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<mods:title id="B638B306EE2A9C835C940253F1387425">Review of the genus Papagona Ball, 1935 (Hemiptera: Caliscelidae) including a new Neotropical species</mods:title>
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<paragraph id="8B9B36A2FFDDE37DFF5CFDE3FDB47556" blockId="7.[151,512,546,609]" box="[151,512,546,573]" pageId="7" pageNumber="114">
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<taxonomicName id="4C244D21FFDDE37DFF5CFDE3FDB47556" ID-CoL="4CJ52" authority="Ball, 1935" authorityName="Ball" authorityYear="1935" box="[151,512,546,573]" class="Insecta" family="Caliscelidae" genus="Papagona" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="papoosa">
<emphasis id="B950EAB0FFDDE37DFF5CFDE3FDB47556" bold="true" box="[151,512,546,573]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF5CFDE3FEC87557" bold="true" box="[151,380,547,573]" italics="true" pageId="7" pageNumber="114">Papagona papoosa</emphasis>
<bibRefCitation id="EFB54B53FFDDE37DFE48FDE2FDB47556" author="Ball, E. D." box="[387,512,546,572]" pageId="7" pageNumber="114" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball, 1935</bibRefCitation>
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<paragraph id="8B9B36A2FFDDE37DFF5CFD86FF5F750B" blockId="7.[151,512,546,609]" box="[151,235,582,609]" pageId="7" pageNumber="114">
(
<figureCitation id="131F2A27FFDDE37DFF6BFD86FF56750B" box="[160,226,582,609]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5</figureCitation>
)
</paragraph>
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<paragraph id="8B9B36A2FFDDE37DFF5CFD4BFD5675C9" blockId="7.[151,738,650,675]" box="[151,738,650,675]" pageId="7" pageNumber="114">
<treatmentCitationGroup id="AB34118CFFDDE37DFF5CFD4BFD5675C9" box="[151,738,650,675]" pageId="7" pageNumber="114">
<taxonomicName id="4C244D21FFDDE37DFF5CFD4BFEE875C8" ID-CoL="4CJ52" authorityName="Ball" authorityYear="1935" box="[151,348,651,674]" class="Insecta" family="Caliscelidae" genus="Papagona" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="papoosa">
<emphasis id="B950EAB0FFDDE37DFF5CFD4BFEE875C8" box="[151,348,651,674]" italics="true" pageId="7" pageNumber="114">Papagona papoosa</emphasis>
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<treatmentCitation id="0A8510B3FFDDE37DFEBDFD4BFE4A75C9" author="Ball, E. D." box="[374,510,650,675]" page="41" pageId="7" pageNumber="114" year="1935">
<bibRefCitation id="EFB54B53FFDDE37DFEBDFD4BFE4A75C9" author="Ball, E. D." box="[374,510,650,675]" pageId="7" pageNumber="114" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball 1935: 41</bibRefCitation>
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(Original description)
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<subSubSection id="C33E6529FFDDE37DFF5CFD11FB807085" pageId="7" pageNumber="114" type="materials_examined">
<paragraph id="8B9B36A2FFDDE37DFF5CFD11FBBB7586" blockId="7.[151,1437,721,2031]" box="[151,1039,721,748]" pageId="7" pageNumber="114">
<materialsCitation id="3B4C3CFFFFDDE37DFF5CFD11FBBE7586" box="[151,1034,721,748]" location="Arizona, Santa Cruz" pageId="7" pageNumber="114" specimenCount="1" stateProvince="Arizona, Santa Cruz" typeStatus="holotype">
<emphasis id="B950EAB0FFDDE37DFF5CFD11FE827581" bold="true" box="[151,310,721,747]" pageId="7" pageNumber="114">
<typeStatus id="549F8800FFDDE37DFF5CFD11FF667581" box="[151,210,721,747]" pageId="7" pageNumber="114">Type</typeStatus>
locality.
</emphasis>
<collectingRegion id="49E0F840FFDDE37DFEF7FD11FD947581" box="[316,544,721,747]" isEnumeration="true" pageId="7" pageNumber="114">Arizona, Santa Cruz</collectingRegion>
River, near Tubac (according to
<bibRefCitation id="EFB54B53FFDDE37DFC5BFD11FBB67586" author="Ball, E. D." box="[912,1026,721,748]" pageId="7" pageNumber="114" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball 1935</bibRefCitation>
)
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.
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CFD33FC217464" blockId="7.[151,1437,721,2031]" box="[199,917,755,782]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF0CFD33FE7A7467" bold="true" box="[199,462,755,781]" pageId="7" pageNumber="114">Amended description.</emphasis>
<emphasis id="B950EAB0FFDDE37DFE1EFD33FDD77467" box="[469,611,755,781]" italics="true" pageId="7" pageNumber="114">Body length.</emphasis>
Male,
<quantity id="4CDC9B47FFDDE37DFD7AFD33FCB37464" box="[689,775,755,782]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="7" pageNumber="114" unit="mm" value="2.6">2.6 mm</quantity>
(
<bibRefCitation id="EFB54B53FFDDE37DFCDDFD33FC3D7464" author="Ball, E. D." box="[790,905,755,782]" pageId="7" pageNumber="114" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball 1935</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CFCD6FD9D736B" blockId="7.[151,1437,721,2031]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF0CFCD6FEB9745A" box="[199,269,790,816]" italics="true" pageId="7" pageNumber="114">Color.</emphasis>
Males (
<figureCitation id="131F2A27FFDDE37DFEA4FCD6FE5A745B" box="[367,494,790,817]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5AC</figureCitation>
). Body mainly black with some regions yellowish-brown. Vertex, pronotum, and mesonotum yellowish-brown with broad median longitudinal white stripe crossing these structures (
<figureCitation id="131F2A27FFDDE37DFB38FCF9FAF9743E" box="[1267,1357,825,852]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
). Forewing with corium mostly black with apex light brown and two small circular pale maculae in between corium and clavus, one at base and another near midlength of claval suture; clavus brown (
<figureCitation id="131F2A27FFDDE37DFB8DFCBEFB1574F3" box="[1094,1185,894,921]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
). Legs mostly yellow; metafemur black (
<figureCitation id="131F2A27FFDDE37DFEACFC61FE5774D1" box="[359,483,929,956]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5B, C</figureCitation>
). Abdomen with tergite III with lateral white macula (
<figureCitation id="131F2A27FFDDE37DFB84FC61FB7974D1" box="[1103,1229,929,956]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A, C</figureCitation>
); each segment of abdomen with dorsal median pair of small and few conspicuous yellow maculae (
<figureCitation id="131F2A27FFDDE37DFBEEFC03FBCB74B4" box="[1061,1151,963,990]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
); in lateral view, sternites with lateral white macula (
<figureCitation id="131F2A27FFDDE37DFE09FC26FDA9736B" box="[450,541,998,1025]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CFBC9FE7973C4" blockId="7.[151,1437,721,2031]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF0CFBC9FDB77349" box="[199,515,1033,1059]" italics="true" pageId="7" pageNumber="114">Structure. Head and thorax.</emphasis>
Vertex (
<figureCitation id="131F2A27FFDDE37DFDA8FBC9FD0F734E" box="[611,699,1033,1060]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
) with anterior margin rounded; as long as basal width at midline; subequal to pronotum length at midline. Frons (
<figureCitation id="131F2A27FFDDE37DFD64FBEBFCB1732C" box="[687,773,1067,1094]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5B</figureCitation>
) without median carina; in lateral view (
<figureCitation id="131F2A27FFDDE37DFB04FBEBFA91732C" box="[1231,1317,1067,1094]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5C</figureCitation>
) each side with row of seven sensory pits bordering sublateral carina, four pits bordering frontogenal carina, and a pair of pits bordering fastigium (linking the other two rows providing a triangle-like arrangement—with two isolated pits within, aligned diagonally).
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CFB77FF4A727F" blockId="7.[151,1437,721,2031]" pageId="7" pageNumber="114">
Pronotum (
<figureCitation id="131F2A27FFDDE37DFE8DFB76FE2B73BB" box="[326,415,1206,1233]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
) with six sensory pits bordering lateral margins of disc and a group of fiver inner ones at posterior half. Mesonotum (
<figureCitation id="131F2A27FFDDE37DFE1AFB19FD9C739E" box="[465,552,1241,1268]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5A</figureCitation>
) without median carina; region outerad of lateral carina with eight to nine sensory pits.
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CFADEFBD372A9" blockId="7.[151,1437,721,2031]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF0CFADEFE8D7252" box="[199,313,1310,1336]" italics="true" pageId="7" pageNumber="114">Abdomen.</emphasis>
Tergite III (
<figureCitation id="131F2A27FFDDE37DFE08FADEFDAF7253" box="[451,539,1310,1337]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5C</figureCitation>
), in lateral view, with pair of sensory pits. Tergite IV (
<figureCitation id="131F2A27FFDDE37DFB4DFADEFB6B7253" box="[1158,1247,1310,1337]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5C</figureCitation>
), in lateral view, with pair of sensory pits followed by an isolated ventral pair. Tergite
<collectionCode id="ED35AE67FFDDE37DFC6DFA81FC0E7231" box="[934,954,1345,1371]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="7" pageNumber="114" type="Museum">V</collectionCode>
(
<figureCitation id="131F2A27FFDDE37DFC01FA81FB967236" box="[970,1058,1345,1372]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5C</figureCitation>
), in lateral view, with one row of three sensory pits followed by an isolated ventral pair. Tergite
<collectionCode id="ED35AE67FFDDE37DFC87FAA4FCDE7214" box="[844,874,1380,1406]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="114">VI</collectionCode>
(
<figureCitation id="131F2A27FFDDE37DFCB2FAA3FC647214" box="[889,976,1379,1406]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5C</figureCitation>
), in lateral view, with pair of sensory pits followed by an isolated ventral pair. Tergite VII (
<figureCitation id="131F2A27FFDDE37DFD0CFA46FCAB72CB" box="[711,799,1414,1441]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5C</figureCitation>
), in lateral view, with one row of three sensory pits followed by an isolated ventral pair. Tergite VIII, in lateral view, with pair of sensory pits.
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CFA0BFC067009" blockId="7.[151,1437,721,2031]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF0CFA0BFE32728F" box="[199,390,1483,1509]" italics="true" pageId="7" pageNumber="114">Male terminalia.</emphasis>
Pygofer (
<figureCitation id="131F2A27FFDDE37DFE3CFA0BFDE4728C" box="[503,592,1483,1510]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5D</figureCitation>
) with anterior margin concave; with posterior margin convex. Connective (
<figureCitation id="131F2A27FFDDE37DFF6BFA2EFF4C7163" box="[160,248,1518,1545]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5E</figureCitation>
) with tectiform structure bearing tectiductus; ventral support inverted Y-shaped. Gonostylus (
<figureCitation id="131F2A27FFDDE37DFAF6FA2EFA277163" box="[1341,1427,1518,1545]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5F</figureCitation>
) hook-like; anterior portion pointed; caudal portion curved anterodorsally; dorsal margin follows almost straight with a rounded protuberance in between anterior and median third; ventral margin mostly rounded; median third longer than high, setose. Endosoma (
<figureCitation id="131F2A27FFDDE37DFDF9F996FD1F711B" box="[562,683,1622,1649]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5G, H</figureCitation>
) enclosing almost all phallobase and aedeagus lengths laterally and ventrally; asymmetrical, with two different sides linked ventrally: one side is longer and with apex curved ventrally, comma-like; and other side is shorter and apically truncated, bearing sub-triangular expansion ventrally directed to the longest side of endosoma at aedeagus midlength (
<figureCitation id="131F2A27FFDDE37DFD2DF97EFC8B71B3" box="[742,831,1726,1753]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5G</figureCitation>
). Phallobase membranous, shorter than endosoma; enclosing aedeagus half-length laterally and ventrally; slightly visible in lateral view, apically and dorsally. Aedeagus (
<figureCitation id="131F2A27FFDDE37DFF6BF8C3FEAC7074" box="[160,280,1795,1822]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5G, H</figureCitation>
) opened dorsally; apically, narrowing and with pair of aedeagal hooks (
<figureCitation id="131F2A27FFDDE37DFBF8F8C3FB3F7074" box="[1075,1163,1795,1822]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5H</figureCitation>
), both with same length, longer and thinner than aedeagus, and curved anterodorsally, reaching the base of phallus. Suspensorium V-shaped (
<figureCitation id="131F2A27FFDDE37DFF6BF889FF4E700E" box="[160,250,1865,1892]" captionStart="FIGURE 5" captionStartId="8.[151,250,1787,1811]" captionTargetBox="[164,1426,322,1747]" captionTargetId="figure-65@8.[151,1437,308,1763]" captionTargetPageId="8" captionText="FIGURE 5. Papagona papoosa Ball, 1935, male holotype (USNM): (A) Dorsal habitus; (B) ventral habitus; (C) lateral habitus; (D) Pygofer, lateral view; (E) connective, anterior view; (F) style, lateral view; (GH) Phallus, dorsal view (G) and lateral view (H). Scale bar: AC = 0.5 mm; DH = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5225563" httpUri="https://zenodo.org/record/5225563/files/figure.png" pageId="7" pageNumber="114">Fig. 5G</figureCitation>
). Anal tube as long as wide; posterior margin rounded; setose.
</paragraph>
<paragraph id="8B9B36A2FFDDE37DFF0CF8ABFB807085" blockId="7.[151,1437,721,2031]" pageId="7" pageNumber="114">
<emphasis id="B950EAB0FFDDE37DFF0CF8ABFE2270EC" bold="true" box="[199,406,1899,1926]" pageId="7" pageNumber="114">Taxonomic notes.</emphasis>
Although
<bibRefCitation id="EFB54B53FFDDE37DFDC5F8ABFD3A70EC" author="Ball, E. D." box="[526,654,1899,1926]" pageId="7" pageNumber="114" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball (1935)</bibRefCitation>
states in the generic description that the vertex of
<taxonomicName id="4C244D21FFDDE37DFB0AF8ACFA8070EF" authorityName="Ball" authorityYear="1935" box="[1217,1332,1900,1925]" class="Insecta" family="Caliscelidae" genus="Papagona" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="B950EAB0FFDDE37DFB0AF8ACFA8070EF" box="[1217,1332,1900,1925]" italics="true" pageId="7" pageNumber="114">Papagona</emphasis>
</taxonomicName>
is longer than pronotum length (see Introduction), in the
<typeStatus id="549F8800FFDDE37DFD6FF84EFCB270C2" box="[676,774,1934,1960]" pageId="7" pageNumber="114" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C244D21FFDDE37DFCE2F84FFC1770C2" authorityName="Ball" authorityYear="1935" box="[809,931,1935,1960]" class="Insecta" family="Caliscelidae" genus="Papagona" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="papoosa">
<emphasis id="B950EAB0FFDDE37DFCE2F84FFC1770C2" box="[809,931,1935,1960]" italics="true" pageId="7" pageNumber="114">P. papoosa</emphasis>
</taxonomicName>
it appears only slightly longer than pronotum length (
<figureCitation id="131F2A27FFDDE37DFF25F871FEFD70A1" box="[238,329,1969,1995]" captionStart="FIGURE 1" captionStartId="3.[151,250,1217,1241]" captionTargetBox="[151,1436,737,1193]" captionTargetId="figure-318@3.[151,1436,737,1193]" captionTargetPageId="3" captionText="FIGURE 1. Live specimens of Papagona dietrichi sp. nov.: (A) male specimen; (B) female specimen. Photographs by Dr. C. H. Dietrich." figureDoi="http://doi.org/10.5281/zenodo.5225553" httpUri="https://zenodo.org/record/5225553/files/figure.png" pageId="7" pageNumber="114">Fig. 1A</figureCitation>
). However, this could be an intraspecific variation or artefact the specimen position when photographed. See taxonomic notes of
<taxonomicName id="4C244D21FFDDE37DFDC7F816FD337085" authority="De Freitas &amp; Zahniser &amp; Takiya, 2021" authorityName="De Freitas &amp; Zahniser &amp; Takiya" authorityYear="2021" box="[524,647,2005,2031]" class="Insecta" family="Caliscelidae" genus="Papagona" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="114" phylum="Arthropoda" rank="species" species="dietrichi" status="sp. nov.">
<emphasis id="B950EAB0FFDDE37DFDC7F816FD337085" box="[524,647,2005,2031]" italics="true" pageId="7" pageNumber="114">P. dietrichi</emphasis>
</taxonomicName>
<emphasis id="B950EAB0FFDDE37DFD45F815FD537085" bold="true" box="[654,743,2005,2031]" pageId="7" pageNumber="114">
<taxonomicNameLabel id="A26357CBFFDDE37DFD45F815FD537085" box="[654,743,2005,2031]" pageId="7" pageNumber="114" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
above for comparative notes.
</paragraph>
</subSubSection>
<subSubSection id="C33E6529FFD2E372FF0CFF56FE027677" pageId="8" pageNumber="115" type="distribution">
<paragraph id="8B9B36A2FFD2E372FF0CFF56FD4077DB" blockId="8.[151,1437,150,285]" box="[199,756,150,177]" pageId="8" pageNumber="115">
<emphasis id="B950EAB0FFD2E372FF0CFF56FED577DA" bold="true" box="[199,353,150,176]" pageId="8" pageNumber="115">Distribution.</emphasis>
<collectingCountry id="F3337632FFD2E372FEACFF56FDB677DA" box="[359,514,150,176]" name="United States of America" pageId="8" pageNumber="115">United States</collectingCountry>
:
<collectingRegion id="49E0F840FFD2E372FDC0FF56FDD177DA" box="[523,613,150,176]" country="United States of America" name="Arizona" pageId="8" pageNumber="115">Arizona</collectingRegion>
(
<bibRefCitation id="EFB54B53FFD2E372FDBFFF56FD5377DB" author="Ball, E. D." box="[628,743,150,177]" pageId="8" pageNumber="115" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball 1935</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B36A2FFD2E372FF0CFF7AFB5D77BF" blockId="8.[151,1437,150,285]" box="[199,1257,186,213]" pageId="8" pageNumber="115">
<emphasis id="B950EAB0FFD2E372FF0CFF7AFE1777BE" bold="true" box="[199,419,186,212]" pageId="8" pageNumber="115">Plant associations.</emphasis>
<taxonomicName id="4C244D21FFD2E372FE61FF7AFCFA77BE" authority="Scribn. ex Beal" authorityName="Beal" box="[426,846,186,213]" class="Liliopsida" family="Poaceae" genus="Muhlenbergia" kingdom="Plantae" order="Poales" pageId="8" pageNumber="115" phylum="Tracheophyta" rank="species" species="porteri">
<emphasis id="B950EAB0FFD2E372FE61FF7AFD2877BE" box="[426,668,186,212]" italics="true" pageId="8" pageNumber="115">Muhlenbergia porteri</emphasis>
Scribn. ex Beal
</taxonomicName>
(muhly grass,
<taxonomicName id="4C244D21FFD2E372FC3EFF7BFBE777BF" authorityName="Barnhart" authorityYear="1895" baseAuthorityName="R.Br." box="[1013,1107,187,213]" class="Liliopsida" family="Poaceae" kingdom="Plantae" order="Poales" pageId="8" pageNumber="115" phylum="Tracheophyta" rank="family">Poaceae</taxonomicName>
) (
<bibRefCitation id="EFB54B53FFD2E372FBA2FF7AFB6877BF" author="Ball, E. D." box="[1129,1244,186,213]" pageId="8" pageNumber="115" pagination="37 - 41" refId="ref7573" refString="Ball, E. D. (1935) Some new Issidae, with notes on others (Homoptera-Fulgoridae). Bulletin of the Brooklyn Entomological Society, 30, 37 - 41." type="journal article" year="1935">Ball 1935</bibRefCitation>
).
</paragraph>
<paragraph id="8B9B36A2FFD2E372FF0CFF1EFE027677" blockId="8.[151,1437,150,285]" pageId="8" pageNumber="115">
<emphasis id="B950EAB0FFD2E372FF0CFF1EFE217792" bold="true" box="[199,405,222,248]" pageId="8" pageNumber="115">Studied material.</emphasis>
<materialsCitation id="3B4C3CFFFFD2E372FE50FF1EFE027677" collectingDate="1932-08-06" collectionCode="USNM" collectorName="E. D. Ball" country="United States of America" location="Santa Cruz River" pageId="8" pageNumber="115" specimenCode="ENT 01513540" specimenCount="1" stateProvince="Arizona" typeStatus="holotype">
<emphasis id="B950EAB0FFD2E372FE50FF1EFDA57792" bold="true" box="[411,529,222,248]" pageId="8" pageNumber="115">
<typeStatus id="549F8800FFD2E372FE50FF1EFDB87792" box="[411,524,222,248]" pageId="8" pageNumber="115" type="holotype">Holotype</typeStatus>
:
</emphasis>
male (dissected herein),
<collectingCountry id="F3337632FFD2E372FCEEFF1EFCD47792" box="[805,864,222,248]" name="United States of America" pageId="8" pageNumber="115">USA</collectingCountry>
,
<collectingRegion id="49E0F840FFD2E372FCA2FF1EFC727792" box="[873,966,222,248]" country="United States of America" name="Arizona" pageId="8" pageNumber="115">Arizona</collectingRegion>
,
<location id="8EFB6079FFD2E372FC1BFF1EFB247792" LSID="urn:lsid:plazi:treatment:038D87B4FFDDE372FF5CFDE3FE027677:8EFB6079FFD2E372FC1BFF1EFB247792" box="[976,1168,222,248]" country="United States of America" name="Santa Cruz River" pageId="8" pageNumber="115" stateProvince="Arizona">Santa Cruz River</location>
,
<date id="FF9A1062FFD2E372FB51FF1EFA917792" box="[1178,1317,222,248]" pageId="8" pageNumber="115" value="1932-08-06">
<collectingDate id="EFDEE98AFFD2E372FB51FF1EFA917792" box="[1178,1317,222,248]" pageId="8" pageNumber="115" value="1932-08-06">6 Aug. 1932</collectingDate>
</date>
,
<collectorName id="26D15374FFD2E372FAE5FF1FFA297792" box="[1326,1437,222,249]" pageId="8" pageNumber="115">E. D. Ball</collectorName>
(
<collectionCode id="ED35AE67FFD2E372FF54FEC2FF467676" box="[159,242,258,284]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="115" type="Museum">USNM</collectionCode>
<specimenCode id="DB829ED9FFD2E372FF33FEC3FE1E7677" box="[248,426,258,285]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="8" pageNumber="115" type="Museum">ENT 01513540</specimenCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>