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<document id="DBF9DBA7A68E21893F35849A5841C49B" ID-DOI="10.5281/zenodo.174573" ID-GBIF-Dataset="8c531d6d-9d32-49fd-8449-12126dac1d5c" ID-ISSN="1175-5326" ID-Zenodo-Dep="174573" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459785485270" checkinUser="plazi" docAuthor="Mesibov, Robert" docDate="2006" docId="038D879B2469FF83FEC0FD527869F8BD" docLanguage="en" docName="zt01354p044.pdf.imd" docOrigin="Zootaxa 1354" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Tholerosoma Mesibov, 2006, n. gen." docType="treatment" docVersion="12" lastPageNumber="37" masterDocId="FFB4FFE3246FFF85FFC8FFAE7B5AFFDC" masterDocTitle="Dirt-encrusted and dragon millipedes (Diplopoda: Polydesmida: Paradoxosomatidae) from Queensland, Australia" masterLastPageNumber="44" masterPageNumber="31" pageNumber="37" updateTime="1698218411109" updateUser="plazi">
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<mods:title id="49D67F7CACC9AAA69A8566A5BFD9A039">Dirt-encrusted and dragon millipedes (Diplopoda: Polydesmida: Paradoxosomatidae) from Queensland, Australia</mods:title>
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<mods:namePart id="5379D3E67E2920EE267087D424592BB4">Mesibov, Robert</mods:namePart>
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<mods:title id="346DC894827B60BC658BAA43365891BF">Zootaxa</mods:title>
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<mods:date id="F66DE36F2750BD22298C69DD1B2D0E5B">2006</mods:date>
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<treatment id="038D879B2469FF83FEC0FD527869F8BD" ID-DOI="http://doi.org/10.5281/zenodo.6261703" ID-GBIF-Taxon="119348952" ID-Zenodo-Dep="6261703" LSID="urn:lsid:plazi:treatment:038D879B2469FF83FEC0FD527869F8BD" httpUri="http://treatment.plazi.org/id/038D879B2469FF83FEC0FD527869F8BD" lastPageNumber="37" pageId="6" pageNumber="37">
<subSubSection id="C33E65062469FF83FEC0FD52795BFCCA" box="[264,513,764,790]" pageId="6" pageNumber="37" type="nomenclature">
<paragraph id="8B9B368D2469FF83FEC0FD52795BFCCA" blockId="6.[264,513,764,790]" box="[264,513,764,790]" pageId="6" pageNumber="37">
<heading id="D0D381E12469FF83FEC0FD52795BFCCA" bold="true" box="[264,513,764,790]" fontSize="11" level="1" pageId="6" pageNumber="37" reason="1">
<emphasis id="B950EA9F2469FF83FEC0FD52795BFCCA" bold="true" box="[264,513,764,790]" pageId="6" pageNumber="37">
<taxonomicName id="4C244D0E2469FF83FEC0FD527AFDFCCA" ID-CoL="7Q4F4" box="[264,423,764,790]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B950EA9F2469FF83FEC0FD527AFDFCCA" bold="true" box="[264,423,764,790]" italics="true" pageId="6" pageNumber="37">Tholerosoma</emphasis>
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<taxonomicNameLabel id="A26357E42469FF83FE67FD52795BFCCA" box="[431,513,764,790]" pageId="6" pageNumber="37" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C33E65062469FF83FEC0FCE979BDFCA2" pageId="6" pageNumber="37" type="materials_examined">
<paragraph id="8B9B368D2469FF83FEC0FCE979BDFCA2" blockId="6.[264,986,838,894]" pageId="6" pageNumber="37">
<emphasis id="B950EA9F2469FF83FEC0FCE97AD7FC82" box="[264,397,839,862]" italics="true" pageId="6" pageNumber="37">
<typeStatus id="549F882F2469FF83FEC0FCE97A60FC82" box="[264,314,839,862]" pageId="6" pageNumber="37">Type</typeStatus>
species
</emphasis>
:
<taxonomicName id="4C244D0E2469FF83FE54FCE979C9FC82" box="[412,659,839,862]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="monteithi" status="sp. nov.">
<emphasis id="B950EA9F2469FF83FE54FCE979C9FC82" box="[412,659,839,862]" italics="true" pageId="6" pageNumber="37">Tholerosoma monteithi</emphasis>
</taxonomicName>
<emphasis id="B950EA9F2469FF83FD51FCE8798FFC82" bold="true" box="[665,725,838,862]" pageId="6" pageNumber="37">
<taxonomicNameLabel id="A26357E42469FF83FD51FCE8798FFC82" box="[665,725,838,862]" pageId="6" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, by present designation.
<emphasis id="B950EA9F2469FF83FEC0FCC97AA4FCA2" box="[264,510,871,894]" italics="true" pageId="6" pageNumber="37">Other assigned species</emphasis>
:
<taxonomicName id="4C244D0E2469FF83FDC4FCC979FEFCA2" box="[524,676,871,894]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="species" species="corrugatum" status="sp. nov.">
<emphasis id="B950EA9F2469FF83FDC4FCC979FEFCA2" box="[524,676,871,894]" italics="true" pageId="6" pageNumber="37">T. corrugatum</emphasis>
</taxonomicName>
<emphasis id="B950EA9F2469FF83FD63FCC879BDFCA2" bold="true" box="[683,743,870,894]" pageId="6" pageNumber="37">
<taxonomicNameLabel id="A26357E42469FF83FD63FCC879BDFCA2" box="[683,743,870,894]" pageId="6" pageNumber="37" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C33E65062469FF83FEC0FC01787FFB9D" pageId="6" pageNumber="37" type="diagnosis">
<paragraph id="8B9B368D2469FF83FEC0FC01787FFB9D" blockId="6.[264,1323,943,1889]" pageId="6" pageNumber="37">
<emphasis id="B950EA9F2469FF83FEC0FC017A25FC14" box="[264,383,943,968]" italics="true" pageId="6" pageNumber="37">Diagnosis</emphasis>
: Medium-sized paradoxosomatids (males
<quantity id="4CDC9B682469FF83FCBBFC0178A8FC15" box="[883,1010,943,969]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="6" pageNumber="37" unit="mm" value="20.0" valueMax="22.0" valueMin="18.0">1822 mm</quantity>
long) with head+20 rings; most of body and appendages with a tenacious, variably thick coating of fine soil particles; paranota wide, inflated, with thick, rounded margins; waist with deep longitudinal corrugations; gonopod telopodite unbranched.
</paragraph>
</subSubSection>
<subSubSection id="C33E65062469FF83FEF0FBE17F79FBB5" box="[312,1059,1103,1129]" pageId="6" pageNumber="37" type="etymology">
<paragraph id="8B9B368D2469FF83FEF0FBE17F79FBB5" blockId="6.[264,1323,943,1889]" box="[312,1059,1103,1129]" pageId="6" pageNumber="37">
<emphasis id="B950EA9F2469FF83FEF0FBE17AEFFBB4" box="[312,437,1103,1128]" italics="true" pageId="6" pageNumber="37">Etymology</emphasis>
: Greek
<emphasis id="B950EA9F2469FF83FDDBFBE17928FBB4" box="[531,626,1103,1128]" italics="true" pageId="6" pageNumber="37">tholeros</emphasis>
(“muddy”) +
<emphasis id="B950EA9F2469FF83FCD0FBE1780CFBB4" box="[792,854,1103,1128]" italics="true" pageId="6" pageNumber="37">soma</emphasis>
(“body”), neuter.
</paragraph>
</subSubSection>
<subSubSection id="C33E65062469FF83FEF0FBD97869F8BD" pageId="6" pageNumber="37" type="discussion">
<paragraph id="8B9B368D2469FF83FEF0FBD979EEFA25" blockId="6.[264,1323,943,1889]" pageId="6" pageNumber="37">
<emphasis id="B950EA9F2469FF83FEF0FBD97AC4FB4C" box="[312,414,1143,1168]" italics="true" pageId="6" pageNumber="37">Remarks</emphasis>
: On well-encrusted adults of
<taxonomicName id="4C244D0E2469FF83FCECFBD978E4FB4C" box="[804,958,1143,1168]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9F2469FF83FCECFBD978E4FB4C" box="[804,958,1143,1168]" italics="true" pageId="6" pageNumber="37">Tholerosoma</emphasis>
</taxonomicName>
, the only parts of the body completely free of soil particles are the prozonites, spiracles and gonopods. A variably thick layer of soil covers the head, collum, metazonites (including sternites), telson, all podomeres and all antennomeres. The coating consists of organic and crystalline soil particles up to 0.2 mm across, as well as tiny fragments of plant debris. On some parts of the body the coating can be pulled off in sheet-like pieces. Ultrasonic cleaning for up to 20 minutes did not entirely remove the coating from two test specimens. The coating appears to have the same colour and composition as
<taxonomicName id="4C244D0E2469FF83FC8AFA217886FA74" box="[834,988,1423,1448]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9F2469FF83FC8AFA217886FA74" box="[834,988,1423,1448]" italics="true" pageId="6" pageNumber="37">Tholerosoma</emphasis>
</taxonomicName>
gut contents. Stadium VII juveniles examined are all dirt-encrusted, although more lightly so than most adults; earlier juveniles are so far unknown.
</paragraph>
<paragraph id="8B9B368D2469FF83FEF0F9A97A83F935" blockId="6.[264,1323,943,1889]" pageId="6" pageNumber="37">
How the coating originates, how and when it is applied and how it sticks so tenaciously are subjects for future research. The rough texture seen on all surfaces under the coating may assist the “grip” of an adhesive material holding the coating in place. The coating may act as camouflage to help the animals avoid detection by predators who sense their prey by sight, smell, taste, or texture. It may also make captured
<taxonomicName id="4C244D0E2469FF83FB93F9097FAFF91C" box="[1115,1269,1703,1728]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9F2469FF83FB93F9097FAFF91C" box="[1115,1269,1703,1728]" italics="true" pageId="6" pageNumber="37">Tholerosoma</emphasis>
</taxonomicName>
less likely to be eaten.
</paragraph>
<paragraph id="8B9B368D2469FF83FEF0F9597869F8BD" blockId="6.[264,1323,943,1889]" pageId="6" pageNumber="37">
There is some variation in gonopod and paranotal details (noted below) within each of the two named
<taxonomicName id="4C244D0E2469FF83FE0AF8B17906F8E4" box="[450,604,1823,1848]" class="Diplopoda" family="Paradoxosomatidae" genus="Tholerosoma" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="37" phylum="Arthropoda" rank="genus">
<emphasis id="B950EA9F2469FF83FE0AF8B17906F8E4" box="[450,604,1823,1848]" italics="true" pageId="6" pageNumber="37">Tholerosoma</emphasis>
</taxonomicName>
taxa, and further study of these peculiar animals may show that splitting into additional species is justified.
</paragraph>
</subSubSection>
</treatment>
</document>