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<document id="5100DA9472CFE60CC324EE822C498953" ID="10.11646/zootaxa.4267.1.1SLASH11021" ID-CLB-Dataset="33061" ID-DOI="10.5281/zenodo.575641" ID-GBIF-Dataset="810b55e4-b8ba-4368-bf9c-8517ee33e7d0" ID-ISSN="1175-5326" ID-Zenodo-Dep="575641" ID-ZooBank="8920893E-E5F8-482A-A60D-7A248D2B0DCB" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1494577389872" checkinUser="plazi" docAuthor="Skelley, Paul, Xu, Guang, Tang, William, Lindström, Anders J., Marler, Thomas, Khuraijam, Jibankumar Singh, Singh, Rita &amp; Rich, Stephen" docDate="2017" docId="03A38781FFE1FF87FF70FF7CFCA7C6C1" docLanguage="en" docName="zootaxa.4267.1.1SLASH11021.pdf" docOrigin="Zootaxa 4267 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cycadophila (Cycadophila) Xu, Tang, Skelley" docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="FF9AFFF9FFE6FF8FFFE7FFEBFFD5C639" masterDocTitle="Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species" masterLastPageNumber="63" masterPageNumber="1" pageNumber="8" updateTime="1698439510239" updateUser="ExternalLinkService">
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<mods:title id="528A1FA3C42838593087DEEFC5B48F49">Review of Cycadophila Xu, Tang &amp; Skelley (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas (Cycadaceae) in Asia, with descriptions of a new subgenus and thirteen new species</mods:title>
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<mods:namePart id="E74CE68967A0E18FB81F883CA7CBB140">Skelley, Paul</mods:namePart>
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<mods:namePart id="0B803F183A09AF74E620E2A2490E4E47">Xu, Guang</mods:namePart>
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<mods:namePart id="135678E23FEAF79372EA0A9B5B6ABACE">Tang, William</mods:namePart>
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<mods:roleTerm id="3CEF193599FB085FDCC26A1B68E91E28">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA46EA57DD16019FA3E87FD73E74AF5B">Lindström, Anders J.</mods:namePart>
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<mods:name id="C35F9DD70ECBA23B1926A7E34B67291F" type="personal">
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<mods:namePart id="5084864C7C6FC10DAE1B496A2147C1C4">Marler, Thomas</mods:namePart>
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<mods:roleTerm id="41EF0C401540308C85B551F61576513E">Author</mods:roleTerm>
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<mods:namePart id="B96DFE8740A563AFC9B676F2BD6B0366">Khuraijam, Jibankumar Singh</mods:namePart>
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<mods:namePart id="4AFFF34B3DFB354D76A57EC62AB24464">Singh, Rita</mods:namePart>
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<mods:date id="89D0863AA7B0EBD6BEEE5D75F474C255">2017</mods:date>
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<mods:identifier id="05F4709DC2C6AE6FAC109AF0041861B1" type="CLB-Dataset">33061</mods:identifier>
<mods:identifier id="59C835A92C165B7468F2568863F88E49" type="DOI">10.5281/zenodo.575641</mods:identifier>
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<treatment id="03A38781FFE1FF87FF70FF7CFCA7C6C1" ID-DOI="http://doi.org/10.5281/zenodo.5999158" ID-GBIF-Taxon="130809499" ID-Zenodo-Dep="5999158" LSID="urn:lsid:plazi:treatment:03A38781FFE1FF87FF70FF7CFCA7C6C1" httpUri="http://treatment.plazi.org/id/03A38781FFE1FF87FF70FF7CFCA7C6C1" lastPageId="8" lastPageNumber="8" pageId="7" pageNumber="8">
<subSubSection id="C310651CFFE1FF88FF70FF7CFE3AC6C9" pageId="7" pageNumber="8" type="nomenclature">
<paragraph id="8BB53697FFE1FF88FF70FF7CFD16C68B" blockId="7.[151,707,151,178]" box="[151,707,151,178]" pageId="7" pageNumber="8">
<heading id="D0FD81FBFFE1FF88FF70FF7CFD16C68B" bold="true" box="[151,707,151,178]" fontSize="11" level="1" pageId="7" pageNumber="8" reason="1">
<taxonomicName id="4C0A4D14FFE1FF88FF70FF7CFD16C68B" authority="Xu, Tang, Skelley" authorityName="Xu, Tang, Skelley" box="[151,707,151,178]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF70FF7CFD16C68B" bold="true" box="[151,707,151,178]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF70FF7CFEFAC688" bold="true" box="[151,303,151,177]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FED8FF7CFE03C688" bold="true" box="[319,470,151,177]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
) Xu, Tang, Skelley
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF20FF3CFE3AC6C9" blockId="7.[199,709,215,349]" box="[199,495,215,240]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FF3CFE3AC6C9" bold="true" box="[199,495,215,240]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FF3CFF34C6C9" bold="true" box="[199,225,215,240]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FF16FF3CFEDEC6C9" bold="true" box="[241,267,215,240]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<taxonomicName id="4C0A4D14FFE1FF88FEFCFF3CFE9DC6C9" box="[283,328,215,240]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lata">lata</taxonomicName>
species group
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C310651CFFE1FF87FF12FF16FCA7C6C1" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8BB53697FFE1FF88FF12FF16FE2BC764" blockId="7.[199,709,215,349]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FF16FD72C72C" authority="Xu, Tang &amp; Skelley" authorityName="Xu, Tang &amp; Skelley" box="[245,679,252,277]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="debaonica" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FF16FEDAC72D" box="[245,271,253,276]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FF16FEECC72D" box="[287,313,253,276]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFF17FE6BC72C" box="[329,446,252,277]" italics="true" pageId="7" pageNumber="8">debaonica</emphasis>
Xu, Tang &amp; Skelley
</taxonomicName>
*
<taxonomicName id="4C0A4D14FFE1FF88FF12FECBFDABC701" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,638,287,312]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="collina" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FECBFEDAC70E" box="[245,271,288,311]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FECBFEECC70E" box="[287,313,288,311]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFEF4FE42C701" box="[329,407,287,312]" italics="true" pageId="7" pageNumber="8">collina</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD6EFECBFD10C70E" box="[649,709,288,311]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FEAEFE2BC764" authority="Grouvelle" authorityName="Grouvelle" box="[245,510,324,349]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lata" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FEAEFEDAC765" box="[245,271,325,348]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FEAEFEECC765" box="[287,313,325,348]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFEAFFEA0C764" box="[329,373,324,349]" italics="true" pageId="7" pageNumber="8">lata</emphasis>
(Grouvelle)
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF20FE6EFD9FC7A7" blockId="7.[199,685,389,450]" box="[199,586,389,414]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FE6EFD9FC7A7" bold="true" box="[199,586,389,414]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FE6EFF34C7A7" bold="true" box="[199,225,389,414]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FF16FE6EFEDEC7A7" bold="true" box="[241,267,389,414]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<taxonomicName id="4C0A4D14FFE1FF88FEFCFE6EFE77C7A7" box="[283,418,389,414]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fupingensis">fupingensis</taxonomicName>
species group
</emphasis>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FE41FD78C7F8" blockId="7.[199,685,389,450]" box="[245,685,425,450]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FE41FD78C7F8" authority="Skelley, Tang &amp; Xu" authorityName="Skelley, Tang &amp; Xu" box="[245,685,425,450]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fupingensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FE41FEDAC7F8" box="[245,271,426,449]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FE41FEECC7F8" box="[287,313,426,449]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFE42FE1DC7F8" box="[329,456,425,449]" italics="true" pageId="7" pageNumber="8">fupingensis</emphasis>
Skelley, Tang &amp; Xu
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF20FE01FDD7C43A" blockId="7.[199,776,490,875]" box="[199,514,490,515]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FE01FDD7C43A" bold="true" box="[199,514,490,515]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FE01FF34C43A" bold="true" box="[199,225,490,515]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FF16FE01FEDEC43A" bold="true" box="[241,267,490,515]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<taxonomicName id="4C0A4D14FFE1FF88FEFCFE01FE8EC43A" box="[283,347,490,515]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigra">nigra</taxonomicName>
species group
</emphasis>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FDE4FD9CC4AA" blockId="7.[199,776,490,875]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FDE4FDA9C41E" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,636,526,551]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="abyssa" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FDE4FEDAC41F" box="[245,271,527,550]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FDE4FEECC41F" box="[287,313,527,550]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFDE5FE40C41E" box="[329,405,526,551]" italics="true" pageId="7" pageNumber="8">abyssa</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD6FFDE4FD11C41F" box="[648,708,527,550]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FDD8FD58C472" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,653,562,587]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="convexa" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FDD8FEDAC473" box="[245,271,563,586]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FDD8FEECC473" box="[287,313,563,586]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFDDFFE70C472" box="[329,421,564,587]" italics="true" pageId="7" pageNumber="8">convexa</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD70FDD8FD06C473" box="[663,723,563,586]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FDBCFD14C456" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,705,598,623]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="cyclochasma" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FDBCFEDAC457" box="[245,271,599,622]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FDBCFEECC457" box="[287,313,599,622]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFDBDFE0FC456" box="[329,474,598,623]" italics="true" pageId="7" pageNumber="8">cyclochasma</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD2BFDBCFCDDC457" box="[716,776,599,622]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FD90FD9CC4AA" authority="Mader" authorityName="Mader" box="[245,585,634,659]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="discimaculata" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FD90FEDAC4AB" box="[245,271,635,658]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FD90FEECC4AB" box="[287,313,635,658]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFD91FE33C4AA" box="[329,486,634,659]" italics="true" pageId="7" pageNumber="8">discimaculata</emphasis>
(Mader)
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FD74FDF4C4E2" blockId="7.[199,776,490,875]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FD74FD46C48E" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,659,670,695]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="eurynota" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FD74FEDAC48F" box="[245,271,671,694]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FD74FEECC48F" box="[287,313,671,694]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFD74FE78C48F" box="[329,429,671,694]" italics="true" pageId="7" pageNumber="8">eurynota</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD78FD74FD0EC48F" box="[671,731,671,694]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FD28FDF4C4E2" authority="Chujo" authorityName="Chujo" box="[245,545,706,731]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="intermedia" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FD28FEDAC4E3" box="[245,271,707,730]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FD28FEECC4E3" box="[287,313,707,730]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFD29FE17C4E2" box="[329,450,706,731]" italics="true" pageId="7" pageNumber="8">intermedia</emphasis>
(Chûjô)
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FD0CFE2EC4C6" blockId="7.[199,776,490,875]" box="[245,507,742,767]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FD0CFE2EC4C6" authority="Gorham" authorityName="Gorham" box="[245,507,742,767]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigra" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FD0CFEDAC4C7" box="[245,271,743,766]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FD0CFEECC4C7" box="[287,313,743,766]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFD0CFE50C4C7" box="[329,389,743,766]" italics="true" pageId="7" pageNumber="8">nigra</emphasis>
(Gorham)
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FCE0FE21C57E" blockId="7.[199,776,490,875]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FCE0FD5AC51A" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,655,778,803]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="torquata" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FCE0FEDAC51B" box="[245,271,779,802]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FCE0FEECC51B" box="[287,313,779,802]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFCE7FE7DC51A" box="[329,424,780,803]" italics="true" pageId="7" pageNumber="8">torquata</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD7CFCE0FD02C51B" box="[667,727,779,802]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FCC4FE21C57E" authority="Arrow" authorityName="Arrow" box="[245,500,815,839]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="vittata" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FCC4FEDAC57F" box="[245,271,815,838]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FCC4FEECC57F" box="[287,313,815,838]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFCC4FE44C57F" box="[329,401,815,838]" italics="true" pageId="7" pageNumber="8">vittata</emphasis>
(Arrow)
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FCB8FD88C552" blockId="7.[199,776,490,875]" box="[245,605,850,875]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FCB8FD88C552" authority="Grouvelle" authorityName="Grouvelle" box="[245,605,850,875]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yunnanensis" subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FCB8FEDAC553" box="[245,271,851,874]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FCB8FEECC553" box="[287,313,851,874]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFCBFFE06C552" box="[329,467,852,875]" italics="true" pageId="7" pageNumber="8">yunnanensis</emphasis>
(Grouvelle)
</taxonomicName>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF20FC7FFDD8C594" blockId="7.[199,716,916,1013]" box="[199,525,916,941]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FC7FFDD8C594" bold="true" box="[199,525,916,941]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF20FC7FFF34C594" bold="true" box="[199,225,916,941]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FF16FC7FFEDEC594" bold="true" box="[241,267,916,941]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<taxonomicName id="4C0A4D14FFE1FF88FEFCFC7FFEB3C594" box="[283,358,916,941]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="papua">papua</taxonomicName>
species group
</emphasis>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FC53FD19C5CD" blockId="7.[199,716,916,1013]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF12FC53FDA3C5E9" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,630,951,976]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="papua" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FC53FEDAC5F6" box="[245,271,952,975]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FC53FEECC5F6" box="[287,313,952,975]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFC52FE5AC5E9" box="[329,399,953,976]" italics="true" pageId="7" pageNumber="8">papua</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD65FC53FD6BC5F6" box="[642,702,952,975]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FC36FD50C5CC" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,645,988,1013]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="samara" status="sp. nov." subGenus="Cycadophila">
<emphasis id="B97EEA85FFE1FF88FF12FC36FEDAC5CD" box="[245,271,989,1012]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FC36FEECC5CD" box="[287,313,989,1012]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEAEFC36FE48C5CD" box="[329,413,989,1012]" italics="true" pageId="7" pageNumber="8">samara</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD77FC36FD19C5CD" box="[656,716,989,1012]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF70FBF6FD7EC244" blockId="7.[151,774,1053,1258]" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FF70FBF6FD6AC20F" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[151,703,1053,1078]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subGenus" subGenus="Strobilophila">
<emphasis id="B97EEA85FFE1FF88FF70FBF6FE03C20F" bold="true" box="[151,470,1053,1078]" pageId="7" pageNumber="8">
<emphasis id="B97EEA85FFE1FF88FF70FBF6FEFDC20F" bold="true" box="[151,296,1053,1078]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEDFFBF6FE1BC20F" bold="true" box="[312,462,1053,1078]" italics="true" pageId="7" pageNumber="8">Strobilophila</emphasis>
)
</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
, n. sg.
<taxonomicName id="4C0A4D14FFE1FF88FF12FBA9FD7CC263" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,681,1089,1114]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="assamensis" status="sp. nov." subGenus="Strobilophila">
<emphasis id="B97EEA85FFE1FF88FF12FBA9FEDAC260" box="[245,271,1090,1113]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FBA9FEE1C260" box="[287,308,1090,1113]" italics="true" pageId="7" pageNumber="8">S.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEA3FBA9FE17C260" box="[324,450,1090,1113]" italics="true" pageId="7" pageNumber="8">assamensis</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD52FBA9FD24C260" box="[693,753,1090,1113]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FB8DFDB1C247" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,612,1125,1150]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="hiepi" status="sp. nov." subGenus="Strobilophila">
<emphasis id="B97EEA85FFE1FF88FF12FB8DFEDAC244" box="[245,271,1126,1149]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FB8DFEE1C244" box="[287,308,1126,1149]" italics="true" pageId="7" pageNumber="8">S.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEA3FB8EFEA9C247" box="[324,380,1125,1150]" italics="true" pageId="7" pageNumber="8">hiepi</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD88FB8DFD7EC244" box="[623,683,1126,1149]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB53697FFE1FF88FF12FB61FD64C2D0" blockId="7.[151,774,1053,1258]" pageId="7" pageNumber="8">
C.
<taxonomicName id="4C0A4D14FFE1FF88FEF2FB61FD4CC29B" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[277,665,1161,1186]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kwaiensis" status="sp. nov." subGenus="Strobilophila">
(
<emphasis id="B97EEA85FFE1FF88FEF8FB61FEF8C298" box="[287,301,1162,1185]" italics="true" pageId="7" pageNumber="8">S</emphasis>
.)
<emphasis id="B97EEA85FFE1FF88FEA3FB62FE64C29B" box="[324,433,1161,1186]" italics="true" pageId="7" pageNumber="8">kwaiensis</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD43FB61FD35C298" box="[676,736,1162,1185]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
<taxonomicName id="4C0A4D14FFE1FF88FF12FB45FD4CC2FF" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,665,1197,1222]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="tansachai" status="sp. nov." subGenus="Strobilophila">
<emphasis id="B97EEA85FFE1FF88FF12FB45FEDAC2FC" box="[245,271,1198,1221]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FB45FEE1C2FC" box="[287,308,1198,1221]" italics="true" pageId="7" pageNumber="8">S.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEA3FB46FE64C2FF" box="[324,433,1197,1222]" italics="true" pageId="7" pageNumber="8">tansachai</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD43FB45FD35C2FF" box="[676,736,1198,1222]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
*
<taxonomicName id="4C0A4D14FFE1FF88FF12FB39FDBFC2D3" authority="Skelley, Xu &amp; Tang" authorityName="Skelley, Xu &amp; Tang" box="[245,618,1233,1258]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yangi" status="sp. nov." subGenus="Strobilophila">
<emphasis id="B97EEA85FFE1FF88FF12FB39FEDAC2D0" box="[245,271,1234,1257]" italics="true" pageId="7" pageNumber="8">C.</emphasis>
(
<emphasis id="B97EEA85FFE1FF88FEF8FB39FEE1C2D0" box="[287,308,1234,1257]" italics="true" pageId="7" pageNumber="8">S.</emphasis>
)
<emphasis id="B97EEA85FFE1FF88FEA3FB39FE57C2D0" box="[324,386,1234,1257]" italics="true" pageId="7" pageNumber="8">yangi</emphasis>
Skelley, Xu &amp; Tang
</taxonomicName>
,
<taxonomicNameLabel id="A24D57FEFFE1FF88FD92FB39FD64C2D0" box="[629,689,1234,1257]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
<paragraph id="8BB53697FFE1FF87FF20FADCFCA7C6C1" blockId="7.[151,1437,1335,2009]" lastBlockId="8.[151,1436,151,248]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
The PCR product of the 16S rRNA gene had a length of 311–313 bp. The alignment matrix was 317 bp long, with 5 gaps, 218 conserved sites, 96 variable sites, 67 parsimony-informative sites and 29 singleton sites. The NJ, MP and ML trees derived from DNA analyses were largely congruent with essentially the same terminal clades and no rearrangement of clades relative to one another. Only the one derived from ML is presented here (
<figureCitation id="13312A12FFE1FF88FAE8FA4FFA84C385" box="[1295,1361,1444,1469]" captionStart="FIGURE 1" captionStartId="6.[151,250,1875,1897]" captionTargetBox="[220,1315,214,1828]" captionTargetId="figure@6.[212,1360,195,1836]" captionTargetPageId="6" captionText="FIGURE 1. Phylogenetic tree based on maximum likelihood analysis of 16 S rRNA gene sequences of four species of New World Pharaxonotha and ten species of Asian Cycadophila. Scale bar indicates base pair substitutions per nucleotide position; numbers on the branches of the trees are bootstrap values." httpUri="https://zenodo.org/record/575642/files/figure.png" pageId="7" pageNumber="8">Fig. 1</figureCitation>
). This tree is similar to those presented by
<bibRefCitation id="EF9B4B66FFE1FF88FDC9FA23FD31C3D9" box="[558,740,1479,1504]" pageId="7" pageNumber="8" refString="Xu, G., Tang, W., Skelley, P., Liu, N., &amp; Rich, S. (2015) Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa, 3986 (3), 251 - 278." type="journal article">
Xu
<emphasis id="B97EEA85FFE1FF88FDB0FA22FD45C3D9" box="[599,656,1479,1504]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2015)
</bibRefCitation>
, but is based on a larger sample of populations and species of
<taxonomicName id="4C0A4D14FFE1FF88FF70FA07FEF3C03C" box="[151,294,1516,1541]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE1FF88FF70FA07FEF3C03C" box="[151,294,1516,1541]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
</taxonomicName>
and includes an additional (undescribed) taxon within the subfamily
<taxonomicName id="4C0A4D14FFE1FF88FBCAFA07FB3CC03C" box="[1069,1257,1516,1541]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
as an outgroup. The same clades and branching patterns are present in our tree as in
<bibRefCitation id="EF9B4B66FFE1FF88FC6AF9FBFB94C011" box="[909,1089,1551,1576]" pageId="7" pageNumber="8" refString="Xu, G., Tang, W., Skelley, P., Liu, N., &amp; Rich, S. (2015) Cycadophila, a new genus (Coleoptera: Erotylidae: Pharaxonothinae) inhabiting Cycas debaoensis (Cycadaceae) in Asia. Zootaxa, 3986 (3), 251 - 278." type="journal article">
Xu
<emphasis id="B97EEA85FFE1FF88FC51F9FAFC3BC011" box="[950,1006,1551,1576]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
(2015)
</bibRefCitation>
, except that no members of the
<emphasis id="B97EEA85FFE1FF88FF70F9DFFEFCC074" box="[151,297,1588,1613]" italics="true" pageId="7" pageNumber="8">Strobilophila</emphasis>
clade and
<taxonomicName id="4C0A4D14FFE1FF88FE44F9DEFE33C075" box="[419,486,1589,1612]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="papua">papua</taxonomicName>
clade were analyzed in the previous study. In the tree presented here
<taxonomicName id="4C0A4D14FFE1FF88FB12F9DFFA51C074" box="[1269,1412,1588,1613]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE1FF88FB12F9DFFA51C074" box="[1269,1412,1588,1613]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
</taxonomicName>
+
<taxonomicName id="4C0A4D14FFE1FF88FF70F9BCFEE2C049" box="[151,311,1623,1648]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE1FF88FF70F9BCFEE2C049" box="[151,311,1623,1648]" italics="true" pageId="7" pageNumber="8">Pharaxonotha</emphasis>
</taxonomicName>
are a monophyletic grouping considered together in relation to the other members of the
<taxonomicName id="4C0A4D14FFE1FF88FF70F997FE86C0AC" box="[151,339,1660,1685]" class="Insecta" family="Erotylidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="subFamily" subFamily="Pharaxonothinae">Pharaxonothinae</taxonomicName>
sampled (bootstrap support = 100). Bootstrap support for the
<taxonomicName id="4C0A4D14FFE1FF88FBF6F997FB64C0AC" box="[1041,1201,1660,1685]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE1FF88FBF6F997FB64C0AC" box="[1041,1201,1660,1685]" italics="true" pageId="7" pageNumber="8">Pharaxonotha</emphasis>
</taxonomicName>
(
<emphasis id="B97EEA85FFE1FF88FB25F996FA9AC0AD" box="[1218,1359,1661,1684]" italics="true" pageId="7" pageNumber="8">sensu stricto</emphasis>
) clade is also strong (&gt;90). Within this New World clade
<taxonomicName id="4C0A4D14FFE1FF88FD2BF94AFCEEC081" box="[716,827,1695,1720]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="kirschii">
<emphasis id="B97EEA85FFE1FF88FD2BF94AFCEEC081" box="[716,827,1695,1720]" italics="true" pageId="7" pageNumber="8">P. kirschii</emphasis>
</taxonomicName>
is sister to
<taxonomicName id="4C0A4D14FFE1FF88FC5BF94AFBFBC081" box="[956,1070,1696,1720]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="confusa">
<emphasis id="B97EEA85FFE1FF88FC5BF94AFBFBC081" box="[956,1070,1696,1720]" italics="true" pageId="7" pageNumber="8">P. confusa</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0A4D14FFE1FF88FBDBF94AFB0EC081" box="[1084,1243,1695,1720]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="portophylla">
<emphasis id="B97EEA85FFE1FF88FBDBF94AFB0EC081" box="[1084,1243,1695,1720]" italics="true" pageId="7" pageNumber="8">P. portophylla</emphasis>
</taxonomicName>
and
<emphasis id="B97EEA85FFE1FF88FAF6F94AFA48C081" box="[1297,1437,1695,1720]" italics="true" pageId="7" pageNumber="8">
<taxonomicName id="4C0A4D14FFE1FF88FAF6F94AFA4CC081" box="[1297,1433,1695,1720]" class="Insecta" family="Erotylidae" genus="Pharaxonotha" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="floridana">P. floridana</taxonomicName>
.
</emphasis>
With the addition of more populations and taxa to the analysis the genus
<taxonomicName id="4C0A4D14FFE1FF88FC1AF92FFB59C0E4" box="[1021,1164,1732,1757]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE1FF88FC1AF92FFB59C0E4" box="[1021,1164,1732,1757]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
</taxonomicName>
remains monophyletic, although bootstrap support is relatively weak (&lt;40). Within the subgenus
<taxonomicName id="4C0A4D14FFE1FF88FC37F90CFB8AC139" box="[976,1119,1767,1792]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFE1FF88FC37F90CFB8AC139" box="[976,1119,1767,1792]" italics="true" pageId="7" pageNumber="8">Cycadophila</emphasis>
</taxonomicName>
this analysis recovered four strongly supported (bootstrap&gt;90) clades: 1) the
<taxonomicName id="4C0A4D14FFE1FF88FD59F8E7FD33C11C" box="[702,742,1804,1829]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lata">lata</taxonomicName>
species group (only
<taxonomicName id="4C0A4D14FFE1FF88FC28F8E6FBB1C11C" box="[975,1124,1804,1829]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="debaonica">
<emphasis id="B97EEA85FFE1FF88FC28F8E6FBB1C11C" box="[975,1124,1804,1829]" italics="true" pageId="7" pageNumber="8">C. debaonica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D14FFE1FF88FB7DF8E6FADCC11C" box="[1178,1289,1804,1829]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="collina">
<emphasis id="B97EEA85FFE1FF88FB7DF8E6FADCC11C" box="[1178,1289,1804,1829]" italics="true" pageId="7" pageNumber="8">C. collina</emphasis>
</taxonomicName>
sampled); 2) the
<taxonomicName id="4C0A4D14FFE1FF88FF58F8C4FF2CC171" box="[191,249,1839,1864]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigra">nigra</taxonomicName>
species group (only
<taxonomicName id="4C0A4D14FFE1FF88FE38F8DBFD5FC171" box="[479,650,1840,1864]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yunnanensis">
<emphasis id="B97EEA85FFE1FF88FE38F8DBFD5FC171" box="[479,650,1840,1864]" italics="true" pageId="7" pageNumber="8">C. yunnanensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0A4D14FFE1FF88FD58F8DBFCC9C171" box="[703,796,1840,1864]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B97EEA85FFE1FF88FD58F8DBFCC9C171" box="[703,796,1840,1864]" italics="true" pageId="7" pageNumber="8">C. nigra</emphasis>
</taxonomicName>
sampled); 3) the
<taxonomicName id="4C0A4D14FFE1FF88FC3AF8C4FB8BC171" box="[989,1118,1839,1864]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fupingensis">fupingensis</taxonomicName>
species group (consisting of a single species
<taxonomicName id="4C0A4D14FFE1FF88FEBFF8BEFE28C155" box="[344,509,1876,1900]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fupingensis">
<emphasis id="B97EEA85FFE1FF88FEBFF8BEFE28C155" box="[344,509,1876,1900]" italics="true" pageId="7" pageNumber="8">C. fupingensis</emphasis>
</taxonomicName>
); and 4) the
<taxonomicName id="4C0A4D14FFE1FF88FD7AF8BEFD35C155" box="[669,736,1877,1900]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="papua">papua</taxonomicName>
species group (only
<taxonomicName id="4C0A4D14FFE1FF88FC3EF8BEFB87C155" box="[985,1106,1877,1900]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="samara">
<emphasis id="B97EEA85FFE1FF88FC3EF8BEFB87C155" box="[985,1106,1877,1900]" italics="true" pageId="7" pageNumber="8">C. samara</emphasis>
</taxonomicName>
sampled). The relationships among these four species groups remains unclear, since bootstrap support at nodes is relatively low (&lt;40). For the subgenus
<emphasis id="B97EEA85FFE1FF88FEE0F877FE4CC18C" box="[263,409,1948,1973]" italics="true" pageId="7" pageNumber="8">Strobilophila</emphasis>
, six of seven populations (three of four species) analyzed form a monophyletic group with one outlier [see more discussion below under Key to larvae and Remarks for
<emphasis id="B97EEA85FFE1FF88FBF2F82AFBFDC1E1" box="[1045,1064,1985,2008]" italics="true" pageId="7" pageNumber="8">C</emphasis>
. (
<emphasis id="B97EEA85FFE1FF88FBA6F82AFB9AC1E1" box="[1089,1103,1985,2008]" italics="true" pageId="7" pageNumber="8">S</emphasis>
).
<taxonomicName id="4C0A4D14FFE1FF88FB8EF82AFB72C1E1" box="[1129,1191,1985,2008]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="yangi">
<emphasis id="B97EEA85FFE1FF88FB8EF82AFB72C1E1" box="[1129,1191,1985,2008]" italics="true" pageId="7" pageNumber="8">yangi</emphasis>
</taxonomicName>
]. Although bootstrap support for many nodes within the genus
<taxonomicName id="4C0A4D14FFEEFF87FD8AFF7CFD29C689" box="[621,764,151,176]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFEEFF87FD8AFF7CFD29C689" box="[621,764,151,176]" italics="true" pageId="8" pageNumber="9">Cycadophila</emphasis>
</taxonomicName>
clade may be considered weak, our molecular data support our morphological hypotheses of relationships for this genus. Together molecular and morphological data support our concept of the genus
<taxonomicName id="4C0A4D14FFEEFF87FE56FF34FD95C6C1" box="[433,576,223,248]" class="Insecta" family="Erotylidae" genus="Cycadophila" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B97EEA85FFEEFF87FE56FF34FD95C6C1" box="[433,576,223,248]" italics="true" pageId="8" pageNumber="9">Cycadophila</emphasis>
</taxonomicName>
as a monophyletic lineage.
</paragraph>
</subSubSection>
</treatment>
</document>
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