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<document id="3CED8E7F24892EC0E5C011782ACD2A81" ID-CLB-Dataset="6101" ID-DOI="10.11646/zootaxa.4980.1.3" ID-GBIF-Dataset="9e5a4897-46e2-4e6f-8dc9-4aad664b979d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4882930" ID-ZooBank="B136A181-ED9A-410D-A9E3-3CEB1F81A4B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1622461250437" checkinUser="plazi" docAuthor="Kovačić, Marcelo, Esmaeili, Hamid Reza, Zarei, Fatah, Abbasi, Keyvan &amp; Schliewen, Ulrich K." docDate="2021" docId="038D87D3A805FF9DC1ECF994718BFE2A" docLanguage="en" docName="zootaxa.4980.1.3.pdf" docOrigin="Zootaxa 4980 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Benthophilus persicus Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="61" masterDocId="FFB4FFABA807FF8DC17BFF867419FF87" masterDocTitle="A new species of tadpole-goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea" masterLastPageNumber="63" masterPageNumber="45" pageNumber="47" updateTime="1698995035287" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="68AF0250B5D776EB4B5CE4C322CF76F6">A new species of tadpole-goby, Benthophilus persicus sp. nov. (Teleostei: Gobiidae) from the southern Caspian Sea</mods:title>
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<mods:namePart id="6732A88889C5185E3BAE11D7304E35C3">Kovačić, Marcelo</mods:namePart>
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<mods:namePart id="B2F75C9632C502C075889C6910D91D20">Esmaeili, Hamid Reza</mods:namePart>
<mods:affiliation id="43B8D960DD7C8CAA8E211E06F16AEBA9">Ichthyology and Molecular Systematics Research laboratory, Zoology Section, Department of Biology, School of Science, Shiraz University, Shiraz, Iran.</mods:affiliation>
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<mods:namePart id="03320591EE86F894C1478F7EF20A9788">Zarei, Fatah</mods:namePart>
<mods:affiliation id="E8BEEC9241773581F579B368FEC483CB">Ichthyology and Molecular Systematics Research laboratory, Zoology Section, Department of Biology, School of Science, Shiraz University, Shiraz, Iran.</mods:affiliation>
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<mods:namePart id="C03E57979EF956844F021D2CFAA41970">Abbasi, Keyvan</mods:namePart>
<mods:affiliation id="A8F9568A29E613DC8E9AD2C6F4E5B503">Inland Waters Aquaculture Research Center, Iranian Fisheries Sciences Research Institute, Agricultural Research, Education and Extension Organization, Bandar Anzali, Iran.</mods:affiliation>
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<mods:namePart id="31E441579F3B455BE979CA7F915F59A4">Schliewen, Ulrich K.</mods:namePart>
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<mods:affiliation id="573DD7F397B4FFB79E4B4F9050D4F88E">Department of Ichthyology, SNSB - Bavarian State Collection of Zoology, Münchhausenstr. 21, 81247 Munich, Germany. schliewen @ snsb. de; https: // orcid. org / 0000 - 0003 - 0187 - 4319</mods:affiliation>
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<taxonomicName id="4C244D46A805FF8FC1ECF99475BCF9AA" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[151,421,1554,1581]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="47" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A805FF8FC1ECF99475BCF9AA" bold="true" box="[151,421,1554,1581]" italics="true" pageId="2" pageNumber="47">Benthophilus persicus</emphasis>
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<paragraph id="8B9B36C5A805FF8FC1ECF9B17560F9D6" blockId="2.[151,522,1554,1617]" box="[151,377,1591,1617]" pageId="2" pageNumber="47">
(
<figureCitation id="131F2A40A805FF8FC1E4F9B1750BF9D6" box="[159,274,1591,1617]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStart-6="FIGURE 8" captionStart-7="FIGURE 9" captionStart-8="FIGURE 10" captionStartId-0="3.[151,250,966,990]" captionStartId-1="3.[151,250,1907,1931]" captionStartId-2="4.[151,250,726,750]" captionStartId-3="4.[151,250,1857,1881]" captionStartId-4="5.[151,250,1909,1933]" captionStartId-5="6.[151,250,1776,1800]" captionStartId-6="10.[151,250,798,822]" captionStartId-7="10.[151,250,1929,1953]" captionStartId-8="11.[151,250,1076,1100]" captionTargetBox-0="[225,1361,181,942]" captionTargetBox-1="[225,1361,1122,1883]" captionTargetBox-2="[248,1340,181,702]" captionTargetBox-3="[248,1339,882,1824]" captionTargetBox-4="[315,1272,181,1885]" captionTargetBox-5="[396,1191,181,1753]" captionTargetBox-6="[262,1333,191,774]" captionTargetBox-7="[265,1323,882,1905]" captionTargetBox-8="[213,1375,181,1052]" captionTargetId-0="figure-20@3.[225,1361,181,942]" captionTargetId-1="figure-79@3.[225,1361,1122,1883]" captionTargetId-2="figure-33@4.[248,1340,181,702]" captionTargetId-3="figure-102@4.[248,1339,882,1833]" captionTargetId-4="figure-20@5.[315,1272,181,1885]" captionTargetId-5="figure-33@6.[396,1191,181,1753]" captionTargetId-6="figure-33@10.[254,1333,181,774]" captionTargetId-7="figure-71@10.[265,1323,882,1905]" captionTargetId-8="figure-20@11.[213,1375,181,1052]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionTargetPageId-4="5" captionTargetPageId-5="6" captionTargetPageId-6="10" captionTargetPageId-7="10" captionTargetPageId-8="11" captionText-0="FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić." captionText-1="FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić." captionText-2="FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić." captionText-3="FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić." captionText-4="FIGURE 6. Benthophilus persicus sp. nov. Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea: A: lateral detail of head, B: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian: C: chin close up with barbel. Photo by M. Kovačić." captionText-5="FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić." captionText-6="FIGURE 8. Head sensory papillae pattern of lateral line system of Benthophilus persicus sp. nov. Paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Terminology in text. Drawing by M. Kovačić." captionText-7="FIGURE 9. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, anterior part of preoperculo-mandibular rows in PMR VP4680, male, 47.0+8.6 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Terminology in text. Photo by M. Kovačić." captionText-8="FIGURE 10. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, sensory papillae rows on anterior flank in PMR VP4681, male, 35.1+8.5 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Sensory papillae rows marked, terminology text. The transversal ltm rows starting anteriorly behind axilla, alternating with three longitudinal llm rows making anterior beginning pattern of -II-I-I. Photo by M. Kovačić." figureDoi-0="http://doi.org/10.5281/zenodo.4882934" figureDoi-1="http://doi.org/10.5281/zenodo.4882936" figureDoi-2="http://doi.org/10.5281/zenodo.4882938" figureDoi-3="http://doi.org/10.5281/zenodo.4882940" figureDoi-4="http://doi.org/10.5281/zenodo.4882942" figureDoi-5="http://doi.org/10.5281/zenodo.4882944" figureDoi-6="http://doi.org/10.5281/zenodo.4882946" figureDoi-7="http://doi.org/10.5281/zenodo.4882948" figureDoi-8="http://doi.org/10.5281/zenodo.4882950" httpUri-0="https://zenodo.org/record/4882934/files/figure.png" httpUri-1="https://zenodo.org/record/4882936/files/figure.png" httpUri-2="https://zenodo.org/record/4882938/files/figure.png" httpUri-3="https://zenodo.org/record/4882940/files/figure.png" httpUri-4="https://zenodo.org/record/4882942/files/figure.png" httpUri-5="https://zenodo.org/record/4882944/files/figure.png" httpUri-6="https://zenodo.org/record/4882946/files/figure.png" httpUri-7="https://zenodo.org/record/4882948/files/figure.png" httpUri-8="https://zenodo.org/record/4882950/files/figure.png" pageId="2" pageNumber="47">Figs 210</figureCitation>
,
<tableCitation id="C6A6037EA805FF8FC066F9B17577F9D6" box="[285,366,1591,1617]" captionStart="TABLE 1" captionStartId="13.[151,239,477,503]" captionTargetBox="[168,2025,656,1650]" captionText="TABLE 1. Benthophilus persicus sp. nov. Morphometric characters as % of standard length or as % of head length. Characters are sorted as in Boldyrev &amp; Bogutskaya (2007) with the new introduced characters added at the end in alphabetic order. The morphometric character and the related values of dimorphic characters with non-overlapping ranges of groups marked with ^ for adult males vs. the immature male and all females, with * for all males and the large female vs. small mature and immature females, and with ˜ for adult males and the large female vs. smaller females and the immature male, see Discussion." httpUri="http://table.plazi.org/id/DF5B664DA80AFF80C1ECFE5B7068FDE4" pageId="2" pageNumber="47" tableUuid="DF5B664DA80AFF80C1ECFE5B7068FDE4">Table 1</tableCitation>
)
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<paragraph id="8B9B36C5A805FF8FC1ECF9F9771FF93A" blockId="2.[151,1437,1662,1941]" pageId="2" pageNumber="47">
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47595, male, 45.2+
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<collectingDate id="EFDEE9EDA805FF8FC37DF9257681F93A" box="[518,664,1699,1725]" pageId="2" pageNumber="47" value="2004-04-01">01 Apr. 2004</collectingDate>
</date>
(
<figureCitation id="131F2A40A805FF8FC3DCF92576E0F93A" box="[679,761,1699,1725]" captionStart="FIGURE 4" captionStartId="4.[151,250,726,750]" captionTargetBox="[248,1340,181,702]" captionTargetId="figure-33@4.[248,1340,181,702]" captionTargetPageId="4" captionText="FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882938" httpUri="https://zenodo.org/record/4882938/files/figure.png" pageId="2" pageNumber="47">Fig. 4a</figureCitation>
).
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</paragraph>
<paragraph id="8B9B36C5A805FF8FC1BCF941777AF812" blockId="2.[151,1437,1662,1941]" pageId="2" pageNumber="47">
<materialsCitation id="3B4C3C98A805FF8FC1BCF94176EDF882" ID-GBIF-Occurrence="3131327301" accessionNumber="ZSM 47596" collectingDate="2002-11-18" collectionCode="ZSM" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" latitude="37.516666" location="southern Caspian Sea" longLatPrecision="1181" longitude="49.5" municipality="Anzali" pageId="2" pageNumber="47" specimenCount="1" stateProvince="Gilan" typeStatus="paratype">
<emphasis id="B950EAD7A805FF8FC1BCF9417526F966" bold="true" box="[199,319,1735,1761]" pageId="2" pageNumber="47">
<typeStatus id="549F8867A805FF8FC1BCF9417526F966" box="[199,319,1735,1761]" pageId="2" pageNumber="47" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectionCode id="ED35AE00A805FF8FC035F9417591F966" box="[334,392,1735,1761]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="47" type="Museum">ZSM</collectionCode>
47596, female, 45.8+
<quantity id="4CDC9B20A805FF8FC3FAF94076C3F966" box="[641,730,1734,1761]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="2" pageNumber="47" unit="mm" value="9.7">9.7 mm</quantity>
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<collectingCountry id="F3337655A805FF8FC39DF941770FF966" box="[742,790,1735,1761]" name="Iran" pageId="2" pageNumber="47">Iran</collectingCountry>
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,
<collectingMunicipality id="6BFFACBFA805FF8FC2A1F941703EF966" box="[986,1063,1735,1761]" pageId="2" pageNumber="47">Anzali</collectingMunicipality>
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&amp;
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</date>
</materialsCitation>
.
<materialsCitation id="3B4C3C98A805FF8FC385F96D70B7F8AE" ID-GBIF-Occurrence="3131327307" accessionNumber="ZSM 47597" collectingDate="2004-01-06" collectionCode="ZSM" collectorName="K. Abbasi &amp; M. Tavakoli" country="Iran" latitude="37.6" location="southern Caspian Sea" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" pageId="2" pageNumber="47" specimenCount="1" stateProvince="Gilan" typeStatus="paratype">
<collectionCode id="ED35AE00A805FF8FC385F96D7721F882" box="[766,824,1771,1797]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="47" type="Museum">ZSM</collectionCode>
47597, juvenile male, 30.4+
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<collectingCountry id="F3337655A805FF8FC5A6F96D7114F882" box="[1245,1293,1771,1797]" name="Iran" pageId="2" pageNumber="47">Iran</collectingCountry>
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<collectingMunicipality id="6BFFACBFA805FF8FC1AAF8897507F8AE" box="[209,286,1807,1833]" pageId="2" pageNumber="47">Anzali</collectingMunicipality>
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<location id="8EFB601EA805FF8FC053F8897600F8AE" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA805FF8FC053F8897600F8AE" box="[296,537,1807,1833]" country="Iran" latitude="37.6" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="2" pageNumber="47" stateProvince="Gilan">southern Caspian Sea</location>
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<collectorName id="26D15313A805FF8FC380F8897773F8AE" box="[763,874,1807,1833]" pageId="2" pageNumber="47">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A805FF8FC2F0F889700CF8AE" box="[907,1045,1807,1833]" pageId="2" pageNumber="47">M. Tavakoli</collectorName>
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</date>
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.
<materialsCitation id="3B4C3C98A805FF8FC5CCF8897573F8F6" ID-GBIF-Occurrence="3131327303" accessionNumber="ZSM 47599" collectingDate="2004-01-05" collectionCode="ZSM" collectorName="K. Abbasi &amp; M. Tavakoli" country="Iran" latitude="37.6" location="southern Caspian Sea" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" pageId="2" pageNumber="47" specimenCount="1" stateProvince="Gilan" typeStatus="paratype">
<collectionCode id="ED35AE00A805FF8FC5CCF88970E8F8AE" box="[1207,1265,1807,1833]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="47" type="Museum">ZSM</collectionCode>
47599, female, 30.8+7.0 mm,
<collectingCountry id="F3337655A805FF8FC041F8B57573F8CA" box="[314,362,1843,1869]" name="Iran" pageId="2" pageNumber="47">Iran</collectingCountry>
,
<collectingRegion id="49E0F827A805FF8FC00EF8B57639F8CA" box="[373,544,1843,1869]" country="Iran" name="Gilan" pageId="2" pageNumber="47">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA805FF8FC352F8B5766FF8CA" box="[553,630,1843,1869]" pageId="2" pageNumber="47">Anzali</collectingMunicipality>
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<location id="8EFB601EA805FF8FC3FAF8B5776DF8CA" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA805FF8FC3FAF8B5776DF8CA" box="[641,884,1843,1869]" country="Iran" latitude="37.6" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="2" pageNumber="47" stateProvince="Gilan">southern Caspian Sea</location>
,
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&amp;
<collectorName id="26D15313A805FF8FC590F8B5716FF8CA" box="[1259,1398,1843,1869]" pageId="2" pageNumber="47">M. Tavakoli</collectorName>
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<collectingDate id="EFDEE9EDA805FF8FC4FBF8B47518F8F6" pageId="2" pageNumber="47" value="2004-01-05">05 Jan. 2004</collectingDate>
</date>
(
<figureCitation id="131F2A40A805FF8FC06BF8D17578F8F6" box="[272,353,1879,1905]" captionStart="FIGURE 4" captionStartId="4.[151,250,726,750]" captionTargetBox="[248,1340,181,702]" captionTargetId="figure-33@4.[248,1340,181,702]" captionTargetPageId="4" captionText="FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882938" httpUri="https://zenodo.org/record/4882938/files/figure.png" pageId="2" pageNumber="47">Fig. 4c</figureCitation>
)
</materialsCitation>
.
<materialsCitation id="3B4C3C98A805FF8FC00FF8D17779F812" ID-GBIF-Occurrence="3131327302" accessionNumber="ZSM 47598" collectingDate="2002-11-01" collectionCode="ZSM" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" latitude="37.6" location="southern Caspian Sea" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" pageId="2" pageNumber="47" specimenCount="1" stateProvince="Gilan" typeStatus="paratype">
<collectionCode id="ED35AE00A805FF8FC00FF8D175B7F8F6" box="[372,430,1879,1905]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="2" pageNumber="47" type="Museum">ZSM</collectionCode>
47598, juvenile female, 27.5+
<quantity id="4CDC9B20A805FF8FC27BF8D0774EF8F6" box="[768,855,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="2" pageNumber="47" unit="mm" value="6.4">6.4 mm</quantity>
,
<collectingCountry id="F3337655A805FF8FC219F8D1778BF8F6" box="[866,914,1879,1905]" name="Iran" pageId="2" pageNumber="47">Iran</collectingCountry>
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<collectingRegion id="49E0F827A805FF8FC2E7F8D1705FF8F6" box="[924,1094,1879,1905]" country="Iran" name="Gilan" pageId="2" pageNumber="47">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA805FF8FC534F8D17085F8F6" box="[1103,1180,1879,1905]" pageId="2" pageNumber="47">Anzali</collectingMunicipality>
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<location id="8EFB601EA805FF8FC5DDF8D17181F8F6" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA805FF8FC5DDF8D17181F8F6" box="[1190,1432,1879,1905]" country="Iran" latitude="37.6" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="2" pageNumber="47" stateProvince="Gilan">southern Caspian Sea</location>
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<geoCoordinate id="EE105002A805FF8FC1ECF8FD74E4F812" box="[151,253,1915,1941]" degrees="37" direction="north" minutes="36" orientation="latitude" pageId="2" pageNumber="47" precision="925" value="37.6">37°36 N</geoCoordinate>
<geoCoordinate id="EE105002A805FF8FC07FF8FD7571F812" box="[260,360,1915,1941]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="2" pageNumber="47" precision="925" value="49.483334">49°29 E</geoCoordinate>
,
<collectorName id="26D15313A805FF8FC00FF8FD75FDF812" box="[372,484,1915,1941]" pageId="2" pageNumber="47">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A805FF8FC373F8FD76A0F812" box="[520,697,1915,1941]" pageId="2" pageNumber="47">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A805FF8FC3BFF8FD7779F812" box="[708,864,1915,1941]" pageId="2" pageNumber="47" value="2002-11-01">
<collectingDate id="EFDEE9EDA805FF8FC3BFF8FD7779F812" box="[708,864,1915,1941]" pageId="2" pageNumber="47" value="2002-11-01">01 Nov. 2002</collectingDate>
</date>
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.
</paragraph>
<caption id="DF5B664DA804FF8EC1ECFC4074EAFBCD" ID-DOI="http://doi.org/10.5281/zenodo.4882934" ID-Zenodo-Dep="4882934" httpUri="https://zenodo.org/record/4882934/files/figure.png" pageId="3" pageNumber="48" startId="3.[151,250,966,990]" targetBox="[225,1361,181,942]" targetPageId="3">
<paragraph id="8B9B36C5A804FF8EC1ECFC4074EAFBCD" blockId="3.[151,1437,966,1098]" pageId="3" pageNumber="48">
<emphasis id="B950EAD7A804FF8EC1ECFC40750EFC59" bold="true" box="[151,279,966,990]" pageId="3" pageNumber="48">FIGURE 2.</emphasis>
<taxonomicName id="4C244D46A804FF8EC05AFC40761DFC59" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[289,516,966,990]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="48" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A804FF8EC05AFC40761DFC59" box="[289,516,966,990]" italics="true" pageId="3" pageNumber="48">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A804FF8EC374FC40767AFC59" bold="true" box="[527,611,966,990]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="A26357ACA804FF8EC374FC40767AFC59" box="[527,611,966,990]" pageId="3" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA804FF8EC1ECF8F57618F870" ID-DOI="http://doi.org/10.5281/zenodo.4882936" ID-Zenodo-Dep="4882936" httpUri="https://zenodo.org/record/4882936/files/figure.png" pageId="3" pageNumber="48" startId="3.[151,250,1907,1931]" targetBox="[225,1361,1122,1883]" targetPageId="3">
<paragraph id="8B9B36C5A804FF8EC1ECF8F57618F870" blockId="3.[151,1436,1907,2040]" pageId="3" pageNumber="48">
<emphasis id="B950EAD7A804FF8EC1ECF8F5750EF80C" bold="true" box="[151,279,1907,1931]" pageId="3" pageNumber="48">FIGURE 3.</emphasis>
<taxonomicName id="4C244D46A804FF8EC05AF8F5761DF80C" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[289,516,1907,1931]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="48" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A804FF8EC05AF8F5761DF80C" box="[289,516,1907,1931]" italics="true" pageId="3" pageNumber="48">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A804FF8EC374F8F2767AF80B" bold="true" box="[527,611,1908,1932]" pageId="3" pageNumber="48">
<taxonomicNameLabel id="A26357ACA804FF8EC374F8F2767AF80B" box="[527,611,1908,1932]" pageId="3" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA803FF89C1ECFD50768EFCDD" ID-DOI="http://doi.org/10.5281/zenodo.4882938" ID-Zenodo-Dep="4882938" httpUri="https://zenodo.org/record/4882938/files/figure.png" pageId="4" pageNumber="49" startId="4.[151,250,726,750]" targetBox="[248,1340,181,702]" targetPageId="4">
<paragraph id="8B9B36C5A803FF89C1ECFD50768EFCDD" blockId="4.[151,1437,726,858]" pageId="4" pageNumber="49">
<emphasis id="B950EAD7A803FF89C1ECFD50750FFD69" bold="true" box="[151,278,726,750]" pageId="4" pageNumber="49">FIGURE 4.</emphasis>
<taxonomicName id="4C244D46A803FF89C05BFD50761DFD69" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[288,516,726,750]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A803FF89C05BFD50761DFD69" box="[288,516,726,750]" italics="true" pageId="4" pageNumber="49">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A803FF89C375FD50767BFD69" bold="true" box="[526,610,726,750]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="A26357ACA803FF89C375FD50767BFD69" box="[526,610,726,750]" pageId="4" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea:
<emphasis id="B950EAD7A803FF89C34FFD7C765FFC95" bold="true" box="[564,582,762,786]" pageId="4" pageNumber="49">A</emphasis>
: lateral view,
<emphasis id="B950EAD7A803FF89C3ADFD7C76FEFC95" bold="true" box="[726,743,762,786]" pageId="4" pageNumber="49">B</emphasis>
: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea:
<emphasis id="B950EAD7A803FF89C2ACFC9877F0FCB1" bold="true" box="[983,1001,798,822]" pageId="4" pageNumber="49">C</emphasis>
: lateral view,
<emphasis id="B950EAD7A803FF89C50CFC987090FCB1" bold="true" box="[1143,1161,798,822]" pageId="4" pageNumber="49">D</emphasis>
: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA803FF89C1ECF8C77060F86E" ID-DOI="http://doi.org/10.5281/zenodo.4882940" ID-Zenodo-Dep="4882940" httpUri="https://zenodo.org/record/4882940/files/figure.png" pageId="4" pageNumber="49" startId="4.[151,250,1857,1881]" targetBox="[248,1339,882,1824]" targetPageId="4">
<paragraph id="8B9B36C5A803FF89C1ECF8C77060F86E" blockId="4.[151,1436,1857,2026]" pageId="4" pageNumber="49">
<emphasis id="B950EAD7A803FF89C1ECF8C7750BF8DE" bold="true" box="[151,274,1857,1881]" pageId="4" pageNumber="49">FIGURE 5.</emphasis>
<taxonomicName id="4C244D46A803FF89C063F8C775E1F8DE" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[280,504,1857,1881]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="4" pageNumber="49" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A803FF89C063F8C775E1F8DE" box="[280,504,1857,1881]" italics="true" pageId="4" pageNumber="49">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A803FF89C085F8C77657F8DE" bold="true" box="[510,590,1857,1881]" pageId="4" pageNumber="49">
<taxonomicNameLabel id="A26357ACA803FF89C085F8C77657F8DE" box="[510,590,1857,1881]" pageId="4" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA802FF88C1ECF8F375EEF87E" ID-DOI="http://doi.org/10.5281/zenodo.4882942" ID-Zenodo-Dep="4882942" httpUri="https://zenodo.org/record/4882942/files/figure.png" pageId="5" pageNumber="50" startId="5.[151,250,1909,1933]" targetBox="[315,1272,181,1885]" targetPageId="5">
<paragraph id="8B9B36C5A802FF88C1ECF8F375EEF87E" blockId="5.[151,1437,1909,2041]" pageId="5" pageNumber="50">
<emphasis id="B950EAD7A802FF88C1ECF8F37508F80A" bold="true" box="[151,273,1909,1933]" pageId="5" pageNumber="50">FIGURE 6.</emphasis>
<taxonomicName id="4C244D46A802FF88C06CF8F375ECF80A" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[279,501,1909,1933]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="5" pageNumber="50" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A802FF88C06CF8F375ECF80A" box="[279,501,1909,1933]" italics="true" pageId="5" pageNumber="50">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A802FF88C080F8F37653F80A" bold="true" box="[507,586,1909,1933]" pageId="5" pageNumber="50">
<taxonomicNameLabel id="A26357ACA802FF88C080F8F37653F80A" box="[507,586,1909,1933]" pageId="5" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea:
<emphasis id="B950EAD7A802FF88C2D5F81F77D9F836" bold="true" box="[942,960,1945,1969]" pageId="5" pageNumber="50">A</emphasis>
: lateral detail of head,
<emphasis id="B950EAD7A802FF88C5D2F81F70A3F836" bold="true" box="[1193,1210,1945,1969]" pageId="5" pageNumber="50">B</emphasis>
: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian:
<emphasis id="B950EAD7A802FF88C583F83B7113F852" bold="true" box="[1272,1290,1981,2005]" pageId="5" pageNumber="50">C</emphasis>
: chin close up with barbel. Photo by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA801FF8BC1ECF9767598F866" ID-DOI="http://doi.org/10.5281/zenodo.4882944" ID-Zenodo-Dep="4882944" httpUri="https://zenodo.org/record/4882944/files/figure.png" pageId="6" pageNumber="51" startId="6.[151,250,1776,1800]" targetBox="[396,1191,181,1753]" targetPageId="6">
<paragraph id="8B9B36C5A801FF8BC1ECF9767598F866" blockId="6.[151,1437,1776,2017]" pageId="6" pageNumber="51">
<emphasis id="B950EAD7A801FF8BC1ECF976750BF88E" bold="true" box="[151,274,1776,1801]" pageId="6" pageNumber="51">FIGURE 7.</emphasis>
<taxonomicName id="4C244D46A801FF8BC063F97775E1F88F" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[280,504,1777,1801]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="6" pageNumber="51" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A801FF8BC063F97775E1F88F" box="[280,504,1777,1801]" italics="true" pageId="6" pageNumber="51">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A801FF8BC085F9777657F88E" bold="true" box="[510,590,1777,1801]" pageId="6" pageNumber="51">
<taxonomicNameLabel id="A26357ACA801FF8BC085F9777657F88E" box="[510,590,1777,1801]" pageId="6" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea.
<emphasis id="B950EAD7A801FF8BC34FF892765FF8AB" bold="true" box="[564,582,1812,1836]" pageId="6" pageNumber="51">A</emphasis>
: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules.
<emphasis id="B950EAD7A801FF8BC367F8BF7634F8D6" bold="true" box="[540,557,1849,1873]" pageId="6" pageNumber="51">B</emphasis>
: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules.
<emphasis id="B950EAD7A801FF8BC399F8DA76EDF8F3" bold="true" box="[738,756,1884,1908]" pageId="6" pageNumber="51">C</emphasis>
: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles.
<emphasis id="B950EAD7A801FF8BC557F8077027F81E" bold="true" box="[1068,1086,1921,1945]" pageId="6" pageNumber="51">D</emphasis>
: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić.
</paragraph>
</caption>
<paragraph id="8B9B36C5A800FF8AC1BCFF11770BFE56" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="52">
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ZM-CBSU 5003-128, female, 30.9+
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ZM-CBSU 5001-1, female, 35.1+8.1,
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ZM-CBSU 5003-60, juvenile female, 25.4+
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ZM-CBSU 5022-23, female, 26.0+5.9,
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ZM-CBSU 5024-1, female, 23.7+
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ZM-CBSU 5003-77, female, 25.1+
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,
<location id="8EFB601EA800FF8AC529FE15715FFE2A" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC529FE15715FFE2A" box="[1106,1350,403,429]" country="Iran" latitude="37.483334" longLatPrecision="1181" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
,
<geoCoordinate id="EE105002A800FF8AC429FE1574B2FE56" degrees="37" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="52" precision="925" value="37.483334">37°29 N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC1C9FE31750EFE56" box="[178,279,439,465]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="52" precision="925" value="49.483334">49°29 E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC058FE31758AFE56" box="[291,403,439,465]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC0CCFE317671FE56" box="[439,616,439,465]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC308FE317717FE56" box="[627,782,439,465]" pageId="7" pageNumber="52" value="2002-11-01">
<collectingDate id="EFDEE9EDA800FF8AC308FE317717FE56" box="[627,782,439,465]" pageId="7" pageNumber="52" value="2002-11-01">01 Nov. 2002</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9B36C5A800FF8AC1BCFE5D7094FD4A" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="52">
<materialsCitation id="3B4C3C98A800FF8AC1BCFE5D7640FD9E" ID-GBIF-Occurrence="3131327309" collectingDate="2005-03-01" collectionCode="PMR" collectorName="K. Abbasi &amp; A. Sarapnah" country="Iran" latitude="37.016666" location="southern Caspian Sea" longLatPrecision="1184" longitude="50.566666" municipality="Chaboksar" pageId="7" pageNumber="52" specimenCode="VP4679" specimenCount="1" stateProvince="Gilan">
<collectionCode id="ED35AE00A800FF8AC1BCFE5D751BFE72" box="[199,258,475,501]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">PMR</collectionCode>
<specimenCode id="DB829EBEA800FF8AC073FE5D757AFE72" box="[264,355,475,501]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">VP4679</specimenCode>
male, 43.3+10.0 mm,
<collectingCountry id="F3337655A800FF8AC327FE5D7695FE72" box="[604,652,475,501]" name="Iran" pageId="7" pageNumber="52">Iran</collectingCountry>
,
<collectingRegion id="49E0F827A800FF8AC3EDFE5D7759FE72" box="[662,832,475,501]" country="Iran" name="Gilan" pageId="7" pageNumber="52">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA800FF8AC230FE5D77DCFE72" box="[843,965,475,501]" pageId="7" pageNumber="52">Chaboksar</collectingMunicipality>
,
<location id="8EFB601EA800FF8AC2B5FE5D70A6FE72" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC2B5FE5D70A6FE72" box="[974,1215,475,501]" country="Iran" latitude="37.016666" longLatPrecision="1184" longitude="50.566666" municipality="Chaboksar" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
,
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<geoCoordinate id="EE105002A800FF8AC44FFE5C718EFE72" box="[1332,1431,474,501]" degrees="50" direction="east" minutes="34" orientation="longitude" pageId="7" pageNumber="52" precision="925" value="50.566666">50°34 E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC1ECFE79751FFD9E" box="[151,262,511,537]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC053FE7975ACFD9E" box="[296,437,511,537]" pageId="7" pageNumber="52">A. Sarapnah</collectorName>
,
<date id="FF9A1005A800FF8AC0BBFE797640FD9E" box="[448,601,510,537]" pageId="7" pageNumber="52" value="2005-03-01">
<collectingDate id="EFDEE9EDA800FF8AC0BBFE797640FD9E" box="[448,601,510,537]" pageId="7" pageNumber="52" value="2005-03-01">01 Mar. 2005</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B4C3C98A800FF8AC318FE79701FFDBA" ID-GBIF-Occurrence="3131327318" collectingDate="2003-03-09" collectionCode="PMR" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" latitude="37.516666" location="southern Caspian Sea" longLatPrecision="1181" longitude="49.5" municipality="Anzali" pageId="7" pageNumber="52" specimenCode="VP4680" specimenCount="1" stateProvince="Gilan">
<collectionCode id="ED35AE00A800FF8AC318FE797687FD9E" box="[611,670,511,537]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">PMR</collectionCode>
<specimenCode id="DB829EBEA800FF8AC3DFFE79771BFD9E" box="[676,770,511,537]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">VP4680</specimenCode>
, male, 47.0+
<quantity id="4CDC9B20A800FF8AC2E8FE7977F0FD9E" box="[915,1001,511,537]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="7" pageNumber="52" unit="mm" value="8.6">8.6 mm</quantity>
,
<collectingCountry id="F3337655A800FF8AC28EFE79703CFD9E" box="[1013,1061,511,537]" name="Iran" pageId="7" pageNumber="52">Iran</collectingCountry>
,
<collectingRegion id="49E0F827A800FF8AC554FE7970C3FD9E" box="[1071,1242,511,537]" country="Iran" name="Gilan" pageId="7" pageNumber="52">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA800FF8AC59FFE797128FD9E" box="[1252,1329,511,537]" pageId="7" pageNumber="52">Anzali</collectingMunicipality>
,
<location id="8EFB601EA800FF8AC447FE79753CFDBA" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC447FE79753CFDBA" country="Iran" latitude="37.516666" longLatPrecision="1181" longitude="49.5" municipality="Anzali" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
,
<geoCoordinate id="EE105002A800FF8AC04AFDA57581FDBA" box="[305,408,547,573]" degrees="37" direction="north" minutes="31" orientation="latitude" pageId="7" pageNumber="52" precision="925" value="37.516666">37°31 N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC0DBFDA5761FFDBA" box="[416,518,547,573]" degrees="49" direction="east" minutes="30" orientation="longitude" pageId="7" pageNumber="52" precision="925" value="49.5">49°30 E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC368FDA5769DFDBA" box="[531,644,547,573]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC3D0FDA57744FDBA" box="[683,861,547,573]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC212FDA5701FFDBA" box="[873,1030,547,573]" pageId="7" pageNumber="52" value="2003-03-09">
<collectingDate id="EFDEE9EDA800FF8AC212FDA5701FFDBA" box="[873,1030,547,573]" pageId="7" pageNumber="52" value="2003-03-09">09 Mar. 2003</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B4C3C98A800FF8AC56AFDA57180FDE6" ID-GBIF-Occurrence="3131327311" collectingDate="2003-03-01" collectionCode="PMR" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" latitude="37.483334" location="southern Caspian Sea" longLatPrecision="1181" longitude="49.483334" municipality="Anzali" pageId="7" pageNumber="52" specimenCode="VP4681" specimenCount="1" stateProvince="Gilan">
<collectionCode id="ED35AE00A800FF8AC56AFDA57055FDBA" box="[1041,1100,547,573]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">PMR</collectionCode>
<specimenCode id="DB829EBEA800FF8AC52FFDA570B6FDBA" box="[1108,1199,547,573]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">VP4681</specimenCode>
male, 35.1+
<quantity id="4CDC9B20A800FF8AC445FDA47181FDBA" box="[1342,1432,546,573]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="7" pageNumber="52" unit="mm" value="8.5">8.5 mm</quantity>
,
<collectingCountry id="F3337655A800FF8AC1ECFDC174DEFDE6" box="[151,199,583,609]" name="Iran" pageId="7" pageNumber="52">Iran</collectingCountry>
,
<collectingRegion id="49E0F827A800FF8AC1A9FDC17565FDE6" box="[210,380,583,609]" country="Iran" name="Gilan" pageId="7" pageNumber="52">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA800FF8AC0FEFDC175CBFDE6" box="[389,466,583,609]" pageId="7" pageNumber="52">Anzali</collectingMunicipality>
,
<location id="8EFB601EA800FF8AC0A6FDC176D6FDE6" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC0A6FDC176D6FDE6" box="[477,719,583,609]" country="Iran" latitude="37.483334" longLatPrecision="1181" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
,
<geoCoordinate id="EE105002A800FF8AC3A1FDC17726FDE6" box="[730,831,583,609]" degrees="37" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="52" precision="925" value="37.483334">37°29 N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC23EFDC177B0FDE6" box="[837,937,583,609]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="52" precision="925" value="49.483334">49°29 E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC2CFFDC1703AFDE6" box="[948,1059,583,609]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC53DFDC170EFFDE6" box="[1094,1270,583,609]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC47AFDC17180FDE6" box="[1281,1433,583,609]" pageId="7" pageNumber="52" value="2003-03-01">
<collectingDate id="EFDEE9EDA800FF8AC47AFDC17180FDE6" box="[1281,1433,583,609]" pageId="7" pageNumber="52" value="2003-03-01">01 Mar. 2003</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B4C3C98A800FF8AC1ECFDED764FFD2E" ID-GBIF-Occurrence="3131327316" collectingDate="2002-09-17" collectionCode="PMR" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" latitude="37.483334" location="southern Caspian Sea" longLatPrecision="1181" longitude="49.483334" municipality="Anzali" pageId="7" pageNumber="52" specimenCode="VP4682" specimenCount="1" stateProvince="Gilan">
<collectionCode id="ED35AE00A800FF8AC1ECFDED74CBFD02" box="[151,210,619,645]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">PMR</collectionCode>
<specimenCode id="DB829EBEA800FF8AC1A1FDED7520FD02" box="[218,313,619,645]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">VP4682</specimenCode>
, female, 30.4+
<quantity id="4CDC9B20A800FF8AC099FDED7625FD02" box="[482,572,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="7" pageNumber="52" unit="mm" value="7.4">7.4 mm</quantity>
,
<collectingCountry id="F3337655A800FF8AC332FDED7660FD02" box="[585,633,619,645]" name="Iran" pageId="7" pageNumber="52">Iran</collectingCountry>
,
<collectingRegion id="49E0F827A800FF8AC3FDFDED772AFD02" box="[646,819,619,645]" country="Iran" name="Gilan" pageId="7" pageNumber="52">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA800FF8AC244FDED7795FD02" box="[831,908,619,645]" pageId="7" pageNumber="52">Anzali</collectingMunicipality>
,
<location id="8EFB601EA800FF8AC2E2FDED7089FD02" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC2E2FDED7089FD02" box="[921,1168,619,645]" country="Iran" latitude="37.483334" longLatPrecision="1181" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
,
<geoCoordinate id="EE105002A800FF8AC5E6FDED711DFD02" box="[1181,1284,619,645]" degrees="37" direction="north" minutes="29" orientation="latitude" pageId="7" pageNumber="52" precision="925" value="37.483334">37°29 N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC476FDED716AFD02" box="[1293,1395,619,645]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="52" precision="925" value="49.483334">49°29 E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC4FAFDED74FFFD2E" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC073FD0975AEFD2E" box="[264,439,655,681]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC0BAFD09764FFD2E" box="[449,598,655,681]" pageId="7" pageNumber="52" value="2002-09-17">
<collectingDate id="EFDEE9EDA800FF8AC0BAFD09764FFD2E" box="[449,598,655,681]" pageId="7" pageNumber="52" value="2002-09-17">17 Sep. 2002</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B4C3C98A800FF8AC325FD097090FD4A" ID-GBIF-Occurrence="3131327306" collectingDate="2004-01-01" collectionCode="PMR" collectorName="K. Abbasi &amp; A. Sarapnah" country="Iran" latitude="37.583332" location="southern Caspian Sea" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" pageId="7" pageNumber="52" specimenCode="VP4683" specimenCount="1" stateProvince="Gilan">
<collectionCode id="ED35AE00A800FF8AC325FD097680FD2E" box="[606,665,655,681]" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">PMR</collectionCode>
<specimenCode id="DB829EBEA800FF8AC3E4FD0976E7FD2E" box="[671,766,655,681]" collectionCode="PMR" country="0" httpUri="http://grbio.org/cool/xq4z-q5gq" name="Prirodoslovni muzej Rijeka" pageId="7" pageNumber="52">VP4683</specimenCode>
, female,
<quantity id="4CDC9B20A800FF8AC21BFD0977DCFD2E" box="[864,965,655,681]" metricMagnitude="-2" metricUnit="m" metricValue="3.42" pageId="7" pageNumber="52" unit="mm" value="34.2">34.2 mm</quantity>
, caudal fin damaged,
<collectingCountry id="F3337655A800FF8AC5CFFD0970FDFD2E" box="[1204,1252,655,681]" name="Iran" pageId="7" pageNumber="52">Iran</collectingCountry>
,
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,
<collectingMunicipality id="6BFFACBFA800FF8AC1ECFD3574FDFD4A" box="[151,228,691,717]" pageId="7" pageNumber="52">Anzali</collectingMunicipality>
,
<location id="8EFB601EA800FF8AC18BFD3575FAFD4A" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC18BFD3575FAFD4A" box="[240,483,691,717]" country="Iran" latitude="37.583332" longLatPrecision="1180" longitude="49.483334" municipality="Anzali" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
,
<geoCoordinate id="EE105002A800FF8AC094FD34764DFD4A" box="[495,596,690,717]" degrees="37" direction="north" minutes="35" orientation="latitude" pageId="7" pageNumber="52" precision="925" value="37.583332">37°35 N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC327FD3576D9FD4A" box="[604,704,691,717]" degrees="49" direction="east" minutes="29" orientation="longitude" pageId="7" pageNumber="52" precision="925" value="49.483334">49°29 E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC3B7FD357725FD4A" box="[716,828,691,717]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC225FD3577F5FD4A" box="[862,1004,691,717]" pageId="7" pageNumber="52">A. Sarapnah</collectorName>
,
<date id="FF9A1005A800FF8AC28CFD357090FD4A" box="[1015,1161,691,717]" pageId="7" pageNumber="52" value="2004-01-01">
<collectingDate id="EFDEE9EDA800FF8AC28CFD357090FD4A" box="[1015,1161,691,717]" pageId="7" pageNumber="52" value="2004-01-01">01 Jan. 2004</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
<paragraph id="8B9B36C5A800FF8AC1BCFD5070E7FC06" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="52">
<emphasis id="B950EAD7A800FF8AC1BCFD5075ACFD77" bold="true" box="[199,437,726,752]" pageId="7" pageNumber="52">Additional material</emphasis>
.
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ZM-CBSU S003-17,
<specimenCount id="9D22FD4CA800FF8AC3C0FD517740FD76" box="[699,857,727,753]" pageId="7" pageNumber="52" type="generic">21 specimens</specimenCount>
,
<quantity id="4CDC9B20A800FF8AC21DFD517012FD76" box="[870,1035,727,753]" metricMagnitude="-2" metricUnit="m" metricValue="3.95" metricValueMax="4.7299999999999995" metricValueMin="3.17" pageId="7" pageNumber="52" unit="mm" value="39.5" valueMax="47.3" valueMin="31.7">31.747.3 mm</quantity>
SL,
<geoCoordinate id="EE105002A800FF8AC53EFD5170E2FD76" box="[1093,1275,727,753]" degrees="37" direction="north" minutes="37" orientation="latitude" pageId="7" pageNumber="52" precision="1" seconds="46.71" value="37.62964">37°3746.71” N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC47FFD5074B3FC92" degrees="49" direction="east" minutes="33" orientation="longitude" pageId="7" pageNumber="52" precision="1" seconds="55.10" value="49.565304">49°3355.10” E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC1C1FD7D7534FC92" box="[186,301,763,789]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC023FD7D7615FC92" box="[344,524,763,789]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC360FD7D76A5FC92" box="[539,700,763,789]" pageId="7" pageNumber="52" value="2003-03-09">
<collectingDate id="EFDEE9EDA800FF8AC360FD7D76A5FC92" box="[539,700,763,789]" pageId="7" pageNumber="52" value="2003-03-09">09 Mar. 2003</collectingDate>
</date>
. ZM-CBSU S003-112113, ZM-CBSU S003-115,
</materialsCitation>
<materialsCitation id="3B4C3C98A800FF8AC473FD7D7519FCDA" ID-GBIF-Occurrence="3131327314" collectingDate="2003-03-09" collectingDateMax="2004-01-10" collectingDateMin="2003-03-09" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" county="All" latitude="37.62964" location="southern Caspian Sea" longLatPrecision="1" longitude="49.565304" municipality="Anzali" pageId="7" pageNumber="52" specimenCount="26" stateProvince="Gilan">
<specimenCount id="9D22FD4CA800FF8AC473FD7D7181FC92" box="[1288,1432,763,789]" pageId="7" pageNumber="52" type="generic">3 specimens</specimenCount>
,
<quantity id="4CDC9B20A800FF8AC1ECFC997523FCBE" box="[151,314,799,825]" metricMagnitude="-2" metricUnit="m" metricValue="3.915" metricValueMax="4.01" metricValueMin="3.82" pageId="7" pageNumber="52" unit="mm" value="39.15" valueMax="40.1" valueMin="38.2">38.240.1 mm</quantity>
SL,
<geoCoordinate id="EE105002A800FF8AC00BFC99763AFCBE" box="[368,547,799,825]" degrees="37" direction="north" minutes="37" orientation="latitude" pageId="7" pageNumber="52" precision="1" seconds="46.71" value="37.62964">37°3746.71” N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC351FC9876C5FCBE" box="[554,732,798,825]" degrees="49" direction="east" minutes="33" orientation="longitude" pageId="7" pageNumber="52" precision="1" seconds="55.10" value="49.565304">49°3355.10” E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC393FC997741FCBE" box="[744,856,799,825]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC206FC997037FCBE" box="[893,1070,799,825]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC542FC9870D5FCBE" box="[1081,1228,798,825]" pageId="7" pageNumber="52" value="2004-01-05">
<collectingDate id="EFDEE9EDA800FF8AC542FC9870D5FCBE" box="[1081,1228,798,825]" pageId="7" pageNumber="52" value="2004-01-05">05 Jan. 2004</collectingDate>
</date>
. ZM-CBSU S003- 134135,
</materialsCitation>
<materialsCitation id="3B4C3C98A800FF8AC073FCC570E3FC06" ID-GBIF-Occurrence="3131327317" collectingDate="2003-03-09" collectingDateMax="2004-01-10" collectingDateMin="2003-03-09" collectorName="K. Abbasi &amp; S. Abdolmaleki" country="Iran" county="All" latitude="37.62964" location="southern Caspian Sea" longLatPrecision="1" longitude="49.565304" municipality="Anzali" pageId="7" pageNumber="52" specimenCount="26" stateProvince="Gilan">
<specimenCount id="9D22FD4CA800FF8AC073FCC5758FFCDA" box="[264,406,835,861]" pageId="7" pageNumber="52" type="generic">2 specimens</specimenCount>
,
<quantity id="4CDC9B20A800FF8AC0DAFCC4765CFCDA" box="[417,581,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="3.715" metricValueMax="4.08" metricValueMin="3.35" pageId="7" pageNumber="52" unit="mm" value="37.15" valueMax="40.8" valueMin="33.5">33.540.8 mm</quantity>
SL,
<geoCoordinate id="EE105002A800FF8AC307FCC57729FCDA" box="[636,816,835,861]" degrees="37" direction="north" minutes="37" orientation="latitude" pageId="7" pageNumber="52" precision="1" seconds="46.71" value="37.62964">37°3746.71” N</geoCoordinate>
<geoCoordinate id="EE105002A800FF8AC243FCC477F2FCDA" box="[824,1003,834,861]" degrees="49" direction="east" minutes="33" orientation="longitude" pageId="7" pageNumber="52" precision="1" seconds="55.10" value="49.565304">49°3355.10” E</geoCoordinate>
,
<collectorName id="26D15313A800FF8AC283FCC57071FCDA" box="[1016,1128,835,861]" pageId="7" pageNumber="52">K. Abbasi</collectorName>
&amp;
<collectorName id="26D15313A800FF8AC5F5FCC57159FCDA" box="[1166,1344,835,861]" pageId="7" pageNumber="52">S. Abdolmaleki</collectorName>
,
<date id="FF9A1005A800FF8AC437FCC574CAFC06" pageId="7" pageNumber="52" value="2004-01-10">
<collectingDate id="EFDEE9EDA800FF8AC437FCC574CAFC06" pageId="7" pageNumber="52" value="2004-01-10">10 Jan. 2004</collectingDate>
</date>
.
<collectingCounty id="62FA4E49A800FF8AC1A7FCE17519FC06" box="[220,256,871,897]" pageId="7" pageNumber="52">All</collectingCounty>
additional material was collected from
<collectingCountry id="F3337655A800FF8AC3C7FCE176F5FC06" box="[700,748,871,897]" name="Iran" pageId="7" pageNumber="52">Iran</collectingCountry>
,
<collectingRegion id="49E0F827A800FF8AC38CFCE177BAFC06" box="[759,931,871,897]" country="Iran" name="Gilan" pageId="7" pageNumber="52">Gilan Province</collectingRegion>
,
<collectingMunicipality id="6BFFACBFA800FF8AC2D7FCE177E0FC06" box="[940,1017,871,897]" pageId="7" pageNumber="52">Anzali</collectingMunicipality>
,
<location id="8EFB601EA800FF8AC57EFCE170E3FC06" LSID="urn:lsid:plazi:treatment:038D87D3A805FF9DC1ECF994718BFE2A:8EFB601EA800FF8AC57EFCE170E3FC06" box="[1029,1274,871,897]" country="Iran" county="All" latitude="37.62964" longLatPrecision="1" longitude="49.565304" municipality="Anzali" name="southern Caspian Sea" pageId="7" pageNumber="52" stateProvince="Gilan">southern Caspian Sea</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E654EA800FF8AC1BCFC0C711CFAFE" pageId="7" pageNumber="52" type="diagnosis">
<paragraph id="8B9B36C5A800FF8AC1BCFC0C711CFAFE" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="52">
<emphasis id="B950EAD7A800FF8AC1BCFC0C7521FC23" bold="true" box="[199,312,906,932]" pageId="7" pageNumber="52">Diagnosis</emphasis>
.
<taxonomicName id="4C244D46A800FF8AC03CFC0D7622FC23" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[327,571,907,933]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A800FF8AC03CFC0D7622FC23" box="[327,571,907,933]" italics="true" pageId="7" pageNumber="52">Benthophilus persicus</emphasis>
</taxonomicName>
is distinguished from all other congeneric species by: (1) dermal fold behind jaws well-developed, large, rectangular, (2) chin barbel of moderate size, 1/32/3 of eye diameter, (3) maximum body width 15.122.9% of SL, (4) mouth width 36.355.8% of head length, (5) second dorsal fin I+78; (6) origin of anal fin in front of vertical through origin of second dorsal fin, (7) dermal tubercles present, clearly larger than granules, with two posterior rows of spinules forming an acute, always less than right angle, (8) dorsal row of tubercles complete, 22 29, (9) ventral row of tubercles 2225, (10) ventrolateral row of tubercles absent, (11) tubercles not present on temporal and occipital head regions, (12) granules not present on flanks, (13) transversal suborbital row
<emphasis id="B950EAD7A800FF8AC5DDFBE470A2FBFB" box="[1190,1211,1122,1148]" italics="true" pageId="7" pageNumber="52">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A800FF8AC19EFB0174EAFB26" box="[229,243,1159,1185]" italics="true" pageId="7" pageNumber="52">b</emphasis>
, (14) anterior interorbital transversal row
<emphasis id="B950EAD7A800FF8AC3C7FB0176C1FB27" box="[700,728,1159,1184]" italics="true" pageId="7" pageNumber="52">pa</emphasis>
with one or two papillae and anterior interorbital transversal row
<emphasis id="B950EAD7A800FF8AC1ECFB2D74AAFB43" box="[151,179,1195,1220]" italics="true" pageId="7" pageNumber="52">pp</emphasis>
with two or three papillae, and (15) body with 2022 transversal
<emphasis id="B950EAD7A800FF8AC2FBFB2D77BAFB42" box="[896,931,1195,1221]" italics="true" pageId="7" pageNumber="52">ltm</emphasis>
rows starting anteriorly behind pectoral axilla and alternating anteriorly with three longitudinal
<emphasis id="B950EAD7A800FF8AC3CEFB4976C1FB6E" box="[693,728,1231,1257]" italics="true" pageId="7" pageNumber="52">llm</emphasis>
rows (characters presented in the order of appearance in the description). Each of the selected diagnostic characters differentiate the new species from between 4 to 15
<taxonomicName id="4C244D46A800FF8AC477FB7574F7FAB6" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="B950EAD7A800FF8AC477FB757184FA8A" box="[1292,1437,1267,1293]" italics="true" pageId="7" pageNumber="52">Benthophilus</emphasis>
species.
</taxonomicName>
Considering only these selected fifteen diagnostic characters, the new species differs from congeneric species in a range from at least five characters (from
<taxonomicName id="4C244D46A800FF8AC3F4FABD7042FAD2" authority="Boldyrev &amp; Bogutskaya, 2004" authorityName="Boldyrev &amp; Bogutskaya" authorityYear="2004" box="[655,1115,1339,1365]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="durrelli">
<emphasis id="B950EAD7A800FF8AC3F4FABD7719FAD2" box="[655,768,1339,1365]" italics="true" pageId="7" pageNumber="52">B. durrelli</emphasis>
<bibRefCitation id="EFB54B34A800FF8AC272FABD7042FAD2" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." box="[777,1115,1339,1365]" pageId="7" pageNumber="52" pagination="129 - 135" refId="ref11803" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2004) Description of two new species of tadpole gobies (Teleostei: Gobiidae: Benthophilus). Zoosystematica Rossica, 13 (1), 129 - 135." type="journal article" year="2004">Boldyrev &amp; Bogutskaya, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C244D46A800FF8AC512FABD74D7FAFE" authority="Ragimov, 1976" authorityName="Ragimov" authorityYear="1976" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="mahmudbejovi">
<emphasis id="B950EAD7A800FF8AC512FABD7132FAD2" box="[1129,1323,1339,1365]" italics="true" pageId="7" pageNumber="52">B. mahmudbejovi</emphasis>
Ragimov, 1976
</taxonomicName>
and
<taxonomicName id="4C244D46A800FF8AC078FAD97636FAFE" authority="Ragimov, 1982" authorityName="Ragimov" authorityYear="1982" box="[259,559,1375,1401]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="pinchuki">
<emphasis id="B950EAD7A800FF8AC078FAD97566FAFE" box="[259,383,1375,1401]" italics="true" pageId="7" pageNumber="52">B. pinchuki</emphasis>
Ragimov, 1982
</taxonomicName>
) up to eleven differential characters (from
<taxonomicName id="4C244D46A800FF8AC578FAD970E0FAFE" authority="Berg, 1927" authorityName="Berg" authorityYear="1927" box="[1027,1273,1375,1401]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="7" pageNumber="52" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="B950EAD7A800FF8AC578FAD9706CFAFE" box="[1027,1141,1375,1401]" italics="true" pageId="7" pageNumber="52">B. kessleri</emphasis>
Berg, 1927
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C33E654EA800FF84C1B4FA0474E6FC6A" lastPageId="9" lastPageNumber="54" pageId="7" pageNumber="52" type="description">
<paragraph id="8B9B36C5A800FF8AC1B4FA0475CDF98E" blockId="7.[151,1437,150,2013]" pageId="7" pageNumber="52">
<emphasis id="B950EAD7A800FF8AC1B4FA047540FA1B" bold="true" box="[207,345,1410,1436]" pageId="7" pageNumber="52">Description</emphasis>
(
<figureCitation id="131F2A40A800FF8AC012FA0575B6FA1A" box="[361,431,1411,1437]" captionStart="FIGURE 4" captionStartId="4.[151,250,726,750]" captionTargetBox="[248,1340,181,702]" captionTargetId="figure-33@4.[248,1340,181,702]" captionTargetPageId="4" captionText="FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882938" httpUri="https://zenodo.org/record/4882938/files/figure.png" pageId="7" pageNumber="52">Fig. 4</figureCitation>
). All morphometric and meristic values in the text are presented first for the
<typeStatus id="549F8867A800FF8AC469FA05716DFA1A" box="[1298,1396,1411,1437]" pageId="7" pageNumber="52" type="holotype">holotype</typeStatus>
for the
<typeStatus id="549F8867A800FF8AC1BFFA217536FA46" box="[196,303,1447,1473]" pageId="7" pageNumber="52" type="paratype">paratypes</typeStatus>
in parentheses. Morphometric data are provided in
<tableCitation id="C6A6037EA800FF8AC2F7FA2177FAFA46" box="[908,995,1447,1473]" captionStart="TABLE 1" captionStartId="13.[151,239,477,503]" captionTargetBox="[168,2025,656,1650]" captionText="TABLE 1. Benthophilus persicus sp. nov. Morphometric characters as % of standard length or as % of head length. Characters are sorted as in Boldyrev &amp; Bogutskaya (2007) with the new introduced characters added at the end in alphabetic order. The morphometric character and the related values of dimorphic characters with non-overlapping ranges of groups marked with ^ for adult males vs. the immature male and all females, with * for all males and the large female vs. small mature and immature females, and with ˜ for adult males and the large female vs. smaller females and the immature male, see Discussion." httpUri="http://table.plazi.org/id/DF5B664DA80AFF80C1ECFE5B7068FDE4" pageId="7" pageNumber="52" tableUuid="DF5B664DA80AFF80C1ECFE5B7068FDE4">Table 1</tableCitation>
. Particularly large variability of some features may be based on sex, developmental stage or size dimorphism and is highlighted in the
<tableCitation id="C6A6037EA800FF8AC5B3FA4D7103FA62" box="[1224,1306,1483,1509]" captionStart="TABLE 1" captionStartId="13.[151,239,477,503]" captionTargetBox="[168,2025,656,1650]" captionText="TABLE 1. Benthophilus persicus sp. nov. Morphometric characters as % of standard length or as % of head length. Characters are sorted as in Boldyrev &amp; Bogutskaya (2007) with the new introduced characters added at the end in alphabetic order. The morphometric character and the related values of dimorphic characters with non-overlapping ranges of groups marked with ^ for adult males vs. the immature male and all females, with * for all males and the large female vs. small mature and immature females, and with ˜ for adult males and the large female vs. smaller females and the immature male, see Discussion." httpUri="http://table.plazi.org/id/DF5B664DA80AFF80C1ECFE5B7068FDE4" pageId="7" pageNumber="52" tableUuid="DF5B664DA80AFF80C1ECFE5B7068FDE4">Table 1</tableCitation>
and further explained in the Discussion.
</paragraph>
<paragraph id="8B9B36C5A800FF85C1BCF9957178FEC6" blockId="7.[151,1437,150,2013]" lastBlockId="8.[151,1437,151,2014]" lastPageId="8" lastPageNumber="53" pageId="7" pageNumber="52">
<emphasis id="B950EAD7A800FF8AC1BCF99575A3F9AA" box="[199,442,1555,1581]" italics="true" pageId="7" pageNumber="52">General morphology.</emphasis>
Head rounded in vertical view, i.e., triangular with well-rounded lateral sides (
<figureCitation id="131F2A40A800FF8AC43CF9957197F9AA" box="[1351,1422,1554,1581]" captionStart="FIGURE 5" captionStartId="4.[151,250,1857,1881]" captionTargetBox="[248,1339,882,1824]" captionTargetId="figure-102@4.[248,1339,882,1833]" captionTargetPageId="4" captionText="FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882940" httpUri="https://zenodo.org/record/4882940/files/figure.png" pageId="7" pageNumber="52">Fig. 5</figureCitation>
). Head large, long and wide, length 2.8 (2.63.0) in SL, width 1.0 (1.01.2) in head length. Head depressed (dorsoventrally compressed), head depth in head width 1.9 (1.72.2), head depth 1.8 (1.62.4) in head length, wider than body, head width 0.7 (0.50.6) in maximum body width. Snout gently rounded, broad and moderately long, larger than eye diameter, 0.5 (0.60.8) in eye diameter, 3.6 (3.64.2) in head length. Eye small, horizontal diameter 7.1 (4.67.1) in head length. Interorbital distance 6.8 (5.210.9) in head length. Eye diameter and interorbital width both size-dependent, i.e., eye diameter negatively correlated and interorbital distance positively correlated with the body size. Dermal fold behind end of jaws well-developed, large, rectangular, elongate (
<figureCitation id="131F2A40A800FF8AC5FBF889712AF8AD" box="[1152,1331,1807,1834]" captionStart="FIGURE 6" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[315,1272,181,1885]" captionTargetId="figure-20@5.[315,1272,181,1885]" captionTargetPageId="5" captionText="FIGURE 6. Benthophilus persicus sp. nov. Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea: A: lateral detail of head, B: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian: C: chin close up with barbel. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882942" httpUri="https://zenodo.org/record/4882942/files/figure.png" pageId="7" pageNumber="52">Figs. 6a and 6b</figureCitation>
). Dermal fold depth in length 2.4 (2.53.7), with more or less rounded angles, along edge undulate or straight. Dermal fold of variable size to eye diameter, length of its base 0.9 (0.81.6) in eye diameter, 6.0 (4.98.2) in head length. Chin barbel of moderate size, 0.6 (0.40.7) in eye diameter, 12.0 (10.413.7) in head length, triangular, widened at base, triangle narrower in larger specimens, broader in smaller fish (
<figureCitation id="131F2A40A800FF8AC230F81977EEF83D" box="[843,1015,1951,1978]" captionStart="FIGURE 6" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[315,1272,181,1885]" captionTargetId="figure-20@5.[315,1272,181,1885]" captionTargetPageId="5" captionText="FIGURE 6. Benthophilus persicus sp. nov. Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea: A: lateral detail of head, B: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian: C: chin close up with barbel. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882942" httpUri="https://zenodo.org/record/4882942/files/figure.png" pageId="7" pageNumber="52">Figs. 6b and 6c</figureCitation>
). Mouth relatively wide, mouth width 2.0 (1.82.8) in head length. Mouth corner below anterior eye margin. Anterior nostril tube without process from rim, reaching upper lip; posterior nostril with raised rim. No medial groove present on temporal and occipital head regions. Body deepest at first dorsal-fin origin or slightly in front of it, depth decreasing towards caudal-fin base. Greatest body width at middle between pectoral-fin bases, 4.4 (4.46.6) in SL, strongly decreasing towards caudalfin base. Caudal peduncle laterally compressed, caudal peduncle width 1.3 (
<quantity id="4CDC9B20A80FFF85C57BFE84706DFE9A" box="[1024,1140,258,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.429" metricValueMax="3.81" metricValueMin="3.048" pageId="8" pageNumber="53" unit="in" value="1.35" valueMax="1.5" valueMin="1.2">1.21.5 in</quantity>
depth), shallow, its depth 17.4 (15.019.3) in SL, and narrow, its width 22.2 (20.225.9) in SL. Maximum size
<quantity id="4CDC9B20A80FFF85C528FEA070AEFEC6" box="[1107,1207,294,321]" metricMagnitude="-2" metricUnit="m" metricValue="5.56" pageId="8" pageNumber="53" unit="mm" value="55.6">55.6 mm</quantity>
in total length.
</paragraph>
<paragraph id="8B9B36C5A80FFF85C1BCFECD70F5FD2E" blockId="8.[151,1437,151,2014]" pageId="8" pageNumber="53">
<emphasis id="B950EAD7A80FFF85C1BCFECD7519FEE3" box="[199,256,331,356]" italics="true" pageId="8" pageNumber="53">Fins.</emphasis>
The poor condition of fins in some cases prevents exact counts. D1 IV (III: 1, IV: 14), D2 I+7 (I+7: 7, I+8: 7; in
<typeStatus id="549F8867A80FFF85C1AFFEE9752DFE0E" box="[212,308,367,393]" pageId="8" pageNumber="53" type="paratype">paratype</typeStatus>
ZM-CBSU 5001-1 positive count was not possible), A I+8 (I+7: 6, I+8: 8; in
<typeStatus id="549F8867A80FFF85C5CCFEE9710EFE0E" box="[1207,1303,367,393]" pageId="8" pageNumber="53" type="paratype">paratype</typeStatus>
ZM-CBSU 5001-1 positive count was not possible), C branched rays 10 (10: 7, 11: 3, 12: 2; in
<specimenCount id="9D22FD4CA80FFF85C537FE1570EEFE2A" box="[1100,1271,403,429]" pageId="8" pageNumber="53" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
count was not possible), segmented 13 (13: 12; in
<specimenCount id="9D22FD4CA80FFF85C352FE3176CAFE56" box="[553,723,439,465]" pageId="8" pageNumber="53" type="generic" typeStatus="paratypes">three paratypes</specimenCount>
count was not possible), P 16 on both sides (16: 20, 17: 9, both sides counted; in
<typeStatus id="549F8867A80FFF85C026FE5D75A4FE72" box="[349,445,475,501]" pageId="8" pageNumber="53" type="paratype">paratype</typeStatus>
PMR VP4681 positive count was not possible on left side), V I+5/5+I (V I+5/5+I: 15). First dorsal fin low, its height 11.7 (
<quantity id="4CDC9B20A80FFF85C35EFE7876ABFD9E" box="[549,690,510,537]" metricMagnitude="-1" metricUnit="m" metricValue="2.8447999999999998" metricValueMax="3.0226" metricValueMin="2.667" pageId="8" pageNumber="53" unit="in" value="11.2" valueMax="11.9" valueMin="10.5">10.511.9 in</quantity>
adult males,
<quantity id="4CDC9B20A80FFF85C23CFE7877CAFD9E" box="[839,979,510,537]" metricMagnitude="-1" metricUnit="m" metricValue="4.4196" metricValueMax="5.4102" metricValueMin="3.429" pageId="8" pageNumber="53" unit="in" value="17.4" valueMax="21.3" valueMin="13.5">13.521.3 in</quantity>
other individuals) in SL, lower than second dorsal fin, 7.8 (6.58.8) in SL. Origin of anal fin in front of vertical through origin of second dorsal fin. Anal-fin height 9.7 (8.410.4) in SL. Pectoral fin long, reaching backwards halfway between the first and second dorsal fins when folded back. Pelvic disc complete and oval with well-developed anterior membrane, anterior membrane with straight edge. Pelvic disc long, 3.2 (3.13.8) in SL, reaching to anal-fin origin. Caudal fin rounded.
</paragraph>
<paragraph id="8B9B36C5A80FFF85C1BCFD3576E4FAFE" blockId="8.[151,1437,151,2014]" pageId="8" pageNumber="53">
<emphasis id="B950EAD7A80FFF85C1BCFD3575A1FD4A" box="[199,440,691,717]" italics="true" pageId="8" pageNumber="53">Dermal ossifications.</emphasis>
Granules present only on head (on snout, between eyes, on temporal and occipital head regions, and around eye in a circle), on predorsal area and between dorsal and dorsolateral rows of tubercles below the first dorsal fin and below interdorsal space (
<figureCitation id="131F2A40A80FFF85C3DCFD7D76EDFC92" box="[679,756,763,789]" captionStart="FIGURE 3" captionStartId="3.[151,250,1907,1931]" captionTargetBox="[225,1361,1122,1883]" captionTargetId="figure-79@3.[225,1361,1122,1883]" captionTargetPageId="3" captionText="FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882936" httpUri="https://zenodo.org/record/4882936/files/figure.png" pageId="8" pageNumber="53">Figs. 3</figureCitation>
and
<figureCitation id="131F2A40A80FFF85C252FD7C7720FC92" box="[809,825,762,789]" captionStart="FIGURE 5" captionStartId="4.[151,250,1857,1881]" captionTargetBox="[248,1339,882,1824]" captionTargetId="figure-102@4.[248,1339,882,1833]" captionTargetPageId="4" captionText="FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882940" httpUri="https://zenodo.org/record/4882940/files/figure.png" pageId="8" pageNumber="53">5</figureCitation>
). No granules on eyes. No granules on gill covers, dorsal body and posterior flank, around pelvic disc, and only a few granules on posterior dorsal part of caudal peduncle (
<figureCitation id="131F2A40A80FFF85C1DBFCC574FCFCDA" box="[160,229,835,861]" captionStart="FIGURE 7" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[396,1191,181,1753]" captionTargetId="figure-33@6.[396,1191,181,1753]" captionTargetPageId="6" captionText="FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882944" httpUri="https://zenodo.org/record/4882944/files/figure.png" pageId="8" pageNumber="53">Fig. 7</figureCitation>
). Granules tiny, simple-structured bumps, sometimes grouped two or three together, small specimens with comparatively large granules (
<figureCitation id="131F2A40A80FFF85C090FCE17621FC06" box="[491,568,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,966,990]" captionTargetBox="[225,1361,181,942]" captionTargetId="figure-20@3.[225,1361,181,942]" captionTargetPageId="3" captionText="FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882934" httpUri="https://zenodo.org/record/4882934/files/figure.png" pageId="8" pageNumber="53">Figs. 2</figureCitation>
and
<figureCitation id="131F2A40A80FFF85C314FCE17666FC06" box="[623,639,871,897]" captionStart="FIGURE 3" captionStartId="3.[151,250,1907,1931]" captionTargetBox="[225,1361,1122,1883]" captionTargetId="figure-79@3.[225,1361,1122,1883]" captionTargetPageId="3" captionText="FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882936" httpUri="https://zenodo.org/record/4882936/files/figure.png" pageId="8" pageNumber="53">3</figureCitation>
). Tubercles distinctively larger than granules (
<figureCitation id="131F2A40A80FFF85C5FDFCE170CDFC06" box="[1158,1236,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,966,990]" captionTargetBox="[225,1361,181,942]" captionTargetId="figure-20@3.[225,1361,181,942]" captionTargetPageId="3" captionText="FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882934" httpUri="https://zenodo.org/record/4882934/files/figure.png" pageId="8" pageNumber="53">Figs. 2</figureCitation>
and
<figureCitation id="131F2A40A80FFF85C471FCE17103FC06" box="[1290,1306,871,897]" captionStart="FIGURE 3" captionStartId="3.[151,250,1907,1931]" captionTargetBox="[225,1361,1122,1883]" captionTargetId="figure-79@3.[225,1361,1122,1883]" captionTargetPageId="3" captionText="FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882936" httpUri="https://zenodo.org/record/4882936/files/figure.png" pageId="8" pageNumber="53">3</figureCitation>
). Tubercles on head appear more or less randomly scattered. Head with a few small tubercles on snout, 24 tubercles present between eyes and 1012 well-developed tubercles on upper cheek, preopercle and opercle (
<figureCitation id="131F2A40A80FFF85C500FC2970D1FC4E" box="[1147,1224,942,969]" captionStart="FIGURE 5" captionStartId="4.[151,250,1857,1881]" captionTargetBox="[248,1339,882,1824]" captionTargetId="figure-102@4.[248,1339,882,1833]" captionTargetPageId="4" captionText="FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882940" httpUri="https://zenodo.org/record/4882940/files/figure.png" pageId="8" pageNumber="53">Figs. 5</figureCitation>
and
<figureCitation id="131F2A40A80FFF85C585FC297102FC4E" box="[1278,1307,943,969]" captionStart="FIGURE 7" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[396,1191,181,1753]" captionTargetId="figure-33@6.[396,1191,181,1753]" captionTargetPageId="6" captionText="FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882944" httpUri="https://zenodo.org/record/4882944/files/figure.png" pageId="8" pageNumber="53">7a</figureCitation>
). Tubercles not present on temporal and occipital head regions, i.e. all dermal ossifications are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e. anteriorly to rear edge of eyes (
<figureCitation id="131F2A40A80FFF85C0FEFB9D75D3FBB2" box="[389,458,1050,1077]" captionStart="FIGURE 5" captionStartId="4.[151,250,1857,1881]" captionTargetBox="[248,1339,882,1824]" captionTargetId="figure-102@4.[248,1339,882,1833]" captionTargetPageId="4" captionText="FIGURE 5. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on dorsal view of head, TCO - tubercles along upper cheek, preopercle and opercle, TI - interorbital tubercles, G - granules. Tubercles not present on temporal and occipital head regions, i.e., all dermal ossifications there are definable as granules according to size and shape; and the first tubercles anteriorly to temporal region are located at interorbital space, i.e., anteriorly to rear edge of eyes. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882940" httpUri="https://zenodo.org/record/4882940/files/figure.png" pageId="8" pageNumber="53">Fig. 5</figureCitation>
). Tubercles on trunk are arranged in longitudinal rows: dorsal, dorsolateral and ventral rows (
<figureCitation id="131F2A40A80FFF85C1A4FBB97534FBDD" box="[223,301,1087,1114]" captionStart="FIGURE 7" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[396,1191,181,1753]" captionTargetId="figure-33@6.[396,1191,181,1753]" captionTargetPageId="6" captionText="FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882944" httpUri="https://zenodo.org/record/4882944/files/figure.png" pageId="8" pageNumber="53">Figs. 7</figureCitation>
b-d). Dorsal row complete, with 2229 tubercles, including 24 tubercles in front of the first dorsal fin. Several anterior tubercles of dorsal row in front and along the first dorsal fin smaller than remaining tubercles and with one radial row of spinules instead of two. Dorsolateral row with 2026 well developed tubercles, starting above pectoral-fin base, ending at posterior part or end of D2, decreasing in size posteriorly. No ventrolateral row (
<figureCitation id="131F2A40A80FFF85C1DBFB4974EDFB6E" box="[160,244,1231,1257]" captionStart="FIGURE 7" captionStartId="6.[151,250,1776,1800]" captionTargetBox="[396,1191,181,1753]" captionTargetId="figure-33@6.[396,1191,181,1753]" captionTargetPageId="6" captionText="FIGURE 7. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. A: Dermal ossifications on lateral view of head, TCO - tubercles along upper cheek, preopercle and opercle, G - granules. B: Dermal ossifications on posterior dorsal body view: D - dorsal row of tubercles, DL - dorsolateral row of tubercles, T - tubercle, G - granules. C: Dermal ossifications on anterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. D: Dermal ossifications on posterior lateral body view, D - dorsal row of tubercles, DL - dorsolateral row of tubercles, V - ventral row of tubercles, T - tubercles. Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882944" httpUri="https://zenodo.org/record/4882944/files/figure.png" pageId="8" pageNumber="53">Fig. 7c</figureCitation>
). Ventral row anteriorly curved upwards with 23 tubercles above others, 2225 tubercles including the 23 anterior upper tubercles. Tubercle bodies poorly defined, dominated by spinules. Tubercles of body rows and of head possess two posterior rows of spinules forming an acute angle, always less than right angle. Exceptions are the anterior tubercles of dorsal row, with one radial row of spinules instead of two (
<figureCitation id="131F2A40A80FFF85C574FABD704BFAD2" box="[1039,1106,1339,1365]" captionStart="FIGURE 3" captionStartId="3.[151,250,1907,1931]" captionTargetBox="[225,1361,1122,1883]" captionTargetId="figure-79@3.[225,1361,1122,1883]" captionTargetPageId="3" captionText="FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882936" httpUri="https://zenodo.org/record/4882936/files/figure.png" pageId="8" pageNumber="53">Fig. 3</figureCitation>
), and tubercles on preopercle and opercle, where spinules look disorganised (
<figureCitation id="131F2A40A80FFF85C3D1FAD976E9FAFE" box="[682,752,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,966,990]" captionTargetBox="[225,1361,181,942]" captionTargetId="figure-20@3.[225,1361,181,942]" captionTargetPageId="3" captionText="FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882934" httpUri="https://zenodo.org/record/4882934/files/figure.png" pageId="8" pageNumber="53">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8B9B36C5A80FFF84C1BCFA0577C9FDBA" blockId="8.[151,1437,151,2014]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="54" pageId="8" pageNumber="53">
<emphasis id="B950EAD7A80FFF85C1BCFA0575B8FA1B" box="[199,417,1411,1437]" italics="true" pageId="8" pageNumber="53">Lateral line system</emphasis>
(
<figureCitation id="131F2A40A80FFF85C0C9FA0575E1FA1A" box="[434,504,1411,1437]" captionStart="FIGURE 8" captionStartId="10.[151,250,798,822]" captionTargetBox="[262,1333,191,774]" captionTargetId="figure-33@10.[254,1333,181,774]" captionTargetPageId="10" captionText="FIGURE 8. Head sensory papillae pattern of lateral line system of Benthophilus persicus sp. nov. Paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Terminology in text. Drawing by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882946" httpUri="https://zenodo.org/record/4882946/files/figure.png" pageId="8" pageNumber="53">Fig. 8</figureCitation>
). No head canals present. Number of papillae in rows are strongly specimen size depending, with larger rows of sensory papillae irregularly doubled or tripled in larger specimens (e.g., suborbital transversal rows or row
<emphasis id="B950EAD7A80FFF85C0DEFA4D75A2FA63" box="[421,443,1483,1508]" italics="true" pageId="8" pageNumber="53">ot</emphasis>
) (
<figureCitation id="131F2A40A80FFF85C0A8FA4D7639FA62" box="[467,544,1483,1509]" captionStart="FIGURE 2" captionStartId="3.[151,250,966,990]" captionTargetBox="[225,1361,181,942]" captionTargetId="figure-20@3.[225,1361,181,942]" captionTargetPageId="3" captionText="FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882934" httpUri="https://zenodo.org/record/4882934/files/figure.png" pageId="8" pageNumber="53">Figs. 2</figureCitation>
and
<figureCitation id="131F2A40A80FFF85C32DFA4D766DFA61" box="[598,628,1483,1510]" captionStart="FIGURE 6" captionStartId="5.[151,250,1909,1933]" captionTargetBox="[315,1272,181,1885]" captionTargetId="figure-20@5.[315,1272,181,1885]" captionTargetPageId="5" captionText="FIGURE 6. Benthophilus persicus sp. nov. Dermal fold (DF) and chin barbell (CB). Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea: A: lateral detail of head, B: ventral view of head. Reversibly stained holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian: C: chin close up with barbel. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882942" httpUri="https://zenodo.org/record/4882942/files/figure.png" pageId="8" pageNumber="53">6a</figureCitation>
). Some rows or parts of rows as ridges with papillae along top, e.g., row
<emphasis id="B950EAD7A80FFF85C1ECFA6974BAF98F" box="[151,163,1519,1544]" italics="true" pageId="8" pageNumber="53">e</emphasis>
(
<figureCitation id="131F2A40A80FFF85C1C8FA6974EEF98E" box="[179,247,1519,1545]" captionStart="FIGURE 9" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[265,1323,882,1905]" captionTargetId="figure-71@10.[265,1323,882,1905]" captionTargetPageId="10" captionText="FIGURE 9. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, anterior part of preoperculo-mandibular rows in PMR VP4680, male, 47.0+8.6 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Terminology in text. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882948" httpUri="https://zenodo.org/record/4882948/files/figure.png" pageId="8" pageNumber="53">Fig. 9</figureCitation>
). Rows with range of number of sensory papillae in parentheses as follows: (1)
<emphasis id="B950EAD7A80FFF85C509FA6970FAF98E" box="[1138,1251,1519,1545]" italics="true" pageId="8" pageNumber="53">preorbital</emphasis>
: snout with four median preorbital series, vertical row
<emphasis id="B950EAD7A80FFF85C333F995764AF9AB" box="[584,595,1555,1580]" italics="true" pageId="8" pageNumber="53">r</emphasis>
(35) slightly above horizontal level of posterior nares, horizontal row
<emphasis id="B950EAD7A80FFF85C4F2F9957185F9A7" box="[1417,1436,1554,1580]" italics="true" pageId="8" pageNumber="53">
s
<superScript id="7C519B8DA80FFF85C4EFF9947185F9A7" attach="left" box="[1428,1436,1554,1568]" fontSize="6" pageId="8" pageNumber="53">1</superScript>
</emphasis>
(35) below horizontal level of posterior nares, horizontal row
<emphasis id="B950EAD7A80FFF85C212F9B17765F9C3" box="[873,892,1590,1616]" italics="true" pageId="8" pageNumber="53">
s
<superScript id="7C519B8DA80FFF85C20FF9B07765F9C3" attach="left" box="[884,892,1590,1604]" fontSize="6" pageId="8" pageNumber="53">2</superScript>
</emphasis>
(34) n the level of anterior nares and below
<emphasis id="B950EAD7A80FFF85C4F2F9B17185F9C3" box="[1417,1436,1590,1616]" italics="true" pageId="8" pageNumber="53">
s
<superScript id="7C519B8DA80FFF85C4EFF9B07185F9C3" attach="left" box="[1428,1436,1590,1604]" fontSize="6" pageId="8" pageNumber="53">1</superScript>
</emphasis>
and vertical
<emphasis id="B950EAD7A80FFF85C05BF9DD752AF9EF" box="[288,307,1626,1652]" italics="true" pageId="8" pageNumber="53">
s
<superScript id="7C519B8DA80FFF85C050F9DC752AF9EF" attach="left" box="[299,307,1626,1640]" fontSize="6" pageId="8" pageNumber="53">3</superScript>
</emphasis>
(24) more medially above upper lip. Lateral series
<emphasis id="B950EAD7A80FFF85C202F9DD779CF9F3" box="[889,901,1627,1652]" italics="true" pageId="8" pageNumber="53">c</emphasis>
in four parts: superior
<emphasis id="B950EAD7A80FFF85C5FFF9DD7080F9EF" box="[1156,1177,1626,1652]" italics="true" pageId="8" pageNumber="53">
c
<superScript id="7C519B8DA80FFF85C5EAF9DC7080F9EF" attach="left" box="[1169,1177,1626,1640]" fontSize="6" pageId="8" pageNumber="53">2</superScript>
</emphasis>
as two horizontal rows between anterior and posterior nostril (2+4 4+8); middle
<emphasis id="B950EAD7A80FFF85C24BF9F9775DF90B" box="[816,836,1662,1688]" italics="true" pageId="8" pageNumber="53">
c
<superScript id="7C519B8DA80FFF85C247F9F8775DF90B" attach="left" box="[828,836,1662,1676]" fontSize="6" pageId="8" pageNumber="53">1</superScript>
</emphasis>
(36) starting at anterior nostril; inferior rows, upper horizontal
<emphasis id="B950EAD7A80FFF85C06AF925753CF944" box="[273,293,1699,1731]" italics="true" pageId="8" pageNumber="53">
c
<subScript id="17A03480A80FFF85C066F933753CF944" attach="left" box="[285,293,1717,1731]" fontSize="6" pageId="8" pageNumber="53">2</subScript>
</emphasis>
(510) and lower horizontal
<emphasis id="B950EAD7A80FFF85C308F925769EF944" box="[627,647,1699,1731]" italics="true" pageId="8" pageNumber="53">
c
<subScript id="17A03480A80FFF85C304F933769EF944" attach="left" box="[639,647,1717,1731]" fontSize="6" pageId="8" pageNumber="53">1</subScript>
</emphasis>
(36) starting anteriorly at upper lip. (2)
<emphasis id="B950EAD7A80FFF85C51BF92570CAF93A" box="[1120,1235,1699,1725]" italics="true" pageId="8" pageNumber="53">suborbital</emphasis>
: seven transverse suborbital rows (
<emphasis id="B950EAD7A80FFF85C021F941759DF966" box="[346,388,1735,1761]" italics="true" pageId="8" pageNumber="53">17</emphasis>
) of sensory papillae: rows
<emphasis id="B950EAD7A80FFF85C3B9F94176F5F966" box="[706,748,1735,1761]" italics="true" pageId="8" pageNumber="53">14</emphasis>
begin distant from orbit, row
<emphasis id="B950EAD7A80FFF85C536F9417042F966" box="[1101,1115,1735,1761]" italics="true" pageId="8" pageNumber="53">4</emphasis>
from anterior end of row
<emphasis id="B950EAD7A80FFF85C4F5F9417185F966" box="[1422,1436,1735,1761]" italics="true" pageId="8" pageNumber="53">b</emphasis>
downwards to posterior end of row
<emphasis id="B950EAD7A80FFF85C354F96D7624F882" box="[559,573,1771,1797]" italics="true" pageId="8" pageNumber="53">d</emphasis>
; superior segments rows
<emphasis id="B950EAD7A80FFF85C21AF96D7763F882" box="[865,890,1771,1797]" italics="true" pageId="8" pageNumber="53">5s</emphasis>
and
<emphasis id="B950EAD7A80FFF85C2C8F96C77D5F883" box="[947,972,1770,1796]" italics="true" pageId="8" pageNumber="53">6s</emphasis>
and row
<emphasis id="B950EAD7A80FFF85C542F96D705EF882" box="[1081,1095,1771,1797]" italics="true" pageId="8" pageNumber="53">7</emphasis>
close to eye, inferior sections of rows
<emphasis id="B950EAD7A80FFF85C18EF889751AF8AE" box="[245,259,1807,1833]" italics="true" pageId="8" pageNumber="53">5</emphasis>
and
<emphasis id="B950EAD7A80FFF85C040F8887550F8AF" box="[315,329,1806,1832]" italics="true" pageId="8" pageNumber="53">6</emphasis>
well developed, row
<emphasis id="B950EAD7A80FFF85C345F889764DF8AE" box="[574,596,1807,1833]" italics="true" pageId="8" pageNumber="53">5i</emphasis>
below middle of row
<emphasis id="B950EAD7A80FFF85C22AF8897746F8AE" box="[849,863,1807,1833]" italics="true" pageId="8" pageNumber="53">b</emphasis>
, row
<emphasis id="B950EAD7A80FFF85C2DAF88877AEF8AF" box="[929,951,1806,1832]" italics="true" pageId="8" pageNumber="53">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80FFF85C582F889711EF8AE" box="[1273,1287,1807,1833]" italics="true" pageId="8" pageNumber="53">b</emphasis>
, both ending downwards below row
<emphasis id="B950EAD7A80FFF85C0E1F8B575B1F8CA" box="[410,424,1843,1869]" italics="true" pageId="8" pageNumber="53">d</emphasis>
in the level and behind dermal fold (
<emphasis id="B950EAD7A80FFF85C246F8B57752F8CA" box="[829,843,1843,1869]" italics="true" pageId="8" pageNumber="53">1</emphasis>
: 10+23,
<emphasis id="B950EAD7A80FFF85C2D6F8B577A2F8CA" box="[941,955,1843,1869]" italics="true" pageId="8" pageNumber="53">2</emphasis>
: 815,
<emphasis id="B950EAD7A80FFF85C575F8B57005F8CA" box="[1038,1052,1843,1869]" italics="true" pageId="8" pageNumber="53">3</emphasis>
: 821,
<emphasis id="B950EAD7A80FFF85C515F8B57065F8CA" box="[1134,1148,1843,1869]" italics="true" pageId="8" pageNumber="53">4</emphasis>
: 8
<geoCoordinate id="EE105002A80FFF85C5D3F8B470F1F8CA" box="[1192,1256,1842,1869]" degrees="15, 5" direction="south" orientation="latitude" pageId="8" pageNumber="53" precision="5555" value="-15.5">
15,
<emphasis id="B950EAD7A80FFF85C5B4F8B570F1F8CA" box="[1231,1256,1843,1869]" italics="true" pageId="8" pageNumber="53">5s</emphasis>
</geoCoordinate>
: 48,
<emphasis id="B950EAD7A80FFF85C457F8B5715BF8CA" box="[1324,1346,1843,1869]" italics="true" pageId="8" pageNumber="53">5i</emphasis>
: 10
<geoCoordinate id="EE105002A80FFF85C407F8B574A9F8F7" degrees="16, 6" direction="south" orientation="latitude" pageId="8" pageNumber="53" precision="5555" value="-16.6">
16,
<emphasis id="B950EAD7A80FFF85C1ECF8D074A9F8F7" box="[151,176,1878,1904]" italics="true" pageId="8" pageNumber="53">6s</emphasis>
</geoCoordinate>
: 47,
<emphasis id="B950EAD7A80FFF85C181F8D07508F8F7" box="[250,273,1878,1904]" italics="true" pageId="8" pageNumber="53">6i</emphasis>
: 1219,
<emphasis id="B950EAD7A80FFF85C00DF8D17597F8F6" box="[374,398,1879,1905]" italics="true" pageId="8" pageNumber="53">7:</emphasis>
12). Longitudinal row
<emphasis id="B950EAD7A80FFF85C3DCF8D176ACF8F6" box="[679,693,1879,1905]" italics="true" pageId="8" pageNumber="53">b</emphasis>
(1016) extending forwards above row
<emphasis id="B950EAD7A80FFF85C5F8F8D17080F8F6" box="[1155,1177,1879,1905]" italics="true" pageId="8" pageNumber="53">5i</emphasis>
to upper end of row
<emphasis id="B950EAD7A80FFF85C4F5F8D17185F8F6" box="[1422,1436,1879,1905]" italics="true" pageId="8" pageNumber="53">4</emphasis>
not reaching below eye. Longitudinal row
<emphasis id="B950EAD7A80FFF85C300F8FD7690F812" box="[635,649,1915,1941]" italics="true" pageId="8" pageNumber="53">d</emphasis>
(9+8 15+11) discontinuous with large gap between supralabial and cheek parts from suborbital row
<emphasis id="B950EAD7A80FFF85C373F819760FF83E" box="[520,534,1951,1977]" italics="true" pageId="8" pageNumber="53">2</emphasis>
to row
<emphasis id="B950EAD7A80FFF85C314F819769DF83E" box="[623,644,1951,1977]" italics="true" pageId="8" pageNumber="53">3.</emphasis>
(3)
<emphasis id="B950EAD7A80FFF85C3CFF81977DEF83E" box="[692,967,1951,1977]" italics="true" pageId="8" pageNumber="53">preoperculo-mandibular</emphasis>
: external row
<emphasis id="B950EAD7A80FFF85C517F8197061F83F" box="[1132,1144,1951,1976]" italics="true" pageId="8" pageNumber="53">e</emphasis>
(27+22 44+34) divided into anterior and posterior sections; internal row
<emphasis id="B950EAD7A80FFF85C3C8F84576A2F85B" box="[691,699,1987,2012]" italics="true" pageId="8" pageNumber="53">i</emphasis>
continuous (4062), mental row
<emphasis id="B950EAD7A80FFF85C551F845702BF85A" box="[1066,1074,1987,2013]" italics="true" pageId="8" pageNumber="53">f</emphasis>
(614) as cluster in front of chin barbel (
<figureCitation id="131F2A40A80EFF84C190FF117536FF36" box="[235,303,151,177]" captionStart="FIGURE 9" captionStartId="10.[151,250,1929,1953]" captionTargetBox="[265,1323,882,1905]" captionTargetId="figure-71@10.[265,1323,882,1905]" captionTargetPageId="10" captionText="FIGURE 9. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, anterior part of preoperculo-mandibular rows in PMR VP4680, male, 47.0+8.6 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Terminology in text. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882948" httpUri="https://zenodo.org/record/4882948/files/figure.png" pageId="9" pageNumber="54">Fig. 9</figureCitation>
). (4)
<emphasis id="B950EAD7A80EFF84C012FF117610FF36" box="[361,521,151,177]" italics="true" pageId="9" pageNumber="54">oculoscapular</emphasis>
: vertical row
<emphasis id="B950EAD7A80EFF84C3D9FF1176DAFF37" box="[674,707,151,176]" italics="true" pageId="9" pageNumber="54">tra</emphasis>
(13) behind lower posterior eye edge with one additional papilla behind it, longitudinal row
<emphasis id="B950EAD7A80EFF84C0AAFF3A75FCFF50" box="[465,485,188,215]" italics="true" pageId="9" pageNumber="54"></emphasis>
placed posteriorly above opercle (1+3 5+3), divided by vertical row
<emphasis id="B950EAD7A80EFF84C46CFF3D7121FF53" box="[1303,1336,187,212]" italics="true" pageId="9" pageNumber="54">trp</emphasis>
(35) in two parts, vertical row
<emphasis id="B950EAD7A80EFF84C0E3FF5975BFFF7F" box="[408,422,223,248]" italics="true" pageId="9" pageNumber="54">q</emphasis>
(25) behind and below row
<emphasis id="B950EAD7A80EFF84C395FF667712FF7C" box="[750,779,224,251]" italics="true" pageId="9" pageNumber="54">x¹,</emphasis>
with one or two additional papillae behind it. Longitudinal row
<emphasis id="B950EAD7A80EFF84C1B2FE8274C7FE98" box="[201,222,260,287]" italics="true" pageId="9" pageNumber="54"></emphasis>
(23) placed above opercular posterior edge, with transversal row
<emphasis id="B950EAD7A80EFF84C2BEFE8577C8FE9B" box="[965,977,259,284]" italics="true" pageId="9" pageNumber="54">y</emphasis>
(23) below it. Axillary vertical rows
<emphasis id="B950EAD7A80EFF84C400FE857185FE97" box="[1403,1436,258,284]" italics="true" pageId="9" pageNumber="54">
as
<superScript id="7C519B8DA80EFF84C4EFFE847185FE97" attach="left" box="[1428,1436,258,272]" fontSize="6" pageId="9" pageNumber="54">1</superScript>
</emphasis>
(37),
<emphasis id="B950EAD7A80EFF84C19AFEA1751BFEB3" box="[225,258,294,320]" italics="true" pageId="9" pageNumber="54">
as
<superScript id="7C519B8DA80EFF84C181FEA0751BFEB3" attach="left" box="[250,258,294,308]" fontSize="6" pageId="9" pageNumber="54">2</superScript>
</emphasis>
(47),
<emphasis id="B950EAD7A80EFF84C029FEA1756AFEB3" box="[338,371,294,320]" italics="true" pageId="9" pageNumber="54">
as
<superScript id="7C519B8DA80EFF84C010FEA0756AFEB3" attach="left" box="[363,371,294,308]" fontSize="6" pageId="9" pageNumber="54">3</superScript>
</emphasis>
(612) present, row
<emphasis id="B950EAD7A80EFF84C322FEA1766EFEB3" box="[601,631,294,321]" italics="true" pageId="9" pageNumber="54">
la
<superScript id="7C519B8DA80EFF84C314FEA0766EFEB3" attach="left" box="[623,631,294,308]" fontSize="6" pageId="9" pageNumber="54">1</superScript>
</emphasis>
(24) above
<emphasis id="B950EAD7A80EFF84C272FEA17733FEB3" box="[777,810,294,320]" italics="true" pageId="9" pageNumber="54">
as
<superScript id="7C519B8DA80EFF84C259FEA07733FEB3" attach="left" box="[802,810,294,308]" fontSize="6" pageId="9" pageNumber="54">2</superScript>
</emphasis>
, row
<emphasis id="B950EAD7A80EFF84C212FEA1779EFEB3" box="[873,903,294,321]" italics="true" pageId="9" pageNumber="54">
la
<superScript id="7C519B8DA80EFF84C204FEA0779EFEB3" attach="left" box="[895,903,294,308]" fontSize="6" pageId="9" pageNumber="54">2</superScript>
</emphasis>
(35) above between
<emphasis id="B950EAD7A80EFF84C507FEA17084FEB3" box="[1148,1181,294,320]" italics="true" pageId="9" pageNumber="54">
as
<superScript id="7C519B8DA80EFF84C5EEFEA07084FEB3" attach="left" box="[1173,1181,294,308]" fontSize="6" pageId="9" pageNumber="54">2</superScript>
</emphasis>
and
<emphasis id="B950EAD7A80EFF84C5A9FEA170EAFEB3" box="[1234,1267,294,320]" italics="true" pageId="9" pageNumber="54">
as
<superScript id="7C519B8DA80EFF84C590FEA070EAFEB3" attach="left" box="[1259,1267,294,308]" fontSize="6" pageId="9" pageNumber="54">3</superScript>
</emphasis>
. (5)
<emphasis id="B950EAD7A80EFF84C45CFEA1718DFEC6" box="[1319,1428,295,321]" italics="true" pageId="9" pageNumber="54">opercular</emphasis>
: transverse row
<emphasis id="B950EAD7A80EFF84C03AFECD754EFEE3" box="[321,343,331,356]" italics="true" pageId="9" pageNumber="54">ot</emphasis>
(2348); superior longitudinal row
<emphasis id="B950EAD7A80EFF84C39DFECD76E6FEE3" box="[742,767,331,356]" italics="true" pageId="9" pageNumber="54">os</emphasis>
(1024); and interior longitudinal row
<emphasis id="B950EAD7A80EFF84C5C8FECD70D0FEE3" box="[1203,1225,331,356]" italics="true" pageId="9" pageNumber="54">oi</emphasis>
(35). (6)
<emphasis id="B950EAD7A80EFF84C43BFECD74C6FE0E" italics="true" pageId="9" pageNumber="54">anterior dorsal</emphasis>
: anterior row
<emphasis id="B950EAD7A80EFF84C0FEFEE9758AFE0F" box="[389,403,367,392]" italics="true" pageId="9" pageNumber="54">n</emphasis>
longitudinal (13) behind upper eye, transverse row
<emphasis id="B950EAD7A80EFF84C28CFEE9701CFE0F" box="[1015,1029,367,392]" italics="true" pageId="9" pageNumber="54">o</emphasis>
(14) distant from fellow in dorsal midline; longitudinal row
<emphasis id="B950EAD7A80EFF84C0C4FE1575D4FE2B" box="[447,461,403,428]" italics="true" pageId="9" pageNumber="54">g</emphasis>
(34) distant behind row
<emphasis id="B950EAD7A80EFF84C388FE157718FE2B" box="[755,769,403,428]" italics="true" pageId="9" pageNumber="54">o</emphasis>
, longitudinal row
<emphasis id="B950EAD7A80EFF84C2A9FE1577FFFE2B" box="[978,998,403,428]" italics="true" pageId="9" pageNumber="54">m</emphasis>
and longitudinal row
<emphasis id="B950EAD7A80EFF84C59AFE1570F6FE2A" box="[1249,1263,403,429]" italics="true" pageId="9" pageNumber="54">h</emphasis>
not visible. (7)
<emphasis id="B950EAD7A80EFF84C1ECFE31750EFE56" box="[151,279,439,465]" italics="true" pageId="9" pageNumber="54">interorbital</emphasis>
: two pairs of interorbital transversal rows, anterior
<emphasis id="B950EAD7A80EFF84C219FE317767FE57" box="[866,894,439,464]" italics="true" pageId="9" pageNumber="54">pa</emphasis>
(12) and posterior
<emphasis id="B950EAD7A80EFF84C51CFE31709AFE57" box="[1127,1155,439,464]" italics="true" pageId="9" pageNumber="54">pp</emphasis>
(23). Body with 1822 transversal
<emphasis id="B950EAD7A80EFF84C063FE5D7525FE72" box="[280,316,475,501]" italics="true" pageId="9" pageNumber="54">ltm</emphasis>
rows starting anteriorly behind axilla and
<emphasis id="B950EAD7A80EFF84C262FE5D772BFE73" box="[793,818,475,500]" italics="true" pageId="9" pageNumber="54">as</emphasis>
rows, alternating anteriorly with three longitudinal
<emphasis id="B950EAD7A80EFF84C403FE5D7185FE72" box="[1400,1436,475,501]" italics="true" pageId="9" pageNumber="54">llm</emphasis>
rows making anterior beginning pattern of -II-I-I (
<figureCitation id="131F2A40A80EFF84C3A9FE79773FFD9E" box="[722,806,511,537]" captionStart="FIGURE 10" captionStartId="11.[151,250,1076,1100]" captionTargetBox="[213,1375,181,1052]" captionTargetId="figure-20@11.[213,1375,181,1052]" captionTargetPageId="11" captionText="FIGURE 10. Benthophilus persicus sp. nov. Sensory papillae pattern of lateral line system in reversibly stained paratypes, sensory papillae rows on anterior flank in PMR VP4681, male, 35.1+8.5 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Sensory papillae rows marked, terminology text. The transversal ltm rows starting anteriorly behind axilla, alternating with three longitudinal llm rows making anterior beginning pattern of -II-I-I. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882950" httpUri="https://zenodo.org/record/4882950/files/figure.png" pageId="9" pageNumber="54">Fig. 10</figureCitation>
). Three transversal
<emphasis id="B950EAD7A80EFF84C57CFE797002FD9E" box="[1031,1051,511,537]" italics="true" pageId="9" pageNumber="54">lv</emphasis>
rows at lower anterior body. Two longitudinal
<emphasis id="B950EAD7A80EFF84C05DFDA57523FDBA" box="[294,314,547,573]" italics="true" pageId="9" pageNumber="54">lc</emphasis>
rows, one along midline of caudal fin, the second above it.
</paragraph>
<paragraph id="8B9B36C5A80EFF84C1BCFDC170DEFCBE" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="54">
<emphasis id="B950EAD7A80EFF84C1BCFDC17526FDE6" box="[199,319,583,609]" italics="true" pageId="9" pageNumber="54">Osteology.</emphasis>
Vertebral column: 9 (810) precaudal and 19 (1920) caudal vertebrae (including urostyle); total vertebral count: 28 (2830). D1 pterygiophore insertion pattern: 322 1*01*1*1* (only from
<typeStatus id="549F8867A80EFF84C5D5FDED710CFD01" box="[1198,1301,619,646]" pageId="9" pageNumber="54" type="holotype">holotype</typeStatus>
); number of anal pterygiophores anterior to the first haemal spine 0 (01). Crest-like remaxillary process present on posterior third of premaxialla, sloping with a steep angle on anterior rim and gently towards posterior tip of premaxialla. Five branchiostegal rays. One epural. Number of C rays inserting in hypural 5: 2 (12), 3+4 (fused): 5 (56), hypural 1+2 (fused): 4 (45) and parhypural: 1 (01), total number of C rays inserting in hypurals, and parhypural: 12 (12; fused hypural 1+2 and 3+4 separated by a large gap, which does not support any branched caudal ray.
</paragraph>
<paragraph id="8B9B36C5A80EFF84C1BCFCC574E6FC6A" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="54">
<emphasis id="B950EAD7A80EFF84C1BCFCC5755EFCDA" box="[199,327,835,861]" italics="true" pageId="9" pageNumber="54">Coloration.</emphasis>
No live coloration recorded. Color of preserved specimens (
<figureCitation id="131F2A40A80EFF84C294FCC5702CFCDA" box="[1007,1077,835,861]" captionStart="FIGURE 4" captionStartId="4.[151,250,726,750]" captionTargetBox="[248,1340,181,702]" captionTargetId="figure-33@4.[248,1340,181,702]" captionTargetPageId="4" captionText="FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882938" httpUri="https://zenodo.org/record/4882938/files/figure.png" pageId="9" pageNumber="54">Fig. 4</figureCitation>
): body opaque fawn, irregularly scattered melanophores present on upper head and body, and also on dorsal, caudal and pectoral fins. In some specimens remaining pigmentation almost invisible. Some specimens with three whitish saddles on back: at D2 anterior beginning, D2 posterior end and on caudal peduncle, and with four pigmented blotches on caudal fin longitudinally arranged.
</paragraph>
</subSubSection>
<subSubSection id="C33E654EA80EFF84C1BCFC7176D9FB96" box="[199,704,1015,1041]" pageId="9" pageNumber="54" type="etymology">
<paragraph id="8B9B36C5A80EFF84C1BCFC7176D9FB96" blockId="9.[151,1437,151,2013]" box="[199,704,1015,1041]" pageId="9" pageNumber="54">
<emphasis id="B950EAD7A80EFF84C1BCFC717551FB96" bold="true" box="[199,328,1015,1041]" pageId="9" pageNumber="54">Etymology</emphasis>
. The species is named for
<collectingCountry id="F3337655A80EFF84C30FFC7176A5FB96" box="[628,700,1015,1041]" name="Iran" pageId="9" pageNumber="54">Persia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C33E654EA80EFF84C1BCFB9C70D9FBFA" pageId="9" pageNumber="54" type="distribution">
<paragraph id="8B9B36C5A80EFF84C1BCFB9C70D9FBFA" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="54">
<emphasis id="B950EAD7A80EFF84C1BCFB9C75F0FBB3" bold="true" box="[199,489,1050,1077]" pageId="9" pageNumber="54">Distribution and habitat</emphasis>
. Southern Caspian Sea basin (
<figureCitation id="131F2A40A80EFF84C241FB9D7763FBB2" box="[826,890,1051,1077]" captionStart="FIGURE 1" captionStartId="1.[151,250,1424,1448]" captionTargetBox="[344,1241,306,1394]" captionTargetId="figure-83@1.[344,1244,306,1398]" captionTargetPageId="1" captionText="FIGURE 1. Bathymetric map of the Caspian Sea, showing the type locality (yellow star) of Benthophilus persicus sp. nov. near Anzali, southern Caspian Sea. The image was produced using Surfer ver. 10.1.561 mapping software package (Golden Software, LLC)." figureDoi="http://doi.org/10.5281/zenodo.4882932" httpUri="https://zenodo.org/record/4882932/files/figure.png" pageId="9" pageNumber="54">Fig. 1</figureCitation>
).
<taxonomicName id="4C244D46A80EFF84C2E9FB9D7090FBB3" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[914,1161,1051,1077]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80EFF84C2E9FB9D7090FBB3" box="[914,1161,1051,1077]" italics="true" pageId="9" pageNumber="54">Benthophilus persicus</emphasis>
</taxonomicName>
inhabits brackish waters and is abundant on sandy bottoms in coastal areas of the southern Caspian Sea. Capture depth ranges from
<quantity id="4CDC9B20A80EFF84C435FBB974A9FBFA" metricMagnitude="1" metricUnit="m" metricValue="3.8" metricValueMax="7.0" metricValueMin="0.6" pageId="9" pageNumber="54" unit="m" value="38.0" valueMax="70.0" valueMin="6.0">
6 to
<specimenCount id="9D22FD4CA80EFF84C4FBFBB974A9FBFA" pageId="9" pageNumber="54" type="generic">70 m</specimenCount>
</quantity>
. However, no specimens have yet been collected in the eastern part of southern Caspian Sea.
</paragraph>
</subSubSection>
<subSubSection id="C33E654EA80EFF9DC1BCFB01718BFE2A" lastPageId="16" lastPageNumber="61" pageId="9" pageNumber="54" type="discussion">
<paragraph id="8B9B36C5A80EFF84C1BCFB017678F9A9" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="54">
<emphasis id="B950EAD7A80EFF84C1BCFB01752DFB26" bold="true" box="[199,308,1159,1185]" pageId="9" pageNumber="54">Remarks</emphasis>
.
<bibRefCitation id="EFB54B34A80EFF84C039FB0176BCFB26" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." box="[322,677,1159,1185]" pageId="9" pageNumber="54" pagination="31 - 96" refId="ref11847" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2007) Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18 (1), 31 - 96." type="journal article" year="2007">Boldyrev &amp; Bogutskaya (2007)</bibRefCitation>
tentatively assigned 20 recognized species of the genus
<taxonomicName id="4C244D46A80EFF84C45BFB0174A1FB42" authorityName="Eichwald" authorityYear="1831" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="genus">
<emphasis id="B950EAD7A80EFF84C45BFB0174A1FB42" italics="true" pageId="9" pageNumber="54">Benthophilus</emphasis>
</taxonomicName>
to four phenotypic groups. The most prominent differences of the new species as compared with members of the four different species groups are as follows. The new species clearly differs from group I members comprising
<taxonomicName id="4C244D46A80EFF84C1ECFB7575C1FA8A" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[151,472,1267,1293]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="granulosus">
<emphasis id="B950EAD7A80EFF84C1ECFB75752AFA8A" box="[151,307,1267,1293]" italics="true" pageId="9" pageNumber="54">B. granulosus</emphasis>
Kessler, 1877
</taxonomicName>
,
<taxonomicName id="4C244D46A80EFF84C09FFB7576E0FA8A" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[484,761,1267,1293]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="grimmi">
<emphasis id="B950EAD7A80EFF84C09FFB75764DFA8B" box="[484,596,1267,1292]" italics="true" pageId="9" pageNumber="54">B. grimmi</emphasis>
Kessler, 1877
</taxonomicName>
,
<taxonomicName id="4C244D46A80EFF84C27FFB7577E7FA8A" authority="Berg, 1927" authorityName="Berg" authorityYear="1927" box="[772,1022,1267,1293]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="kessleri">
<emphasis id="B950EAD7A80EFF84C27FFB757763FA8A" box="[772,890,1267,1293]" italics="true" pageId="9" pageNumber="54">B. kessleri</emphasis>
Berg, 1927
</taxonomicName>
,
<taxonomicName id="4C244D46A80EFF84C576FB757175FA8A" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[1037,1388,1267,1293]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="leptorhynchus">
<emphasis id="B950EAD7A80EFF84C576FB7570D2FA8A" box="[1037,1227,1267,1293]" italics="true" pageId="9" pageNumber="54">B. leptorhynchus</emphasis>
Kessler, 1877
</taxonomicName>
and
<taxonomicName id="4C244D46A80EFF84C1ECFA917668FAB6" authority="Pinchuk &amp; Ragimov, 1979" authorityName="Pinchuk &amp; Ragimov" authorityYear="1979" box="[151,625,1303,1329]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="svetovidovi">
<emphasis id="B950EAD7A80EFF84C1ECFA91752EFAB6" box="[151,311,1303,1329]" italics="true" pageId="9" pageNumber="54">B. svetovidovi</emphasis>
<bibRefCitation id="EFB54B34A80EFF84C03BFA917668FAB6" author="Pinchuk, V. I. &amp; Ragimov, D. B." box="[320,625,1303,1329]" pageId="9" pageNumber="54" pagination="515 - 519" refId="ref12224" refString="Pinchuk, V. I. &amp; Ragimov, D. B. (1979) Benthophilus svetovidovi Pinchuk et Ragimov, sp. n. (Pisces, Gobiidae) from the Caspian Sea and a key of species of the genus Benthophilus. Zoologicheskii Zhurnal, 58, 515 - 519. [in Russian]" type="journal article" year="1979">Pinchuk &amp; Ragimov, 1979</bibRefCitation>
</taxonomicName>
by having tubercles on the body (
<emphasis id="B950EAD7A80EFF84C284FA917007FAB7" box="[1023,1054,1303,1328]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
bony plates rather than tubercles on body).
<taxonomicName id="4C244D46A80EFF84C071FABD761AFAD3" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[266,515,1339,1365]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80EFF84C071FABD761AFAD3" box="[266,515,1339,1365]" italics="true" pageId="9" pageNumber="54">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80EFF84C3E3FABD778DFAD2" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[664,916,1339,1365]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="baeri">
<emphasis id="B950EAD7A80EFF84C3E3FABD76EAFAD2" box="[664,755,1339,1365]" italics="true" pageId="9" pageNumber="54">B. baeri</emphasis>
Kessler, 1877
</taxonomicName>
and
<taxonomicName id="4C244D46A80EFF84C2B0FABD70F5FAD2" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[971,1260,1339,1365]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="spinosus">
<emphasis id="B950EAD7A80EFF84C2B0FABD7052FAD3" box="[971,1099,1339,1364]" italics="true" pageId="9" pageNumber="54">B. spinosus</emphasis>
Kessler, 1877
</taxonomicName>
of group IV by having comparatively small dorsal tubercles with body poorly defined, dominated by clearly visible spinules (
<figureCitation id="131F2A40A80EFF84C41FFAD974BCFA1A" captionStart="FIGURE 2" captionStartId="3.[151,250,966,990]" captionTargetBox="[225,1361,181,942]" captionTargetId="figure-20@3.[225,1361,181,942]" captionTargetPageId="3" captionText="FIGURE 2. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Preopercle with dermal ossifications: T tubercles with spinules on tubercles appear irregularly arranged, G1 - single granule, G2 and G3 - granules grouped two and three together, respectively. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882934" httpUri="https://zenodo.org/record/4882934/files/figure.png" pageId="9" pageNumber="54">Figs. 2</figureCitation>
and
<figureCitation id="131F2A40A80EFF84C1A7FA0574F3FA1A" box="[220,234,1411,1437]" captionStart="FIGURE 3" captionStartId="3.[151,250,1907,1931]" captionTargetBox="[225,1361,1122,1883]" captionTargetId="figure-79@3.[225,1361,1122,1883]" captionTargetPageId="3" captionText="FIGURE 3. Benthophilus persicus sp. nov. Reversibly stained paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Dermal ossifications on anterior dorsal body view: D - dorsal row of tubercles: T1- tubercle with one radial row of spinules and T2 - tubercle with two radial rows of spinules, DL - dorsolateral row of tubercles, G - granules. Photo by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882936" httpUri="https://zenodo.org/record/4882936/files/figure.png" pageId="9" pageNumber="54">3</figureCitation>
) (
<emphasis id="B950EAD7A80EFF84C07FFA05753BFA1B" box="[260,290,1411,1436]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
very large dorsal tubercles with clear polygonal conical erected body with small or hardly visible spinules,
<figureCitation id="131F2A40A80EFF84C19BFA217524FA46" box="[224,317,1447,1473]" captionStart="FIGURE 4" captionStartId="4.[151,250,726,750]" captionTargetBox="[248,1340,181,702]" captionTargetId="figure-33@4.[248,1340,181,702]" captionTargetPageId="4" captionText="FIGURE 4. Benthophilus persicus sp. nov. preserved specimens. Holotype, ZSM 47595, male, 45.2+9.9 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: A: lateral view, B: urogenital papilla of adult male (marked by arrow). Paratype, ZSM 47599, female, 30.8+7.0 mm, Iran, Gilan Province, Anzali, southern Caspian Sea: C: lateral view, D: urogenital papilla of adult female (marked by arrow). Photos by M. Kovačić." figureDoi="http://doi.org/10.5281/zenodo.4882938" httpUri="https://zenodo.org/record/4882938/files/figure.png" pageId="9" pageNumber="54">
Figure
<quantity id="4CDC9B20A80EFF84C054FA217524FA46" box="[303,317,1447,1473]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="9" pageNumber="54" unit="in" value="4.0">4</quantity>
</figureCitation>
in
<bibRefCitation id="EFB54B34A80EFF84C024FA2176D3FA46" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." box="[351,714,1447,1473]" pageId="9" pageNumber="54" pagination="31 - 96" refId="ref11847" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2007) Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18 (1), 31 - 96." type="journal article" year="2007">Boldyrev &amp; Bogutskaya (2007))</bibRefCitation>
, 2225 tubercles in ventral row (
<emphasis id="B950EAD7A80EFF84C543FA21704FFA47" box="[1080,1110,1447,1472]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
920), numerous tiny simple structured granules, sometimes grouped two or three together (
<emphasis id="B950EAD7A80EFF84C219FA4D7799FA63" box="[866,896,1483,1508]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
large and sparse granules with spinules), a few granules present on the posterior dorsal part of the caudal peduncle (
<emphasis id="B950EAD7A80EFF84C2F1FA6977B1F98F" box="[906,936,1519,1544]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
granules restricted to the upper head surface, gill covers and anterior part of the back).
</paragraph>
<paragraph id="8B9B36C5A80EFF84C1BCF9B1776CF85A" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="54">
<taxonomicName id="4C244D46A80EFF84C1BCF9B175A7F9D7" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,446,1591,1617]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80EFF84C1BCF9B175A7F9D7" box="[199,446,1591,1617]" italics="true" pageId="9" pageNumber="54">Benthophilus persicus</emphasis>
</taxonomicName>
cannot be unambiguously assigned to phenotypic groups II or III of
<bibRefCitation id="EFB54B34A80EFF84C5C2F9B17504F9F2" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." pageId="9" pageNumber="54" pagination="31 - 96" refId="ref11847" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2007) Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18 (1), 31 - 96." type="journal article" year="2007">Boldyrev &amp; Bogutskaya (2007)</bibRefCitation>
since it features a mix of characters that were used to distinguish these two groups. The new species has tubercles with two posterior rows of spinules forming an acute angle (
<emphasis id="B950EAD7A80EFF84C2D8F9F977D8F91F" box="[931,961,1663,1688]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
almost right angle of two rows of spinules on dorsal tubercules in group III), tubercles present between eyes (
<emphasis id="B950EAD7A80EFF84C2F2F92577B1F93B" box="[905,936,1699,1724]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
tubercles absent between eyes in group III), a complete dorsal row (
<emphasis id="B950EAD7A80EFF84C0DAF94175A6F967" box="[417,447,1735,1760]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
dorsal row incomplete in group III), and a low count of 2229 dorsal row tubercles (
<emphasis id="B950EAD7A80EFF84C405F9417185F967" box="[1406,1436,1735,1760]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
two out of four species of group III,
<taxonomicName id="4C244D46A80EFF84C347F96D776BF882" authority="Ragimov, 1982" authorityName="Ragimov" authorityYear="1982" box="[572,882,1771,1797]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="9" pageNumber="54" phylum="Chordata" rank="species" species="pinchuki">
<emphasis id="B950EAD7A80EFF84C347F96D76A4F882" box="[572,701,1771,1797]" italics="true" pageId="9" pageNumber="54">B. pinchuki</emphasis>
Ragimov, 1982
</taxonomicName>
and
<taxonomicName id="4C244D46A80EFF84C2D7F96D7181F882" authority="Boldyrev &amp; Bogutskaya, 2004" authorityName="Boldyrev &amp; Bogutskaya" authorityYear="2004" box="[940,1432,1771,1797]" genus="Bogutskaya" pageId="9" pageNumber="54" rank="species" species="ragimovi">
<emphasis id="B950EAD7A80EFF84C2D7F96D702BF883" box="[940,1074,1771,1796]" italics="true" pageId="9" pageNumber="54">B. ragimovi</emphasis>
<bibRefCitation id="EFB54B34A80EFF84C541F96D7181F882" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." box="[1082,1432,1771,1797]" pageId="9" pageNumber="54" pagination="129 - 135" refId="ref11803" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2004) Description of two new species of tadpole gobies (Teleostei: Gobiidae: Benthophilus). Zoosystematica Rossica, 13 (1), 129 - 135." type="journal article" year="2004">Boldyrev &amp; Bogutskaya, 2004</bibRefCitation>
</taxonomicName>
, having higher counts of dorsal row tubercles). However, it has no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80EFF84C1DAF8B574A6F8CB" box="[161,191,1843,1868]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
usually four tubercles in row on each side of head in temporal and occipital regions of head, one unpaired temporal tubercle in group II) and no blotches on body in a preserved state (
<emphasis id="B950EAD7A80EFF84C2B9F8D177F8F8F7" box="[962,993,1879,1904]" italics="true" pageId="9" pageNumber="54">vs.</emphasis>
blotches on body in group II, except
<taxonomicName id="4C244D46A80EFF84C4FFF8D17618F812" authority="Ragimov, 1978" authorityName="Ragimov" authorityYear="1978" genus="Bogutskaya" pageId="9" pageNumber="54" rank="species" species="abdurahmanovi">
<emphasis id="B950EAD7A80EFF84C4FFF8D17551F812" italics="true" pageId="9" pageNumber="54">B. abdurahmanovi</emphasis>
<bibRefCitation id="EFB54B34A80EFF84C02AF8FD7618F812" author="Ragimov, D. B." box="[337,513,1915,1941]" pageId="9" pageNumber="54" pagination="791 - 798" refId="ref12612" refString="Ragimov, D. B. (1978) On the taxonomic status of some species of the genus Benthophilus (Gobiidae) from the Caspian and Azov Seas. Voprosy Ikhtiologii, 18, 791 - 798. [in Russian]" type="journal article" year="1978">Ragimov, 1978</bibRefCitation>
</taxonomicName>
). The phenotypic groups II and III of
<bibRefCitation id="EFB54B34A80EFF84C2CAF8FD710EF812" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." box="[945,1303,1915,1941]" pageId="9" pageNumber="54" pagination="31 - 96" refId="ref11847" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2007) Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18 (1), 31 - 96." type="journal article" year="2007">Boldyrev &amp; Bogutskaya (2007)</bibRefCitation>
appear less well defined morphologically than the other two groups. Therefore, the new species is here compared with each of the 13 recognized species of groups II and III in alphabetic order:
</paragraph>
<caption id="DF5B664DA80DFF87C1ECFC98719AFCDD" ID-DOI="http://doi.org/10.5281/zenodo.4882946" ID-Zenodo-Dep="4882946" httpUri="https://zenodo.org/record/4882946/files/figure.png" pageId="10" pageNumber="55" startId="10.[151,250,798,822]" targetBox="[262,1333,191,774]" targetPageId="10">
<paragraph id="8B9B36C5A80DFF87C1ECFC98719AFCDD" blockId="10.[151,1436,798,858]" pageId="10" pageNumber="55">
<emphasis id="B950EAD7A80DFF87C1ECFC98750CFCB1" bold="true" box="[151,277,798,822]" pageId="10" pageNumber="55">FIGURE 8.</emphasis>
Head sensory papillae pattern of lateral line system of
<taxonomicName id="4C244D46A80DFF87C223FC987023FCB1" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[856,1082,798,822]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="55" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A80DFF87C223FC987023FCB1" box="[856,1082,798,822]" italics="true" pageId="10" pageNumber="55">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A80DFF87C538FC98708CFCB1" bold="true" box="[1091,1173,798,822]" pageId="10" pageNumber="55">
<taxonomicNameLabel id="A26357ACA80DFF87C538FC98708CFCB1" box="[1091,1173,798,822]" pageId="10" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Paratype, PMR VP4679, male, 43.3+10.0 mm, Iran, Gilan Province, Chaboksar, southern Caspian Sea. Terminology in text. Drawing by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA80DFF87C1ECF80F7667F86E" ID-DOI="http://doi.org/10.5281/zenodo.4882948" ID-Zenodo-Dep="4882948" httpUri="https://zenodo.org/record/4882948/files/figure.png" pageId="10" pageNumber="55" startId="10.[151,250,1929,1953]" targetBox="[265,1323,882,1905]" targetPageId="10">
<paragraph id="8B9B36C5A80DFF87C1ECF80F7667F86E" blockId="10.[151,1437,1929,2026]" pageId="10" pageNumber="55">
<emphasis id="B950EAD7A80DFF87C1ECF80F750AF826" bold="true" box="[151,275,1929,1953]" pageId="10" pageNumber="55">FIGURE 9.</emphasis>
<taxonomicName id="4C244D46A80DFF87C062F80F75E0F826" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[281,505,1929,1953]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="55" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A80DFF87C062F80F75E0F826" box="[281,505,1929,1953]" italics="true" pageId="10" pageNumber="55">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A80DFF87C084F80C7656F825" bold="true" box="[511,591,1930,1954]" pageId="10" pageNumber="55">
<taxonomicNameLabel id="A26357ACA80DFF87C084F80C7656F825" box="[511,591,1930,1954]" pageId="10" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Sensory papillae pattern of lateral line system in reversibly stained paratypes, anterior part of
<emphasis id="B950EAD7A80DFF87C071F82B761AF842" box="[266,515,1965,1989]" italics="true" pageId="10" pageNumber="55">preoperculo-mandibular</emphasis>
rows in PMR VP4680, male, 47.0+8.6 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Terminology in text. Photo by M. Kovačić.
</paragraph>
</caption>
<caption id="DF5B664DA80CFF86C1ECFBB2704BFB3F" ID-DOI="http://doi.org/10.5281/zenodo.4882950" ID-Zenodo-Dep="4882950" httpUri="https://zenodo.org/record/4882950/files/figure.png" pageId="11" pageNumber="56" startId="11.[151,250,1076,1100]" targetBox="[213,1375,181,1052]" targetPageId="11">
<paragraph id="8B9B36C5A80CFF86C1ECFBB2704BFB3F" blockId="11.[151,1437,1076,1208]" pageId="11" pageNumber="56">
<emphasis id="B950EAD7A80CFF86C1ECFBB2753BFBCB" bold="true" box="[151,290,1076,1100]" pageId="11" pageNumber="56">FIGURE 10.</emphasis>
<taxonomicName id="4C244D46A80CFF86C050FBB27617FBCB" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[299,526,1076,1100]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A80CFF86C050FBB27617FBCB" box="[299,526,1076,1100]" italics="true" pageId="11" pageNumber="56">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A80CFF86C36CFBB27673FBCB" bold="true" box="[535,618,1076,1100]" pageId="11" pageNumber="56">
<taxonomicNameLabel id="A26357ACA80CFF86C36CFBB27673FBCB" box="[535,618,1076,1100]" pageId="11" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Sensory papillae pattern of lateral line system in reversibly stained paratypes, sensory papillae rows on anterior flank in PMR VP4681, male, 35.1+8.5 mm, Iran, Gilan Province, Anzali, southern Caspian Sea. Sensory papillae rows marked, terminology text. The transversal
<emphasis id="B950EAD7A80CFF86C22CFBFA776EFB13" box="[855,887,1148,1172]" italics="true" pageId="11" pageNumber="56">ltm</emphasis>
rows starting anteriorly behind axilla, alternating with three longitudinal
<emphasis id="B950EAD7A80CFF86C02AFB267568FB3F" box="[337,369,1184,1208]" italics="true" pageId="11" pageNumber="56">llm</emphasis>
rows making anterior beginning pattern of -II-I-I. Photo by M. Kovačić.
</paragraph>
</caption>
<paragraph id="8B9B36C5A80CFF86C1BCFB607659FA5E" blockId="11.[151,1437,1254,2001]" pageId="11" pageNumber="56">
<taxonomicName id="4C244D46A80CFF86C1BCFB6075D9FA87" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,448,1254,1280]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80CFF86C1BCFB6075D9FA87" box="[199,448,1254,1280]" italics="true" pageId="11" pageNumber="56">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80CFF86C323FB6177C7FA86" authority="Ragimov, 1978" authorityName="Ragimov" authorityYear="1978" box="[600,990,1254,1281]" genus="Bogutskaya" pageId="11" pageNumber="56" rank="species" species="abdurahmanovi">
<emphasis id="B950EAD7A80CFF86C323FB617733FA87" box="[600,810,1254,1280]" italics="true" pageId="11" pageNumber="56">B. abdurahmanovi</emphasis>
<bibRefCitation id="EFB54B34A80CFF86C249FB6077C7FA86" author="Ragimov, D. B." box="[818,990,1254,1281]" pageId="11" pageNumber="56" pagination="791 - 798" refId="ref12612" refString="Ragimov, D. B. (1978) On the taxonomic status of some species of the genus Benthophilus (Gobiidae) from the Caspian and Azov Seas. Voprosy Ikhtiologii, 18, 791 - 798. [in Russian]" type="journal article" year="1978">Ragimov, 1978</bibRefCitation>
</taxonomicName>
by having origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80CFF86C3E2FA8D76AEFAA3" box="[665,695,1291,1316]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
under origin of the second dorsal fin), tubercles present, clearly larger than granules, with two posterior rows of spinules forming an acute angle (
<emphasis id="B950EAD7A80CFF86C54FFAA9704BFACF" box="[1076,1106,1327,1352]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
tubercles slightly larger than granules, with weakly developed spinules), no ventrolateral row of tubercles (
<emphasis id="B950EAD7A80CFF86C280FAD57003FAEB" box="[1019,1050,1363,1388]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
present), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80CFF86C0B6FAF175F2FA17" box="[461,491,1399,1424]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
weak tubercles present there), no granules on flanks (
<emphasis id="B950EAD7A80CFF86C533FAF1707FFA17" box="[1096,1126,1399,1424]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
present), transversal suborbital row
<emphasis id="B950EAD7A80CFF86C078FA1C7500FA33" box="[259,281,1434,1460]" italics="true" pageId="11" pageNumber="56">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80CFF86C323FA1C767FFA33" box="[600,614,1434,1460]" italics="true" pageId="11" pageNumber="56">b</emphasis>
(
<emphasis id="B950EAD7A80CFF86C30CFA1D768CFA33" box="[631,661,1435,1460]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80CFF86C2F4FA1C7784FA33" box="[911,925,1434,1460]" italics="true" pageId="11" pageNumber="56">b</emphasis>
), and the anterior interorbital transversal row
<emphasis id="B950EAD7A80CFF86C1ECFA3974AAFA5F" box="[151,179,1471,1496]" italics="true" pageId="11" pageNumber="56">pa</emphasis>
with 12 papilla (
<emphasis id="B950EAD7A80CFF86C0FBFA397586FA5F" box="[384,415,1471,1496]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
35 papillae).
</paragraph>
<paragraph id="8B9B36C5A80CFF86C1BCFA647536F97E" blockId="11.[151,1437,1254,2001]" pageId="11" pageNumber="56">
<taxonomicName id="4C244D46A80CFF86C1BCFA6475D9FA7B" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,448,1506,1532]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80CFF86C1BCFA6475D9FA7B" box="[199,448,1506,1532]" italics="true" pageId="11" pageNumber="56">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80CFF86C323FA6577B1FA7A" authority="Ragimov, 1978" authorityName="Ragimov" authorityYear="1978" box="[600,936,1506,1533]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="casachicus">
<emphasis id="B950EAD7A80CFF86C323FA6576EDFA7B" box="[600,756,1506,1532]" italics="true" pageId="11" pageNumber="56">B. casachicus</emphasis>
<bibRefCitation id="EFB54B34A80CFF86C387FA6477B1FA7A" author="Ragimov, D. B." box="[764,936,1506,1533]" pageId="11" pageNumber="56" pagination="791 - 798" refId="ref12612" refString="Ragimov, D. B. (1978) On the taxonomic status of some species of the genus Benthophilus (Gobiidae) from the Caspian and Azov Seas. Voprosy Ikhtiologii, 18, 791 - 798. [in Russian]" type="journal article" year="1978">Ragimov, 1978</bibRefCitation>
</taxonomicName>
by having the dermal fold rectangular large (
<emphasis id="B950EAD7A80CFF86C1DAF98174A6F9A7" box="[161,191,1543,1568]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
triangular large), the origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80CFF86C597F9817113F9A7" box="[1260,1290,1543,1568]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
under origin of the second dorsal fin), tubercles present with two posterior rows of spinules forming an acute angle (
<emphasis id="B950EAD7A80CFF86C469F9AD7128F9C3" box="[1298,1329,1579,1604]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
tubercles with numerous radial rows of spinules), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80CFF86C5B0F9C970F3F9EF" box="[1227,1258,1615,1640]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
weak tubercles present on head), anterior interorbital transversal row
<emphasis id="B950EAD7A80CFF86C27CF9F5773AF90B" box="[775,803,1651,1676]" italics="true" pageId="11" pageNumber="56">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A80CFF86C572F9F57031F90B" box="[1033,1064,1651,1676]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
35 papillae), body with 1822 transversal
<emphasis id="B950EAD7A80CFF86C061F9107527F937" box="[282,318,1686,1712]" italics="true" pageId="11" pageNumber="56">ltm</emphasis>
rows starting anteriorly behind axilla and alternating anteriorly with three longitudinal
<emphasis id="B950EAD7A80CFF86C44AF910714CF937" box="[1329,1365,1686,1712]" italics="true" pageId="11" pageNumber="56">llm</emphasis>
rows, having a total of 2125
<emphasis id="B950EAD7A80CFF86C0D2F93C75DCF953" box="[425,453,1722,1748]" italics="true" pageId="11" pageNumber="56">lm</emphasis>
rows (
<emphasis id="B950EAD7A80CFF86C36EF93D762AF953" box="[533,563,1723,1748]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
1718
<emphasis id="B950EAD7A80CFF86C3F1F93C76BFF953" box="[650,678,1722,1748]" italics="true" pageId="11" pageNumber="56">lm</emphasis>
rows in total), and a maximum body width 15.122.9% of SL (
<emphasis id="B950EAD7A80CFF86C405F93D7185F953" box="[1406,1436,1723,1748]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
23.227.8%).
</paragraph>
<paragraph id="8B9B36C5A80CFF86C1BCF88475F7F856" blockId="11.[151,1437,1254,2001]" pageId="11" pageNumber="56">
<taxonomicName id="4C244D46A80CFF86C1BCF88475DBF89B" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,450,1794,1820]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80CFF86C1BCF88475DBF89B" box="[199,450,1794,1820]" italics="true" pageId="11" pageNumber="56">Benthophilus persicus</emphasis>
</taxonomicName>
is different from
<taxonomicName id="4C244D46A80CFF86C3EAF88577FFF89A" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[657,998,1794,1821]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="56" phylum="Chordata" rank="species" species="ctenolepidus">
<emphasis id="B950EAD7A80CFF86C3EAF8857758F89B" box="[657,833,1794,1820]" italics="true" pageId="11" pageNumber="56">B. ctenolepidus</emphasis>
Kessler,
<quantity id="4CDC9B20A80CFF86C2D5F88577FFF89A" box="[942,998,1795,1821]" metricMagnitude="1" metricUnit="m" metricValue="4.76758" pageId="11" pageNumber="56" unit="in" value="1877.0">1877</quantity>
</taxonomicName>
in having the dermal fold rectangular large (
<emphasis id="B950EAD7A80CFF86C19BF8A174E6F8C7" box="[224,255,1831,1856]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
curved large), origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80CFF86C590F8A17110F8C7" box="[1259,1289,1831,1856]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
under origin of the second dorsal fin), tubercles present with two posterior rows of spinules forming an acute, always less than right, angle (
<emphasis id="B950EAD7A80CFF86C05DF8E9755DF80F" box="[294,324,1903,1928]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
about right angle), dorsal row of tubercles complete, 2229 (
<emphasis id="B950EAD7A80CFF86C295F8E97014F80F" box="[1006,1037,1903,1928]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
incomplete dorsal row), transversal suborbital row
<emphasis id="B950EAD7A80CFF86C03AF8147541F82B" box="[321,344,1938,1964]" italics="true" pageId="11" pageNumber="56">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80CFF86C3EDF81476BDF82B" box="[662,676,1938,1964]" italics="true" pageId="11" pageNumber="56">b</emphasis>
(
<emphasis id="B950EAD7A80CFF86C3CEF81576CDF82B" box="[693,724,1939,1964]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80CFF86C2B6F81477C2F82B" box="[973,987,1938,1964]" italics="true" pageId="11" pageNumber="56">b</emphasis>
), and second dorsal fin with I+78 rays (
<emphasis id="B950EAD7A80CFF86C1DAF83174A6F857" box="[161,191,1975,2000]" italics="true" pageId="11" pageNumber="56">vs.</emphasis>
second dorsal fin I+910).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFF117719FEE1" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFF1175D8FF37" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,449,151,177]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFF1175D8FF37" box="[199,449,151,177]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C326FF1176CDFF36" authorityName="Boldyrev &amp; Bogutskaya" authorityYear="2004" box="[605,724,151,177]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="durrelli">
<emphasis id="B950EAD7A80BFF81C326FF1176CDFF36" box="[605,724,151,177]" italics="true" pageId="12" pageNumber="57">B. durrelli</emphasis>
</taxonomicName>
by being distributed in southern Caspian Sea (
<emphasis id="B950EAD7A80BFF81C588FF11710BFF37" box="[1267,1298,151,176]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
distribution in the Taganrog Bay of the Sea of Azov and Don River from mouth upstream to the upper stretch of Tsymlyansk Reservoir) and by having no ventrolateral row of tubercles (
<emphasis id="B950EAD7A80BFF81C23EFF59777AFF7F" box="[837,867,223,248]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
large tubercles present), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80BFF81C0B2FE8575F1FE9B" box="[457,488,259,284]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
present in two radial rows), no granules on flanks (
<emphasis id="B950EAD7A80BFF81C55AFE857026FE9B" box="[1057,1087,259,284]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
sparsely scattered but present), transversal suborbital row
<emphasis id="B950EAD7A80BFF81C0BBFEA075CFFEC7" box="[448,470,294,320]" italics="true" pageId="12" pageNumber="57">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80BFF81C269FEA17739FEC6" box="[786,800,295,321]" italics="true" pageId="12" pageNumber="57">b</emphasis>
(
<emphasis id="B950EAD7A80BFF81C24BFEA17757FEC7" box="[816,846,295,320]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80BFF81C53EFEA1704AFEC6" box="[1093,1107,295,321]" italics="true" pageId="12" pageNumber="57">b</emphasis>
), and the anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C031FECD757FFEE3" box="[330,358,331,356]" italics="true" pageId="12" pageNumber="57">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A80BFF81C33BFECD7647FEE3" box="[576,606,331,356]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
35 papillae).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFEE9761CFDBA" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFEE975A6FE0F" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,447,367,393]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFEE975A6FE0F" box="[199,447,367,393]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C329FEE97770FE0E" authority="Berg, 1949" authorityName="Berg" authorityYear="1949" box="[594,873,367,393]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="leobergius">
<emphasis id="B950EAD7A80BFF81C329FEE976FFFE0E" box="[594,742,367,393]" italics="true" pageId="12" pageNumber="57">B. leobergius</emphasis>
Berg,
<quantity id="4CDC9B20A80BFF81C24AFEE97770FE0E" box="[817,873,367,393]" metricMagnitude="1" metricUnit="m" metricValue="4.95046" pageId="12" pageNumber="57" unit="in" value="1949.0">1949</quantity>
</taxonomicName>
in having the dermal fold rectangular (
<emphasis id="B950EAD7A80BFF81C465FEE97124FE0F" box="[1310,1341,367,392]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
triangular), origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80BFF81C291FE157011FE2B" box="[1002,1032,403,428]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
origin of anal fin under origin of the second dorsal fin), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80BFF81C2CDFE3177CDFE57" box="[950,980,439,464]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
tubercles present), transversal suborbital row
<emphasis id="B950EAD7A80BFF81C1B1FE5C74F9FE73" box="[202,224,474,500]" italics="true" pageId="12" pageNumber="57">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80BFF81C35AFE5D7636FE72" box="[545,559,475,501]" italics="true" pageId="12" pageNumber="57">b</emphasis>
(
<emphasis id="B950EAD7A80BFF81C33BFE5D7647FE73" box="[576,606,475,500]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80BFF81C222FE5D777EFE72" box="[857,871,475,501]" italics="true" pageId="12" pageNumber="57">b</emphasis>
), anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C46EFE5D7128FE73" box="[1301,1329,475,500]" italics="true" pageId="12" pageNumber="57">pa</emphasis>
with 12 papilla (
<emphasis id="B950EAD7A80BFF81C188FE79750BFD9F" box="[243,274,511,536]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
35 papillae), maximum body width 15.122.9% of SL (
<emphasis id="B950EAD7A80BFF81C2F4FE7977B7FD9F" box="[911,942,511,536]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
24.531.1%), and mouth width 36.355.8% of head length (
<emphasis id="B950EAD7A80BFF81C033FDA5757FFDBB" box="[328,358,547,572]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
65.371.3%).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFDC17586FC92" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFDC175A7FDE7" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,446,583,609]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFDC175A7FDE7" box="[199,446,583,609]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C335FDC177BCFDE6" authority="Kessler, 1877" authorityName="Kessler" authorityYear="1877" box="[590,933,583,609]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="leptocephalus">
<emphasis id="B950EAD7A80BFF81C335FDC17711FDE6" box="[590,776,583,609]" italics="true" pageId="12" pageNumber="57">B. leptocephalus</emphasis>
Kessler,
<quantity id="4CDC9B20A80BFF81C216FDC177BCFDE6" box="[877,933,583,609]" metricMagnitude="1" metricUnit="m" metricValue="4.76758" pageId="12" pageNumber="57" unit="in" value="1877.0">1877</quantity>
</taxonomicName>
in having the dermal fold large and rectangular (
<emphasis id="B950EAD7A80BFF81C1BEFDED74FAFD03" box="[197,227,619,644]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
dermal fold absent), a moderate chin barbel, 1/32/3 of eye diameter in length (
<emphasis id="B950EAD7A80BFF81C51BFDED7066FD03" box="[1120,1151,619,644]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
very small barbel, hardly visible or absent), tubercles present with two posterior rows of spinules forming an acute, always less than right, angle (
<emphasis id="B950EAD7A80BFF81C19FFD35751BFD4B" box="[228,258,691,716]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
nearly right angle), a complete dorsal row of tubercles, 2229 (
<emphasis id="B950EAD7A80BFF81C2B7FD3577F3FD4B" box="[972,1002,691,716]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
incomplete dorsal row), anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C0E6FD5175A0FD77" box="[413,441,727,752]" italics="true" pageId="12" pageNumber="57">pp</emphasis>
with 23 papillae (
<emphasis id="B950EAD7A80BFF81C3EDFD5176ADFD77" box="[662,692,727,752]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
single papilla), and second dorsal fin with I+78 rays (
<emphasis id="B950EAD7A80BFF81C453FD51715EFD77" box="[1320,1351,727,752]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
second dorsal fin rays I+911).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFC9977E0FC6A" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFC9975A6FCBF" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,447,799,825]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFC9975A6FCBF" box="[199,447,799,825]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C32FFC9977D9FCBE" authority="(Pallas, 1787)" baseAuthorityName="Pallas" baseAuthorityYear="1787" box="[596,960,799,825]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="macrocephalus">
<emphasis id="B950EAD7A80BFF81C32FFC997704FCBE" box="[596,797,799,825]" italics="true" pageId="12" pageNumber="57">B. macrocephalus</emphasis>
(Pallas, 1787)
</taxonomicName>
in having a moderate chin barbel, 1/32/3 of eye diameter (
<emphasis id="B950EAD7A80BFF81C02EFCC5756DFCDB" box="[341,372,835,860]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
large barbel, about equal eye diameter), the origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80BFF81C0FEFCE175BDFC07" box="[389,420,871,896]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
under origin of the second dorsal fin), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80BFF81C1DAFC0D74A6FC23" box="[161,191,907,932]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
tubercles present), no granules on flanks (
<emphasis id="B950EAD7A80BFF81C3E3FC0D76AFFC23" box="[664,694,907,932]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
thorn like granules on flanks), transversal suborbital row
<emphasis id="B950EAD7A80BFF81C440FC0C714BFC23" box="[1339,1362,906,932]" italics="true" pageId="12" pageNumber="57">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80BFF81C0FCFC29758CFC4E" box="[391,405,943,969]" italics="true" pageId="12" pageNumber="57">b</emphasis>
(
<emphasis id="B950EAD7A80BFF81C0DCFC2975DCFC4F" box="[423,453,943,968]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80BFF81C3B8FC2976C8FC4E" box="[707,721,943,969]" italics="true" pageId="12" pageNumber="57">b</emphasis>
), anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C5F8FC297086FC4F" box="[1155,1183,943,968]" italics="true" pageId="12" pageNumber="57">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A80BFF81C405FC297185FC4F" box="[1406,1436,943,968]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
36 papillae), and mouth width 36.355.8% of head length (
<emphasis id="B950EAD7A80BFF81C247FC557743FC6B" box="[828,858,979,1004]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
56.267.8%).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFC71718DFB41" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFC7175D9FB97" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,448,1015,1041]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFC7175D9FB97" box="[199,448,1015,1041]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C244D46A80BFF81C3C6FC7177AEFB96" authority="Iljin, 1927" authorityName="Iljin" authorityYear="1927" box="[701,951,1015,1041]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="magistri">
<emphasis id="B950EAD7A80BFF81C3C6FC717720FB97" box="[701,825,1015,1040]" italics="true" pageId="12" pageNumber="57">B. magistri</emphasis>
Iljin, 1927
</taxonomicName>
by its distribution in southern Caspian Sea (
<emphasis id="B950EAD7A80BFF81C1DAFB9D74A6FBB3" box="[161,191,1051,1076]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
distribution in the eastern part of the Sea of Azov) and by having the origin of the anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80BFF81C351FBB97651FBDF" box="[554,584,1087,1112]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
origin of anal fin under origin of the second dorsal fin), no ventrolateral row of tubercles (
<emphasis id="B950EAD7A80BFF81C050FBE57553FBFB" box="[299,330,1123,1148]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
large tubercles present), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80BFF81C5C3FBE570CFFBFB" box="[1208,1238,1123,1148]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
very small tubercles present), no granules on flanks (
<emphasis id="B950EAD7A80BFF81C348FB017648FB27" box="[563,593,1159,1184]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
granules present), transversal suborbital row
<emphasis id="B950EAD7A80BFF81C52BFB00707FFB27" box="[1104,1126,1158,1184]" italics="true" pageId="12" pageNumber="57">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80BFF81C1ECFB2D74BCFB42" box="[151,165,1195,1221]" italics="true" pageId="12" pageNumber="57">b</emphasis>
(
<emphasis id="B950EAD7A80BFF81C1CDFB2D74CDFB43" box="[182,212,1195,1220]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80BFF81C0B7FB2D75C3FB42" box="[460,474,1195,1221]" italics="true" pageId="12" pageNumber="57">b</emphasis>
), and the anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C2A5FB2D77E3FB43" box="[990,1018,1195,1220]" italics="true" pageId="12" pageNumber="57">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A80BFF81C5AFFB2D70EBFB43" box="[1236,1266,1195,1220]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
35 papillae).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFB49708BFAD1" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFB4975D8FB6F" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,449,1231,1257]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFB4975D8FB6F" box="[199,449,1231,1257]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C327FB49773BFB6E" authorityName="Ragimov" authorityYear="1976" box="[604,802,1231,1257]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="mahmudbejovi">
<emphasis id="B950EAD7A80BFF81C327FB49773BFB6E" box="[604,802,1231,1257]" italics="true" pageId="12" pageNumber="57">B. mahmudbejovi</emphasis>
</taxonomicName>
by lacking a ventrolateral row of tubercles (
<emphasis id="B950EAD7A80BFF81C45FFB49715BFB6F" box="[1316,1346,1231,1256]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
present with large tubercles), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80BFF81C2A0FB7577E0FA8B" box="[987,1017,1267,1292]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
tubercles present), granules not present on flanks (
<emphasis id="B950EAD7A80BFF81C04CFA91754CFAB7" box="[311,341,1303,1328]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
granules present), transversal suborbital row
<emphasis id="B950EAD7A80BFF81C229FA907771FAB7" box="[850,872,1302,1328]" italics="true" pageId="12" pageNumber="57">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80BFF81C5D8FA9170A8FAB6" box="[1187,1201,1303,1329]" italics="true" pageId="12" pageNumber="57">b</emphasis>
(
<emphasis id="B950EAD7A80BFF81C5BAFA9170C6FAB7" box="[1217,1247,1303,1328]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80BFF81C1B1FABD74C1FAD2" box="[202,216,1339,1365]" italics="true" pageId="12" pageNumber="57">b</emphasis>
), and the anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C3A7FABD76E1FAD3" box="[732,760,1339,1364]" italics="true" pageId="12" pageNumber="57">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A80BFF81C2A9FABD77E9FAD3" box="[978,1008,1339,1364]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
35 papillae).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCFAD977F8F9D5" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCFAD975A7FAFF" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,446,1375,1401]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCFAD975A7FAFF" box="[199,446,1375,1401]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C334FAD97729FAFE" authority="Berg, 1998" authorityName="Berg" authorityYear="1998" box="[591,816,1375,1401]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="nudus">
<emphasis id="B950EAD7A80BFF81C334FAD976A9FAFE" box="[591,688,1375,1401]" italics="true" pageId="12" pageNumber="57">B. nudus</emphasis>
Berg, 1998
</taxonomicName>
by its distribution in southern Caspian Sea (
<emphasis id="B950EAD7A80BFF81C462FAD97121FAFF" box="[1305,1336,1375,1400]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
distribution in the lagoons, lakes and rivers of the north-western Black Sea) and by having the dermal fold rectangular and large (
<emphasis id="B950EAD7A80BFF81C1A5FA2174E4FA47" box="[222,253,1447,1472]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
dermal fold roughly triangular, large), origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80BFF81C1BEFA4D74FDFA63" box="[197,228,1483,1508]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
behind vertical through the origin of the second dorsal fin), no ventrolateral row of tubercles (
<emphasis id="B950EAD7A80BFF81C58BFA4D7117FA63" box="[1264,1294,1483,1508]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
present with large tubercles), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A80BFF81C2D6FA6977D2F98F" box="[941,971,1519,1544]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
tubercles present), no granules on flanks (
<emphasis id="B950EAD7A80BFF81C1DAF99574A6F9AB" box="[161,191,1555,1580]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
granules present), and transversal suborbital row
<emphasis id="B950EAD7A80BFF81C38BF994771EF9AB" box="[752,775,1554,1580]" italics="true" pageId="12" pageNumber="57">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A80BFF81C53DF995704DF9AA" box="[1094,1108,1555,1581]" italics="true" pageId="12" pageNumber="57">b</emphasis>
(
<emphasis id="B950EAD7A80BFF81C51EF995709DF9AB" box="[1125,1156,1555,1580]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A80BFF81C405F9957195F9AA" box="[1406,1420,1555,1581]" italics="true" pageId="12" pageNumber="57">b</emphasis>
), anterior interorbital transversal row
<emphasis id="B950EAD7A80BFF81C350F9B1765EF9D7" box="[555,583,1591,1616]" italics="true" pageId="12" pageNumber="57">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A80BFF81C25AF9B17759F9D7" box="[801,832,1591,1616]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
35 papillae).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCF9DD713AF882" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCF9DD75A6F9F3" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,447,1627,1653]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCF9DD75A6F9F3" box="[199,447,1627,1653]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C32EF9DD76CCF9F2" authorityName="Ragimov" authorityYear="1982" box="[597,725,1627,1653]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="pinchuki">
<emphasis id="B950EAD7A80BFF81C32EF9DD76CCF9F2" box="[597,725,1627,1653]" italics="true" pageId="12" pageNumber="57">B. pinchuki</emphasis>
</taxonomicName>
in having the dermal fold well developed, rectangular (
<emphasis id="B950EAD7A80BFF81C431F9DD7171F9F3" box="[1354,1384,1627,1652]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
dermal fold triangular, small), origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80BFF81C468F9F97128F91F" box="[1299,1329,1663,1688]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
origin of anal fin under origin of the second dorsal fin), tubercles present with two posterior rows of spinules forming an acute, always less than right, angle (
<emphasis id="B950EAD7A80BFF81C357F9417652F967" box="[556,587,1735,1760]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
nearly right angle), dorsal row of tubercles complete, 2229 (
<emphasis id="B950EAD7A80BFF81C47BF9417106F967" box="[1280,1311,1735,1760]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
dorsal row complete, 3133 tubercles), and a second dorsal fin with I+78 rays (
<emphasis id="B950EAD7A80BFF81C2DBF96D77A6F883" box="[928,959,1771,1796]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
second dorsal fin rays I+910).
</paragraph>
<paragraph id="8B9B36C5A80BFF81C1BCF88976D0F83E" blockId="12.[151,1437,151,1977]" pageId="12" pageNumber="57">
<taxonomicName id="4C244D46A80BFF81C1BCF88975DBF8AF" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,450,1807,1833]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="57" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A80BFF81C1BCF88975DBF8AF" box="[199,450,1807,1833]" italics="true" pageId="12" pageNumber="57">Benthophilus persicus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C244D46A80BFF81C31AF88976F0F8AF" authorityName="Boldyrev &amp; Bogutskaya" authorityYear="2004" box="[609,745,1807,1832]" genus="Bogutskaya" pageId="12" pageNumber="57" rank="species" species="ragimovi">
<emphasis id="B950EAD7A80BFF81C31AF88976F0F8AF" box="[609,745,1807,1832]" italics="true" pageId="12" pageNumber="57">B. ragimovi</emphasis>
</taxonomicName>
in having the dermal fold rectangular, large (
<emphasis id="B950EAD7A80BFF81C47AF8897106F8AF" box="[1281,1311,1807,1832]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
triangular, small), the origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A80BFF81C536F8B57075F8CB" box="[1101,1132,1843,1868]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
under origin of the second dorsal fin), tubercles present with two posterior rows of spinules forming an acute, always less than right angle (
<emphasis id="B950EAD7A80BFF81C405F8D17185F8F7" box="[1406,1436,1879,1904]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
nearly right angle), dorsal row of tubercles 2229 (
<emphasis id="B950EAD7A80BFF81C3AAF8FD76E9F813" box="[721,752,1915,1940]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
3033), no ventrolateral row of tubercles (
<emphasis id="B950EAD7A80BFF81C5AAF8FD70F6F813" box="[1233,1263,1915,1940]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
present, large), and the second dorsal fin with I+78 (
<emphasis id="B950EAD7A80BFF81C339F8197678F83F" box="[578,609,1951,1976]" italics="true" pageId="12" pageNumber="57">vs.</emphasis>
I+910).
</paragraph>
<caption id="DF5B664DA80AFF80C1ECFE5B7068FDE4" ID-Table-UUID="DF5B664DA80AFF80C1ECFE5B7068FDE4" httpUri="http://table.plazi.org/id/DF5B664DA80AFF80C1ECFE5B7068FDE4" pageId="13" pageNumber="58" startId="13.[151,239,477,503]" targetBox="[168,2025,656,1650]" targetIsTable="true" targetPageId="13">
<paragraph id="8B9B36C5A80AFF80C1ECFE5B7068FDE4" blockId="13.[151,2037,477,611]" pageId="13" pageNumber="58">
<emphasis id="B950EAD7A80AFF80C1ECFE5B7512FE70" bold="true" box="[151,267,477,503]" pageId="13" pageNumber="58">TABLE 1.</emphasis>
<taxonomicName id="4C244D46A80AFF80C069FE5B75E6FE70" authority="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen, 2021" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[274,511,477,503]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="58" phylum="Chordata" rank="species" species="persicus" status="sp. nov.">
<emphasis id="B950EAD7A80AFF80C069FE5B75E6FE70" box="[274,511,477,503]" italics="true" pageId="13" pageNumber="58">Benthophilus persicus</emphasis>
</taxonomicName>
<emphasis id="B950EAD7A80AFF80C37DFE5B7644FE70" bold="true" box="[518,605,477,503]" pageId="13" pageNumber="58">
<taxonomicNameLabel id="A26357ACA80AFF80C37DFE5B7644FE70" box="[518,605,477,503]" pageId="13" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Morphometric characters as % of standard length or as % of head length. Characters are sorted as in
<bibRefCitation id="EFB54B34A80AFF80C7D9FE5B73EDFE70" author="Boldyrev, V. S. &amp; Bogutskaya, N. G." box="[1698,2036,477,503]" pageId="13" pageNumber="58" pagination="31 - 96" refId="ref11847" refString="Boldyrev, V. S. &amp; Bogutskaya, N. G. (2007) Revision of the tadpole-gobies of the genus Benthophilus (Teleostei: Gobiidae). Ichthyological Exploration of Freshwaters, 18 (1), 31 - 96." type="journal article" year="2007">Boldyrev &amp; Bogutskaya (2007)</bibRefCitation>
with the new introduced characters added at the end in alphabetic order. The morphometric character and the related values of dimorphic characters with non-overlapping ranges of groups marked with ^ for adult males
<emphasis id="B950EAD7A80AFF80C335FDA37675FDB8" box="[590,620,549,575]" italics="true" pageId="13" pageNumber="58">vs.</emphasis>
the immature male and all females, with * for all males and the large female
<emphasis id="B950EAD7A80AFF80C4D5FDA371D5FDB8" box="[1454,1484,549,575]" italics="true" pageId="13" pageNumber="58">vs.</emphasis>
small mature and immature females, and with ˜ for adult males and the large female
<emphasis id="B950EAD7A80AFF80C08CFDCF760DFDE4" box="[503,532,585,611]" italics="true" pageId="13" pageNumber="58">vs.</emphasis>
smaller females and the immature male, see Discussion.
</paragraph>
</caption>
<paragraph id="8B9B36C5A80AFF80C1CFFD4C73CFF9D2" pageId="13" pageNumber="58">
<table id="F924C465A80A0072C1D3FD1673F0F9F5" box="[168,2025,656,1650]" gridcols="17" gridrows="11" pageId="13" pageNumber="58">
<tr id="35143487A80A0072C1D3FD1673F0FC9C" box="[168,2025,656,795]" gridrow="0" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FD167500FC9C" box="[168,281,656,795]" gridcol="0" gridrow="0" pageId="13" pageNumber="58">Specimen</th>
<th id="76C55DFBA80A0072C043FD16759FFC9C" box="[312,390,656,795]" gridcol="1" gridrow="0" pageId="13" pageNumber="58">ZSM 47595 holotype</th>
<th id="76C55DFBA80A0072C0DDFD1675EAFC9C" box="[422,499,656,795]" gridcol="2" gridrow="0" pageId="13" pageNumber="58">ZSM 47596 paratype</th>
<th id="76C55DFBA80A0072C360FD167671FC9C" box="[539,616,656,795]" gridcol="3" gridrow="0" pageId="13" pageNumber="58">ZSM 47597 paratype</th>
<th id="76C55DFBA80A0072C3F5FD1676C2FC9C" box="[654,731,656,795]" gridcol="4" gridrow="0" pageId="13" pageNumber="58">ZSM 47599 paratype</th>
<th id="76C55DFBA80A0072C382FD16775FFC9C" box="[761,838,656,795]" gridcol="5" gridrow="0" pageId="13" pageNumber="58">
ZSM 47598
<typeStatus id="549F8867A80AFF80C382FD6E775FFD79" box="[761,838,744,766]" pageId="13" pageNumber="58" type="paratype">paratype</typeStatus>
</th>
<th id="76C55DFBA80A0072C21FFD1677A8FC9C" box="[868,945,656,795]" gridcol="6" gridrow="0" pageId="13" pageNumber="58">
ZM - CBSU
<uuid id="FF820C10A80AFF80C20BFD4C7787FD67" box="[880,926,714,736]" pageId="13" pageNumber="58">5003</uuid>
- 128 paratype
</th>
<th id="76C55DFBA80A0072C2AAFD167007FC9C" box="[977,1054,656,795]" gridcol="7" gridrow="0" pageId="13" pageNumber="58">ZM - CBSU 5001-1 paratype</th>
<th id="76C55DFBA80A0072C545FD167092FC9C" box="[1086,1163,656,795]" gridcol="8" gridrow="0" pageId="13" pageNumber="58">ZM - CBSU 5003-60 paratype</th>
<th id="76C55DFBA80A0072C5D0FD1670E1FC9C" box="[1195,1272,656,795]" gridcol="9" gridrow="0" pageId="13" pageNumber="58">ZM - CBSU 5022-23 paratype</th>
<th id="76C55DFBA80A0072C463FD16717CFC9C" box="[1304,1381,656,795]" gridcol="10" gridrow="0" pageId="13" pageNumber="58">ZM - CBSU 5024-1 paratype</th>
<th id="76C55DFBA80A0072C4F8FD1671C9FC9C" box="[1411,1488,656,795]" gridcol="11" gridrow="0" pageId="13" pageNumber="58">
ZM - CBSU 5003-
<specimenCount id="9D22FD4CA80AFF80C4C3FD5F71D6FD68" box="[1464,1487,729,751]" pageId="13" pageNumber="58" type="generic" typeStatus="holotype">77</specimenCount>
paratype
</th>
<th id="76C55DFBA80A0072C496FD167223FC9C" box="[1517,1594,656,795]" gridcol="12" gridrow="0" pageId="13" pageNumber="58">PMR VP4679 paratype</th>
<th id="76C55DFBA80A0072C723FD1672BCFC9C" box="[1624,1701,656,795]" gridcol="13" gridrow="0" pageId="13" pageNumber="58">
PMR VP4680
<specimenCount id="9D22FD4CA80AFF80C723FD6E72BCFD79" box="[1624,1701,744,766]" pageId="13" pageNumber="58" type="generic" typeStatus="paratype">paratype</specimenCount>
</th>
<th id="76C55DFBA80A0072C7B9FD167316FC9C" box="[1730,1807,656,795]" gridcol="14" gridrow="0" pageId="13" pageNumber="58">PMR VP4681 paratype</th>
<th id="76C55DFBA80A0072C657FD167360FC9C" box="[1836,1913,656,795]" gridcol="15" gridrow="0" pageId="13" pageNumber="58">PMR VP4682 paratype</th>
<th id="76C55DFBA80A0072C6E7FD1673F0FC9C" box="[1948,2025,656,795]" gridcol="16" gridrow="0" pageId="13" pageNumber="58">
PMR VP4683
<typeStatus id="549F8867A80AFF80C6E7FD6E73F0FD79" box="[1948,2025,744,766]" pageId="13" pageNumber="58" type="paratype">paratype</typeStatus>
</th>
</tr>
<tr id="35143487A80A0072C1D3FCAE73F0FCDB" box="[168,2025,808,860]" gridrow="1" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FCAE7500FCDB" box="[168,281,808,860]" gridcol="0" gridrow="1" pageId="13" pageNumber="58">Sex</th>
<td id="76C55DFBA80A0072C043FCAE759FFCDB" box="[312,390,808,860]" gridcol="1" gridrow="1" pageId="13" pageNumber="58">male</td>
<td id="76C55DFBA80A0072C0DDFCAE75EAFCDB" box="[422,499,808,860]" gridcol="2" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C360FCAE7671FCDB" box="[539,616,808,860]" gridcol="3" gridrow="1" pageId="13" pageNumber="58">juvenile male</td>
<td id="76C55DFBA80A0072C3F5FCAE76C2FCDB" box="[654,731,808,860]" gridcol="4" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C382FCAE775FFCDB" box="[761,838,808,860]" gridcol="5" gridrow="1" pageId="13" pageNumber="58">juvenile female</td>
<td id="76C55DFBA80A0072C21FFCAE77A8FCDB" box="[868,945,808,860]" gridcol="6" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C2AAFCAE7007FCDB" box="[977,1054,808,860]" gridcol="7" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C545FCAE7092FCDB" box="[1086,1163,808,860]" gridcol="8" gridrow="1" pageId="13" pageNumber="58">juvenile female</td>
<td id="76C55DFBA80A0072C5D0FCAE70E1FCDB" box="[1195,1272,808,860]" gridcol="9" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C463FCAE717CFCDB" box="[1304,1381,808,860]" gridcol="10" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C4F8FCAE71C9FCDB" box="[1411,1488,808,860]" gridcol="11" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C496FCAE7223FCDB" box="[1517,1594,808,860]" gridcol="12" gridrow="1" pageId="13" pageNumber="58">male</td>
<td id="76C55DFBA80A0072C723FCAE72BCFCDB" box="[1624,1701,808,860]" gridcol="13" gridrow="1" pageId="13" pageNumber="58">male</td>
<td id="76C55DFBA80A0072C7B9FCAE7316FCDB" box="[1730,1807,808,860]" gridcol="14" gridrow="1" pageId="13" pageNumber="58">male</td>
<td id="76C55DFBA80A0072C657FCAE7360FCDB" box="[1836,1913,808,860]" gridcol="15" gridrow="1" pageId="13" pageNumber="58">female</td>
<td id="76C55DFBA80A0072C6E7FCAE73F0FCDB" box="[1948,2025,808,860]" gridcol="16" gridrow="1" pageId="13" pageNumber="58">female</td>
</tr>
<tr id="35143487A80A0072C1D3FCE573F0FC33" box="[168,2025,867,948]" gridrow="2" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FCE57500FC33" box="[168,281,867,948]" gridcol="0" gridrow="2" pageId="13" pageNumber="58">Standard length (SL) inmm</th>
<td id="76C55DFBA80A0072C043FCE5759FFC33" box="[312,390,867,948]" gridcol="1" gridrow="2" pageId="13" pageNumber="58">45.2</td>
<td id="76C55DFBA80A0072C0DDFCE575EAFC33" box="[422,499,867,948]" gridcol="2" gridrow="2" pageId="13" pageNumber="58">45.8</td>
<td id="76C55DFBA80A0072C360FCE57671FC33" box="[539,616,867,948]" gridcol="3" gridrow="2" pageId="13" pageNumber="58">30.4</td>
<td id="76C55DFBA80A0072C3F5FCE576C2FC33" box="[654,731,867,948]" gridcol="4" gridrow="2" pageId="13" pageNumber="58">30.8</td>
<td id="76C55DFBA80A0072C382FCE5775FFC33" box="[761,838,867,948]" gridcol="5" gridrow="2" pageId="13" pageNumber="58">27.5</td>
<td id="76C55DFBA80A0072C21FFCE577A8FC33" box="[868,945,867,948]" gridcol="6" gridrow="2" pageId="13" pageNumber="58">30.9</td>
<td id="76C55DFBA80A0072C2AAFCE57007FC33" box="[977,1054,867,948]" gridcol="7" gridrow="2" pageId="13" pageNumber="58">35.1</td>
<td id="76C55DFBA80A0072C545FCE57092FC33" box="[1086,1163,867,948]" gridcol="8" gridrow="2" pageId="13" pageNumber="58">25.4</td>
<td id="76C55DFBA80A0072C5D0FCE570E1FC33" box="[1195,1272,867,948]" gridcol="9" gridrow="2" pageId="13" pageNumber="58">26.0</td>
<td id="76C55DFBA80A0072C463FCE5717CFC33" box="[1304,1381,867,948]" gridcol="10" gridrow="2" pageId="13" pageNumber="58">23.7</td>
<td id="76C55DFBA80A0072C4F8FCE571C9FC33" box="[1411,1488,867,948]" gridcol="11" gridrow="2" pageId="13" pageNumber="58">25.1</td>
<td id="76C55DFBA80A0072C496FCE57223FC33" box="[1517,1594,867,948]" gridcol="12" gridrow="2" pageId="13" pageNumber="58">43.3</td>
<td id="76C55DFBA80A0072C723FCE572BCFC33" box="[1624,1701,867,948]" gridcol="13" gridrow="2" pageId="13" pageNumber="58">47.0</td>
<td id="76C55DFBA80A0072C7B9FCE57316FC33" box="[1730,1807,867,948]" gridcol="14" gridrow="2" pageId="13" pageNumber="58">35.1</td>
<td id="76C55DFBA80A0072C657FCE57360FC33" box="[1836,1913,867,948]" gridcol="15" gridrow="2" pageId="13" pageNumber="58">30.4</td>
<td id="76C55DFBA80A0072C6E7FCE573F0FC33" box="[1948,2025,867,948]" gridcol="16" gridrow="2" pageId="13" pageNumber="58">34.2</td>
</tr>
<tr id="35143487A80A0072C1D3FC4C73F0FC67" box="[168,2025,970,992]" gridrow="3" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FC4C73F0FC67" box="[168,2025,970,992]" colspan="17" colspanRight="16" gridcol="0" gridrow="3" pageId="13" pageNumber="58">% ofSL</th>
</tr>
<tr id="35143487A80A0072C1D3FB8573F0FBB1" box="[168,2025,1027,1078]" gridrow="4" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FB857500FBB1" box="[168,281,1027,1078]" gridcol="0" gridrow="4" pageId="13" pageNumber="58">Maximum body depth*</th>
<td id="76C55DFBA80A0072C043FB85759FFBB1" box="[312,390,1027,1078]" gridcol="1" gridrow="4" pageId="13" pageNumber="58">22.2*</td>
<td id="76C55DFBA80A0072C0DDFB8575EAFBB1" box="[422,499,1027,1078]" gridcol="2" gridrow="4" pageId="13" pageNumber="58">19.8*</td>
<td id="76C55DFBA80A0072C360FB857671FBB1" box="[539,616,1027,1078]" gridcol="3" gridrow="4" pageId="13" pageNumber="58">21.0*</td>
<td id="76C55DFBA80A0072C3F5FB8576C2FBB1" box="[654,731,1027,1078]" gridcol="4" gridrow="4" pageId="13" pageNumber="58">14.6</td>
<td id="76C55DFBA80A0072C382FB85775FFBB1" box="[761,838,1027,1078]" gridcol="5" gridrow="4" pageId="13" pageNumber="58">14.6</td>
<td id="76C55DFBA80A0072C21FFB8577A8FBB1" box="[868,945,1027,1078]" gridcol="6" gridrow="4" pageId="13" pageNumber="58">17.6</td>
<td id="76C55DFBA80A0072C2AAFB857007FBB1" box="[977,1054,1027,1078]" gridcol="7" gridrow="4" pageId="13" pageNumber="58">16.7</td>
<td id="76C55DFBA80A0072C545FB857092FBB1" box="[1086,1163,1027,1078]" gridcol="8" gridrow="4" pageId="13" pageNumber="58">16.9</td>
<td id="76C55DFBA80A0072C5D0FB8570E1FBB1" box="[1195,1272,1027,1078]" gridcol="9" gridrow="4" pageId="13" pageNumber="58">15.6</td>
<td id="76C55DFBA80A0072C463FB85717CFBB1" box="[1304,1381,1027,1078]" gridcol="10" gridrow="4" pageId="13" pageNumber="58">17.0</td>
<td id="76C55DFBA80A0072C4F8FB8571C9FBB1" box="[1411,1488,1027,1078]" gridcol="11" gridrow="4" pageId="13" pageNumber="58">16.5</td>
<td id="76C55DFBA80A0072C496FB857223FBB1" box="[1517,1594,1027,1078]" gridcol="12" gridrow="4" pageId="13" pageNumber="58">19.7*</td>
<td id="76C55DFBA80A0072C723FB8572BCFBB1" box="[1624,1701,1027,1078]" gridcol="13" gridrow="4" pageId="13" pageNumber="58">20.7*</td>
<td id="76C55DFBA80A0072C7B9FB857316FBB1" box="[1730,1807,1027,1078]" gridcol="14" gridrow="4" pageId="13" pageNumber="58">22.8*</td>
<td id="76C55DFBA80A0072C657FB857360FBB1" box="[1836,1913,1027,1078]" gridcol="15" gridrow="4" pageId="13" pageNumber="58">16.6</td>
<td id="76C55DFBA80A0072C6E7FB8573F0FBB1" box="[1948,2025,1027,1078]" gridcol="16" gridrow="4" pageId="13" pageNumber="58">14.6</td>
</tr>
<tr id="35143487A80A0072C1D3FBCD73F0FBF9" box="[168,2025,1099,1150]" gridrow="5" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FBCD7500FBF9" box="[168,281,1099,1150]" gridcol="0" gridrow="5" pageId="13" pageNumber="58">Maximum body width*</th>
<td id="76C55DFBA80A0072C043FBCD759FFBF9" box="[312,390,1099,1150]" gridcol="1" gridrow="5" pageId="13" pageNumber="58">22.6*</td>
<td id="76C55DFBA80A0072C0DDFBCD75EAFBF9" box="[422,499,1099,1150]" gridcol="2" gridrow="5" pageId="13" pageNumber="58">22.9*</td>
<td id="76C55DFBA80A0072C360FBCD7671FBF9" box="[539,616,1099,1150]" gridcol="3" gridrow="5" pageId="13" pageNumber="58">20.9*</td>
<td id="76C55DFBA80A0072C3F5FBCD76C2FBF9" box="[654,731,1099,1150]" gridcol="4" gridrow="5" pageId="13" pageNumber="58">16.7</td>
<td id="76C55DFBA80A0072C382FBCD775FFBF9" box="[761,838,1099,1150]" gridcol="5" gridrow="5" pageId="13" pageNumber="58">16.0</td>
<td id="76C55DFBA80A0072C21FFBCD77A8FBF9" box="[868,945,1099,1150]" gridcol="6" gridrow="5" pageId="13" pageNumber="58">15.1</td>
<td id="76C55DFBA80A0072C2AAFBCD7007FBF9" box="[977,1054,1099,1150]" gridcol="7" gridrow="5" pageId="13" pageNumber="58">18.0</td>
<td id="76C55DFBA80A0072C545FBCD7092FBF9" box="[1086,1163,1099,1150]" gridcol="8" gridrow="5" pageId="13" pageNumber="58">18.5</td>
<td id="76C55DFBA80A0072C5D0FBCD70E1FBF9" box="[1195,1272,1099,1150]" gridcol="9" gridrow="5" pageId="13" pageNumber="58">17.6</td>
<td id="76C55DFBA80A0072C463FBCD717CFBF9" box="[1304,1381,1099,1150]" gridcol="10" gridrow="5" pageId="13" pageNumber="58">17.5</td>
<td id="76C55DFBA80A0072C4F8FBCD71C9FBF9" box="[1411,1488,1099,1150]" gridcol="11" gridrow="5" pageId="13" pageNumber="58">18.9</td>
<td id="76C55DFBA80A0072C496FBCD7223FBF9" box="[1517,1594,1099,1150]" gridcol="12" gridrow="5" pageId="13" pageNumber="58">19.2*</td>
<td id="76C55DFBA80A0072C723FBCD72BCFBF9" box="[1624,1701,1099,1150]" gridcol="13" gridrow="5" pageId="13" pageNumber="58">20.0*</td>
<td id="76C55DFBA80A0072C7B9FBCD7316FBF9" box="[1730,1807,1099,1150]" gridcol="14" gridrow="5" pageId="13" pageNumber="58">21.9*</td>
<td id="76C55DFBA80A0072C657FBCD7360FBF9" box="[1836,1913,1099,1150]" gridcol="15" gridrow="5" pageId="13" pageNumber="58">18.4</td>
<td id="76C55DFBA80A0072C6E7FBCD73F0FBF9" box="[1948,2025,1099,1150]" gridcol="16" gridrow="5" pageId="13" pageNumber="58">17.2</td>
</tr>
<tr id="35143487A80A0072C1D3FB1073F0FB61" box="[168,2025,1174,1254]" gridrow="6" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FB107500FB61" box="[168,281,1174,1254]" gridcol="0" gridrow="6" pageId="13" pageNumber="58">Depth of caudal peduncle</th>
<td id="76C55DFBA80A0072C043FB10759FFB61" box="[312,390,1174,1254]" gridcol="1" gridrow="6" pageId="13" pageNumber="58">5.8</td>
<td id="76C55DFBA80A0072C0DDFB1075EAFB61" box="[422,499,1174,1254]" gridcol="2" gridrow="6" pageId="13" pageNumber="58">6.3</td>
<td id="76C55DFBA80A0072C360FB107671FB61" box="[539,616,1174,1254]" gridcol="3" gridrow="6" pageId="13" pageNumber="58">6.7</td>
<td id="76C55DFBA80A0072C3F5FB1076C2FB61" box="[654,731,1174,1254]" gridcol="4" gridrow="6" pageId="13" pageNumber="58">5.7</td>
<td id="76C55DFBA80A0072C382FB10775FFB61" box="[761,838,1174,1254]" gridcol="5" gridrow="6" pageId="13" pageNumber="58">5.4</td>
<td id="76C55DFBA80A0072C21FFB1077A8FB61" box="[868,945,1174,1254]" gridcol="6" gridrow="6" pageId="13" pageNumber="58">5.3</td>
<td id="76C55DFBA80A0072C2AAFB107007FB61" box="[977,1054,1174,1254]" gridcol="7" gridrow="6" pageId="13" pageNumber="58">6.0</td>
<td id="76C55DFBA80A0072C545FB107092FB61" box="[1086,1163,1174,1254]" gridcol="8" gridrow="6" pageId="13" pageNumber="58">5.9</td>
<td id="76C55DFBA80A0072C5D0FB1070E1FB61" box="[1195,1272,1174,1254]" gridcol="9" gridrow="6" pageId="13" pageNumber="58">5.5</td>
<td id="76C55DFBA80A0072C463FB10717CFB61" box="[1304,1381,1174,1254]" gridcol="10" gridrow="6" pageId="13" pageNumber="58">5.6</td>
<td id="76C55DFBA80A0072C4F8FB1071C9FB61" box="[1411,1488,1174,1254]" gridcol="11" gridrow="6" pageId="13" pageNumber="58">5.5</td>
<td id="76C55DFBA80A0072C496FB107223FB61" box="[1517,1594,1174,1254]" gridcol="12" gridrow="6" pageId="13" pageNumber="58">6.5</td>
<td id="76C55DFBA80A0072C723FB1072BCFB61" box="[1624,1701,1174,1254]" gridcol="13" gridrow="6" pageId="13" pageNumber="58">6.3</td>
<td id="76C55DFBA80A0072C7B9FB107316FB61" box="[1730,1807,1174,1254]" gridcol="14" gridrow="6" pageId="13" pageNumber="58">5.4</td>
<td id="76C55DFBA80A0072C657FB107360FB61" box="[1836,1913,1174,1254]" gridcol="15" gridrow="6" pageId="13" pageNumber="58">5.7</td>
<td id="76C55DFBA80A0072C6E7FB1073F0FB61" box="[1948,2025,1174,1254]" gridcol="16" gridrow="6" pageId="13" pageNumber="58">5.2</td>
</tr>
<tr id="35143487A80A0072C1D3FA8773F0FAD5" box="[168,2025,1281,1362]" gridrow="7" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FA877500FAD5" box="[168,281,1281,1362]" gridcol="0" gridrow="7" pageId="13" pageNumber="58">Width of caudal peduncle</th>
<td id="76C55DFBA80A0072C043FA87759FFAD5" box="[312,390,1281,1362]" gridcol="1" gridrow="7" pageId="13" pageNumber="58">4.5</td>
<td id="76C55DFBA80A0072C0DDFA8775EAFAD5" box="[422,499,1281,1362]" gridcol="2" gridrow="7" pageId="13" pageNumber="58">4.3</td>
<td id="76C55DFBA80A0072C360FA877671FAD5" box="[539,616,1281,1362]" gridcol="3" gridrow="7" pageId="13" pageNumber="58">4.6</td>
<td id="76C55DFBA80A0072C3F5FA8776C2FAD5" box="[654,731,1281,1362]" gridcol="4" gridrow="7" pageId="13" pageNumber="58">3.9</td>
<td id="76C55DFBA80A0072C382FA87775FFAD5" box="[761,838,1281,1362]" gridcol="5" gridrow="7" pageId="13" pageNumber="58">3.9</td>
<td id="76C55DFBA80A0072C21FFA8777A8FAD5" box="[868,945,1281,1362]" gridcol="6" gridrow="7" pageId="13" pageNumber="58">3.9</td>
<td id="76C55DFBA80A0072C2AAFA877007FAD5" box="[977,1054,1281,1362]" gridcol="7" gridrow="7" pageId="13" pageNumber="58">4.2</td>
<td id="76C55DFBA80A0072C545FA877092FAD5" box="[1086,1163,1281,1362]" gridcol="8" gridrow="7" pageId="13" pageNumber="58">5.0</td>
<td id="76C55DFBA80A0072C5D0FA8770E1FAD5" box="[1195,1272,1281,1362]" gridcol="9" gridrow="7" pageId="13" pageNumber="58">4.0</td>
<td id="76C55DFBA80A0072C463FA87717CFAD5" box="[1304,1381,1281,1362]" gridcol="10" gridrow="7" pageId="13" pageNumber="58">4.0</td>
<td id="76C55DFBA80A0072C4F8FA8771C9FAD5" box="[1411,1488,1281,1362]" gridcol="11" gridrow="7" pageId="13" pageNumber="58">3.9</td>
<td id="76C55DFBA80A0072C496FA877223FAD5" box="[1517,1594,1281,1362]" gridcol="12" gridrow="7" pageId="13" pageNumber="58">4.3</td>
<td id="76C55DFBA80A0072C723FA8772BCFAD5" box="[1624,1701,1281,1362]" gridcol="13" gridrow="7" pageId="13" pageNumber="58">4.2</td>
<td id="76C55DFBA80A0072C7B9FA877316FAD5" box="[1730,1807,1281,1362]" gridcol="14" gridrow="7" pageId="13" pageNumber="58">4.0</td>
<td id="76C55DFBA80A0072C657FA877360FAD5" box="[1836,1913,1281,1362]" gridcol="15" gridrow="7" pageId="13" pageNumber="58">4.3</td>
<td id="76C55DFBA80A0072C6E7FA8773F0FAD5" box="[1948,2025,1281,1362]" gridcol="16" gridrow="7" pageId="13" pageNumber="58">3.9</td>
</tr>
<tr id="35143487A80A0072C1D3FADA73F0FA2B" box="[168,2025,1372,1452]" gridrow="8" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FADA7500FA2B" box="[168,281,1372,1452]" gridcol="0" gridrow="8" pageId="13" pageNumber="58">First predorsal distance</th>
<td id="76C55DFBA80A0072C043FADA759FFA2B" box="[312,390,1372,1452]" gridcol="1" gridrow="8" pageId="13" pageNumber="58">40.1</td>
<td id="76C55DFBA80A0072C0DDFADA75EAFA2B" box="[422,499,1372,1452]" gridcol="2" gridrow="8" pageId="13" pageNumber="58">42.3</td>
<td id="76C55DFBA80A0072C360FADA7671FA2B" box="[539,616,1372,1452]" gridcol="3" gridrow="8" pageId="13" pageNumber="58">42.8</td>
<td id="76C55DFBA80A0072C3F5FADA76C2FA2B" box="[654,731,1372,1452]" gridcol="4" gridrow="8" pageId="13" pageNumber="58">39.8</td>
<td id="76C55DFBA80A0072C382FADA775FFA2B" box="[761,838,1372,1452]" gridcol="5" gridrow="8" pageId="13" pageNumber="58">39.3</td>
<td id="76C55DFBA80A0072C21FFADA77A8FA2B" box="[868,945,1372,1452]" gridcol="6" gridrow="8" pageId="13" pageNumber="58">40.7</td>
<td id="76C55DFBA80A0072C2AAFADA7007FA2B" box="[977,1054,1372,1452]" gridcol="7" gridrow="8" pageId="13" pageNumber="58">41.3</td>
<td id="76C55DFBA80A0072C545FADA7092FA2B" box="[1086,1163,1372,1452]" gridcol="8" gridrow="8" pageId="13" pageNumber="58">39.3</td>
<td id="76C55DFBA80A0072C5D0FADA70E1FA2B" box="[1195,1272,1372,1452]" gridcol="9" gridrow="8" pageId="13" pageNumber="58">39.5</td>
<td id="76C55DFBA80A0072C463FADA717CFA2B" box="[1304,1381,1372,1452]" gridcol="10" gridrow="8" pageId="13" pageNumber="58">39.7</td>
<td id="76C55DFBA80A0072C4F8FADA71C9FA2B" box="[1411,1488,1372,1452]" gridcol="11" gridrow="8" pageId="13" pageNumber="58">39.6</td>
<td id="76C55DFBA80A0072C496FADA7223FA2B" box="[1517,1594,1372,1452]" gridcol="12" gridrow="8" pageId="13" pageNumber="58">39.9</td>
<td id="76C55DFBA80A0072C723FADA72BCFA2B" box="[1624,1701,1372,1452]" gridcol="13" gridrow="8" pageId="13" pageNumber="58">41.1</td>
<td id="76C55DFBA80A0072C7B9FADA7316FA2B" box="[1730,1807,1372,1452]" gridcol="14" gridrow="8" pageId="13" pageNumber="58">41.2</td>
<td id="76C55DFBA80A0072C657FADA7360FA2B" box="[1836,1913,1372,1452]" gridcol="15" gridrow="8" pageId="13" pageNumber="58">38.9</td>
<td id="76C55DFBA80A0072C6E7FADA73F0FA2B" box="[1948,2025,1372,1452]" gridcol="16" gridrow="8" pageId="13" pageNumber="58">42.7</td>
</tr>
<tr id="35143487A80A0072C1D3FA3073F0F980" box="[168,2025,1462,1543]" gridrow="9" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3FA307500F980" box="[168,281,1462,1543]" gridcol="0" gridrow="9" pageId="13" pageNumber="58">Second predorsal distance</th>
<td id="76C55DFBA80A0072C043FA30759FF980" box="[312,390,1462,1543]" gridcol="1" gridrow="9" pageId="13" pageNumber="58">67.1</td>
<td id="76C55DFBA80A0072C0DDFA3075EAF980" box="[422,499,1462,1543]" gridcol="2" gridrow="9" pageId="13" pageNumber="58">68.0</td>
<td id="76C55DFBA80A0072C360FA307671F980" box="[539,616,1462,1543]" gridcol="3" gridrow="9" pageId="13" pageNumber="58">61.7</td>
<td id="76C55DFBA80A0072C3F5FA3076C2F980" box="[654,731,1462,1543]" gridcol="4" gridrow="9" pageId="13" pageNumber="58">63.2</td>
<td id="76C55DFBA80A0072C382FA30775FF980" box="[761,838,1462,1543]" gridcol="5" gridrow="9" pageId="13" pageNumber="58">62.1</td>
<td id="76C55DFBA80A0072C21FFA3077A8F980" box="[868,945,1462,1543]" gridcol="6" gridrow="9" pageId="13" pageNumber="58">63.0</td>
<td id="76C55DFBA80A0072C2AAFA307007F980" box="[977,1054,1462,1543]" gridcol="7" gridrow="9" pageId="13" pageNumber="58">65.1</td>
<td id="76C55DFBA80A0072C545FA307092F980" box="[1086,1163,1462,1543]" gridcol="8" gridrow="9" pageId="13" pageNumber="58">62.1</td>
<td id="76C55DFBA80A0072C5D0FA3070E1F980" box="[1195,1272,1462,1543]" gridcol="9" gridrow="9" pageId="13" pageNumber="58">62.0</td>
<td id="76C55DFBA80A0072C463FA30717CF980" box="[1304,1381,1462,1543]" gridcol="10" gridrow="9" pageId="13" pageNumber="58">60.4</td>
<td id="76C55DFBA80A0072C4F8FA3071C9F980" box="[1411,1488,1462,1543]" gridcol="11" gridrow="9" pageId="13" pageNumber="58">61.9</td>
<td id="76C55DFBA80A0072C496FA307223F980" box="[1517,1594,1462,1543]" gridcol="12" gridrow="9" pageId="13" pageNumber="58">66.6</td>
<td id="76C55DFBA80A0072C723FA3072BCF980" box="[1624,1701,1462,1543]" gridcol="13" gridrow="9" pageId="13" pageNumber="58">65.0</td>
<td id="76C55DFBA80A0072C7B9FA307316F980" box="[1730,1807,1462,1543]" gridcol="14" gridrow="9" pageId="13" pageNumber="58">66.2</td>
<td id="76C55DFBA80A0072C657FA307360F980" box="[1836,1913,1462,1543]" gridcol="15" gridrow="9" pageId="13" pageNumber="58">63.3</td>
<td id="76C55DFBA80A0072C6E7FA3073F0F980" box="[1948,2025,1462,1543]" gridcol="16" gridrow="9" pageId="13" pageNumber="58">64.3</td>
</tr>
<tr id="35143487A80A0072C1D3F9A473F0F9F5" box="[168,2025,1570,1650]" gridrow="10" pageId="13" pageNumber="58">
<th id="76C55DFBA80A0072C1D3F9A47500F9F5" box="[168,281,1570,1650]" gridcol="0" gridrow="10" pageId="13" pageNumber="58">Caudal peduncle length</th>
<td id="76C55DFBA80A0072C043F9A4759FF9F5" box="[312,390,1570,1650]" gridcol="1" gridrow="10" pageId="13" pageNumber="58">18.0</td>
<td id="76C55DFBA80A0072C0DDF9A475EAF9F5" box="[422,499,1570,1650]" gridcol="2" gridrow="10" pageId="13" pageNumber="58">17.9</td>
<td id="76C55DFBA80A0072C360F9A47671F9F5" box="[539,616,1570,1650]" gridcol="3" gridrow="10" pageId="13" pageNumber="58">19.9</td>
<td id="76C55DFBA80A0072C3F5F9A476C2F9F5" box="[654,731,1570,1650]" gridcol="4" gridrow="10" pageId="13" pageNumber="58">20.5</td>
<td id="76C55DFBA80A0072C382F9A4775FF9F5" box="[761,838,1570,1650]" gridcol="5" gridrow="10" pageId="13" pageNumber="58">19.5</td>
<td id="76C55DFBA80A0072C21FF9A477A8F9F5" box="[868,945,1570,1650]" gridcol="6" gridrow="10" pageId="13" pageNumber="58">19.4</td>
<td id="76C55DFBA80A0072C2AAF9A47007F9F5" box="[977,1054,1570,1650]" gridcol="7" gridrow="10" pageId="13" pageNumber="58">20.3</td>
<td id="76C55DFBA80A0072C545F9A47092F9F5" box="[1086,1163,1570,1650]" gridcol="8" gridrow="10" pageId="13" pageNumber="58">20.0</td>
<td id="76C55DFBA80A0072C5D0F9A470E1F9F5" box="[1195,1272,1570,1650]" gridcol="9" gridrow="10" pageId="13" pageNumber="58">19.1</td>
<td id="76C55DFBA80A0072C463F9A4717CF9F5" box="[1304,1381,1570,1650]" gridcol="10" gridrow="10" pageId="13" pageNumber="58">18.2</td>
<td id="76C55DFBA80A0072C4F8F9A471C9F9F5" box="[1411,1488,1570,1650]" gridcol="11" gridrow="10" pageId="13" pageNumber="58">20.7</td>
<td id="76C55DFBA80A0072C496F9A47223F9F5" box="[1517,1594,1570,1650]" gridcol="12" gridrow="10" pageId="13" pageNumber="58">18.1</td>
<td id="76C55DFBA80A0072C723F9A472BCF9F5" box="[1624,1701,1570,1650]" gridcol="13" gridrow="10" pageId="13" pageNumber="58">20.3</td>
<td id="76C55DFBA80A0072C7B9F9A47316F9F5" box="[1730,1807,1570,1650]" gridcol="14" gridrow="10" pageId="13" pageNumber="58">19.2</td>
<td id="76C55DFBA80A0072C657F9A47360F9F5" box="[1836,1913,1570,1650]" gridcol="15" gridrow="10" pageId="13" pageNumber="58">19.0</td>
<td id="76C55DFBA80A0072C6E7F9A473F0F9F5" box="[1948,2025,1570,1650]" gridcol="16" gridrow="10" pageId="13" pageNumber="58">19.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9B36C5A80AFF80C79CF91C73F6F937" blockId="13.[1767,2031,1690,1712]" box="[1767,2031,1690,1712]" pageId="13" pageNumber="58">
<emphasis id="B950EAD7A80AFF80C79CF91C73F6F937" box="[1767,2031,1690,1712]" italics="true" pageId="13" pageNumber="58">
<tableNote id="76C2374BA80AFF80C79CF91C73F6F937" box="[1767,2031,1690,1712]" pageId="13" pageNumber="58">...Continued on the next page</tableNote>
</emphasis>
</paragraph>
<caption id="DF5B664DA809FF83C1ECFE707581FD97" box="[151,408,502,528]" isContinuationCaption="true" pageId="14" pageNumber="59" startId="14.[151,239,502,528]" targetBox="[152,2010,548,1634]" targetIsTable="true" targetPageId="14">
<paragraph id="8B9B36C5A809FF83C1ECFE707581FD97" blockId="14.[151,408,502,528]" box="[151,408,502,528]" pageId="14" pageNumber="59">
<emphasis id="B950EAD7A809FF83C1ECFE707513FD97" bold="true" box="[151,266,502,528]" pageId="14" pageNumber="59">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B9B36C5A809FF83C1DEFDD973DBF9C2" pageId="14" pageNumber="59">
<table id="F924C465A8090072C1E3FDA273C3F9E5" box="[152,2010,548,1634]" gridcols="17" gridrows="13" pageId="14" pageNumber="59">
<tr id="35143487A8090072C1E3FDA273C3FD42" box="[152,2010,548,709]" gridrow="0" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FDA27512FD42" box="[152,267,548,709]" gridcol="0" gridrow="0" pageId="14" pageNumber="59">Specimen Depth of</th>
<th id="76C55DFBA8090072C051FDA27561FD42" box="[298,376,548,709]" gridcol="1" gridrow="0" pageId="14" pageNumber="59">ZSM 47595 holotype</th>
<th id="76C55DFBA8090072C0E3FDA275FCFD42" box="[408,485,548,709]" gridcol="2" gridrow="0" pageId="14" pageNumber="59">ZSM 47596 paratype</th>
<th id="76C55DFBA8090072C377FDA27640FD42" box="[524,601,548,709]" gridcol="3" gridrow="0" pageId="14" pageNumber="59">ZSM 47597 paratype</th>
<th id="76C55DFBA8090072C304FDA276D5FD42" box="[639,716,548,709]" gridcol="4" gridrow="0" pageId="14" pageNumber="59">ZSM 47599 paratype</th>
<th id="76C55DFBA8090072C390FDA27721FD42" box="[747,824,548,709]" gridcol="5" gridrow="0" pageId="14" pageNumber="59">ZSM 47598 paratype</th>
<th id="76C55DFBA8090072C22EFDA277BBFD42" box="[853,930,548,709]" gridcol="6" gridrow="0" pageId="14" pageNumber="59">ZM - CBSU 5003- 128 paratype</th>
<th id="76C55DFBA8090072C2BAFDA2700BFD42" box="[961,1042,548,709]" gridcol="7" gridrow="0" pageId="14" pageNumber="59">ZM - CBSU 5001-1 paratype</th>
<th id="76C55DFBA8090072C54BFDA27064FD42" box="[1072,1149,548,709]" gridcol="8" gridrow="0" pageId="14" pageNumber="59">ZM - CBSU 5003-60 paratype</th>
<th id="76C55DFBA8090072C5E6FDA270F3FD42" box="[1181,1258,548,709]" gridcol="9" gridrow="0" pageId="14" pageNumber="59">ZM - CBSU 5022-23 paratype</th>
<th id="76C55DFBA8090072C471FDA2714EFD42" box="[1290,1367,548,709]" gridcol="10" gridrow="0" pageId="14" pageNumber="59">ZM - CBSU 5024-1 paratype</th>
<th id="76C55DFBA8090072C40FFDA271D8FD42" box="[1396,1473,548,709]" gridcol="11" gridrow="0" pageId="14" pageNumber="59">ZM - CBSU 5003-77 paratype</th>
<th id="76C55DFBA8090072C4A4FDA27235FD42" box="[1503,1580,548,709]" gridcol="12" gridrow="0" pageId="14" pageNumber="59">PMR VP4679 paratype</th>
<th id="76C55DFBA8090072C732FDA2728FFD42" box="[1609,1686,548,709]" gridcol="13" gridrow="0" pageId="14" pageNumber="59">PMR VP4680 paratype</th>
<th id="76C55DFBA8090072C7CFFDA27318FD42" box="[1716,1793,548,709]" gridcol="14" gridrow="0" pageId="14" pageNumber="59">PMR VP4681 paratype</th>
<th id="76C55DFBA8090072C665FDA27372FD42" box="[1822,1899,548,709]" gridcol="15" gridrow="0" pageId="14" pageNumber="59">PMR VP4682 paratype</th>
<th id="76C55DFBA8090072C6F6FDA273C3FD42" box="[1933,2010,548,709]" gridcol="16" gridrow="0" pageId="14" pageNumber="59">PMR VP4683 paratype</th>
</tr>
<tr id="35143487A8090072C1E3FD4A73C3FC96" box="[152,2010,716,785]" gridrow="1" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FD4A7512FC96" box="[152,267,716,785]" gridcol="0" gridrow="1" pageId="14" pageNumber="59">first dorsal fin^ Length</th>
<td id="76C55DFBA8090072C051FD4A7561FC96" box="[298,376,716,785]" gridcol="1" gridrow="1" pageId="14" pageNumber="59">8.5^</td>
<td id="76C55DFBA8090072C0E3FD4A75FCFC96" box="[408,485,716,785]" gridcol="2" gridrow="1" pageId="14" pageNumber="59">7.2</td>
<td id="76C55DFBA8090072C377FD4A7640FC96" box="[524,601,716,785]" gridcol="3" gridrow="1" pageId="14" pageNumber="59">6.9</td>
<td id="76C55DFBA8090072C304FD4A76D5FC96" box="[639,716,716,785]" gridcol="4" gridrow="1" pageId="14" pageNumber="59">6.3</td>
<td id="76C55DFBA8090072C390FD4A7721FC96" box="[747,824,716,785]" gridcol="5" gridrow="1" pageId="14" pageNumber="59">5.7</td>
<td id="76C55DFBA8090072C22EFD4A77BBFC96" box="[853,930,716,785]" gridcol="6" gridrow="1" pageId="14" pageNumber="59">7.3</td>
<td id="76C55DFBA8090072C2BAFD4A700BFC96" box="[961,1042,716,785]" gridcol="7" gridrow="1" pageId="14" pageNumber="59">6.9</td>
<td id="76C55DFBA8090072C54BFD4A7064FC96" box="[1072,1149,716,785]" gridcol="8" gridrow="1" pageId="14" pageNumber="59">4.7</td>
<td id="76C55DFBA8090072C5E6FD4A70F3FC96" box="[1181,1258,716,785]" gridcol="9" gridrow="1" pageId="14" pageNumber="59">5.5</td>
<td id="76C55DFBA8090072C471FD4A714EFC96" box="[1290,1367,716,785]" gridcol="10" gridrow="1" pageId="14" pageNumber="59">5.1</td>
<td id="76C55DFBA8090072C40FFD4A71D8FC96" box="[1396,1473,716,785]" gridcol="11" gridrow="1" pageId="14" pageNumber="59">6.9</td>
<td id="76C55DFBA8090072C4A4FD4A7235FC96" box="[1503,1580,716,785]" gridcol="12" gridrow="1" pageId="14" pageNumber="59">9.5^</td>
<td id="76C55DFBA8090072C732FD4A728FFC96" box="[1609,1686,716,785]" gridcol="13" gridrow="1" pageId="14" pageNumber="59">9.0^</td>
<td id="76C55DFBA8090072C7CFFD4A7318FC96" box="[1716,1793,716,785]" gridcol="14" gridrow="1" pageId="14" pageNumber="59">8.4^</td>
<td id="76C55DFBA8090072C665FD4A7372FC96" box="[1822,1899,716,785]" gridcol="15" gridrow="1" pageId="14" pageNumber="59">7.4</td>
<td id="76C55DFBA8090072C6F6FD4A73C3FC96" box="[1933,2010,716,785]" gridcol="16" gridrow="1" pageId="14" pageNumber="59">7.1</td>
</tr>
<tr id="35143487A8090072C1E3FC9E73C3FCEE" box="[152,2010,792,873]" gridrow="2" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FC9E7512FCEE" box="[152,267,792,873]" gridcol="0" gridrow="2" pageId="14" pageNumber="59">of second dorsal fin base</th>
<td id="76C55DFBA8090072C051FC9E7561FCEE" box="[298,376,792,873]" gridcol="1" gridrow="2" pageId="14" pageNumber="59">17.4</td>
<td id="76C55DFBA8090072C0E3FC9E75FCFCEE" box="[408,485,792,873]" gridcol="2" gridrow="2" pageId="14" pageNumber="59">16.7</td>
<td id="76C55DFBA8090072C377FC9E7640FCEE" box="[524,601,792,873]" gridcol="3" gridrow="2" pageId="14" pageNumber="59">18.3</td>
<td id="76C55DFBA8090072C304FC9E76D5FCEE" box="[639,716,792,873]" gridcol="4" gridrow="2" pageId="14" pageNumber="59">18.5</td>
<td id="76C55DFBA8090072C390FC9E7721FCEE" box="[747,824,792,873]" gridcol="5" gridrow="2" pageId="14" pageNumber="59">18.8</td>
<td id="76C55DFBA8090072C22EFC9E77BBFCEE" box="[853,930,792,873]" gridcol="6" gridrow="2" pageId="14" pageNumber="59">19.9</td>
<td id="76C55DFBA8090072C2BAFC9E700BFCEE" box="[961,1042,792,873]" gridcol="7" gridrow="2" pageId="14" pageNumber="59">17.3</td>
<td id="76C55DFBA8090072C54BFC9E7064FCEE" box="[1072,1149,792,873]" gridcol="8" gridrow="2" pageId="14" pageNumber="59">18.6</td>
<td id="76C55DFBA8090072C5E6FC9E70F3FCEE" box="[1181,1258,792,873]" gridcol="9" gridrow="2" pageId="14" pageNumber="59">16.7</td>
<td id="76C55DFBA8090072C471FC9E714EFCEE" box="[1290,1367,792,873]" gridcol="10" gridrow="2" pageId="14" pageNumber="59">18.4</td>
<td id="76C55DFBA8090072C40FFC9E71D8FCEE" box="[1396,1473,792,873]" gridcol="11" gridrow="2" pageId="14" pageNumber="59">17.9</td>
<td id="76C55DFBA8090072C4A4FC9E7235FCEE" box="[1503,1580,792,873]" gridcol="12" gridrow="2" pageId="14" pageNumber="59">18.5</td>
<td id="76C55DFBA8090072C732FC9E728FFCEE" box="[1609,1686,792,873]" gridcol="13" gridrow="2" pageId="14" pageNumber="59">19.8</td>
<td id="76C55DFBA8090072C7CFFC9E7318FCEE" box="[1716,1793,792,873]" gridcol="14" gridrow="2" pageId="14" pageNumber="59">20.1</td>
<td id="76C55DFBA8090072C665FC9E7372FCEE" box="[1822,1899,792,873]" gridcol="15" gridrow="2" pageId="14" pageNumber="59">16.7</td>
<td id="76C55DFBA8090072C6F6FC9E73C3FCEE" box="[1933,2010,792,873]" gridcol="16" gridrow="2" pageId="14" pageNumber="59">17.7</td>
</tr>
<tr id="35143487A8090072C1E3FCF373C3FC42" box="[152,2010,885,965]" gridrow="3" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FCF37512FC42" box="[152,267,885,965]" gridcol="0" gridrow="3" pageId="14" pageNumber="59">Depth of second dorsal fin</th>
<td id="76C55DFBA8090072C051FCF37561FC42" box="[298,376,885,965]" gridcol="1" gridrow="3" pageId="14" pageNumber="59">12.9</td>
<td id="76C55DFBA8090072C0E3FCF375FCFC42" box="[408,485,885,965]" gridcol="2" gridrow="3" pageId="14" pageNumber="59">12.0</td>
<td id="76C55DFBA8090072C377FCF37640FC42" box="[524,601,885,965]" gridcol="3" gridrow="3" pageId="14" pageNumber="59">15.3</td>
<td id="76C55DFBA8090072C304FCF376D5FC42" box="[639,716,885,965]" gridcol="4" gridrow="3" pageId="14" pageNumber="59">13.2</td>
<td id="76C55DFBA8090072C390FCF37721FC42" box="[747,824,885,965]" gridcol="5" gridrow="3" pageId="14" pageNumber="59">11.3</td>
<td id="76C55DFBA8090072C22EFCF377BBFC42" box="[853,930,885,965]" gridcol="6" gridrow="3" pageId="14" pageNumber="59">13.2</td>
<td id="76C55DFBA8090072C2BAFCF3700BFC42" box="[961,1042,885,965]" gridcol="7" gridrow="3" pageId="14" pageNumber="59">12.1</td>
<td id="76C55DFBA8090072C54BFCF37064FC42" box="[1072,1149,885,965]" gridcol="8" gridrow="3" pageId="14" pageNumber="59">12.3</td>
<td id="76C55DFBA8090072C5E6FCF370F3FC42" box="[1181,1258,885,965]" gridcol="9" gridrow="3" pageId="14" pageNumber="59">12.2</td>
<td id="76C55DFBA8090072C471FCF3714EFC42" box="[1290,1367,885,965]" gridcol="10" gridrow="3" pageId="14" pageNumber="59">12.7</td>
<td id="76C55DFBA8090072C40FFCF371D8FC42" box="[1396,1473,885,965]" gridcol="11" gridrow="3" pageId="14" pageNumber="59">11.6</td>
<td id="76C55DFBA8090072C4A4FCF37235FC42" box="[1503,1580,885,965]" gridcol="12" gridrow="3" pageId="14" pageNumber="59">13.4</td>
<td id="76C55DFBA8090072C732FCF3728FFC42" box="[1609,1686,885,965]" gridcol="13" gridrow="3" pageId="14" pageNumber="59">12.4</td>
<td id="76C55DFBA8090072C7CFFCF37318FC42" box="[1716,1793,885,965]" gridcol="14" gridrow="3" pageId="14" pageNumber="59">14.0</td>
<td id="76C55DFBA8090072C665FCF37372FC42" box="[1822,1899,885,965]" gridcol="15" gridrow="3" pageId="14" pageNumber="59">12.1</td>
<td id="76C55DFBA8090072C6F6FCF373C3FC42" box="[1933,2010,885,965]" gridcol="16" gridrow="3" pageId="14" pageNumber="59">12.7</td>
</tr>
<tr id="35143487A8090072C1E3FC5B73C3FB97" box="[152,2010,989,1040]" gridrow="4" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FC5B7512FB97" box="[152,267,989,1040]" gridcol="0" gridrow="4" pageId="14" pageNumber="59">Length of anal-fin base</th>
<td id="76C55DFBA8090072C051FC5B7561FB97" box="[298,376,989,1040]" gridcol="1" gridrow="4" pageId="14" pageNumber="59">19.8</td>
<td id="76C55DFBA8090072C0E3FC5B75FCFB97" box="[408,485,989,1040]" gridcol="2" gridrow="4" pageId="14" pageNumber="59">23.1</td>
<td id="76C55DFBA8090072C377FC5B7640FB97" box="[524,601,989,1040]" gridcol="3" gridrow="4" pageId="14" pageNumber="59">20.4</td>
<td id="76C55DFBA8090072C304FC5B76D5FB97" box="[639,716,989,1040]" gridcol="4" gridrow="4" pageId="14" pageNumber="59">19.5</td>
<td id="76C55DFBA8090072C390FC5B7721FB97" box="[747,824,989,1040]" gridcol="5" gridrow="4" pageId="14" pageNumber="59">19.5</td>
<td id="76C55DFBA8090072C22EFC5B77BBFB97" box="[853,930,989,1040]" gridcol="6" gridrow="4" pageId="14" pageNumber="59">20.7</td>
<td id="76C55DFBA8090072C2BAFC5B700BFB97" box="[961,1042,989,1040]" gridcol="7" gridrow="4" pageId="14" pageNumber="59">20.2</td>
<td id="76C55DFBA8090072C54BFC5B7064FB97" box="[1072,1149,989,1040]" gridcol="8" gridrow="4" pageId="14" pageNumber="59">22.1</td>
<td id="76C55DFBA8090072C5E6FC5B70F3FB97" box="[1181,1258,989,1040]" gridcol="9" gridrow="4" pageId="14" pageNumber="59">21.6</td>
<td id="76C55DFBA8090072C471FC5B714EFB97" box="[1290,1367,989,1040]" gridcol="10" gridrow="4" pageId="14" pageNumber="59">19.2</td>
<td id="76C55DFBA8090072C40FFC5B71D8FB97" box="[1396,1473,989,1040]" gridcol="11" gridrow="4" pageId="14" pageNumber="59">20.8</td>
<td id="76C55DFBA8090072C4A4FC5B7235FB97" box="[1503,1580,989,1040]" gridcol="12" gridrow="4" pageId="14" pageNumber="59">21.5</td>
<td id="76C55DFBA8090072C732FC5B728FFB97" box="[1609,1686,989,1040]" gridcol="13" gridrow="4" pageId="14" pageNumber="59">21.9</td>
<td id="76C55DFBA8090072C7CFFC5B7318FB97" box="[1716,1793,989,1040]" gridcol="14" gridrow="4" pageId="14" pageNumber="59">22.3</td>
<td id="76C55DFBA8090072C665FC5B7372FB97" box="[1822,1899,989,1040]" gridcol="15" gridrow="4" pageId="14" pageNumber="59">23.8</td>
<td id="76C55DFBA8090072C6F6FC5B73C3FB97" box="[1933,2010,989,1040]" gridcol="16" gridrow="4" pageId="14" pageNumber="59">19.7</td>
</tr>
<tr id="35143487A8090072C1E3FBA373C3FBDF" box="[152,2010,1061,1112]" gridrow="5" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FBA37512FBDF" box="[152,267,1061,1112]" gridcol="0" gridrow="5" pageId="14" pageNumber="59">Depth of anal fin</th>
<td id="76C55DFBA8090072C051FBA37561FBDF" box="[298,376,1061,1112]" gridcol="1" gridrow="5" pageId="14" pageNumber="59">10.3</td>
<td id="76C55DFBA8090072C0E3FBA375FCFBDF" box="[408,485,1061,1112]" gridcol="2" gridrow="5" pageId="14" pageNumber="59">10.3</td>
<td id="76C55DFBA8090072C377FBA37640FBDF" box="[524,601,1061,1112]" gridcol="3" gridrow="5" pageId="14" pageNumber="59">11.1</td>
<td id="76C55DFBA8090072C304FBA376D5FBDF" box="[639,716,1061,1112]" gridcol="4" gridrow="5" pageId="14" pageNumber="59">10.7</td>
<td id="76C55DFBA8090072C390FBA37721FBDF" box="[747,824,1061,1112]" gridcol="5" gridrow="5" pageId="14" pageNumber="59">9.9</td>
<td id="76C55DFBA8090072C22EFBA377BBFBDF" box="[853,930,1061,1112]" gridcol="6" gridrow="5" pageId="14" pageNumber="59">10.2</td>
<td id="76C55DFBA8090072C2BAFBA3700BFBDF" box="[961,1042,1061,1112]" gridcol="7" gridrow="5" pageId="14" pageNumber="59">11.4</td>
<td id="76C55DFBA8090072C54BFBA37064FBDF" box="[1072,1149,1061,1112]" gridcol="8" gridrow="5" pageId="14" pageNumber="59">10.9</td>
<td id="76C55DFBA8090072C5E6FBA370F3FBDF" box="[1181,1258,1061,1112]" gridcol="9" gridrow="5" pageId="14" pageNumber="59">9.6</td>
<td id="76C55DFBA8090072C471FBA3714EFBDF" box="[1290,1367,1061,1112]" gridcol="10" gridrow="5" pageId="14" pageNumber="59">11.0</td>
<td id="76C55DFBA8090072C40FFBA371D8FBDF" box="[1396,1473,1061,1112]" gridcol="11" gridrow="5" pageId="14" pageNumber="59">10.2</td>
<td id="76C55DFBA8090072C4A4FBA37235FBDF" box="[1503,1580,1061,1112]" gridcol="12" gridrow="5" pageId="14" pageNumber="59">10.5</td>
<td id="76C55DFBA8090072C732FBA3728FFBDF" box="[1609,1686,1061,1112]" gridcol="13" gridrow="5" pageId="14" pageNumber="59">10.7</td>
<td id="76C55DFBA8090072C7CFFBA37318FBDF" box="[1716,1793,1061,1112]" gridcol="14" gridrow="5" pageId="14" pageNumber="59">10.1</td>
<td id="76C55DFBA8090072C665FBA37372FBDF" box="[1822,1899,1061,1112]" gridcol="15" gridrow="5" pageId="14" pageNumber="59">11.5</td>
<td id="76C55DFBA8090072C6F6FBA373C3FBDF" box="[1933,2010,1061,1112]" gridcol="16" gridrow="5" pageId="14" pageNumber="59">11.9</td>
</tr>
<tr id="35143487A8090072C1E3FBEB73C3FB27" box="[152,2010,1133,1184]" gridrow="6" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FBEB7512FB27" box="[152,267,1133,1184]" gridcol="0" gridrow="6" pageId="14" pageNumber="59">Preanal distance</th>
<td id="76C55DFBA8090072C051FBEB7561FB27" box="[298,376,1133,1184]" gridcol="1" gridrow="6" pageId="14" pageNumber="59">62.8</td>
<td id="76C55DFBA8090072C0E3FBEB75FCFB27" box="[408,485,1133,1184]" gridcol="2" gridrow="6" pageId="14" pageNumber="59">57.4</td>
<td id="76C55DFBA8090072C377FBEB7640FB27" box="[524,601,1133,1184]" gridcol="3" gridrow="6" pageId="14" pageNumber="59">60.2</td>
<td id="76C55DFBA8090072C304FBEB76D5FB27" box="[639,716,1133,1184]" gridcol="4" gridrow="6" pageId="14" pageNumber="59">59.9</td>
<td id="76C55DFBA8090072C390FBEB7721FB27" box="[747,824,1133,1184]" gridcol="5" gridrow="6" pageId="14" pageNumber="59">57.2</td>
<td id="76C55DFBA8090072C22EFBEB77BBFB27" box="[853,930,1133,1184]" gridcol="6" gridrow="6" pageId="14" pageNumber="59">59.7</td>
<td id="76C55DFBA8090072C2BAFBEB700BFB27" box="[961,1042,1133,1184]" gridcol="7" gridrow="6" pageId="14" pageNumber="59">59.6</td>
<td id="76C55DFBA8090072C54BFBEB7064FB27" box="[1072,1149,1133,1184]" gridcol="8" gridrow="6" pageId="14" pageNumber="59">57.1</td>
<td id="76C55DFBA8090072C5E6FBEB70F3FB27" box="[1181,1258,1133,1184]" gridcol="9" gridrow="6" pageId="14" pageNumber="59">58.9</td>
<td id="76C55DFBA8090072C471FBEB714EFB27" box="[1290,1367,1133,1184]" gridcol="10" gridrow="6" pageId="14" pageNumber="59">60.2</td>
<td id="76C55DFBA8090072C40FFBEB71D8FB27" box="[1396,1473,1133,1184]" gridcol="11" gridrow="6" pageId="14" pageNumber="59">59.2</td>
<td id="76C55DFBA8090072C4A4FBEB7235FB27" box="[1503,1580,1133,1184]" gridcol="12" gridrow="6" pageId="14" pageNumber="59">57.6</td>
<td id="76C55DFBA8090072C732FBEB728FFB27" box="[1609,1686,1133,1184]" gridcol="13" gridrow="6" pageId="14" pageNumber="59">60.4</td>
<td id="76C55DFBA8090072C7CFFBEB7318FB27" box="[1716,1793,1133,1184]" gridcol="14" gridrow="6" pageId="14" pageNumber="59">60.7</td>
<td id="76C55DFBA8090072C665FBEB7372FB27" box="[1822,1899,1133,1184]" gridcol="15" gridrow="6" pageId="14" pageNumber="59">58.3</td>
<td id="76C55DFBA8090072C6F6FBEB73C3FB27" box="[1933,2010,1133,1184]" gridcol="16" gridrow="6" pageId="14" pageNumber="59">58.6</td>
</tr>
<tr id="35143487A8090072C1E3FB3373C3FB6F" box="[152,2010,1205,1256]" gridrow="7" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FB337512FB6F" box="[152,267,1205,1256]" gridcol="0" gridrow="7" pageId="14" pageNumber="59">Prepectoral distance</th>
<td id="76C55DFBA8090072C051FB337561FB6F" box="[298,376,1205,1256]" gridcol="1" gridrow="7" pageId="14" pageNumber="59">36.5</td>
<td id="76C55DFBA8090072C0E3FB3375FCFB6F" box="[408,485,1205,1256]" gridcol="2" gridrow="7" pageId="14" pageNumber="59">37.1</td>
<td id="76C55DFBA8090072C377FB337640FB6F" box="[524,601,1205,1256]" gridcol="3" gridrow="7" pageId="14" pageNumber="59">39.9</td>
<td id="76C55DFBA8090072C304FB3376D5FB6F" box="[639,716,1205,1256]" gridcol="4" gridrow="7" pageId="14" pageNumber="59">35.4</td>
<td id="76C55DFBA8090072C390FB337721FB6F" box="[747,824,1205,1256]" gridcol="5" gridrow="7" pageId="14" pageNumber="59">32.9</td>
<td id="76C55DFBA8090072C22EFB3377BBFB6F" box="[853,930,1205,1256]" gridcol="6" gridrow="7" pageId="14" pageNumber="59">38.4</td>
<td id="76C55DFBA8090072C2BAFB33700BFB6F" box="[961,1042,1205,1256]" gridcol="7" gridrow="7" pageId="14" pageNumber="59">36.8</td>
<td id="76C55DFBA8090072C54BFB337064FB6F" box="[1072,1149,1205,1256]" gridcol="8" gridrow="7" pageId="14" pageNumber="59">37.9</td>
<td id="76C55DFBA8090072C5E6FB3370F3FB6F" box="[1181,1258,1205,1256]" gridcol="9" gridrow="7" pageId="14" pageNumber="59">33.2</td>
<td id="76C55DFBA8090072C471FB33714EFB6F" box="[1290,1367,1205,1256]" gridcol="10" gridrow="7" pageId="14" pageNumber="59">35.6</td>
<td id="76C55DFBA8090072C40FFB3371D8FB6F" box="[1396,1473,1205,1256]" gridcol="11" gridrow="7" pageId="14" pageNumber="59">34.6</td>
<td id="76C55DFBA8090072C4A4FB337235FB6F" box="[1503,1580,1205,1256]" gridcol="12" gridrow="7" pageId="14" pageNumber="59">35.5</td>
<td id="76C55DFBA8090072C732FB33728FFB6F" box="[1609,1686,1205,1256]" gridcol="13" gridrow="7" pageId="14" pageNumber="59">37.4</td>
<td id="76C55DFBA8090072C7CFFB337318FB6F" box="[1716,1793,1205,1256]" gridcol="14" gridrow="7" pageId="14" pageNumber="59">37.6</td>
<td id="76C55DFBA8090072C665FB337372FB6F" box="[1822,1899,1205,1256]" gridcol="15" gridrow="7" pageId="14" pageNumber="59">37.7</td>
<td id="76C55DFBA8090072C6F6FB3373C3FB6F" box="[1933,2010,1205,1256]" gridcol="16" gridrow="7" pageId="14" pageNumber="59">38.7</td>
</tr>
<tr id="35143487A8090072C1E3FB7B73C3FAB7" box="[152,2010,1277,1328]" gridrow="8" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FB7B7512FAB7" box="[152,267,1277,1328]" gridcol="0" gridrow="8" pageId="14" pageNumber="59">Prepelvic distance*</th>
<td id="76C55DFBA8090072C051FB7B7561FAB7" box="[298,376,1277,1328]" gridcol="1" gridrow="8" pageId="14" pageNumber="59">32.2*</td>
<td id="76C55DFBA8090072C0E3FB7B75FCFAB7" box="[408,485,1277,1328]" gridcol="2" gridrow="8" pageId="14" pageNumber="59">35.1*</td>
<td id="76C55DFBA8090072C377FB7B7640FAB7" box="[524,601,1277,1328]" gridcol="3" gridrow="8" pageId="14" pageNumber="59">32.1*</td>
<td id="76C55DFBA8090072C304FB7B76D5FAB7" box="[639,716,1277,1328]" gridcol="4" gridrow="8" pageId="14" pageNumber="59">31.0</td>
<td id="76C55DFBA8090072C390FB7B7721FAB7" box="[747,824,1277,1328]" gridcol="5" gridrow="8" pageId="14" pageNumber="59">29.1</td>
<td id="76C55DFBA8090072C22EFB7B77BBFAB7" box="[853,930,1277,1328]" gridcol="6" gridrow="8" pageId="14" pageNumber="59">31.6</td>
<td id="76C55DFBA8090072C2BAFB7B700BFAB7" box="[961,1042,1277,1328]" gridcol="7" gridrow="8" pageId="14" pageNumber="59">30.8</td>
<td id="76C55DFBA8090072C54BFB7B7064FAB7" box="[1072,1149,1277,1328]" gridcol="8" gridrow="8" pageId="14" pageNumber="59">30.6</td>
<td id="76C55DFBA8090072C5E6FB7B70F3FAB7" box="[1181,1258,1277,1328]" gridcol="9" gridrow="8" pageId="14" pageNumber="59">31.6</td>
<td id="76C55DFBA8090072C471FB7B714EFAB7" box="[1290,1367,1277,1328]" gridcol="10" gridrow="8" pageId="14" pageNumber="59">30.7</td>
<td id="76C55DFBA8090072C40FFB7B71D8FAB7" box="[1396,1473,1277,1328]" gridcol="11" gridrow="8" pageId="14" pageNumber="59">29.7</td>
<td id="76C55DFBA8090072C4A4FB7B7235FAB7" box="[1503,1580,1277,1328]" gridcol="12" gridrow="8" pageId="14" pageNumber="59">32.5*</td>
<td id="76C55DFBA8090072C732FB7B728FFAB7" box="[1609,1686,1277,1328]" gridcol="13" gridrow="8" pageId="14" pageNumber="59">35.9*</td>
<td id="76C55DFBA8090072C7CFFB7B7318FAB7" box="[1716,1793,1277,1328]" gridcol="14" gridrow="8" pageId="14" pageNumber="59">35.2*</td>
<td id="76C55DFBA8090072C665FB7B7372FAB7" box="[1822,1899,1277,1328]" gridcol="15" gridrow="8" pageId="14" pageNumber="59">28.8</td>
<td id="76C55DFBA8090072C6F6FB7B73C3FAB7" box="[1933,2010,1277,1328]" gridcol="16" gridrow="8" pageId="14" pageNumber="59">27.2</td>
</tr>
<tr id="35143487A8090072C1E3FAC373C3FAFF" box="[152,2010,1349,1400]" gridrow="9" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FAC37512FAFF" box="[152,267,1349,1400]" gridcol="0" gridrow="9" pageId="14" pageNumber="59">Pelvic disc length</th>
<td id="76C55DFBA8090072C051FAC37561FAFF" box="[298,376,1349,1400]" gridcol="1" gridrow="9" pageId="14" pageNumber="59">31.1</td>
<td id="76C55DFBA8090072C0E3FAC375FCFAFF" box="[408,485,1349,1400]" gridcol="2" gridrow="9" pageId="14" pageNumber="59">28.6</td>
<td id="76C55DFBA8090072C377FAC37640FAFF" box="[524,601,1349,1400]" gridcol="3" gridrow="9" pageId="14" pageNumber="59">31.4</td>
<td id="76C55DFBA8090072C304FAC376D5FAFF" box="[639,716,1349,1400]" gridcol="4" gridrow="9" pageId="14" pageNumber="59">29.4</td>
<td id="76C55DFBA8090072C390FAC37721FAFF" box="[747,824,1349,1400]" gridcol="5" gridrow="9" pageId="14" pageNumber="59">30.2</td>
<td id="76C55DFBA8090072C22EFAC377BBFAFF" box="[853,930,1349,1400]" gridcol="6" gridrow="9" pageId="14" pageNumber="59">32.1</td>
<td id="76C55DFBA8090072C2BAFAC3700BFAFF" box="[961,1042,1349,1400]" gridcol="7" gridrow="9" pageId="14" pageNumber="59">damaged</td>
<td id="76C55DFBA8090072C54BFAC37064FAFF" box="[1072,1149,1349,1400]" gridcol="8" gridrow="9" pageId="14" pageNumber="59">29.3</td>
<td id="76C55DFBA8090072C5E6FAC370F3FAFF" box="[1181,1258,1349,1400]" gridcol="9" gridrow="9" pageId="14" pageNumber="59">29.4</td>
<td id="76C55DFBA8090072C471FAC3714EFAFF" box="[1290,1367,1349,1400]" gridcol="10" gridrow="9" pageId="14" pageNumber="59">29.4</td>
<td id="76C55DFBA8090072C40FFAC371D8FAFF" box="[1396,1473,1349,1400]" gridcol="11" gridrow="9" pageId="14" pageNumber="59">26.2</td>
<td id="76C55DFBA8090072C4A4FAC37235FAFF" box="[1503,1580,1349,1400]" gridcol="12" gridrow="9" pageId="14" pageNumber="59">29.8</td>
<td id="76C55DFBA8090072C732FAC3728FFAFF" box="[1609,1686,1349,1400]" gridcol="13" gridrow="9" pageId="14" pageNumber="59">28.2</td>
<td id="76C55DFBA8090072C7CFFAC37318FAFF" box="[1716,1793,1349,1400]" gridcol="14" gridrow="9" pageId="14" pageNumber="59">28.7</td>
<td id="76C55DFBA8090072C665FAC37372FAFF" box="[1822,1899,1349,1400]" gridcol="15" gridrow="9" pageId="14" pageNumber="59">29.9</td>
<td id="76C55DFBA8090072C6F6FAC373C3FAFF" box="[1933,2010,1349,1400]" gridcol="16" gridrow="9" pageId="14" pageNumber="59">27.0</td>
</tr>
<tr id="35143487A8090072C1E3FA0B73C3FA5A" box="[152,2010,1421,1501]" gridrow="10" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FA0B7512FA5A" box="[152,267,1421,1501]" gridcol="0" gridrow="10" pageId="14" pageNumber="59">Lateral head length Body depth</th>
<td id="76C55DFBA8090072C051FA0B7561FA5A" box="[298,376,1421,1501]" gridcol="1" gridrow="10" pageId="14" pageNumber="59">35.3</td>
<td id="76C55DFBA8090072C0E3FA0B75FCFA5A" box="[408,485,1421,1501]" gridcol="2" gridrow="10" pageId="14" pageNumber="59">35.1</td>
<td id="76C55DFBA8090072C377FA0B7640FA5A" box="[524,601,1421,1501]" gridcol="3" gridrow="10" pageId="14" pageNumber="59">37.8</td>
<td id="76C55DFBA8090072C304FA0B76D5FA5A" box="[639,716,1421,1501]" gridcol="4" gridrow="10" pageId="14" pageNumber="59">35.4</td>
<td id="76C55DFBA8090072C390FA0B7721FA5A" box="[747,824,1421,1501]" gridcol="5" gridrow="10" pageId="14" pageNumber="59">33.4</td>
<td id="76C55DFBA8090072C22EFA0B77BBFA5A" box="[853,930,1421,1501]" gridcol="6" gridrow="10" pageId="14" pageNumber="59">36.1</td>
<td id="76C55DFBA8090072C2BAFA0B700BFA5A" box="[961,1042,1421,1501]" gridcol="7" gridrow="10" pageId="14" pageNumber="59">37.6</td>
<td id="76C55DFBA8090072C54BFA0B7064FA5A" box="[1072,1149,1421,1501]" gridcol="8" gridrow="10" pageId="14" pageNumber="59">36.8</td>
<td id="76C55DFBA8090072C5E6FA0B70F3FA5A" box="[1181,1258,1421,1501]" gridcol="9" gridrow="10" pageId="14" pageNumber="59">33.4</td>
<td id="76C55DFBA8090072C471FA0B714EFA5A" box="[1290,1367,1421,1501]" gridcol="10" gridrow="10" pageId="14" pageNumber="59">33.2</td>
<td id="76C55DFBA8090072C40FFA0B71D8FA5A" box="[1396,1473,1421,1501]" gridcol="11" gridrow="10" pageId="14" pageNumber="59">33.9</td>
<td id="76C55DFBA8090072C4A4FA0B7235FA5A" box="[1503,1580,1421,1501]" gridcol="12" gridrow="10" pageId="14" pageNumber="59">37.0</td>
<td id="76C55DFBA8090072C732FA0B728FFA5A" box="[1609,1686,1421,1501]" gridcol="13" gridrow="10" pageId="14" pageNumber="59">36.0</td>
<td id="76C55DFBA8090072C7CFFA0B7318FA5A" box="[1716,1793,1421,1501]" gridcol="14" gridrow="10" pageId="14" pageNumber="59">38.1</td>
<td id="76C55DFBA8090072C665FA0B7372FA5A" box="[1822,1899,1421,1501]" gridcol="15" gridrow="10" pageId="14" pageNumber="59">35.4</td>
<td id="76C55DFBA8090072C6F6FA0B73C3FA5A" box="[1933,2010,1421,1501]" gridcol="16" gridrow="10" pageId="14" pageNumber="59">36.9</td>
</tr>
<tr id="35143487A8090072C1E3FA6373C3F9AF" box="[152,2010,1509,1576]" gridrow="11" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3FA637512F9AF" box="[152,267,1509,1576]" gridcol="0" gridrow="11" pageId="14" pageNumber="59">at anal-fin origin* Body width</th>
<td id="76C55DFBA8090072C051FA637561F9AF" box="[298,376,1509,1576]" gridcol="1" gridrow="11" pageId="14" pageNumber="59">13.8*</td>
<td id="76C55DFBA8090072C0E3FA6375FCF9AF" box="[408,485,1509,1576]" gridcol="2" gridrow="11" pageId="14" pageNumber="59">13.8*</td>
<td id="76C55DFBA8090072C377FA637640F9AF" box="[524,601,1509,1576]" gridcol="3" gridrow="11" pageId="14" pageNumber="59">13.2*</td>
<td id="76C55DFBA8090072C304FA6376D5F9AF" box="[639,716,1509,1576]" gridcol="4" gridrow="11" pageId="14" pageNumber="59">11.7</td>
<td id="76C55DFBA8090072C390FA637721F9AF" box="[747,824,1509,1576]" gridcol="5" gridrow="11" pageId="14" pageNumber="59">11.1</td>
<td id="76C55DFBA8090072C22EFA6377BBF9AF" box="[853,930,1509,1576]" gridcol="6" gridrow="11" pageId="14" pageNumber="59">12.2</td>
<td id="76C55DFBA8090072C2BAFA63700BF9AF" box="[961,1042,1509,1576]" gridcol="7" gridrow="11" pageId="14" pageNumber="59">12.0</td>
<td id="76C55DFBA8090072C54BFA637064F9AF" box="[1072,1149,1509,1576]" gridcol="8" gridrow="11" pageId="14" pageNumber="59">12.7</td>
<td id="76C55DFBA8090072C5E6FA6370F3F9AF" box="[1181,1258,1509,1576]" gridcol="9" gridrow="11" pageId="14" pageNumber="59">12.4</td>
<td id="76C55DFBA8090072C471FA63714EF9AF" box="[1290,1367,1509,1576]" gridcol="10" gridrow="11" pageId="14" pageNumber="59">12.7</td>
<td id="76C55DFBA8090072C40FFA6371D8F9AF" box="[1396,1473,1509,1576]" gridcol="11" gridrow="11" pageId="14" pageNumber="59">11.8</td>
<td id="76C55DFBA8090072C4A4FA637235F9AF" box="[1503,1580,1509,1576]" gridcol="12" gridrow="11" pageId="14" pageNumber="59">13.0*</td>
<td id="76C55DFBA8090072C732FA63728FF9AF" box="[1609,1686,1509,1576]" gridcol="13" gridrow="11" pageId="14" pageNumber="59">13.4*</td>
<td id="76C55DFBA8090072C7CFFA637318F9AF" box="[1716,1793,1509,1576]" gridcol="14" gridrow="11" pageId="14" pageNumber="59">12.9*</td>
<td id="76C55DFBA8090072C665FA637372F9AF" box="[1822,1899,1509,1576]" gridcol="15" gridrow="11" pageId="14" pageNumber="59">12.7</td>
<td id="76C55DFBA8090072C6F6FA6373C3F9AF" box="[1933,2010,1509,1576]" gridcol="16" gridrow="11" pageId="14" pageNumber="59">11.5</td>
</tr>
<tr id="35143487A8090072C1E3F9A973C3F9E5" box="[152,2010,1583,1634]" gridrow="12" pageId="14" pageNumber="59">
<th id="76C55DFBA8090072C1E3F9A97512F9E5" box="[152,267,1583,1634]" gridcol="0" gridrow="12" pageId="14" pageNumber="59">at anal-fin origin</th>
<td id="76C55DFBA8090072C051F9A97561F9E5" box="[298,376,1583,1634]" gridcol="1" gridrow="12" pageId="14" pageNumber="59">11.4</td>
<td id="76C55DFBA8090072C0E3F9A975FCF9E5" box="[408,485,1583,1634]" gridcol="2" gridrow="12" pageId="14" pageNumber="59">12.0</td>
<td id="76C55DFBA8090072C377F9A97640F9E5" box="[524,601,1583,1634]" gridcol="3" gridrow="12" pageId="14" pageNumber="59">12.0</td>
<td id="76C55DFBA8090072C304F9A976D5F9E5" box="[639,716,1583,1634]" gridcol="4" gridrow="12" pageId="14" pageNumber="59">10.2</td>
<td id="76C55DFBA8090072C390F9A97721F9E5" box="[747,824,1583,1634]" gridcol="5" gridrow="12" pageId="14" pageNumber="59">10.2</td>
<td id="76C55DFBA8090072C22EF9A977BBF9E5" box="[853,930,1583,1634]" gridcol="6" gridrow="12" pageId="14" pageNumber="59">9.6</td>
<td id="76C55DFBA8090072C2BAF9A9700BF9E5" box="[961,1042,1583,1634]" gridcol="7" gridrow="12" pageId="14" pageNumber="59">10.7</td>
<td id="76C55DFBA8090072C54BF9A97064F9E5" box="[1072,1149,1583,1634]" gridcol="8" gridrow="12" pageId="14" pageNumber="59">10.6</td>
<td id="76C55DFBA8090072C5E6F9A970F3F9E5" box="[1181,1258,1583,1634]" gridcol="9" gridrow="12" pageId="14" pageNumber="59">9.7</td>
<td id="76C55DFBA8090072C471F9A9714EF9E5" box="[1290,1367,1583,1634]" gridcol="10" gridrow="12" pageId="14" pageNumber="59">10.3</td>
<td id="76C55DFBA8090072C40FF9A971D8F9E5" box="[1396,1473,1583,1634]" gridcol="11" gridrow="12" pageId="14" pageNumber="59">9.2</td>
<td id="76C55DFBA8090072C4A4F9A97235F9E5" box="[1503,1580,1583,1634]" gridcol="12" gridrow="12" pageId="14" pageNumber="59">10.1</td>
<td id="76C55DFBA8090072C732F9A9728FF9E5" box="[1609,1686,1583,1634]" gridcol="13" gridrow="12" pageId="14" pageNumber="59">10.5</td>
<td id="76C55DFBA8090072C7CFF9A97318F9E5" box="[1716,1793,1583,1634]" gridcol="14" gridrow="12" pageId="14" pageNumber="59">10.6</td>
<td id="76C55DFBA8090072C665F9A97372F9E5" box="[1822,1899,1583,1634]" gridcol="15" gridrow="12" pageId="14" pageNumber="59">10.9</td>
<td id="76C55DFBA8090072C6F6F9A973C3F9E5" box="[1933,2010,1583,1634]" gridcol="16" gridrow="12" pageId="14" pageNumber="59">9.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9B36C5A809FF83C7A2F9FF73F9F908" blockId="14.[1753,2016,1661,1679]" box="[1753,2016,1657,1679]" pageId="14" pageNumber="59">
<emphasis id="B950EAD7A809FF83C7A2F9FF73F9F908" box="[1753,2016,1657,1679]" italics="true" pageId="14" pageNumber="59">
<tableNote id="76C2374BA809FF83C7A2F9FF73F9F908" box="[1753,2016,1657,1679]" pageId="14" pageNumber="59" targetBox="[152,2010,548,1634]" targetPageId="14">...Continued on the next page</tableNote>
</emphasis>
</paragraph>
<caption id="DF5B664DA808FF82C1ECFE757581FD8A" box="[151,408,499,525]" isContinuationCaption="true" pageId="15" pageNumber="60" startId="15.[151,239,499,525]" targetBox="[151,2013,545,1685]" targetIsTable="true" targetPageId="15">
<paragraph id="8B9B36C5A808FF82C1ECFE757581FD8A" blockId="15.[151,408,499,525]" box="[151,408,499,525]" pageId="15" pageNumber="60">
<emphasis id="B950EAD7A808FF82C1ECFE757513FD8A" bold="true" box="[151,266,499,525]" pageId="15" pageNumber="60">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8B9B36C5A808FF82C1DEFDDA73DEF900" pageId="15" pageNumber="60">
<table id="F924C465A8080072C1ECFDA773C4F912" box="[151,2013,545,1685]" gridcols="17" gridrows="17" pageId="15" pageNumber="60">
<tr id="35143487A8080072C1ECFDA773C4FD2A" box="[151,2013,545,685]" gridrow="0" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFDA77515FD2A" box="[151,268,545,685]" gridcol="0" gridrow="0" pageId="15" pageNumber="60">Specimen</th>
<th id="76C55DFBA8080072C051FDA77561FD2A" box="[298,376,545,685]" gridcol="1" gridrow="0" pageId="15" pageNumber="60">ZSM 47595 holotype</th>
<th id="76C55DFBA8080072C0EDFDA775FEFD2A" box="[406,487,545,685]" gridcol="2" gridrow="0" pageId="15" pageNumber="60">ZSM 47596 paratype</th>
<th id="76C55DFBA8080072C377FDA77640FD2A" box="[524,601,545,685]" gridcol="3" gridrow="0" pageId="15" pageNumber="60">ZSM 47597 paratype</th>
<th id="76C55DFBA8080072C304FDA776D5FD2A" box="[639,716,545,685]" gridcol="4" gridrow="0" pageId="15" pageNumber="60">ZSM 47599 paratype</th>
<th id="76C55DFBA8080072C390FDA77721FD2A" box="[747,824,545,685]" gridcol="5" gridrow="0" pageId="15" pageNumber="60">ZSM 47598 paratype</th>
<th id="76C55DFBA8080072C22EFDA777BBFD2A" box="[853,930,545,685]" gridcol="6" gridrow="0" pageId="15" pageNumber="60">ZM - CBSU 5003- 128 paratype</th>
<th id="76C55DFBA8080072C2B9FDA77016FD2A" box="[962,1039,545,685]" gridcol="7" gridrow="0" pageId="15" pageNumber="60">ZM - CBSU 5001-1 paratype</th>
<th id="76C55DFBA8080072C54BFDA77064FD2A" box="[1072,1149,545,685]" gridcol="8" gridrow="0" pageId="15" pageNumber="60">ZM - CBSU 5003-60 paratype</th>
<th id="76C55DFBA8080072C5E6FDA770F3FD2A" box="[1181,1258,545,685]" gridcol="9" gridrow="0" pageId="15" pageNumber="60">ZM - CBSU 5022-23 paratype</th>
<th id="76C55DFBA8080072C471FDA7714EFD2A" box="[1290,1367,545,685]" gridcol="10" gridrow="0" pageId="15" pageNumber="60">ZM - CBSU 5024-1 paratype</th>
<th id="76C55DFBA8080072C40FFDA771D8FD2A" box="[1396,1473,545,685]" gridcol="11" gridrow="0" pageId="15" pageNumber="60">ZM - CBSU 5003-77 paratype</th>
<th id="76C55DFBA8080072C4A4FDA77235FD2A" box="[1503,1580,545,685]" gridcol="12" gridrow="0" pageId="15" pageNumber="60">PMR VP4679 paratype</th>
<th id="76C55DFBA8080072C732FDA7728FFD2A" box="[1609,1686,545,685]" gridcol="13" gridrow="0" pageId="15" pageNumber="60">PMR VP4680 paratype</th>
<th id="76C55DFBA8080072C7CFFDA77318FD2A" box="[1716,1793,545,685]" gridcol="14" gridrow="0" pageId="15" pageNumber="60">PMR VP4681 paratype</th>
<th id="76C55DFBA8080072C665FDA77372FD2A" box="[1822,1899,545,685]" gridcol="15" gridrow="0" pageId="15" pageNumber="60">PMR VP4682 paratype</th>
<th id="76C55DFBA8080072C6F7FDA773C4FD2A" box="[1932,2013,545,685]" gridcol="16" gridrow="0" pageId="15" pageNumber="60">PMR VP4683 paratype</th>
</tr>
<tr id="35143487A8080072C1ECFD3F73C4FC8D" box="[151,2013,697,778]" gridrow="1" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFD3F7515FC8D" box="[151,268,697,778]" gridcol="0" gridrow="1" pageId="15" pageNumber="60">Caudal-fin length Length of</th>
<td id="76C55DFBA8080072C051FD3F7561FC8D" box="[298,376,697,778]" gridcol="1" gridrow="1" pageId="15" pageNumber="60">21.8</td>
<td id="76C55DFBA8080072C0EDFD3F75FEFC8D" box="[406,487,697,778]" gridcol="2" gridrow="1" pageId="15" pageNumber="60">21.3</td>
<td id="76C55DFBA8080072C377FD3F7640FC8D" box="[524,601,697,778]" gridcol="3" gridrow="1" pageId="15" pageNumber="60">24.9</td>
<td id="76C55DFBA8080072C304FD3F76D5FC8D" box="[639,716,697,778]" gridcol="4" gridrow="1" pageId="15" pageNumber="60">22.8</td>
<td id="76C55DFBA8080072C390FD3F7721FC8D" box="[747,824,697,778]" gridcol="5" gridrow="1" pageId="15" pageNumber="60">23.4</td>
<td id="76C55DFBA8080072C22EFD3F77BBFC8D" box="[853,930,697,778]" gridcol="6" gridrow="1" pageId="15" pageNumber="60">26.5</td>
<td id="76C55DFBA8080072C2B9FD3F7016FC8D" box="[962,1039,697,778]" gridcol="7" gridrow="1" pageId="15" pageNumber="60">23.1</td>
<td id="76C55DFBA8080072C54BFD3F7064FC8D" box="[1072,1149,697,778]" gridcol="8" gridrow="1" pageId="15" pageNumber="60">23.8</td>
<td id="76C55DFBA8080072C5E6FD3F70F3FC8D" box="[1181,1258,697,778]" gridcol="9" gridrow="1" pageId="15" pageNumber="60">22.6</td>
<td id="76C55DFBA8080072C471FD3F714EFC8D" box="[1290,1367,697,778]" gridcol="10" gridrow="1" pageId="15" pageNumber="60">22.3</td>
<td id="76C55DFBA8080072C40FFD3F71D8FC8D" box="[1396,1473,697,778]" gridcol="11" gridrow="1" pageId="15" pageNumber="60">25.0</td>
<td id="76C55DFBA8080072C4A4FD3F7235FC8D" box="[1503,1580,697,778]" gridcol="12" gridrow="1" pageId="15" pageNumber="60">23.0</td>
<td id="76C55DFBA8080072C732FD3F728FFC8D" box="[1609,1686,697,778]" gridcol="13" gridrow="1" pageId="15" pageNumber="60">18.2</td>
<td id="76C55DFBA8080072C7CFFD3F7318FC8D" box="[1716,1793,697,778]" gridcol="14" gridrow="1" pageId="15" pageNumber="60">24.1</td>
<td id="76C55DFBA8080072C665FD3F7372FC8D" box="[1822,1899,697,778]" gridcol="15" gridrow="1" pageId="15" pageNumber="60">24.3</td>
<td id="76C55DFBA8080072C6F7FD3F73C4FC8D" box="[1932,2013,697,778]" gridcol="16" gridrow="1" pageId="15" pageNumber="60">damaged</td>
</tr>
<tr id="35143487A8080072C1ECFC9773C4FCC2" box="[151,2013,785,837]" gridrow="2" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFC977515FCC2" box="[151,268,785,837]" gridcol="0" gridrow="2" pageId="15" pageNumber="60">first dorsal- fin base</th>
<td id="76C55DFBA8080072C051FC977561FCC2" box="[298,376,785,837]" gridcol="1" gridrow="2" pageId="15" pageNumber="60">5.6</td>
<td id="76C55DFBA8080072C0EDFC9775FEFCC2" box="[406,487,785,837]" gridcol="2" gridrow="2" pageId="15" pageNumber="60">4.6</td>
<td id="76C55DFBA8080072C377FC977640FCC2" box="[524,601,785,837]" gridcol="3" gridrow="2" pageId="15" pageNumber="60">5.4</td>
<td id="76C55DFBA8080072C304FC9776D5FCC2" box="[639,716,785,837]" gridcol="4" gridrow="2" pageId="15" pageNumber="60">5.4</td>
<td id="76C55DFBA8080072C390FC977721FCC2" box="[747,824,785,837]" gridcol="5" gridrow="2" pageId="15" pageNumber="60">4.1</td>
<td id="76C55DFBA8080072C22EFC9777BBFCC2" box="[853,930,785,837]" gridcol="6" gridrow="2" pageId="15" pageNumber="60">5.3</td>
<td id="76C55DFBA8080072C2B9FC977016FCC2" box="[962,1039,785,837]" gridcol="7" gridrow="2" pageId="15" pageNumber="60">4.4</td>
<td id="76C55DFBA8080072C54BFC977064FCC2" box="[1072,1149,785,837]" gridcol="8" gridrow="2" pageId="15" pageNumber="60">4.6</td>
<td id="76C55DFBA8080072C5E6FC9770F3FCC2" box="[1181,1258,785,837]" gridcol="9" gridrow="2" pageId="15" pageNumber="60">5.1</td>
<td id="76C55DFBA8080072C471FC97714EFCC2" box="[1290,1367,785,837]" gridcol="10" gridrow="2" pageId="15" pageNumber="60">4.5</td>
<td id="76C55DFBA8080072C40FFC9771D8FCC2" box="[1396,1473,785,837]" gridcol="11" gridrow="2" pageId="15" pageNumber="60">5.3</td>
<td id="76C55DFBA8080072C4A4FC977235FCC2" box="[1503,1580,785,837]" gridcol="12" gridrow="2" pageId="15" pageNumber="60">5.8</td>
<td id="76C55DFBA8080072C732FC97728FFCC2" box="[1609,1686,785,837]" gridcol="13" gridrow="2" pageId="15" pageNumber="60">6.0</td>
<td id="76C55DFBA8080072C7CFFC977318FCC2" box="[1716,1793,785,837]" gridcol="14" gridrow="2" pageId="15" pageNumber="60">5.2</td>
<td id="76C55DFBA8080072C665FC977372FCC2" box="[1822,1899,785,837]" gridcol="15" gridrow="2" pageId="15" pageNumber="60">5.9</td>
<td id="76C55DFBA8080072C6F7FC9773C4FCC2" box="[1932,2013,785,837]" gridcol="16" gridrow="2" pageId="15" pageNumber="60">5.4</td>
</tr>
<tr id="35143487A8080072C1ECFCCA73C4FCF8" box="[151,2013,844,895]" gridrow="3" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFCCA7515FCF8" box="[151,268,844,895]" gridcol="0" gridrow="3" pageId="15" pageNumber="60">Postanal-fin distance</th>
<td id="76C55DFBA8080072C051FCCA7561FCF8" box="[298,376,844,895]" gridcol="1" gridrow="3" pageId="15" pageNumber="60">18.0</td>
<td id="76C55DFBA8080072C0EDFCCA75FEFCF8" box="[406,487,844,895]" gridcol="2" gridrow="3" pageId="15" pageNumber="60">17.1</td>
<td id="76C55DFBA8080072C377FCCA7640FCF8" box="[524,601,844,895]" gridcol="3" gridrow="3" pageId="15" pageNumber="60">17.8</td>
<td id="76C55DFBA8080072C304FCCA76D5FCF8" box="[639,716,844,895]" gridcol="4" gridrow="3" pageId="15" pageNumber="60">18.0</td>
<td id="76C55DFBA8080072C390FCCA7721FCF8" box="[747,824,844,895]" gridcol="5" gridrow="3" pageId="15" pageNumber="60">18.7</td>
<td id="76C55DFBA8080072C22EFCCA77BBFCF8" box="[853,930,844,895]" gridcol="6" gridrow="3" pageId="15" pageNumber="60">17.8</td>
<td id="76C55DFBA8080072C2B9FCCA7016FCF8" box="[962,1039,844,895]" gridcol="7" gridrow="3" pageId="15" pageNumber="60">18.2</td>
<td id="76C55DFBA8080072C54BFCCA7064FCF8" box="[1072,1149,844,895]" gridcol="8" gridrow="3" pageId="15" pageNumber="60">18.4</td>
<td id="76C55DFBA8080072C5E6FCCA70F3FCF8" box="[1181,1258,844,895]" gridcol="9" gridrow="3" pageId="15" pageNumber="60">18.8</td>
<td id="76C55DFBA8080072C471FCCA714EFCF8" box="[1290,1367,844,895]" gridcol="10" gridrow="3" pageId="15" pageNumber="60">17.6</td>
<td id="76C55DFBA8080072C40FFCCA71D8FCF8" box="[1396,1473,844,895]" gridcol="11" gridrow="3" pageId="15" pageNumber="60">18.9</td>
<td id="76C55DFBA8080072C4A4FCCA7235FCF8" box="[1503,1580,844,895]" gridcol="12" gridrow="3" pageId="15" pageNumber="60">17.5</td>
<td id="76C55DFBA8080072C732FCCA728FFCF8" box="[1609,1686,844,895]" gridcol="13" gridrow="3" pageId="15" pageNumber="60">19.7</td>
<td id="76C55DFBA8080072C7CFFCCA7318FCF8" box="[1716,1793,844,895]" gridcol="14" gridrow="3" pageId="15" pageNumber="60">17.8</td>
<td id="76C55DFBA8080072C665FCCA7372FCF8" box="[1822,1899,844,895]" gridcol="15" gridrow="3" pageId="15" pageNumber="60">17.1</td>
<td id="76C55DFBA8080072C6F7FCCA73C4FCF8" box="[1932,2013,844,895]" gridcol="16" gridrow="3" pageId="15" pageNumber="60">18.6</td>
</tr>
<tr id="35143487A8080072C1ECFC1273C4FC40" box="[151,2013,916,967]" gridrow="4" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFC127515FC40" box="[151,268,916,967]" gridcol="0" gridrow="4" pageId="15" pageNumber="60">Postdorsal- fin distance</th>
<td id="76C55DFBA8080072C051FC127561FC40" box="[298,376,916,967]" gridcol="1" gridrow="4" pageId="15" pageNumber="60">16.3</td>
<td id="76C55DFBA8080072C0EDFC1275FEFC40" box="[406,487,916,967]" gridcol="2" gridrow="4" pageId="15" pageNumber="60">17.8</td>
<td id="76C55DFBA8080072C377FC127640FC40" box="[524,601,916,967]" gridcol="3" gridrow="4" pageId="15" pageNumber="60">19.0</td>
<td id="76C55DFBA8080072C304FC1276D5FC40" box="[639,716,916,967]" gridcol="4" gridrow="4" pageId="15" pageNumber="60">18.2</td>
<td id="76C55DFBA8080072C390FC127721FC40" box="[747,824,916,967]" gridcol="5" gridrow="4" pageId="15" pageNumber="60">18.4</td>
<td id="76C55DFBA8080072C22EFC1277BBFC40" box="[853,930,916,967]" gridcol="6" gridrow="4" pageId="15" pageNumber="60">18.5</td>
<td id="76C55DFBA8080072C2B9FC127016FC40" box="[962,1039,916,967]" gridcol="7" gridrow="4" pageId="15" pageNumber="60">18.1</td>
<td id="76C55DFBA8080072C54BFC127064FC40" box="[1072,1149,916,967]" gridcol="8" gridrow="4" pageId="15" pageNumber="60">17.4</td>
<td id="76C55DFBA8080072C5E6FC1270F3FC40" box="[1181,1258,916,967]" gridcol="9" gridrow="4" pageId="15" pageNumber="60">16.4</td>
<td id="76C55DFBA8080072C471FC12714EFC40" box="[1290,1367,916,967]" gridcol="10" gridrow="4" pageId="15" pageNumber="60">17.9</td>
<td id="76C55DFBA8080072C40FFC1271D8FC40" box="[1396,1473,916,967]" gridcol="11" gridrow="4" pageId="15" pageNumber="60">18.4</td>
<td id="76C55DFBA8080072C4A4FC127235FC40" box="[1503,1580,916,967]" gridcol="12" gridrow="4" pageId="15" pageNumber="60">15.2</td>
<td id="76C55DFBA8080072C732FC12728FFC40" box="[1609,1686,916,967]" gridcol="13" gridrow="4" pageId="15" pageNumber="60">16.1</td>
<td id="76C55DFBA8080072C7CFFC127318FC40" box="[1716,1793,916,967]" gridcol="14" gridrow="4" pageId="15" pageNumber="60">17.0</td>
<td id="76C55DFBA8080072C665FC127372FC40" box="[1822,1899,916,967]" gridcol="15" gridrow="4" pageId="15" pageNumber="60">16.9</td>
<td id="76C55DFBA8080072C6F7FC1273C4FC40" box="[1932,2013,916,967]" gridcol="16" gridrow="4" pageId="15" pageNumber="60">17.9</td>
</tr>
<tr id="35143487A8080072C1ECFC5A73C4FB88" box="[151,2013,988,1039]" gridrow="5" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFC5A73C4FB88" box="[151,2013,988,1039]" colspan="17" colspanRight="16" gridcol="0" gridrow="5" pageId="15" pageNumber="60">% of head length</th>
</tr>
<tr id="35143487A8080072C1ECFBA473C4FBBF" box="[151,2013,1058,1080]" gridrow="6" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFBA47515FBBF" box="[151,268,1058,1080]" gridcol="0" gridrow="6" pageId="15" pageNumber="60">Head depth*</th>
<td id="76C55DFBA8080072C051FBA47561FBBF" box="[298,376,1058,1080]" gridcol="1" gridrow="6" pageId="15" pageNumber="60">56.7*</td>
<td id="76C55DFBA8080072C0EDFBA475FEFBBF" box="[406,487,1058,1080]" gridcol="2" gridrow="6" pageId="15" pageNumber="60">54.7*</td>
<td id="76C55DFBA8080072C377FBA47640FBBF" box="[524,601,1058,1080]" gridcol="3" gridrow="6" pageId="15" pageNumber="60">55.8*</td>
<td id="76C55DFBA8080072C304FBA476D5FBBF" box="[639,716,1058,1080]" gridcol="4" gridrow="6" pageId="15" pageNumber="60">44.9</td>
<td id="76C55DFBA8080072C390FBA47721FBBF" box="[747,824,1058,1080]" gridcol="5" gridrow="6" pageId="15" pageNumber="60">46.2</td>
<td id="76C55DFBA8080072C22EFBA477BBFBBF" box="[853,930,1058,1080]" gridcol="6" gridrow="6" pageId="15" pageNumber="60">47.0</td>
<td id="76C55DFBA8080072C2B9FBA47016FBBF" box="[962,1039,1058,1080]" gridcol="7" gridrow="6" pageId="15" pageNumber="60">42.2</td>
<td id="76C55DFBA8080072C54BFBA47064FBBF" box="[1072,1149,1058,1080]" gridcol="8" gridrow="6" pageId="15" pageNumber="60">43.0</td>
<td id="76C55DFBA8080072C5E6FBA470F3FBBF" box="[1181,1258,1058,1080]" gridcol="9" gridrow="6" pageId="15" pageNumber="60">44.8</td>
<td id="76C55DFBA8080072C471FBA4714EFBBF" box="[1290,1367,1058,1080]" gridcol="10" gridrow="6" pageId="15" pageNumber="60">47.7</td>
<td id="76C55DFBA8080072C40FFBA471D8FBBF" box="[1396,1473,1058,1080]" gridcol="11" gridrow="6" pageId="15" pageNumber="60">49.8</td>
<td id="76C55DFBA8080072C4A4FBA47235FBBF" box="[1503,1580,1058,1080]" gridcol="12" gridrow="6" pageId="15" pageNumber="60">56.5*</td>
<td id="76C55DFBA8080072C732FBA4728FFBBF" box="[1609,1686,1058,1080]" gridcol="13" gridrow="6" pageId="15" pageNumber="60">61.0*</td>
<td id="76C55DFBA8080072C7CFFBA47318FBBF" box="[1716,1793,1058,1080]" gridcol="14" gridrow="6" pageId="15" pageNumber="60">56.6*</td>
<td id="76C55DFBA8080072C665FBA47372FBBF" box="[1822,1899,1058,1080]" gridcol="15" gridrow="6" pageId="15" pageNumber="60">48.2</td>
<td id="76C55DFBA8080072C6F7FBA473C4FBBF" box="[1932,2013,1058,1080]" gridcol="16" gridrow="6" pageId="15" pageNumber="60">41.0</td>
</tr>
<tr id="35143487A8080072C1ECFBCE73C4FBFE" box="[151,2013,1096,1145]" gridrow="7" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFBCE7515FBFE" box="[151,268,1096,1145]" gridcol="0" gridrow="7" pageId="15" pageNumber="60">Head width* Horizontal</th>
<td id="76C55DFBA8080072C051FBCE7561FBFE" box="[298,376,1096,1145]" gridcol="1" gridrow="7" pageId="15" pageNumber="60">95.8*</td>
<td id="76C55DFBA8080072C0EDFBCE75FEFBFE" box="[406,487,1096,1145]" gridcol="2" gridrow="7" pageId="15" pageNumber="60">101.2*</td>
<td id="76C55DFBA8080072C377FBCE7640FBFE" box="[524,601,1096,1145]" gridcol="3" gridrow="7" pageId="15" pageNumber="60">94.4*</td>
<td id="76C55DFBA8080072C304FBCE76D5FBFE" box="[639,716,1096,1145]" gridcol="4" gridrow="7" pageId="15" pageNumber="60">86.8</td>
<td id="76C55DFBA8080072C390FBCE7721FBFE" box="[747,824,1096,1145]" gridcol="5" gridrow="7" pageId="15" pageNumber="60">87.3</td>
<td id="76C55DFBA8080072C22EFBCE77BBFBFE" box="[853,930,1096,1145]" gridcol="6" gridrow="7" pageId="15" pageNumber="60">90.7</td>
<td id="76C55DFBA8080072C2B9FBCE7016FBFE" box="[962,1039,1096,1145]" gridcol="7" gridrow="7" pageId="15" pageNumber="60">85.8</td>
<td id="76C55DFBA8080072C54BFBCE7064FBFE" box="[1072,1149,1096,1145]" gridcol="8" gridrow="7" pageId="15" pageNumber="60">84.5</td>
<td id="76C55DFBA8080072C5E6FBCE70F3FBFE" box="[1181,1258,1096,1145]" gridcol="9" gridrow="7" pageId="15" pageNumber="60">86.4</td>
<td id="76C55DFBA8080072C471FBCE714EFBFE" box="[1290,1367,1096,1145]" gridcol="10" gridrow="7" pageId="15" pageNumber="60">89.1</td>
<td id="76C55DFBA8080072C40FFBCE71D8FBFE" box="[1396,1473,1096,1145]" gridcol="11" gridrow="7" pageId="15" pageNumber="60">94.8</td>
<td id="76C55DFBA8080072C4A4FBCE7235FBFE" box="[1503,1580,1096,1145]" gridcol="12" gridrow="7" pageId="15" pageNumber="60">103.7*</td>
<td id="76C55DFBA8080072C732FBCE728FFBFE" box="[1609,1686,1096,1145]" gridcol="13" gridrow="7" pageId="15" pageNumber="60">105.0*</td>
<td id="76C55DFBA8080072C7CFFBCE7318FBFE" box="[1716,1793,1096,1145]" gridcol="14" gridrow="7" pageId="15" pageNumber="60">103.7*</td>
<td id="76C55DFBA8080072C665FBCE7372FBFE" box="[1822,1899,1096,1145]" gridcol="15" gridrow="7" pageId="15" pageNumber="60">89.7</td>
<td id="76C55DFBA8080072C6F7FBCE73C4FBFE" box="[1932,2013,1096,1145]" gridcol="16" gridrow="7" pageId="15" pageNumber="60">89.1</td>
</tr>
<tr id="35143487A8080072C1ECFB0673C4FB11" box="[151,2013,1152,1174]" gridrow="8" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFB067515FB11" box="[151,268,1152,1174]" gridcol="0" gridrow="8" pageId="15" pageNumber="60">eye</th>
<td id="76C55DFBA8080072C051FB067561FB11" box="[298,376,1152,1174]" gridcol="1" gridrow="8" pageId="15" pageNumber="60">14.1˜</td>
<td id="76C55DFBA8080072C0EDFB0675FEFB11" box="[406,487,1152,1174]" gridcol="2" gridrow="8" pageId="15" pageNumber="60">14.2˜</td>
<td id="76C55DFBA8080072C377FB067640FB11" box="[524,601,1152,1174]" gridcol="3" gridrow="8" pageId="15" pageNumber="60">17.3</td>
<td id="76C55DFBA8080072C304FB0676D5FB11" box="[639,716,1152,1174]" gridcol="4" gridrow="8" pageId="15" pageNumber="60">17.4</td>
<td id="76C55DFBA8080072C390FB067721FB11" box="[747,824,1152,1174]" gridcol="5" gridrow="8" pageId="15" pageNumber="60">16.7</td>
<td id="76C55DFBA8080072C22EFB0677BBFB11" box="[853,930,1152,1174]" gridcol="6" gridrow="8" pageId="15" pageNumber="60">16.0</td>
<td id="76C55DFBA8080072C2B9FB067016FB11" box="[962,1039,1152,1174]" gridcol="7" gridrow="8" pageId="15" pageNumber="60">16.8</td>
<td id="76C55DFBA8080072C54BFB067064FB11" box="[1072,1149,1152,1174]" gridcol="8" gridrow="8" pageId="15" pageNumber="60">18.9</td>
<td id="76C55DFBA8080072C5E6FB0670F3FB11" box="[1181,1258,1152,1174]" gridcol="9" gridrow="8" pageId="15" pageNumber="60">17.4</td>
<td id="76C55DFBA8080072C471FB06714EFB11" box="[1290,1367,1152,1174]" gridcol="10" gridrow="8" pageId="15" pageNumber="60">21.6</td>
<td id="76C55DFBA8080072C40FFB0671D8FB11" box="[1396,1473,1152,1174]" gridcol="11" gridrow="8" pageId="15" pageNumber="60">19.7</td>
<td id="76C55DFBA8080072C4A4FB067235FB11" box="[1503,1580,1152,1174]" gridcol="12" gridrow="8" pageId="15" pageNumber="60">15.2˜</td>
<td id="76C55DFBA8080072C732FB06728FFB11" box="[1609,1686,1152,1174]" gridcol="13" gridrow="8" pageId="15" pageNumber="60">15.4˜</td>
<td id="76C55DFBA8080072C7CFFB067318FB11" box="[1716,1793,1152,1174]" gridcol="14" gridrow="8" pageId="15" pageNumber="60">16.1 ˜</td>
<td id="76C55DFBA8080072C665FB067372FB11" box="[1822,1899,1152,1174]" gridcol="15" gridrow="8" pageId="15" pageNumber="60">16.4</td>
<td id="76C55DFBA8080072C6F7FB0673C4FB11" box="[1932,2013,1152,1174]" gridcol="16" gridrow="8" pageId="15" pageNumber="60">15.1</td>
</tr>
<tr id="35143487A8080072C1ECFB1873C4FB69" box="[151,2013,1182,1262]" gridrow="9" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFB187515FB69" box="[151,268,1182,1262]" gridcol="0" gridrow="9" pageId="15" pageNumber="60">diameter˜ Interorbital distance˜</th>
<td id="76C55DFBA8080072C051FB187561FB69" box="[298,376,1182,1262]" gridcol="1" gridrow="9" pageId="15" pageNumber="60">14.8˜</td>
<td id="76C55DFBA8080072C0EDFB1875FEFB69" box="[406,487,1182,1262]" gridcol="2" gridrow="9" pageId="15" pageNumber="60">16.5˜</td>
<td id="76C55DFBA8080072C377FB187640FB69" box="[524,601,1182,1262]" gridcol="3" gridrow="9" pageId="15" pageNumber="60">10.5</td>
<td id="76C55DFBA8080072C304FB1876D5FB69" box="[639,716,1182,1262]" gridcol="4" gridrow="9" pageId="15" pageNumber="60">10.6</td>
<td id="76C55DFBA8080072C390FB187721FB69" box="[747,824,1182,1262]" gridcol="5" gridrow="9" pageId="15" pageNumber="60">11.3</td>
<td id="76C55DFBA8080072C22EFB1877BBFB69" box="[853,930,1182,1262]" gridcol="6" gridrow="9" pageId="15" pageNumber="60">10.7</td>
<td id="76C55DFBA8080072C2B9FB187016FB69" box="[962,1039,1182,1262]" gridcol="7" gridrow="9" pageId="15" pageNumber="60">11.4</td>
<td id="76C55DFBA8080072C54BFB187064FB69" box="[1072,1149,1182,1262]" gridcol="8" gridrow="9" pageId="15" pageNumber="60">9.2</td>
<td id="76C55DFBA8080072C5E6FB1870F3FB69" box="[1181,1258,1182,1262]" gridcol="9" gridrow="9" pageId="15" pageNumber="60">9.5</td>
<td id="76C55DFBA8080072C471FB18714EFB69" box="[1290,1367,1182,1262]" gridcol="10" gridrow="9" pageId="15" pageNumber="60">9.3</td>
<td id="76C55DFBA8080072C40FFB1871D8FB69" box="[1396,1473,1182,1262]" gridcol="11" gridrow="9" pageId="15" pageNumber="60">9.2</td>
<td id="76C55DFBA8080072C4A4FB187235FB69" box="[1503,1580,1182,1262]" gridcol="12" gridrow="9" pageId="15" pageNumber="60">17.3˜</td>
<td id="76C55DFBA8080072C732FB18728FFB69" box="[1609,1686,1182,1262]" gridcol="13" gridrow="9" pageId="15" pageNumber="60">19.3˜</td>
<td id="76C55DFBA8080072C7CFFB187318FB69" box="[1716,1793,1182,1262]" gridcol="14" gridrow="9" pageId="15" pageNumber="60">13.8˜</td>
<td id="76C55DFBA8080072C665FB187372FB69" box="[1822,1899,1182,1262]" gridcol="15" gridrow="9" pageId="15" pageNumber="60">11.9</td>
<td id="76C55DFBA8080072C6F7FB1873C4FB69" box="[1932,2013,1182,1262]" gridcol="16" gridrow="9" pageId="15" pageNumber="60">10.7</td>
</tr>
<tr id="35143487A8080072C1ECFA8773C4FA90" box="[151,2013,1281,1303]" gridrow="10" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFA877515FA90" box="[151,268,1281,1303]" gridcol="0" gridrow="10" pageId="15" pageNumber="60">Snout length</th>
<td id="76C55DFBA8080072C051FA877561FA90" box="[298,376,1281,1303]" gridcol="1" gridrow="10" pageId="15" pageNumber="60">27.8</td>
<td id="76C55DFBA8080072C0EDFA8775FEFA90" box="[406,487,1281,1303]" gridcol="2" gridrow="10" pageId="15" pageNumber="60">25.4</td>
<td id="76C55DFBA8080072C377FA877640FA90" box="[524,601,1281,1303]" gridcol="3" gridrow="10" pageId="15" pageNumber="60">24.6</td>
<td id="76C55DFBA8080072C304FA8776D5FA90" box="[639,716,1281,1303]" gridcol="4" gridrow="10" pageId="15" pageNumber="60">24.0</td>
<td id="76C55DFBA8080072C390FA877721FA90" box="[747,824,1281,1303]" gridcol="5" gridrow="10" pageId="15" pageNumber="60">26.8</td>
<td id="76C55DFBA8080072C22EFA8777BBFA90" box="[853,930,1281,1303]" gridcol="6" gridrow="10" pageId="15" pageNumber="60">25.2</td>
<td id="76C55DFBA8080072C2B9FA877016FA90" box="[962,1039,1281,1303]" gridcol="7" gridrow="10" pageId="15" pageNumber="60">27.9</td>
<td id="76C55DFBA8080072C54BFA877064FA90" box="[1072,1149,1281,1303]" gridcol="8" gridrow="10" pageId="15" pageNumber="60">24.6</td>
<td id="76C55DFBA8080072C5E6FA8770F3FA90" box="[1181,1258,1281,1303]" gridcol="9" gridrow="10" pageId="15" pageNumber="60">26.0</td>
<td id="76C55DFBA8080072C471FA87714EFA90" box="[1290,1367,1281,1303]" gridcol="10" gridrow="10" pageId="15" pageNumber="60">26.7</td>
<td id="76C55DFBA8080072C40FFA8771D8FA90" box="[1396,1473,1281,1303]" gridcol="11" gridrow="10" pageId="15" pageNumber="60">24.6</td>
<td id="76C55DFBA8080072C4A4FA877235FA90" box="[1503,1580,1281,1303]" gridcol="12" gridrow="10" pageId="15" pageNumber="60">24.3</td>
<td id="76C55DFBA8080072C732FA87728FFA90" box="[1609,1686,1281,1303]" gridcol="13" gridrow="10" pageId="15" pageNumber="60">27.9</td>
<td id="76C55DFBA8080072C7CFFA877318FA90" box="[1716,1793,1281,1303]" gridcol="14" gridrow="10" pageId="15" pageNumber="60">26.2</td>
<td id="76C55DFBA8080072C665FA877372FA90" box="[1822,1899,1281,1303]" gridcol="15" gridrow="10" pageId="15" pageNumber="60">24.4</td>
<td id="76C55DFBA8080072C6F7FA8773C4FA90" box="[1932,2013,1281,1303]" gridcol="16" gridrow="10" pageId="15" pageNumber="60">25.5</td>
</tr>
<tr id="35143487A8080072C1ECFAAF73C4FADB" box="[151,2013,1321,1372]" gridrow="11" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFAAF7515FADB" box="[151,268,1321,1372]" gridcol="0" gridrow="11" pageId="15" pageNumber="60">Postorbital distance</th>
<td id="76C55DFBA8080072C051FAAF7561FADB" box="[298,376,1321,1372]" gridcol="1" gridrow="11" pageId="15" pageNumber="60">60.1</td>
<td id="76C55DFBA8080072C0EDFAAF75FEFADB" box="[406,487,1321,1372]" gridcol="2" gridrow="11" pageId="15" pageNumber="60">59.1</td>
<td id="76C55DFBA8080072C377FAAF7640FADB" box="[524,601,1321,1372]" gridcol="3" gridrow="11" pageId="15" pageNumber="60">56.8</td>
<td id="76C55DFBA8080072C304FAAF76D5FADB" box="[639,716,1321,1372]" gridcol="4" gridrow="11" pageId="15" pageNumber="60">57.6</td>
<td id="76C55DFBA8080072C390FAAF7721FADB" box="[747,824,1321,1372]" gridcol="5" gridrow="11" pageId="15" pageNumber="60">54.9</td>
<td id="76C55DFBA8080072C22EFAAF77BBFADB" box="[853,930,1321,1372]" gridcol="6" gridrow="11" pageId="15" pageNumber="60">56.6</td>
<td id="76C55DFBA8080072C2B9FAAF7016FADB" box="[962,1039,1321,1372]" gridcol="7" gridrow="11" pageId="15" pageNumber="60">57.8</td>
<td id="76C55DFBA8080072C54BFAAF7064FADB" box="[1072,1149,1321,1372]" gridcol="8" gridrow="11" pageId="15" pageNumber="60">54.8</td>
<td id="76C55DFBA8080072C5E6FAAF70F3FADB" box="[1181,1258,1321,1372]" gridcol="9" gridrow="11" pageId="15" pageNumber="60">54.5</td>
<td id="76C55DFBA8080072C471FAAF714EFADB" box="[1290,1367,1321,1372]" gridcol="10" gridrow="11" pageId="15" pageNumber="60">58.8</td>
<td id="76C55DFBA8080072C40FFAAF71D8FADB" box="[1396,1473,1321,1372]" gridcol="11" gridrow="11" pageId="15" pageNumber="60">54.3</td>
<td id="76C55DFBA8080072C4A4FAAF7235FADB" box="[1503,1580,1321,1372]" gridcol="12" gridrow="11" pageId="15" pageNumber="60">62.5</td>
<td id="76C55DFBA8080072C732FAAF728FFADB" box="[1609,1686,1321,1372]" gridcol="13" gridrow="11" pageId="15" pageNumber="60">59.8</td>
<td id="76C55DFBA8080072C7CFFAAF7318FADB" box="[1716,1793,1321,1372]" gridcol="14" gridrow="11" pageId="15" pageNumber="60">59.3</td>
<td id="76C55DFBA8080072C665FAAF7372FADB" box="[1822,1899,1321,1372]" gridcol="15" gridrow="11" pageId="15" pageNumber="60">59.4</td>
<td id="76C55DFBA8080072C6F7FAAF73C4FADB" box="[1932,2013,1321,1372]" gridcol="16" gridrow="11" pageId="15" pageNumber="60">56.1</td>
</tr>
<tr id="35143487A8080072C1ECFAE973C4FA02" box="[151,2013,1391,1413]" gridrow="12" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFAE97515FA02" box="[151,268,1391,1413]" gridcol="0" gridrow="12" pageId="15" pageNumber="60">Mouth width</th>
<td id="76C55DFBA8080072C051FAE97561FA02" box="[298,376,1391,1413]" gridcol="1" gridrow="12" pageId="15" pageNumber="60">50.0*</td>
<td id="76C55DFBA8080072C0EDFAE975FEFA02" box="[406,487,1391,1413]" gridcol="2" gridrow="12" pageId="15" pageNumber="60">54.9*</td>
<td id="76C55DFBA8080072C377FAE97640FA02" box="[524,601,1391,1413]" gridcol="3" gridrow="12" pageId="15" pageNumber="60">50.7*</td>
<td id="76C55DFBA8080072C304FAE976D5FA02" box="[639,716,1391,1413]" gridcol="4" gridrow="12" pageId="15" pageNumber="60">37.9</td>
<td id="76C55DFBA8080072C390FAE97721FA02" box="[747,824,1391,1413]" gridcol="5" gridrow="12" pageId="15" pageNumber="60">45.8</td>
<td id="76C55DFBA8080072C22EFAE977BBFA02" box="[853,930,1391,1413]" gridcol="6" gridrow="12" pageId="15" pageNumber="60">41.4</td>
<td id="76C55DFBA8080072C2B9FAE97016FA02" box="[962,1039,1391,1413]" gridcol="7" gridrow="12" pageId="15" pageNumber="60">39.8</td>
<td id="76C55DFBA8080072C54BFAE97064FA02" box="[1072,1149,1391,1413]" gridcol="8" gridrow="12" pageId="15" pageNumber="60">36.3</td>
<td id="76C55DFBA8080072C5E6FAE970F3FA02" box="[1181,1258,1391,1413]" gridcol="9" gridrow="12" pageId="15" pageNumber="60">37.4</td>
<td id="76C55DFBA8080072C471FAE9714EFA02" box="[1290,1367,1391,1413]" gridcol="10" gridrow="12" pageId="15" pageNumber="60">41.0</td>
<td id="76C55DFBA8080072C40FFAE971D8FA02" box="[1396,1473,1391,1413]" gridcol="11" gridrow="12" pageId="15" pageNumber="60">44.5</td>
<td id="76C55DFBA8080072C4A4FAE97235FA02" box="[1503,1580,1391,1413]" gridcol="12" gridrow="12" pageId="15" pageNumber="60">55.2*</td>
<td id="76C55DFBA8080072C732FAE9728FFA02" box="[1609,1686,1391,1413]" gridcol="13" gridrow="12" pageId="15" pageNumber="60">55.8*</td>
<td id="76C55DFBA8080072C7CFFAE97318FA02" box="[1716,1793,1391,1413]" gridcol="14" gridrow="12" pageId="15" pageNumber="60">49.1*</td>
<td id="76C55DFBA8080072C665FAE97372FA02" box="[1822,1899,1391,1413]" gridcol="15" gridrow="12" pageId="15" pageNumber="60">43.6</td>
<td id="76C55DFBA8080072C6F7FAE973C4FA02" box="[1932,2013,1391,1413]" gridcol="16" gridrow="12" pageId="15" pageNumber="60">38.2</td>
</tr>
<tr id="35143487A8080072C1ECFA1173C4FA4C" box="[151,2013,1431,1483]" gridrow="13" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFA117515FA4C" box="[151,268,1431,1483]" gridcol="0" gridrow="13" pageId="15" pageNumber="60">Dermal fold length</th>
<td id="76C55DFBA8080072C051FA117561FA4C" box="[298,376,1431,1483]" gridcol="1" gridrow="13" pageId="15" pageNumber="60">16.6</td>
<td id="76C55DFBA8080072C0EDFA1175FEFA4C" box="[406,487,1431,1483]" gridcol="2" gridrow="13" pageId="15" pageNumber="60">damaged</td>
<td id="76C55DFBA8080072C377FA117640FA4C" box="[524,601,1431,1483]" gridcol="3" gridrow="13" pageId="15" pageNumber="60">19.7</td>
<td id="76C55DFBA8080072C304FA1176D5FA4C" box="[639,716,1431,1483]" gridcol="4" gridrow="13" pageId="15" pageNumber="60">15.9</td>
<td id="76C55DFBA8080072C390FA117721FA4C" box="[747,824,1431,1483]" gridcol="5" gridrow="13" pageId="15" pageNumber="60">20.5</td>
<td id="76C55DFBA8080072C22EFA1177BBFA4C" box="[853,930,1431,1483]" gridcol="6" gridrow="13" pageId="15" pageNumber="60">15.5</td>
<td id="76C55DFBA8080072C2B9FA117016FA4C" box="[962,1039,1431,1483]" gridcol="7" gridrow="13" pageId="15" pageNumber="60">13.4</td>
<td id="76C55DFBA8080072C54BFA117064FA4C" box="[1072,1149,1431,1483]" gridcol="8" gridrow="13" pageId="15" pageNumber="60">12.2</td>
<td id="76C55DFBA8080072C5E6FA1170F3FA4C" box="[1181,1258,1431,1483]" gridcol="9" gridrow="13" pageId="15" pageNumber="60">14.9</td>
<td id="76C55DFBA8080072C471FA11714EFA4C" box="[1290,1367,1431,1483]" gridcol="10" gridrow="13" pageId="15" pageNumber="60">18.8</td>
<td id="76C55DFBA8080072C40FFA1171D8FA4C" box="[1396,1473,1431,1483]" gridcol="11" gridrow="13" pageId="15" pageNumber="60">17.7</td>
<td id="76C55DFBA8080072C4A4FA117235FA4C" box="[1503,1580,1431,1483]" gridcol="12" gridrow="13" pageId="15" pageNumber="60">15.4</td>
<td id="76C55DFBA8080072C732FA11728FFA4C" box="[1609,1686,1431,1483]" gridcol="13" gridrow="13" pageId="15" pageNumber="60">17.1</td>
<td id="76C55DFBA8080072C7CFFA117318FA4C" box="[1716,1793,1431,1483]" gridcol="14" gridrow="13" pageId="15" pageNumber="60">15.5</td>
<td id="76C55DFBA8080072C665FA117372FA4C" box="[1822,1899,1431,1483]" gridcol="15" gridrow="13" pageId="15" pageNumber="60">14.1</td>
<td id="76C55DFBA8080072C6F7FA1173C4FA4C" box="[1932,2013,1431,1483]" gridcol="16" gridrow="13" pageId="15" pageNumber="60">14.0</td>
</tr>
<tr id="35143487A8080072C1ECFA5973C4F994" box="[151,2013,1503,1555]" gridrow="14" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECFA597515F994" box="[151,268,1503,1555]" gridcol="0" gridrow="14" pageId="15" pageNumber="60">Dermal fold depth</th>
<td id="76C55DFBA8080072C051FA597561F994" box="[298,376,1503,1555]" gridcol="1" gridrow="14" pageId="15" pageNumber="60">6.8</td>
<td id="76C55DFBA8080072C0EDFA5975FEF994" box="[406,487,1503,1555]" gridcol="2" gridrow="14" pageId="15" pageNumber="60">damaged</td>
<td id="76C55DFBA8080072C377FA597640F994" box="[524,601,1503,1555]" gridcol="3" gridrow="14" pageId="15" pageNumber="60">6.9</td>
<td id="76C55DFBA8080072C304FA5976D5F994" box="[639,716,1503,1555]" gridcol="4" gridrow="14" pageId="15" pageNumber="60">6.1</td>
<td id="76C55DFBA8080072C390FA597721F994" box="[747,824,1503,1555]" gridcol="5" gridrow="14" pageId="15" pageNumber="60">5.9</td>
<td id="76C55DFBA8080072C22EFA5977BBF994" box="[853,930,1503,1555]" gridcol="6" gridrow="14" pageId="15" pageNumber="60">6.1</td>
<td id="76C55DFBA8080072C2B9FA597016F994" box="[962,1039,1503,1555]" gridcol="7" gridrow="14" pageId="15" pageNumber="60">5.0</td>
<td id="76C55DFBA8080072C54BFA597064F994" box="[1072,1149,1503,1555]" gridcol="8" gridrow="14" pageId="15" pageNumber="60">4.7</td>
<td id="76C55DFBA8080072C5E6FA5970F3F994" box="[1181,1258,1503,1555]" gridcol="9" gridrow="14" pageId="15" pageNumber="60">5.5</td>
<td id="76C55DFBA8080072C471FA59714EF994" box="[1290,1367,1503,1555]" gridcol="10" gridrow="14" pageId="15" pageNumber="60">5.6</td>
<td id="76C55DFBA8080072C40FFA5971D8F994" box="[1396,1473,1503,1555]" gridcol="11" gridrow="14" pageId="15" pageNumber="60">4.8</td>
<td id="76C55DFBA8080072C4A4FA597235F994" box="[1503,1580,1503,1555]" gridcol="12" gridrow="14" pageId="15" pageNumber="60">5.8</td>
<td id="76C55DFBA8080072C732FA59728FF994" box="[1609,1686,1503,1555]" gridcol="13" gridrow="14" pageId="15" pageNumber="60">5.3</td>
<td id="76C55DFBA8080072C7CFFA597318F994" box="[1716,1793,1503,1555]" gridcol="14" gridrow="14" pageId="15" pageNumber="60">5.2</td>
<td id="76C55DFBA8080072C665FA597372F994" box="[1822,1899,1503,1555]" gridcol="15" gridrow="14" pageId="15" pageNumber="60">5.4</td>
<td id="76C55DFBA8080072C6F7FA5973C4F994" box="[1932,2013,1503,1555]" gridcol="16" gridrow="14" pageId="15" pageNumber="60">5.2</td>
</tr>
<tr id="35143487A8080072C1ECF9A173C4F9DC" box="[151,2013,1575,1627]" gridrow="15" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECF9A17515F9DC" box="[151,268,1575,1627]" gridcol="0" gridrow="15" pageId="15" pageNumber="60">Chin barbel length</th>
<td id="76C55DFBA8080072C051F9A17561F9DC" box="[298,376,1575,1627]" gridcol="1" gridrow="15" pageId="15" pageNumber="60">8.3</td>
<td id="76C55DFBA8080072C0EDF9A175FEF9DC" box="[406,487,1575,1627]" gridcol="2" gridrow="15" pageId="15" pageNumber="60">9.4</td>
<td id="76C55DFBA8080072C377F9A17640F9DC" box="[524,601,1575,1627]" gridcol="3" gridrow="15" pageId="15" pageNumber="60">9.3</td>
<td id="76C55DFBA8080072C304F9A176D5F9DC" box="[639,716,1575,1627]" gridcol="4" gridrow="15" pageId="15" pageNumber="60">8.6</td>
<td id="76C55DFBA8080072C390F9A17721F9DC" box="[747,824,1575,1627]" gridcol="5" gridrow="15" pageId="15" pageNumber="60">7.8</td>
<td id="76C55DFBA8080072C22EF9A177BBF9DC" box="[853,930,1575,1627]" gridcol="6" gridrow="15" pageId="15" pageNumber="60">9.5</td>
<td id="76C55DFBA8080072C2B9F9A17016F9DC" box="[962,1039,1575,1627]" gridcol="7" gridrow="15" pageId="15" pageNumber="60">7.4</td>
<td id="76C55DFBA8080072C54BF9A17064F9DC" box="[1072,1149,1575,1627]" gridcol="8" gridrow="15" pageId="15" pageNumber="60">8.2</td>
<td id="76C55DFBA8080072C5E6F9A170F3F9DC" box="[1181,1258,1575,1627]" gridcol="9" gridrow="15" pageId="15" pageNumber="60">9.3</td>
<td id="76C55DFBA8080072C471F9A1714EF9DC" box="[1290,1367,1575,1627]" gridcol="10" gridrow="15" pageId="15" pageNumber="60">8.0</td>
<td id="76C55DFBA8080072C40FF9A171D8F9DC" box="[1396,1473,1575,1627]" gridcol="11" gridrow="15" pageId="15" pageNumber="60">7.3</td>
<td id="76C55DFBA8080072C4A4F9A17235F9DC" box="[1503,1580,1575,1627]" gridcol="12" gridrow="15" pageId="15" pageNumber="60">9.5</td>
<td id="76C55DFBA8080072C732F9A1728FF9DC" box="[1609,1686,1575,1627]" gridcol="13" gridrow="15" pageId="15" pageNumber="60">8.9</td>
<td id="76C55DFBA8080072C7CFF9A17318F9DC" box="[1716,1793,1575,1627]" gridcol="14" gridrow="15" pageId="15" pageNumber="60">8.2</td>
<td id="76C55DFBA8080072C665F9A17372F9DC" box="[1822,1899,1575,1627]" gridcol="15" gridrow="15" pageId="15" pageNumber="60">9.6</td>
<td id="76C55DFBA8080072C6F7F9A173C4F9DC" box="[1932,2013,1575,1627]" gridcol="16" gridrow="15" pageId="15" pageNumber="60">8.6</td>
</tr>
<tr id="35143487A8080072C1ECF9E473C4F912" box="[151,2013,1634,1685]" gridrow="16" pageId="15" pageNumber="60">
<th id="76C55DFBA8080072C1ECF9E47515F912" box="[151,268,1634,1685]" gridcol="0" gridrow="16" pageId="15" pageNumber="60">Lower jaw length</th>
<td id="76C55DFBA8080072C051F9E47561F912" box="[298,376,1634,1685]" gridcol="1" gridrow="16" pageId="15" pageNumber="60">37.6</td>
<td id="76C55DFBA8080072C0EDF9E475FEF912" box="[406,487,1634,1685]" gridcol="2" gridrow="16" pageId="15" pageNumber="60">36.2</td>
<td id="76C55DFBA8080072C377F9E47640F912" box="[524,601,1634,1685]" gridcol="3" gridrow="16" pageId="15" pageNumber="60">37.4</td>
<td id="76C55DFBA8080072C304F9E476D5F912" box="[639,716,1634,1685]" gridcol="4" gridrow="16" pageId="15" pageNumber="60">36.3</td>
<td id="76C55DFBA8080072C390F9E47721F912" box="[747,824,1634,1685]" gridcol="5" gridrow="16" pageId="15" pageNumber="60">34.5</td>
<td id="76C55DFBA8080072C22EF9E477BBF912" box="[853,930,1634,1685]" gridcol="6" gridrow="16" pageId="15" pageNumber="60">33.8</td>
<td id="76C55DFBA8080072C2B9F9E47016F912" box="[962,1039,1634,1685]" gridcol="7" gridrow="16" pageId="15" pageNumber="60">37.6</td>
<td id="76C55DFBA8080072C54BF9E47064F912" box="[1072,1149,1634,1685]" gridcol="8" gridrow="16" pageId="15" pageNumber="60">34.5</td>
<td id="76C55DFBA8080072C5E6F9E470F3F912" box="[1181,1258,1634,1685]" gridcol="9" gridrow="16" pageId="15" pageNumber="60">33.5</td>
<td id="76C55DFBA8080072C471F9E4714EF912" box="[1290,1367,1634,1685]" gridcol="10" gridrow="16" pageId="15" pageNumber="60">36.2</td>
<td id="76C55DFBA8080072C40FF9E471D8F912" box="[1396,1473,1634,1685]" gridcol="11" gridrow="16" pageId="15" pageNumber="60">34.4</td>
<td id="76C55DFBA8080072C4A4F9E47235F912" box="[1503,1580,1634,1685]" gridcol="12" gridrow="16" pageId="15" pageNumber="60">37.8</td>
<td id="76C55DFBA8080072C732F9E4728FF912" box="[1609,1686,1634,1685]" gridcol="13" gridrow="16" pageId="15" pageNumber="60">34.7</td>
<td id="76C55DFBA8080072C7CFF9E47318F912" box="[1716,1793,1634,1685]" gridcol="14" gridrow="16" pageId="15" pageNumber="60">33.2</td>
<td id="76C55DFBA8080072C665F9E47372F912" box="[1822,1899,1634,1685]" gridcol="15" gridrow="16" pageId="15" pageNumber="60">38.1</td>
<td id="76C55DFBA8080072C6F7F9E473C4F912" box="[1932,2013,1634,1685]" gridcol="16" gridrow="16" pageId="15" pageNumber="60">33.6</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9B36C5A817FF9DC1BCFF11718BFE2A" blockId="16.[151,1437,151,429]" pageId="16" pageNumber="61">
<taxonomicName id="4C244D46A817FF9DC1BCFF1175A7FF37" authorityName="Kovačić &amp; Esmaeili &amp; Zarei &amp; Abbasi &amp; Schliewen" authorityYear="2021" box="[199,446,151,177]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="61" phylum="Chordata" rank="species" species="persicus">
<emphasis id="B950EAD7A817FF9DC1BCFF1175A7FF37" box="[199,446,151,177]" italics="true" pageId="16" pageNumber="61">Benthophilus persicus</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C244D46A817FF9DC3CEFF1177F7FF36" authority="(Sauvage, 1874)" baseAuthorityName="Sauvage" baseAuthorityYear="1874" box="[693,1006,151,177]" class="Actinopterygii" family="Gobiidae" genus="Benthophilus" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="61" phylum="Chordata" rank="species" species="stellatus">
<emphasis id="B950EAD7A817FF9DC3CEFF117728FF36" box="[693,817,151,177]" italics="true" pageId="16" pageNumber="61">B. stellatus</emphasis>
(Sauvage, 1874)
</taxonomicName>
by its distribution in the southern Caspian Sea (
<emphasis id="B950EAD7A817FF9DC073FF3D753FFF53" box="[264,294,187,212]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
distribution in the Taganrog Bay of the Sea of Azov and Don river from mouth upstream to the upper stretch of Tsymlyansk reservoir) and by having the dermal fold rectangular (
<emphasis id="B950EAD7A817FF9DC281FF597001FF7F" box="[1018,1048,223,248]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
dermal fold roughly triangular), a moderate chin barbel, 1/32/3 of eye diameter in length (
<emphasis id="B950EAD7A817FF9DC262FE85772EFE9B" box="[793,823,259,284]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
large, about equal eye diameter), the origin of anal fin in front of vertical through origin of second dorsal fin (
<emphasis id="B950EAD7A817FF9DC274FEA17734FEC7" box="[783,813,295,320]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
under origin of the second dorsal fin), no tubercles on temporal and occipital head regions (
<emphasis id="B950EAD7A817FF9DC34FFECD764BFEE3" box="[564,594,331,356]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
tubercles present), transversal suborbital row
<emphasis id="B950EAD7A817FF9DC528FECC7073FEE3" box="[1107,1130,330,356]" italics="true" pageId="16" pageNumber="61">6i</emphasis>
below posterior end of row
<emphasis id="B950EAD7A817FF9DC1ECFEE974BCFE0E" box="[151,165,367,393]" italics="true" pageId="16" pageNumber="61">b</emphasis>
(
<emphasis id="B950EAD7A817FF9DC1CFFEE974CAFE0F" box="[180,211,367,392]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
below middle of row
<emphasis id="B950EAD7A817FF9DC0BFFEE975CBFE0E" box="[452,466,367,393]" italics="true" pageId="16" pageNumber="61">b</emphasis>
), anterior interorbital transversal row
<emphasis id="B950EAD7A817FF9DC203FEE9778DFE0F" box="[888,916,367,392]" italics="true" pageId="16" pageNumber="61">pa</emphasis>
with 12 papillae (
<emphasis id="B950EAD7A817FF9DC513FEE9709EFE0F" box="[1128,1159,367,392]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
35 papillae), maximum body width 15.122.9% of SL (
<emphasis id="B950EAD7A817FF9DC081FE157601FE2B" box="[506,536,403,428]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
22.828.3%), and a mouth width 36.355.8% of head length (
<emphasis id="B950EAD7A817FF9DC5AEFE1570EAFE2B" box="[1237,1267,403,428]" italics="true" pageId="16" pageNumber="61">vs.</emphasis>
55.862.2%).
</paragraph>
</subSubSection>
</treatment>
</document>