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<document id="2BA17B20BB2203EA9CA0068A6BE74374" ID-DOI="10.5281/zenodo.35388" ID-GBIF-Dataset="b184dfcc-9b5f-4abb-b401-0f45f76006f6" checkinTime="1450029366780" checkinUser="donat" docAuthor="Dijkstra, Klaas-Douwe B., Kipping, Jens &amp; Mézière, Nicolas" docDate="2015" docId="03A25264CA4CFFBDEF00FF10405AFE3A" docLanguage="en" docName="Odonatologica_44-4-low_res.pdf.imf" docOrigin="Odonatologica 44 (4)" docTitle="Pseudagrion pacale Dijkstra, sp. nov." docType="treatment" docVersion="20" lastPageNumber="532" masterDocId="FF9B2A1CCA19FFEAEE22FFED4105FFB2" masterDocTitle="Sixty new dragonfly and damselfly species from Africa (Odonata)" masterLastPageNumber="678" masterPageNumber="447" pageNumber="530" updateTime="1695001031402" updateUser="admin">
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<mods:titleInfo id="E207B6FED27268F218358AA04D3EC229">
<mods:title id="EBF74778BED9603B88125ECA28C817E7">Sixty new dragonfly and damselfly species from Africa (Odonata)</mods:title>
</mods:titleInfo>
<mods:name id="90D81D28B7F3B70F217F388AD2ABE2B2" type="personal">
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<mods:namePart id="24F4A601E1E6BB5B1BF623FAE2FF9C39">Dijkstra, Klaas-Douwe B.</mods:namePart>
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<mods:namePart id="F5E5ABA741819E1E8E893E0639567881">Kipping, Jens</mods:namePart>
</mods:name>
<mods:name id="78EC3D8A7EC6CB8A3CB93CCA977F4572" type="personal">
<mods:role id="394550E57E1BBB0A2E481D702F0C2686">
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<mods:namePart id="6CC9FC8258493C3550E2BFD85817FB1F">Mézière, Nicolas</mods:namePart>
</mods:name>
<mods:typeOfResource id="5F21692331AEFC408749DD5821EAD1A6">text</mods:typeOfResource>
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<mods:titleInfo id="D2313C1E7C39ACE81D55037D6FDFAF8C">
<mods:title id="8977C92CAF19214D8AA9FB5E9A500DED">Odonatologica</mods:title>
</mods:titleInfo>
<mods:part id="3885A4C9137F2ABEEC83A1EF4CA97EC8">
<mods:date id="4778C6B757E2610D08ACBCB1E5B9B4C2">2015</mods:date>
<mods:detail id="8FE245219694B5AD2793E99A751A265F" type="volume">
<mods:number id="377C40012EBAE5D090218C6904A72DAC">44</mods:number>
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<mods:classification id="AED5DB813DF74E3F3A03CD837E5B70F2">journal article</mods:classification>
<mods:identifier id="E45161E8BE17B25E7CD529BB2B39B50F" type="DOI">10.5281/zenodo.35388</mods:identifier>
</mods:mods>
<treatment id="03A25264CA4CFFBDEF00FF10405AFE3A" ID-DOI="http://doi.org/10.5281/zenodo.5640232" ID-GBIF-Taxon="114923428" ID-Zenodo-Dep="5640232" LSID="urn:lsid:zoobank.org:act:03A25264-CA4C-FFBD-EF00-FF10405AFE3A" httpUri="http://treatment.plazi.org/id/03A25264CA4CFFBDEF00FF10405AFE3A" lastPageId="87" lastPageNumber="532" pageId="85" pageNumber="530">
<subSubSection id="C311B0F9CA4CFFBFEF00FF10423CFEF1" pageId="85" pageNumber="530" type="nomenclature">
<paragraph id="8BB4E372CA4CFFBFEF00FF10423CFEF1" pageId="85" pageNumber="530">
<heading id="D0FC541ECA4CFFBFEF00FF10423CFEF1" bold="true" centered="true" fontSize="11" level="2" pageId="85" pageNumber="530" reason="4">
<emphasis id="B97F3F60CA4CFFBFEF00FF10428CFEAA" bold="true" box="[290,905,252,280]" pageId="85" pageNumber="530">
<taxonomicName id="4C0B98F1CA4CFFBFEF00FF104376FEAA" LSID-ZBK="urn:lsid:zoobank.org:act:03A25264-CA4C-FFBD-EF00-FF10405AFE3A" authority="Dijkstra" authorityName="Dijkstra" box="[290,627,253,280]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="pacale" status="sp. nov.">
<emphasis id="B97F3F60CA4CFFBFEF00FF104315FEAA" bold="true" box="[290,528,253,280]" italics="true" pageId="85" pageNumber="530">Pseudagrion pacale</emphasis>
Dijkstra
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA4CFFBFEC58FF1043D2FEAA" box="[634,727,253,280]" pageId="85" pageNumber="530" rank="species">sp. nov.</taxonomicNameLabel>
Peace Sprite
</emphasis>
(
<typeStatus id="54B05DD0CA4CFFBFEF55FECA40B7FEF1" box="[375,434,295,323]" pageId="85" pageNumber="530">Type</typeStatus>
<figureCitation id="1330FFF7CA4CFFBFEF9BFECA432DFEF0" box="[441,552,295,322]" captionStart="Photo 21" captionStartId="64.[159,225,1030,1055]" captionTargetBox="[158,1035,461,989]" captionTargetId="figure@64.[156,1038,458,991]" captionTargetPageId="64" captionText="Photo 21. Ceriagrion banditum, holotype male; Lake Chila, Mbala, Zambia. Photo: JK (08 - xii- 2014)" pageId="85" pageNumber="530">Photo 21</figureCitation>
,
<figureCitation id="1330FFF7CA4CFFBFEC10FECA43A8FEF0" box="[562,685,295,322]" captionStart="Photo 32" captionStartId="86.[159,225,1381,1405]" captionText="Photo 32. Rocky section of Moa River at Tilorma, Sierra Leone. Type locality of Pseudagrion pacale. Photo: KDBD (25 - ii- 2011)" pageId="85" pageNumber="530">Photos 32</figureCitation>
,
<figureCitation id="1330FFF7CA4CFFBFEC95FECA43DCFEF0" box="[695,729,295,322]" captionStart="Photo 75" captionStartId="218.[159,225,1322,1347]" captionText="Photo 75. Rocky stretch of the Moa River at Tiwai Island, Sierra Leone. Type locality of Zygonyx annika and habitat of Pseudagrion pacale. Photo: KDBD (14 - iii- 2011)" pageId="85" pageNumber="530">75</figureCitation>
,
<figureCitation id="1330FFF7CA4CFFBFECC1FECA4237FEF1" box="[739,818,295,323]" captionStart="Figure 12" captionStartId="78.[159,227,1266,1291]" captionTargetBox="[464,1034,602,1553]" captionTargetId="figure@78.[460,1038,598,1560]" captionTargetPageId="78" captionText="Figure 12. Penis in lateral and ventral view (left) and male appendages in dorsal (middle) and lateral view (right) of selection of Pseudagrion B-group species; P. tanganyicum sp. nov. is structurally identical to P. massaicum." pageId="85" pageNumber="530">Fig. 12</figureCitation>
)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBFEEBDFEBB401AFEC2" box="[159,287,342,368]" pageId="85" pageNumber="530" type="description">
<paragraph id="8BB4E372CA4CFFBFEEBDFEBB401AFEC2" box="[159,287,342,368]" pageId="85" pageNumber="530">
<heading id="D0FC541ECA4CFFBFEEBDFEBB401AFEC2" bold="true" box="[159,287,342,368]" fontSize="11" level="3" pageId="85" pageNumber="530" reason="6">
<emphasis id="B97F3F60CA4CFFBFEEBDFEBB401AFEC2" bold="true" box="[159,287,342,368]" pageId="85" pageNumber="530">Taxonomy</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBFEEBDFE9741D2FD98" pageId="85" pageNumber="530" type="discussion">
<paragraph id="8BB4E372CA4CFFBFEEBDFE9741D2FD98" pageId="85" pageNumber="530">
Belongs to a subgroup of
<taxonomicName id="4C0B98F1CA4CFFBFEFE3FE97434AFE27" box="[449,591,378,405]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA4CFFBFEFE3FE97434AFE27" box="[449,591,378,405]" italics="true" pageId="85" pageNumber="530">Pseudagrion</emphasis>
</taxonomicName>
B-group species that favour larger and swifter flowing waters. These include
<taxonomicName id="4C0B98F1CA4CFFBFEC76FE4C4266FE08" authority="Forster, 1906" authorityName="Forster" authorityYear="1906" box="[596,867,416,442]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="acaciae">
<emphasis id="B97F3F60CA4CFFBFEC76FE4C43C6FE08" box="[596,707,417,442]" italics="true" pageId="85" pageNumber="530">P. acaciae</emphasis>
Förster, 1906
</taxonomicName>
,
<taxonomicName id="4C0B98F1CA4CFFBFED4DFE4C404EFE52" authority="Karsch, 1893" authorityName="Karsch" authorityYear="1893" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="sublacteum">
<emphasis id="B97F3F60CA4CFFBFED4DFE4C450EFE08" box="[879,1035,416,442]" italics="true" pageId="85" pageNumber="530">P. sublacteum</emphasis>
(Karsch, 1893)
</taxonomicName>
,
<taxonomicName id="4C0B98F1CA4CFFBFEF77FE2B4377FE52" authority="Chutter, 1962" authorityName="Chutter" authorityYear="1962" box="[341,626,453,480]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="vaalense">
<emphasis id="B97F3F60CA4CFFBFEF77FE2B40CAFE52" box="[341,463,453,480]" italics="true" pageId="85" pageNumber="530">P. vaalense</emphasis>
Chutter, 1962
</taxonomicName>
and probably
<taxonomicName id="4C0B98F1CA4CFFBFED37FE2841DFFDB7" authority="Dumont, 1978" authorityName="Dumont" authorityYear="1978" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="niloticum">
<emphasis id="B97F3F60CA4CFFBFED37FE28429EFE52" box="[789,923,453,480]" italics="true" pageId="85" pageNumber="530">P.niloticum</emphasis>
Dumont, 1978
</taxonomicName>
. Present species is morphologically and genetically clearly differentiated.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBFEEBDFDB5406DFDC0" box="[159,360,600,626]" pageId="85" pageNumber="530" type="description">
<paragraph id="8BB4E372CA4CFFBFEEBDFDB5406DFDC0" box="[159,360,600,626]" pageId="85" pageNumber="530">
<heading id="D0FC541ECA4CFFBFEEBDFDB5406DFDC0" bold="true" box="[159,360,600,626]" fontSize="11" level="3" pageId="85" pageNumber="530" reason="6">
<emphasis id="B97F3F60CA4CFFBFEEBDFDB5406DFDC0" bold="true" box="[159,360,600,626]" pageId="85" pageNumber="530">Material studied</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBFEEBDFD93422CFCCA" pageId="85" pageNumber="530" type="materials_examined">
<paragraph id="8BB4E372CA4CFFBFEEBDFD9340A1FCBA" pageId="85" pageNumber="530">
<materialsCitation id="3B63E92FCA4CFFBFEEBDFD9340A5FCBA" ID-GBIF-Occurrence="1224547800" collectingDate="2011-02-26" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Sierra Leone" elevation="147 m" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF104-15" latitude="7.8123" location="S of Kenema, Moa River at Tilorma" longLatPrecision="7" longitude="-11.1779" pageId="85" pageNumber="530" specimenCode="RMNH.INS.554425" specimenCount="1" specimenCount-male="1" stateProvince="Eastern Province" typeStatus="holotype">
<typeStatus id="54B05DD0CA4CFFBFEEBDFD934015FD2A" box="[159,272,638,664]" pageId="85" pageNumber="530">
<emphasis id="B97F3F60CA4CFFBFEEBDFD934015FD2A" bold="true" box="[159,272,638,664]" pageId="85" pageNumber="530">Holotype</emphasis>
</typeStatus>
<emphasis id="B97F3F60CA4CFFBFEF38FD90402BFD25" bold="true" box="[282,302,637,663]" pageId="85" pageNumber="530">
<specimenCount id="24D81F8BD5006C75FB3A8C446D76E35F" type="male"></specimenCount>
</emphasis>
<emphasis id="B97F3F60CA4CFFBFEF0CFD934030FD2A" bold="true" box="[302,309,638,664]" pageId="85" pageNumber="530">.</emphasis>
<specimenCode id="B409D09C234E998033D56B33C528CE9E">RMNH.INS.501668</specimenCode>
,
<collectingCountry id="F31CA3E2CA4CFFBFEC15FD9043CAFD2A" box="[567,719,637,664]" name="Sierra Leone" pageId="85" pageNumber="530" value="Sierra Leone">Sierra Leone</collectingCountry>
,
<collectingRegion id="49CF2D90CA4CFFBFECFEFD9342ADFD2A" box="[732,936,637,664]" pageId="85" pageNumber="530">Eastern Province</collectingRegion>
,
<quantity id="4CF34E97CA4CFFBFED94FD9342F1FD2A" box="[950,1012,637,664]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="85" pageNumber="530" unit="km" value="7.0">
<locationDeviation id="F61BDF4F85BEACB97A0567E6A276CB33" box="[950,1012,637,664]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="85" pageNumber="530" unit="km" value="7.0">7 km</locationDeviation>
</quantity>
<location id="9761AA5905A6E53F76155BD2AA8BBB15" LSID="urn:lsid:plazi:treatment:03A25264CA4CFFBDEF00FF10405AFE3A:9761AA5905A6E53F76155BD2AA8BBB15" country="Sierra Leone" latitude="7.8123" longLatPrecision="7" longitude="-11.1779" name="S of Kenema, Moa River at Tilorma" stateProvince="Eastern Province">S of Kenema, Moa River at Tilorma</location>
, large river with gallery forest and rocky rapids (
<figureCitation id="1330FFF7CA4CFFBFEEDBFD25406CFD50" box="[249,361,712,738]" captionStart="Photo 32" captionStartId="86.[159,225,1381,1405]" captionText="Photo 32. Rocky section of Moa River at Tilorma, Sierra Leone. Type locality of Pseudagrion pacale. Photo: KDBD (25 - ii- 2011)" pageId="85" pageNumber="530">Photo 32</figureCitation>
),
<quantity id="4CF34E97CA4CFFBFEFA2FD2540CEFD50" box="[384,459,712,738]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="85" pageNumber="530" unit="m" value="147.0">
<elevation id="13BB7F6A8E1CBCC75C33BE8265476CAA" box="[384,459,712,738]" metricMagnitude="2" metricUnit="m" metricValue="1.47" pageId="85" pageNumber="530" unit="m" value="147.0">147 m</elevation>
</quantity>
a.s.l. (
<geoCoordinate id="EE3F85B5CA4CFFBFEC3BFD254382FD50" box="[537,647,712,738]" direction="north" orientation="latitude" pageId="85" pageNumber="530" precision="5" value="7.8123">7.8123 ° N</geoCoordinate>
<geoCoordinate id="EE3F85B5CA4CFFBFECB3FD254213FD50" box="[657,790,712,738]" direction="west" orientation="longitude" pageId="85" pageNumber="530" precision="5" value="-11.1779">11.1779 ° W</geoCoordinate>
),
<date id="FFB5C5B2CA4CFFBFED0EFD2542ADFD50" box="[812,936,712,738]" pageId="85" pageNumber="530" value="2011-02-26">
<collectingDate id="DB536000263F5EB8F6FB8E49AFE7595B" box="[812,936,712,738]" pageId="85" pageNumber="530" value="2011-02-26">26 -ii- 2011</collectingDate>
</date>
, leg.
<collectorName id="723CEF387F86637FB35D0C6972477AE6">K.- D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA4CFFBFEF66FD0340A5FCBA" box="[324,416,750,776]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="85" pageNumber="530">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BB4E372CA4CFFBFEEBDFCFE422CFCCA" pageId="85" pageNumber="530">
<emphasis id="B97F3F60CA4CFFBFEEBDFCFE4085FC9F" bold="true" box="[159,384,787,813]" pageId="85" pageNumber="530">Further material.</emphasis>
<materialsCitation id="3B63E92FCA4CFFBFEFB7FCFE439AFCE0" ID-GBIF-Occurrence="1224547780" collectingDate="2011-02-26" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Sierra Leone" elevation="147 m" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF102-15" httpUri-1="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF106-15" httpUri-2="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF105-15" latitude="7.8123" location="S of Kenema, Moa River at Tilorma" longLatPrecision="7" longitude="-11.1779" pageId="85" pageNumber="530" specimenCode="RMNH.INS.501661, RMNH.INS.501670, RMNH.INS.501669" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Eastern Province">
<specimenCount id="9D0D28FBCA4CFFBFEFB7FCFE40B2FC9E" box="[405,439,786,813]" pageId="85" pageNumber="530" type="generic">2 ♂</specimenCount>
(
<specimenCode id="E7DCAD8E05324514AACF11B063B44C57">RMNH.INS.501661</specimenCode>
,
<specimenCount id="432BE790D9C541B4E8DE0086EC554BC4">RMNH.INS.501670</specimenCount>
),
<specimenCount id="C0CCFC8D6F95A1D169099971702891B0" type="female">1 ♀</specimenCount>
(
<specimenCode id="FFF2E22DA975963389F4370B908E532A">RMNH.INS.501669</specimenCode>
), as holotype,
<collectionCode id="ED1A7BB7CA4CFFBFEC61FCD5439AFCE0" box="[579,671,824,850]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="85" pageNumber="530">RMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B63E92FCA4CFFBFEC89FCD54220FCCA" ID-GBIF-Occurrence="1224547796" collectingDate="2011-03-15" collectionCode="RMNH" collectorName="K. - D. B. Dijkstra" country="Sierra Leone" elevation="147 m" httpUri-0="http://www.boldsystems.org/index.php/Public_RecordView?processid=NSPAF107-15" latitude="7.8123" location="S of Kenema, Moa River at Tilorma" longLatPrecision="7" longitude="-11.1779" pageId="85" pageNumber="530" specimenCode="RMNH.INS.554425" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="Eastern Province">
<specimenCount id="9D0D28FBCA4CFFBFEC89FCD543C8FCE3" box="[683,717,824,850]" pageId="85" pageNumber="530" type="generic">3 ♂</specimenCount>
(
<specimenCode id="7B210A05B06C234FF2EF36713174639B">RMNH.INS.501674</specimenCode>
),
<specimenCount id="E9B80E1AEB85CED128D3911E4F060B4C" type="female">1 ♀</specimenCount>
, same locality,
<date id="FFB5C5B2CA4CFFBFEF60FCB040C0FCCA" box="[322,453,861,888]" pageId="85" pageNumber="530" value="2011-03-15">
<collectingDate id="6760C95DC6AD04594F2CED3FF0FCC7A0" box="[322,453,861,888]" pageId="85" pageNumber="530" value="2011-03-15">15 -iii- 2011</collectingDate>
</date>
, leg.
<collectorName id="7E39C11735AC6671D2C3A97DC87EC9FB">K.-D.B. Dijkstra</collectorName>
,
<collectionCode id="ED1A7BB7CA4CFFBFECEBFCB34220FCCA" box="[713,805,862,888]" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum (&quot;Naturalis&quot;) [formerly Rijksmuseum van Natuurlijke Historie]" pageId="85" pageNumber="530">RMNH</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBFEEBDFC4B400CFC72" box="[159,265,934,960]" pageId="85" pageNumber="530" type="description">
<paragraph id="8BB4E372CA4CFFBFEEBDFC4B400CFC72" box="[159,265,934,960]" pageId="85" pageNumber="530">
<heading id="D0FC541ECA4CFFBFEEBDFC4B400CFC72" bold="true" box="[159,265,934,960]" fontSize="11" level="3" pageId="85" pageNumber="530" reason="6">
<emphasis id="B97F3F60CA4CFFBFEEBDFC4B400CFC72" bold="true" box="[159,265,934,960]" pageId="85" pageNumber="530">Genetics</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBFEEBDFC2642ABFBB8" pageId="85" pageNumber="530" type="discussion">
<paragraph id="8BB4E372CA4CFFBFEEBDFC2642ABFBB8" pageId="85" pageNumber="530">
Two unique haplotypes (n = 5) distant from but nearest to
<taxonomicName id="4C0B98F1CA4CFFBFED47FC264500FC57" box="[869,1029,970,997]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="sublacteum">
<emphasis id="B97F3F60CA4CFFBFED47FC264500FC57" box="[869,1029,970,997]" italics="true" pageId="85" pageNumber="530">P. sublacteum</emphasis>
</taxonomicName>
; more distant to
<taxonomicName id="4C0B98F1CA4CFFBFEF7AFC1C40C3FBB8" box="[344,454,1009,1034]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="acaciae">
<emphasis id="B97F3F60CA4CFFBFEF7AFC1C40C3FBB8" box="[344,454,1009,1034]" italics="true" pageId="85" pageNumber="530">P. acaciae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0B98F1CA4CFFBFEC22FC1C437EFBB8" box="[512,635,1008,1034]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="vaalense">
<emphasis id="B97F3F60CA4CFFBFEC22FC1C437EFBB8" box="[512,635,1008,1034]" italics="true" pageId="85" pageNumber="530">P. vaalense</emphasis>
</taxonomicName>
;
<taxonomicName id="4C0B98F1CA4CFFBFECAAFC1C4215FBB8" box="[648,784,1008,1034]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="niloticum">
<emphasis id="B97F3F60CA4CFFBFECAAFC1C4215FBB8" box="[648,784,1008,1034]" italics="true" pageId="85" pageNumber="530">P. niloticum</emphasis>
</taxonomicName>
not sampled.
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4CFFBCEEBDFBD44266FE8F" lastPageId="86" lastPageNumber="531" pageId="85" pageNumber="530" type="diagnosis">
<paragraph id="8BB4E372CA4CFFBFEEBDFBD44317FBE0" box="[159,530,1080,1106]" pageId="85" pageNumber="530">
<heading id="D0FC541ECA4CFFBFEEBDFBD44317FBE0" bold="true" box="[159,530,1080,1106]" fontSize="11" level="3" pageId="85" pageNumber="530" reason="6">
<emphasis id="B97F3F60CA4CFFBFEEBDFBD44317FBE0" bold="true" box="[159,530,1080,1106]" pageId="85" pageNumber="530">Male morphological diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA4CFFBCEEBDFBB34266FE8F" lastPageId="86" lastPageNumber="531" pageId="85" pageNumber="530">
Nearest to
<taxonomicName id="4C0B98F1CA4CFFBFEF00FBB340C5FBCA" box="[290,448,1117,1144]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="sublacteum">
<emphasis id="B97F3F60CA4CFFBFEF00FBB340C5FBCA" box="[290,448,1117,1144]" italics="true" pageId="85" pageNumber="530">P. sublacteum</emphasis>
</taxonomicName>
by (a) the essentially similar markings including reddish eyes, face and ante-humeral stripes; (b) the broad lateral and apical lobes of the penis; (c) the narrow and slender cerci that clearly surpass the paraprocts; and (d) the hatchet-shaped paraprocts, with a prominent dorsal knob in lateral view (
<figureCitation id="1330FFF7CA4CFFBFEFE2FB1F4311FABF" box="[448,532,1266,1293]" captionStart="Figure 12" captionStartId="78.[159,227,1266,1291]" captionTargetBox="[464,1034,602,1553]" captionTargetId="figure@78.[460,1038,598,1560]" captionTargetPageId="78" captionText="Figure 12. Penis in lateral and ventral view (left) and male appendages in dorsal (middle) and lateral view (right) of selection of Pseudagrion B-group species; P. tanganyicum sp. nov. is structurally identical to P. massaicum." pageId="85" pageNumber="530">Fig. 12</figureCitation>
). However, (1) is larger, Hw 21.5 24.0mm (n = 6) rather than 18.5 21.0 mm (n = 6); (2) the postocular spots and antehumeral stripes become deep orange and the thorax sides pale green with maturity, rather than all turning dark and eventually uniformly pruinose; (3) the postocular spots are clearly framed with black; (4) the apical cleft of the penis is deep and narrow, while its wide lateral lobes bear tiny projections, thus the penis most recalls that of
<taxonomicName id="4C0B98F1CA4CFFBFECA2FA3E42B5FA5F" authority="Longfield, 1947" authorityName="Longfield" authorityYear="1947" box="[640,944,1490,1517]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="85" pageNumber="530" phylum="Arthropoda" rank="species" species="coeleste">
<emphasis id="B97F3F60CA4CFFBFECA2FA3E43EAFA5F" box="[640,751,1490,1517]" italics="true" pageId="85" pageNumber="530">P. coeleste</emphasis>
Longfield, 1947
</taxonomicName>
; (5)the paraprocts are pointed rather than rounded in dorsal view; and (6) the dorsal knob of the paraprocts is much higher than deep and thus more pronounced than in other African
<taxonomicName id="4C0B98F1CA4FFFBCEC2EFECF439FFE8F" box="[524,666,290,317]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="86" pageNumber="531" phylum="Arthropoda" rank="genus">
<emphasis id="B97F3F60CA4FFFBCEC2EFECF439FFE8F" box="[524,666,290,317]" italics="true" pageId="86" pageNumber="531">Pseudagrion</emphasis>
</taxonomicName>
species (
<figureCitation id="1330FFF7CA4FFFBCED20FECF4250FE8F" box="[770,853,290,317]" captionStart="Figure 12" captionStartId="78.[159,227,1266,1291]" captionTargetBox="[464,1034,602,1553]" captionTargetId="figure@78.[460,1038,598,1560]" captionTargetPageId="78" captionText="Figure 12. Penis in lateral and ventral view (left) and male appendages in dorsal (middle) and lateral view (right) of selection of Pseudagrion B-group species; P. tanganyicum sp. nov. is structurally identical to P. massaicum." pageId="86" pageNumber="531">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C311B0F9CA4FFFBCEEBDFE834294FE4A" pageId="86" pageNumber="531" type="etymology">
<paragraph id="8BB4E372CA4FFFBCEEBDFE834021FE3A" box="[159,292,366,392]" pageId="86" pageNumber="531">
<heading id="D0FC541ECA4FFFBCEEBDFE834021FE3A" bold="true" box="[159,292,366,392]" fontSize="11" level="3" pageId="86" pageNumber="531" reason="6">
<emphasis id="B97F3F60CA4FFFBCEEBDFE834021FE3A" bold="true" box="[159,292,366,392]" pageId="86" pageNumber="531">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA4FFFBCEEBDFE7F4294FE4A" pageId="86" pageNumber="531">Latin “belonging to peace” to contrast with the 19912002 civil war in which the Moa River was a frontline and the Ebola epidemic since 2013 of which the Kenema region was the national epicentre (neuter adjective).</paragraph>
</subSubSection>
<caption id="DF74B3FACA4FFFBCEEBDFBA64253FBD1" box="[159,854,1099,1124]" pageId="86" pageNumber="531" targetBox="[158,1035,539,1058]" targetPageId="86">
<paragraph id="8BB4E372CA4FFFBCEEBDFBA64253FBD1" box="[159,854,1099,1124]" pageId="86" pageNumber="531">
<emphasis id="B97F3F60CA4FFFBCEEBDFBA64041FBD1" bold="true" box="[159,324,1099,1124]" pageId="86" pageNumber="531">
Type
<figureCitation id="1330FFF7CA4FFFBCEEF8FBA6403AFBD1" box="[218,319,1099,1123]" captionStart="Photo 21" captionStartId="64.[159,225,1030,1055]" captionTargetBox="[158,1035,461,989]" captionTargetId="figure@64.[156,1038,458,991]" captionTargetPageId="64" captionText="Photo 21. Ceriagrion banditum, holotype male; Lake Chila, Mbala, Zambia. Photo: JK (08 - xii- 2014)" pageId="86" pageNumber="531">Photo 21</figureCitation>
.
</emphasis>
Holotype of
<taxonomicName id="4C0B98F1CA4FFFBCEFEDFBA643A6FBD1" box="[463,675,1099,1123]" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="86" pageNumber="531" phylum="Arthropoda" rank="species" species="pacale" status="sp. nov.">
<emphasis id="B97F3F60CA4FFFBCEFEDFBA643A6FBD1" box="[463,675,1099,1123]" italics="true" pageId="86" pageNumber="531">Pseudagrion pacale</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24C821BCA4FFFBCEC8BFBA643FAFBD1" box="[681,767,1099,1123]" pageId="86" pageNumber="531" rank="species">sp. nov.</taxonomicNameLabel>
, RMNH.
</paragraph>
</caption>
<caption id="DF74B3FACA4FFFBCEEBDFA884065F9A2" pageId="86" pageNumber="531">
<paragraph id="8BB4E372CA4FFFBCEEBDFA884065F9A2" pageId="86" pageNumber="531">
<emphasis id="B97F3F60CA4FFFBCEEBDFA884015FACF" bold="true" box="[159,272,1381,1405]" pageId="86" pageNumber="531">Photo 32.</emphasis>
Rocky section of Moa River at Tilorma, Sierra Leone. Type locality of
<taxonomicName id="4C0B98F1CA4FFFBCEF69FA50403CFA40" class="Insecta" family="Coenagrionidae" genus="Pseudagrion" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Odonata" pageId="86" pageNumber="531" phylum="Arthropoda" rank="species" species="pacale">
<emphasis id="B97F3F60CA4FFFBCEF69FA50403CFA40" italics="true" pageId="86" pageNumber="531">Pseudagrion pacale</emphasis>
</taxonomicName>
. Photo: KDBD (25-ii-2011)
</paragraph>
</caption>
<subSubSection id="C311B0F9CA4EFFBDEEBDFF13405AFE3A" pageId="87" pageNumber="532" type="distribution">
<paragraph id="8BB4E372CA4EFFBDEEBDFF134081FEAA" box="[159,388,254,280]" pageId="87" pageNumber="532">
<heading id="D0FC541ECA4EFFBDEEBDFF134081FEAA" bold="true" box="[159,388,254,280]" fontSize="11" level="3" pageId="87" pageNumber="532" reason="6">
<emphasis id="B97F3F60CA4EFFBDEEBDFF134081FEAA" bold="true" box="[159,388,254,280]" pageId="87" pageNumber="532">Range and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB4E372CA4EFFBDEEBDFECF405AFE3A" pageId="87" pageNumber="532">
Rocky sections with rapid flow of the broad Moa River between 120 and
<quantity id="4CF34E97CA4EFFBDEEBDFEA541E2FED0" box="[159,231,328,354]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="87" pageNumber="532" unit="m" value="150.0">150m</quantity>
a.s.l. in Sierra Leone; also seen at Tiwai Island,
<quantity id="4CF34E97CA4EFFBDED2FFEA54253FED0" box="[781,854,328,354]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="87" pageNumber="532" unit="km" value="35.0">35 km</quantity>
downstream of the
<typeStatus id="54B05DD0CA4EFFBDEEE9FE8341F8FE3A" box="[203,253,366,392]" pageId="87" pageNumber="532">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>