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<document id="A817042B9FDF3DB67E7E448B5157FF14" ID-DOI="10.11646/zootaxa.2279.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5307538" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629653572613" checkinUser="felipe" docAuthor="Brown, John W." docDate="2009" docId="038E7928FFD0FF8DFDABE135FAEFFEB1" docLanguage="en" docName="zt02279p050.pdf" docOrigin="Zootaxa 2279 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2279.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Megalota spinulosa Brown 2009, new species" docType="treatment" docVersion="3" lastPageNumber="19" masterDocId="FFB70150FFC0FF9FFFBCE420FFFBFFB3" masterDocTitle="The discovery of Megalota in the Neotropics, with a revision of the New World species (Lepidoptera: Tortricidae: Olethreutini)" masterLastPageNumber="50" masterPageNumber="1" pageNumber="17" updateTime="1699089488996" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="10E1ECDA012C28F575616FBF7590404C">The discovery of Megalota in the Neotropics, with a revision of the New World species (Lepidoptera: Tortricidae: Olethreutini)</mods:title>
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<mods:namePart id="2A7D9D9616DB15AFAE617D5B89AB4685">Brown, John W.</mods:namePart>
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<mods:date id="7B0BF2C9E206EE0F85EB900E6AED28CC">2009</mods:date>
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<mods:identifier id="AFDE79A502C9253652BE9D4D87C559C7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="038E7928FFD0FF8DFDABE135FAEFFEB1" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1" httpUri="http://treatment.plazi.org/id/038E7928FFD0FF8DFDABE135FAEFFEB1" lastPageId="18" lastPageNumber="19" pageId="16" pageNumber="17">
<subSubSection id="C33D9BB5FFD0FF8FFDABE135FC96FAE1" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B98C83EFFD0FF8FFDABE135FC96FAE1" blockId="16.[535,1051,1301,1362]" pageId="16" pageNumber="17">
<emphasis id="B953142CFFD0FF8FFDABE135FBE0FA9C" bold="true" box="[535,1051,1301,1327]" pageId="16" pageNumber="17">
9.
<taxonomicName id="4C27B3BDFFD0FF8FFD88E135FC85FA9C" ID-CoL="B3WZG" authority="Brown" authorityName="Brown" authorityYear="2009" box="[564,894,1301,1327]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="spinulosa" status="sp. nov.">
<emphasis id="B953142CFFD0FF8FFD88E135FCDAFA9C" bold="true" box="[564,801,1301,1327]" italics="true" pageId="16" pageNumber="17">Megalota spinulosa</emphasis>
Brown
</taxonomicName>
,
<taxonomicNameLabel id="A260A957FFD0FF8FFC37E135FBE0FA9C" box="[907,1051,1301,1327]" pageId="16" pageNumber="17" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation id="131CD4BBFFD0FF8FFD79E118FCE3FAE1" box="[709,792,1336,1362]" captionStart="FIGURES 112" captionStartId="36.[151,269,1661,1685]" captionTargetBox="[199,1384,202,1634]" captionTargetId="figure-73@36.[195,1392,194,1637]" captionTargetPageId="36" captionText="FIGURES 112. Adults of Megalota. 1. M. submicans (holotype), 2. M. synchysis (paratype), 3. M. peruviana (paratype), 4. M. aquilonaris (holotype), 5. M. vulgaris (paratype), 6. M. cacaulana (holotype), 7. M. macrosocia (holotype), 8. M. ochreoapex (holotype), 9. M. spinulosa (holotype), 10. M. simpliciana (paratype), 11. M. delphinosema (Costa Rica), 12. M. jamaicana (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5307542" httpUri="https://zenodo.org/record/5307542/files/figure.png" pageId="16" pageNumber="17">Figs. 9</figureCitation>
,
<figureCitation id="131CD4BBFFD0FF8FFC98E118FCBFFAE1" box="[804,836,1336,1362]" captionStart="FIGURES 3133" captionStartId="40.[151,269,1976,2000]" captionTargetBox="[192,1397,178,1968]" captionTargetId="figure-56@40.[192,1398,138,2000]" captionTargetPageId="40" captionText="FIGURES 3133. Male genitalia of Megalota; valva spread, phallus removed. 31. M. macrosocia, image of USNM slide 124,139; 32. M. ochreoapex, image of USNM slide 85,863; 33. M. spinulosa, image of USNM slide 84,895." figureDoi="http://doi.org/10.5281/zenodo.5307552" httpUri="https://zenodo.org/record/5307552/files/figure.png" pageId="16" pageNumber="17">33</figureCitation>
,
<figureCitation id="131CD4BBFFD0FF8FFCECE118FC96FAE1" box="[848,877,1336,1362]" captionStart="FIGURES 5255" captionStartId="47.[151,269,1939,1963]" captionTargetBox="[231,1365,184,1931]" captionTargetId="figure-57@47.[231,1365,184,1936]" captionTargetPageId="47" captionText="FIGURES 5255. Female genitalia of Megalota. 52. M. aquilonaris, image of USNM slide 124,109; 53. M. vulgaris, image of USNM slide 95,323; 54. M. ochreoapex, image of USNM slide 85,864; 55. M. spinulosa, image of USNM slide 84,874." figureDoi="http://doi.org/10.5281/zenodo.5307564" httpUri="https://zenodo.org/record/5307564/files/figure.png" pageId="16" pageNumber="17">55</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFD0FF8FFF2BE1A2FEEEF907" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B98C83EFFD0FF8FFF2BE1A2FEEEF907" blockId="16.[151,1436,1410,2036]" pageId="16" pageNumber="17">
<emphasis id="B953142CFFD0FF8FFF2BE1A2FEF4FA2F" bold="true" box="[151,271,1410,1436]" pageId="16" pageNumber="17">Diagnosis</emphasis>
. Adults of
<taxonomicName id="4C27B3BDFFD0FF8FFE28E1A3FDD0FA2F" authorityName="Brown" authorityYear="2009" box="[404,555,1411,1436]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="spinulosa">
<emphasis id="B953142CFFD0FF8FFE28E1A3FDD0FA2F" box="[404,555,1411,1436]" italics="true" pageId="16" pageNumber="17">M. spinulosa</emphasis>
</taxonomicName>
are superficially similar to most congeners, but the forewing maculation is slightly darker. Males are easily distinguished from the similar and sympatric
<taxonomicName id="4C27B3BDFFD0FF8FFB91E18BFB4FFA77" authorityName="Brown" authorityYear="2009" box="[1069,1204,1451,1476]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="vulgaris">
<emphasis id="B953142CFFD0FF8FFB91E18BFB4FFA77" box="[1069,1204,1451,1476]" italics="true" pageId="16" pageNumber="17">M. vulgaris</emphasis>
</taxonomicName>
by the distinct anal roll of the hindwing, which is poorly developed or absent in the latter. The male genitalia are easily distinguished by the comparatively small uncus; the relatively straight basal process of the valva; the conspicuous, narrow, flangelike projection from the valva about 0.65 the distance from the base; and the absence of spiniform setae from the basal 0.5 of the valva. The female genitalia are characterized by a rugose bandlike sterigma with a trapezoidal mesal lobe and a sclerotized colliculum that occupies the posterior 0.6 of the ductus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFD0FF8EFF7AE2E2FAD1FD21" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="description">
<paragraph id="8B98C83EFFD0FF8EFF7AE2E2FAD1FD21" blockId="16.[151,1436,1410,2036]" lastBlockId="17.[151,1436,152,2018]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B953142CFFD0FF8FFF7AE2E2FEACF96F" bold="true" box="[198,343,1730,1756]" pageId="16" pageNumber="17">Description</emphasis>
.
<emphasis id="B953142CFFD0FF8FFED7E2E3FE51F96F" box="[363,426,1731,1756]" italics="true" pageId="16" pageNumber="17">Head</emphasis>
: Vertex mostly brown with some tawny brown, frons tawny brown; labial palpus tawny brown to creamy white, with small areas of brown and red brown.
<emphasis id="B953142CFFD0FF8FFBBEE2CBFBADF8B7" box="[1026,1110,1771,1796]" italics="true" pageId="16" pageNumber="17">Thorax</emphasis>
: Dorsum brown and tawny brown; metascutum with tuft of bronze brown scales. Hind tibia in male with dense patch of white sex scales concealing hairpencil. Forewing length 6.07.0 mm (mean = 6.5); ground color reticulated brown and tawny brown, with lesser amounts of pinkish ocherous and red brown; ill-defined region of gray and creamy white scales separating basal blotch from median fascia; median fascia ill-defined, dark brown at costa, fading to olive brown at dorsum; oblique blotch arched from mid-termen, intersecting median fascia near distal end of discal cell, bordered by silvery gray. Fringe brown. Hindwing dark brown, anal margin in male with well- developed roll bearing sex scales.
<emphasis id="B953142CFFD1FF8EFD94E4B8FD6CFF02" box="[552,663,152,177]" italics="true" pageId="17" pageNumber="18">Abdomen</emphasis>
: Shiny bronze gray. Male genitalia (
<figureCitation id="131CD4BBFFD1FF8EFBFDE4B8FB63FF01" box="[1089,1176,152,178]" captionStart="FIGURES 3133" captionStartId="40.[151,269,1976,2000]" captionTargetBox="[192,1397,178,1968]" captionTargetId="figure-56@40.[192,1398,138,2000]" captionTargetPageId="40" captionText="FIGURES 3133. Male genitalia of Megalota; valva spread, phallus removed. 31. M. macrosocia, image of USNM slide 124,139; 32. M. ochreoapex, image of USNM slide 85,863; 33. M. spinulosa, image of USNM slide 84,895." figureDoi="http://doi.org/10.5281/zenodo.5307552" httpUri="https://zenodo.org/record/5307552/files/figure.png" pageId="17" pageNumber="18">Fig. 33</figureCitation>
;
<figureCitation id="131CD4BBFFD1FF8EFB19E4B8FB4FFF01" box="[1189,1204,152,178]" captionStart="FIGURES 112" captionStartId="36.[151,269,1661,1685]" captionTargetBox="[199,1384,202,1634]" captionTargetId="figure-73@36.[195,1392,194,1637]" captionTargetPageId="36" captionText="FIGURES 112. Adults of Megalota. 1. M. submicans (holotype), 2. M. synchysis (paratype), 3. M. peruviana (paratype), 4. M. aquilonaris (holotype), 5. M. vulgaris (paratype), 6. M. cacaulana (holotype), 7. M. macrosocia (holotype), 8. M. ochreoapex (holotype), 9. M. spinulosa (holotype), 10. M. simpliciana (paratype), 11. M. delphinosema (Costa Rica), 12. M. jamaicana (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5307542" httpUri="https://zenodo.org/record/5307542/files/figure.png" pageId="17" pageNumber="18">6</figureCitation>
preparations examined) with tegumen subrectangular, slightly narrowed dorsally, with weak triangular expansion immediately before junction with uncus; uncus somewhat rounded, comparatively short, only slightly wider than tegumen; socius membranous, inconspicuous; valva parallel-sided, slightly more narrow in distal 0.33, with slender, attenuate flange ca. 0.65 distance from base to apex, with patch of elongate setae from venter of valva just basad of flange; basal process of valva rather broad, straight, as in
<taxonomicName id="4C27B3BDFFD1FF8EFC0DE540FBC3FECA" authorityName="Brown" authorityYear="2009" box="[945,1080,352,377]" class="Insecta" family="Tortricidae" genus="Megalota" kingdom="Animalia" order="Lepidoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="ochreoapex">
<emphasis id="B953142CFFD1FF8EFC0DE540FBC3FECA" box="[945,1080,352,377]" italics="true" pageId="17" pageNumber="18">ochreoapex</emphasis>
</taxonomicName>
, without elbow, attenuate and spined in distal 0.2. Phallus rather short, curved, with a short external subdorsal thorn subapically; vesica with two cornuti, one short and stout, the other twice as long, slender. Female genitalia (
<figureCitation id="131CD4BBFFD1FF8EFB36E590FB18FE79" box="[1162,1251,432,458]" captionStart="FIGURES 5255" captionStartId="47.[151,269,1939,1963]" captionTargetBox="[231,1365,184,1931]" captionTargetId="figure-57@47.[231,1365,184,1936]" captionTargetPageId="47" captionText="FIGURES 5255. Female genitalia of Megalota. 52. M. aquilonaris, image of USNM slide 124,109; 53. M. vulgaris, image of USNM slide 95,323; 54. M. ochreoapex, image of USNM slide 85,864; 55. M. spinulosa, image of USNM slide 84,874." figureDoi="http://doi.org/10.5281/zenodo.5307564" httpUri="https://zenodo.org/record/5307564/files/figure.png" pageId="17" pageNumber="18">Fig. 55</figureCitation>
;
<figureCitation id="131CD4BBFFD1FF8EFB4FE590FAFAFE79" box="[1267,1281,432,458]" captionStart="FIGURES 112" captionStartId="36.[151,269,1661,1685]" captionTargetBox="[199,1384,202,1634]" captionTargetId="figure-73@36.[195,1392,194,1637]" captionTargetPageId="36" captionText="FIGURES 112. Adults of Megalota. 1. M. submicans (holotype), 2. M. synchysis (paratype), 3. M. peruviana (paratype), 4. M. aquilonaris (holotype), 5. M. vulgaris (paratype), 6. M. cacaulana (holotype), 7. M. macrosocia (holotype), 8. M. ochreoapex (holotype), 9. M. spinulosa (holotype), 10. M. simpliciana (paratype), 11. M. delphinosema (Costa Rica), 12. M. jamaicana (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5307542" httpUri="https://zenodo.org/record/5307542/files/figure.png" pageId="17" pageNumber="18">4</figureCitation>
preparations examined) with papillae anales relatively broad; sterigma a parallel-sided, spinulose band, divided mesally by invaginated, subrectangular lobe with blunt posterior end; colliculum lightly sclerotized, occupying posterior ca. 0.6 of ductus bursae, with pair of long, strongly sclerotized internal struts along edge of ductus bursae, one ca. 0.9 length of colliculum, the other ca. 0.7 length of colliculum; remainder of ductus bursae membranous; corpus bursae rounded-oblong; signum a subrectangular, finlike plate from a small, ill-defined base.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFD1FF8EFF7AE6BFFD15F821" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8B98C83EFFD1FF8EFF7AE6BFFD44FD51" blockId="17.[151,1436,152,2018]" pageId="17" pageNumber="18">
<materialsCitation id="3B4FC263FFD1FF8EFF7AE6BFFD44FD51" collectingDate="2003-02-09" collectionCode="INBio" collectorName="D. Wagner" country="Costa Rica" latitude="10.433333" location="La Selva Biological Station" longLatPrecision="1298" longitude="-84.01667" municipality="Male" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Heredia" typeStatus="holotype">
<emphasis id="B953142CFFD1FF8EFF7AE6BFFEC3FD0A" bold="true" box="[198,312,671,697]" pageId="17" pageNumber="18">
<typeStatus id="549C769CFFD1FF8EFF7AE6BFFEC3FD0A" box="[198,312,671,697]" pageId="17" pageNumber="18" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectingMunicipality id="6BFC5244FFD1FF8EFEF5E680FE73FD09" box="[329,392,672,698]" pageId="17" pageNumber="18">Male</collectingMunicipality>
,
<collectingCountry id="F33088AEFFD1FF8EFE2BE680FDE7FD09" box="[407,540,672,698]" name="Costa Rica" pageId="17" pageNumber="18">Costa Rica</collectingCountry>
,
<collectingRegion id="49E306DCFFD1FF8EFD97E680FD71FD09" box="[555,650,672,698]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="18">Heredia</collectingRegion>
,
<location id="8EF89EE5FFD1FF8EFD25E680FC1DFD09" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFD25E680FC1DFD09" box="[665,998,672,698]" country="Costa Rica" latitude="10.433333" longLatPrecision="1298" longitude="-84.01667" municipality="Male" name="La Selva Biological Station" pageId="17" pageNumber="18" stateProvince="Heredia">La Selva Biological Station</location>
,
<geoCoordinate id="EE13AEF9FFD1FF8EFC48E680FBA8FD09" box="[1012,1107,671,698]" degrees="10" direction="north" minutes="26" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="10.433333">10°26N</geoCoordinate>
,
<geoCoordinate id="EE13AEF9FFD1FF8EFBDEE680FB3FFD09" box="[1122,1220,671,698]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<date id="FF99EEFEFFD1FF8EFB6EE680FAA5FD09" box="[1234,1374,672,698]" pageId="17" pageNumber="18" value="2003-02-09">
<collectingDate id="EFDD1716FFD1FF8EFB6EE680FAA5FD09" box="[1234,1374,672,698]" pageId="17" pageNumber="18" value="2003-02-09">9 Feb 2003</collectingDate>
</date>
, MV light,
<collectorName id="26D2ADE8FFD1FF8EFF67E6E8FEA2FD51" box="[219,345,712,738]" pageId="17" pageNumber="18">D. Wagner</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFED5E6E8FE43FD51" box="[361,440,712,738]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
), USNM slide 95,334.
</materialsCitation>
</paragraph>
<paragraph id="8B98C83EFFD1FF8EFF7AE6CFFD15F821" blockId="17.[151,1436,152,2018]" pageId="17" pageNumber="18">
<materialsCitation id="3B4FC263FFD1FF8EFF7AE6CFFC5CFCB9" box="[198,935,751,778]" country="Costa Rica" location="Turrialba" pageId="17" pageNumber="18" specimenCount="162" specimenCount-female="15" specimenCount-male="147" stateProvince="Cartago" typeStatus="paratype">
<emphasis id="B953142CFFD1FF8EFF7AE6CFFEB8FCBA" bold="true" box="[198,323,751,777]" pageId="17" pageNumber="18">
<typeStatus id="549C769CFFD1FF8EFF7AE6CFFEB8FCBA" box="[198,323,751,777]" pageId="17" pageNumber="18" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D2103B7FFD1FF8EFEEBE6D0FE63FCBB" box="[343,408,752,778]" pageId="17" pageNumber="18" type="male">147♂</specimenCount>
,
<specimenCount id="9D2103B7FFD1FF8EFE17E6D0FE25FCBA" box="[427,478,751,778]" pageId="17" pageNumber="18" type="female">15♀</specimenCount>
).
<collectingCountry id="F33088AEFFD1FF8EFE47E6D0FD55FCB9" box="[507,686,752,778]" name="Costa Rica" pageId="17" pageNumber="18">COSTA RICA</collectingCountry>
:
<emphasis id="B953142CFFD1FF8EFD01E6CFFCD6FCBA" bold="true" box="[701,813,751,777]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFD01E6CFFCD3FCBA" box="[701,808,751,777]" country="Costa Rica" name="Cartago" pageId="17" pageNumber="18">Cartago</collectingRegion>
:
</emphasis>
<location id="8EF89EE5FFD1FF8EFC84E6D0FC5CFCB9" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFC84E6D0FC5CFCB9" box="[824,935,752,778]" country="Costa Rica" name="Turrialba" pageId="17" pageNumber="18" stateProvince="Cartago">Turrialba</location>
</materialsCitation>
,
<date id="FF99EEFEFFD1FF8EFC0BE6D0FB7AFCB9" box="[951,1153,752,778]" pageId="17" pageNumber="18" value="1965-03-13" valueMax="1965-03-17" valueMin="1965-03-13">1317 Mar 1965</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB2BE6D0FB42FCBB" box="[1175,1209,752,778]" pageId="17" pageNumber="18" type="male">8♂</specimenCount>
),
<materialsCitation id="3B4FC263FFD1FF8EFB6AE6D0FD80FC81" collectingDate="1965-02-17" collectingDateMax="1965-02-21" collectingDateMin="1965-02-17" collectionCode="USNM" collectorName="Duckworth" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="4" specimenCount-male="4" stateProvince="Cartago" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFB6AE6D0FA60FCB9" box="[1238,1435,752,778]" pageId="17" pageNumber="18" value="1965-02-17" valueMax="1965-02-21" valueMin="1965-02-17">
<collectingDate id="EFDD1716FFD1FF8EFB6AE6D0FA60FCB9" box="[1238,1435,752,778]" pageId="17" pageNumber="18" value="1965-02-17" valueMax="1965-02-21" valueMin="1965-02-17">1721 Feb 1965</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFF23E738FF38FC83" box="[159,195,792,818]" pageId="17" pageNumber="18" type="male">4♂</specimenCount>
), S. S. &amp; W. D.
<collectorName id="26D2ADE8FFD1FF8EFE38E738FDF3FC81" box="[388,520,792,818]" pageId="17" pageNumber="18">Duckworth</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFDA4E738FD88FC81" box="[536,627,792,818]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFD3BE738FF11FCE9" collectingDate="1985-05-15" collectionCode="EME" collectorName="Chemsak" country="Costa Rica" elevation="600" location="3 km SE Turrialba" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="paratype">
CATIE,
<location id="8EF89EE5FFD1FF8EFD55E738FC3EFC81" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFD55E738FC3EFC81" box="[745,965,792,818]" country="Costa Rica" name="3 km SE Turrialba" pageId="17" pageNumber="18" stateProvince="Cartago">
<quantity id="4CDF65DBFFD1FF8EFD55E738FCDFFC81" box="[745,804,792,818]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="18" unit="km" value="3.0">3 km</quantity>
SE Turrialba
</location>
,
<quantity id="4CDF65DBFFD1FF8EFC6CE738FBE6FC81" box="[976,1053,792,818]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="18" unit="m" value="600.0">
<elevation id="000A2F0DFFD1FF8EFC6CE738FBE6FC81" box="[976,1053,792,818]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="18" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<date id="FF99EEFEFFD1FF8EFB96E738FB3DFC81" box="[1066,1222,792,818]" pageId="17" pageNumber="18" value="1985-05-15">
<collectingDate id="EFDD1716FFD1FF8EFB96E738FB3DFC81" box="[1066,1222,792,818]" pageId="17" pageNumber="18" value="1985-05-15">15 May 1985</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB65E738FB07FC82" box="[1241,1276,791,818]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
), J.
<collectorName id="26D2ADE8FFD1FF8EFA93E738FA67FC81" box="[1327,1436,792,818]" pageId="17" pageNumber="18">Chemsak</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFF23E760FF19FCE9" box="[159,226,832,858]" pageId="17" pageNumber="18">EME</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFD1FF8EFF4BE760FC9DFCE9" box="[247,870,831,858]" collectingDate="1985-05-29" collectingDateMax="1985-05-30" collectingDateMin="1985-05-29" collectionCode="EME" collectorName="Powell &amp; Doyen" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Cartago" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFF4BE760FE3BFCE9" box="[247,448,832,858]" pageId="17" pageNumber="18" value="1985-05-29" valueMax="1985-05-30" valueMin="1985-05-29">
<collectingDate id="EFDD1716FFD1FF8EFF4BE760FE3BFCE9" box="[247,448,832,858]" pageId="17" pageNumber="18" value="1985-05-29" valueMax="1985-05-30" valueMin="1985-05-29">2930 May 1985</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE6FE760FE0DFCEA" box="[467,502,831,858]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
), J.
<collectorName id="26D2ADE8FFD1FF8EFD95E760FD81FCE9" box="[553,634,832,858]" pageId="17" pageNumber="18">Powell</collectorName>
&amp; J.
<collectorName id="26D2ADE8FFD1FF8EFD07E760FCF2FCE9" box="[699,777,832,858]" pageId="17" pageNumber="18">Doyen</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFCA5E760FCA6FCE9" box="[793,861,832,858]" pageId="17" pageNumber="18">EME</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFCCDE71FFF1FFC31" country="Costa Rica" elevation="700" location="Est. Pitilla" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Guanacaste" typeStatus="paratype">
<emphasis id="B953142CFFD1FF8EFCCDE71FFBF8FCEA" bold="true" box="[881,1027,831,857]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFCCDE71FFBF8FCEA" box="[881,1027,831,857]" country="Costa Rica" name="Guanacaste" pageId="17" pageNumber="18">Guanacaste</collectingRegion>
</emphasis>
:
<location id="8EF89EE5FFD1FF8EFBA8E760FB6AFCE9" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFBA8E760FB6AFCE9" box="[1044,1169,832,858]" country="Costa Rica" name="Est. Pitilla" pageId="17" pageNumber="18" stateProvince="Guanacaste">Est. Pitilla</location>
,
<locationDeviation id="9A2AD11CFFD1FF8EFB21E760FA63FCE9" box="[1181,1432,832,858]" location="Santa Cecilia" pageId="17" pageNumber="18">
<quantity id="4CDF65DBFFD1FF8EFB21E760FB22FCE9" box="[1181,1241,832,858]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="17" pageNumber="18" unit="km" value="9.0">9 km</quantity>
S Santa Cecilia
</locationDeviation>
,
<quantity id="4CDF65DBFFD1FF8EFF2BE748FF1FFC31" box="[151,228,872,898]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="m" value="700.0">
<elevation id="000A2F0DFFD1FF8EFF2BE748FF1FFC31" box="[151,228,872,898]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="17" pageNumber="18" unit="m" value="700.0">700 m</elevation>
</quantity>
</materialsCitation>
,
<date id="FF99EEFEFFD1FF8EFF4EE748FE56FC31" box="[242,429,872,898]" pageId="17" pageNumber="18" value="1993-01-12" valueMax="1993-01-30" valueMin="1993-01-12">1230 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE7EE748FE19FC33" box="[450,482,872,898]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), C. Moraga,
<materialsCitation id="3B4FC263FFD1FF8EFD36E748FBB7FC31" box="[650,1100,872,898]" collectingDate="1993-08-06" collectingDateMax="1993-08-18" collectingDateMin="1993-08-06" collectionCode="INBio" collectorName="Rios" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Guanacaste" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFD36E748FCB9FC31" box="[650,834,872,898]" pageId="17" pageNumber="18" value="1993-08-06" valueMax="1993-08-18" valueMin="1993-08-06">
<collectingDate id="EFDD1716FFD1FF8EFD36E748FCB9FC31" box="[650,834,872,898]" pageId="17" pageNumber="18" value="1993-08-06" valueMax="1993-08-18" valueMin="1993-08-06">618 Aug 1993</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFCEBE748FC83FC33" box="[855,888,872,898]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), P.
<collectorName id="26D2ADE8FFD1FF8EFC12E748FC1FFC31" box="[942,996,872,898]" pageId="17" pageNumber="18">Rios</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFC48E748FBB8FC31" box="[1012,1091,872,898]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFBE4E747FDC5FC19" country="Costa Rica" elevation="100" latitude="10.416667" location="Est. Biologica La Selva" longLatPrecision="1298" longitude="-84.01667" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Heredia" typeStatus="paratype">
<emphasis id="B953142CFFD1FF8EFBE4E747FB47FC32" bold="true" box="[1112,1212,871,897]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFBE4E747FB47FC32" box="[1112,1212,871,897]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="18">Heredia</collectingRegion>
</emphasis>
:
<location id="8EF89EE5FFD1FF8EFB71E748FF20FC19" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFB71E748FF20FC19" country="Costa Rica" latitude="10.416667" longLatPrecision="1298" longitude="-84.01667" name="Est. Biologica La Selva" pageId="17" pageNumber="18" stateProvince="Heredia">Est. Biologica La Selva</location>
,
<geoCoordinate id="EE13AEF9FFD1FF8EFF54E7B0FEBDFC19" box="[232,326,911,938]" degrees="10" direction="north" minutes="25" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="10.416667">10°25N</geoCoordinate>
,
<geoCoordinate id="EE13AEF9FFD1FF8EFEE8E7B0FE4CFC19" box="[340,439,911,938]" degrees="84" direction="west" minutes="01" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-84.01667">84°01W</geoCoordinate>
,
<quantity id="4CDF65DBFFD1FF8EFE78E7B0FDC5FC19" box="[452,574,912,938]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="17" pageNumber="18" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">
<elevation id="000A2F0DFFD1FF8EFE78E7B0FDC5FC19" box="[452,574,912,938]" metricMagnitude="2" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="17" pageNumber="18" unit="m" value="100.0" valueMax="150.0" valueMin="50.0">50150 m</elevation>
</quantity>
</materialsCitation>
,
<date id="FF99EEFEFFD1FF8EFDF0E7B0FD34FC19" box="[588,719,912,938]" pageId="17" pageNumber="18" value="1993-01-09">9 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD5DE7B0FCFFFC1B" box="[737,772,912,938]" pageId="17" pageNumber="18" type="male">3♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFCA2E7B0FC54FC19" box="[798,943,912,938]" pageId="17" pageNumber="18" value="1993-01-10">10 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC7EE7B0FC1EFC1B" box="[962,997,912,938]" pageId="17" pageNumber="18" type="male">3♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFC43E7B0FB47FC19" box="[1023,1212,912,938]" pageId="17" pageNumber="18" value="1993-01-10" valueMax="1993-01-17" valueMin="1993-01-10">1017 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB6DE7B0FB0AFC1B" box="[1233,1265,912,938]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFAB0E7B0FA60FC19" box="[1292,1435,912,938]" pageId="17" pageNumber="18" value="1993-01-11">11 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFF1EE798FF38FC63" box="[162,195,952,978]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFF61E798FE95FC61" box="[221,366,952,978]" pageId="17" pageNumber="18" value="1993-01-12">12 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE3CE798FE5FFC63" box="[384,420,952,978]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
,
<specimenCount id="9D2103B7FFD1FF8EFE08E798FE22FC62" box="[436,473,951,978]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFE4FE798FD78FC61" box="[499,643,952,978]" pageId="17" pageNumber="18" value="1993-01-13">13 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD2AE798FD42FC63" box="[662,697,952,978]" pageId="17" pageNumber="18" type="male">3♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFD6FE798FC9FFC61" box="[723,868,952,978]" pageId="17" pageNumber="18" value="1993-01-14">14 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFCCBE798FC61FC63" box="[887,922,952,978]" pageId="17" pageNumber="18" type="male">3♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFC08E798FBBFFC61" box="[948,1092,952,978]" pageId="17" pageNumber="18" value="1993-01-15">15 Jan 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFBE5E798FB81FC63" box="[1113,1146,952,978]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFB28E798FAD5FC61" box="[1172,1326,952,978]" pageId="17" pageNumber="18" value="1993-03-20">20 Mar 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFAFDE798FA9FFC63" box="[1345,1380,952,978]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFAC3E798FEF3FC49" pageId="17" pageNumber="18" value="1993-03-30">30 Mar 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFEA7E7C0FEC7FC4B" box="[283,316,992,1018]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFEE8E7C0FE1DFC49" box="[340,486,992,1018]" pageId="17" pageNumber="18" value="1993-04-13">13 Apr 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE45E7C0FDE2FC4B" box="[505,537,992,1018]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFD8DE7C0FD3AFC49" box="[561,705,992,1018]" pageId="17" pageNumber="18" value="1993-09-11">11 Sep 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD6FE7C0FD0FFC4B" box="[723,756,992,1018]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFCB0E7C0FC60FC49" box="[780,923,992,1018]" pageId="17" pageNumber="18" value="1993-10-12">12 Oct 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC12E7C0FC34FC4B" box="[942,975,992,1018]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFC5AE7C0FB86FC49" box="[998,1149,992,1018]" pageId="17" pageNumber="18" value="1993-11-11">11 Nov 1993</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB33E7C0FB4BFC4B" box="[1167,1200,992,1018]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFB74E7C0FAAEFC49" box="[1224,1365,992,1018]" pageId="17" pageNumber="18" value="1994-06-27">27 Jun 1994</date>
(
<specimenCount id="9D2103B7FFD1FF8EFAD4E7C0FA70FC4A" box="[1384,1419,991,1018]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFF2BE028FEDDFB91" box="[151,294,1032,1058]" pageId="17" pageNumber="18" value="1994-06-28">28 Jun 1994</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE85E028FEA1FB93" box="[313,346,1032,1058]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFECFE028FE0DFB91" box="[371,502,1032,1058]" pageId="17" pageNumber="18" value="1996-02-05">5 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFDB5E028FDD7FB92" box="[521,556,1031,1058]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFDF9E028FD33FB91" box="[581,712,1032,1058]" pageId="17" pageNumber="18" value="1996-02-06">6 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD60E028FD06FB93" box="[732,765,1032,1058]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFCA9E028FC63FB91" box="[789,920,1032,1058]" pageId="17" pageNumber="18" value="1996-02-07">7 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC10E028FC36FB93" box="[940,973,1032,1058]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFC59E028FB8CFB91" box="[997,1143,1032,1058]" pageId="17" pageNumber="18" value="1996-02-10">10 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB37E028FB57FB93" box="[1163,1196,1032,1058]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFB78E028FAADFB91" box="[1220,1366,1032,1058]" pageId="17" pageNumber="18" value="1996-02-12">12 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFAD6E028FA70FB93" box="[1386,1419,1032,1058]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFF2BE010FED0FBF9" box="[151,299,1072,1098]" pageId="17" pageNumber="18" value="1996-02-15">15 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE83E010FE9BFBFB" box="[319,352,1072,1098]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
,
<specimenCount id="9D2103B7FFD1FF8EFED3E010FE6FFBFA" box="[367,404,1071,1098]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFE11E010FDBAFBF9" box="[429,577,1072,1098]" pageId="17" pageNumber="18" value="1996-02-19">19 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFDE9E010FD8DFBFB" box="[597,630,1072,1098]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFD2CE010FCDFFBF9" box="[656,804,1072,1098]" pageId="17" pageNumber="18" value="1996-02-22">22 Feb 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC84E010FCA2FBFB" box="[824,857,1072,1098]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFCCEE010FBF7FBF9" box="[882,1036,1072,1098]" pageId="17" pageNumber="18" value="1996-03-13">13 Mar 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFBA3E010FBBBFBFB" box="[1055,1088,1072,1098]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFBE5E010FB09FBF9" box="[1113,1266,1072,1098]" pageId="17" pageNumber="18" value="1996-03-17">17 Mar 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFABAE010FADCFBFB" box="[1286,1319,1072,1098]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFAFDE010FF2AFBC1" pageId="17" pageNumber="18" value="1996-05-10">10 May 1996</date>
(
<specimenCount id="9D2103B7FFD1FF8EFF59E078FEFDFBC3" box="[229,262,1112,1138]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFEA2E078FE40FBC1" box="[286,443,1112,1138]" pageId="17" pageNumber="18" value="1996-05-12">12 May 1996</date>
(
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(
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(
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(
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(
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(
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(
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(
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(
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),
<date id="FF99EEFEFFD1FF8EFAE6E0A0FF2AFB71" pageId="17" pageNumber="18" value="1998-02-09">9 Feb 1998</date>
(
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),
<date id="FF99EEFEFFD1FF8EFE9CE088FE4EFB71" box="[288,437,1192,1218]" pageId="17" pageNumber="18" value="1998-02-10">10 Feb 1998</date>
(
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<specimenCount id="9D2103B7FFD1FF8EFE47E088FDE4FB72" box="[507,543,1191,1218]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
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(
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),
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(
<specimenCount id="9D2103B7FFD1FF8EFC7BE088FC13FB73" box="[967,1000,1192,1218]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
,
<specimenCount id="9D2103B7FFD1FF8EFC44E088FBE7FB72" box="[1016,1052,1191,1218]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFB8AE088FB3AFB71" box="[1078,1217,1192,1218]" pageId="17" pageNumber="18" value="1998-03-04">4 Mar 1998</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB68E088FB0CFB73" box="[1236,1271,1192,1218]" pageId="17" pageNumber="18" type="male">3♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFAACE088FA60FB71" box="[1296,1435,1192,1218]" pageId="17" pageNumber="18" value="1998-03-05">5 Mar 1998</date>
(
<specimenCount id="9D2103B7FFD1FF8EFF1EE0F0FF38FB5B" box="[162,195,1232,1258]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFF61E0F0FE8CFB59" box="[221,375,1232,1258]" pageId="17" pageNumber="18" value="1998-03-16">16 Mar 1998</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE30E0F0FE56FB5B" box="[396,429,1232,1258]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
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(
<specimenCount id="9D2103B7FFD1FF8EFDC8E0F0FD6CFB5B" box="[628,663,1232,1258]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFD0DE0F0FCC2FB59" box="[689,825,1232,1258]" pageId="17" pageNumber="18" value="1998-04-01">1 Apr 1998</date>
(
<specimenCount id="9D2103B7FFD1FF8EFCF2E0F0FC94FB5B" box="[846,879,1232,1258]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
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(
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(
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),
<date id="FF99EEFEFFD1FF8EFAF4E0F0FF2AFAA1" pageId="17" pageNumber="18" value="1998-04-29">29 Apr 1998</date>
(
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),
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(
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(
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<date id="FF99EEFEFFD1FF8EFD4CE0D8FC83FAA1" box="[752,888,1272,1298]" pageId="17" pageNumber="18" value="1998-07-14">14 Jul 1998</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC30E0D8FC57FAA3" box="[908,940,1272,1298]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
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(
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),
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(
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),
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(
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(
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(
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(
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(
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(
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(
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(
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(
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(
<specimenCount id="9D2103B7FFD1FF8EFBE7E168FB87FAD3" box="[1115,1148,1352,1378]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
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(
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(
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(
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(
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(
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(
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(
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<date id="FF99EEFEFFD1FF8EFF2BE1B8FEE5FA01" box="[151,286,1432,1458]" pageId="17" pageNumber="18" value="1999-06-01">1 Jun 1999</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE8AE1B8FEACFA03" box="[310,343,1432,1458]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
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<date id="FF99EEFEFFD1FF8EFECFE1B8FE00FA01" box="[371,507,1432,1458]" pageId="17" pageNumber="18" value="1999-06-09">9 Jun 1999</date>
(
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(
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(
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),
<date id="FF99EEFEFFD1FF8EFBDDE1E0FB0CFA69" box="[1121,1271,1472,1498]" pageId="17" pageNumber="18" value="2003-02-19">19 Feb 2003</date>
(
<specimenCount id="9D2103B7FFD1FF8EFAB1E1E0FAD5FA6B" box="[1293,1326,1472,1498]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFAF5E1E0FF2AF9B1" pageId="17" pageNumber="18" value="2003-02-20">20 Feb 2003</date>
(
<specimenCount id="9D2103B7FFD1FF8EFF55E1C8FEF1F9B3" box="[233,266,1512,1538]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFE9BE1C8FE3CF9B1" box="[295,455,1512,1538]" pageId="17" pageNumber="18" value="2003-03-21">21 Mar 2003</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE63E1C8FDFBF9B3" box="[479,512,1512,1538]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFDA1E1C8FD1EF9B1" box="[541,741,1512,1538]" pageId="17" pageNumber="18" value="2004-02-23" valueMax="2004-02-29" valueMin="2004-02-23">2329 Feb 2004</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD41E1C8FCE6F9B3" box="[765,797,1512,1538]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<materialsCitation id="3B4FC263FFD1FF8EFC87E1C8FF17F999" collectingDate="2006-03-07" collectionCode="INBio" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFC87E1C8FC37F9B1" box="[827,972,1512,1538]" pageId="17" pageNumber="18" value="2006-03-07">
<collectingDate id="EFDD1716FFD1FF8EFC87E1C8FC37F9B1" box="[827,972,1512,1538]" pageId="17" pageNumber="18" value="2006-03-07">7 Mar 2006</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC58E1C8FBFEF9B3" box="[996,1029,1512,1538]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), INBio-OET-ALAS transect (all
<collectionCode id="ED3650FBFFD1FF8EFF2BE230FF1FF999" box="[151,228,1552,1578]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFD1FF8EFF46E230FD5DF999" box="[250,678,1552,1578]" collectingDate="1998-01-17" collectionCode="EME" collectorName="Powell" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFF46E230FE70F999" box="[250,395,1552,1578]" pageId="17" pageNumber="18" value="1998-01-17">
<collectingDate id="EFDD1716FFD1FF8EFF46E230FE70F999" box="[250,395,1552,1578]" pageId="17" pageNumber="18" value="1998-01-17">17 Jan 1998</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE22E230FE3AF99B" box="[414,449,1552,1578]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
), J.
<collectorName id="26D2ADE8FFD1FF8EFE44E230FDB2F999" box="[504,585,1552,1578]" pageId="17" pageNumber="18">Powell</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFDE7E230FD65F999" box="[603,670,1552,1578]" pageId="17" pageNumber="18">EME</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFD1FF8EFD08E230FABFF999" box="[692,1348,1552,1578]" collectingDate="1992-03-27" collectingDateMax="1992-03-28" collectingDateMin="1992-03-27" collectionCode="EME" collectorName="McCarty &amp; Powell" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFD08E230FC80F999" box="[692,891,1552,1578]" pageId="17" pageNumber="18" value="1992-03-27" valueMax="1992-03-28" valueMin="1992-03-27">
<collectingDate id="EFDD1716FFD1FF8EFD08E230FC80F999" box="[692,891,1552,1578]" pageId="17" pageNumber="18" value="1992-03-27" valueMax="1992-03-28" valueMin="1992-03-27">2728 Mar 1992</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC31E230FC4AF99B" box="[909,945,1552,1578]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
), J.
<collectorName id="26D2ADE8FFD1FF8EFC5BE230FBB4F999" box="[999,1103,1552,1578]" pageId="17" pageNumber="18">McCarty</collectorName>
&amp; J.
<collectorName id="26D2ADE8FFD1FF8EFB29E230FB1DF999" box="[1173,1254,1552,1578]" pageId="17" pageNumber="18">Powell</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFB44E230FAC7F999" box="[1272,1340,1552,1578]" pageId="17" pageNumber="18">EME</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFAEDE230FA63F9E1" collectingDate="2004-03-19" collectionCode="INBio" country="Costa Rica" elevation="300" latitude="10.35" location="La Virgen" longLatPrecision="1298" longitude="-84.05" pageId="17" pageNumber="18" specimenCount="1" specimenCount-female="1" stateProvince="Heredia" typeStatus="paratype">
<quantity id="4CDF65DBFFD1FF8EFAEDE230FA67F999" box="[1361,1436,1552,1578]" metricMagnitude="4" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="km" value="11.0">11 km</quantity>
ESE
<location id="8EF89EE5FFD1FF8EFF6AE218FEA9F9E1" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFF6AE218FEA9F9E1" box="[214,338,1592,1618]" country="Costa Rica" latitude="10.35" longLatPrecision="1298" longitude="-84.05" name="La Virgen" pageId="17" pageNumber="18" stateProvince="Heredia">La Virgen</location>
,
<quantity id="4CDF65DBFFD1FF8EFEDEE218FE16F9E1" box="[354,493,1592,1618]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="17" pageNumber="18" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">
<elevation id="000A2F0DFFD1FF8EFEDEE218FE16F9E1" box="[354,493,1592,1618]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="17" pageNumber="18" unit="m" value="300.0" valueMax="350.0" valueMin="250.0">250350 m</elevation>
</quantity>
,
<geoCoordinate id="EE13AEF9FFD1FF8EFE42E218FD9EF9E1" box="[510,613,1591,1618]" degrees="10" direction="north" minutes="21" orientation="latitude" pageId="17" pageNumber="18" precision="925" value="10.35">10°21N</geoCoordinate>
,
<geoCoordinate id="EE13AEF9FFD1FF8EFDCAE218FD1AF9E1" box="[630,737,1591,1618]" degrees="84" direction="west" minutes="03" orientation="longitude" pageId="17" pageNumber="18" precision="925" value="-84.05">84°03W</geoCoordinate>
,
<date id="FF99EEFEFFD1FF8EFD4DE218FC6BF9E1" box="[753,912,1592,1618]" pageId="17" pageNumber="18" value="2004-03-19">
<collectingDate id="EFDD1716FFD1FF8EFD4DE218FC6BF9E1" box="[753,912,1592,1618]" pageId="17" pageNumber="18" value="2004-03-19">19 Mar 2004</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC1BE218FC31F9E2" box="[935,970,1591,1618]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
), INBio-OET-ALAS transect (
<collectionCode id="ED3650FBFFD1FF8EFAFCE218FA6BF9E1" box="[1344,1424,1592,1618]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFF2BE240FBBCF9C9" box="[151,1095,1632,1658]" collectingDate="1991-01" collectionCode="INBio" collectorName="Braulio Carrillo N. P. &amp; Aguilar" country="Costa Rica" elevation="200" location="Est. Magsasay" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<collectorName id="26D2ADE8FFD1FF8EFF2BE240FE76F9C9" box="[151,397,1632,1658]" pageId="17" pageNumber="18">Braulio Carrillo N.P.</collectorName>
,
<location id="8EF89EE5FFD1FF8EFE26E240FDBEF9C9" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFE26E240FDBEF9C9" box="[410,581,1632,1658]" country="Costa Rica" name="Est. Magsasay" pageId="17" pageNumber="18" stateProvince="Heredia">Est. Magsasay</location>
,
<quantity id="4CDF65DBFFD1FF8EFDEDE240FD64F9C9" box="[593,671,1632,1658]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="200.0">
<elevation id="000A2F0DFFD1FF8EFDEDE240FD64F9C9" box="[593,671,1632,1658]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="FF99EEFEFFD1FF8EFD17E240FCEEF9C9" box="[683,789,1632,1658]" pageId="17" pageNumber="18" value="1991-01">
<collectingDate id="EFDD1716FFD1FF8EFD17E240FCEEF9C9" box="[683,789,1632,1658]" pageId="17" pageNumber="18" value="1991-01">Jan 1991</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC9AE240FCB2F9CB" box="[806,841,1632,1658]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
), R.
<collectorName id="26D2ADE8FFD1FF8EFC3AE240FC24F9C9" box="[902,991,1632,1658]" pageId="17" pageNumber="18">Aguilar</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFC53E240FBC5F9C9" box="[1007,1086,1632,1658]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFBEFE27FFBAEF911" collectingDate="2001-03" collectionCode="INBio" collectorName="Valle de la Estrella, R. B. &amp; Hitoy Cerere, E. &amp; Chavarria" country="Costa Rica" elevation="100" location="Hitoy Cerere" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<emphasis id="B953142CFFD1FF8EFBEFE27FFB5DF9CA" bold="true" box="[1107,1190,1631,1657]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFBEFE27FFB5DF9CA" box="[1107,1190,1631,1657]" country="Costa Rica" name="Limon" pageId="17" pageNumber="18">Limón</collectingRegion>
</emphasis>
:
<collectorName id="26D2ADE8FFD1FF8EFB0AE240FE6FF911" pageId="17" pageNumber="18">Valle de la Estrella, R.B. Hitoy Cerere, E.</collectorName>
<location id="8EF89EE5FFD1FF8EFE27E2A8FDCDF911" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFE27E2A8FDCDF911" box="[411,566,1672,1698]" country="Costa Rica" name="Hitoy Cerere" pageId="17" pageNumber="18" stateProvince="Limon">Hitoy Cerere</location>
,
<quantity id="4CDF65DBFFD1FF8EFDFEE2A8FD75F911" box="[578,654,1672,1698]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">
<elevation id="000A2F0DFFD1FF8EFDFEE2A8FD75F911" box="[578,654,1672,1698]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="m" value="100.0">100 m</elevation>
</quantity>
,
<date id="FF99EEFEFFD1FF8EFD27E2A8FCF6F911" box="[667,781,1672,1698]" pageId="17" pageNumber="18" value="2001-03">
<collectingDate id="EFDD1716FFD1FF8EFD27E2A8FCF6F911" box="[667,781,1672,1698]" pageId="17" pageNumber="18" value="2001-03">Mar 2001</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC9CE2A8FCBAF913" box="[800,833,1672,1698]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), L.
<collectorName id="26D2ADE8FFD1FF8EFCC6E2A8FC16F911" box="[890,1005,1672,1698]" pageId="17" pageNumber="18">Chavarria</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFC41E2A8FBB6F911" box="[1021,1101,1672,1698]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFBDCE2A8FE80F979" collectorName="E. Rojas" country="Costa Rica" elevation="150" location="Finca" municipality="Sector Cerro Cocori" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
<collectingMunicipality id="6BFC5244FFD1FF8EFBDCE2A8FAB4F911" box="[1120,1359,1672,1698]" pageId="17" pageNumber="18">Sector Cerro Cocori</collectingMunicipality>
,
<location id="8EF89EE5FFD1FF8EFAE7E2A8FA67F911" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFAE7E2A8FA67F911" box="[1371,1436,1672,1698]" country="Costa Rica" municipality="Sector Cerro Cocori" name="Finca" pageId="17" pageNumber="18" stateProvince="Limon">Finca</location>
de
<collectorName id="26D2ADE8FFD1FF8EFF06E290FED9F979" box="[186,290,1712,1738]" pageId="17" pageNumber="18">E. Rojas</collectorName>
,
<quantity id="4CDF65DBFFD1FF8EFE92E290FE80F979" box="[302,379,1712,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="m" value="150.0">
<elevation id="000A2F0DFFD1FF8EFE92E290FE80F979" box="[302,379,1712,1738]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="17" pageNumber="18" unit="m" value="150.0">150 m</elevation>
</quantity>
</materialsCitation>
,
<date id="FF99EEFEFFD1FF8EFE34E290FE06F979" box="[392,509,1712,1738]" pageId="17" pageNumber="18" value="1991-11">Nov 1991</date>
(
<specimenCount id="9D2103B7FFD1FF8EFDADE290FDC9F97B" box="[529,562,1712,1738]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<materialsCitation id="3B4FC263FFD1FF8EFDF7E290FC34F979" box="[587,975,1712,1738]" collectingDate="1992-01" collectionCode="INBio" collectorName="Rojas" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFDF7E290FD4FF979" box="[587,692,1712,1738]" pageId="17" pageNumber="18" value="1992-01">
<collectingDate id="EFDD1716FFD1FF8EFDF7E290FD4FF979" box="[587,692,1712,1738]" pageId="17" pageNumber="18" value="1992-01">Jan 1992</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD74E290FD12F97B" box="[712,745,1712,1738]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), E.
<collectorName id="26D2ADE8FFD1FF8EFC98E290FC9CF979" box="[804,871,1712,1738]" pageId="17" pageNumber="18">Rojas</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFCCBE290FC3CF979" box="[887,967,1712,1738]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFC5FE28FFB5DF941" collectingDate="1981-01-05" collectingDateMax="1981-01-11" collectingDateMin="1981-01-05" collectionCode="INBio" collectorName="Est. Sirena, P. N &amp; Janzen &amp; Hallwachs" country="Costa Rica" elevation="50" location="Oso Peninsula" municipality="Corcovado" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<emphasis id="B953142CFFD1FF8EFC5FE28FFB89F97A" bold="true" box="[995,1138,1711,1737]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFC5FE28FFB89F97A" box="[995,1138,1711,1737]" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="18">Puntarenas</collectingRegion>
</emphasis>
:
<collectorName id="26D2ADE8FFD1FF8EFB3EE290FAC5F979" box="[1154,1342,1712,1738]" pageId="17" pageNumber="18">Est. Sirena, P.N</collectorName>
.
<collectingMunicipality id="6BFC5244FFD1FF8EFAF6E290FF28F941" pageId="17" pageNumber="18">Corcovado</collectingMunicipality>
,
<location id="8EF89EE5FFD1FF8EFF63E2F8FE70F941" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFF63E2F8FE70F941" box="[223,395,1752,1778]" country="Costa Rica" municipality="Corcovado" name="Oso Peninsula" pageId="17" pageNumber="18" stateProvince="Puntarenas">Oso Peninsula</location>
,
<quantity id="4CDF65DBFFD1FF8EFE2BE2F8FE07F941" box="[407,508,1752,1778]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="17" pageNumber="18" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
<elevation id="000A2F0DFFD1FF8EFE2BE2F8FE07F941" box="[407,508,1752,1778]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="17" pageNumber="18" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0-100 m</elevation>
</quantity>
,
<date id="FF99EEFEFFD1FF8EFDB5E2F8FD48F941" box="[521,691,1752,1778]" pageId="17" pageNumber="18" value="1981-01-05" valueMax="1981-01-11" valueMin="1981-01-05">
<collectingDate id="EFDD1716FFD1FF8EFDB5E2F8FD48F941" box="[521,691,1752,1778]" pageId="17" pageNumber="18" value="1981-01-05" valueMax="1981-01-11" valueMin="1981-01-05">511 Jan 1981</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFD7BE2F8FD13F943" box="[711,744,1752,1778]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), D.
<collectorName id="26D2ADE8FFD1FF8EFC99E2F8FC8EF941" box="[805,885,1752,1778]" pageId="17" pageNumber="18">Janzen</collectorName>
&amp; W.
<collectorName id="26D2ADE8FFD1FF8EFC7FE2F8FBC5F941" box="[963,1086,1752,1778]" pageId="17" pageNumber="18">Hallwachs</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFBF3E2F8FB65F941" box="[1103,1182,1752,1778]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
,
<date id="FF99EEFEFFD1FF8EFB0FE2F8FAE4F941" box="[1203,1311,1752,1778]" pageId="17" pageNumber="18" value="1990-10">Oct 1990</date>
(
<specimenCount id="9D2103B7FFD1FF8EFA8FE2F8FAAFF943" box="[1331,1364,1752,1778]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFAD1E2F8FF2AF8A9" pageId="17" pageNumber="18" value="1990-12">Dec 1990</date>
(
<specimenCount id="9D2103B7FFD1FF8EFF58E320FEFCF8AB" box="[228,263,1792,1818]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFE9EE320FE6DF8A9" box="[290,406,1792,1818]" pageId="17" pageNumber="18" value="1991-03">Mar 1991</date>
(
<specimenCount id="9D2103B7FFD1FF8EFE17E320FE37F8AB" box="[427,460,1792,1818]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFE5BE320FDA3F8A9" box="[487,600,1792,1818]" pageId="17" pageNumber="18" value="1991-04">Apr 1991</date>
(
<specimenCount id="9D2103B7FFD1FF8EFDD1E320FD75F8AB" box="[621,654,1792,1818]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFD15E320FCEDF8A9" box="[681,790,1792,1818]" pageId="17" pageNumber="18" value="1991-10">Oct 1991</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC90E320FCB6F8AB" box="[812,845,1792,1818]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<date id="FF99EEFEFFD1FF8EFCDBE320FC26F8A9" box="[871,989,1792,1818]" pageId="17" pageNumber="18" value="1991-11">Nov 1991</date>
(
<specimenCount id="9D2103B7FFD1FF8EFC4FE320FBE8F8AB" box="[1011,1043,1792,1818]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<materialsCitation id="3B4FC263FFD1FF8EFB92E320FF17F8F1" collectingDate="1994-01" collectionCode="INBio" collectorName="Fonseca" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFB92E320FB62F8A9" box="[1070,1177,1792,1818]" pageId="17" pageNumber="18" value="1994-01">
<collectingDate id="EFDD1716FFD1FF8EFB92E320FB62F8A9" box="[1070,1177,1792,1818]" pageId="17" pageNumber="18" value="1994-01">Jan 1994</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB12E320FB34F8AB" box="[1198,1231,1792,1818]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), G.
<collectorName id="26D2ADE8FFD1FF8EFAB7E320FA90F8A9" box="[1291,1387,1792,1818]" pageId="17" pageNumber="18">Fonseca</collectorName>
(all
<collectionCode id="ED3650FBFFD1FF8EFF2BE308FF1FF8F1" box="[151,228,1832,1858]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
,
<materialsCitation id="3B4FC263FFD1FF8EFF46E308FD3AF8F1" box="[250,705,1832,1858]" collectingDate="1991-12" collectionCode="INBio" collectorName="Saborio" country="Costa Rica" pageId="17" pageNumber="18" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFF46E308FE96F8F1" box="[250,365,1832,1858]" pageId="17" pageNumber="18" value="1991-12">
<collectingDate id="EFDD1716FFD1FF8EFF46E308FE96F8F1" box="[250,365,1832,1858]" pageId="17" pageNumber="18" value="1991-12">Dec 1991</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFEC3E308FE58F8F3" box="[383,419,1832,1858]" pageId="17" pageNumber="18" type="male">2♂</specimenCount>
), J. C.
<collectorName id="26D2ADE8FFD1FF8EFE41E308FDA3F8F1" box="[509,600,1832,1858]" pageId="17" pageNumber="18">Saborio</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFDD5E308FD43F8F1" box="[617,696,1832,1858]" country="Costa Rica" httpUri="http://grbio.org/cool/931n-s1te" name="National Biodiversity Institute, Costa Rica" pageId="17" pageNumber="18">INBio</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFD72E308FD97F8D9" collectingDate="1978-03-09" collectionCode="USNM" collectorName="Yurubi N. P. &amp; J. B. Heppner" country="Venezuela" location="20 km SE Yumare" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Yaracuy" typeStatus="paratype">
<collectingCountry id="F33088AEFFD1FF8EFD72E308FC7AF8F1" box="[718,897,1832,1858]" name="Venezuela" pageId="17" pageNumber="18">VENEZUELA</collectingCountry>
:
<emphasis id="B953142CFFD1FF8EFC32E307FC0DF8F2" bold="true" box="[910,1014,1831,1857]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFC32E307FC0DF8F2" box="[910,1014,1831,1857]" country="Venezuela" name="Yaracuy" pageId="17" pageNumber="18">Yaracuy</collectingRegion>
</emphasis>
:
<collectorName id="26D2ADE8FFD1FF8EFBBBE308FB68F8F1" box="[1031,1171,1832,1858]" pageId="17" pageNumber="18">Yurubi N.P.</collectorName>
,
<location id="8EF89EE5FFD1FF8EFB1DE308FA7BF8F1" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFB1DE308FA7BF8F1" box="[1185,1408,1832,1858]" country="Venezuela" name="20 km SE Yumare" pageId="17" pageNumber="18" stateProvince="Yaracuy">
<quantity id="4CDF65DBFFD1FF8EFB1DE308FB16F8F1" box="[1185,1261,1832,1858]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="18" unit="km" value="20.0">20 km</quantity>
SE Yumare
</location>
,
<date id="FF99EEFEFFD1FF8EFA31E308FEF2F8D9" pageId="17" pageNumber="18" value="1978-03-09">
<collectingDate id="EFDD1716FFD1FF8EFA31E308FEF2F8D9" pageId="17" pageNumber="18" value="1978-03-09">9 Mar 1978</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFEA1E370FEC5F8DB" box="[285,318,1872,1898]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
),
<collectorName id="26D2ADE8FFD1FF8EFEEBE370FE02F8D9" box="[343,505,1872,1898]" pageId="17" pageNumber="18">J. B. Heppner</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFDB5E370FD9FF8D9" box="[521,612,1872,1898]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B4FC263FFD1FF8EFDC4E36FFBF2F8D9" box="[632,1033,1871,1898]" country="Venezuela" elevation="1100" location="Rancho Grande" pageId="17" pageNumber="18" specimenCount="1" stateProvince="Aragua" typeStatus="paratype">
<emphasis id="B953142CFFD1FF8EFDC4E36FFD1BF8DA" bold="true" box="[632,736,1871,1897]" pageId="17" pageNumber="18">
<collectingRegion id="49E306DCFFD1FF8EFDC4E36FFD20F8DA" box="[632,731,1871,1897]" country="Venezuela" name="Aragua" pageId="17" pageNumber="18">Aragua</collectingRegion>
:
</emphasis>
<location id="8EF89EE5FFD1FF8EFD54E370FC59F8D9" LSID="urn:lsid:plazi:treatment:038E7928FFD0FF8DFDABE135FAEFFEB1:8EF89EE5FFD1FF8EFD54E370FC59F8D9" box="[744,930,1872,1898]" country="Venezuela" name="Rancho Grande" pageId="17" pageNumber="18" stateProvince="Aragua">Rancho Grande</location>
,
<quantity id="4CDF65DBFFD1FF8EFC12E370FBF2F8D9" box="[942,1033,1872,1898]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="m" value="1100.0">
<elevation id="000A2F0DFFD1FF8EFC12E370FBF2F8D9" box="[942,1033,1872,1898]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="17" pageNumber="18" unit="m" value="1100.0">1100 m</elevation>
</quantity>
</materialsCitation>
,
<date id="FF99EEFEFFD1FF8EFBAAE370FB28F8D9" box="[1046,1235,1872,1898]" pageId="17" pageNumber="18" value="1966-10-16" valueMax="1966-10-23" valueMin="1966-10-16">1623 Oct 1966</date>
(
<specimenCount id="9D2103B7FFD1FF8EFB5AE370FAFCF8DB" box="[1254,1287,1872,1898]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
,
<specimenCount id="9D2103B7FFD1FF8EFAAAE370FAC0F8DA" box="[1302,1339,1871,1898]" pageId="17" pageNumber="18" type="female">1♀</specimenCount>
),
<materialsCitation id="3B4FC263FFD1FF8EFAE8E370FD11F821" collectingDate="1966-01-11" collectingDateMax="1966-01-15" collectingDateMin="1966-01-11" collectionCode="USNM" collectorName="Duckworth" country="Venezuela" pageId="17" pageNumber="18" specimenCount="1" specimenCount-male="1" stateProvince="Aragua" typeStatus="paratype">
<date id="FF99EEFEFFD1FF8EFAE8E370FF04F821" pageId="17" pageNumber="18" value="1966-01-11" valueMax="1966-01-15" valueMin="1966-01-11">
<collectingDate id="EFDD1716FFD1FF8EFAE8E370FF04F821" pageId="17" pageNumber="18" value="1966-01-11" valueMax="1966-01-15" valueMin="1966-01-11">1115 Jan 1966</collectingDate>
</date>
(
<specimenCount id="9D2103B7FFD1FF8EFEAFE358FEC8F823" box="[275,307,1912,1938]" pageId="17" pageNumber="18" type="male">1♂</specimenCount>
), S. S. &amp; W. D.
<collectorName id="26D2ADE8FFD1FF8EFE4FE358FD8CF821" box="[499,631,1912,1938]" pageId="17" pageNumber="18">Duckworth</collectorName>
(
<collectionCode id="ED3650FBFFD1FF8EFD3BE358FD19F821" box="[647,738,1912,1938]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="18" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFD1FF8DFF7AE3BFFDB7FF69" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="distribution">
<paragraph id="8B98C83EFFD1FF8DFF7AE3BFFDB7FF69" blockId="17.[151,1436,152,2018]" lastBlockId="18.[151,1436,152,258]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B953142CFFD1FF8EFF7AE3BFFE01F80A" bold="true" box="[198,506,1951,1977]" pageId="17" pageNumber="18">Distribution and Biology</emphasis>
. This species is known from the
<collectingRegion id="49E306DCFFD1FF8EFC3CE380FC39F809" box="[896,962,1952,1978]" country="New Zealand" name="West Coast" pageId="17" pageNumber="18">Costa</collectingRegion>
Rican provinces of
<collectingRegion id="49E306DCFFD1FF8EFB0CE380FAF7F809" box="[1200,1292,1952,1978]" country="Costa Rica" name="Heredia" pageId="17" pageNumber="18">Heredia</collectingRegion>
and
<collectingRegion id="49E306DCFFD1FF8EFAFAE380FF30F851" country="Costa Rica" name="Puntarenas" pageId="17" pageNumber="18">Puntarenas</collectingRegion>
below
<quantity id="4CDF65DBFFD1FF8EFE9CE3E8FE90F851" box="[288,363,1992,2018]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="17" pageNumber="18" unit="m" value="500.0">500 m</quantity>
elevation, and from the Venezuelan provinces of
<collectingRegion id="49E306DCFFD1FF8EFC11E3E8FBF6F851" box="[941,1037,1992,2018]" country="Venezuela" name="Yaracuy" pageId="17" pageNumber="18">Yaracuy</collectingRegion>
and
<collectingRegion id="49E306DCFFD1FF8EFBFAE3E8FB67F851" box="[1094,1180,1992,2018]" country="Venezuela" name="Aragua" pageId="17" pageNumber="18">Aragua</collectingRegion>
at slightly higher elevations. It is recorded throughout much of the year, but records are absent from January-February and July. Nothing is known of the early stages.
</paragraph>
</subSubSection>
<subSubSection id="C33D9BB5FFD2FF8DFF7AE4C7FAEFFEB1" box="[198,1300,231,258]" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8B98C83EFFD2FF8DFF7AE4C7FAEFFEB1" blockId="18.[151,1436,152,258]" box="[198,1300,231,258]" pageId="18" pageNumber="19">
<emphasis id="B953142CFFD2FF8DFF7AE4C7FEB0FEB2" bold="true" box="[198,331,231,257]" pageId="18" pageNumber="19">Etymology</emphasis>
. The species name refers to the flangelike spine on the valva of the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>