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<document id="1F983016BE2FE15D4B572661AAEC8A8C" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CA92FF65EF22FDA94274FD00" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Paragomphus cammaertsi Dijkstra &amp; Papazian, sp. nov." docType="treatment" docVersion="19" lastPageNumber="588" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="584" updateTime="1695001031402" updateUser="admin">
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<mods:title id="B51590FF94AE089D684C7F1AF62726BF">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
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<mods:namePart id="C378ACF9739C476F864710E402E3E342">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="6BEDCE61B8C9E542FBC87378F5B1A22E">Kipping, Jens</mods:namePart>
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<mods:namePart id="5D5C8BC7CB40A856646AB3B78D8D48F8">Mézière, Nicolas</mods:namePart>
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<mods:title id="C023C7701AF2FEC661AACF4026C5723B">Odonatologica</mods:title>
</mods:titleInfo>
<mods:part id="619CF6E333AEBDF8C4A9F618AF992440">
<mods:date id="934E75B8C35390CD37AA788EFF6045BF">2015</mods:date>
<mods:detail id="8F1398CE9F5C6DC6C9404F77EB65DF09" type="volume">
<mods:number id="333F02D347A4FF940823858C1F0E3567">44</mods:number>
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<mods:classification id="2F1EA2759AF762A397D3BB22CFFC8B82">journal article</mods:classification>
<mods:identifier id="F08C8B135ECC67C7C1B0ED529B974D74" type="DOI">10.5281/zenodo.35388</mods:identifier>
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<treatment id="03A25264CA92FF65EF22FDA94274FD00" ID-DOI="http://doi.org/10.5281/zenodo.5640266" ID-GBIF-Taxon="114923475" ID-Zenodo-Dep="5640266" LSID="urn:lsid:zoobank.org:act:03A25264-CA92-FF65-EF22-FDA94274FD00" httpUri="http://treatment.plazi.org/id/03A25264CA92FF65EF22FDA94274FD00" lastPageId="143" lastPageNumber="588" pageId="139" pageNumber="584">
<subSubSection id="C311B0F9CA92FF61EF22FDA943DFFD30" pageId="139" pageNumber="584" type="nomenclature">
<paragraph id="8BB4E372CA92FF61EF22FDA943DFFD30" pageId="139" pageNumber="584">
<heading id="D0FC541ECA92FF61EF22FDA943DFFD30" bold="true" centered="true" fontSize="11" level="2" pageId="139" pageNumber="584" reason="4">
<emphasis id="B97F3F60CA92FF61EF22FDA942AFFDED" bold="true" box="[256,938,580,608]" pageId="139" pageNumber="584">
<taxonomicName id="4C0B98F1CA92FF61EF22FDA94237FDED" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CA92-FF65-EF22-FDA94274FD00" authority="Dijkstra &amp; Papazian" authorityName="Dijkstra &amp; Papazian" box="[256,818,580,608]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="cammaertsi" status="sp. nov.">
<emphasis id="B97F3F60CA92FF61EF22FDA9433AFDED" bold="true" box="[256,575,580,608]" italics="true" pageId="139" pageNumber="584">Paragomphus cammaertsi</emphasis>
Dijkstra &amp; Papazian
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA92FF61ED1AFDA94290FDED" box="[824,917,580,607]" pageId="139" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
–
</emphasis>
<emphasis id="B97F3F60CA92FF61EFF2FD8843DFFD30" bold="true" box="[464,730,613,642]" pageId="139" pageNumber="584">Cammaerts’s Hooktail</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA92FF61EFEBFD7C401AFD68" pageId="139" pageNumber="584" type="description">
<paragraph id="8BB4E372CA92FF61EFEBFD7C43E4FD1F" box="[457,737,657,685]" pageId="139" pageNumber="584">
<heading id="D0FC541ECA92FF61EFEBFD7C43E4FD1F" box="[457,737,657,685]" centered="true" fontSize="11" level="4" pageId="139" pageNumber="584" reason="4">
(
<typeStatus id="54B05DD0CA92FF61EFF2FD7C430FFD1F" box="[464,522,657,685]" pageId="139" pageNumber="584">Type</typeStatus>
<figureCitation id="1330FFF7CA92FF61EC33FD7C4385FD1E" box="[529,640,657,684]" captionStart="Photo 37" captionStartId="97.[674,741,1381,1405]" captionText="Photo 37. Anax gladiator, male resting near the Mansha River at Kapishya Hot Springs, Zambia. The blade-like spike at the abdomen tip is well visible. Photo: JK (13 - xii- 2014)" pageId="139" pageNumber="584">Photo 37</figureCitation>
,
<figureCitation id="1330FFF7CA92FF61ECA8FD7C43DFFD1F" box="[650,730,657,685]" captionStart="Figure 22" captionStartId="143.[781,850,1264,1289]" captionTargetBox="[159,743,720,1550]" captionTargetId="figure@143.[156,746,715,1554]" captionTargetPageId="143" captionText="Figure 22. Male appendages in dorsal (left) and lateral view (right) of the African Paragomphus species with thick diverging tips of the cerci and a black thorax with five well-defined yellowish stripes on each side." pageId="139" pageNumber="584">Fig. 22</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BB4E372CA92FF61EEBDFD2D401AFD68" box="[159,287,704,730]" pageId="139" pageNumber="584">
<heading id="D0FC541ECA92FF61EEBDFD2D401AFD68" bold="true" box="[159,287,704,730]" fontSize="11" level="3" pageId="139" pageNumber="584" reason="6">
<emphasis id="B97F3F60CA92FF61EEBDFD2D401AFD68" bold="true" box="[159,287,704,730]" pageId="139" pageNumber="584">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA92FF67EEBDFD0943E0FE60" lastPageId="141" lastPageNumber="586" pageId="139" pageNumber="584" type="materials_examined">
<paragraph id="8BB4E372CA92FF66EEBDFD0940B4FDAF" lastPageId="140" lastPageNumber="585" pageId="139" pageNumber="584">
<bibRefCitation id="D50BFF3C99BB34A45E8089FB6E987856" author="Dijkstra K. - D. B. &amp; Clausnitzer V." journalOrPublisher="Royal Museum for Central Africa, Tervuren" refId="ref66897" refString="Dijkstra K. - D. B. &amp; Clausnitzer V. 2014. The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe. Royal Museum for Central Africa, Tervuren" title="The dragonflies and damselflies of eastern Africa: handbook for all Odonata from Sudan to Zimbabwe" type="book" year="2014">Dijkstra &amp; Clausnitzer (2014)</bibRefCitation>
noted that the taxonomic problems in
<taxonomicName id="4C0B98F1CA92FF61EEBDFCE740E4FC96" authority="Cowley, 1934" authorityName="Cowley" authorityYear="1934" box="[159,481,778,804]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA92FF61EEBDFCE7403FFC96" box="[159,314,778,804]" italics="true" pageId="139" pageNumber="584">Paragomphus</emphasis>
Cowley, 1934
</taxonomicName>
reside largely among the species with truncat- ed diverging cerci, which they dubbed the
<taxonomicName id="4C0B98F1CA92FF61ECB2FCDD43F0FCF8" box="[656,757,816,842]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B97F3F60CA92FF61ECB2FCDD43F0FCF8" box="[656,757,816,842]" italics="true" pageId="139" pageNumber="584">cognatus</emphasis>
</taxonomicName>
-group, although not all related species may share this character (see
<taxonomicName id="4C0B98F1CA92FF61EC9CFCB84293FCDD" box="[702,918,852,879]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="clausnitzerorum" status="sp. nov.">
<emphasis id="B97F3F60CA92FF61EC9CFCB84293FCDD" box="[702,918,852,879]" italics="true" pageId="139" pageNumber="584">P. clausnitzerorum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA92FF61ED82FCB842FAFCDD" box="[928,1023,852,879]" pageId="139" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
).
<bibRefCitation id="92B3396EB410B54820A8772BA0E553E7" author="Cammaerts R." journalOrPublisher="Bulletin et Annales de la Societe royale entomologique de Belgique" pagination="43 - 59" part="104" refId="ref66208" refString="Cammaerts R. 1968. Paragomphus cogna­ tus (Rambur, 1842) et quelques espèces voisines, en Afrique Centrale. Bulletin et Annales de la Société royale entomologique de Belgique 104: 43 - 59" title="Paragomphus cogna- tus (Rambur, 1842) et quelques especes voisines, en Afrique Centrale" type="journal article" year="1968">Cammaerts (1968)</bibRefCitation>
clarified the identity of
<taxonomicName id="4C0B98F1CA92FF61ECBFFC9642DCFC26" authority="Rambur, 1842" authorityName="Rambur" authorityYear="1842" box="[669,985,890,916]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B97F3F60CA92FF61ECBFFC96421EFC26" box="[669,795,891,916]" italics="true" pageId="139" pageNumber="584">P. cognatus</emphasis>
(Rambur, 1842)
</taxonomicName>
and compared the
<typeStatus id="54B05DD0CA92FF61EF64FC7240BDFC08" box="[326,440,927,954]" pageId="139" pageNumber="584">holotypes</typeStatus>
of
<taxonomicName id="4C0B98F1CA92FF61EFFEFC4D422CFC08" authority="Schouteden, 1934" authorityName="Schouteden" authorityYear="1934" box="[476,809,927,954]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="bredoi">
<emphasis id="B97F3F60CA92FF61EFFEFC4D433BFC08" box="[476,574,927,954]" italics="true" pageId="139" pageNumber="584">P. bredoi</emphasis>
(Schouteden, 1934)
</taxonomicName>
,
<taxonomicName id="4C0B98F1CA92FF61ED17FC4D4003FC6D" authority="Fraser, 1955" authorityName="Fraser" authorityYear="1955" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="longiventris">
<emphasis id="B97F3F60CA92FF61ED17FC4D42D1FC08" box="[821,980,927,954]" italics="true" pageId="139" pageNumber="584">P. longiventris</emphasis>
Fraser, 1955
</taxonomicName>
and
<emphasis id="B97F3F60CA92FF61EF60FC2840B1FC6D" box="[322,436,964,991]" italics="true" pageId="139" pageNumber="584">P. x a n t h u s</emphasis>
Pinhey, 1966 that all originate from north-eastern Congo-Kinshasa and are kept in
<collectionCode id="ED1A7BB7CA92FF61EC00FC07437CFBB6" box="[546,633,1002,1028]" collectionName="Belgium, Tervuren, Musee Royal de l'Afrique Centrale" pageId="139" pageNumber="584">MRAC</collectionCode>
. He synonymised the first two but was hesitant to include the latter, although it matches well morphologically and its paler appearance is explained by its more teneral state. Similar males have been found in Gabon, Nigeria, Benin, Togo, Ghana, Liberia and Sierra Leone: the species thus ranges widely across central and western Africa, warranting re-examination of the
<taxonomicName id="4C0B98F1CA92FF61EC2EFB494509FB0D" authority="Baumann, 1898" authorityName="Baumann" authorityYear="1898" box="[524,1036,1188,1215]" class="Insecta" family="Gomphidae" genus="Onychogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="serrulatus">
<emphasis id="B97F3F60CA92FF61EC2EFB494242FB0D" box="[524,839,1188,1215]" italics="true" pageId="139" pageNumber="584">Onychogomphus serrulatus</emphasis>
Baumann, 1898
</taxonomicName>
<typeStatus id="54B05DD0CA92FF61EEBDFB274003FB56" box="[159,262,1226,1252]" pageId="139" pageNumber="584">holotype</typeStatus>
from Togo in
<collectionCode id="ED1A7BB7CA92FF61EF89FB264305FB56" box="[427,512,1227,1252]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="139" pageNumber="584">ZMHB</collectionCode>
, which was recognised as pertaining to
<taxonomicName id="4C0B98F1CA92FF61EDEEFB264000FAB8" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA92FF61EDEEFB264000FAB8" italics="true" pageId="139" pageNumber="584">Paragomphus</emphasis>
</taxonomicName>
by Erich Schmidt (in lit. 1943). This female has an occiput adorned with two series of six large denticles (hence the specific name) and markings like those of females associated with the
<taxonomicName id="4C0B98F1CA92FF61EC58FAD643D9FAE6" box="[634,732,1338,1364]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="bredoi">
<emphasis id="B97F3F60CA92FF61EC58FAD643D9FAE6" box="[634,732,1338,1364]" italics="true" pageId="139" pageNumber="584">P. bredoi</emphasis>
</taxonomicName>
and
<emphasis id="B97F3F60CA92FF61ED34FAD6428DFAE6" box="[790,904,1338,1364]" italics="true" pageId="139" pageNumber="584">P. x a n t h u s</emphasis>
<typeStatus id="54B05DD0CA92FF61EDB1FAD74502FAE6" box="[915,1031,1338,1364]" pageId="139" pageNumber="584">holotypes</typeStatus>
, and we place all under the name
<taxonomicName id="4C0B98F1CA92FF61EC34FA8D43A4FAC8" box="[534,673,1375,1402]" class="Insecta" family="Gomphidae" genus="Onychogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="serrulatus">
<emphasis id="B97F3F60CA92FF61EC34FA8D43A4FAC8" box="[534,673,1375,1402]" italics="true" pageId="139" pageNumber="584">P. serrulatus</emphasis>
</taxonomicName>
[new synonymies]. The species differs from the largely allopatric
<taxonomicName id="4C0B98F1CA92FF61EC01FA6843A4FA2D" box="[547,673,1413,1439]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="139" pageNumber="584" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B97F3F60CA92FF61EC01FA6843A4FA2D" box="[547,673,1413,1439]" italics="true" pageId="139" pageNumber="584">P. cognatus</emphasis>
</taxonomicName>
of eastern and southern Africa (but see Tree 6) by the (1) complete pale metepisternal stripe interrupted only in very dark specimens, rather than typically interrupted at least above metastigma and often broken up into three spots; (2) apical segment of the penis bearing a small but prominent dorsal hook on each side; (3) narrower tips of the cerci, with one prominent and often spine-like posterior tooth and several often indistinct anterior ones, rather than several coarse similarly-sized teeth (
<figureCitation id="1330FFF7CA95FF66EF4BFE8040B8FE3A" box="[361,445,365,392]" captionStart="Figure 22" captionStartId="143.[781,850,1264,1289]" captionTargetBox="[159,743,720,1550]" captionTargetId="figure@143.[156,746,715,1554]" captionTargetPageId="143" captionText="Figure 22. Male appendages in dorsal (left) and lateral view (right) of the African Paragomphus species with thick diverging tips of the cerci and a black thorax with five well-defined yellowish stripes on each side." pageId="140" pageNumber="585">Fig. 22</figureCitation>
); and (4) clearly longer than wide epiproct in ventral view, rather than almost as wide as long. By this definition, a similarly pale species that is almost topotypical with the
<taxonomicName id="4C0B98F1CA95FF66ECF5FE544260FE60" box="[727,869,440,466]" class="Insecta" family="Gomphidae" genus="Onychogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="140" pageNumber="585" phylum="Arthropoda" rank="species" species="serrulatus">
<emphasis id="B97F3F60CA95FF66ECF5FE544260FE60" box="[727,869,440,466]" italics="true" pageId="140" pageNumber="585">P. serrulatus</emphasis>
</taxonomicName>
<typeStatus id="54B05DD0CA95FF66ED4CFE5542D2FE60" box="[878,983,440,466]" pageId="140" pageNumber="585">holotype</typeStatus>
, but has distinct markings and male appendages, although no genetic data, thus represents a new taxon.
</paragraph>
<paragraph id="8BB4E372CA95FF66EEBDFDA3406DFDDA" box="[159,360,590,616]" pageId="140" pageNumber="585">
<heading id="D0FC541ECA95FF66EEBDFDA3406DFDDA" bold="true" box="[159,360,590,616]" fontSize="11" level="3" pageId="140" pageNumber="585" reason="6">
<emphasis id="B97F3F60CA95FF66EEBDFDA3406DFDDA" bold="true" box="[159,360,590,616]" pageId="140" pageNumber="585">Material studied</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA95FF66EEBDFD99403DFD6A" pageId="140" pageNumber="585">
<materialsCitation id="3B63E92FCA95FF66EEBDFD994031FD6A" ID-GBIF-Occurrence="1224547947" collectingDate="2000-04-26" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Ghana" elevation="340 m" latitude="7.1166" location="Agumatsa River between Wli waterfalls and village" longLatPrecision="7" longitude="0.5833" pageId="140" pageNumber="585" specimenCount="1" specimenCount-male="1" stateProvince="Volta Region" typeStatus="holotype">
<typeStatus id="54B05DD0CA95FF66EEBDFD994015FD3F" box="[159,272,628,653]" pageId="140" pageNumber="585">
<emphasis id="B97F3F60CA95FF66EEBDFD994015FD3F" bold="true" box="[159,272,628,653]" pageId="140" pageNumber="585">Holotype</emphasis>
</typeStatus>
<emphasis id="B97F3F60CA95FF66EF3AFD9F4029FD3E" bold="true" box="[280,300,626,652]" pageId="140" pageNumber="585">
<specimenCount id="934EB2C1FBF2B545502536F1DEDCAFB7" type="male">♂</specimenCount>
</emphasis>
<emphasis id="B97F3F60CA95FF66EF0EFD994036FD3F" bold="true" box="[300,307,628,653]" pageId="140" pageNumber="585">.</emphasis>
<collectingCountry id="F31CA3E2CA95FF66EF18FD9F408FFD3F" box="[314,394,626,653]" name="Ghana" pageId="140" pageNumber="585" value="Ghana">Ghana</collectingCountry>
,
<collectingRegion id="49CF2D90CA95FF66EFB4FD9F4334FD3F" box="[406,561,626,653]" pageId="140" pageNumber="585">Volta Region</collectingRegion>
,
<location id="AA2C13A9416904766299A9CD71980638" LSID="urn:lsid:plazi:treatment:03A25264CA92FF65EF22FDA94274FD00:AA2C13A9416904766299A9CD71980638" country="Ghana" latitude="7.1166" longLatPrecision="7" longitude="0.5833" name="Agumatsa River between Wli waterfalls and village" stateProvince="Volta Region">Agumatsa River between Wli waterfalls and village</location>
,
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<elevation id="EFF64F12CF1D977B7FE60A8924B67FB4" box="[299,372,664,690]" metricMagnitude="2" metricUnit="m" metricValue="3.4" pageId="140" pageNumber="585" unit="m" value="340.0">340 m</elevation>
</quantity>
a.s.l. (
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<geoCoordinate id="EE3F85B5CA95FF66EC16FD7543A5FD00" box="[564,672,664,690]" direction="east" orientation="longitude" pageId="140" pageNumber="585" precision="5" value="0.5833">0.5833 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA95FF66EC96FD754231FD00" box="[692,820,664,690]" pageId="140" pageNumber="585" value="2000-04-26">
<collectingDate id="F6B599124910FC543796F146E5C5918F" box="[692,820,664,690]" pageId="140" pageNumber="585" value="2000-04-26">26 -iv- 2000</collectingDate>
</date>
, leg.
<collectorName id="C1AB415AD81FD64857224D0F9CA0559E">K.-D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA95FF66EEFAFD534031FD6A" box="[216,308,702,728]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="140" pageNumber="585">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<caption id="DF74B3FACA95FF66EEBDFA3540D1F9BD" pageId="140" pageNumber="585" targetBox="[158,1035,766,1456]" targetPageId="140">
<paragraph id="8BB4E372CA95FF66EEBDFA3540D1F9BD" pageId="140" pageNumber="585">
<emphasis id="B97F3F60CA95FF66EEBDFA354043FA43" bold="true" box="[159,326,1496,1521]" pageId="140" pageNumber="585">
Type
<figureCitation id="1330FFF7CA95FF66EEF9FA354047FA43" box="[219,322,1496,1521]" captionStart="Photo 37" captionStartId="97.[674,741,1381,1405]" captionText="Photo 37. Anax gladiator, male resting near the Mansha River at Kapishya Hot Springs, Zambia. The blade-like spike at the abdomen tip is well visible. Photo: JK (13 - xii- 2014)" pageId="140" pageNumber="585">Photo 37</figureCitation>
.
</emphasis>
Holotype of
<taxonomicName id="4C0B98F1CA95FF66EFF4FA3543F6FA42" box="[470,755,1496,1520]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="140" pageNumber="585" phylum="Arthropoda" rank="species" species="cammaertsi" status="sp. nov.">
<emphasis id="B97F3F60CA95FF66EFF4FA3543F6FA42" box="[470,755,1496,1520]" italics="true" pageId="140" pageNumber="585">Paragomphus cammaertsi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA95FF66ECD8FA354257FA43" box="[762,850,1496,1521]" pageId="140" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
, RMNH (horizontally mirrored from original).
</paragraph>
</caption>
<paragraph id="8BB4E372CA94FF67EEBDFF1343E0FE60" pageId="141" pageNumber="586">
<emphasis id="B97F3F60CA94FF67EEBDFF134071FEAA" bold="true" box="[159,372,254,280]" pageId="141" pageNumber="586">Further material.</emphasis>
<materialsCitation id="3B63E92FCA94FF67EF5EFF134061FE8F" ID-GBIF-Occurrence="1224547934" collectingDate="2000-04-26" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Ghana" elevation="340 m" latitude="7.1166" location="Volta Region" longLatPrecision="7" longitude="0.5833" pageId="141" pageNumber="586" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Volta Region">
<collectingCountry id="F31CA3E2CA94FF67EF5EFF1340E1FEAA" box="[380,484,254,280]" name="Ghana" pageId="141" pageNumber="586" value="Ghana">GHANA</collectingCountry>
(
<collectingRegion id="49CF2D90CA94FF67EFD6FF104391FEAA" box="[500,660,253,280]" pageId="141" pageNumber="586">Volta Region</collectingRegion>
):
<specimenCount id="9D0D28FBCA94FF67EC8BFF1343C8FEA5" box="[681,717,253,280]" pageId="141" pageNumber="586" type="generic">2 ♂</specimenCount>
<specimenCount id="F51924B09B8AA7E6DC0869AFF9DE9425" type="female">♀</specimenCount>
, as holotype, leg.
<collectorName id="C9E2B864C78D6241D03459FD00D88589">K.-D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA94FF67EF2AFECE4061FE8F" box="[264,356,291,317]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="141" pageNumber="586">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA94FF67EF4EFECE450DFED0" ID-GBIF-Occurrence="1224547960" collectingDate="1962-03-08" collectionCode="BMNH" collectorName="R. M. Gambles" country="Nigeria" elevation="995 m" latitude="9.5973" location="hill stream near Kagoro" longLatPrecision="7" longitude="8.4014" pageId="141" pageNumber="586" specimenCount="1" specimenCount-male="1" stateProvince="Kaduna Province">
<collectingCountry id="F31CA3E2CA94FF67EF4EFECE40E5FE8F" box="[364,480,291,317]" name="Nigeria" pageId="141" pageNumber="586" value="Nigeria">NIGERIA</collectingCountry>
(
<collectingRegion id="49CF2D90CA94FF67EFCFFECF43B9FE8F" box="[493,700,290,317]" pageId="141" pageNumber="586">Kaduna Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA94FF67ECEEFECE43EBFE8E" box="[716,750,290,317]" pageId="141" pageNumber="586" type="generic">1 ♂</specimenCount>
,
<location id="61F314B0DE106E524DB6CB9D5A3EA0AF" LSID="urn:lsid:plazi:treatment:03A25264CA92FF65EF22FDA94274FD00:61F314B0DE106E524DB6CB9D5A3EA0AF" country="Nigeria" latitude="9.5973" longLatPrecision="7" longitude="8.4014" name="hill stream near Kagoro" stateProvince="Kaduna Province">hill stream near Kagoro</location>
,
<quantity id="4CF34E97CA94FF67EEBDFEA541EFFED0" box="[159,234,328,354]" metricMagnitude="2" metricUnit="m" metricValue="9.95" pageId="141" pageNumber="586" unit="m" value="995.0">
<elevation id="5A91A316AE63259D26D6892DBC470B1A" box="[159,234,328,354]" metricMagnitude="2" metricUnit="m" metricValue="9.95" pageId="141" pageNumber="586" unit="m" value="995.0">995 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA94FF67EF1AFEA540A3FED0" box="[312,422,328,354]" direction="north" orientation="latitude" pageId="141" pageNumber="586" precision="5" value="9.5973">9.5973 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA94FF67EF92FEA5431EFED0" box="[432,539,328,354]" direction="east" orientation="longitude" pageId="141" pageNumber="586" precision="5" value="8.4014">8.4014 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA94FF67EC13FEA543B1FED0" box="[561,692,328,354]" pageId="141" pageNumber="586" value="1962-03-08">
<collectingDate id="B96A55AD091CEF03EF732FF6F56A7FB4" box="[561,692,328,354]" pageId="141" pageNumber="586" value="1962-03-08">08-iii- 1962</collectingDate>
</date>
, leg.
<collectorName id="91888C5E54FA74C264BBDDFF66BBC14D">R.M. Gambles</collectorName>
,
<collectionCode id="ED1A7BB7CA94FF67ED8FFEA4450DFED0" box="[941,1032,329,354]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="141" pageNumber="586">BMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA94FF67EEBDFE834033FE1F" ID-GBIF-Occurrence="1224547937" collectingDate="1962-04-23" collectionCode="BMNH" collectorName="R. M. Gambles" country="Nigeria" elevation="525 m" latitude="9.4596" location="Jemaa" longLatPrecision="7" longitude="8.3796" pageId="141" pageNumber="586" specimenCount="1" specimenCount-male="1" stateProvince="Kaduna Province">
<specimenCount id="9D0D28FBCA94FF67EEBDFE8341C4FE35" box="[159,193,365,392]" pageId="141" pageNumber="586" type="generic">1 ♂</specimenCount>
,
<location id="FD0D0D1F71E6A8B87036DDA0FAC9EDD5" LSID="urn:lsid:plazi:treatment:03A25264CA92FF65EF22FDA94274FD00:FD0D0D1F71E6A8B87036DDA0FAC9EDD5" country="Nigeria" latitude="9.4596" longLatPrecision="7" longitude="8.3796" name="Jemaa" stateProvince="Kaduna Province">Jemaa</location>
,
<quantity id="4CF34E97CA94FF67EF08FE834073FE3A" box="[298,374,366,392]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="141" pageNumber="586" unit="m" value="525.0">
<elevation id="48CAB41897048BCB43259385E622473E" box="[298,374,366,392]" metricMagnitude="2" metricUnit="m" metricValue="5.25" pageId="141" pageNumber="586" unit="m" value="525.0">525 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA94FF67EFE7FE804331FE3A" box="[453,564,365,392]" direction="north" orientation="latitude" pageId="141" pageNumber="586" precision="5" value="9.4596">9.4596 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA94FF67EC1DFE8043AEFE3A" box="[575,683,365,392]" direction="east" orientation="longitude" pageId="141" pageNumber="586" precision="5" value="8.3796">8.3796 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA94FF67ECE3FE804247FE3A" box="[705,834,365,392]" pageId="141" pageNumber="586" value="1962-04-23">
<collectingDate id="6EA9C0C218DAED8D47353425BE5E471D" box="[705,834,365,392]" pageId="141" pageNumber="586" value="1962-04-23">23 -iv- 1962</collectingDate>
</date>
, leg.
<collectorName id="3BE7E4F944879914E18505AEB4C411F6">R.M. Gambles</collectorName>
,
<collectionCode id="ED1A7BB7CA94FF67EEF9FE7E4033FE1F" box="[219,310,403,429]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="141" pageNumber="586">BMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA94FF67EF60FE7E43E4FE60" ID-GBIF-Occurrence="1224547946" collectingDate="1952-05-25" collectionCode="BMNH" collectorName="R. M. Gambles" country="Nigeria" elevation="924 m" latitude="9.6634" location="Jal" longLatPrecision="7" longitude="8.6328" pageId="141" pageNumber="586" specimenCount="1" specimenCount-male="1" stateProvince="Plateau Province">
<collectingCountry id="F31CA3E2CA94FF67EF60FE7E40B2FE1F" box="[322,439,403,429]" name="Nigeria" pageId="141" pageNumber="586" value="Nigeria">NIGERIA</collectingCountry>
(
<collectingRegion id="49CF2D90CA94FF67EFE5FE7F4391FE1F" box="[455,660,402,429]" pageId="141" pageNumber="586">Plateau Province</collectingRegion>
):
<specimenCount id="9D0D28FBCA94FF67EC8AFE7E43CFFE1E" box="[680,714,402,429]" pageId="141" pageNumber="586" type="generic">1 ♂</specimenCount>
,
<location id="C0277B00BD14A4014501336401829F45" LSID="urn:lsid:plazi:treatment:03A25264CA92FF65EF22FDA94274FD00:C0277B00BD14A4014501336401829F45" country="Nigeria" latitude="9.6634" longLatPrecision="7" longitude="8.6328" name="Jal" stateProvince="Plateau Province">Jal</location>
,
<quantity id="4CF34E97CA94FF67ED24FE7E4254FE1F" box="[774,849,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.24" pageId="141" pageNumber="586" unit="m" value="924.0">
<elevation id="E257877C07D5D4C4D1117771CC7A2B32" box="[774,849,403,429]" metricMagnitude="2" metricUnit="m" metricValue="9.24" pageId="141" pageNumber="586" unit="m" value="924.0">924 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA94FF67EDBFFE7F4509FE1F" box="[925,1036,402,429]" direction="north" orientation="latitude" pageId="141" pageNumber="586" precision="5" value="9.6634">9.6634 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA94FF67EEBDFE55400EFE60" box="[159,267,440,466]" direction="east" orientation="longitude" pageId="141" pageNumber="586" precision="5" value="8.6328">8.6328 ° E</geoCoordinate>
),
<date id="FFB5C5B2CA94FF67EF3CFE554092FE60" box="[286,407,440,466]" pageId="141" pageNumber="586" value="1952-05-25">
<collectingDate id="38676C5D34DC5507B99C203320A0C701" box="[286,407,440,466]" pageId="141" pageNumber="586" value="1952-05-25">25 -v- 1952</collectingDate>
</date>
, leg.
<collectorName id="7ED571C7D0872558F55F35F1B6ED1D2F">R.M. Gambles</collectorName>
,
<collectionCode id="ED1A7BB7CA94FF67ECA4FE5443E4FE60" box="[646,737,441,466]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="141" pageNumber="586">BMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA94FF67EEBDFDEE406AFDF0" box="[159,367,515,578]" pageId="141" pageNumber="586" type="description">
<paragraph id="8BB4E372CA94FF67EEBDFDEE400CFDAF" box="[159,265,515,541]" pageId="141" pageNumber="586">
<heading id="D0FC541ECA94FF67EEBDFDEE400CFDAF" bold="true" box="[159,265,515,541]" fontSize="11" level="3" pageId="141" pageNumber="586" reason="6">
<emphasis id="B97F3F60CA94FF67EEBDFDEE400CFDAF" bold="true" box="[159,265,515,541]" pageId="141" pageNumber="586">Genetics</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA94FF67EEBDFDC4406AFDF0" box="[159,367,552,578]" pageId="141" pageNumber="586">No data available.</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA94FF65EEBDFD99404FFE8F" lastPageId="143" lastPageNumber="588" pageId="141" pageNumber="586" type="diagnosis">
<paragraph id="8BB4E372CA94FF67EEBDFD994317FD3F" box="[159,530,627,653]" pageId="141" pageNumber="586">
<heading id="D0FC541ECA94FF67EEBDFD994317FD3F" bold="true" box="[159,530,627,653]" fontSize="11" level="3" pageId="141" pageNumber="586" reason="6">
<emphasis id="B97F3F60CA94FF67EEBDFD994317FD3F" bold="true" box="[159,530,627,653]" pageId="141" pageNumber="586">Male morphological diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA94FF65EEBDFD75404FFE8F" lastPageId="143" lastPageNumber="588" pageId="141" pageNumber="586">
Recalls
<taxonomicName id="4C0B98F1CA94FF67EED7FD744077FD00" box="[245,370,665,690]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B97F3F60CA94FF67EED7FD744077FD00" box="[245,370,665,690]" italics="true" pageId="141" pageNumber="586">P.cognatus</emphasis>
</taxonomicName>
and the potentially sympatric
<taxonomicName id="4C0B98F1CA94FF67ECEEFD744253FD00" box="[716,854,664,690]" class="Insecta" family="Gomphidae" genus="Onychogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="serrulatus">
<emphasis id="B97F3F60CA94FF67ECEEFD744253FD00" box="[716,854,664,690]" italics="true" pageId="141" pageNumber="586">P.serrulatus</emphasis>
</taxonomicName>
by (a) the moderate size, Hw
<quantity id="4CF34E97CA94FF67EF6BFD5340F5FD6A" box="[329,496,702,728]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="2.35" metricValueMin="2.25" pageId="141" pageNumber="586" unit="mm" value="23.0" valueMax="23.5" valueMin="22.5">22.5–23.5 mm</quantity>
(n= 3) and rather intermediate extent of dark markings, with the (b) pale occiput and narrowly black-edged and -based labrum contrasting with the largely dark vertex, antefrons and postclypeus; (c) ante-humeral and metepisternal stripes almost as prominent as the oth- er pale thoracic markings; and (d) S 3–6 laterally pale over much of their length; as well as (e) apices of the cerci that diverge in dorsal view (
<figureCitation id="1330FFF7CA94FF67ED88FC9542FAFC20" box="[938,1023,888,914]" captionStart="Figure 22" captionStartId="143.[781,850,1264,1289]" captionTargetBox="[159,743,720,1550]" captionTargetId="figure@143.[156,746,715,1554]" captionTargetPageId="143" captionText="Figure 22. Male appendages in dorsal (left) and lateral view (right) of the African Paragomphus species with thick diverging tips of the cerci and a black thorax with five well-defined yellowish stripes on each side." pageId="141" pageNumber="586">Fig. 22</figureCitation>
). However, (1) the pale ante-humeral and metepistermal markings consist of a stripe and a separate dorsal spot, while typically in
<taxonomicName id="4C0B98F1CA94FF67ED62FC2E42CAFC6F" box="[832,975,962,989]" class="Insecta" family="Gomphidae" genus="Onychogomphus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="serrulatus">
<emphasis id="B97F3F60CA94FF67ED62FC2E42CAFC6F" box="[832,975,962,989]" italics="true" pageId="141" pageNumber="586">P. serrulatus</emphasis>
</taxonomicName>
they form complete stripes and in
<taxonomicName id="4C0B98F1CA94FF67EFD3FC04436BFBB1" box="[497,622,1001,1027]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B97F3F60CA94FF67EFD3FC04436BFBB1" box="[497,622,1001,1027]" italics="true" pageId="141" pageNumber="586">P. cognatus</emphasis>
</taxonomicName>
they are broken up into three spots; (2)the black S 8–10 and appendages are marked laterally with some reddish brown, rather than being contrastingly marked with yellow at least on the appendages; (3) the appendages are short, the cerci being less than twice as long as S 10 and the epiproct about half as long S 10, rather than at least twice and just as long respectively, thus being intermediate between
<taxonomicName id="4C0B98F1CA94FF67ED8FFB4E4078FB50" authority="Kimmins, 1955" authorityName="Kimmins" authorityYear="1955" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="nyasicus">
<emphasis id="B97F3F60CA94FF67ED8FFB4E41C1FB50" italics="true" pageId="141" pageNumber="586">P. nyasicus</emphasis>
Kimmins, 1955
</taxonomicName>
(
<figureCitation id="1330FFF7CA94FF67EFAFFB2540DBFB50" box="[397,478,1224,1250]" captionStart="Figure 25" captionStartId="156.[159,227,1089,1114]" captionTargetBox="[448,1033,712,1550]" captionTargetId="figure@156.[445,1038,709,1553]" captionTargetPageId="156" captionText="Figure 25. Male appendages in dorsal (left) and lateral view (right) of the African Paragomphus species with parallel tips of the cerci, the epiproct about a third the length of the cerci, and a largely green thorax with limited brown markings; not shown is P. cataractae with very thin cerci and an (almost) unmarked thorax." pageId="141" pageNumber="586">Fig. 25</figureCitation>
) and
<taxonomicName id="4C0B98F1CA94FF67EC02FB2441D2FABA" authority="Clausnitzer &amp; Dijkstra, 2005" authorityName="Clausnitzer &amp; Dijkstra" authorityYear="2005" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="crenigomphoides">
<emphasis id="B97F3F60CA94FF67EC02FB2443FDFB51" box="[544,760,1224,1251]" italics="true" pageId="141" pageNumber="586">P. crenigomphoides</emphasis>
Clausnitzer &amp;
<bibRefCitation id="C69474019C3C62C431ED13A9A126B201" author="Dijkstra K. - D. B." journalOrPublisher="Journal of Afrotropical Zoology" pagination="3 - 14" part="2" refId="ref66743" refString="Dijkstra K. - D. B. 2005 a. A review of continental Afrotropical Ceriagrion (Odonata: Coenagrionidae). Journal of Afrotropical Zoology 2: 3 - 14" title="A review of continental Afrotropical Ceriagrion (Odonata: Coenagrionidae)" type="journal article" year="2005">Dijkstra, 2005</bibRefCitation>
</taxonomicName>
(
<figureCitation id="1330FFF7CA94FF67EECAFB004039FABA" box="[232,316,1261,1288]" captionStart="Figure 22" captionStartId="143.[781,850,1264,1289]" captionTargetBox="[159,743,720,1550]" captionTargetId="figure@143.[156,746,715,1554]" captionTargetPageId="143" captionText="Figure 22. Male appendages in dorsal (left) and lateral view (right) of the African Paragomphus species with thick diverging tips of the cerci and a black thorax with five well-defined yellowish stripes on each side." pageId="141" pageNumber="586">Fig. 22</figureCitation>
); and (4) the curved section of the cerci is notably short and sharply bent in lateral view, with thick and rounded tips with 6–7 rather in- distinct teeth, rather than square-ended with quite distinct and well-spaced teeth (
<figureCitation id="1330FFF7CA96FF65EECBFECF4038FE8F" box="[233,317,290,317]" captionStart="Figure 22" captionStartId="143.[781,850,1264,1289]" captionTargetBox="[159,743,720,1550]" captionTargetId="figure@143.[156,746,715,1554]" captionTargetPageId="143" captionText="Figure 22. Male appendages in dorsal (left) and lateral view (right) of the African Paragomphus species with thick diverging tips of the cerci and a black thorax with five well-defined yellowish stripes on each side." pageId="143" pageNumber="588">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF74B3FACA94FF67EEBDFA6F4094F9A2" pageId="141" pageNumber="586">
<paragraph id="8BB4E372CA94FF67EEBDFA6F4094F9A2" pageId="141" pageNumber="586">
<emphasis id="B97F3F60CA94FF67EEBDFA6F4084FA28" bold="true" box="[159,385,1410,1435]" pageId="141" pageNumber="586">Right page – Tree 6.</emphasis>
COI gene tree from Bayesian inference analysis for the extended
<taxonomicName id="4C0B98F1CA94FF67EED5FA72405FFA0A" box="[247,346,1439,1464]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="species" species="cognatus">
<emphasis id="B97F3F60CA94FF67EED5FA72405FFA0A" box="[247,346,1439,1464]" italics="true" pageId="141" pageNumber="586">cognatus</emphasis>
</taxonomicName>
-group of
<taxonomicName id="4C0B98F1CA94FF67EFEFFA724366FA0A" box="[461,611,1439,1464]" class="Insecta" family="Gomphidae" genus="Paragomphus" higherTaxonomySource="CoL" kingdom="Animalia" order="Odonata" pageId="141" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA94FF67EFEFFA724366FA0A" box="[461,611,1439,1464]" italics="true" pageId="141" pageNumber="586">Paragomphus</emphasis>
</taxonomicName>
and two outgroup taxa. For relevant branches posterior probabilities are shown first, as is bootstrap support from a 50% majority rule consensus of 1 0 0 0 trees from Maximum Likelihood analysis (both as percentages).
</paragraph>
</caption>
<subSubSection id="C311B0F9CA96FF65EEBDFE83433AFE4A" pageId="143" pageNumber="588" type="etymology">
<paragraph id="8BB4E372CA96FF65EEBDFE834021FE3A" box="[159,292,366,392]" pageId="143" pageNumber="588">
<heading id="D0FC541ECA96FF65EEBDFE834021FE3A" bold="true" box="[159,292,366,392]" fontSize="11" level="3" pageId="143" pageNumber="588" reason="6">
<emphasis id="B97F3F60CA96FF65EEBDFE834021FE3A" bold="true" box="[159,292,366,392]" pageId="143" pageNumber="588">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA96FF65EEBDFE7F433AFE4A" pageId="143" pageNumber="588">
Named in honour of Dr Roger Cammaerts who made great contributions to gomphid taxonomy and first recognised this species (in lit.) in
<collectionCode id="ED1A7BB7CA96FF65ED96FE544509FE60" box="[948,1036,441,466]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="143" pageNumber="588">BMNH</collectionCode>
(masculine singular genitive noun).
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA96FF65EEBDFDC44274FD00" pageId="143" pageNumber="588" type="distribution">
<paragraph id="8BB4E372CA96FF65EEBDFDC44081FDF1" box="[159,388,553,579]" pageId="143" pageNumber="588">
<heading id="D0FC541ECA96FF65EEBDFDC44081FDF1" bold="true" box="[159,388,553,579]" fontSize="11" level="3" pageId="143" pageNumber="588" reason="6">
<emphasis id="B97F3F60CA96FF65EEBDFDC44081FDF1" bold="true" box="[159,388,553,579]" pageId="143" pageNumber="588">Range and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA96FF65EEBDFDA34274FD00" pageId="143" pageNumber="588">
Known between 340 and
<quantity id="4CF34E97CA96FF65EFF6FDA34325FDDA" box="[468,544,590,616]" metricMagnitude="2" metricUnit="m" metricValue="9.95" pageId="143" pageNumber="588" unit="m" value="995.0">995 m</quantity>
a.s.l. from hilly regions in the transition from rainforest to humid savannah in Ghana and Nigeria; in Ghana found by a rocky stream at the base of a waterfall in a forested gorge.
</paragraph>
</subSubSection>
</treatment>
</document>
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