackish, almost totally obfuscating longitudinal striping, ground color barely extending to lateral edges of ventrals.
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</paragraph>
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<paragraph id="8BB536B23F4EFFBD2FE5F8F1FBF072B2" blockId="1.[151,1437,1397,2034]" pageId="1" pageNumber="166">
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<taxonomicName id="4C0A4D313F4EFFBD2FE5F8F1FE457221" box="[199,430,1865,1889]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="dysmica">
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<emphasis id="B97EEAA03F4EFFBD2FE5F8F1FE457221" box="[199,430,1865,1889]" italics="true" pageId="1" pageNumber="166">Rhadinella dysmica</emphasis>
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</taxonomicName>
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may be distinguished from many congeners (i.e.,
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<taxonomicName id="4C0A4D313F4EFFBD2B0EF8F2FB5C7221" box="[1068,1207,1865,1889]" class="Reptilia" family="Colubridae" genus="Rhadinaea" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="godmani">
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<emphasis id="B97EEAA03F4EFFBD2B0EF8F2FB5C7221" box="[1068,1207,1865,1889]" italics="true" pageId="1" pageNumber="166">R. godmani</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2BEDF8F2FA877221" box="[1231,1388,1865,1889]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="hannsteini">
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<emphasis id="B97EEAA03F4EFFBD2BEDF8F2FA877221" box="[1231,1388,1865,1889]" italics="true" pageId="1" pageNumber="166">R. hannsteini</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2AA6F8F2FEC272C5" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="hempsteadae">
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<emphasis id="B97EEAA03F4EFFBD2AA6F8F2FEC272C5" italics="true" pageId="1" pageNumber="166">R. hempsteadae</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2E18F8D6FE1172C5" box="[314,506,1901,1925]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="kanalchuchan">
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||
<emphasis id="B97EEAA03F4EFFBD2E18F8D6FE1172C5" box="[314,506,1901,1925]" italics="true" pageId="1" pageNumber="166">R. kanalchuchan</emphasis>
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||
</taxonomicName>
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||
,
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||
<taxonomicName id="4C0A4D313F4EFFBD2D29F8D6FD6572C5" box="[523,654,1901,1925]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="kinkelini">
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||
<emphasis id="B97EEAA03F4EFFBD2D29F8D6FD6572C5" box="[523,654,1901,1925]" italics="true" pageId="1" pageNumber="166">R. kinkelini</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2DBDF8D6FCAE72C5" box="[671,837,1901,1925]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="lachrymans">
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||
<emphasis id="B97EEAA03F4EFFBD2DBDF8D6FCAE72C5" box="[671,837,1901,1925]" italics="true" pageId="1" pageNumber="166">R. lachrymans</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2C74F8D6FBEF72C5" box="[854,1028,1902,1925]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="montecristoi">
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||
<emphasis id="B97EEAA03F4EFFBD2C74F8D6FBEF72C5" box="[854,1028,1902,1925]" italics="true" pageId="1" pageNumber="166">R. montecristoi</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2B34F8D6FB5A72C5" box="[1046,1201,1901,1925]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="pegosalyta">
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<emphasis id="B97EEAA03F4EFFBD2B34F8D6FB5A72C5" box="[1046,1201,1901,1925]" italics="true" pageId="1" pageNumber="166">R. pegosalyta</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2BE1F8D6FA9972C5" box="[1219,1394,1902,1925]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="rogerromani">
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<emphasis id="B97EEAA03F4EFFBD2BE1F8D6FA9972C5" box="[1219,1394,1902,1925]" italics="true" pageId="1" pageNumber="166">R. rogerromani</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C0A4D313F4EFFBD2AA6F8D6FEFE72E9" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="serperaster">
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||
<emphasis id="B97EEAA03F4EFFBD2AA6F8D6FEFE72E9" italics="true" pageId="1" pageNumber="166">R. serperaster</emphasis>
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</taxonomicName>
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, and
|
||
<taxonomicName id="4C0A4D313F4EFFBD2E71F82AFE1D72E9" box="[339,502,1937,1961]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="tolpanorum">
|
||
<emphasis id="B97EEAA03F4EFFBD2E71F82AFE1D72E9" box="[339,502,1937,1961]" italics="true" pageId="1" pageNumber="166">R. tolpanorum</emphasis>
|
||
</taxonomicName>
|
||
) by dark dorsal body coloration and absence of conspicuous striping. Additionally, the new species differs from
|
||
<taxonomicName id="4C0A4D313F4EFFBD2EF9F80EFDB7728D" box="[475,604,1973,1997]" class="Reptilia" family="Colubridae" genus="Rhadinaea" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="godmani">
|
||
<emphasis id="B97EEAA03F4EFFBD2EF9F80EFDB7728D" box="[475,604,1973,1997]" italics="true" pageId="1" pageNumber="166">R. godmani</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4EFFBD2D48F80EFCF7728D" box="[618,796,1973,1997]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="hempsteadae">
|
||
<emphasis id="B97EEAA03F4EFFBD2D48F80EFCF7728D" box="[618,796,1973,1997]" italics="true" pageId="1" pageNumber="166">R. hempsteadae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4EFFBD2C08F80EFC0D728D" box="[810,998,1973,1997]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="kanalchuchan">
|
||
<emphasis id="B97EEAA03F4EFFBD2C08F80EFC0D728D" box="[810,998,1973,1997]" italics="true" pageId="1" pageNumber="166">R. kanalchuchan</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4EFFBD2CD6F80EFB4B728D" box="[1012,1184,1974,1997]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="montecristoi">
|
||
<emphasis id="B97EEAA03F4EFFBD2CD6F80EFB4B728D" box="[1012,1184,1974,1997]" italics="true" pageId="1" pageNumber="166">R. montecristoi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4EFFBD2B8FF80EFAAD728D" box="[1197,1350,1973,1997]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="pegosalyta">
|
||
<emphasis id="B97EEAA03F4EFFBD2B8FF80EFAAD728D" box="[1197,1350,1973,1997]" italics="true" pageId="1" pageNumber="166">R. pegosalyta</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0A4D313F4EFFBD2AA6F80EFEFC72B1" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="166" phylum="Chordata" rank="species" species="serperaster">
|
||
<emphasis id="B97EEAA03F4EFFBD2AA6F80EFEFC72B1" italics="true" pageId="1" pageNumber="166">R. serperaster</emphasis>
|
||
</taxonomicName>
|
||
by having 17 dorsal scale rows (vs. more than 17 dorsal scale rows).
|
||
</paragraph>
|
||
<caption id="DF75663A3F4DFFBE2FB5FCDAFB4376D6" httpUri="https://zenodo.org/record/262519/files/figure.png" pageId="2" pageNumber="167" targetBox="[176,1408,196,846]" targetPageId="2">
|
||
<paragraph id="8BB536B23F4DFFBE2FB5FCDAFB4376D6" blockId="2.[151,1436,866,918]" pageId="2" pageNumber="167">
|
||
<emphasis id="B97EEAA03F4DFFBE2FB5FCDAFEFD7637" bold="true" box="[151,278,866,888]" pageId="2" pageNumber="167">FIGURE 1.</emphasis>
|
||
Dorsal view in life of the holotype of
|
||
<taxonomicName id="4C0A4D313F4DFFBE2D9EFCDAFC637637" box="[700,904,866,887]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4DFFBE2D9EFCDAFC637637" box="[700,904,866,887]" italics="true" pageId="2" pageNumber="167">Rhadinella dysmica</emphasis>
|
||
</taxonomicName>
|
||
(ENCB 18951) from near Cueva de Tepozonales, Cuajilotla, 20 km S Mochitlán, Guerrero, Mexico. Snout-vent length 256 mm, total length 341+N mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536B23F4DFFBE2FE5FC06FC4B70EA" blockId="2.[151,1437,957,2026]" pageId="2" pageNumber="167">
|
||
<taxonomicName id="4C0A4D313F4DFFBE2FE5FC06FE4F7696" box="[199,420,958,982]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4DFFBE2FE5FC06FE4F7696" box="[199,420,958,982]" italics="true" pageId="2" pageNumber="167">Rhadinella dysmica</emphasis>
|
||
</taxonomicName>
|
||
is morphologically similar to other species having mostly uniformly blackish dorsum, with or without indistinct dark vertebral line and pale dashes on some dorsal scales (i.e.,
|
||
<taxonomicName id="4C0A4D313F4DFFBE2B4CFC5AFB1C76BA" box="[1134,1271,994,1018]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F4DFFBE2B4CFC5AFB1C76BA" box="[1134,1271,994,1018]" italics="true" pageId="2" pageNumber="167">R. schistosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4DFFBE2A2AFC5AFA9976BA" box="[1288,1394,994,1018]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F4DFFBE2A2AFC5AFA9976BA" box="[1288,1394,994,1018]" italics="true" pageId="2" pageNumber="167">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4DFFBE2AA6FC5AFEF1715E" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="pilonaorum">
|
||
<emphasis id="B97EEAA03F4DFFBE2AA6FC5AFEF1715E" italics="true" pageId="2" pageNumber="167">R. pilonaorum</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0A4D313F4DFFBE2E7EFBBFFE09715E" box="[348,482,1030,1054]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="posadasi">
|
||
<emphasis id="B97EEAA03F4DFFBE2E7EFBBFFE09715E" box="[348,482,1030,1054]" italics="true" pageId="2" pageNumber="167">R. posadasi</emphasis>
|
||
</taxonomicName>
|
||
). In
|
||
<taxonomicName id="4C0A4D313F4DFFBE2D3FFBBFFD70715E" box="[541,667,1030,1054]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4DFFBE2D3FFBBFFD70715E" box="[541,667,1030,1054]" italics="true" pageId="2" pageNumber="167">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
the color and configuration of the collar alone diagnostic from all other species of dark
|
||
<taxonomicName id="4C0A4D313F4DFFBE2EAFFB92FDE37102" box="[397,520,1066,1090]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F4DFFBE2EAFFB92FDE37102" box="[397,520,1066,1090]" italics="true" pageId="2" pageNumber="167">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
, extending from about posterior tips of parietals for several scale lengths, being white, and distinctly set off from mostly black head-cap. In
|
||
<taxonomicName id="4C0A4D313F4DFFBE2C4EFBF7FC077126" box="[876,1004,1102,1126]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="shistosa">
|
||
<emphasis id="B97EEAA03F4DFFBE2C4EFBF7FC077126" box="[876,1004,1102,1126]" italics="true" pageId="2" pageNumber="167">R. shistosa</emphasis>
|
||
</taxonomicName>
|
||
collar U-shaped and usually broken medially (
|
||
<figureCitation id="13312A373F4DFFBE2E2AFBC9FEA771CA" box="[264,332,1137,1162]" captionStart="FIGURE 3" captionStartId="4.[151,250,1929,1951]" captionTargetBox="[157,1395,739,1874]" captionTargetId="figure@4.[152,1435,697,1908]" captionTargetPageId="4" captionText="FIGURE 3. Dorsal (A) and (B) lateral aspects of Rhadinella schistosa (FMNH 100297, paratype) from Cuautlapan, Veracruz, Mexico, showing incomplete collar across nape. Head length 6.5 mm." httpUri="https://zenodo.org/record/262521/files/figure.png" pageId="2" pageNumber="167">Fig. 3</figureCitation>
|
||
); in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2E5DFBCAFE0E71CA" box="[383,485,1138,1162]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F4DFFBE2E5DFBCAFE0E71CA" box="[383,485,1138,1162]" italics="true" pageId="2" pageNumber="167">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
white coloration of collar extends on to head, involving most of parietals and part of frontal; in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2E2EFB2FFE4571EE" box="[268,430,1174,1198]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="pilonaorum">
|
||
<emphasis id="B97EEAA03F4DFFBE2E2EFB2FFE4571EE" box="[268,430,1174,1198]" italics="true" pageId="2" pageNumber="167">R. pilonaorum</emphasis>
|
||
</taxonomicName>
|
||
collar orange with top of head mostly auburn mottled with black; in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2B89FB2FFAC671EE" box="[1195,1325,1174,1198]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="posadasi">
|
||
<emphasis id="B97EEAA03F4DFFBE2B89FB2FFAC671EE" box="[1195,1325,1174,1198]" italics="true" pageId="2" pageNumber="167">R. posadasi</emphasis>
|
||
</taxonomicName>
|
||
the collar is dark orange or auburn. Other features distinguishing these dark-colored species of
|
||
<taxonomicName id="4C0A4D313F4DFFBE2B58FB02FB1E7192" box="[1146,1269,1210,1234]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F4DFFBE2B58FB02FB1E7192" box="[1146,1269,1210,1234]" italics="true" pageId="2" pageNumber="167">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
include fewer ventrals (
|
||
<quantity id="4CF29B573F4DFFBE2E23FB65FE6F71B6" box="[257,388,1245,1270]" metricMagnitude="0" metricUnit="m" metricValue="3.8989" metricValueMax="3.9624" metricValueMin="3.8354" pageId="2" pageNumber="167" unit="in" value="153.5" valueMax="156.0" valueMin="151.0">151–156 in</quantity>
|
||
females), fewer subcaudals (
|
||
<quantity id="4CF29B573F4DFFBE2DF7FB66FCD771B6" box="[725,828,1245,1270]" metricMagnitude="-1" metricUnit="m" metricValue="8.254999999999999" metricValueMax="8.636" metricValueMin="7.874" pageId="2" pageNumber="167" unit="in" value="32.5" valueMax="34.0" valueMin="31.0">31–34 in</quantity>
|
||
females), and 1+1 temporals in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2B97FB66FA7F71B6" box="[1205,1428,1246,1270]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="shistosa">
|
||
<emphasis id="B97EEAA03F4DFFBE2B97FB66FA7F71B6" box="[1205,1428,1246,1270]" italics="true" pageId="2" pageNumber="167">Rhadinella shistosa</emphasis>
|
||
</taxonomicName>
|
||
; more subcaudals (
|
||
<quantity id="4CF29B573F4DFFBE2E49FABAFE04705A" box="[363,495,1281,1306]" metricMagnitude="0" metricUnit="m" metricValue="2.6797" metricValueMax="2.8194" metricValueMin="2.54" pageId="2" pageNumber="167" unit="in" value="105.5" valueMax="111.0" valueMin="100.0">100–111 in</quantity>
|
||
females) in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2DABFABAFCDB705A" box="[649,816,1282,1306]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="pilonaorum">
|
||
<emphasis id="B97EEAA03F4DFFBE2DABFABAFCDB705A" box="[649,816,1282,1306]" italics="true" pageId="2" pageNumber="167">R. pilonaorum</emphasis>
|
||
</taxonomicName>
|
||
; and fewer ventrals (
|
||
<quantity id="4CF29B573F4DFFBE2B12FAB9FB5D705A" box="[1072,1206,1281,1306]" metricMagnitude="0" metricUnit="m" metricValue="3.6703" metricValueMax="3.7084" metricValueMin="3.6322" pageId="2" pageNumber="167" unit="in" value="144.5" valueMax="146.0" valueMin="143.0">143–146 in</quantity>
|
||
females) and more subcaudals (
|
||
<quantity id="4CF29B573F4DFFBE2E01FA9DFE62707E" box="[291,393,1317,1342]" metricMagnitude="0" metricUnit="m" metricValue="2.2733" metricValueMax="2.3622" metricValueMin="2.1844" pageId="2" pageNumber="167" unit="in" value="89.5" valueMax="93.0" valueMin="86.0">86–93 in</quantity>
|
||
females) in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2D3BFA9FFD75707E" box="[537,670,1318,1342]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="posadasi">
|
||
<emphasis id="B97EEAA03F4DFFBE2D3BFA9FFD75707E" box="[537,670,1318,1342]" italics="true" pageId="2" pageNumber="167">R. posadasi</emphasis>
|
||
</taxonomicName>
|
||
. Fourth and fifth infralabials contact posterior pair of genials in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2AA6FA9FFF157022" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F4DFFBE2AA6FA9FFF157022" italics="true" pageId="2" pageNumber="167">R. schistosa</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C0A4D313F4DFFBE2E2FFAF2FE5B7022" box="[269,432,1354,1378]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="pilonaorum">
|
||
<emphasis id="B97EEAA03F4DFFBE2E2FFAF2FE5B7022" box="[269,432,1354,1378]" italics="true" pageId="2" pageNumber="167">R. pilonaorum</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C0A4D313F4DFFBE2ED2FAF2FD9F7022" box="[496,628,1354,1378]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="posadasi">
|
||
<emphasis id="B97EEAA03F4DFFBE2ED2FAF2FD9F7022" box="[496,628,1354,1378]" italics="true" pageId="2" pageNumber="167">R. posadasi</emphasis>
|
||
</taxonomicName>
|
||
(vs. the third and fourth in
|
||
<taxonomicName id="4C0A4D313F4DFFBE2C92FAF2FBC67022" box="[944,1069,1354,1378]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4DFFBE2C92FAF2FBC67022" box="[944,1069,1354,1378]" italics="true" pageId="2" pageNumber="167">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D313F4DFFBE2B47FAF2FB267022" box="[1125,1229,1354,1378]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F4DFFBE2B47FAF2FB267022" box="[1125,1229,1354,1378]" italics="true" pageId="2" pageNumber="167">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
); distal portion of tails incomplete in
|
||
<typeStatus id="54B188103F4DFFBE2E55FAD5FE0F70C6" box="[375,484,1389,1414]" pageId="2" pageNumber="167" type="holotype">holotypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C0A4D313F4DFFBE2D32FAD7FD6570C6" box="[528,654,1390,1414]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4DFFBE2D32FAD7FD6570C6" box="[528,654,1390,1414]" italics="true" pageId="2" pageNumber="167">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
(a female) and
|
||
<taxonomicName id="4C0A4D313F4DFFBE2C69FAD7FC5E70C6" box="[843,949,1390,1414]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F4DFFBE2C69FAD7FC5E70C6" box="[843,949,1390,1414]" italics="true" pageId="2" pageNumber="167">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
(a male), however with 66 subcaudals
|
||
<taxonomicName id="4C0A4D313F4DFFBE2AA6FAD7FF1870EA" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4DFFBE2AA6FAD7FF1870EA" italics="true" pageId="2" pageNumber="167">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
appears to have higher number than
|
||
<emphasis id="B97EEAA03F4DFFBE2DB3FA2AFD1470EA" box="[657,767,1426,1450]" italics="true" pageId="2" pageNumber="167">
|
||
<taxonomicName id="4C0A4D313F4DFFBE2DB3FA2AFD1070EA" box="[657,763,1426,1450]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="167" phylum="Chordata" rank="species" species="donaji">R. donaji</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which has 40.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31065393F4DFFB82FE5FA0DFE5C77C5" lastPageId="4" lastPageNumber="169" pageId="2" pageNumber="167" type="description">
|
||
<paragraph id="8BB536B23F4DFFBE2FE5FA0DFAF3721A" blockId="2.[151,1437,957,2026]" pageId="2" pageNumber="167">
|
||
<emphasis id="B97EEAA03F4DFFBE2FE5FA0DFDFA708E" bold="true" box="[199,529,1461,1486]" pageId="2" pageNumber="167">
|
||
Description of the
|
||
<typeStatus id="54B188103F4DFFBE2E81FA0DFDE7708E" box="[419,524,1461,1486]" pageId="2" pageNumber="167" type="holotype">holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Adult female with few small supracloacal tubercles; head length 9.5; head width 4.0; neck 3.0; snout-vent length 256; tail missing perhaps no more than 2–3 distal subcaudals, length 85 + N; eye diameter 1.1; naris to anterior edge of eye 3.0; head scales arranged in typical colubrid fashion with paired internasals, paired prefrontals, broad median frontal, narrow paired supraoculars, and elongate paired parietals; internasal suture 0.6, about 33% length of prefrontal suture; frontal length 3.0, about 120% as long as its distance from tip of snout, 67% as long as parietals; interocular distance 3.5, 63% occupied by frontal; mental broad, 1.5 x 1.0; two pairs of genials, subequal in size, posterior pair wing-shaped, deeply incised along median suture. Dorsal scales smooth, in 17/17/17 rows; four rows at midlength of tail; apical pits appear to be absent; supralabials 7/7, third and fourth entering orbit; infralabials 7/8, first pair in contact behind mental along midventer, first to third/ first to fourth in contact with anterior genials, respectively; third and fourth/fourth and fifth infralabials in contact with posterior genials, respectively; preoculars 1/1, subpreocular absent; postoculars 1/1; temporals 1+2; preventrals 2; ventrals 172; cloacal plate divided; subcaudals 63+N (incomplete), divided; pupil round.
|
||
</paragraph>
|
||
<paragraph id="8BB536B23F4DFFBE2FE5F8DDFD7F72AA" blockId="2.[151,1437,957,2026]" pageId="2" pageNumber="167">Left maxilla bearing 12 teeth, increasing in size posteriorly to ninth tooth, with 4–5 teeth in middle of series noticeably enlarged, several posteriormost teeth smaller; tooth nine even with anterior edge of ectopterygoid process; teeth arranged in straight line except for slight medial curvature of anterior of maxilla; no tooth noticeably offset; no pronounced diastema within series.</paragraph>
|
||
<caption id="DF75663A3F4CFFBF2FB5F8ADFD96720A" httpUri="https://zenodo.org/record/262520/files/figure.png" pageId="3" pageNumber="168" targetBox="[154,1348,269,1732]" targetPageId="3">
|
||
<paragraph id="8BB536B23F4CFFBF2FB5F8ADFD96720A" blockId="3.[151,1435,1813,1866]" pageId="3" pageNumber="168">
|
||
<emphasis id="B97EEAA03F4CFFBF2FB5F8ADFEF9726A" bold="true" box="[151,274,1813,1835]" pageId="3" pageNumber="168">FIGURE 2.</emphasis>
|
||
Dorsal (A), lateral (B), and ventral (C) views of head of the holotype of
|
||
<taxonomicName id="4C0A4D313F4CFFBF2CD0F8ADFB51726A" box="[1010,1210,1813,1834]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="168" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4CFFBF2CD0F8ADFB51726A" box="[1010,1210,1813,1834]" italics="true" pageId="3" pageNumber="168">Rhadinella dysmica</emphasis>
|
||
</taxonomicName>
|
||
(ENCB 18951). Head length 9.5 mm. Photos by N. V. López Valentín.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536B23F4CFFB82FE5F8CAFE5C77C5" blockId="3.[151,1436,1905,2038]" lastBlockId="4.[151,1436,151,645]" lastPageId="4" lastPageNumber="169" pageId="3" pageNumber="168">In life, black head cap extending from internasals to near posterior edges of parietals, including prefrontals, frontal, supraoculars; rostral with inverted, black U-shaped marking bordering lingual groove, bordered above with dusky white on edges of first supralabials, anterior nasals, and internasals; very small amount of beige mottling on posterior nasal, loreal and anterolateral portion of prefrontal; parietals with pale spot lying to either side of common parietal suture; lateral and posterior edges of parietals with dusky white borders; side of head including nasals, loreal, preocular, and postocular blackish; upper edges of most supralabials blackish, whitish below, with lower white area of supralabials sometimes with dark brown to blackish spotting; most of primary, secondary, and upper tertiary temporals with distinctive dark brown borders and whitish centers; mental with dark brown lateral borders; infralabials 1–5 with irregular dark brown vertical streaks either centrally or along suture; anterior one-third to onehalf of anterior genial with dark brown pigment; nuchal white collar extending from posterior edge of parietals posteriorly 2.0–2.5 scale lengths, about 3–4 scale lengths wide on side of head, involving posterior portion of ultimate supralabials; dorsal pigment dark brown, barely encroaching on lateral portion of ventrals, more so on subcaudals; first two rows of dorsals with whitish central dashes; dorsal scale rows 4–5 with inconspicuous brown stripe, bordered below by a darker brown line on adjacent portions of scale rows 3–4; scale row 6 with inconspicuous dark brown stripe extending to adjacent portions of scale rows 5 and 7; subcaudals white with irregular brown pigment along midventral area; distal one-third of tail mostly brown; iris dark bronze. In preservative (ethanol after formalin) dark pigment of head and body mostly dark brown; striping of body obscured; collar and ventrum white.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF75663A3F4BFFB82FB5F831FCB772FE" httpUri="https://zenodo.org/record/262521/files/figure.png" pageId="4" pageNumber="169" targetBox="[157,1395,739,1874]" targetPageId="4">
|
||
<paragraph id="8BB536B23F4BFFB82FB5F831FCB772FE" blockId="4.[151,1436,1929,1982]" pageId="4" pageNumber="169">
|
||
<emphasis id="B97EEAA03F4BFFB82FB5F831FEF872DE" bold="true" box="[151,275,1929,1951]" pageId="4" pageNumber="169">FIGURE 3.</emphasis>
|
||
Dorsal (A) and (B) lateral aspects of
|
||
<taxonomicName id="4C0A4D313F4BFFB82DB0F831FC8E72DE" box="[658,869,1929,1950]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="169" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F4BFFB82DB0F831FC8E72DE" box="[658,869,1929,1950]" italics="true" pageId="4" pageNumber="169">Rhadinella schistosa</emphasis>
|
||
</taxonomicName>
|
||
(FMNH 100297, paratype) from Cuautlapan, Veracruz, Mexico, showing incomplete collar across nape. Head length 6.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF75663A3F4AFFB92FB5FC56FE1E7162" httpUri="https://zenodo.org/record/262522/files/figure.png" pageId="5" pageNumber="170" targetBox="[156,1429,411,979]" targetPageId="5">
|
||
<paragraph id="8BB536B23F4AFFB92FB5FC56FE1E7162" blockId="5.[151,1436,1006,1058]" pageId="5" pageNumber="170">
|
||
<emphasis id="B97EEAA03F4AFFB92FB5FC56FEFF7143" bold="true" box="[151,276,1006,1028]" pageId="5" pageNumber="170">FIGURE 4.</emphasis>
|
||
Landscape view taken of type-locality of
|
||
<taxonomicName id="4C0A4D313F4AFFB92DEBFC56FC787143" box="[713,915,1006,1027]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="170" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F4AFFB92DEBFC56FC787143" box="[713,915,1006,1027]" italics="true" pageId="5" pageNumber="170">Rhadinella dysmica</emphasis>
|
||
</taxonomicName>
|
||
, looking north from Cuajilotla, Guerrero, Mexico, showing tropical deciduous forest.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF75663A3F4AFFB92FB5F81EFC1872FC" box="[151,1011,1958,1980]" httpUri="https://zenodo.org/record/262523/files/figure.png" pageId="5" pageNumber="170" targetBox="[211,1366,1155,1885]" targetPageId="5">
|
||
<paragraph id="8BB536B23F4AFFB92FB5F81EFC1872FC" blockId="5.[151,1011,1958,1980]" box="[151,1011,1958,1980]" pageId="5" pageNumber="170">
|
||
<emphasis id="B97EEAA03F4AFFB92FB5F81EFEF972FC" bold="true" box="[151,274,1958,1980]" pageId="5" pageNumber="170">FIGURE 5.</emphasis>
|
||
Distribution of dark-colored species of
|
||
<taxonomicName id="4C0A4D313F4AFFB92D85F81EFCFD72FB" box="[679,790,1958,1979]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="170" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F4AFFB92D85F81EFCFD72FB" box="[679,790,1958,1979]" italics="true" pageId="5" pageNumber="170">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
occurring in Mexico.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31065393F49FFBA2FE5FF2FFB0275B8" pageId="6" pageNumber="171" type="etymology">
|
||
<paragraph id="8BB536B23F49FFBA2FE5FF2FFB0275B8" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="171">
|
||
<emphasis id="B97EEAA03F49FFBA2FE5FF2FFEA575F0" bold="true" box="[199,334,151,176]" pageId="6" pageNumber="171">Etymology.</emphasis>
|
||
The specific epithet is derived from the Greek ἐφέσπερος (
|
||
<emphasis id="B97EEAA03F49FFBA2B0AFF20FB7375F0" box="[1064,1176,152,176]" italics="true" pageId="6" pageNumber="171">dysmikos)</emphasis>
|
||
, meaning western, in reference to the westernmost distribution of this species of the genus. We propose standard English and Spanish names of Guerrero Slender Leaf Litter Snake and Hojarasquera Delgada de Guerrero, respectively.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31065393F49FFBB2FE5FEBCFBD97117" lastPageId="7" lastPageNumber="172" pageId="6" pageNumber="171" type="distribution">
|
||
<paragraph id="8BB536B23F49FFBA2FE5FEBCFD877490" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="171">
|
||
<emphasis id="B97EEAA03F49FFBA2FE5FEBCFE19745D" bold="true" box="[199,498,260,285]" pageId="6" pageNumber="171">Distribution and habitat.</emphasis>
|
||
The type-locality lies at
|
||
<quantity id="4CF29B573F49FFBA2C2FFEBCFCBF745C" box="[781,852,260,284]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="6" pageNumber="171" unit="m" value="432.0">432 m</quantity>
|
||
asl at the edge of a small valley occupied the small village of Cuajilotla (
|
||
<figureCitation id="13312A373F49FFBA2EA8FE9FFE257400" box="[394,462,295,320]" captionStart="FIGURE 4" captionStartId="5.[151,250,1006,1028]" captionTargetBox="[156,1429,411,979]" captionTargetId="figure@5.[152,1435,193,985]" captionTargetPageId="5" captionText="FIGURE 4. Landscape view taken of type-locality of Rhadinella dysmica, looking north from Cuajilotla, Guerrero, Mexico, showing tropical deciduous forest." httpUri="https://zenodo.org/record/262522/files/figure.png" pageId="6" pageNumber="171">Fig. 4</figureCitation>
|
||
) in the foothills of the eastern portion of Sierra Madre del Sur in Guerrero in tropical deciduous forest. The predominant
|
||
<typeStatus id="54B188103F49FFBA2D03FEF5FDB77425" box="[545,604,333,357]" pageId="6" pageNumber="171">types</typeStatus>
|
||
of vegetation at higher elevations in the region are oak forest and pine-oak forest, grading into tropical deciduous forest, which is restricted to the lowlands of foothills, valleys, and slopes (
|
||
<bibRefCitation id="EF9B4B433F49FFBA2FBCFE2CFEAA74ED" author="Leopold" box="[158,321,404,429]" pageId="6" pageNumber="171" refString="Leopold, A. S. (1950) Vegetation zones of Mexico. Ecology, 31, 507 - 518. http: // dx. doi. org / 10.2307 / 1931569" type="journal article" year="1950">Leopold 1950</bibRefCitation>
|
||
). The type-locality (
|
||
<figureCitation id="13312A373F49FFBA2D06FE2CFD8C74ED" box="[548,615,404,429]" captionStart="FIGURE 5" captionStartId="5.[151,250,1958,1980]" captionTargetBox="[211,1366,1155,1885]" captionTargetId="figure@5.[150,1436,1108,1936]" captionTargetPageId="5" captionText="FIGURE 5. Distribution of dark-colored species of Rhadinella occurring in Mexico." httpUri="https://zenodo.org/record/262523/files/figure.png" pageId="6" pageNumber="171">Fig. 5</figureCitation>
|
||
) is drained by the Río Chapolapa, an upper tributary of the Río Omitlán- Azul, which flows into the Río Papagayo.
|
||
</paragraph>
|
||
<paragraph id="8BB536B23F49FFBA2FE5FE64FAC17720" blockId="6.[151,1437,151,608]" pageId="6" pageNumber="171">
|
||
Other species of amphibian and reptiles recorded between
|
||
<quantity id="4CF29B573F49FFBA2C7EFE64FC3774B4" box="[860,988,476,500]" metricMagnitude="2" metricUnit="m" metricValue="3.86" metricValueMax="4.32" metricValueMin="3.4" pageId="6" pageNumber="171" unit="m" value="386.0" valueMax="432.0" valueMin="340.0">340–432 m</quantity>
|
||
in the vicinity of the type-locality were
|
||
<emphasis id="B97EEAA03F49FFBA2FB5FDB8FD5F771F" italics="true" pageId="6" pageNumber="171">
|
||
<taxonomicName id="4C0A4D313F49FFBA2FB5FDB8FE037757" authority="Craugastor" authorityName="Craugastor" box="[151,488,512,536]" class="Amphibia" family="Bufonidae" genus="Rhinella" kingdom="Animalia" order="Anura" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="marina">Rhinella marina, Craugastor</taxonomicName>
|
||
rugulosus, Agalychnis dacnicolor, Smilisca baudinii, Anolis dunni, Aspidoscelis guttata, Holcosus undulatus, Phyllodactylus lanei, Phyllodactylus tuberculosus, Ctenosaura pectinata, Sceloporus siniferus, Urosaurus bicarinatus,
|
||
<taxonomicName id="4C0A4D313F49FFBA2D32FDF1FD5F771F" box="[528,692,584,608]" class="Reptilia" family="Boidae" genus="Boa" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="imperator">Boa imperator</taxonomicName>
|
||
</emphasis>
|
||
(sensu
|
||
<bibRefCitation id="EF9B4B433F49FFBA2C28FDFFFC137720" author="Hynkova" box="[778,1016,583,608]" pageId="6" pageNumber="171" refString="Hynkova, I, Starostova, Z. & Frynta, D. (2009) Mitochondrial DNA variation reveals recent evolutionary history of main Boa constrictor clades. Zoological Science, 26, 623 - 631." type="journal article" year="2009">
|
||
Hynková
|
||
<emphasis id="B97EEAA03F49FFBA2C5AFDF1FC5B7720" box="[888,944,584,608]" italics="true" pageId="6" pageNumber="171">et al.</emphasis>
|
||
, 2009
|
||
</bibRefCitation>
|
||
), and
|
||
<emphasis id="B97EEAA03F49FFBA2B1EFDF0FAC17720" box="[1084,1322,584,608]" italics="true" pageId="6" pageNumber="171">
|
||
<taxonomicName id="4C0A4D313F49FFBA2B1EFDF0FACE7720" box="[1084,1317,584,608]" class="Reptilia" family="Colubridae" genus="Leptodeira" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="annulata">Leptodeira annulata</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<caption id="DF75663A3F49FFBA2FB5FD28FAFE77E7" ID-Table-UUID="DF75663A3F49FFBA2FB5FD28FAFE77E7" box="[151,1301,655,679]" httpUri="http://table.plazi.org/id/DF75663A3F49FFBA2FB5FD28FAFE77E7" pageId="6" pageNumber="171" targetBox="[151,1411,707,1381]" targetIsTable="true" targetPageId="6">
|
||
<paragraph id="8BB536B23F49FFBA2FB5FD28FAFE77E7" blockId="6.[151,1301,655,679]" box="[151,1301,655,679]" pageId="6" pageNumber="171">
|
||
<emphasis id="B97EEAA03F49FFBA2FB5FD28FEE177E7" bold="true" box="[151,266,656,679]" pageId="6" pageNumber="171">TABLE 1.</emphasis>
|
||
Comparison of selected features from Mexican species of
|
||
<taxonomicName id="4C0A4D313F49FFBA2C5FFD37FC1977E7" box="[893,1010,655,679]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F49FFBA2C5FFD37FC1977E7" box="[893,1010,655,679]" italics="true" pageId="6" pageNumber="171">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
with dark-colored dorsum.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB536B23F49FFBA2EBDFD7BFEF07025" pageId="6" pageNumber="171">
|
||
<table id="F90AC4123F4900432FB5FD7BFA687025" box="[151,1411,707,1381]" gridcols="5" gridrows="9" pageId="6" pageNumber="171">
|
||
<tr id="353A34F03F4900432FB5FD7BFA687798" box="[151,1411,707,728]" gridrow="0" pageId="6" pageNumber="171" rowspan-0="1">
|
||
<th id="76EB5D8C3F4900432EBDFD7BFD947798" box="[415,639,707,728]" gridcol="1" gridrow="0" pageId="6" pageNumber="171">
|
||
<taxonomicName id="4C0A4D313F49FFBA2EBDFD7BFD8C7798" box="[415,615,707,728]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F49FFBA2EBDFD7BFD8C7798" box="[415,615,707,728]" italics="true" pageId="6" pageNumber="171">Rhadinella dysmica</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76EB5D8C3F4900432DB7FD7BFC807798" box="[661,875,707,728]" gridcol="2" gridrow="0" pageId="6" pageNumber="171">
|
||
<taxonomicName id="4C0A4D313F49FFBA2DB7FD7BFCA77798" box="[661,844,707,728]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F49FFBA2DB7FD7BFCA77798" box="[661,844,707,728]" italics="true" pageId="6" pageNumber="171">Rhadinella donaji</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76EB5D8C3F4900432CB3FD7BFB9E7798" box="[913,1141,707,728]" gridcol="3" gridrow="0" pageId="6" pageNumber="171">
|
||
<taxonomicName id="4C0A4D313F49FFBA2CB3FD7BFB887798" box="[913,1123,707,728]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F49FFBA2CB3FD7BFB887798" box="[913,1123,707,728]" italics="true" pageId="6" pageNumber="171">Rhadinella schistosa</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76EB5D8C3F4900432BA9FD7BFA687798" box="[1163,1411,707,728]" gridcol="4" gridrow="0" pageId="6" pageNumber="171">
|
||
<taxonomicName id="4C0A4D313F49FFBA2BA9FD7BFAB07798" box="[1163,1371,707,728]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="posadasi">
|
||
<emphasis id="B97EEAA03F49FFBA2BA9FD7BFAB07798" box="[1163,1371,707,728]" italics="true" pageId="6" pageNumber="171">Rhadinella posadasi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FD54FA68763C" box="[151,1411,748,892]" gridrow="1" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FD54FE69763C" box="[151,386,748,892]" gridcol="0" gridrow="1" pageId="6" pageNumber="171">Head pattern</th>
|
||
<td id="76EB5D8C3F4900432EBDFD54FD94763C" box="[415,639,748,892]" gridcol="1" gridrow="1" pageId="6" pageNumber="171">Head mostly black followed by white collar; lateral edges of parietals dingy white</td>
|
||
<td id="76EB5D8C3F4900432DB7FD54FC80763C" box="[661,875,748,892]" gridcol="2" gridrow="1" pageId="6" pageNumber="171">Head mostly white with scant dark mottling followed by white collar</td>
|
||
<td id="76EB5D8C3F4900432CB3FD54FB9E763C" box="[913,1141,748,892]" gridcol="3" gridrow="1" pageId="6" pageNumber="171">Top of head uniformly dark followed by U- shaped white collar, usually broken medially</td>
|
||
<td id="76EB5D8C3F4900432BA9FD54FA68763C" box="[1163,1411,748,892]" gridcol="4" gridrow="1" pageId="6" pageNumber="171">Top of head with uniformly dark head cap followed by orange or auburn collar</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FC28FA687684" box="[151,1411,912,964]" gridrow="2" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FC28FE697684" box="[151,386,912,964]" gridcol="0" gridrow="2" pageId="6" pageNumber="171">Infralabials contacting posterior genials</th>
|
||
<td id="76EB5D8C3F4900432EBDFC28FD947684" box="[415,639,912,964]" gridcol="1" gridrow="2" pageId="6" pageNumber="171">Third and fourth/ fourth and fifth</td>
|
||
<td id="76EB5D8C3F4900432DB7FC28FC807684" box="[661,875,912,964]" gridcol="2" gridrow="2" pageId="6" pageNumber="171">Third and fourth</td>
|
||
<td id="76EB5D8C3F4900432CB3FC28FB9E7684" box="[913,1141,912,964]" gridcol="3" gridrow="2" pageId="6" pageNumber="171">Fourth and fifth</td>
|
||
<td id="76EB5D8C3F4900432BA9FC28FA687684" box="[1163,1411,912,964]" gridcol="4" gridrow="2" pageId="6" pageNumber="171">Fourth and fifth</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FC60FA6876AE" box="[151,1411,984,1006]" gridrow="3" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FC60FE6976AE" box="[151,386,984,1006]" gridcol="0" gridrow="3" pageId="6" pageNumber="171">Supralabials</th>
|
||
<td id="76EB5D8C3F4900432EBDFC60FD9476AE" box="[415,639,984,1006]" gridcol="1" gridrow="3" pageId="6" pageNumber="171">7</td>
|
||
<td id="76EB5D8C3F4900432DB7FC60FC8076AE" box="[661,875,984,1006]" gridcol="2" gridrow="3" pageId="6" pageNumber="171">7</td>
|
||
<td id="76EB5D8C3F4900432CB3FC60FB9E76AE" box="[913,1141,984,1006]" gridcol="3" gridrow="3" pageId="6" pageNumber="171">8</td>
|
||
<td id="76EB5D8C3F4900432BA9FC60FA6876AE" box="[1163,1411,984,1006]" gridcol="4" gridrow="3" pageId="6" pageNumber="171">7</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FBB9FA687157" box="[151,1411,1025,1047]" gridrow="4" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FBB9FE697157" box="[151,386,1025,1047]" gridcol="0" gridrow="4" pageId="6" pageNumber="171">Temporals</th>
|
||
<td id="76EB5D8C3F4900432EBDFBB9FD947157" box="[415,639,1025,1047]" gridcol="1" gridrow="4" pageId="6" pageNumber="171">1 + 2</td>
|
||
<td id="76EB5D8C3F4900432DB7FBB9FC807157" box="[661,875,1025,1047]" gridcol="2" gridrow="4" pageId="6" pageNumber="171">1 + 2</td>
|
||
<td id="76EB5D8C3F4900432CB3FBB9FB9E7157" box="[913,1141,1025,1047]" gridcol="3" gridrow="4" pageId="6" pageNumber="171">1 + 1</td>
|
||
<td id="76EB5D8C3F4900432BA9FBB9FA687157" box="[1163,1411,1025,1047]" gridcol="4" gridrow="4" pageId="6" pageNumber="171">1 + 2</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FB92FA68711F" box="[151,1411,1066,1119]" gridrow="5" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FB92FE69711F" box="[151,386,1066,1119]" gridcol="0" gridrow="5" pageId="6" pageNumber="171">Ventrals</th>
|
||
<td id="76EB5D8C3F4900432EBDFB92FD94711F" box="[415,639,1066,1119]" gridcol="1" gridrow="5" pageId="6" pageNumber="171">Males: unknown Female: 172</td>
|
||
<td id="76EB5D8C3F4900432DB7FB92FC80711F" box="[661,875,1066,1119]" gridcol="2" gridrow="5" pageId="6" pageNumber="171">Male: 166 Females: unknown</td>
|
||
<td id="76EB5D8C3F4900432CB3FB92FB9E711F" box="[913,1141,1066,1119]" gridcol="3" gridrow="5" pageId="6" pageNumber="171">Males: 145–147 Females: 151–156</td>
|
||
<td id="76EB5D8C3F4900432BA9FB92FA68711F" box="[1163,1411,1066,1119]" gridcol="4" gridrow="5" pageId="6" pageNumber="171">Males: 136–141 Females: 143–146</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FBCAFA6871E7" box="[151,1411,1138,1191]" gridrow="6" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FBCAFE6971E7" box="[151,386,1138,1191]" gridcol="0" gridrow="6" pageId="6" pageNumber="171">Subcaudals</th>
|
||
<td id="76EB5D8C3F4900432EBDFBCAFD9471E7" box="[415,639,1138,1191]" gridcol="1" gridrow="6" pageId="6" pageNumber="171">Males: unknown Female: 63+ N</td>
|
||
<td id="76EB5D8C3F4900432DB7FBCAFC8071E7" box="[661,875,1138,1191]" gridcol="2" gridrow="6" pageId="6" pageNumber="171">Male: unknown Females: unknown</td>
|
||
<td id="76EB5D8C3F4900432CB3FBCAFB9E71E7" box="[913,1141,1138,1191]" gridcol="3" gridrow="6" pageId="6" pageNumber="171">Males: 40–42 Females: 31–34</td>
|
||
<td id="76EB5D8C3F4900432BA9FBCAFA6871E7" box="[1163,1411,1138,1191]" gridcol="4" gridrow="6" pageId="6" pageNumber="171">Male: 92–95 Females: 86–93</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FB02FA6871AF" box="[151,1411,1210,1263]" gridrow="7" pageId="6" pageNumber="171">
|
||
<th id="76EB5D8C3F4900432FB5FB02FE6971AF" box="[151,386,1210,1263]" gridcol="0" gridrow="7" pageId="6" pageNumber="171">No. maxillary teeth (sample size)</th>
|
||
<td id="76EB5D8C3F4900432EBDFB02FD9471AF" box="[415,639,1210,1263]" gridcol="1" gridrow="7" pageId="6" pageNumber="171">12 (1)</td>
|
||
<td id="76EB5D8C3F4900432DB7FB02FC8071AF" box="[661,875,1210,1263]" gridcol="2" gridrow="7" pageId="6" pageNumber="171">11 (1)</td>
|
||
<td id="76EB5D8C3F4900432CB3FB02FB9E71AF" box="[913,1141,1210,1263]" gridcol="3" gridrow="7" pageId="6" pageNumber="171">16–17 (5)</td>
|
||
<td id="76EB5D8C3F4900432BA9FB02FA6871AF" box="[1163,1411,1210,1263]" gridcol="4" gridrow="7" pageId="6" pageNumber="171">11 (2)</td>
|
||
</tr>
|
||
<tr id="353A34F03F4900432FB5FAF3FA687025" box="[151,1411,1355,1381]" gridrow="8" pageId="6" pageNumber="171" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1">
|
||
<th id="76EB5D8C3F4900432FB5FAF3FE697025" box="[151,386,1355,1381]" gridcol="0" gridrow="8" pageId="6" pageNumber="171">Discussion</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB536B23F49FFBA2FB5FA29FE9E73AE" blockId="6.[151,1436,1425,2026]" pageId="6" pageNumber="171">
|
||
The taxonomic histories of snakes placed in the genus
|
||
<taxonomicName id="4C0A4D313F49FFBA2C25FA29FC6970E9" box="[775,898,1425,1449]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F49FFBA2C25FA29FC6970E9" box="[775,898,1425,1449]" italics="true" pageId="6" pageNumber="171">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
were most recently discussed by
|
||
<bibRefCitation id="EF9B4B433F49FFBA2A20FA2AFA7770EA" author="Myers" box="[1282,1436,1426,1450]" pageId="6" pageNumber="171" refString="Myers, C. W. (2011) A new genus and new tribe for Enicognathus melanauchen Jan 1863, a neglected South American snake (Colubridae: Xenodontidae), with taxonomic notes on some Dipsadinae. American Museum Novitates, 3715, 1 - 33. http: // dx. doi. org / 10.1206 / 3715.2" type="journal article" year="2011">Myers (2011)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF9B4B433F49FFBA2FE5FA0DFE61708E" author="Campbell" box="[199,394,1461,1486]" pageId="6" pageNumber="171" refString="Campbell, J. A. (2015) A new species of Rhadinella (Serpentes: Colubridae) from the Pacific versant of Oaxaca, Mexico. Zootaxa, 3918 (3), 397 - 405. http: // dx. doi. org / 10.11646 / zootaxa. 3918.3.3" type="journal article" year="2015">Campbell (2015)</bibRefCitation>
|
||
. Most of the species in the genus occur in Nuclear Central
|
||
<collectingCountry id="F31D76223F49FFBA2B08FA0DFB66708E" box="[1066,1165,1461,1486]" name="United States of America" pageId="6" pageNumber="171">America</collectingCountry>
|
||
, with a number of them inhabiting Chiapas,
|
||
<collectingCountry id="F31D76223F49FFBA2E5AFA61FE3A70B2" box="[376,465,1497,1522]" name="Mexico" pageId="6" pageNumber="171">Mexico</collectingCountry>
|
||
. Of the species having a dark, blackish dorsum, with or without inconspicuous stripes, only two species where known from
|
||
<collectingCountry id="F31D76223F49FFBA2D67FA45FD717356" box="[581,666,1533,1558]" name="Mexico" pageId="6" pageNumber="171">Mexico</collectingCountry>
|
||
until recently:
|
||
<taxonomicName id="4C0A4D313F49FFBA2C6EFA46FC3E7355" box="[844,981,1533,1557]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F49FFBA2C6EFA46FC3E7355" box="[844,981,1533,1557]" italics="true" pageId="6" pageNumber="171">R. schistosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9B4B433F49FFBA2CC4FA45FAC37356" author="Smith" box="[998,1320,1533,1558]" pageId="6" pageNumber="171" refString="Smith, H. M. (1941) A new genus of Mexican snakes related to Rhadinaea. Copeia, 1941, 7 - 10. http: // dx. doi. org / 10.2307 / 1437694" type="journal article" year="1941">Smith 1941; the generotype</bibRefCitation>
|
||
) from the eastern versant of Veracruz and
|
||
<taxonomicName id="4C0A4D313F49FFBA2ED9F99AFD957379" box="[507,638,1569,1593]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="posadasi">
|
||
<emphasis id="B97EEAA03F49FFBA2ED9F99AFD957379" box="[507,638,1569,1593]" italics="true" pageId="6" pageNumber="171">R. posadasi</emphasis>
|
||
</taxonomicName>
|
||
from extreme southern Chiapas. The genus was seemingly absent from the Pacific versant of southern
|
||
<collectingCountry id="F31D76223F49FFBA2EDAF9FDFDA5731E" box="[504,590,1605,1630]" name="Mexico" pageId="6" pageNumber="171">Mexico</collectingCountry>
|
||
until the recent description of
|
||
<taxonomicName id="4C0A4D313F49FFBA2C8FF9FEFB36731E" authority="Campbell (2015)" authorityName="Campbell" authorityYear="2015" box="[941,1245,1605,1630]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F49FFBA2C8FF9FEFBFF731D" box="[941,1044,1605,1629]" italics="true" pageId="6" pageNumber="171">R. donaji</emphasis>
|
||
<bibRefCitation id="EF9B4B433F49FFBA2B3FF9FDFB36731E" author="Campbell" box="[1053,1245,1605,1630]" pageId="6" pageNumber="171" refString="Campbell, J. A. (2015) A new species of Rhadinella (Serpentes: Colubridae) from the Pacific versant of Oaxaca, Mexico. Zootaxa, 3918 (3), 397 - 405. http: // dx. doi. org / 10.11646 / zootaxa. 3918.3.3" type="journal article" year="2015">Campbell (2015)</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Pacific highlands of Oaxaca. The subsequent discovery of
|
||
<taxonomicName id="4C0A4D313F49FFBA2DFAF9D2FCBF73C1" box="[728,852,1641,1665]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F49FFBA2DFAF9D2FCBF73C1" box="[728,852,1641,1665]" italics="true" pageId="6" pageNumber="171">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
at a relatively low elevation (
|
||
<quantity id="4CF29B573F49FFBA2B87F9D2FB0473C2" box="[1189,1263,1642,1666]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="6" pageNumber="171" unit="m" value="432.0">432 m</quantity>
|
||
) in Guerrero is somewhat of a surprise. Snakes occurring at low elevations in tropical deciduous forest usually have rather extensive distributions in this widespread and continuous lowland habitat, not being affected by the isolating effects of elevation.
|
||
</paragraph>
|
||
<paragraph id="8BB536B23F49FFBA2FE5F942FB2A72AA" blockId="6.[151,1436,1425,2026]" pageId="6" pageNumber="171">
|
||
Many species of
|
||
<taxonomicName id="4C0A4D313F49FFBA2EA9F941FDED7251" box="[395,518,1785,1809]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F49FFBA2EA9F941FDED7251" box="[395,518,1785,1809]" italics="true" pageId="6" pageNumber="171">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
have distinctive longitudinal body striping that serves as a useful identifier (see
|
||
<bibRefCitation id="EF9B4B433F49FFBA2FB5F8A6FECA7276" author="Myers" box="[151,289,1822,1846]" pageId="6" pageNumber="171" refString="Myers, C. W. (1974) The systematics of Rhadinaea (Colubridae), a genus of New World snakes. Bulletin of the American Museum of Natural History, 133, 1 - 262." type="journal article" year="1974">Myers 1974</bibRefCitation>
|
||
: fig. 28). The pattern in
|
||
<taxonomicName id="4C0A4D313F49FFBA2D13F8A6FD477275" box="[561,684,1821,1845]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F49FFBA2D13F8A6FD477275" box="[561,684,1821,1845]" italics="true" pageId="6" pageNumber="171">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
with the combination of dark striping on adjacent portions of scale rows 3–4 and on the sixth scale row and adjacent portions of fifth and seventh scale rows is not found in any other species of
|
||
<taxonomicName id="4C0A4D313F49FFBA2E2EF8DDFE6C723D" box="[268,391,1893,1917]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F49FFBA2E2EF8DDFE6C723D" box="[268,391,1893,1917]" italics="true" pageId="6" pageNumber="171">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
. Unfortunately, the dorsal patterning in the dark-colored species is often difficult to discern or obscured altogether. We are able to address the striping in
|
||
<taxonomicName id="4C0A4D313F49FFBA2C14F832FC5872E1" box="[822,947,1929,1953]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="171" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F49FFBA2C14F832FC5872E1" box="[822,947,1929,1953]" italics="true" pageId="6" pageNumber="171">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
only because of the excellent photographs taken in life. Under certain light conditions, striping pattern in life is obscured owing to the highly iridescent scales of this snake, and in preservative there is a tendency for striping to become even more obscure.
|
||
</paragraph>
|
||
<paragraph id="8BB536B23F48FFBB2FE5FF2FFCF0777C" blockId="7.[151,1437,151,1112]" pageId="7" pageNumber="172">
|
||
The geographically closest congeners to
|
||
<taxonomicName id="4C0A4D313F48FFBB2D8FFF21FCC475F0" box="[685,815,152,176]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2D8FFF21FCC475F0" box="[685,815,152,176]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
are
|
||
<taxonomicName id="4C0A4D313F48FFBB2C49FF21FC1C75F0" box="[875,1015,152,176]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F48FFBB2C49FF21FC1C75F0" box="[875,1015,152,176]" italics="true" pageId="7" pageNumber="172">R. schistosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D313F48FFBB2B1BFF21FB4E75F0" box="[1081,1189,152,176]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2B1BFF21FB4E75F0" box="[1081,1189,152,176]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13312A373F48FFBB2B99FF2FFAED75F0" box="[1211,1286,151,176]" captionStart="FIGURE 5" captionStartId="5.[151,250,1958,1980]" captionTargetBox="[211,1366,1155,1885]" captionTargetId="figure@5.[150,1436,1108,1936]" captionTargetPageId="5" captionText="FIGURE 5. Distribution of dark-colored species of Rhadinella occurring in Mexico." httpUri="https://zenodo.org/record/262523/files/figure.png" pageId="7" pageNumber="172">Fig. 5</figureCitation>
|
||
).
|
||
<taxonomicName id="4C0A4D313F48FFBB2A03FF20FF157594" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="schistosa">
|
||
<emphasis id="B97EEAA03F48FFBB2A03FF20FF157594" italics="true" pageId="7" pageNumber="172">Rhadinella schistosa</emphasis>
|
||
</taxonomicName>
|
||
occurs about
|
||
<quantity id="4CF29B573F48FFBB2EBAFF04FE067595" box="[408,493,188,213]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="172" unit="km" value="300.0">300 km</quantity>
|
||
to the northeast, separated by the xeric Tehuacán-Cuicatlán Valley. Most individuals have been taken from near
|
||
<quantity id="4CF29B573F48FFBB2EE8FF58FDCB75B7" box="[458,544,224,248]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="172" unit="m" value="1000.0">1000 m</quantity>
|
||
asl in mesic tropical evergreen forest or cloud forest.
|
||
<taxonomicName id="4C0A4D313F48FFBB2BA3FF58FAA075B8" box="[1153,1355,224,248]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2BA3FF58FAA075B8" box="[1153,1355,224,248]" italics="true" pageId="7" pageNumber="172">Rhadinella donaji</emphasis>
|
||
</taxonomicName>
|
||
occurs about
|
||
<quantity id="4CF29B573F48FFBB2FC1FEBCFED4745D" box="[227,319,260,285]" metricMagnitude="5" metricUnit="m" metricValue="2.75" pageId="7" pageNumber="172" unit="km" value="275.0">275 km</quantity>
|
||
to the East and, similarly to
|
||
<taxonomicName id="4C0A4D313F48FFBB2D90FEBDFCDF745C" box="[690,820,260,284]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2D90FEBDFCDF745C" box="[690,820,260,284]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
, inhabits the Pacific versant, but their habitats are dramatically different:
|
||
<taxonomicName id="4C0A4D313F48FFBB2EB5FE91FDFA7400" box="[407,529,296,320]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2EB5FE91FDFA7400" box="[407,529,296,320]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
occurs in tropical deciduous forest at
|
||
<quantity id="4CF29B573F48FFBB2C95FE90FC15747F" box="[951,1022,296,320]" metricMagnitude="2" metricUnit="m" metricValue="4.32" pageId="7" pageNumber="172" unit="m" value="432.0">432 m</quantity>
|
||
asl, whereas
|
||
<taxonomicName id="4C0A4D313F48FFBB2BB0FE91FB137400" box="[1170,1272,296,320]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2BB0FE91FB137400" box="[1170,1272,296,320]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
is only known from pine-oak forest at
|
||
<quantity id="4CF29B573F48FFBB2E83FEF4FE1C7424" box="[417,503,332,357]" metricMagnitude="3" metricUnit="m" metricValue="2.195" pageId="7" pageNumber="172" unit="m" value="2195.0">2195 m</quantity>
|
||
asl. The elevational distribution of these species remains to be ascertained but it is unlikely that their ranges overlap given habitat and elevational differences between the two species, and the fact that distribution of other species of
|
||
<taxonomicName id="4C0A4D313F48FFBB2D16FE2CFD4474EC" box="[564,687,404,428]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F48FFBB2D16FE2CFD4474EC" box="[564,687,404,428]" italics="true" pageId="7" pageNumber="172">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
are geographically restricted. There are several traits apparent in members of the genus
|
||
<taxonomicName id="4C0A4D313F48FFBB2E82FE00FDF07490" box="[416,539,440,464]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97EEAA03F48FFBB2E82FE00FDF07490" box="[416,539,440,464]" italics="true" pageId="7" pageNumber="172">Rhadinella</emphasis>
|
||
</taxonomicName>
|
||
with a dark dorsum. First, each species seems to be restricted to a particular habitat, whether it be cloud forest, pine-oak, rainforest, or in the case of
|
||
<taxonomicName id="4C0A4D313F48FFBB2CC8FE65FB8374B4" box="[1002,1128,476,500]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2CC8FE65FB8374B4" box="[1002,1128,476,500]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
, tropical deciduous forest. Secondly, the pale head pattern shows little variation within species (
|
||
<bibRefCitation id="EF9B4B433F48FFBB2C9BFDB8FBA27758" author="Myers" box="[953,1097,512,536]" pageId="7" pageNumber="172" refString="Myers, C. W. (1974) The systematics of Rhadinaea (Colubridae), a genus of New World snakes. Bulletin of the American Museum of Natural History, 133, 1 - 262." type="journal article" year="1974">Myers 1974</bibRefCitation>
|
||
). Both of these traits provide evidence for the distinction of
|
||
<taxonomicName id="4C0A4D313F48FFBB2ECCFD9DFD82777C" box="[494,617,548,572]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2ECCFD9DFD82777C" box="[494,617,548,572]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C0A4D313F48FFBB2D8FFD9DFCF8777C" box="[685,787,548,572]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2D8FFD9DFCF8777C" box="[685,787,548,572]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB536B23F48FFBB2FE5FDFFFBD97117" blockId="7.[151,1437,151,1112]" pageId="7" pageNumber="172">
|
||
The segmental counts for
|
||
<taxonomicName id="4C0A4D313F48FFBB2EC5FDF1FDA77720" box="[487,588,584,608]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2EC5FDF1FDA77720" box="[487,588,584,608]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0A4D313F48FFBB2DA0FDF1FD177720" box="[642,764,584,608]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2DA0FDF1FD177720" box="[642,764,584,608]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
are high compared to other members of the genus and given their distributions it is likely they are closely related. With a sample of one for each of these species, it is not possible to address variation. Nonetheless, these species sufficiently differ from each other that their heterospecificity can hardly be doubted. Beside features outlined in the diagnosis, careful examination of these species reveals the following differences in color pattern. The most distinctive element of pattern is the bright white collar which in
|
||
<taxonomicName id="4C0A4D313F48FFBB2E65FD45FE287654" box="[327,451,764,788]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2E65FD45FE287654" box="[327,451,764,788]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
begins behind the parietals, although it is contiguous with parts of the parietals that are edged with dingy white. In
|
||
<taxonomicName id="4C0A4D313F48FFBB2EEAFC99FDC47678" box="[456,559,800,824]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2EEAFC99FDC47678" box="[456,559,800,824]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
the white collar is continuous with the mostly pale parietals. In R.
|
||
<taxonomicName id="4C0A4D313F48FFBB2A35FC98FA987678" box="[1303,1395,800,824]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2A35FC98FA987678" box="[1303,1395,800,824]" italics="true" pageId="7" pageNumber="172">dysmica</emphasis>
|
||
</taxonomicName>
|
||
the temporals have mostly dull white centers with irregularly black edging and the loreal region is dark with small tan flecks, whereas in
|
||
<taxonomicName id="4C0A4D313F48FFBB2E4CFCD1FE3D76C0" box="[366,470,872,896]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2E4CFCD1FE3D76C0" box="[366,470,872,896]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
the temporals are mostly white with irregular dark mottling and the loreal region is mostly white. The dark dorsal ground coloration barely reaches the lateral portions of the ventrals but strongly encroaches on lateral portions of the subcaudals in
|
||
<taxonomicName id="4C0A4D313F48FFBB2DFCFC09FCB17688" box="[734,858,944,968]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2DFCFC09FCB17688" box="[734,858,944,968]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
, whereas in
|
||
<taxonomicName id="4C0A4D313F48FFBB2CCEFC09FBBF7688" box="[1004,1108,944,968]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2CCEFC09FBBF7688" box="[1004,1108,944,968]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
the dorsal coloration fails to reach the ventrals and barely encroaches on the subcaudals. In
|
||
<taxonomicName id="4C0A4D313F48FFBB2C75FC6DFC3976AC" box="[855,978,980,1004]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2C75FC6DFC3976AC" box="[855,978,980,1004]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
dark pigment along the midventer of the tail forms a grayish band, but in
|
||
<taxonomicName id="4C0A4D313F48FFBB2D27FC41FD877150" box="[517,620,1016,1040]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2D27FC41FD877150" box="[517,620,1016,1040]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
the subcaudals are immaculate. The lower two scale rows in
|
||
<taxonomicName id="4C0A4D313F48FFBB2A03FC41FA777150" box="[1313,1436,1016,1040]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="dysmica">
|
||
<emphasis id="B97EEAA03F48FFBB2A03FC41FA777150" box="[1313,1436,1016,1040]" italics="true" pageId="7" pageNumber="172">R. dysmica</emphasis>
|
||
</taxonomicName>
|
||
have very faint and irregular indications of pale dashes in their center and in
|
||
<taxonomicName id="4C0A4D313F48FFBB2CD8FBA5FB8A7174" box="[1018,1121,1052,1076]" class="Reptilia" family="Colubridae" genus="Rhadinella" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="172" phylum="Chordata" rank="species" species="donaji">
|
||
<emphasis id="B97EEAA03F48FFBB2CD8FBA5FB8A7174" box="[1018,1121,1052,1076]" italics="true" pageId="7" pageNumber="172">R. donaji</emphasis>
|
||
</taxonomicName>
|
||
the lower fourth scale rows have distinct pale dashes which are especially conspicuous on the lower two rows.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |