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<document id="B200F0E66ED7C563253D77771BE56B5C" ID-ISSN="2336-9744" ID-ZooBank="urn:lsid:zoobank.org:pub:86F17262-17A8-40FF-88B9-2D4552A92F12" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1720662477596" checkinUser="felipe" docAuthor="Volynkin, Anton V., Huang, Si-Yao &amp; Ivanova, Maria S." docDate="2019" docId="03A2A079553B8045805346A2A3151140" docLanguage="en" docName="EcolMontenegrina.26.14-92.pdf" docOrigin="Ecologica Montenegrina 26" docStyle="DocumentStyle:3485E46B0EDC64DBF2992EDE30BFB5AE.2:EcolMontenegrina.2017-.journal_article" docStyleId="3485E46B0EDC64DBF2992EDE30BFB5AE" docStyleName="EcolMontenegrina.2017-.journal_article" docStyleVersion="2" docTitle="Barsine Walker 1854" docType="treatment" docVersion="1" lastPageNumber="17" masterDocId="FF9BD80155398046823D4076A72A127E" masterDocTitle="An overview of genera and subgenera of the Asura / Miltochrista generic complex (Lepidoptera, Erebidae, Arctiinae). Part 1. Barsine Walker, 1854 sensu lato, Asura Walker, 1854 and related genera, with descriptions of twenty new genera, ten new subgenera and a check list of taxa of the Asura / Miltochrista generic complex" masterLastPageNumber="92" masterPageNumber="14" pageNumber="16" updateTime="1720672203402" updateUser="GgImagineBatch" zenodo-license-document="CC-BY-4.0">
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<mods:title id="E26B21281103BBC4DEFD2C476BB8082E">An overview of genera and subgenera of the Asura / Miltochrista generic complex (Lepidoptera, Erebidae, Arctiinae). Part 1. Barsine Walker, 1854 sensu lato, Asura Walker, 1854 and related genera, with descriptions of twenty new genera, ten new subgenera and a check list of taxa of the Asura / Miltochrista generic complex</mods:title>
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<mods:namePart id="D452FD170DD032CAE0E4B2832DCB9BC4">Volynkin, Anton V.</mods:namePart>
<mods:affiliation id="99AF445B09D8ED8FEB6D3C4F38439531">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia &amp; National Research Tomsk State University, Lenina Avenue, 36, RF- 634050, Tomsk, Russia</mods:affiliation>
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<mods:namePart id="581FED6892BED4A4D8B64E4A72BDF4E9">Huang, Si-Yao</mods:namePart>
<mods:affiliation id="1D4DC325483B332577843E5088B130D4">Department of Entomology, College of Agriculture, South China Agricultural University, Guangzhou, 510642, Guangdong, China * Corresponding author. E-mail: volynkin _ a @ mail. ru</mods:affiliation>
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<mods:namePart id="EB267D9A21697B342DC9E5EA9EB8F59C">Ivanova, Maria S.</mods:namePart>
<mods:affiliation id="D16F0BBF114D347E9CA5951D63A39208">Altai State University, Lenina Avenue, 61, RF- 656049, Barnaul, Russia</mods:affiliation>
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<paragraph id="8BB4116F553B8044805346A2A4EF1491" blockId="2.[622,965,1748,1808]" box="[622,965,1748,1775]" pageId="2" pageNumber="16">
<heading id="D0FCA603553B8044805346A2A4EF1491" box="[622,965,1748,1775]" centered="true" fontSize="36" level="2" pageId="2" pageNumber="16" reason="4">
Genus
<taxonomicName id="4C0B6AEC553B804480FD46A2A4EF1491" ID-CoL="93ND4" authority="Walker, 1854" authorityName="Walker" authorityYear="1854" box="[704,965,1748,1775]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B97FCD7D553B804480FD46A2A4351490" bold="true" box="[704,799,1748,1774]" italics="true" pageId="2" pageNumber="16">Barsine</emphasis>
Walker, 1854
</taxonomicName>
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</paragraph>
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<subSubSection id="C31142E4553B8044809F4683A3201597" pageId="2" pageNumber="16" type="description">
<paragraph id="8BB4116F553B8044809F4683A4BB156E" blockId="2.[622,965,1748,1808]" box="[674,913,1781,1808]" pageId="2" pageNumber="16">
(
<figureCitation id="13300DEA553B804480974683A43E156E" box="[682,788,1781,1808]" captionStart="Figures 116" captionStartId="4.[151,236,1914,1938]" captionTargetBox="[152,1436,189,1858]" captionTargetId="figure-19@4.[151,1436,189,1877]" captionTargetPageId="4" captionText="Figures 116. Type species of genera and subgenera of the Asura / Miltochrista generic complex: adults. Depositories of specimens: 13, 79, 1216 MWM/ZSM; 4, 10 NHMUK; 5, 6, 11 CKC." pageId="2" pageNumber="16">Figs 1, 2</figureCitation>
,
<figureCitation id="13300DEA553B8044811D4683A465156E" box="[800,847,1781,1808]" captionStart="Figures 104107" captionStartId="26.[151,236,1964,1988]" captionTargetBox="[154,1432,362,1884]" captionTargetId="figure-58@26.[151,1436,357,1896]" captionTargetPageId="26" captionText="Figures 104107. Type species of genera and subgenera of the Asura / Miltochrista generic complex: male genitalia." pageId="2" pageNumber="16">104</figureCitation>
,
<figureCitation id="13300DEA553B804481664683A4A2156E" box="[859,904,1781,1808]" captionStart="Figures 160165" captionStartId="41.[151,236,1938,1962]" captionTargetBox="[161,1408,290,1852]" captionTargetId="figure-19@41.[151,1436,290,1870]" captionTargetPageId="41" captionText="Figures 160165. Type species of genera and subgenera of the Asura / Miltochrista generic complex: female genitalia." pageId="2" pageNumber="16">160</figureCitation>
)
</paragraph>
<paragraph id="8BB4116F553B80448280474FA34B1510" blockId="2.[151,1436,1847,2026]" pageId="2" pageNumber="16">
<taxonomicName id="4C0B6AEC553B80448280474FA687152E" authority="Walker, 1854" authorityName="Walker" authorityYear="1854" box="[189,429,1848,1872]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B97FCD7D553B80448280474FA63A152E" box="[189,272,1849,1872]" italics="true" pageId="2" pageNumber="16">Barsine</emphasis>
Walker, 1854
</taxonomicName>
,
<emphasis id="B97FCD7D553B80448384474FA265152E" box="[441,1359,1847,1872]" italics="true" pageId="2" pageNumber="16">List of the specimens of lepidopterous insects in the collection of the British Museum</emphasis>
<emphasis id="B97FCD7D553B80448765474EA24F152E" bold="true" box="[1368,1381,1848,1872]" pageId="2" pageNumber="16">2</emphasis>
: 546 (
<typeStatus id="54B0AFCD553B8044829D4721A7FC1511" box="[160,214,1879,1903]" pageId="2" pageNumber="16">Type</typeStatus>
species:
<taxonomicName id="4C0B6AEC553B804483054721A5501510" authority="Walker, 1854" authorityName="Walker" authorityYear="1854" box="[312,634,1878,1902]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="species" species="defecta">
<emphasis id="B97FCD7D553B804483054721A6F51510" box="[312,479,1878,1902]" italics="true" pageId="2" pageNumber="16">Barsine defecta</emphasis>
Walker, 1854
</taxonomicName>
, by subsequent designation by
<bibRefCitation id="EF9A6C9E553B804481FB4720A3771510" author="Kirby, W. F." box="[966,1117,1878,1902]" pageId="2" pageNumber="16" pagination="1 - 951" refId="ref32248" refString="Kirby, W. F. (1892) Sphinges and Bombyces. A synonymic catalogue of Lepidoptera Heterocera. (Moths), 1, 1 - 951." type="journal article" year="1892">Kirby (1892))</bibRefCitation>
.
</paragraph>
<paragraph id="8BB4116F553B804482804700A44415D2" blockId="2.[151,1436,1847,2026]" pageId="2" pageNumber="16">
=
<taxonomicName id="4C0B6AEC553B804482EE4703A6F115F3" authority="Walker, 1864" authorityName="Walker" authorityYear="1864" box="[211,475,1909,1933]" class="Insecta" family="Erebidae" genus="Castabala" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B97FCD7D553B804482EE4703A66A15F3" box="[211,320,1909,1933]" italics="true" pageId="2" pageNumber="16">Castabala</emphasis>
Walker, 1864
</taxonomicName>
,
<emphasis id="B97FCD7D553B804483DA4700A25815F3" box="[487,1394,1909,1933]" italics="true" pageId="2" pageNumber="16">List of the specimens of lepidopterous insects in the collection of the British Museum</emphasis>
<emphasis id="B97FCD7D553B804487474703A2BF15F3" bold="true" box="[1402,1429,1909,1933]" pageId="2" pageNumber="16">31</emphasis>
: 270 (
<typeStatus id="54B0AFCD553B804482F247E3A62F15D3" box="[207,261,1941,1965]" pageId="2" pageNumber="16">Type</typeStatus>
species:
<taxonomicName id="4C0B6AEC553B8044835A47E2A5EC15D2" authority="Walker, 1864" authorityName="Walker" authorityYear="1864" box="[359,710,1940,1964]" class="Insecta" family="Erebidae" genus="Castabala" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="species" species="roseata">
<emphasis id="B97FCD7D553B8044835A47E2A50015D2" box="[359,554,1940,1964]" italics="true" pageId="2" pageNumber="16">Castabala roseata</emphasis>
Walker, 1864
</taxonomicName>
, by monotypy).
</paragraph>
<paragraph id="8BB4116F553B8044828047C5A3201597" blockId="2.[151,1436,1847,2026]" pageId="2" pageNumber="16">
=
<taxonomicName id="4C0B6AEC553B804482E847C4A6FC15B4" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[213,470,1970,1994]" class="Insecta" family="Erebidae" genus="Mahavira" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="genus">
<emphasis id="B97FCD7D553B804482E847C4A61415B4" box="[213,318,1970,1994]" italics="true" pageId="2" pageNumber="16">Mahavira</emphasis>
<materialsCitation id="3B631B32553B8044837A47C5A6FC15B4" box="[327,470,1970,1994]" location="Proceedings" municipality="Zoological Society of London" pageId="2" pageNumber="16" specimenCount="1">Moore, 1878</materialsCitation>
</taxonomicName>
,
<emphasis id="B97FCD7D553B804483DE47C4A7C31597" italics="true" pageId="2" pageNumber="16">
<location id="8ED447B4553B804483DE47C4A54D15B4" LSID="urn:lsid:plazi:treatment:03A2A079553B8045805346A2A3151140:8ED447B4553B804483DE47C4A54D15B4" box="[483,615,1970,1994]" municipality="Zoological Society of London" name="Proceedings" pageId="2" pageNumber="16">Proceedings</location>
of the general meetings for scientific business of the
<collectingMunicipality id="6BD08B15553B8044868847C4A7C31597" pageId="2" pageNumber="16">Zoological Society of London</collectingMunicipality>
</emphasis>
<emphasis id="B97FCD7D553B804482CD47A4A60F1594" bold="true" box="[240,293,2002,2026]" pageId="2" pageNumber="16">1878</emphasis>
: 11 (
<typeStatus id="54B0AFCD553B8044836047A4A6B91594" box="[349,403,2002,2026]" pageId="2" pageNumber="16">Type</typeStatus>
species:
<taxonomicName id="4C0B6AEC553B804483C847A7A4481597" authority="Moore, 1878" authorityName="Moore" authorityYear="1878" box="[501,866,2001,2025]" class="Insecta" family="Erebidae" genus="Mahavira" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="16" phylum="Arthropoda" rank="species" species="flavicollis">
<emphasis id="B97FCD7D553B804483C847A7A5E51597" box="[501,719,2001,2025]" italics="true" pageId="2" pageNumber="16">Mahavira flavicollis</emphasis>
Moore, 1878
</taxonomicName>
, by monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C31142E4553A8045828040B2A65F10C9" pageId="3" pageNumber="17" type="diagnosis">
<paragraph id="8BB4116F553A8045828040B2A65F10C9" blockId="3.[151,1436,195,830]" pageId="3" pageNumber="17">
<emphasis id="B97FCD7D553A8045828040B2A61C12A0" bold="true" box="[189,310,196,222]" pageId="3" pageNumber="17">Diagnosis</emphasis>
. Medium-sized moths with yellow, orange, red or white ground colour and pattern consisting of black elements on veins and reddish spots between veins. In some species yellow forms without red spots are known. In
<taxonomicName id="4C0B6AEC553A8045837D417EA69F135C" authorityName="Walker" authorityYear="1854" box="[320,437,263,290]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="defecta">
<emphasis id="B97FCD7D553A8045837D417EA69F135C" box="[320,437,263,290]" italics="true" pageId="3" pageNumber="17">B. defecta</emphasis>
</taxonomicName>
all females have no red spots. In some populations of some other taxa (
<taxonomicName id="4C0B6AEC553A8045873D417EA2B6135F" baseAuthorityName="de Joannis" baseAuthorityYear="1930" box="[1280,1436,264,289]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="gratissima">
<emphasis id="B97FCD7D553A8045873D417EA2B6135F" box="[1280,1436,264,289]" italics="true" pageId="3" pageNumber="17">B. gratissima</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B6AEC553A804582F7415CA6FB133A" authorityName="Černý &amp; Pinratana" authorityYear="2009" box="[202,465,297,324]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="17" phylum="Arthropoda" rank="subSpecies" species="orientalis" subSpecies="bigamica">
<emphasis id="B97FCD7D553A804582F7415CA6FB133A" box="[202,465,297,324]" italics="true" pageId="3" pageNumber="17">B. orientalis bigamica</emphasis>
</taxonomicName>
) such yellow coloured females are also common. In some species (
<taxonomicName id="4C0B6AEC553A804586C5415CA2BE133A" baseAuthorityName="Hampson" baseAuthorityYear="1900" box="[1272,1428,297,324]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="perpallida">
<emphasis id="B97FCD7D553A804586C5415CA2BE133A" box="[1272,1428,297,324]" italics="true" pageId="3" pageNumber="17">B. perpallida</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0B6AEC553A804582AA413DA614131B" authorityName="Volynkin, Černý &amp; Huang" authorityYear="2019" box="[151,318,330,357]" class="Insecta" family="Erebidae" genus="Barsine" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="andromeda">
<emphasis id="B97FCD7D553A804582AA413DA614131B" box="[151,318,330,357]" italics="true" pageId="3" pageNumber="17">B. andromeda</emphasis>
</taxonomicName>
and members of the
<taxonomicName id="4C0B6AEC553A80458001413DA5F8131B" authorityName="Moore" authorityYear="1878" box="[572,722,330,357]" class="Insecta" family="Erebidae" genus="Mahavira" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="17" phylum="Arthropoda" rank="species" species="flavicollis">
<emphasis id="B97FCD7D553A80458001413DA5F8131B" box="[572,722,330,357]" italics="true" pageId="3" pageNumber="17">B. flavicollis</emphasis>
</taxonomicName>
species-group) the wing pattern is completely reduced. The main autapomorphic feature of the male genitalia is the presence of a long and robust basal saccular process. In addition, the male genitalia are characterised by the combination of the following features: (1) the weakly sclerotised, more or less X-shaped juxta consisting of two lateral parts connected by a weak membrane; (2) the presence of a robust medial costal process, which is usually broadened basally, apically curved and weakly setose; (3) the narrow distal section of costa protruding distally beyond the medial process and sometimes having a small distal process; (4) the presence of a distal membranous lobe of valva having a broad and elongated base; (5) the sacculus being distally narrow and having a robust distal process being well-separated from the valva surface; in many species this process has an additional dorsal lobe subbasally; (6) the aedeagus vesica having several well-separated diverticula bearing fields of small stout cornuti or robust granulation.
</paragraph>
</subSubSection>
<subSubSection id="C31142E4553A8045828042C8A3151140" pageId="3" pageNumber="17" type="distribution">
<paragraph id="8BB4116F553A8045828042C8A4AB1162" blockId="3.[151,1436,195,830]" pageId="3" pageNumber="17">
<emphasis id="B97FCD7D553A8045828042C8A67710A6" bold="true" box="[189,349,702,728]" pageId="3" pageNumber="17">Distribution.</emphasis>
The genus is widespread from
<collectingCountry id="F31C51FF553A804580E742CBA43C10A6" box="[730,790,701,728]" name="India" pageId="3" pageNumber="17">India</collectingCountry>
and
<collectingCountry id="F31C51FF553A8045816942CBA4B010A6" box="[852,922,701,728]" name="Nepal" pageId="3" pageNumber="17">Nepal</collectingCountry>
through Mainland
<collectingCountry id="F31C51FF553A804586BF42CBA3ED10A6" box="[1154,1223,701,728]" name="China" pageId="3" pageNumber="17">China</collectingCountry>
and Indochina to Sundaland including the Philippine island of Palawan. Some species are known from Russian Far East, Korean Peninsula,
<collectingCountry id="F31C51FF553A804583484374A6921162" box="[373,440,770,796]" name="Japan" pageId="3" pageNumber="17">Japan</collectingCountry>
and the islands of
<collectingCountry id="F31C51FF553A804580A84377A5C61162" box="[661,748,769,796]" name="Taiwan" pageId="3" pageNumber="17">Taiwan</collectingCountry>
and Hainan.
</paragraph>
<paragraph id="8BB4116F553A804582804355A3151140" blockId="3.[151,1436,195,830]" box="[189,1087,803,830]" pageId="3" pageNumber="17">
<emphasis id="B97FCD7D553A804582804355A6891143" bold="true" box="[189,419,803,829]" pageId="3" pageNumber="17">Number of species</emphasis>
. The genus comprises 63 valid species and 7 subspecies.
</paragraph>
</subSubSection>
</treatment>
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