344 lines
53 KiB
XML
344 lines
53 KiB
XML
<document id="0DCE14CC1664815F5A2094327AF598EE" ID-DOI="10.11646/zootaxa.3948.3.5" ID-GBIF-Dataset="48fd46c1-a333-45ef-85fd-bb89451998da" ID-ISSN="1175-5326" ID-Zenodo-Dep="241857" ID-ZooBank="809AC895-779E-419D-8EBE-071F0ACCD72E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461044448529" checkinUser="plazi" docAuthor="Desutter-Grandcolas, Laure" docDate="2015" docId="038D8783FF8A3261FF7DFF39FB76FD97" docLanguage="en" docName="zt03948p496.pdf" docOrigin="Zootaxa 3948 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Upupagryllus Desutter-Grandcolas" docType="treatment" docVersion="9" lastPageNumber="486" masterDocId="FFB4FFFBFF953242FFEAFFA1FF9EFF8F" masterDocTitle="Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera" masterLastPageNumber="496" masterPageNumber="451" pageNumber="482" updateTime="1698576427866" updateUser="plazi">
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<mods:mods id="A9126E041C67B8E334A99815D0E141BD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DEB711053F6350234FE3D735D80DB75C">
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<mods:title id="CC4C88C40F3D318BFD18743671E81B61">Phalangopsidae crickets from Tropical Africa (Orthoptera, Grylloidea), with descriptions of new taxa and an identification key for African genera</mods:title>
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</mods:titleInfo>
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<mods:name id="980060B777783304B5486CED345E9193" type="personal">
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<mods:roleTerm id="A22CA5EB3EAE014E4DC3B584C08A77F2">Author</mods:roleTerm>
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<mods:namePart id="2BE4F92C15C006104D9E4579E2F8BD99">Desutter-Grandcolas, Laure</mods:namePart>
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<mods:typeOfResource id="76A0CCB2DEB821BE1B3D139D7547B511">text</mods:typeOfResource>
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<mods:title id="1E5B7BC655E8CE24A6A39FAF3B7D54D3">Zootaxa</mods:title>
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<mods:part id="3C30AB86662584475858E0457303C984">
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<mods:date id="7C02CDFF24068C8502A0BEB7BE508E81">2015</mods:date>
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<mods:detail id="3F27B023A1A6C8DAE5329D11DAF6CFF3" type="volume">
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<mods:number id="13E6E9BD8E7CBE56CCD14F5170EC24C1">3948</mods:number>
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<mods:detail id="5A8E244E17D7573E4038C030B9591E68" type="issue">
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<mods:number id="077E82135F418AAA6617A33F8BBC9E33">3</mods:number>
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<mods:start id="EC91D06C5A5F73957C8C4298C7E9574F">451</mods:start>
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<mods:end id="19E7901957FF1899E9508BB2FA1B4D36">496</mods:end>
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<mods:classification id="B6271E054AE1737D8E57B9C07D35C292">journal article</mods:classification>
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<mods:identifier id="DFEF552410B56EC8C9C02D3DAEF37E84" type="DOI">10.11646/zootaxa.3948.3.5</mods:identifier>
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<mods:identifier id="00EFDDC762503B62133A1A0327A355F6" type="GBIF-Dataset">48fd46c1-a333-45ef-85fd-bb89451998da</mods:identifier>
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<mods:identifier id="4C9D508CD3DB2414757064EBB1C7AAEA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="881E3788B7C6DB2D61D31DDAF1FEFEA9" type="Zenodo-Dep">241857</mods:identifier>
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<mods:identifier id="037E838EC1AA8FD0DF88682DF5E4CDA2" type="ZooBank">809AC895-779E-419D-8EBE-071F0ACCD72E</mods:identifier>
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<treatment id="038D8783FF8A3261FF7DFF39FB76FD97" ID-DOI="http://doi.org/10.5281/zenodo.5687709" ID-GBIF-Taxon="119610267" ID-Zenodo-Dep="5687709" LSID="urn:lsid:plazi:treatment:038D8783FF8A3261FF7DFF39FB76FD97" httpUri="http://treatment.plazi.org/id/038D8783FF8A3261FF7DFF39FB76FD97" lastPageId="35" lastPageNumber="486" pageId="31" pageNumber="482">
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<subSubSection id="C33E651EFF8A325DFF7DFF39FEB9FF5C" pageId="31" pageNumber="482" type="nomenclature">
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<paragraph id="8B9B3695FF8A325DFF7DFF39FD61FF3D" blockId="31.[151,767,151,211]" box="[151,767,151,178]" pageId="31" pageNumber="482">
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<heading id="D0D381F9FF8A325DFF7DFF39FD61FF3D" bold="true" box="[151,767,151,178]" fontSize="11" level="1" pageId="31" pageNumber="482" reason="1">
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<emphasis id="B950EA87FF8A325DFF7DFF39FD61FF3D" bold="true" box="[151,767,151,178]" pageId="31" pageNumber="482">
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Genus
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<taxonomicName id="4C244D16FF8A325DFF04FF36FD3FFF3D" ID-CoL="8P2LH" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[238,673,151,178]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="genus">
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<emphasis id="B950EA87FF8A325DFF04FF36FE0DFF3E" bold="true" box="[238,403,151,177]" italics="true" pageId="31" pageNumber="482">Upupagryllus</emphasis>
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Desutter-Grandcolas
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</taxonomicName>
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, n. gen.
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9B3695FF8A325DFF7DFF1BFEB9FF5C" blockId="31.[151,767,151,211]" box="[151,295,186,211]" pageId="31" pageNumber="482">
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(
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<figureCitation id="131F2A10FF8A325DFF75FF1BFE81FF5C" box="[159,287,186,211]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStartId-0="32.[151,250,1795,1817]" captionStartId-1="33.[151,250,1845,1867]" captionStartId-2="34.[151,250,1842,1864]" captionTargetBox-0="[152,1434,199,1773]" captionTargetBox-1="[155,1423,205,1815]" captionTargetBox-2="[179,1423,202,1802]" captionTargetId-0="figure@32.[151,1436,193,1773]" captionTargetId-1="figure@33.[150,1436,193,1824]" captionTargetId-2="figure@34.[178,1427,192,1804]" captionTargetPageId-0="32" captionTargetPageId-1="33" captionTargetPageId-2="34" captionText-0="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." captionText-1="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." captionText-2="FIGURE 14. Upupagryllus Desutter-Grandcolas, n. gen., Male genitalia: A – C, U. subapterus Desutter-Grandcolas, n. sp.; D – F, U. alatus Desutter-Grandcolas, n. sp. in dorsal (A, D), ventral (B, E) and lateral (C, F) views. Abbreviations: see page 4." httpUri-0="https://zenodo.org/record/241869/files/figure.png" httpUri-1="https://zenodo.org/record/241870/files/figure.png" httpUri-2="https://zenodo.org/record/241871/files/figure.png" pageId="31" pageNumber="482">Figs 12–14</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E651EFF8A325DFF7DFEA3FB9FFEB1" pageId="31" pageNumber="482" type="materials_examined">
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<paragraph id="8B9B3695FF8A325DFF7DFEA3FC13FE95" blockId="31.[151,1437,258,2011]" box="[151,909,258,283]" pageId="31" pageNumber="482">
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<emphasis id="B950EA87FF8A325DFF7DFEA3FEACFE94" bold="true" box="[151,306,258,283]" pageId="31" pageNumber="482">
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<typeStatus id="549F8837FF8A325DFF7DFEA3FF4CFE94" box="[151,210,258,283]" pageId="31" pageNumber="482">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C244D16FF8A325DFED3FEA3FCDBFE94" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[313,837,258,283]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
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<emphasis id="B950EA87FF8A325DFED3FEA3FDCBFE94" box="[313,597,258,283]" italics="true" pageId="31" pageNumber="482">Upupagryllus subapterus</emphasis>
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Desutter-Grandcolas
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26357FCFF8A325DFCBAFEA2FC13FE95" box="[848,909,259,282]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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<paragraph id="8B9B3695FF8A325DFF2DFE87FB9FFEB1" blockId="31.[151,1437,258,2011]" box="[199,1025,294,319]" pageId="31" pageNumber="482">
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<emphasis id="B950EA87FF8A325DFF2DFE87FE43FEB0" bold="true" box="[199,477,294,319]" pageId="31" pageNumber="482">Other species included.</emphasis>
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<taxonomicName id="4C244D16FF8A325DFE0EFE87FC24FEB0" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[484,954,294,319]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
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<emphasis id="B950EA87FF8A325DFE0EFE87FD57FEB0" box="[484,713,294,319]" italics="true" pageId="31" pageNumber="482">Upupagryllus alatus</emphasis>
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Desutter-Grandcolas
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26357FCFF8A325DFC2FFE86FB9FFEB1" box="[965,1025,295,318]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E651EFF8A325DFF2DFEEBFDBDFE09" pageId="31" pageNumber="482" type="etymology">
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<paragraph id="8B9B3695FF8A325DFF2DFEEBFDBDFE09" blockId="31.[151,1437,258,2011]" pageId="31" pageNumber="482">
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<emphasis id="B950EA87FF8A325DFF2DFEEBFED0FEEC" bold="true" box="[199,334,330,355]" pageId="31" pageNumber="482">Etymology.</emphasis>
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Genus named after the large amount of large setae occurring on the fastigium of the observed specimens, making a kind of crest.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E651EFF8A325DFF2DFE33FD3FFE24" box="[199,673,402,427]" pageId="31" pageNumber="482" type="distribution">
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<paragraph id="8B9B3695FF8A325DFF2DFE33FD3FFE24" blockId="31.[151,1437,258,2011]" box="[199,673,402,427]" pageId="31" pageNumber="482">
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<emphasis id="B950EA87FF8A325DFF2DFE33FEFEFE24" bold="true" box="[199,352,402,427]" pageId="31" pageNumber="482">Distribution.</emphasis>
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Known from
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<collectingCountry id="F3337605FF8A325DFE14FE33FDFDFE24" box="[510,611,402,427]" name="Tanzania" pageId="31" pageNumber="482">Tanzania</collectingCountry>
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only.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33E651EFF8A325DFF2DFE17FC6EFC64" pageId="31" pageNumber="482" type="diagnosis">
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<paragraph id="8B9B3695FF8A325DFF2DFE17FC6EFC64" blockId="31.[151,1437,258,2011]" pageId="31" pageNumber="482">
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<emphasis id="B950EA87FF8A325DFF2DFE17FEDFFE40" bold="true" box="[199,321,438,463]" pageId="31" pageNumber="482">Diagnosis.</emphasis>
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Small species covered with many, strong setae, ressembling
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<taxonomicName id="4C244D16FF8A325DFBE8FE17FB15FE40" box="[1026,1163,438,463]" class="Insecta" family="Trigonidiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="subFamily" subFamily="Nemobiinae">Nemobiinae</taxonomicName>
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crickets. Size small for
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<taxonomicName id="4C244D16FF8A325DFF7DFE7BFED4FE7C" box="[151,330,474,499]" class="Insecta" family="Phalangopsidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="family">Phalangopsidae</taxonomicName>
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, legs short (
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<figureCitation id="131F2A10FF8A325DFE37FE7BFDACFE7C" box="[477,562,474,499]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
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A–C). Head rounded dorsally, fastigium not separated from the vertex and wider than the scape (
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<figureCitation id="131F2A10FF8A325DFE65FE5FFE7CFD98" box="[399,482,510,535]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
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D, E); ocelli well developed, the lateral ones more widely separated from each other than the median ocellus from one lateral ocellus (
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<figureCitation id="131F2A10FF8A325DFD22FD83FC85FDB4" box="[712,795,546,571]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
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D). Eyes small, only slightly protruding, widely separate from each other (
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<figureCitation id="131F2A10FF8A325DFE88FDE7FE28FDD0" box="[354,438,582,607]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
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D, F). Maxillary palpi short (
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<figureCitation id="131F2A10FF8A325DFCE7FDE7FCF8FDD0" box="[781,870,582,607]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Figs 12</figureCitation>
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C, 13A, J). TI without tympanum (
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<taxonomicName id="4C244D16FF8A325DFAE8FDE7FD9AFD0C" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
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<emphasis id="B950EA87FF8A325DFAE8FDE7FE8CFD0C" italics="true" pageId="31" pageNumber="482">Upupagryllus subapterus</emphasis>
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Desutter-Grandcolas
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</taxonomicName>
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,
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<taxonomicNameLabel id="A26357FCFF8A325DFDFAFDCAFDD1FD0C" box="[528,591,619,643]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
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), or with one inner and one outer tympana (
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<taxonomicName id="4C244D16FF8A325DFBAFFDCBFAB2FD0C" box="[1093,1324,618,643]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="alatus">
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<emphasis id="B950EA87FF8A325DFBAFFDCBFAB2FD0C" box="[1093,1324,618,643]" italics="true" pageId="31" pageNumber="482">Upupagryllus alatus</emphasis>
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||
</taxonomicName>
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Desutter- Grandcolas, n. sp.); two long, ventral apical spurs. FIII thick, without a filiform apical part (
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<figureCitation id="131F2A10FF8A325DFB70FD2FFB74FD28" box="[1178,1258,654,679]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
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A, C). TIII with four pairs of short, subapical spurs; serrulation absent between subapical spurs, absent or scarce above them; three inner and three outer apical spurs. Basitarsomeres III with only one row of dorsal spines.
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<emphasis id="B950EA87FF8A325DFB73FD77FB7FFD60" bold="true" box="[1177,1249,726,751]" pageId="31" pageNumber="482">Males</emphasis>
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with reduced or well-developed FWs; in the latter case, FWs wide and short, with a complete stridulatory apparatus (
|
||
<figureCitation id="131F2A10FF8A325DFACFFD5BFAE6FC9C" box="[1317,1400,762,787]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="31" pageNumber="482">Fig. 13</figureCitation>
|
||
K). Male genitalia (
|
||
<figureCitation id="131F2A10FF8A325DFEACFCBFFE09FCB8" box="[326,407,798,823]" captionStart="FIGURE 14" captionStartId="34.[151,250,1842,1864]" captionTargetBox="[179,1423,202,1802]" captionTargetId="figure@34.[178,1427,192,1804]" captionTargetPageId="34" captionText="FIGURE 14. Upupagryllus Desutter-Grandcolas, n. gen., Male genitalia: A – C, U. subapterus Desutter-Grandcolas, n. sp.; D – F, U. alatus Desutter-Grandcolas, n. sp. in dorsal (A, D), ventral (B, E) and lateral (C, F) views. Abbreviations: see page 4." httpUri="https://zenodo.org/record/241871/files/figure.png" pageId="31" pageNumber="482">Fig. 14</figureCitation>
|
||
) characterized by the great development of a pair of acute, distal lobes directly connected to the rami; upper part of pseudepiphallus transverse; rami with a subdistal transverse crest; pseudepiphallic parameres, ectophallic fold and ectophallic dorsal valves reduced.
|
||
<emphasis id="B950EA87FF8A325DFC5DFCC7FB87FCF0" bold="true" box="[951,1049,870,895]" pageId="31" pageNumber="482">Females</emphasis>
|
||
apterous (
|
||
<figureCitation id="131F2A10FF8A325DFB76FCC7FB6BFCF0" box="[1180,1269,870,895]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
B). Ovipositor flattened laterally, without ornementation (
|
||
<taxonomicName id="4C244D16FF8A325DFD6EFC2BFB06FC2C" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[644,1176,906,931]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFD6EFC2BFC3AFC2C" box="[644,932,906,931]" italics="true" pageId="31" pageNumber="482">Upupagryllus subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFB4CFC2AFB76FC2C" box="[1190,1256,907,931]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
), or very small transverse crests (
|
||
<taxonomicName id="4C244D16FF8A325DFE8FFC0FFCDFFC48" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[357,833,942,967]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFE8FFC0FFDD3FC48" box="[357,589,942,967]" italics="true" pageId="31" pageNumber="482">Upupagryllus alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFCA5FC0EFC0EFC48" box="[847,912,943,967]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
). Female genitalia with a small, more or less cylindrical copulatory papilla, almost entirely sclerotized (
|
||
<figureCitation id="131F2A10FF8A325DFCCFFC73FCEBFC64" box="[805,885,978,1003]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="31" pageNumber="482">Fig. 13</figureCitation>
|
||
C–I, N–P).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E651EFF8A3261FF7DFC57FB76FD97" lastPageId="35" lastPageNumber="486" pageId="31" pageNumber="482" type="description">
|
||
<paragraph id="8B9B3695FF8A325DFF7DFC57FEB7FB80" blockId="31.[151,1437,258,2011]" box="[151,297,1014,1039]" pageId="31" pageNumber="482">
|
||
<emphasis id="B950EA87FF8A325DFF7DFC57FEB7FB80" bold="true" box="[151,297,1014,1039]" pageId="31" pageNumber="482">Description.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FF8A325DFF2DFBBBFCACF9A4" blockId="31.[151,1437,258,2011]" pageId="31" pageNumber="482">
|
||
Size small, body shape stout with short legs (
|
||
<figureCitation id="131F2A10FF8A325DFD2EFBBBFC8BFBBC" box="[708,789,1050,1075]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
A). Head and pronotum with many long and strong setae; legs with few, very long ones, body densely covered with small setae. Head small and vertical, the face oblique (
|
||
<figureCitation id="131F2A10FF8A325DFF75FBC3FF6AFBF4" box="[159,244,1122,1147]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
E). Fastigium rounded, not separated from the vertex, wider than the scape (
|
||
<figureCitation id="131F2A10FF8A325DFB9CFBC3FB54FBF4" box="[1142,1226,1122,1147]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
D, E). Ocelli well developed, set in a wide triangle, the distance between the median and one lateral ocelli somewhat smaller than the distance between the two lateral ocelli (
|
||
<figureCitation id="131F2A10FF8A325DFD89FB0BFD29FB4C" box="[611,695,1194,1219]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
F). Eyes only slightly protruding (
|
||
<figureCitation id="131F2A10FF8A325DFBA8FB0BFB08FB4C" box="[1090,1174,1194,1219]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
D). Scapes longer than wide. Maxillary palpi short; joint 5 little, but regularly widened toward apex, apex obliquely truncate (
|
||
<figureCitation id="131F2A10FF8A325DFAE5FB6FFAC1FB68" box="[1295,1375,1230,1255]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="31" pageNumber="482">Fig. 13</figureCitation>
|
||
A, J). Pronotum transverse, rounded (
|
||
<figureCitation id="131F2A10FF8A325DFE15FB53FDCDFA84" box="[511,595,1266,1291]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
D); lateral lobes quite short, clearly inflated anteriorly. TI with two long, ventral, apical spurs; tympana lacking (
|
||
<taxonomicName id="4C244D16FF8A325DFDB2FAB7FBF6FAA0" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[600,1128,1302,1327]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFDB2FAB7FCE8FAA0" box="[600,886,1302,1327]" italics="true" pageId="31" pageNumber="482">Upupagryllus subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFB9FFAB6FB2AFAA0" box="[1141,1204,1303,1327]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
), or present on each side (
|
||
<taxonomicName id="4C244D16FF8A325DFF3FFA9BFD30FADC" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[213,686,1338,1363]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFF3FFA9BFE25FADC" box="[213,443,1338,1363]" italics="true" pageId="31" pageNumber="482">Upupagryllus alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFD50FA9AFD67FADC" box="[698,761,1339,1363]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
); in the latter species, tympana small and obliterate, and TI not inflated. TII with three apical spurs, the outer dorsal spur lacking. TIII with three inner, and three outer apical spurs; inner spur: dorsal spur the longest, more than half basitarsomere III in length; outer spurs: shorter, the median the longest. TIII with four pairs of subapical spurs, not very long; inner spurs much longer than outer spurs, increasing in size toward TIII apex; outer spurs shorter, the first or the second the longest (see infra). TIII not serrulated between subapical spurs; few or no spines above subapical spurs. Basitarsomeres with only one row of dorsal spines, on outer side. Cerci not particularly elongate.
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FF8A325DFF2DF997FEDBF9FC" blockId="31.[151,1437,258,2011]" pageId="31" pageNumber="482">
|
||
<emphasis id="B950EA87FF8A325DFF2DF997FED1F9C0" bold="true" box="[199,335,1590,1615]" pageId="31" pageNumber="482">Coloration.</emphasis>
|
||
<figureCitation id="131F2A10FF8A325DFEB1F997FE2FF9C0" box="[347,433,1590,1615]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
. Light yellowish brown variegated with brown; setae on head and body dark brown. Tergites darker.
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FF8A325DFF2DF9DFFE44F8A8" blockId="31.[151,1437,258,2011]" pageId="31" pageNumber="482">
|
||
<emphasis id="B950EA87FF8A325DFF2DF9DFFE95F918" bold="true" box="[199,267,1662,1687]" pageId="31" pageNumber="482">Male.</emphasis>
|
||
Metanotum and tergites without glandular structures (
|
||
<figureCitation id="131F2A10FF8A325DFC9AF9DFFC5CF918" box="[880,962,1662,1687]" captionStart="FIGURE 12" captionStartId="32.[151,250,1795,1817]" captionTargetBox="[152,1434,199,1773]" captionTargetId="figure@32.[151,1436,193,1773]" captionTargetPageId="32" captionText="FIGURE 12. Upupagryllus subapterus Desutter-Grandcolas, n. sp., male holotype (A, D – H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm." httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="31" pageNumber="482">Fig. 12</figureCitation>
|
||
A). FWs either well-developed and with a stridulatory apparatus (
|
||
<taxonomicName id="4C244D16FF8A325DFE4EF903FC1BF934" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[420,901,1698,1723]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFE4EF903FD10F934" box="[420,654,1698,1723]" italics="true" pageId="31" pageNumber="482">Upupagryllus alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFC7FF902FC49F935" box="[917,983,1699,1722]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
,
|
||
<figureCitation id="131F2A10FF8A325DFC02F903FBA3F934" box="[1000,1085,1698,1723]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="31" pageNumber="482">Fig. 13</figureCitation>
|
||
K), or very short and without acoustic structure (
|
||
<taxonomicName id="4C244D16FF8A325DFE99F967FC18F950" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[371,902,1734,1759]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFE99F967FD0CF950" box="[371,658,1734,1759]" italics="true" pageId="31" pageNumber="482">Upupagryllus subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFC7FF966FC4AF950" box="[917,980,1735,1759]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
) (see infra, species description for FW characteristics); HWs vestigial or lacking, respectively. Supra anal plate without denticles. Subgenital plate short and high, V-truncate distally.
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FF8A325DFF2DF893FE6DF854" blockId="31.[151,1437,258,2011]" pageId="31" pageNumber="482">
|
||
<emphasis id="B950EA87FF8A325DFF2DF893FEE5F8C4" bold="true" box="[199,379,1842,1867]" pageId="31" pageNumber="482">Male genitalia.</emphasis>
|
||
<figureCitation id="131F2A10FF8A325DFE6FF893FE44F8C4" box="[389,474,1842,1867]" captionStart="FIGURE 14" captionStartId="34.[151,250,1842,1864]" captionTargetBox="[179,1423,202,1802]" captionTargetId="figure@34.[178,1427,192,1804]" captionTargetPageId="34" captionText="FIGURE 14. Upupagryllus Desutter-Grandcolas, n. gen., Male genitalia: A – C, U. subapterus Desutter-Grandcolas, n. sp.; D – F, U. alatus Desutter-Grandcolas, n. sp. in dorsal (A, D), ventral (B, E) and lateral (C, F) views. Abbreviations: see page 4." httpUri="https://zenodo.org/record/241871/files/figure.png" pageId="31" pageNumber="482">Fig. 14</figureCitation>
|
||
. Upper part of pseudepiphallic sclerite transverse; lower part with two long, distal, almost triangular acute processes; rami short, with a transverse apical or subapical apodeme. Pseudepiphallic parameres compact, projected dorsally as foliaceous sclerites (
|
||
<taxonomicName id="4C244D16FF8A325DFCBAF8DBFAD2F81C" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[848,1356,1914,1939]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="subalatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFCBAF8DBFBC2F81C" box="[848,1116,1914,1939]" italics="true" pageId="31" pageNumber="482">Upupagryllus subalatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFABDF8DAFA0DF81C" box="[1367,1427,1915,1939]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
) or not (
|
||
<taxonomicName id="4C244D16FF8A325DFF07F83FFD56F838" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[237,712,1950,1975]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="31" pageNumber="482" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FF8A325DFF07F83FFE4AF838" box="[237,468,1950,1975]" italics="true" pageId="31" pageNumber="482">Upupagryllus alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFF8A325DFD3FF83EFC8BF838" box="[725,789,1951,1975]" pageId="31" pageNumber="482" rank="species">n. sp.</taxonomicNameLabel>
|
||
). Endophallic sclerite narrow and more or less elongate. Endophallic apodeme variable.
|
||
</paragraph>
|
||
<caption id="DF5B661DFFB53262FF7DF8A2FD17F8D9" httpUri="https://zenodo.org/record/241869/files/figure.png" pageId="32" pageNumber="483" targetBox="[152,1434,199,1773]" targetPageId="32">
|
||
<paragraph id="8B9B3695FFB53262FF7DF8A2FD17F8D9" blockId="32.[151,1436,1795,1878]" pageId="32" pageNumber="483">
|
||
<emphasis id="B950EA87FFB53262FF7DF8A2FEBEF896" bold="true" box="[151,288,1795,1817]" pageId="32" pageNumber="483">FIGURE 12.</emphasis>
|
||
<taxonomicName id="4C244D16FFB53262FECDF8A2FC9AF896" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[295,772,1795,1817]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="32" pageNumber="483" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB53262FECDF8A2FDB7F896" box="[295,553,1795,1817]" italics="true" pageId="32" pageNumber="483">Upupagryllus subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB53262FCE5F8A5FCD6F896" box="[783,840,1796,1817]" pageId="32" pageNumber="483" rank="species">n. sp.</taxonomicNameLabel>
|
||
, male holotype (A, D–H) and female allotype (B, C, I, J): habitus in dorsal (A, B) and lateral (C) views; head in dorsal (D), lateral (E) and frontal (F) views; supra anal (G, I) and subgenital (H, J) plates. Scales for habitus 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5B661DFFB43263FF7DF894FCC6F849" httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="33" pageNumber="484" targetBox="[155,1423,205,1815]" targetPageId="33">
|
||
<paragraph id="8B9B3695FFB43263FF7DF894FCC6F849" blockId="33.[151,1436,1845,1990]" pageId="33" pageNumber="484">
|
||
<emphasis id="B950EA87FFB43263FF7DF894FE81F8C4" bold="true" box="[151,287,1845,1867]" pageId="33" pageNumber="484">FIGURE 13.</emphasis>
|
||
<taxonomicName id="4C244D16FFB43263FECFF894FD15F8C4" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[293,651,1845,1867]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B950EA87FFB43263FECFF894FE2FF8C4" box="[293,433,1845,1867]" italics="true" pageId="33" pageNumber="484">Upupagryllus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
, n. gen., A–I,
|
||
<taxonomicName id="4C244D16FFB43263FCF3F896FB1EF8C4" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[793,1152,1845,1868]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB43263FCF3F896FC38F8C4" box="[793,934,1845,1868]" italics="true" pageId="33" pageNumber="484">U. subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB43263FB60F897FB5CF8C4" box="[1162,1218,1846,1867]" pageId="33" pageNumber="484" rank="species">n. sp.</taxonomicNameLabel>
|
||
, A, maxillary palpus; B, apex of female ovipositor; C–I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn5, 23.v.1995 (C–E), female fn13, 17.v.1995 (F–H) and female fn19, 19.v.1995 (I). J–P,
|
||
<taxonomicName id="4C244D16FFB43263FBEAF8D5FAA7F806" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[1024,1337,1907,1929]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB43263FBEAF8D5FBC1F806" box="[1024,1119,1907,1929]" italics="true" pageId="33" pageNumber="484">U. alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB43263FAAFF8D5FA1EF806" box="[1349,1408,1908,1929]" pageId="33" pageNumber="484" rank="species">n. sp.</taxonomicNameLabel>
|
||
: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N–P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5B661DFFB73260FF7DF893FAECF8E4" httpUri="https://zenodo.org/record/241871/files/figure.png" pageId="34" pageNumber="485" targetBox="[179,1423,202,1802]" targetPageId="34">
|
||
<paragraph id="8B9B3695FFB73260FF7DF893FAECF8E4" blockId="34.[151,1436,1842,1899]" pageId="34" pageNumber="485">
|
||
<emphasis id="B950EA87FFB73260FF7DF893FEBEF8C7" bold="true" box="[151,288,1842,1864]" pageId="34" pageNumber="485">FIGURE 14.</emphasis>
|
||
<taxonomicName id="4C244D16FFB73260FECCF893FD12F8C7" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[294,652,1842,1864]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B950EA87FFB73260FECCF893FE2FF8C7" box="[294,433,1842,1864]" italics="true" pageId="34" pageNumber="485">Upupagryllus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
, n. gen., Male genitalia: A–C,
|
||
<taxonomicName id="4C244D16FFB73260FC2FF892FAB0F8C7" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[965,1326,1842,1864]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB73260FC2FF892FBCDF8C7" box="[965,1107,1842,1864]" italics="true" pageId="34" pageNumber="485">U. subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB73260FAD3F892FAECF8C7" box="[1337,1394,1843,1864]" pageId="34" pageNumber="485" rank="species">n. sp.</taxonomicNameLabel>
|
||
; D– F,
|
||
<taxonomicName id="4C244D16FFB73260FF45F8F7FE79F8E4" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[175,487,1877,1899]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="34" pageNumber="485" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB73260FF45F8F7FE92F8E4" box="[175,268,1877,1899]" italics="true" pageId="34" pageNumber="485">U. alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB73260FE1AF8F7FDB9F8E4" box="[496,551,1878,1899]" pageId="34" pageNumber="485" rank="species">n. sp.</taxonomicNameLabel>
|
||
in dorsal (A, D), ventral (B, E) and lateral (C, F) views. Abbreviations: see page 4.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B3695FFB63261FF2DFF36FC72FF77" blockId="35.[151,1436,151,536]" pageId="35" pageNumber="486">
|
||
<emphasis id="B950EA87FFB63261FF2DFF36FEBBFF3F" bold="true" box="[199,293,151,176]" pageId="35" pageNumber="486">Female.</emphasis>
|
||
Apterous. Subgenital plate transverse, distal margin more or less bisinuate. Ovipositor flattened laterally, without strong apical ornementation; apex elongate: apex of dorsal valve short and greatly widened basally, apex of ventral valve longer and more narrow; no notch (
|
||
<figureCitation id="131F2A10FFB63261FC98FF7EFC5DFF77" box="[882,963,223,248]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="35" pageNumber="486">Fig. 13</figureCitation>
|
||
M).
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FFB63261FF2DFEA5FEEAFEEB" blockId="35.[151,1436,151,536]" pageId="35" pageNumber="486">
|
||
<emphasis id="B950EA87FFB63261FF2DFEA5FE0BFE92" bold="true" box="[199,405,260,285]" pageId="35" pageNumber="486">Female genitalia.</emphasis>
|
||
Copulatory papilla small and more or less cylindrical (
|
||
<figureCitation id="131F2A10FFB63261FBFFFEA5FBF9FE93" box="[1045,1127,260,285]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="35" pageNumber="486">Fig. 13</figureCitation>
|
||
C–I, N–P), with (
|
||
<taxonomicName id="4C244D16FFB63261FAD9FEA4FE1FFECF" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="alatus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB63261FAD9FEA4FA02FE92" box="[1331,1436,260,285]" italics="true" pageId="35" pageNumber="486">U. alatus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB63261FE64FE89FE53FECF" box="[398,461,296,320]" pageId="35" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
|
||
) or without (
|
||
<taxonomicName id="4C244D16FFB63261FD82FE88FC65FECF" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[616,1019,295,320]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="species" species="subapterus" status="sp. nov.">
|
||
<emphasis id="B950EA87FFB63261FD82FE88FC96FECF" box="[616,776,295,320]" italics="true" pageId="35" pageNumber="486">U. subapterus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A26357FCFFB63261FBE3FE89FBD9FECF" box="[1033,1095,296,320]" pageId="35" pageNumber="486" rank="species">n. sp.</taxonomicNameLabel>
|
||
) a pair of membranous distal lobes (
|
||
<figureCitation id="131F2A10FFB63261FF0BFEEDFEAEFEEB" box="[225,304,332,357]" captionStart="FIGURE 13" captionStartId="33.[151,250,1845,1867]" captionTargetBox="[155,1423,205,1815]" captionTargetId="figure@33.[150,1436,193,1824]" captionTargetPageId="33" captionText="FIGURE 13. Upupagryllus Desutter-Grandcolas, n. gen., A – I, U. subapterus Desutter-Grandcolas, n. sp., A, maxillary palpus; B, apex of female ovipositor; C – I, female copulatory papilla, dorsal (C, F), ventral (D, G), lateral (E, H, I), in female fn 5, 23. v. 1995 (C – E), female fn 13, 17. v. 1995 (F – H) and female fn 19, 19. v. 1995 (I). J – P, U. alatus Desutter-Grandcolas, n. sp.: J, maxillary palpus; K, L, dorsal and lateral field of male holotype right FW; M, female subgenital plate; N – P, female copulatory papilla, dorsal (N), ventral (O) and lateral (P) views. Scale bars 1 mm." httpUri="https://zenodo.org/record/241870/files/figure.png" pageId="35" pageNumber="486">Fig. 13</figureCitation>
|
||
N–P).
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FFB63261FF2DFECEFEE0FE7A" blockId="35.[151,1436,151,536]" pageId="35" pageNumber="486">
|
||
<emphasis id="B950EA87FFB63261FF2DFECEFD89FE07" bold="true" box="[199,535,367,392]" pageId="35" pageNumber="486">Phylogenetic relationships.</emphasis>
|
||
Morphologically
|
||
<taxonomicName id="4C244D16FFB63261FCE8FECEFB03FE07" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[770,1181,367,392]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B950EA87FFB63261FCE8FECEFC02FE07" box="[770,924,367,392]" italics="true" pageId="35" pageNumber="486">Upupagryllus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
, n. gen. resembles
|
||
<taxonomicName id="4C244D16FFB63261FF7DFE35FE84FE22" box="[151,282,404,429]" class="Insecta" family="Phalangopsidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="subFamily" subFamily="Phaloriinae">Phaloriinae</taxonomicName>
|
||
, especially by TIII spines and spurs, and by the shape of the head. The shape of the fastigium, male FWs and/or stridulum, and male and female genitalia constitute as many apomorphies to define the new genus as a monophyletic entity.
|
||
</paragraph>
|
||
<paragraph id="8B9B3695FFB63261FF2DFE5EFB76FD97" blockId="35.[151,1436,151,536]" box="[199,1256,511,536]" pageId="35" pageNumber="486">
|
||
<emphasis id="B950EA87FFB63261FF2DFE5EFEB4FD97" bold="true" box="[199,298,511,536]" pageId="35" pageNumber="486">Habitat.</emphasis>
|
||
<taxonomicName id="4C244D16FFB63261FEDBFE5EFD22FD97" authority="Desutter-Grandcolas" authorityName="Desutter-Grandcolas" box="[305,700,511,536]" class="Insecta" family="Phalangopsidae" genus="Upupagryllus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Orthoptera" pageId="35" pageNumber="486" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B950EA87FFB63261FEDBFE5EFE55FD97" box="[305,459,511,536]" italics="true" pageId="35" pageNumber="486">Upupagryllus</emphasis>
|
||
Desutter-Grandcolas
|
||
</taxonomicName>
|
||
, n. gen. forages in the leaf litter of tropical forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |