147 lines
18 KiB
XML
147 lines
18 KiB
XML
<document id="E2AC89C144D35A70F99A88C9A321AA49" ID-CLB-Dataset="46266" ID-DOI="10.5281/zenodo.278832" ID-GBIF-Dataset="9d274a58-46e3-471c-8945-bbf84a9b1ba1" ID-ISSN="1175-5326" ID-Zenodo-Dep="278832" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460393709508" checkinUser="plazi" docAuthor="Mahony, Stephen" docDate="2011" docId="485787BFFFBBC324FF0BFB05FEBAF870" docLanguage="en" docName="zt03042p067.pdf" docOrigin="Zootaxa 3042" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Hemidactylus subtriedroides Annandale 1905" docType="treatment" docVersion="9" lastPageNumber="43" masterDocId="B46EFFC7FFBDC322FF9CFFCBFF87FFA9" masterDocTitle="Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae)" masterLastPageNumber="67" masterPageNumber="37" pageNumber="43" updateTime="1698298507063" updateUser="ExternalLinkService">
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<mods:title id="6E5B0BA06DCE3AFEEFC34985F1AC3F74">Taxonomic revision of Hemidactylus brookii Gray: a re-examination of the type series and some Asian synonyms, and a discussion of the obscure species Hemidactylus subtriedrus Jerdon (Reptilia: Gekkonidae)</mods:title>
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<mods:namePart id="E46D2B92F5FA9FA267C4BC2A16242DA4">Mahony, Stephen</mods:namePart>
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<mods:title id="27CFEDFC638CCA61AC5DCC79023B4B42">Zootaxa</mods:title>
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<mods:date id="D65CA71286B394718963A3882A3379C1">2011</mods:date>
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<mods:number id="736F42BB73E4296CB80171558BB3CEF1">3042</mods:number>
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<treatment id="485787BFFFBBC324FF0BFB05FEBAF870" ID-DOI="http://doi.org/10.5281/zenodo.5621720" ID-GBIF-Taxon="119417552" ID-Zenodo-Dep="5621720" LSID="urn:lsid:plazi:treatment:485787BFFFBBC324FF0BFB05FEBAF870" httpUri="http://treatment.plazi.org/id/485787BFFFBBC324FF0BFB05FEBAF870" lastPageNumber="43" pageId="6" pageNumber="43">
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<subSubSection id="88E46522FFBBC324FF0BFB05FD4EFB41" box="[151,713,1230,1256]" pageId="6" pageNumber="43" type="nomenclature">
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<paragraph id="C04136A9FFBBC324FF0BFB05FD4EFB41" blockId="6.[151,713,1230,1256]" box="[151,713,1230,1256]" pageId="6" pageNumber="43">
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<heading id="9B0981C5FFBBC324FF0BFB05FD4EFB41" bold="true" box="[151,713,1230,1256]" fontSize="11" level="1" pageId="6" pageNumber="43" reason="1">
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<taxonomicName id="07FE4D2AFFBBC324FF0BFB05FD4EFB41" authority="Annandale, 1905" authorityName="Annandale" authorityYear="1905" box="[151,713,1230,1256]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFBBC324FF0BFB05FD4EFB41" bold="true" box="[151,713,1230,1256]" pageId="6" pageNumber="43">
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<emphasis id="F28AEABBFFBBC324FF0BFB05FE68FB41" bold="true" box="[151,495,1230,1256]" italics="true" pageId="6" pageNumber="43">Hemidactylus subtriedroides</emphasis>
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Annandale, 1905
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="88E46522FFBBC324FF0BFADEFEBAF870" pageId="6" pageNumber="43" type="reference_group">
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<paragraph id="C04136A9FFBBC324FF0BFADEFEBAF870" blockId="6.[151,1437,1301,2009]" pageId="6" pageNumber="43">
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<taxonomicName id="07FE4D2AFFBBC324FF0BFADEFE5CFA84" box="[151,475,1301,1325]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFBBC324FF0BFADEFE5CFA84" box="[151,475,1301,1325]" italics="true" pageId="6" pageNumber="43">Hemidactylus subtriedroides</emphasis>
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</taxonomicName>
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was described from northern
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<collectingCountry id="B8E97639FFBBC324FCAFFADDFCF8FA84" box="[819,895,1302,1325]" name="Myanmar" pageId="6" pageNumber="43">Burma</collectingCountry>
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(=
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<collectingCountry id="B8E97639FFBBC324FC36FADEFB9EFA84" box="[938,1049,1301,1325]" name="Myanmar" pageId="6" pageNumber="43">Myanmar</collectingCountry>
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) based on two male specimens in the collection of the ZSI, which were previously misidentified as
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<taxonomicName id="07FE4D2AFFBBC324FCE2FAF1FB92FAF8" box="[894,1045,1337,1361]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="maculatus">
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<emphasis id="F28AEABBFFBBC324FCE2FAF1FB92FAF8" box="[894,1045,1337,1361]" italics="true" pageId="6" pageNumber="43">H. maculatus</emphasis>
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</taxonomicName>
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by
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<bibRefCitation id="A46F4B58FFBBC324FBDDFAF2FA84FAF8" author="Anderson" box="[1089,1283,1337,1361]" pageId="6" pageNumber="43" refString="Anderson, J. (1879) Anatomical and Zoological Researches: Comprising an Account of the Zoological Results of the Two Expeditions to Western Yunnan in 1866 and 1875; and a Monograph of the Two Cetacean Genera Platanista and Orcella. Bernard Quaritch, London " 1878 ". Two volumes (Text: 985 pp. [herpetology: pages 703 - 860, 969 - 975]; Atlas: 85 plates [herpetological plates 55 - 78, 75 A, 75 B])." type="book chapter" year="1879">Anderson (1879)</bibRefCitation>
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, the collector of the
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<typeStatus id="1F45880BFFBBC324FF7FFA95FECEFADF" box="[227,329,1374,1398]" pageId="6" pageNumber="43" type="syntype">syntypes</typeStatus>
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. One specimen was transferred to BMNH in 1906. Annandale’s short description was composed primarily of comparing
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<taxonomicName id="07FE4D2AFFBBC324FE3EFA49FDE2FA30" box="[418,613,1409,1433]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFBBC324FE3EFA49FDE2FA30" box="[418,613,1409,1433]" italics="true" pageId="6" pageNumber="43">H. subtriedroides</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="07FE4D2AFFBBC324FD38FA49FCC2FA30" box="[676,837,1409,1433]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="subtriedrus">
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<emphasis id="F28AEABBFFBBC324FD38FA49FCC2FA30" box="[676,837,1409,1433]" italics="true" pageId="6" pageNumber="43">H. subtriedrus</emphasis>
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</taxonomicName>
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, a species which he considered to be the most similar in appearance. He even acknowledged
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<taxonomicName id="07FE4D2AFFBBC324FDD5FA6DFD3DFA14" box="[585,698,1445,1469]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFBBC324FDD5FA6DFD3DFA14" box="[585,698,1445,1469]" italics="true" pageId="6" pageNumber="43">H. brookii</emphasis>
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</taxonomicName>
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, vaguely stating that his new species differed by possessing a flatter and broader tail base, and that “The bulk is greater than that of
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<taxonomicName id="07FE4D2AFFBBC324FC17FA01FB86FA48" box="[907,1025,1481,1505]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="67" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFBBC324FC17FA01FB86FA48" box="[907,1025,1481,1505]" italics="true" pageId="6" pageNumber="43">H. Brookii</emphasis>
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</taxonomicName>
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Gray, the dorsal tubercles are larger and the whole animal is more depressed. The proportions are also different.” Annandale was mostly correct in this statement if considering
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<taxonomicName id="07FE4D2AFFBBC324FE36F9D9FD9BF980" box="[426,540,1553,1577]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" sensu="stricto" species="brookii">
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<emphasis id="F28AEABBFFBBC324FE36F9D9FE39F980" box="[426,446,1554,1577]" italics="true" pageId="6" pageNumber="43">H</emphasis>
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.
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<emphasis id="F28AEABBFFBBC324FE50F9DAFD9BF980" box="[460,540,1553,1577]" italics="true" pageId="6" pageNumber="43">brookii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="E9B957C0FFBBC324FDBFF9D9FDC1F980" box="[547,582,1554,1577]" pageId="6" pageNumber="43" sensu="stricto">s.s.</taxonomicNameLabel>
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, however due to the subjectivity of his comparison without quantifiable evidence, and perhaps more importantly, because one of the
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<typeStatus id="1F45880BFFBBC324FC86F9FDFCFBF9E7" box="[794,892,1590,1614]" pageId="6" pageNumber="43" type="syntype">syntypes</typeStatus>
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of
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<emphasis id="F28AEABBFFBBC324FC3EF9FDFBBCF9E4" box="[930,1083,1589,1613]" italics="true" pageId="6" pageNumber="43">
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<taxonomicName id="07FE4D2AFFBBC324FC3EF9FDFB93F9E4" box="[930,1044,1589,1613]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" sensu="lato" species="brookii">H. brookii</taxonomicName>
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<taxonomicNameLabel id="E9B957C0FFBBC324FB87F9FEFBBCF9E4" box="[1051,1083,1589,1613]" pageId="6" pageNumber="43" sensu="lato">s.l.</taxonomicNameLabel>
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</emphasis>
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represented this species, it was later synonymised without discussion (
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<bibRefCitation id="A46F4B58FFBBC324FDD2F992FD51F9D8" author="Smith" box="[590,726,1625,1649]" pageId="6" pageNumber="43" refString="Smith, M. A. (1935) The Fauna of British India, Including Ceylon and Burma. Reptilia and Amphibia. Vol. II. - Sauria. Taylor and Francis, London. xiii + 440 pp., 2 folding maps, 1 pl." type="book" year="1935">Smith 1935</bibRefCitation>
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).
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<bibRefCitation id="A46F4B58FFBBC324FD76F992FC37F9D8" author="Loveridge" box="[746,944,1625,1649]" pageId="6" pageNumber="43" refString="Loveridge, A. (1941) Certain Afro-American geckos of the genus Hemidactylus. Copeia, 4, 245 - 248." type="journal article" year="1941">Loveridge (1941)</bibRefCitation>
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appeared to recognise that this species was not typical of his concept of
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<taxonomicName id="07FE4D2AFFBBC324FE75F9B5FDD9F93C" box="[489,606,1661,1685]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="brookii">
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<emphasis id="F28AEABBFFBBC324FE75F9B5FDD9F93C" box="[489,606,1661,1685]" italics="true" pageId="6" pageNumber="43">H. brookii</emphasis>
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</taxonomicName>
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and elevated it from synonymy to subspecific status as
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<taxonomicName id="07FE4D2AFFBBC324FB71F9B5FE96F910" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="subSpecies" species="brookii" subSpecies="subtriedroides">
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<emphasis id="F28AEABBFFBBC324FB71F9B5FA86F93C" box="[1261,1281,1662,1685]" italics="true" pageId="6" pageNumber="43">H</emphasis>
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.
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<emphasis id="F28AEABBFFBBC324FA8EF9B6FE96F910" italics="true" pageId="6" pageNumber="43">brookii subtriedroides</emphasis>
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</taxonomicName>
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, but also provided no justification for his action. Having examined both
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<typeStatus id="1F45880BFFBBC324FBDDF969FB24F913" box="[1089,1187,1698,1722]" pageId="6" pageNumber="43" type="syntype">syntypes</typeStatus>
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as part of this study it is clear that this is a distinct species from
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<emphasis id="F28AEABBFFBBC324FDF3F90DFC8AF974" box="[623,781,1733,1757]" italics="true" pageId="6" pageNumber="43">
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<taxonomicName id="07FE4D2AFFBBC324FDF3F90DFD65F974" box="[623,738,1733,1757]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" sensu="stricto" species="brookii">H. brookii</taxonomicName>
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<taxonomicNameLabel id="E9B957C0FFBBC324FD76F90DFC8AF974" box="[746,781,1734,1757]" pageId="6" pageNumber="43" sensu="stricto">s.s.</taxonomicNameLabel>
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</emphasis>
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It is here redescribed based on the BMNH
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<typeStatus id="1F45880BFFBBC324FB65F90DFAD7F977" box="[1273,1360,1734,1758]" pageId="6" pageNumber="43" type="syntype">syntype</typeStatus>
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which is designated as the
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<typeStatus id="1F45880BFFBBC324FEEBF922FE5AF8A8" box="[375,477,1769,1793]" pageId="6" pageNumber="43" type="lectotype">lectotype</typeStatus>
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in accordance with Article 75.1 of the
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<emphasis id="F28AEABBFFBBC324FC0FF922FC49F8A8" box="[915,974,1769,1793]" italics="true" pageId="6" pageNumber="43">Code</emphasis>
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. The figure of
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<taxonomicName id="07FE4D2AFFBBC324FBEAF922FF3AF88C" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="subtriedroides">
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<emphasis id="F28AEABBFFBBC324FBEAF922FF3AF88C" italics="true" pageId="6" pageNumber="43">Hemidactylus subtriedroides</emphasis>
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</taxonomicName>
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published soon after the original description (
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<bibRefCitation id="A46F4B58FFBBC324FD26F8C6FB8AF88C" author="Annandale" box="[698,1037,1805,1829]" pageId="6" pageNumber="43" refString="Annandale, N. (1905 b) Contributions to Oriental Herpetology. Supplement II. Notes on the Oriental lizards in the Indian Museum, with a list of the species recorded from British India and Ceylon. Journal Proceedings of the Asiatic Society Bengal, 1 (3), 81 - 93 + Pl. I - II." type="journal article" year="1905" yearSuffix="b">
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Annandale 1905b:Pl. II,
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<figureCitation id="58C52A2CFFBBC324FC52F8C6FB8AF88C" box="[974,1037,1805,1829]" captionStart="FIGURE 1" captionStartId="5.[151,250,1169,1192]" captionTargetBox="[151,1436,447,1147]" captionTargetId="figure@5.[151,1436,447,1147]" captionTargetPageId="5" captionText="FIGURE 1. Historical figures of the Hemidactylus brookii group taxa: A. H. brookii adult male lectotype (BMNH 1947.3.6.47) by present designation, from Gray (1867: Pl. 15, Fig. 2), later reprinted in Gray (1875: Pl. 15, Fig. 2); B. H. subtriedroides adult male paralectotype (ZSI 4135) by present designation, from Annandale (1905 b: Pl. II, Fig. 1); C. H. gleadowi not based on the original type series, from Boulenger (1890: 86, Fig. 27), later reprinted as H. brookii (Boulenger 1912: 42, Fig. 14; Smith 1935: 90, Fig. 28); D. H. gleadowi a syntype from Murray (1884 a: Fig. 3)." httpUri="https://zenodo.org/record/278833/files/figure.png" pageId="6" pageNumber="43">Fig. 1</figureCitation>
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</bibRefCitation>
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) clearly depicts a specimen with an original tail, and thus is likely based on the
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<typeStatus id="1F45880BFFBBC324FD1AF8FAFC9AF8E0" box="[646,797,1841,1865]" pageId="6" pageNumber="43" type="paralectotype">paralectotype</typeStatus>
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specimen designated here (ZSI 4135:
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<figureCitation id="58C52A2CFFBBC324FB4DF8FAFA93F8E0" box="[1233,1300,1841,1865]" captionStart="FIGURE 2. A" captionStartId="12.[151,250,1676,1699]" captionTargetBox="[151,1436,337,1655]" captionTargetId="figure@12.[151,1436,337,1655]" captionTargetPageId="12" captionText="FIGURE 2. A. Hemidactylus brookii adult male lectotype (BMNH 1947.3.6.47) by present designation; B. H. gleadowi adult male neotype (BMNH [18] 84.7.25.8) by present designation; C. H. subtriedroides adult male lectotype (BMNH 1946.8.25.54) by present designation; D. H. kushmorensis adult male neotype (BMNH [18] 87.9.22.8) by present designation. E & F. Ventral view of the original tails showing the difference of subcaudal proportions and tail shape between the “ litter-dwelling ” and “ rock-dwelling ” morphotypes: E. H. kushmorensis adult male neotype (BMNH [18] 87.9.22.8); F. H. tenkatei adult male (BMNH 1926.10.30.47) from Timor." httpUri="https://zenodo.org/record/278834/files/figure.png" pageId="6" pageNumber="43">Fig. 2</figureCitation>
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B). The reason for choosing the BMNH specimen as the
|
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<typeStatus id="1F45880BFFBBC324FD3BF89EFC96F8C4" box="[679,785,1877,1901]" pageId="6" pageNumber="43" type="lectotype">lectotype</typeStatus>
|
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, is that the preservation of the ZSI
|
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<typeStatus id="1F45880BFFBBC324FB36F89DFA86F8C7" box="[1194,1281,1878,1902]" pageId="6" pageNumber="43" type="syntype">syntype</typeStatus>
|
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is of a lower quality, with signs that this specimen is now, or was in the past not maintained in 70% ethanol, leading to minor soft tissue degradation. See subsequent section of
|
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<taxonomicName id="07FE4D2AFFBBC324FD56F855FCC2F81C" box="[714,837,1949,1973]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="43" phylum="Chordata" rank="species" species="tenkatei">
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<emphasis id="F28AEABBFFBBC324FD56F855FCC2F81C" box="[714,837,1949,1973]" italics="true" pageId="6" pageNumber="43">H. tenkatei</emphasis>
|
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</taxonomicName>
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for further discussion regarding the taxonomic status of this species.
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</paragraph>
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</subSubSection>
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</treatment>
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||
</document> |