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<document id="8450FE13D7F2462A7D8F09A910520462" ID-CLB-Dataset="22540" ID-DOI="10.11646/zootaxa.4657.2.7" ID-GBIF-Dataset="b2d3cc3b-6448-4235-b74f-b0524bb9f918" ID-ISSN="1175-5326" ID-Zenodo-Dep="3772337" ID-ZooBank="9CF65718-E007-4B9D-B9B4-A88401DB971E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588057346543" checkinUser="plazi" docAuthor="Shear, William A. &amp; Marek, Paul E." docDate="2019" docId="038ECD2FCB6B2C79FF7BFA4DFCAF365E" docLanguage="en" docName="zootaxa.4657.2.7.pdf" docOrigin="Zootaxa 4657 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Urochordeuma bumpusi Silvestri 1909" docType="treatment" docVersion="6" lastPageNumber="356" masterDocId="FFB7B557CB6A2C7EFFECFFB7FFD33741" masterDocTitle="Urochordeumatidae Silvestri, 1909, a millipede family endemic to Washington State, USA (Chordeumatida, Striariidea, Striarioidea)" masterLastPageNumber="360" masterPageNumber="352" pageNumber="353" updateTime="1698837621999" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="AE4F4C1D4A92EE3C41B41A06A1C7C0A4">Urochordeumatidae Silvestri, 1909, a millipede family endemic to Washington State, USA (Chordeumatida, Striariidea, Striarioidea)</mods:title>
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<mods:namePart id="DFB2D7DFE632C5B387F3286A1F6E4373">Shear, William A.</mods:namePart>
<mods:affiliation id="D5EF000EEA30FE8B8C8B3CD0C765B53D">Professor Emeritus, Biology Department, Hampden-Sydney College, Hampden-Sydney, VA 23943 USA. E-mail wshear @ hsc. edu Entomology Department, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061</mods:affiliation>
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<mods:namePart id="34ACA220F0A5F422AB45D11958423E83">Marek, Paul E.</mods:namePart>
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<treatment id="038ECD2FCB6B2C79FF7BFA4DFCAF365E" ID-DOI="http://doi.org/10.5281/zenodo.3798216" ID-GBIF-Taxon="163829526" ID-Zenodo-Dep="3798216" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E" httpUri="http://treatment.plazi.org/id/038ECD2FCB6B2C79FF7BFA4DFCAF365E" lastPageId="7" lastPageNumber="356" pageId="1" pageNumber="353">
<subSubSection id="C33D2FB2CB6B2C7FFF7BFA4DFDA43155" box="[151,631,1530,1557]" pageId="1" pageNumber="353" type="nomenclature">
<paragraph id="8B987C39CB6B2C7FFF7BFA4DFDA43155" blockId="1.[151,631,1530,1592]" box="[151,631,1530,1557]" pageId="1" pageNumber="353">
<heading id="D0D0CB55CB6B2C7FFF7BFA4DFDA43155" bold="true" box="[151,631,1530,1557]" fontSize="11" level="1" pageId="1" pageNumber="353" reason="1">
<taxonomicName id="4C2707BACB6B2C7FFF7BFA4DFDA43155" ID-CoL="7DTTJ" authority="Silvestri, 1909" authorityName="Silvestri" authorityYear="1909" box="[151,631,1530,1557]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="353" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB6B2C7FFF7BFA4DFDA43155" bold="true" box="[151,631,1530,1557]" pageId="1" pageNumber="353">
<emphasis id="B953A02BCB6B2C7FFF7BFA4DFE6C3154" bold="true" box="[151,447,1530,1557]" italics="true" pageId="1" pageNumber="353">Urochordeuma bumpusi</emphasis>
<bibRefCitation id="EFB601C8CB6B2C7FFE2AFA4DFDA43155" author="Silvestri, F." box="[454,631,1530,1556]" pageId="1" pageNumber="353" pagination="229 - 233" refId="ref3903" refString="Silvestri, F. (1909) Descrizioni preliminari di varii Artropodi, specialmente d' America. III. Nuovi Chordeumoidea (Diplopoda). Rendiconti, Accademia Nazionale dei Lincei, 5. Ser. / Classe di Scienze Fisiche, Matematiche e Naturali, 18, 229 - 233. Available from: https: // biodiversitylibrary. org / page / 44698853 (Accessed 16 Aug. 2019)" type="journal article" year="1909">Silvestri, 1909</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33D2FB2CB6B2C7FFF7BF9A9FED53179" box="[151,262,1566,1592]" pageId="1" pageNumber="353" type="description">
<paragraph id="8B987C39CB6B2C7FFF7BF9A9FED53179" blockId="1.[151,631,1530,1592]" box="[151,262,1566,1592]" pageId="1" pageNumber="353">
<figureCitation id="131C60BCCB6B2C7FFF7BF9A9FED53179" box="[151,262,1566,1592]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 24" captionStart-2="FIGURES 5, 6" captionStart-3="FIGURES 711" captionStart-4="FIGURES 12" captionStartId-0="2.[151,250,1881,1906]" captionStartId-1="3.[151,264,1809,1834]" captionStartId-2="5.[151,264,1974,1999]" captionStartId-3="6.[151,264,1900,1925]" captionStartId-4="7.[151,264,1480,1505]" captionTargetBox-0="[151,1436,704,1862]" captionTargetBox-1="[170,1419,181,1763]" captionTargetBox-2="[189,1377,181,1949]" captionTargetBox-3="[169,1419,181,1846]" captionTargetBox-4="[163,1425,309,1461]" captionTargetId-0="figure@2.[151,1436,704,1862]" captionTargetId-1="figure@3.[169,1419,181,1786]" captionTargetId-2="figure@5.[188,1402,181,1949]" captionTargetId-3="figure@6.[169,1419,181,1877]" captionTargetId-4="figure@7.[163,1425,309,1461]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionText-0="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." captionText-1="FIGURES 24. Urochordeuma bumpusi Silvestri, 1909. 2: Rings 1012 (10, 11, 12) of male. 3: Rings 28, 29 and telson (28, 29, t) of male. 4: Small cuticular nodules of metazonite, each with two acute, posteriorly pointing processes." captionText-2="FIGURES 5, 6. Urochordeuma bumpusi Silvestri, 1909. 5: First legpair, posterior view. 6: Third legpair, posterior view." captionText-3="FIGURES 711. Urochordeuma bumpusi. 7: Gonopods, anterior view. 8: Left gonopod, mesal view. 9: Tip of right gonopod, lateral view. 10: Ninth legpair, anterior view. 11: Left vulva, ventral view. Abbreviations: acp, anterior coxal process; cx, coxa; cx9, coxa, cp9, coxal process, pcp, posterior coxal process; st, sternum; t9, telopodite" captionText-4="FIGURES 12. Northwest Washington, showing records of Urochordeuma bumpusi. Circles, new records. Triangle, type locality of U. bumpusi. Square, type locality of U. porona." figureDoi-0="http://doi.org/10.5281/zenodo.3772339" figureDoi-1="http://doi.org/10.5281/zenodo.3772341" figureDoi-2="http://doi.org/10.5281/zenodo.3772343" figureDoi-3="http://doi.org/10.5281/zenodo.3772345" figureDoi-4="http://doi.org/10.5281/zenodo.3772347" httpUri-0="https://zenodo.org/record/3772339/files/figure.png" httpUri-1="https://zenodo.org/record/3772341/files/figure.png" httpUri-2="https://zenodo.org/record/3772343/files/figure.png" httpUri-3="https://zenodo.org/record/3772345/files/figure.png" httpUri-4="https://zenodo.org/record/3772347/files/figure.png" pageId="1" pageNumber="353">Figs 112</figureCitation>
</paragraph>
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<subSubSection id="C33D2FB2CB6B2C7FFF7BF9D5FAB331D8" pageId="1" pageNumber="353" type="reference_group">
<paragraph id="8B987C39CB6B2C7FFF7BF9D5FC21313A" blockId="1.[151,1376,1634,1689]" box="[151,1010,1634,1659]" pageId="1" pageNumber="353">
<taxonomicName id="4C2707BACB6B2C7FFF7BF9D5FDBC313A" ID-CoL="7DTTJ" authority="Silvestri 1909, p. 230" authorityName="Silvestri" authorityPageNumber="230" authorityYear="1909" box="[151,623,1634,1659]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="353" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB6B2C7FFF7BF9D5FE5F313B" box="[151,396,1634,1658]" italics="true" pageId="1" pageNumber="353">Urochordeuma bumpusi</emphasis>
<bibRefCitation id="EFB601C8CB6B2C7FFE7FF9D5FDF1313A" author="Silvestri, F." box="[403,546,1634,1659]" pageId="1" pageNumber="353" pagination="229 - 233" refId="ref3903" refString="Silvestri, F. (1909) Descrizioni preliminari di varii Artropodi, specialmente d' America. III. Nuovi Chordeumoidea (Diplopoda). Rendiconti, Accademia Nazionale dei Lincei, 5. Ser. / Classe di Scienze Fisiche, Matematiche e Naturali, 18, 229 - 233. Available from: https: // biodiversitylibrary. org / page / 44698853 (Accessed 16 Aug. 2019)" type="journal article" year="1909">Silvestri 1909</bibRefCitation>
, p. 230
</taxonomicName>
; 1913, p. 305.
<bibRefCitation id="EFB601C8CB6B2C7FFCE9F9D5FC72313A" author="Hoffman, R. L." box="[773,929,1634,1659]" pageId="1" pageNumber="353" refId="ref3671" refString="Hoffman, R. L. (1999) Checklist of Millipeds of North and Middle America. Virginia Museum of Natural History, Special Publication No. 8, 584 pp. Available from: https: // www. fieldmuseum. org / sites / default / files / hoffman _ checklist _ 1999. pdf (Accessed 16 Aug. 2019)" type="book" year="1999">Hoffman, 1999</bibRefCitation>
, p. 256.
</paragraph>
<paragraph id="8B987C39CB6B2C7FFF7BF936FAB331D8" blockId="1.[151,1376,1634,1689]" box="[151,1376,1664,1689]" pageId="1" pageNumber="353">
<taxonomicName id="4C2707BACB6B2C7FFF7BF936FDAB31D8" ID-CoL="7DTTH" authority="Chamberlin 1941, p. 23" authorityName="Chamberlin" authorityPageNumber="23" authorityYear="1941" box="[151,632,1664,1689]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="353" phylum="Arthropoda" rank="species" species="porona">
<emphasis id="B953A02BCB6B2C7FFF7BF936FEAD31D8" box="[151,382,1665,1689]" italics="true" pageId="1" pageNumber="353">Urochordeuma porona</emphasis>
<bibRefCitation id="EFB601C8CB6B2C7FFE69F937FDEB31D8" author="Chamberlin, R. V." box="[389,568,1664,1689]" pageId="1" pageNumber="353" pagination="1 - 389" refId="ref3557" refString="Chamberlin, R. V. (1941) New American millipeds. Bulletin of the University of Utah, 6 (4) / 31 (11), 1 - 389. Available from: https: // collections. lib. utah. edu / details? id = 708781 (Accessed 16 Aug. 2019)" type="journal article" year="1941">Chamberlin 1941</bibRefCitation>
, p. 23
</taxonomicName>
.
<bibRefCitation id="EFB601C8CB6B2C7FFD6EF937FD2F31D8" author="Shear, W. A." box="[642,764,1664,1689]" pageId="1" pageNumber="353" pagination="151 - 352" refId="ref3794" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda). A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. - Bulletin of the Museum of Comparative Zoology, 144, 151 - 352. Available from: https: // biodiversitylibrary. org / page / 4227660 (Accessed 16 Aug. 2019)" type="journal article" year="1972">Shear, 1972</bibRefCitation>
, p. 260.
<bibRefCitation id="EFB601C8CB6B2C7FFCBEF937FC3C31D8" author="Hoffman, R. L." box="[850,1007,1664,1689]" pageId="1" pageNumber="353" refId="ref3671" refString="Hoffman, R. L. (1999) Checklist of Millipeds of North and Middle America. Virginia Museum of Natural History, Special Publication No. 8, 584 pp. Available from: https: // www. fieldmuseum. org / sites / default / files / hoffman _ checklist _ 1999. pdf (Accessed 16 Aug. 2019)" type="book" year="1999">Hoffman, 1999</bibRefCitation>
, p. 256.
<emphasis id="B953A02BCB6B2C7FFBA9F936FAB331D8" bold="true" box="[1093,1376,1664,1689]" pageId="1" pageNumber="353">New subjective synonymy.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33D2FB2CB6B2C7FFF7BF976FCED3009" pageId="1" pageNumber="353" type="materials_examined">
<paragraph id="8B987C39CB6B2C7FFF7BF976FCED3009" blockId="1.[151,1437,1729,2045]" pageId="1" pageNumber="353">
<emphasis id="B953A02BCB6B2C7FFF7BF976FE51319D" bold="true" box="[151,386,1729,1756]" pageId="1" pageNumber="353">Material examined:</emphasis>
Of
<emphasis id="B953A02BCB6B2C7FFE41F974FDE7319D" box="[429,564,1730,1756]" italics="true" pageId="1" pageNumber="353">
<taxonomicName id="4C2707BACB6B2C7FFE41F974FDE2319D" authorityName="Silvestri" authorityYear="1909" box="[429,561,1730,1756]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="353" phylum="Arthropoda" rank="species" species="bumpusi">U. bumpusi</taxonomicName>
,
</emphasis>
male
<typeStatus id="549CC29BCB6B2C7FFD94F975FD09319D" box="[632,730,1730,1756]" pageId="1" pageNumber="353" type="holotype">holotype</typeStatus>
from Longmire Springs, Pierce County,
<collectingRegion id="49E3B2DBCB6B2C7FFB4CF975FAFB319D" box="[1184,1320,1730,1756]" country="United States of America" name="Washington" pageId="1" pageNumber="353">Washington</collectingRegion>
,
<collectingCountry id="F3303CA9CB6B2C7FFADFF975FABD319D" box="[1331,1390,1730,1756]" name="United States of America" pageId="1" pageNumber="353">USA</collectingCountry>
, deposited in the Museo Civico di Storia Naturale de Genova,
<collectingCountry id="F3303CA9CB6B2C7FFCCFF951FC843041" box="[803,855,1766,1792]" name="Italy" pageId="1" pageNumber="353">Italy</collectingCountry>
, examined by
<collectionCode id="ED36E4FCCB6B2C7FFC1AF951FBE23041" box="[1014,1073,1766,1792]" pageId="1" pageNumber="353">WAS</collectionCode>
January, 2019. Male
<typeStatus id="549CC29BCB6B2C7FFAF1F951FAAC3041" box="[1309,1407,1766,1792]" pageId="1" pageNumber="353" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C2707BACB6B2C7FFF7BF8BCFEDB3065" authorityName="Chamberlin" authorityYear="1941" box="[151,264,1803,1828]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="353" phylum="Arthropoda" rank="species" species="porona">
<emphasis id="B953A02BCB6B2C7FFF7BF8BCFEDB3065" box="[151,264,1803,1828]" italics="true" pageId="1" pageNumber="353">U. porona</emphasis>
</taxonomicName>
from Snoqualmie Pass, King County,
<collectingRegion id="49E3B2DBCB6B2C7FFD43F8BDFCE43065" box="[687,823,1802,1828]" country="United States of America" name="Washington" pageId="1" pageNumber="353">Washington</collectingRegion>
,
<collectingCountry id="F3303CA9CB6B2C7FFCACF8BDFCA83065" box="[832,891,1802,1828]" name="United States of America" pageId="1" pageNumber="353">USA</collectingCountry>
, deposited in the
<collectingCountry id="F3303CA9CB6B2C7FFBD6F8BDFB1D3065" box="[1082,1230,1802,1828]" name="United States of America" pageId="1" pageNumber="353">United States</collectingCountry>
National Museum of Natural History, examined by
<collectionCode id="ED36E4FCCB6B2C7FFDEBF899FD913009" box="[519,578,1838,1864]" pageId="1" pageNumber="353">WAS</collectionCode>
in 1970 (
<bibRefCitation id="EFB601C8CB6B2C7FFD40F899FCE23009" author="Shear, W. A." box="[684,817,1838,1864]" pageId="1" pageNumber="353" pagination="151 - 352" refId="ref3794" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda). A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. - Bulletin of the Museum of Comparative Zoology, 144, 151 - 352. Available from: https: // biodiversitylibrary. org / page / 4227660 (Accessed 16 Aug. 2019)" type="journal article" year="1972">Shear 1972</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D2FB2CB6B2C7FFF2BF8E6FB283099" pageId="1" pageNumber="353" type="diagnosis">
<paragraph id="8B987C39CB6B2C7FFF2BF8E6FB283099" blockId="1.[151,1437,1729,2045]" pageId="1" pageNumber="353">
<emphasis id="B953A02BCB6B2C7FFF2BF8E6FDB5302D" bold="true" box="[199,614,1873,1900]" pageId="1" pageNumber="353">Diagnosis (family, genus, species):</emphasis>
Distinct from all other North American chordeumatidans in the broad (ca. 1/4 width of trunk ring), flat, polydesmid-like paranota (
<figureCitation id="131C60BCCB6B2C7FFCA1F8C1FC4130D1" box="[845,914,1910,1936]" captionStart="FIGURES 24" captionStartId="3.[151,264,1809,1834]" captionTargetBox="[170,1419,181,1763]" captionTargetId="figure@3.[169,1419,181,1786]" captionTargetPageId="3" captionText="FIGURES 24. Urochordeuma bumpusi Silvestri, 1909. 2: Rings 1012 (10, 11, 12) of male. 3: Rings 28, 29 and telson (28, 29, t) of male. 4: Small cuticular nodules of metazonite, each with two acute, posteriorly pointing processes." figureDoi="http://doi.org/10.5281/zenodo.3772341" httpUri="https://zenodo.org/record/3772341/files/figure.png" pageId="1" pageNumber="353">Fig. 2</figureCitation>
) beginning on the third or fourth post-collum ring—not as in
<taxonomicName id="4C2707BACB6B2C7FFEA8F82DFDD630F5" authorityName="Silvestri" authorityYear="1909" box="[324,517,1946,1972]" class="Diplopoda" family="Rhiscosomididae" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="353" phylum="Arthropoda" rank="family">Rhiscosomididae</taxonomicName>
with narrow paranota (ca. 1/6 width of ring). Male first legpair uniquely modified, with a long mesobasal process on the femur nearly three times as long as the femur itself (
<figureCitation id="131C60BCCB6B2C7FFB64F809FB1C3099" box="[1160,1231,1982,2008]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="1" pageNumber="353">Figs 1</figureCitation>
,
<figureCitation id="131C60BCCB6B2C7FFB31F80AFB3E3099" box="[1245,1261,1981,2008]" captionStart="FIGURES 5, 6" captionStartId="5.[151,264,1974,1999]" captionTargetBox="[189,1377,181,1949]" captionTargetId="figure@5.[188,1402,181,1949]" captionTargetPageId="5" captionText="FIGURES 5, 6. Urochordeuma bumpusi Silvestri, 1909. 5: First legpair, posterior view. 6: Third legpair, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3772343" httpUri="https://zenodo.org/record/3772343/files/figure.png" pageId="1" pageNumber="353">5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C33D2FB2CB6B2C7CFF2BF855FACD3602" lastPageId="2" lastPageNumber="354" pageId="1" pageNumber="353" type="etymology">
<paragraph id="8B987C39CB6B2C7CFF2BF855FACD3602" blockId="1.[151,1437,1729,2045]" lastBlockId="2.[151,1437,152,683]" lastPageId="2" lastPageNumber="354" pageId="1" pageNumber="353">
<emphasis id="B953A02BCB6B2C7FFF2BF855FE8130BD" bold="true" box="[199,338,2018,2044]" pageId="1" pageNumber="353">Etymology:</emphasis>
Silvestri named the species to honor Hermon Carey Bumpus (
<date id="FF995AF9CB6B2C7FFBF2F855FB7230BC" box="[1054,1185,2018,2045]" pageId="1" pageNumber="353" value="1862" valueMax="1943">18621943</date>
), who at the time was the Director of the American Museum of Natural History in
<collectingRegion id="49E3B2DBCB682C7CFCD8FF2EFC7A37F3" box="[820,937,152,179]" country="United States of America" name="New York" pageId="2" pageNumber="354">New York</collectingRegion>
. Bumpus had a distinguished career, heading the AMNH, the Marine Biological Laboratory, and the
<collectingCountry id="F3303CA9CB682C7CFCEAFF0AFCEB3796" box="[774,824,189,215]" name="United States of America" pageId="2" pageNumber="354">U.S.</collectingCountry>
Bureau of Fisheries Laboratory (Woods Hole,
<collectingRegion id="49E3B2DBCB682C7CFAA7FF0AFF2537BA" country="United States of America" name="Massachusetts" pageId="2" pageNumber="354">Massachusetts</collectingRegion>
). He later served as president of Tufts University (http://en.wikipedia.org/wiki/Hermon_Carey_Bumpus). Chamberlin did not explain his choice of a species name, and there is no Greek or Latin word
<emphasis id="B953A02BCB682C7CFB56FEB2FAC2365F" box="[1210,1297,261,286]" italics="true" pageId="2" pageNumber="354">
<taxonomicName id="4C2707BACB682C7CFB56FEB2FADD365F" authorityName="Chamberlin" authorityYear="1941" box="[1210,1294,261,286]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="354" phylum="Arthropoda" rank="species" species="porona">porona</taxonomicName>
.
</emphasis>
Chamberlin likely adapted the word from ancient Greek,
<emphasis id="B953A02BCB682C7CFD61FE9EFD003602" box="[653,723,297,323]" italics="true" pageId="2" pageNumber="354">póros,</emphasis>
“passage” perhaps in reference to Snoqualmie Pass.
</paragraph>
</subSubSection>
<subSubSection id="C33D2FB2CB682C79FF2BFEFBFCAF365E" lastPageId="7" lastPageNumber="359" pageId="2" pageNumber="354" type="description">
<paragraph id="8B987C39CB682C7CFF2BFEFBFC083627" blockId="2.[151,1437,152,683]" box="[199,987,332,359]" pageId="2" pageNumber="354">
<emphasis id="B953A02BCB682C7CFF2BFEFBFE883626" bold="true" box="[199,347,332,359]" pageId="2" pageNumber="354">Description:</emphasis>
Male from Horseshoe Lake, Pierce County,
<collectingRegion id="49E3B2DBCB682C7CFCA2FEFBFC043627" box="[846,983,332,358]" country="United States of America" name="Washington" pageId="2" pageNumber="354">Washington</collectingRegion>
.
</paragraph>
<paragraph id="8B987C39CB682C7CFF2BFEC7FD1236CA" blockId="2.[151,1437,152,683]" box="[199,705,368,395]" pageId="2" pageNumber="354">
Length about
<quantity id="4CDFD1DCCB682C7CFE8FFEC7FE6736CA" box="[355,436,368,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="354" unit="mm" value="15.0">15 mm</quantity>
, greatest width
<quantity id="4CDFD1DCCB682C7CFD88FEC6FD6E36CA" box="[612,701,369,395]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="2" pageNumber="354" unit="mm" value="2.7">2.7 mm</quantity>
.
</paragraph>
<paragraph id="8B987C39CB682C7CFF2BFE23FA593692" blockId="2.[151,1437,152,683]" pageId="2" pageNumber="354">
Head densely setose. Subtriangular eyepatches with 1416 ocelli, well pigmented. Labrum (
<emphasis id="B953A02BCB682C7CFB50FE22FB3236EE" bold="true" box="[1212,1249,405,431]" pageId="2" pageNumber="354">lab</emphasis>
,
<figureCitation id="131C60BCCB682C7CFB01FE23FAF836EE" box="[1261,1323,404,431]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="2" pageNumber="354">Fig. 1</figureCitation>
) strongly, broadly rebordered, with paired subacute nodules. Stipes of mandibles (
<emphasis id="B953A02BCB682C7CFC50FE0EFC0D3692" bold="true" box="[956,990,441,467]" pageId="2" pageNumber="354">ms</emphasis>
,
<figureCitation id="131C60BCCB682C7CFC00FE0FFBFF3692" box="[1004,1068,440,467]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="2" pageNumber="354">Fig. 1</figureCitation>
) with distal triangular process.
</paragraph>
<paragraph id="8B987C39CB682C7CFF2BFE6BFB4735EB" blockId="2.[151,1437,152,683]" pageId="2" pageNumber="354">Collum as long as wide, extended ventrolaterally, highly arched. Ring 2 with small paranota, paranota increasing in size on more posterior rings, reaching maximum width by ring 7; strongly rebordered, rebordering extends across width of metazonite. Metazonites set with many small tubercles, each bearing two acute projections, somewhat obscured by cerotegument. Metazonites posteriorly with four low, rounded bumps more prominent on midbody to posterior rings. Arrangement of metazonital setae unique: lateralmost seta in notch on paranotal margin, paralateral seta a short distance mesad, mesal seta much more anterior, near median sulcus.</paragraph>
<caption id="DF582CB1CB682C7CFF7BF8EEFCD6309F" ID-DOI="http://doi.org/10.5281/zenodo.3772339" ID-Zenodo-Dep="3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="2" pageNumber="354" startId="2.[151,250,1881,1906]" targetBox="[151,1436,704,1862]" targetPageId="2">
<paragraph id="8B987C39CB682C7CFF7BF8EEFCD6309F" blockId="2.[151,1436,1881,2014]" pageId="2" pageNumber="354">
<emphasis id="B953A02BCB682C7CFF7BF8EEFEC33033" bold="true" box="[151,272,1881,1906]" pageId="2" pageNumber="354">FIGURE 1</emphasis>
. Head and first four rings of
<taxonomicName id="4C2707BACB682C7CFDA6F8EDFC373033" authority="Silvestri, 1909" authorityName="Silvestri" authorityYear="1909" box="[586,996,1881,1906]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="2" pageNumber="354" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB682C7CFDA6F8EDFC913033" box="[586,834,1882,1906]" italics="true" pageId="2" pageNumber="354">Urochordeuma bumpusi</emphasis>
<bibRefCitation id="EFB601C8CB682C7CFCA7F8EEFC373033" author="Silvestri, F." box="[843,996,1881,1906]" pageId="2" pageNumber="354" pagination="229 - 233" refId="ref3903" refString="Silvestri, F. (1909) Descrizioni preliminari di varii Artropodi, specialmente d' America. III. Nuovi Chordeumoidea (Diplopoda). Rendiconti, Accademia Nazionale dei Lincei, 5. Ser. / Classe di Scienze Fisiche, Matematiche e Naturali, 18, 229 - 233. Available from: https: // biodiversitylibrary. org / page / 44698853 (Accessed 16 Aug. 2019)" type="journal article" year="1909">Silvestri, 1909</bibRefCitation>
</taxonomicName>
, male, ventral view.
<emphasis id="B953A02BCB682C7CFB2CF8EEFA453033" bold="true" box="[1216,1430,1881,1906]" pageId="2" pageNumber="354">Abbreviations: Lab</emphasis>
, labrum;
<emphasis id="B953A02BCB682C7CFF00F8C9FED830D7" bold="true" box="[236,267,1918,1942]" pageId="2" pageNumber="354">ms</emphasis>
, mandibular stipes;
<emphasis id="B953A02BCB682C7CFE34F8CAFE2430D7" bold="true" box="[472,503,1917,1942]" pageId="2" pageNumber="354">st1</emphasis>
, sternum of first legpair;
<emphasis id="B953A02BCB682C7CFD14F8CAFC9330D7" bold="true" box="[760,832,1917,1942]" pageId="2" pageNumber="354">L1L4</emphasis>
, legpairs 14 respectively;
<emphasis id="B953A02BCB682C7CFBBAF8CAFBAA30D7" bold="true" box="[1110,1145,1917,1942]" pageId="2" pageNumber="354">fp1</emphasis>
, femoral process of first leg;
<emphasis id="B953A02BCB682C7CFF7BF816FF6F30FB" bold="true" box="[151,188,1953,1978]" pageId="2" pageNumber="354">cx3</emphasis>
, coxa of third leg;
<emphasis id="B953A02BCB682C7CFE93F816FE4530FB" bold="true" box="[383,406,1953,1978]" pageId="2" pageNumber="354">s4</emphasis>
, sternal knob of legpair 4;
<emphasis id="B953A02BCB682C7CFD47F815FD6C30FB" bold="true" box="[683,703,1954,1978]" pageId="2" pageNumber="354">ct</emphasis>
, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue.
</paragraph>
</caption>
<caption id="DF582CB1CB692C7DFF7BF8A6FB6F300C" ID-DOI="http://doi.org/10.5281/zenodo.3772341" ID-Zenodo-Dep="3772341" httpUri="https://zenodo.org/record/3772341/files/figure.png" pageId="3" pageNumber="355" startId="3.[151,264,1809,1834]" targetBox="[170,1419,181,1763]" targetPageId="3">
<paragraph id="8B987C39CB692C7DFF7BF8A6FB6F300C" blockId="3.[151,1436,1809,1870]" pageId="3" pageNumber="355">
<emphasis id="B953A02BCB692C7DFF7BF8A6FEE0306B" bold="true" box="[151,307,1809,1834]" pageId="3" pageNumber="355">FIGURES 24</emphasis>
.
<taxonomicName id="4C2707BACB692C7DFED3F8A5FD1F306B" authority="Silvestri, 1909" authorityName="Silvestri" authorityYear="1909" box="[319,716,1809,1834]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="3" pageNumber="355" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB692C7DFED3F8A5FDE0306B" box="[319,563,1810,1834]" italics="true" pageId="3" pageNumber="355">Urochordeuma bumpusi</emphasis>
<bibRefCitation id="EFB601C8CB692C7DFDD5F8A6FD1F306B" author="Silvestri, F." box="[569,716,1809,1834]" pageId="3" pageNumber="355" pagination="229 - 233" refId="ref3903" refString="Silvestri, F. (1909) Descrizioni preliminari di varii Artropodi, specialmente d' America. III. Nuovi Chordeumoidea (Diplopoda). Rendiconti, Accademia Nazionale dei Lincei, 5. Ser. / Classe di Scienze Fisiche, Matematiche e Naturali, 18, 229 - 233. Available from: https: // biodiversitylibrary. org / page / 44698853 (Accessed 16 Aug. 2019)" type="journal article" year="1909">Silvestri, 1909</bibRefCitation>
</taxonomicName>
.
<emphasis id="B953A02BCB692C7DFD3BF8A6FD37306B" bold="true" box="[727,740,1809,1834]" pageId="3" pageNumber="355">2</emphasis>
: Rings 1012 (
<emphasis id="B953A02BCB692C7DFC92F8A6FC33306B" bold="true" box="[894,992,1809,1834]" pageId="3" pageNumber="355">10, 11, 12</emphasis>
) of male.
<emphasis id="B953A02BCB692C7DFBAAF8A6FB80306B" bold="true" box="[1094,1107,1809,1834]" pageId="3" pageNumber="355">3</emphasis>
: Rings 28, 29 and telson (
<emphasis id="B953A02BCB692C7DFABBF8A6FF4C300F" bold="true" pageId="3" pageNumber="355">28, 29, t</emphasis>
) of male.
<emphasis id="B953A02BCB692C7DFEE4F882FEC6300F" bold="true" box="[264,277,1845,1870]" pageId="3" pageNumber="355">4</emphasis>
: Small cuticular nodules of metazonite, each with two acute, posteriorly pointing processes.
</paragraph>
</caption>
<paragraph id="8B987C39CB692C7DFF2BF8CBFC1530D7" blockId="3.[199,1436,1916,2015]" box="[199,966,1916,1942]" pageId="3" pageNumber="355">
Epiproct shallowly trilobed (
<emphasis id="B953A02BCB692C7DFDE6F8CBFDC930D7" bold="true" box="[522,538,1916,1942]" pageId="3" pageNumber="355">t,</emphasis>
<figureCitation id="131C60BCCB692C7DFDCDF8CBFDB730D7" box="[545,612,1916,1942]" captionStart="FIGURES 24" captionStartId="3.[151,264,1809,1834]" captionTargetBox="[170,1419,181,1763]" captionTargetId="figure@3.[169,1419,181,1786]" captionTargetPageId="3" captionText="FIGURES 24. Urochordeuma bumpusi Silvestri, 1909. 2: Rings 1012 (10, 11, 12) of male. 3: Rings 28, 29 and telson (28, 29, t) of male. 4: Small cuticular nodules of metazonite, each with two acute, posteriorly pointing processes." figureDoi="http://doi.org/10.5281/zenodo.3772341" httpUri="https://zenodo.org/record/3772341/files/figure.png" pageId="3" pageNumber="355">Fig. 3</figureCitation>
), spinnerets directed anteriorly.
</paragraph>
<paragraph id="8B987C39CB692C7DFF2BF817FCD030FB" blockId="3.[199,1436,1916,2015]" box="[199,771,1952,1979]" pageId="3" pageNumber="355">
First sternum broad (
<emphasis id="B953A02BCB692C7DFE5FF817FE0630FB" bold="true" box="[435,469,1952,1978]" pageId="3" pageNumber="355">st1</emphasis>
,
<figureCitation id="131C60BCCB692C7DFE0FF817FDF030FB" box="[483,547,1952,1978]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="3" pageNumber="355">Fig. 1</figureCitation>
), greatly expanded.
</paragraph>
<paragraph id="8B987C39CB692C7AFF2BF873FB9D3626" blockId="3.[199,1436,1916,2015]" lastBlockId="4.[151,1437,152,2015]" lastPageId="4" lastPageNumber="356" pageId="3" pageNumber="355">
First legpair (
<emphasis id="B953A02BCB692C7DFEB2F873FE56309F" bold="true" box="[350,389,1988,2014]" pageId="3" pageNumber="355">L1,</emphasis>
<figureCitation id="131C60BCCB692C7DFE67F873FE1E309F" box="[395,461,1988,2014]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="3" pageNumber="355">Fig. 1</figureCitation>
;
<figureCitation id="131C60BCCB692C7DFE36F873FDC9309F" box="[474,538,1987,2014]" captionStart="FIGURES 5, 6" captionStartId="5.[151,264,1974,1999]" captionTargetBox="[189,1377,181,1949]" captionTargetId="figure@5.[188,1402,181,1949]" captionTargetPageId="5" captionText="FIGURES 5, 6. Urochordeuma bumpusi Silvestri, 1909. 5: First legpair, posterior view. 6: Third legpair, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3772343" httpUri="https://zenodo.org/record/3772343/files/figure.png" pageId="3" pageNumber="355">Fig. 5</figureCitation>
) with six podomeres (lacking trochanter), as long as subsequent legpairs, femora with long basomesal processes (
<emphasis id="B953A02BCB6E2C7AFE12FF2FFDFF37F2" bold="true" box="[510,556,152,179]" pageId="4" pageNumber="356">fp1,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFDDEFF2EFDA637F2" box="[562,629,153,179]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
:
<figureCitation id="131C60BCCB6E2C7AFD6DFF2EFD1237F2" box="[641,705,152,179]" captionStart="FIGURES 5, 6" captionStartId="5.[151,264,1974,1999]" captionTargetBox="[189,1377,181,1949]" captionTargetId="figure@5.[188,1402,181,1949]" captionTargetPageId="5" captionText="FIGURES 5, 6. Urochordeuma bumpusi Silvestri, 1909. 5: First legpair, posterior view. 6: Third legpair, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3772343" httpUri="https://zenodo.org/record/3772343/files/figure.png" pageId="4" pageNumber="356">Fig. 5</figureCitation>
) three times the length of the femora proper, tapering to blunt tip; anteriodistal surface of femur with many small teeth. Second legpair (
<emphasis id="B953A02BCB6E2C7AFC59FF0AFC0F3796" bold="true" box="[949,988,189,215]" pageId="4" pageNumber="356">L2,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFC09FF0AFBF63796" box="[997,1061,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
) unmodified but with prominent spermatic processes. Third legpair (
<emphasis id="B953A02BCB6E2C7AFDC7FF56FD8137BA" bold="true" box="[555,594,225,251]" pageId="4" pageNumber="356">L3,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFDB6FF56FD4E37BA" box="[602,669,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
;
<figureCitation id="131C60BCCB6E2C7AFD40FF56FD3C37BD" box="[684,751,225,252]" captionStart="FIGURES 5, 6" captionStartId="5.[151,264,1974,1999]" captionTargetBox="[189,1377,181,1949]" captionTargetId="figure@5.[188,1402,181,1949]" captionTargetPageId="5" captionText="FIGURES 5, 6. Urochordeuma bumpusi Silvestri, 1909. 5: First legpair, posterior view. 6: Third legpair, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3772343" httpUri="https://zenodo.org/record/3772343/files/figure.png" pageId="4" pageNumber="356">Fig. 6</figureCitation>
) coxae (
<emphasis id="B953A02BCB6E2C7AFCBEFF56FC5137BA" bold="true" box="[850,898,225,251]" pageId="4" pageNumber="356">cx3,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFC65FF56FC1F37BA" box="[905,972,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
;
<figureCitation id="131C60BCCB6E2C7AFC37FF56FBCD37BD" box="[987,1054,225,252]" captionStart="FIGURES 5, 6" captionStartId="5.[151,264,1974,1999]" captionTargetBox="[189,1377,181,1949]" captionTargetId="figure@5.[188,1402,181,1949]" captionTargetPageId="5" captionText="FIGURES 5, 6. Urochordeuma bumpusi Silvestri, 1909. 5: First legpair, posterior view. 6: Third legpair, posterior view." figureDoi="http://doi.org/10.5281/zenodo.3772343" httpUri="https://zenodo.org/record/3772343/files/figure.png" pageId="4" pageNumber="356">Fig. 6</figureCitation>
) extended as long lobes nearly as long as legs themselves, finely toothed on anterodistal surface; femora distally inflated. Fourth sternum bearing anterior knob (
<emphasis id="B953A02BCB6E2C7AFED2FE9EFE8D3602" bold="true" box="[318,350,297,323]" pageId="4" pageNumber="356">s4,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFE8AFE9EFE763602" box="[358,421,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
) projecting between coxae of legpair three. Prefemora of legpair four (
<emphasis id="B953A02BCB6E2C7AFB24FE9EFB233602" bold="true" box="[1224,1264,297,323]" pageId="4" pageNumber="356">L4,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFB1BFE9EFAE43602" box="[1271,1335,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
) distally inflated. Legpairs five and six unmodified. Legpair seven with lamellate coxal lobes.
</paragraph>
<paragraph id="8B987C39CB6E2C7AFF2BFEC6FEBA35C6" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="356">
Gonopod sternum (
<figureCitation id="131C60BCCB6E2C7AFE4DFEC6FE3536CA" box="[417,486,369,395]" captionStart="FIGURES 711" captionStartId="6.[151,264,1900,1925]" captionTargetBox="[169,1419,181,1846]" captionTargetId="figure@6.[169,1419,181,1877]" captionTargetPageId="6" captionText="FIGURES 711. Urochordeuma bumpusi. 7: Gonopods, anterior view. 8: Left gonopod, mesal view. 9: Tip of right gonopod, lateral view. 10: Ninth legpair, anterior view. 11: Left vulva, ventral view. Abbreviations: acp, anterior coxal process; cx, coxa; cx9, coxa, cp9, coxal process, pcp, posterior coxal process; st, sternum; t9, telopodite" figureDoi="http://doi.org/10.5281/zenodo.3772345" httpUri="https://zenodo.org/record/3772345/files/figure.png" pageId="4" pageNumber="356">Fig. 7</figureCitation>
) large, complete, spiracles slitlike, possibly vestigial. Gonopod coxae basally fused, with anterior groups of setae. Gonopods with two distinct processes, each divided, tips projecting anteriorly (
<figureCitation id="131C60BCCB6E2C7AFA86FE22FF1E3692" captionStart="FIGURES 711" captionStartId="6.[151,264,1900,1925]" captionTargetBox="[169,1419,181,1846]" captionTargetId="figure@6.[169,1419,181,1877]" captionTargetPageId="6" captionText="FIGURES 711. Urochordeuma bumpusi. 7: Gonopods, anterior view. 8: Left gonopod, mesal view. 9: Tip of right gonopod, lateral view. 10: Ninth legpair, anterior view. 11: Left vulva, ventral view. Abbreviations: acp, anterior coxal process; cx, coxa; cx9, coxa, cp9, coxal process, pcp, posterior coxal process; st, sternum; t9, telopodite" figureDoi="http://doi.org/10.5281/zenodo.3772345" httpUri="https://zenodo.org/record/3772345/files/figure.png" pageId="4" pageNumber="356">Figs 7 9</figureCitation>
). Anterior process with lateral acuminate tip, mesal part broader, with appressed fimbria. Posterior process anterior tip with ventral ridge finely fimbriate, terminating in four small, subtriangular teeth; posterior tip deeply divided into many highly branched, finely acute processes. Ninth legpair (
<figureCitation id="131C60BCCB6E2C7AFC1FFDB6FB94355A" box="[1011,1095,513,539]" captionStart="FIGURES 711" captionStartId="6.[151,264,1900,1925]" captionTargetBox="[169,1419,181,1846]" captionTargetId="figure@6.[169,1419,181,1877]" captionTargetPageId="6" captionText="FIGURES 711. Urochordeuma bumpusi. 7: Gonopods, anterior view. 8: Left gonopod, mesal view. 9: Tip of right gonopod, lateral view. 10: Ninth legpair, anterior view. 11: Left vulva, ventral view. Abbreviations: acp, anterior coxal process; cx, coxa; cx9, coxa, cp9, coxal process, pcp, posterior coxal process; st, sternum; t9, telopodite" figureDoi="http://doi.org/10.5281/zenodo.3772345" httpUri="https://zenodo.org/record/3772345/files/figure.png" pageId="4" pageNumber="356">Fig. 10</figureCitation>
) reduced to coxae and single telopodite articles; coxae bearing prominent terminal hooks anteriorly directed. Telopodite articles densely setose mesally, lateral surfaces granular, with small, nipple-like projections. Coxae of legpair 10 with small glands, otherwise not modified.
</paragraph>
<paragraph id="8B987C39CB6E2C7AFF2BFD26FA87358E" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="356">
Female similar to male but lacking secondary sexual modification of anterior legs. Collum significantly smaller than in male, slightly longer than wide, not strongly arched or extended ventrolaterally. Vulvae as in
<figureCitation id="131C60BCCB6E2C7AFB13FD02FA83358E" box="[1279,1360,693,719]" captionStart="FIGURES 711" captionStartId="6.[151,264,1900,1925]" captionTargetBox="[169,1419,181,1846]" captionTargetId="figure@6.[169,1419,181,1877]" captionTargetPageId="6" captionText="FIGURES 711. Urochordeuma bumpusi. 7: Gonopods, anterior view. 8: Left gonopod, mesal view. 9: Tip of right gonopod, lateral view. 10: Ninth legpair, anterior view. 11: Left vulva, ventral view. Abbreviations: acp, anterior coxal process; cx, coxa; cx9, coxa, cp9, coxal process, pcp, posterior coxal process; st, sternum; t9, telopodite" figureDoi="http://doi.org/10.5281/zenodo.3772345" httpUri="https://zenodo.org/record/3772345/files/figure.png" pageId="4" pageNumber="356">Fig. 11</figureCitation>
.
</paragraph>
<paragraph id="8B987C39CB6E2C7AFF2BFD6EFD5733AA" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="356">
<emphasis id="B953A02BCB6E2C7AFF2BFD6EFEB635B2" bold="true" box="[199,357,729,755]" pageId="4" pageNumber="356">New Records</emphasis>
: See
<figureCitation id="131C60BCCB6E2C7AFE4EFD6EFE2135B2" box="[418,498,729,755]" captionStart="FIGURES 12" captionStartId="7.[151,264,1480,1505]" captionTargetBox="[163,1425,309,1461]" captionTargetId="figure@7.[163,1425,309,1461]" captionTargetPageId="7" captionText="FIGURES 12. Northwest Washington, showing records of Urochordeuma bumpusi. Circles, new records. Triangle, type locality of U. bumpusi. Square, type locality of U. porona." figureDoi="http://doi.org/10.5281/zenodo.3772347" httpUri="https://zenodo.org/record/3772347/files/figure.png" pageId="4" pageNumber="356">Fig. 12</figureCitation>
for the complete distribution as presently understood, including the two
<typeStatus id="549CC29BCB6E2C7AFAF2FD6EFA9D35B2" box="[1310,1358,729,755]" pageId="4" pageNumber="356">type</typeStatus>
localities. Unless noted, all collections by Rod Crawford and all material deposited in the University of
<collectingRegion id="49E3B2DBCB6E2C7AFB21FD4AFA813456" box="[1229,1362,765,791]" country="United States of America" name="Washington" pageId="4" pageNumber="356">Washington</collectingRegion>
Burke Museum.
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:
<emphasis id="B953A02BCB6E2C7AFE23FC97FDE0347A" bold="true" box="[463,563,800,827]" pageId="4" pageNumber="356">King Co</emphasis>
.: Bridle Trails State Park, from maple leaf litter, 500 (
<quantity id="4CDFD1DCCB6E2C7AFB7AFC97FB3E347A" box="[1174,1261,800,827]" metricMagnitude="2" metricUnit="m" metricValue="1.524" pageId="4" pageNumber="356" unit="m" value="152.4">152.4m</quantity>
) asl,
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,
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,
<date id="FF995AF9CB6E2C7AFEC8FCF2FE01341E" box="[292,466,836,863]" pageId="4" pageNumber="356" value="1985-03-16">16 March 1985</date>
,
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m;
<location id="8EF82AE2CB6E2C7AFDEFFCF2FC99341E" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFDEFFCF2FC99341E" box="[515,842,837,863]" latitude="47.291" longLatPrecision="66" longitude="-121.985" name="Southeast of Black Diamond" pageId="4" pageNumber="356">Southeast of Black Diamond</location>
, from mixed leaf litter, 720 (
<quantity id="4CDFD1DCCB6E2C7AFB78FCF3FB3F341E" box="[1172,1260,836,863]" metricMagnitude="2" metricUnit="m" metricValue="2.195" pageId="4" pageNumber="356" unit="m" value="219.5">
<elevation id="000A9B0ACB6E2C7AFB78FCF3FB3F341E" box="[1172,1260,836,863]" metricMagnitude="2" metricUnit="m" metricValue="2.195" pageId="4" pageNumber="356" unit="m" value="219.5">219.5m</elevation>
</quantity>
) asl,
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,
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,
<date id="FF995AF9CB6E2C7AFECEFCDEFE3E34C2" box="[290,493,873,899]" pageId="4" pageNumber="356" value="2006-11-01">
<collectingDate id="EFDDA311CB6E2C7AFECEFCDEFE3E34C2" box="[290,493,873,899]" pageId="4" pageNumber="356" value="2006-11-01">1 November 2006</collectingDate>
</date>
</materialsCitation>
,
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m;
<location id="8EF82AE2CB6E2C7AFDF7FCDEFD7734C2" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFDF7FCDEFD7734C2" box="[539,676,873,899]" latitude="47.5" longLatPrecision="6704" longitude="-121.8" name="Snoqualmie" pageId="4" pageNumber="356">Snoqualmie</location>
,
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,
<geoCoordinate id="EE131AFECB6E2C7AFCE7FCDEFCBC34C2" box="[779,879,873,899]" degrees="121.8" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="5555" value="-121.8">121.8°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFC96FCDEFB9134C2" box="[890,1090,873,899]" pageId="4" pageNumber="356" value="1940-02-22">
<collectingDate id="EFDDA311CB6E2C7AFC96FCDEFB9134C2" box="[890,1090,873,899]" pageId="4" pageNumber="356" value="1940-02-22">22 February 1940</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4F7664CB6E2C7AFBA0FCDEFCEA34E6" ID-GBIF-Occurrence="2609011259" collectingDate="2004-02-25" collectorName="N. Kobylk" latitude="121.668" location="Twin Falls" municipality="Iron Horse" pageId="4" pageNumber="356" specimenCount="1">
<collectorName id="26D219EFCB6E2C7AFBA0FCDEFB1734C2" box="[1100,1220,873,899]" pageId="4" pageNumber="356">N. Kobylk</collectorName>
, m,
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/
<collectingMunicipality id="6BFCE643CB6E2C7AFA81FCDEFF0934E6" pageId="4" pageNumber="356">Iron Horse</collectingMunicipality>
trailhead,
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,
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,
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<collectingDate id="EFDDA311CB6E2C7AFD83FC3BFCEA34E6" box="[623,825,908,935]" pageId="4" pageNumber="356" value="2004-02-25">25 February 2004</collectingDate>
</date>
</materialsCitation>
,
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W.
<location id="8EF82AE2CB6E2C7AFC86FC3AFC1934E6" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFC86FC3AFC1934E6" box="[874,970,909,935]" county="Schmitz Park" latitude="47.574" longLatPrecision="66" longitude="-122.398" municipality="Richart" name="Leonard" pageId="4" pageNumber="356">Leonard</location>
, C.
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, ff;
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, from maple litter, 150 (
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) asl,
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,
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,
<date id="FF995AF9CB6E2C7AFD50FC06FCAD348A" box="[700,894,944,971]" pageId="4" pageNumber="356" value="1985-10-13">
<collectingDate id="EFDDA311CB6E2C7AFD50FC06FCAD348A" box="[700,894,944,971]" pageId="4" pageNumber="356" value="1985-10-13">13 October 1985</collectingDate>
</date>
</materialsCitation>
,
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m, f;
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, 440460 (
<quantity id="4CDFD1DCCB6E2C7AFB3BFC06FA46348A" box="[1239,1429,945,971]" metricMagnitude="2" metricUnit="m" metricValue="1.3714999999999997" metricValueMax="1.402" metricValueMin="1.341" pageId="4" pageNumber="356" unit="m" value="137.14999999999998" valueMax="140.2" valueMin="134.1">
<elevation id="000A9B0ACB6E2C7AFB3BFC06FA46348A" box="[1239,1429,945,971]" metricMagnitude="2" metricUnit="m" metricValue="1.3714999999999997" metricValueMax="1.402" metricValueMin="1.341" pageId="4" pageNumber="356" unit="m" value="137.14999999999998" valueMax="140.2" valueMin="134.1">134.1 to 140.2m</elevation>
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) asl,
<geoCoordinate id="EE131AFECB6E2C7AFF2AFC62FEF434AE" box="[198,295,981,1007]" degrees="47.49" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="555" value="47.49">47.49°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFED9FC62FE7434AE" box="[309,423,981,1007]" degrees="121.78" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="555" value="-121.78">121.78°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFE58FC62FD8434AE" box="[436,599,981,1007]" pageId="4" pageNumber="356" value="1896-04-21">
<collectingDate id="EFDDA311CB6E2C7AFE58FC62FD8434AE" box="[436,599,981,1007]" pageId="4" pageNumber="356" value="1896-04-21">21 April 1896</collectingDate>
</date>
,
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</materialsCitation>
.
<materialsCitation id="3B4F7664CB6E2C7AFD6EFC62FE1E3352" ID-GBIF-Occurrence="2609011269" collectingDate="2003-10-25" collectorName="Pratt Lake Trailhead, N &amp; W. Leonard" latitude="-121.4852" location="Kincaid" pageId="4" pageNumber="356" specimenCount="1">
<location id="8EF82AE2CB6E2C7AFD6EFC62FD0D34AE" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFD6EFC62FD0D34AE" box="[642,734,981,1007]" latitude="-121.4852" name="Kincaid" pageId="4" pageNumber="356">Kincaid</location>
;
<collectorName id="26D219EFCB6E2C7AFD00FC62FC2B34AE" box="[748,1016,981,1007]" pageId="4" pageNumber="356">Pratt Lake Trailhead, N</collectorName>
of I-90,
<geoCoordinate id="EE131AFECB6E2C7AFB8CFC62FB0E34AE" box="[1120,1245,981,1007]" degrees="47.3972" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="5" value="47.3972">47.3972°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFB06FC63FAA234AE" box="[1258,1393,980,1007]" degrees="121.4852" direction="south" orientation="latitude" pageId="4" pageNumber="356" precision="5" value="-121.4852">121.4852°S</geoCoordinate>
,
<collectorName id="26D219EFCB6E2C7AFA91FC62FF243352" pageId="4" pageNumber="356">W. Leonard</collectorName>
,
<date id="FF995AF9CB6E2C7AFEEAFC4FFE1E3352" box="[262,461,1016,1043]" pageId="4" pageNumber="356" value="2003-10-25">
<collectingDate id="EFDDA311CB6E2C7AFEEAFC4FFE1E3352" box="[262,461,1016,1043]" pageId="4" pageNumber="356" value="2003-10-25">25 October 2003</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4F7664CB6E2C7AFE30FC4EFD993376" ID-GBIF-Occurrence="2609011265" collectingDate="2006-06-07" latitude="47.3973" location="Tinken Road" longLatPrecision="6" longitude="-121.4853" municipality="Snoqualmie National Forest" pageId="4" pageNumber="356" specimenCount="1">
juvs;
<location id="8EF82AE2CB6E2C7AFDF1FC4EFD623352" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFDF1FC4EFD623352" box="[541,689,1017,1043]" latitude="47.3973" longLatPrecision="6" longitude="-121.4853" municipality="Snoqualmie National Forest" name="Tinken Road" pageId="4" pageNumber="356">Tinken Road</location>
exit, north side of I-90,
<collectingMunicipality id="6BFCE643CB6E2C7AFC34FC4EFACE3352" box="[984,1309,1017,1043]" pageId="4" pageNumber="356">Snoqualmie National Forest</collectingMunicipality>
, 1750 asl,
<geoCoordinate id="EE131AFECB6E2C7AFF7BFBAAFEC73376" box="[151,276,1053,1079]" degrees="47.3973" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="5" value="47.3973">47.3973°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFECDFBABFE633376" box="[289,432,1052,1079]" degrees="121.4853" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="5" value="-121.4853">121.4853°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFE50FBAAFD993376" box="[444,586,1053,1079]" pageId="4" pageNumber="356" value="2006-06-07">
<collectingDate id="EFDDA311CB6E2C7AFE50FBAAFD993376" box="[444,586,1053,1079]" pageId="4" pageNumber="356" value="2006-06-07">7 June 2006</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B4F7664CB6E2C7AFDB9FBAAFC8F331A" ID-GBIF-Occurrence="2609011264" collectingDate="1980-07-12" county="Pierce Co." elevation="671" latitude="46.817" location="Leonard" longLatPrecision="67" longitude="-122.151" municipality="North of Little Mashel River" pageId="4" pageNumber="356" specimenCount="1">
W.
<location id="8EF82AE2CB6E2C7AFD90FBAAFD0F3376" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFD90FBAAFD0F3376" box="[636,732,1053,1079]" county="Pierce Co." latitude="46.817" longLatPrecision="67" longitude="-122.151" municipality="North of Little Mashel River" name="Leonard" pageId="4" pageNumber="356">Leonard</location>
, juvenile.
<collectingCounty id="62F904B5CB6E2C7AFCBEFBABFC1E3376" box="[850,973,1052,1079]" pageId="4" pageNumber="356">
<emphasis id="B953A02BCB6E2C7AFCBEFBABFC153376" bold="true" box="[850,966,1052,1079]" pageId="4" pageNumber="356">Pierce Co</emphasis>
.
</collectingCounty>
:
<collectingMunicipality id="6BFCE643CB6E2C7AFC31FBAAFAF53376" box="[989,1318,1053,1079]" pageId="4" pageNumber="356">North of Little Mashel River</collectingMunicipality>
, under cut wood, 2200 (
<quantity id="4CDFD1DCCB6E2C7AFEDDFBF6FE5A331D" box="[305,393,1089,1116]" metricMagnitude="2" metricUnit="m" metricValue="6.706" pageId="4" pageNumber="356" unit="m" value="670.6">
<elevation id="000A9B0ACB6E2C7AFEDDFBF6FE5A331D" box="[305,393,1089,1116]" metricMagnitude="2" metricUnit="m" metricValue="6.706" pageId="4" pageNumber="356" unit="m" value="670.6">670.6m</elevation>
</quantity>
) asl,
<geoCoordinate id="EE131AFECB6E2C7AFE28FBF6FDE0331A" box="[452,563,1089,1115]" degrees="46.817" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="55" value="46.817">46.817°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFDD2FBF7FD6D331A" box="[574,702,1088,1115]" degrees="122.151" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="55" value="-122.151">122.151°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFD26FBF6FC8F331A" box="[714,860,1089,1115]" pageId="4" pageNumber="356" value="1980-07-12">
<collectingDate id="EFDDA311CB6E2C7AFD26FBF6FC8F331A" box="[714,860,1089,1115]" pageId="4" pageNumber="356" value="1980-07-12">12 July 1980</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4F7664CB6E2C7AFC8BFBF6FD45333E" ID-GBIF-Occurrence="2609011268" collectingDate="1977-09-04" elevation="229" latitude="46.915" location="Horseshoe Lake" longLatPrecision="67" longitude="-122.271" pageId="4" pageNumber="356" specimenCount="1">
m;
<location id="8EF82AE2CB6E2C7AFC67FBF6FB90331A" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFC67FBF6FB90331A" box="[907,1091,1089,1115]" latitude="46.915" longLatPrecision="67" longitude="-122.271" name="Horseshoe Lake" pageId="4" pageNumber="356">Horseshoe Lake</location>
, from leaf litter, 750 (
<quantity id="4CDFD1DCCB6E2C7AFAD0FBF6FA46331D" box="[1340,1429,1089,1116]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="356" unit="m" value="228.6">
<elevation id="000A9B0ACB6E2C7AFAD0FBF6FA46331D" box="[1340,1429,1089,1116]" metricMagnitude="2" metricUnit="m" metricValue="2.286" pageId="4" pageNumber="356" unit="m" value="228.6">228.6m</elevation>
</quantity>
) asl,
<geoCoordinate id="EE131AFECB6E2C7AFF28FBD3FEE0333E" box="[196,307,1124,1151]" degrees="46.915" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="55" value="46.915">46.915°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFED1FBD2FE6D333E" box="[317,446,1125,1151]" degrees="122.271" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="55" value="-122.271">122.271°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFE25FBD2FD45333E" box="[457,662,1125,1151]" pageId="4" pageNumber="356" value="1977-09-04">
<collectingDate id="EFDDA311CB6E2C7AFE25FBD2FD45333E" box="[457,662,1125,1151]" pageId="4" pageNumber="356" value="1977-09-04">4 September 1977</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4F7664CB6E2C7AFD73FBD2FCE533E2" ID-GBIF-Occurrence="2609011267" collectingDate="2010-07-23" elevation="1463" latitude="47.008" location="Ruggles" longLatPrecision="67" longitude="-121.662" municipality="Lonesome Lake" pageId="4" pageNumber="356" specimenCount="1">
A.
<location id="8EF82AE2CB6E2C7AFD2CFBD2FCCC333E" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFD2CFBD2FCCC333E" box="[704,799,1125,1151]" latitude="47.008" longLatPrecision="67" longitude="-121.662" municipality="Lonesome Lake" name="Ruggles" pageId="4" pageNumber="356">Ruggles</location>
, m, f;
<collectingMunicipality id="6BFCE643CB6E2C7AFC88FBD2FBC9333E" box="[868,1050,1125,1151]" pageId="4" pageNumber="356">Lonesome Lake</collectingMunicipality>
, from dead wood fragments in forest, 4800 (
<quantity id="4CDFD1DCCB6E2C7AFEFEFB3EFEB133E5" box="[274,354,1161,1188]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="4" pageNumber="356" unit="m" value="1463.0">
<elevation id="000A9B0ACB6E2C7AFEFEFB3EFEB133E5" box="[274,354,1161,1188]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="4" pageNumber="356" unit="m" value="1463.0">1463m</elevation>
</quantity>
) asl,
<geoCoordinate id="EE131AFECB6E2C7AFE71FB3EFDDF33E2" box="[413,524,1161,1187]" degrees="47.008" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="55" value="47.008">47.008°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFDFBFB3EFD4A33E2" box="[535,665,1161,1187]" degrees="121.662" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="55" value="-121.662">121.662°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFD4FFB3EFCE533E2" box="[675,822,1161,1187]" pageId="4" pageNumber="356" value="2010-07-23">
<collectingDate id="EFDDA311CB6E2C7AFD4FFB3EFCE533E2" box="[675,822,1161,1187]" pageId="4" pageNumber="356" value="2010-07-23">23 July 2010</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4F7664CB6E2C7AFCACFB3EFE5C3386" ID-GBIF-Occurrence="2609011263" collectingDate="2003-10-12" county="Thurston Co." location="Hospital Creek" pageId="4" pageNumber="356" specimenCount="1">
m, f.
<collectingCounty id="62F904B5CB6E2C7AFC97FB3EFBC933E2" box="[891,1050,1160,1187]" pageId="4" pageNumber="356">
<emphasis id="B953A02BCB6E2C7AFC97FB3EFBC033E2" bold="true" box="[891,1043,1160,1187]" pageId="4" pageNumber="356">Thurston Co</emphasis>
.
</collectingCounty>
:
<location id="8EF82AE2CB6E2C7AFBC5FB3EFB0733E2" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFBC5FB3EFB0733E2" box="[1065,1236,1161,1187]" county="Thurston Co." name="Hospital Creek" pageId="4" pageNumber="356">Hospital Creek</location>
,
<quantity id="4CDFD1DCCB6E2C7AFB33FB3FFAC233E2" box="[1247,1297,1160,1187]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="4" pageNumber="356" unit="mi" value="5.0">5 mi</quantity>
S,
<locationDeviation id="9A2A651BCB6E2C7AFAD8FB3EFF143386" location="Vail" pageId="4" pageNumber="356">
<quantity id="4CDFD1DCCB6E2C7AFAD8FB3EFAB433E2" box="[1332,1383,1161,1187]" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="4" pageNumber="356" unit="mi" value="3.0">3 mi</quantity>
E of Vail
</locationDeviation>
,
<date id="FF995AF9CB6E2C7AFF3DFB1AFE5C3386" box="[209,399,1197,1223]" pageId="4" pageNumber="356" value="2003-10-12">
<collectingDate id="EFDDA311CB6E2C7AFF3DFB1AFE5C3386" box="[209,399,1197,1223]" pageId="4" pageNumber="356" value="2003-10-12">12 October 2003</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B4F7664CB6E2C7AFE74FB1AFD8F33AA" ID-GBIF-Occurrence="2609011260" collectingDate="1986-10-11" collectorName="W. Leonard" county="Whatcom Co." elevation="213" latitude="48.934" location="Sumas Mountain" longLatPrecision="66" longitude="-122.155" pageId="4" pageNumber="356" specimenCount="1">
<collectorName id="26D219EFCB6E2C7AFE74FB1AFDCB3386" box="[408,536,1197,1223]" pageId="4" pageNumber="356">W.Leonard</collectorName>
, juvs.
<emphasis id="B953A02BCB6E2C7AFDB1FB1AFCD23386" bold="true" box="[605,769,1196,1223]" pageId="4" pageNumber="356">
<collectingCounty id="62F904B5CB6E2C7AFDB1FB1AFCD23386" box="[605,769,1196,1223]" pageId="4" pageNumber="356">Whatcom Co.</collectingCounty>
</emphasis>
:
<location id="8EF82AE2CB6E2C7AFCE3FB1AFC033386" LSID="urn:lsid:plazi:treatment:038ECD2FCB6B2C79FF7BFA4DFCAF365E:8EF82AE2CB6E2C7AFCE3FB1AFC033386" box="[783,976,1197,1223]" county="Whatcom Co." latitude="48.934" longLatPrecision="66" longitude="-122.155" name="Sumas Mountain" pageId="4" pageNumber="356">Sumas Mountain</location>
, from maple leaf litter, 700 (
<quantity id="4CDFD1DCCB6E2C7AFAFCFB1AFABA3386" box="[1296,1385,1197,1223]" metricMagnitude="2" metricUnit="m" metricValue="2.134" pageId="4" pageNumber="356" unit="m" value="213.4">
<elevation id="000A9B0ACB6E2C7AFAFCFB1AFABA3386" box="[1296,1385,1197,1223]" metricMagnitude="2" metricUnit="m" metricValue="2.134" pageId="4" pageNumber="356" unit="m" value="213.4">213.4m</elevation>
</quantity>
) asl,
<geoCoordinate id="EE131AFECB6E2C7AFF7BFB66FED533AA" box="[151,262,1233,1259]" degrees="48.934" direction="north" orientation="latitude" pageId="4" pageNumber="356" precision="55" value="48.934">48.934°N</geoCoordinate>
,
<geoCoordinate id="EE131AFECB6E2C7AFEFEFB67FE4033AA" box="[274,403,1232,1259]" degrees="122.155" direction="west" orientation="longitude" pageId="4" pageNumber="356" precision="55" value="-122.155">122.155°W</geoCoordinate>
,
<date id="FF995AF9CB6E2C7AFE72FB66FD8F33AA" box="[414,604,1233,1259]" pageId="4" pageNumber="356" value="1986-10-11">
<collectingDate id="EFDDA311CB6E2C7AFE72FB66FD8F33AA" box="[414,604,1233,1259]" pageId="4" pageNumber="356" value="1986-10-11">11 October 1986</collectingDate>
</date>
</materialsCitation>
, m.
</paragraph>
<paragraph id="8B987C39CB6E2C7AFF2BFB42FBB631DA" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="356">
<emphasis id="B953A02BCB6E2C7AFF2BFB42FED9324E" bold="true" box="[199,266,1269,1295]" pageId="4" pageNumber="356">Notes</emphasis>
: The differences cited by Chamberlin in describing
<taxonomicName id="4C2707BACB6E2C7AFCB5FB41FC1F324E" authorityName="Chamberlin" authorityYear="1941" box="[857,972,1270,1295]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="356" phylum="Arthropoda" rank="species" species="porona">
<emphasis id="B953A02BCB6E2C7AFCB5FB41FC1F324E" box="[857,972,1270,1295]" italics="true" pageId="4" pageNumber="356">U. porona</emphasis>
</taxonomicName>
are not consistent when more specimens from multiple localities are examined. The maximum width of all mature specimens studied here ranged from
<quantity id="4CDFD1DCCB6E2C7AFA95FAAEFED83216" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="4" pageNumber="356" unit="mm" value="2.85" valueMax="3.1" valueMin="2.6">2.6 to 3.1 mm</quantity>
, and the proportions of the collum (of males) were similar in all. However, the collum is sexually dimorphic in the species, and is larger and more highly arched in males to accommodate the greatly enlarged sternum of the first legpair. This creates a commodious space (
<figureCitation id="131C60BCCB6E2C7AFD1AFA32FCE432DE" box="[758,823,1413,1439]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
) defined laterally by the femoral processes of the first legs and the coxal processes of the third legs; we speculate that this space and the first and third leg modifications are adaptations to firmly hold the females head during mating. Silvestri had both sexes available for study, while Chamberlin had only a single male, which may explain his statement regarding the length of the collum; but this highly three-dimensional structure can appear differently depending on viewing angle.
<bibRefCitation id="EFB601C8CB6E2C7AFB83F9A2FADD316E" author="Shear, W. A." box="[1135,1294,1557,1583]" pageId="4" pageNumber="356" pagination="151 - 352" refId="ref3794" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda). A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. - Bulletin of the Museum of Comparative Zoology, 144, 151 - 352. Available from: https: // biodiversitylibrary. org / page / 4227660 (Accessed 16 Aug. 2019)" type="journal article" year="1972">Shears (1972</bibRefCitation>
, fig. 446) illustration misplaces the paralateral setae, but
<bibRefCitation id="EFB601C8CB6E2C7AFD7CF98EFCED3112" author="Silvestri, F." box="[656,830,1593,1619]" pageId="4" pageNumber="356" pagination="303 - 310" refId="ref3975" refString="Silvestri, F. (1913) Illustrazione di due famiglie di Chordeumoidea (Diplopoda) del Nord America. Bollettino del Laboratorio di zoologia generale e agraria della Facolta agraria in Portici, 7, 303 - 310. Available from: https: // biodiversitylibrary. org / page / 13979142 (Accessed 16 Aug. 2019)" type="journal article" year="1913">Silvestri (1913)</bibRefCitation>
illustrated them correctly. The setae have been rubbed off the specimen used for
<figureCitation id="131C60BCCB6E2C7AFE56F9EAFE283136" box="[442,507,1629,1655]" captionStart="FIGURES 24" captionStartId="3.[151,264,1809,1834]" captionTargetBox="[170,1419,181,1763]" captionTargetId="figure@3.[169,1419,181,1786]" captionTargetPageId="3" captionText="FIGURES 24. Urochordeuma bumpusi Silvestri, 1909. 2: Rings 1012 (10, 11, 12) of male. 3: Rings 28, 29 and telson (28, 29, t) of male. 4: Small cuticular nodules of metazonite, each with two acute, posteriorly pointing processes." figureDoi="http://doi.org/10.5281/zenodo.3772341" httpUri="https://zenodo.org/record/3772341/files/figure.png" pageId="4" pageNumber="356">Fig. 2</figureCitation>
and the sockets are obscured by cerotegument (
<emphasis id="B953A02BCB6E2C7AFBF9F9EAFBE13136" bold="true" box="[1045,1074,1629,1655]" pageId="4" pageNumber="356">ct,</emphasis>
<figureCitation id="131C60BCCB6E2C7AFBD3F9EAFBAC3136" box="[1087,1151,1629,1655]" captionStart="FIGURE 1" captionStartId="2.[151,250,1881,1906]" captionTargetBox="[151,1436,704,1862]" captionTargetId="figure@2.[151,1436,704,1862]" captionTargetPageId="2" captionText="FIGURE 1. Head and first four rings of Urochordeuma bumpusi Silvestri, 1909, male, ventral view. Abbreviations: Lab, labrum; ms, mandibular stipes; st1, sternum of first legpair; L1L4, legpairs 14 respectively; fp1, femoral process of first leg; cx3, coxa of third leg; s4, sternal knob of legpair 4; ct, fragments of cerotegument pulled loose from cuticle. First legs shaded red, second legs yellow, third legs green and fourth legs blue." figureDoi="http://doi.org/10.5281/zenodo.3772339" httpUri="https://zenodo.org/record/3772339/files/figure.png" pageId="4" pageNumber="356">Fig. 1</figureCitation>
). While many specimens have a definite cerotegument, some males do not—possibly they have recently molted.
</paragraph>
<paragraph id="8B987C39CB6E2C7AFF2BF912FB7D31A2" blockId="4.[151,1437,152,2015]" pageId="4" pageNumber="356">
<bibRefCitation id="EFB601C8CB6E2C7AFF2BF912FEA431FE" author="Silvestri, F." box="[199,375,1701,1727]" pageId="4" pageNumber="356" pagination="303 - 310" refId="ref3975" refString="Silvestri, F. (1913) Illustrazione di due famiglie di Chordeumoidea (Diplopoda) del Nord America. Bollettino del Laboratorio di zoologia generale e agraria della Facolta agraria in Portici, 7, 303 - 310. Available from: https: // biodiversitylibrary. org / page / 13979142 (Accessed 16 Aug. 2019)" type="journal article" year="1913">Silvestri (1913)</bibRefCitation>
and
<bibRefCitation id="EFB601C8CB6E2C7AFE42F912FDE931FE" author="Shear, W. A." box="[430,570,1701,1727]" pageId="4" pageNumber="356" pagination="151 - 352" refId="ref3794" refString="Shear, W. A. (1972) Studies in the milliped order Chordeumida (Diplopoda). A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. - Bulletin of the Museum of Comparative Zoology, 144, 151 - 352. Available from: https: // biodiversitylibrary. org / page / 4227660 (Accessed 16 Aug. 2019)" type="journal article" year="1972">Shear (1972</bibRefCitation>
, fig. 447) show a single flagellocoxite, probably sheathed by the anterior angiocoxite process. While present, the flagellocoxite was not visible in our SEM preparations.
</paragraph>
<paragraph id="8B987C39CB6E2C79FF2BF95AFCAF365E" blockId="4.[151,1437,152,2015]" lastBlockId="7.[151,1437,153,287]" lastPageId="7" lastPageNumber="359" pageId="4" pageNumber="356">
From label data, it appears that
<taxonomicName id="4C2707BACB6E2C7AFDD1F959FD123046" authorityName="Silvestri" authorityYear="1909" box="[573,705,1773,1799]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="356" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB6E2C7AFDD1F959FD123046" box="[573,705,1773,1799]" italics="true" pageId="4" pageNumber="356">U. bumpusi</emphasis>
</taxonomicName>
is most often associated with deciduous leaf litter, particularly from maple. Elevations of collecting sites range from 150 to 4800 (
<quantity id="4CDFD1DCCB6E2C7AFC45F8A7FC28306A" box="[937,1019,1808,1835]" metricMagnitude="1" metricUnit="m" metricValue="4.57" pageId="4" pageNumber="356" unit="m" value="45.7">45.7 m</quantity>
to
<quantity id="4CDFD1DCCB6E2C7AFBCEF8A6FBA9306A" box="[1058,1146,1809,1835]" metricMagnitude="3" metricUnit="m" metricValue="1.463" pageId="4" pageNumber="356" unit="m" value="1463.0">1463 m</quantity>
) asl, a broad range given the northerly latitude. The species is not common; collections mostly consist of single specimens or no more than three specimens, while collections of other chordeumatidans from the same localities may include 10
<specimenCount id="9D21B7B0CB6E2C7AFADFF8EEFF0530D6" pageId="4" pageNumber="356" type="generic">20 specimens</specimenCount>
. The Whatcom County collection is a considerable extension, being some
<quantity id="4CDFD1DCCB6E2C7AFBE7F8CAFBA430D6" box="[1035,1143,1917,1943]" metricMagnitude="5" metricUnit="m" metricValue="1.86683904" pageId="4" pageNumber="356" unit="mi" value="116.0">116 miles</quantity>
(
<quantity id="4CDFD1DCCB6E2C7AFB69F8CAFB0F30D9" box="[1157,1244,1917,1944]" metricMagnitude="5" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="356" unit="km" value="187.0">187 km</quantity>
) north-northwest of Snoqualmie Pass, the previous most northerly locality, and suggests that
<taxonomicName id="4C2707BACB6E2C7AFBE6F815FB5C30FA" authorityName="Silvestri" authorityYear="1909" box="[1034,1167,1953,1979]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="4" pageNumber="356" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB6E2C7AFBE6F815FB5C30FA" box="[1034,1167,1953,1979]" italics="true" pageId="4" pageNumber="356">U. bumpusi</emphasis>
</taxonomicName>
will be found in intervening locations and perhaps northward into British
<collectingCountry id="F3303CA9CB6E2C7AFD1AF872FCB4309E" box="[758,871,1989,2015]" name="Colombia" pageId="4" pageNumber="356">Colombia</collectingCountry>
,
<collectingCountry id="F3303CA9CB6E2C7AFC99F872FC1E309E" box="[885,973,1989,2015]" name="Canada" pageId="4" pageNumber="356">Canada</collectingCountry>
. Longmire Springs, the
<typeStatus id="549CC29BCB6E2C7AFB08F872FAC7309E" box="[1252,1300,1989,2015]" pageId="4" pageNumber="356">type</typeStatus>
locality for
<taxonomicName id="4C2707BACB6D2C79FF7BFF2DFEC437F2" authorityName="Silvestri" authorityYear="1909" box="[151,279,153,179]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="359" phylum="Arthropoda" rank="species" species="bumpusi">
<emphasis id="B953A02BCB6D2C79FF7BFF2DFEC437F2" box="[151,279,153,179]" italics="true" pageId="7" pageNumber="359">U. bumpusi</emphasis>
</taxonomicName>
, was a long-popular hot springs resort not far from Tacoma, Washington, at the time of Silvestris visit. It was later incorporated into Mount Rainier National Park and is now an entry station into the park. Snoqualmie Pass,
<typeStatus id="549CC29BCB6D2C79FF7BFF56FF1437BA" box="[151,199,225,251]" pageId="7" pageNumber="359">type</typeStatus>
locality for
<emphasis id="B953A02BCB6D2C79FEA1FF55FE1637BA" box="[333,453,226,251]" italics="true" pageId="7" pageNumber="359">
<taxonomicName id="4C2707BACB6D2C79FEA1FF55FE1237BA" authorityName="Chamberlin" authorityYear="1941" box="[333,449,226,251]" class="Diplopoda" family="Urochordeumatidae" genus="Urochordeuma" kingdom="Animalia" order="Chordeumatida" pageId="7" pageNumber="359" phylum="Arthropoda" rank="species" species="porona">U. porona</taxonomicName>
,
</emphasis>
is a large pass through the Cascade Mountains where Interstate 90 uses this geographical gap as the only four-lane east-west route across Washington State.
</paragraph>
</subSubSection>
</treatment>
</document>