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<document id="69B84EF5A492EC6254A9670BD4C6EAB3" ID-DOI="10.13133/2284-4880/1526" ID-ISSN="2284-4880" ID-Zenodo-Dep="12763176" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721241101321" checkinUser="felipe" docAuthor="Reding, Jean-Paul G." docDate="2023" docId="482C87DD3151A85AFF3FFC3FFBA6F928" docLanguage="en" docName="FragEntomol.55.2.221-262.pdf" docOrigin="Fragmenta entomologica (Frag. Entomol., Roma) 55 (2)" docSource="http://dx.doi.org/10.13133/2284-4880/1526" docStyle="DocumentStyle:8499FF4308F512386C0E43DC290A16BD.1:FragEntomol.2021-.journal_article.0cover" docStyleId="8499FF4308F512386C0E43DC290A16BD" docStyleName="FragEntomol.2021-.journal_article.0cover" docStyleVersion="1" docTitle="Perla pyrenaica Reding, 2023, sp. n." docType="treatment" docUuid="7F837839-7A51-475E-9CC7-A4D528F09E6E" docUuidSource="ZooBank" docVersion="2" lastPageNumber="229" masterDocId="B415FFA53159A852FFA8FFABFFFAFFC4" masterDocTitle="Steps towards a revision of the Perla bipunctata Pictet, 1833 species complex (Plecoptera: Perlidae)" masterLastPageNumber="262" masterPageNumber="221" pageNumber="229" updateTime="1721242779436" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
<mods:mods id="65D17E5F32705F9B15BCF62522B5ECB0" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="4C46834C8A749685D2C3C31D9B84AA31">Steps towards a revision of the Perla bipunctata Pictet, 1833 species complex (Plecoptera: Perlidae)</mods:title>
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<mods:name id="9C7F65EBF4C1F8EEB9152CB912B87B3C" type="personal">
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<mods:namePart id="237F7720A00CD044ADDBECFFA7A8780C">Reding, Jean-Paul G.</mods:namePart>
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<mods:title id="7AA67442C6D7A0564C96ED166CC79D74">Fragmenta entomologica</mods:title>
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<mods:date id="68BD3F68DB742E23648D19FCD0FE41B9">2023</mods:date>
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<mods:title id="7D19FD02B2272368CFADA6B907F6A416">Frag. Entomol., Roma</mods:title>
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<mods:number id="8C917B433D55856FAC5888099156CBED">2023-12-21</mods:number>
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<mods:classification id="8259553D0F75D126EAD3244E23D8E6C9">journal article</mods:classification>
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<treatment id="482C87DD3151A85AFF3FFC3FFBA6F928" LSID="urn:lsid:zoobank.org:act:7F837839-7A51-475E-9CC7-A4D528F09E6E" httpUri="http://treatment.plazi.org/id/482C87DD3151A85AFF3FFC3FFBA6F928" lastPageNumber="229" pageId="8" pageNumber="229">
<subSubSection id="889F65403151A85AFF3FFC3FFE72FC68" box="[151,392,916,940]" pageId="8" pageNumber="229" type="nomenclature">
<paragraph id="C03A36CB3151A85AFF3FFC3FFE72FC68" blockId="8.[151,772,884,1036]" box="[151,392,916,940]" pageId="8" pageNumber="229">
<heading id="9B7281A73151A85AFF3FFC3FFE72FC68" bold="true" box="[151,392,916,940]" fontSize="10" level="2" pageId="8" pageNumber="229" reason="3">
<emphasis id="F2F1EAD93151A85AFF3FFC3FFE72FC68" bold="true" box="[151,392,916,940]" pageId="8" pageNumber="229">
<taxonomicName id="07854D483151A85AFF3FFC3FFEBFFC68" ID-CoL="CDV5P" authorityName="Reding" authorityYear="2023" box="[151,325,916,940]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="pyrenaica" status="sp. nov.">
<emphasis id="F2F1EAD93151A85AFF3FFC3FFEBFFC68" bold="true" box="[151,325,916,940]" italics="true" pageId="8" pageNumber="229">Perla pyrenaica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="E9C257A23151A85AFEE4FC3FFE72FC68" box="[332,392,916,940]" pageId="8" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="889F65403151A85AFF3FFC1FFEB1FBC8" pageId="8" pageNumber="229" type="description">
<paragraph id="C03A36CB3151A85AFF3FFC1FFEB1FBC8" blockId="8.[151,772,884,1036]" pageId="8" pageNumber="229">
(
<figureCitation id="58BE2A4E3151A85AFF37FC1FFEE3FC08" box="[159,281,948,972]" captionStart="Plate I" captionStartId="14.[151,197,1805,1824]" captionTargetBox="[249,1342,166,1766]" captionTargetId="figure-10@14.[247,1342,166,1766]" captionTargetPageId="14" captionText="Plate I, Figs A.1–A.9 – Type A (Italian Alps and Apennines). A.1, hemitergal lobes, mesal field with sensilla and sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.2, sclerotized apex of everted aedeagus of an adult ♂ (Staffora); A.3, sclerotized apex of extracted aedeagus of an adult ♂ (Stura di Ala); A.4, sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.5, aedeagal tube of adult ♂; A.6, aedeagal tube and hemitergal lobes of adult ♂; A.7, adult ♂, hemitergal lobes and mesal field with sensilla (Southern Apennines); A.8, sclerotized apex of extracted aedeagus of an adult ♂ (Southern Apennines); A.9, adult ♂, hemitergal lobes and mesal field with sensilla, dorsal view (Stura di Demonte)." figureDoi="http://doi.org/10.5281/zenodo.12763178" httpUri="https://zenodo.org/record/12763178/files/figure.png" pageId="8" pageNumber="229">Figs E.1–E</figureCitation>
.15)
<uri id="B4143AC93151A85AFF3FFC7FFEB1FBC8" pageId="8" pageNumber="229">
urn:lsid:zoobank.org:act:
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</paragraph>
</subSubSection>
<subSubSection id="889F65403151A85AFF3FFBFFFEDCFB28" pageId="8" pageNumber="229" type="reference_group">
<paragraph id="C03A36CB3151A85AFF3FFBFFFE01FB48" blockId="8.[151,771,1108,1260]" pageId="8" pageNumber="229">
<treatmentCitationGroup id="E09511E53151A85AFF3FFBFFFE01FB48" pageId="8" pageNumber="229">
<taxonomicName id="07854D483151A85AFF3FFBFFFECEFBA8" ID-CoL="8N45N" authorityName="sensu Klapalek" authorityYear="1923" box="[151,308,1108,1132]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="maxima">
<emphasis id="F2F1EAD93151A85AFF3FFBFFFECEFBA8" box="[151,308,1108,1132]" italics="true" pageId="8" pageNumber="229">Perla maxima</emphasis>
</taxonomicName>
;
<treatmentCitation id="412410DA3151A85AFEE1FBFEFDE6FBA8" author="Despax R." box="[329,540,1108,1132]" page="190" pageId="8" pageNumber="229" year="1951">
<bibRefCitation id="A4144B3A3151A85AFEE1FBFEFDE6FBA8" author="Despax R." box="[329,540,1108,1132]" pageId="8" pageNumber="229" refId="ref21706" refString="Despax R. 1951. Plecopteres. P. Lechevalier, Paris. 280 pp. (Faune de France v. 55) Available from: http: // faunedefrance. org / bibliotheque / docs / R. DESPAX (FdeFr 55) Plecopteres. pdf (accessed April 20 th 2023)" type="book" year="1951">Despax 1951: 190</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="412410DA3151A85AFD86FBFFFF03FB48" author="Bertrand H. &amp; Aubert J." page="96" pageId="8" pageNumber="229" year="1952">
<bibRefCitation id="A4144B3A3151A85AFD86FBFFFF03FB48" author="Bertrand H. &amp; Aubert J." pageId="8" pageNumber="229" pagination="93 - 106" refId="ref20985" refString="Bertrand H., Aubert J. 1952. Recoltes de Plecopteres dans les Pyrenees. Annales de la Societe entomologique de France, 121: 93 - 106." type="journal article" year="1952">Bertrand &amp; Aubert 1952: 96</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="A4144B3A3151A85AFEABFBDFFEC1FB48" author="Bertrand H. &amp; Aubert J." box="[259,315,1140,1164]" pageId="8" pageNumber="229" pagination="146 - 149" refId="ref21015" refString="Bertrand H., Aubert J. 1955. Recoltes de Plecopteres dans les Pyrenees (Note complementaire). Bulletin de la Societe entomologique de France, 60 (9 / 10): 146 - 149. Available from: https: // www. persee. fr / doc / bsef _ 0037 - 928 x _ 1955 _ num _ 60 _ 9 _ 18796 (accessed April 20 th 2023)" type="journal article" year="1955">1955</bibRefCitation>
;
<treatmentCitation id="412410DA3151A85AFEEDFBDFFE0DFB48" author="Aubert J." box="[325,503,1140,1164]" page="79" pageId="8" pageNumber="229" year="1963">
<bibRefCitation id="A4144B3A3151A85AFEEDFBDFFE0DFB48" author="Aubert J." box="[325,503,1140,1164]" pageId="8" pageNumber="229" pagination="23 - 107" refId="ref20812" refString="Aubert J. 1963. Les Plecopteres de la Peninsule Iberique. Eos, 39: 23 - 107. Available from: http: // hdl. handle. net / 10261 / 158133 (accessed April 20 th 2023)" type="journal article" year="1963">Aubert 1963: 79</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="C03A36CB3151A85AFF3FFB3FFEDCFB28" blockId="8.[151,771,1108,1260]" pageId="8" pageNumber="229">
<treatmentCitationGroup id="E09511E53151A85AFF3FFB3FFEDCFB28" pageId="8" pageNumber="229">
<taxonomicName id="07854D483151A85AFF3FFB3FFED4FB68" ID-CoL="775K4" authorityName=", sensu Sivec &amp; Stark" authorityYear="2002" box="[151,302,1172,1196]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="F2F1EAD93151A85AFF3FFB3FFED4FB68" box="[151,302,1172,1196]" italics="true" pageId="8" pageNumber="229">Perla grandis</emphasis>
</taxonomicName>
;
<treatmentCitation id="412410DA3151A85AFEE8FB3FFD69FB68" author="Berthelemy C. &amp; Laur C." box="[320,659,1172,1196]" page="278" pageId="8" pageNumber="229" year="1975">
<bibRefCitation id="A4144B3A3151A85AFEE8FB3FFD69FB68" author="Berthelemy C. &amp; Laur C." box="[320,659,1172,1196]" pageId="8" pageNumber="229" pagination="263 - 285" refId="ref20916" refString="Berthelemy C., Laur C. 1975. Plecopteres et Coleopteres aquatiques du Lot (Massif central Francais). Annales de Limnologie, 11 (3): 263 - 285. Available from: https: // www. limnology-journal. org / articles / limn / pdf / 1975 / 03 / limn 1975113 p 263. pdf (accessed April 20 th 2023)" type="journal article" year="1975">Berthélemy &amp; Laur 1975: 278</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="412410DA3151A85AFD0AFB3FFEADFB08" author="Oscoz J. &amp; Duran C." page="188" pageId="8" pageNumber="229" year="2004">
<bibRefCitation id="A4144B3A3151A85AFD0AFB3FFEADFB08" author="Oscoz J. &amp; Duran C." pageId="8" pageNumber="229" pagination="183 - 196" refId="ref24202" refString="Oscoz J., Duran C. 2004. Contribucion al conocimiento de los plecopteros (Insecta: Plecoptera) en la cuenca del Ebro = Contribution to the knowledge of stoneflies (Insecta: Plecoptera) in the Ebro River Basin. Munibe Ciencias Naturales. Natur zientziak, 55: 183 - 196." type="journal article" year="2004">Oscoz &amp; Durán 2004: 188</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="412410DA3151A85AFECDFB1FFD05FB08" author="Tierno de Figueroa J. M. &amp; Sanchez-Ortega A. &amp; Membiela-Iglesias P. &amp; Luzon-Ortega J. M." box="[357,767,1204,1228]" page="112" pageId="8" pageNumber="229" year="2003">
<bibRefCitation id="A4144B3A3151A85AFECDFB1FFD05FB08" author="Tierno de Figueroa J. M. &amp; Sanchez-Ortega A. &amp; Membiela-Iglesias P. &amp; Luzon-Ortega J. M." box="[357,767,1204,1228]" pageId="8" pageNumber="229" pagination="4000 - 8198" refId="ref25534" refString="Tierno de Figueroa J. M., Sanchez-Ortega A., Membiela-Iglesias P., Luzon-Ortega J. M. 2003. Plecoptera. In: Ramos, M. A., Alba-Tercedor, J., Belles, X., Gosalbez, J., Guerra, A., Macpherson, E., Serrano, J. &amp; Templado, J. (Eds.), Fauna Iberica. Vol. 22. Museo Nacional de Ciencias Naturales. CSIC, Madrid, 404 pp. ISBN 978 - 8 - 4000 - 8198 - 0" type="book chapter" year="2003">Tierno de Figueroa et al. (2003: 112</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="A4144B3A3151A85AFF3FFB7FFF27FB28" author="Tierno de Figueroa, J. M. &amp; Luzon-Ortega, J. M. &amp; Lopez-Rodriguez, M. J." box="[151,221,1236,1260]" pageId="8" pageNumber="229" refId="ref25806" refString="Tierno de Figueroa, J. M.; Luzon-Ortega, J. M.; Lopez-Rodriguez, M. J. 2015 b. Species Perla grandis. Rambur, 1842. En: IBERFAUNA. El Banco de Datos de la Fauna Iberica. Museo Nacional de Ciencias Naturales (CSIC). Available from: http: // iberfauna. mncn. csic. es / showficha. aspx? rank = T &amp; id- tax = 26040 (accessed April 20 th 2023)." type="url" year="2015">2015b</bibRefCitation>
;
<bibRefCitation id="A4144B3A3151A85AFF40FB7FFEE5FB28" author="Tierno de Figueroa J. M. &amp; Luzon-Ortega J. M. &amp; Lopez-Rodriguez M. J." box="[232,287,1236,1260]" pageId="8" pageNumber="229" refId="ref25895" refString="Tierno de Figueroa J. M., Luzon-Ortega J. M., Lopez-Rodriguez M. J. 2018. Checklist de Fauna Iberica. Orden Plecoptera (Arthropoda: Insecta) en la peninsula iberica e islas Baleares (edicion 2018). In: Ramos, M. A. &amp; Sanchez Ruiz (Eds.), Documentos Fauna Iberica, 5. Museo Nacional de Ciencias Naturales, CSIC. Madrid: 2 (sn) + 15 pp." type="book" year="2018">2018</bibRefCitation>
)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="889F65403151A85AFF3FFABFFE7DF989" pageId="8" pageNumber="229" type="diagnosis">
<paragraph id="C03A36CB3151A85AFF3FFABFFE7DF989" blockId="8.[151,772,1300,1932]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFF3FFABFFE4CFAE8" bold="true" box="[151,438,1300,1324]" pageId="8" pageNumber="229">Morphological diagnosis.</emphasis>
Males and females macropterous. Body length of males
<quantity id="077D9B2E3151A85AFE68FA9FFDC3FA88" box="[448,569,1332,1356]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.6" metricValueMin="1.8" pageId="8" pageNumber="229" unit="mm" value="22.0" valueMax="26.0" valueMin="18.0">18–26 mm</quantity>
; of females
<quantity id="077D9B2E3151A85AFD68FA9FFF3EFAA8" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.6" metricValueMin="2.0" pageId="8" pageNumber="229" unit="mm" value="28.0" valueMax="36.0" valueMin="20.0">20–36 mm</quantity>
. Body colour dark brown. Head mostly yellow with tawny markings; M-line clearly delineated (
<figureCitation id="58BE2A4E3151A85AFDC1FADFFD3BFA48" box="[617,705,1396,1420]" captionStart="Plate I" captionStartId="14.[151,197,1805,1824]" captionTargetBox="[249,1342,166,1766]" captionTargetId="figure-10@14.[247,1342,166,1766]" captionTargetPageId="14" captionText="Plate I, Figs A.1–A.9 – Type A (Italian Alps and Apennines). A.1, hemitergal lobes, mesal field with sensilla and sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.2, sclerotized apex of everted aedeagus of an adult ♂ (Staffora); A.3, sclerotized apex of extracted aedeagus of an adult ♂ (Stura di Ala); A.4, sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.5, aedeagal tube of adult ♂; A.6, aedeagal tube and hemitergal lobes of adult ♂; A.7, adult ♂, hemitergal lobes and mesal field with sensilla (Southern Apennines); A.8, sclerotized apex of extracted aedeagus of an adult ♂ (Southern Apennines); A.9, adult ♂, hemitergal lobes and mesal field with sensilla, dorsal view (Stura di Demonte)." figureDoi="http://doi.org/10.5281/zenodo.12763178" httpUri="https://zenodo.org/record/12763178/files/figure.png" pageId="8" pageNumber="229">Fig. E.1</figureCitation>
). Pronotum pale yellow, with a large dark central band flanked with a symmetrical darker design (
<figureCitation id="58BE2A4E3151A85AFDA6FA1FFD9CFA08" box="[526,614,1460,1484]" captionStart="Plate I" captionStartId="14.[151,197,1805,1824]" captionTargetBox="[249,1342,166,1766]" captionTargetId="figure-10@14.[247,1342,166,1766]" captionTargetPageId="14" captionText="Plate I, Figs A.1–A.9 – Type A (Italian Alps and Apennines). A.1, hemitergal lobes, mesal field with sensilla and sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.2, sclerotized apex of everted aedeagus of an adult ♂ (Staffora); A.3, sclerotized apex of extracted aedeagus of an adult ♂ (Stura di Ala); A.4, sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.5, aedeagal tube of adult ♂; A.6, aedeagal tube and hemitergal lobes of adult ♂; A.7, adult ♂, hemitergal lobes and mesal field with sensilla (Southern Apennines); A.8, sclerotized apex of extracted aedeagus of an adult ♂ (Southern Apennines); A.9, adult ♂, hemitergal lobes and mesal field with sensilla, dorsal view (Stura di Demonte)." figureDoi="http://doi.org/10.5281/zenodo.12763178" httpUri="https://zenodo.org/record/12763178/files/figure.png" pageId="8" pageNumber="229">Fig. E.1</figureCitation>
). Inner half of the two lower ocelli circled by a narrow black band (
<figureCitation id="58BE2A4E3151A85AFD70FA7FFF41F9C8" captionStart="Plate I" captionStartId="14.[151,197,1805,1824]" captionTargetBox="[249,1342,166,1766]" captionTargetId="figure-10@14.[247,1342,166,1766]" captionTargetPageId="14" captionText="Plate I, Figs A.1–A.9 – Type A (Italian Alps and Apennines). A.1, hemitergal lobes, mesal field with sensilla and sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.2, sclerotized apex of everted aedeagus of an adult ♂ (Staffora); A.3, sclerotized apex of extracted aedeagus of an adult ♂ (Stura di Ala); A.4, sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.5, aedeagal tube of adult ♂; A.6, aedeagal tube and hemitergal lobes of adult ♂; A.7, adult ♂, hemitergal lobes and mesal field with sensilla (Southern Apennines); A.8, sclerotized apex of extracted aedeagus of an adult ♂ (Southern Apennines); A.9, adult ♂, hemitergal lobes and mesal field with sensilla, dorsal view (Stura di Demonte)." figureDoi="http://doi.org/10.5281/zenodo.12763178" httpUri="https://zenodo.org/record/12763178/files/figure.png" pageId="8" pageNumber="229">Fig. E.1</figureCitation>
) on most, and by a large black band (Fig. E.2) on some specimens. This
<typeStatus id="1F3E88693151A85AFEE4F9BEFE80F9E9" box="[332,378,1557,1581]" pageId="8" pageNumber="229">type</typeStatus>
corresponds to putative
<taxonomicName id="07854D483151A85AFD2DF9BFFF4CF988" authorityName=", sensu Sivec &amp; Stark" authorityYear="2002" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="F2F1EAD93151A85AFD2DF9BFFF4CF988" italics="true" pageId="8" pageNumber="229">Perla grandis</emphasis>
</taxonomicName>
from the Pyrenees.
</paragraph>
</subSubSection>
<subSubSection id="889F65403151A85AFF15F9FFFAADF9C8" pageId="8" pageNumber="229" type="materials_examined">
<paragraph id="C03A36CB3151A85AFF15F9FFFD97F968" blockId="8.[151,772,1300,1932]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFF15F9FFFEA6F9A8" bold="true" box="[189,348,1620,1644]" pageId="8" pageNumber="229">Type material</emphasis>
.
<materialsCitation id="70ED3C963151A85AFEC3F9FFFD97F968" collectingDate="2022-06-21" collectionCode="MZL" collectorName="B. Launay" country="France" elevation="2193" location="Estaragne" pageId="8" pageNumber="229" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="1F3E88693151A85AFEC3F9FFFE34F9A8" box="[363,462,1620,1644]" pageId="8" pageNumber="229" type="holotype">Holotype</typeStatus>
male:
<emphasis id="F2F1EAD93151A85AFDB1F9FFFD72F9A8" bold="true" box="[537,648,1620,1644]" pageId="8" pageNumber="229">
<collectingCountry id="B892765B3151A85AFDB1F9FFFD72F9A8" box="[537,648,1620,1644]" name="France" pageId="8" pageNumber="229">FRANCE</collectingCountry>
</emphasis>
. Occitania. Hautes-Pyrénées; tributary to
<location id="C55A60103151A85AFE74F9DEFDB2F949" LSID="urn:lsid:plazi:treatment:482C87DD3151A85AFF3FFC3FFBA6F928:C55A60103151A85AFE74F9DEFDB2F949" box="[476,584,1653,1677]" country="France" name="Estaragne" pageId="8" pageNumber="229">Estaragne</location>
, cote
<quantity id="077D9B2E3151A85AFD21F9DFFF54F968" metricMagnitude="3" metricUnit="m" metricValue="2.193224" pageId="8" pageNumber="229" unit="m" value="2193.224">
<elevation id="4BA8D1F83151A85AFD21F9DFFF54F968" metricMagnitude="3" metricUnit="m" metricValue="2.193224" pageId="8" pageNumber="229" unit="m" value="2193.224">2193, 2240 m</elevation>
</quantity>
,
<date id="B43B100B3151A85AFF11F93FFEC4F968" box="[185,318,1684,1708]" pageId="8" pageNumber="229" value="2022-06-21">
<collectingDate id="A47FE9E33151A85AFF11F93FFEC4F968" box="[185,318,1684,1708]" pageId="8" pageNumber="229" value="2022-06-21">21 Jun 2022</collectingDate>
</date>
,
<specimenCount id="D683FD423151A85AFEE1F93FFE96F968" box="[329,364,1684,1708]" count="1" pageId="8" pageNumber="229" type="male">1♂</specimenCount>
, leg.
<collectorName id="6D70531D3151A85AFE0EF93EFDEFF969" box="[422,533,1685,1709]" pageId="8" pageNumber="229">B. Launay</collectorName>
(
<collectionCode id="A694AE0E3151A85AFD8BF93FFD9BF968" box="[547,609,1684,1708]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="8" pageNumber="229">MZL</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="C03A36CB3151A85AFF15F91EFE8EF848" blockId="8.[151,772,1300,1932]" pageId="8" pageNumber="229">
<materialsCitation id="70ED3C963151A85AFF15F91EFE8AF8E8" collectingDate="1986-07-21" collectionCode="MZL" collectorName="J. Aubert" country="France" elevation="790" location="Albies" municipality="Gargante" pageId="8" pageNumber="229" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<typeStatus id="1F3E88693151A85AFF15F91EFED3F909" box="[189,297,1717,1741]" pageId="8" pageNumber="229" type="paratype">Paratypes</typeStatus>
:
<emphasis id="F2F1EAD93151A85AFEEBF91FFE48F908" bold="true" box="[323,434,1716,1740]" pageId="8" pageNumber="229">
<collectingCountry id="B892765B3151A85AFEEBF91FFE48F908" box="[323,434,1716,1740]" name="France" pageId="8" pageNumber="229">FRANCE</collectingCountry>
</emphasis>
. Occitania. Ariège,
<location id="C55A60103151A85AFD1FF91FFD05F908" LSID="urn:lsid:plazi:treatment:482C87DD3151A85AFF3FFC3FFBA6F928:C55A60103151A85AFD1FF91FFD05F908" box="[695,767,1716,1740]" country="France" municipality="Gargante" name="Albies" pageId="8" pageNumber="229">Albiès</location>
,
<collectingMunicipality id="205EACB13151A85AFF3FF97FFF02F928" box="[151,248,1748,1772]" pageId="8" pageNumber="229">Gargante</collectingMunicipality>
brook,
<quantity id="077D9B2E3151A85AFEFCF97FFE65F928" box="[340,415,1748,1772]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="229" unit="m" value="790.0">
<elevation id="4BA8D1F83151A85AFEFCF97FFE65F928" box="[340,415,1748,1772]" metricMagnitude="2" metricUnit="m" metricValue="7.9" pageId="8" pageNumber="229" unit="m" value="790.0">790 m</elevation>
</quantity>
,
<date id="B43B100B3151A85AFE07F97FFDCDF928" box="[431,567,1748,1772]" pageId="8" pageNumber="229" value="1986-07-21">
<collectingDate id="A47FE9E33151A85AFE07F97FFDCDF928" box="[431,567,1748,1772]" pageId="8" pageNumber="229" value="1986-07-21">21 Jul 1986</collectingDate>
</date>
,
<specimenCount id="D683FD423151A85AFDEFF97FFD92F928" box="[583,616,1748,1772]" count="1" pageId="8" pageNumber="229" type="male">1♂</specimenCount>
with separat- ed aedeagus, leg.
<collectorName id="6D70531D3151A85AFEC2F95EFE28F8C8" box="[362,466,1780,1804]" pageId="8" pageNumber="229">J. Aubert</collectorName>
(
<collectionCode id="A694AE0E3151A85AFE4FF95FFDD9F8C8" box="[487,547,1780,1804]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="8" pageNumber="229">MZL</collectionCode>
, catalogue number: GBIFCH00913253)
</materialsCitation>
.
<materialsCitation id="70ED3C963151A85AFE20F8BFFE8AF848" collectingDate="1951-07" collectionCode="MZL" collectorName="H. Bertrand" country="Spain" elevation="1100" location="Noguera Ribagorzana" pageId="8" pageNumber="229" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="F2F1EAD93151A85AFE20F8BFFE2CF8E8" bold="true" box="[392,470,1812,1836]" pageId="8" pageNumber="229">
<collectingCountry id="B892765B3151A85AFE20F8BFFE2CF8E8" box="[392,470,1812,1836]" name="Spain" pageId="8" pageNumber="229">SPAIN</collectingCountry>
</emphasis>
. Lleida. Bono,
<location id="C55A60103151A85AFD0FF8BEFEDBF888" LSID="urn:lsid:plazi:treatment:482C87DD3151A85AFF3FFC3FFBA6F928:C55A60103151A85AFD0FF8BEFEDBF888" country="Spain" name="Noguera Ribagorzana" pageId="8" pageNumber="229">Noguera Ribagorzana</location>
,
<quantity id="077D9B2E3151A85AFE87F89FFE7CF888" box="[303,390,1844,1868]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="229" unit="m" value="1100.0">
<elevation id="4BA8D1F83151A85AFE87F89FFE7CF888" box="[303,390,1844,1868]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="229" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="B43B100B3151A85AFE3DF89FFE0DF888" box="[405,503,1844,1868]" pageId="8" pageNumber="229" value="1951-07">
<collectingDate id="A47FE9E33151A85AFE3DF89FFE0DF888" box="[405,503,1844,1868]" pageId="8" pageNumber="229" value="1951-07">Jul 1951</collectingDate>
</date>
,
<specimenCount id="D683FD423151A85AFDAEF89FFDDDF888" box="[518,551,1844,1868]" count="1" pageId="8" pageNumber="229" type="male">1♂</specimenCount>
microscopic prepa- ration, leg.
<collectorName id="6D70531D3151A85AFE87F8FEFE41F8A8" box="[303,443,1876,1900]" pageId="8" pageNumber="229">H. Bertrand</collectorName>
(
<collectionCode id="A694AE0E3151A85AFE70F8FFFDEEF8A8" box="[472,532,1876,1900]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="8" pageNumber="229">MZL</collectionCode>
, catalogue number: GBIFCH00913240)
</materialsCitation>
.
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDFF1FFC71FF28" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<typeStatus id="1F3E88693151A85AFCCDFF1FFBCDFF08" box="[869,1079,180,204]" isOtherTypeStatus="true" pageId="8" pageNumber="229">Additional material</typeStatus>
is listed in the Appendix 1, under
<typeStatus id="1F3E88693151A85AFCE8FF7EFC94FF29" box="[832,878,213,237]" pageId="8" pageNumber="229">type</typeStatus>
E.
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDFF5FFB11FDA8" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDFF5FFC03FEC8" bold="true" box="[869,1017,244,268]" pageId="8" pageNumber="229">Male adults.</emphasis>
Sclerotized apex of aedeagus slightly bulging out in its middle section (
<figureCitation id="58BE2A4E3151A85AFB03FEBFFAFEFEE8" box="[1195,1284,276,300]" captionStart="Plate XV" captionStartId="27.[136,181,1885,1904]" captionTargetBox="[244,1328,806,1862]" captionTargetId="figure-6@27.[244,1328,803,1862]" captionTargetPageId="27" captionText="Plate XV, Figs F.3–F.8 – Type F (putative Perla bipunctata from the foothills of the Pyrenees, Río Aragón). F.3, head and pronotum of nymph; F.4, tergites of nymph, dorsal view; F.5, tergites of nymph, dorsal view; F.6, markings on femora; F.7, paraprocts of nymph, ventral view; F.8, cerci of nymph, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12763207" httpUri="https://zenodo.org/record/12763207/files/figure.png" pageId="8" pageNumber="229">Fig. E.3</figureCitation>
). Proximal part of aedeagus with a strong row of aligned brush-like setae (
<figureCitation id="58BE2A4E3151A85AFCEFFEFFFC65FEA8" box="[839,927,340,364]" captionStart="Plate XV" captionStartId="27.[136,181,1885,1904]" captionTargetBox="[244,1328,806,1862]" captionTargetId="figure-6@27.[244,1328,803,1862]" captionTargetPageId="27" captionText="Plate XV, Figs F.3–F.8 – Type F (putative Perla bipunctata from the foothills of the Pyrenees, Río Aragón). F.3, head and pronotum of nymph; F.4, tergites of nymph, dorsal view; F.5, tergites of nymph, dorsal view; F.6, markings on femora; F.7, paraprocts of nymph, ventral view; F.8, cerci of nymph, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12763207" httpUri="https://zenodo.org/record/12763207/files/figure.png" pageId="8" pageNumber="229">Fig. E.3</figureCitation>
); distal part terminated by two narrow blunt ends, medially interrupted (
<figureCitation id="58BE2A4E3151A85AFB82FEDFFB7EFE48" box="[1066,1156,372,396]" captionStart="Plate XV" captionStartId="27.[136,181,1885,1904]" captionTargetBox="[244,1328,806,1862]" captionTargetId="figure-6@27.[244,1328,803,1862]" captionTargetPageId="27" captionText="Plate XV, Figs F.3–F.8 – Type F (putative Perla bipunctata from the foothills of the Pyrenees, Río Aragón). F.3, head and pronotum of nymph; F.4, tergites of nymph, dorsal view; F.5, tergites of nymph, dorsal view; F.6, markings on femora; F.7, paraprocts of nymph, ventral view; F.8, cerci of nymph, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12763207" httpUri="https://zenodo.org/record/12763207/files/figure.png" pageId="8" pageNumber="229">Fig. E.3</figureCitation>
). Length of aedeagus: 0.65 mm; width: 0.30 mm. Aedeagal tube with parallel sides (
<figureCitation id="58BE2A4E3151A85AFCEFFE1FFC65FE08" box="[839,927,436,460]" captionStart="Plate III" captionStartId="16.[151,197,1446,1465]" captionTargetBox="[151,1451,166,1411]" captionTargetId="figure-10@16.[151,1453,164,1412]" captionTargetPageId="16" captionText="Plate III, Figs A.16–A.21 – Type A (French and Swiss Alps). A.16, head and pronotum of nymph (Swiss Alps); A.17, markings on tergites of nymph (Gryonne valley); A.18, markings on tergites of nymph (Sense); A.19, markings on the femora of legs, dorsal view (Gryonne Valley); A.20, sclerotized apex of extracted aedeagus of an adult ♂ (Vercors); A.21, sclerotized apex of aedeagus of Perla grandis, from Aubert 1949: 225, his Fig. 4." figureDoi="http://doi.org/10.5281/zenodo.12763182" httpUri="https://zenodo.org/record/12763182/files/figure.png" pageId="8" pageNumber="229">Fig. E.4</figureCitation>
) and covered by a dense grid of short setae (
<figureCitation id="58BE2A4E3151A85AFA28FE1FFC9CFE28" captionStart="Plate IX" captionStartId="22.[151,197,1447,1466]" captionTargetBox="[151,1451,166,1414]" captionTargetId="figure-10@22.[151,1454,163,1414]" captionTargetPageId="22" captionText="Plate IX, Figs C.13–C.18 – Type C (nymphs, Nure, Rivers Aare and Emme). C.13, head and pronotum of nymph (Nure); C.14, mediodorsal setation on tergites, lateral view (Nure); C.15, paraprocts, ventral view (Aare); C.16, head and pronotum of nymph (Aare); C.17, oval markings on femora (Emme); C.18, sclerotized apex of aedeagus of Perla bipunctata, from Aubert 1949: 225, his Fig. 5." figureDoi="http://doi.org/10.5281/zenodo.12763195" httpUri="https://zenodo.org/record/12763195/files/figure.png" pageId="8" pageNumber="229">Fig. E.5</figureCitation>
). Mesal field on tergite IX short and narrow, with few spiny projections, and shallow distal division (Figs E.6, E.7). Distal side of hemitergal lobes of tergite X apically enlarged, in lateral view (Fig. E.8). Apex of hemitergal lobes with very strong spines (Fig. E.8).
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDFDDFFBE4FC48" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDFDDFFB87FD48" bold="true" box="[869,1149,628,652]" pageId="8" pageNumber="229">Morphological affinities.</emphasis>
Adult males of
<typeStatus id="1F3E88693151A85AFA84FDDEFA9EFD49" box="[1324,1380,629,653]" pageId="8" pageNumber="229">types</typeStatus>
E and F are separable by the shape of the sclerotized apex of their aedeagus. Whereas the one of
<taxonomicName id="07854D483151A85AFB2DFD1FFACBFD08" authorityName="Reding" authorityYear="2023" box="[1157,1329,692,716]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="pyrenaica" status="sp. nov.">
<emphasis id="F2F1EAD93151A85AFB2DFD1FFACBFD08" box="[1157,1329,692,716]" italics="true" pageId="8" pageNumber="229">Perla pyrenaica</emphasis>
</taxonomicName>
<emphasis id="F2F1EAD93151A85AFA9FFD1FFA89FD08" bold="true" box="[1335,1395,692,716]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="E9C257A23151A85AFA9FFD1FFA89FD08" box="[1335,1395,692,716]" pageId="8" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
ends into two blunt extensions (
<figureCitation id="58BE2A4E3151A85AFBF1FD7FFB4AFD28" box="[1113,1200,724,748]" captionStart="Plate XV" captionStartId="27.[136,181,1885,1904]" captionTargetBox="[244,1328,806,1862]" captionTargetId="figure-6@27.[244,1328,803,1862]" captionTargetPageId="27" captionText="Plate XV, Figs F.3–F.8 – Type F (putative Perla bipunctata from the foothills of the Pyrenees, Río Aragón). F.3, head and pronotum of nymph; F.4, tergites of nymph, dorsal view; F.5, tergites of nymph, dorsal view; F.6, markings on femora; F.7, paraprocts of nymph, ventral view; F.8, cerci of nymph, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12763207" httpUri="https://zenodo.org/record/12763207/files/figure.png" pageId="8" pageNumber="229">Fig. E.3</figureCitation>
), putative specimens of
<taxonomicName id="07854D483151A85AFCE8FD5FFC0EFCC8" baseAuthorityName="Pictet" baseAuthorityYear="1841" box="[832,1012,756,780]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="F2F1EAD93151A85AFCE8FD5FFC0EFCC8" box="[832,1012,756,780]" italics="true" pageId="8" pageNumber="229">Perla bipunctata</emphasis>
</taxonomicName>
from the Pyrenees (
<typeStatus id="1F3E88693151A85AFB65FD5FFB06FCC8" box="[1229,1276,756,780]" pageId="8" pageNumber="229">type</typeStatus>
F) have a much shorter, and distally indented aedeagus (
<figureCitation id="58BE2A4E3151A85AFB57FCBFFAA2FCE8" box="[1279,1368,788,812]" captionStart="Plate I" captionStartId="14.[151,197,1805,1824]" captionTargetBox="[249,1342,166,1766]" captionTargetId="figure-10@14.[247,1342,166,1766]" captionTargetPageId="14" captionText="Plate I, Figs A.1–A.9 – Type A (Italian Alps and Apennines). A.1, hemitergal lobes, mesal field with sensilla and sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.2, sclerotized apex of everted aedeagus of an adult ♂ (Staffora); A.3, sclerotized apex of extracted aedeagus of an adult ♂ (Stura di Ala); A.4, sclerotized apex of everted aedeagus of an adult ♂ (Soana); A.5, aedeagal tube of adult ♂; A.6, aedeagal tube and hemitergal lobes of adult ♂; A.7, adult ♂, hemitergal lobes and mesal field with sensilla (Southern Apennines); A.8, sclerotized apex of extracted aedeagus of an adult ♂ (Southern Apennines); A.9, adult ♂, hemitergal lobes and mesal field with sensilla, dorsal view (Stura di Demonte)." figureDoi="http://doi.org/10.5281/zenodo.12763178" httpUri="https://zenodo.org/record/12763178/files/figure.png" pageId="8" pageNumber="229">Fig. F.1</figureCitation>
).
<taxonomicName id="07854D483151A85AFAD8FCBFFC50FC88" authorityName="Reding" authorityYear="2023" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="pyrenaica" status="sp. nov.">
<emphasis id="F2F1EAD93151A85AFAD8FCBFFC50FC88" italics="true" pageId="8" pageNumber="229">Perla pyrenaica</emphasis>
</taxonomicName>
<emphasis id="F2F1EAD93151A85AFC1AFC9FFC15FC88" bold="true" box="[946,1007,820,844]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="E9C257A23151A85AFC1AFC9FFC15FC88" box="[946,1007,820,844]" pageId="8" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="58BE2A4E3151A85AFBA8FC9FFBA3FC88" box="[1024,1113,820,844]" captionStart="Plate XV" captionStartId="27.[136,181,1885,1904]" captionTargetBox="[244,1328,806,1862]" captionTargetId="figure-6@27.[244,1328,803,1862]" captionTargetPageId="27" captionText="Plate XV, Figs F.3–F.8 – Type F (putative Perla bipunctata from the foothills of the Pyrenees, Río Aragón). F.3, head and pronotum of nymph; F.4, tergites of nymph, dorsal view; F.5, tergites of nymph, dorsal view; F.6, markings on femora; F.7, paraprocts of nymph, ventral view; F.8, cerci of nymph, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12763207" httpUri="https://zenodo.org/record/12763207/files/figure.png" pageId="8" pageNumber="229">Fig. E.3</figureCitation>
) differs from alpine
<taxonomicName id="07854D483151A85AFA96FC9EFA51FC88" authorityName=", sensu Sivec &amp; Stark" authorityYear="2002" box="[1342,1451,820,844]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="F2F1EAD93151A85AFA96FC9EFA51FC88" box="[1342,1451,820,844]" italics="true" pageId="8" pageNumber="229">P. grandis</emphasis>
</taxonomicName>
(
<figureCitation id="58BE2A4E3151A85AFCEFFCFFFC51FCA8" box="[839,939,852,876]" captionStart="Plate III" captionStartId="16.[151,197,1446,1465]" captionTargetBox="[151,1451,166,1411]" captionTargetId="figure-10@16.[151,1453,164,1412]" captionTargetPageId="16" captionText="Plate III, Figs A.16–A.21 – Type A (French and Swiss Alps). A.16, head and pronotum of nymph (Swiss Alps); A.17, markings on tergites of nymph (Gryonne valley); A.18, markings on tergites of nymph (Sense); A.19, markings on the femora of legs, dorsal view (Gryonne Valley); A.20, sclerotized apex of extracted aedeagus of an adult ♂ (Vercors); A.21, sclerotized apex of aedeagus of Perla grandis, from Aubert 1949: 225, his Fig. 4." figureDoi="http://doi.org/10.5281/zenodo.12763182" httpUri="https://zenodo.org/record/12763182/files/figure.png" pageId="8" pageNumber="229">Figs. A.4</figureCitation>
, A.8) by its much shorter and medially enlarged apex of its aedeagus.
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDFC3FFB81FC08" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDFC3FFBF7FC68" bold="true" box="[869,1037,916,940]" pageId="8" pageNumber="229">Female adults.</emphasis>
Subgenital plate with two distinct triangular extensions (Fig. E.9).
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDFC7FFB3FFAA8" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDFC7FFC31FC28" bold="true" box="[869,971,980,1004]" pageId="8" pageNumber="229">Nymphs.</emphasis>
Length:
<quantity id="077D9B2E3151A85AFB81FC7FFB4BFC28" box="[1065,1201,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.6" metricValueMin="2.4" pageId="8" pageNumber="229" unit="mm" value="30.0" valueMax="36.0" valueMin="24.0">24 to 36 mm</quantity>
. Pronotum with a large, dark, central stripe flanked by two irregularly shaped dark bands loosely connected to the upper and to the lower edges of the pronotum (
<figureCitation id="58BE2A4E3151A85AFB8AFB9FFB6FFB88" box="[1058,1173,1076,1100]" captionStart="Plate II" captionStartId="15.[136,182,1446,1465]" captionTargetBox="[136,1436,166,1412]" captionTargetId="figure-4@15.[134,1436,163,1412]" captionTargetPageId="15" captionText="Plate II, Figs A.10–A.15 – Type A (male and female adults, nymphs). A.10, adult ♂, hemitergal lobes, lateral view (Southern Apennines); A.11, head and pronotum of an adult ♂ (Stura di Demonte); A.12, subgenital plate of adult ♀ (Stura di Demonte); A.13, subgenital plate of adult ♀ (Haute-Savoie); A.14, basal section of cerci of nymph (Staffora); A.15, basal section of cerci of nymph (Gryonne valley)." figureDoi="http://doi.org/10.5281/zenodo.12763180" httpUri="https://zenodo.org/record/12763180/files/figure.png" pageId="8" pageNumber="229">Figs. E.10</figureCitation>
, E.
<figureCitation id="58BE2A4E3151A85AFB13FB9FFB2CFB88" box="[1211,1238,1076,1100]" captionStart="Plate XIV" captionStartId="27.[136,182,679,698]" captionTargetBox="[283,1313,166,646]" captionTargetId="figure-4@27.[241,1313,164,647]" captionTargetPageId="27" captionText="Plate XIV, Figs F.1–F.2 – Type F (putative Perla bipunctata from the foothills of the French Pyrenees). F.1, illustrations of Despax (1942, his Figs 11, 12. Fig. 11 = adult ♂b with everted aedeagus, lateral view; Fig. 12 = adult ♂b, sclerotized apex of aedeagus); F.2, illustrations of Despax (1951, his Fig. 86, D, E, F. D = adult ♂b, mesal field with sensilla; E = adult ♂b, mesal field; F = adult ♂b, hemitergite)." figureDoi="http://doi.org/10.5281/zenodo.12763205" httpUri="https://zenodo.org/record/12763205/files/figure.png" pageId="8" pageNumber="229">11</figureCitation>
). Rim of pronotum with a wide, half-circled dark band (
<figureCitation id="58BE2A4E3151A85AFB75FBFFFAB2FBA8" box="[1245,1352,1108,1132]" captionStart="Plate II" captionStartId="15.[136,182,1446,1465]" captionTargetBox="[136,1436,166,1412]" captionTargetId="figure-4@15.[134,1436,163,1412]" captionTargetPageId="15" captionText="Plate II, Figs A.10–A.15 – Type A (male and female adults, nymphs). A.10, adult ♂, hemitergal lobes, lateral view (Southern Apennines); A.11, head and pronotum of an adult ♂ (Stura di Demonte); A.12, subgenital plate of adult ♀ (Stura di Demonte); A.13, subgenital plate of adult ♀ (Haute-Savoie); A.14, basal section of cerci of nymph (Staffora); A.15, basal section of cerci of nymph (Gryonne valley)." figureDoi="http://doi.org/10.5281/zenodo.12763180" httpUri="https://zenodo.org/record/12763180/files/figure.png" pageId="8" pageNumber="229">Fig. E.10</figureCitation>
). Lateral edges of pronotum yellow (
<figureCitation id="58BE2A4E3151A85AFBDBFBDFFB26FB48" box="[1139,1244,1140,1164]" captionStart="Plate II" captionStartId="15.[136,182,1446,1465]" captionTargetBox="[136,1436,166,1412]" captionTargetId="figure-4@15.[134,1436,163,1412]" captionTargetPageId="15" captionText="Plate II, Figs A.10–A.15 – Type A (male and female adults, nymphs). A.10, adult ♂, hemitergal lobes, lateral view (Southern Apennines); A.11, head and pronotum of an adult ♂ (Stura di Demonte); A.12, subgenital plate of adult ♀ (Stura di Demonte); A.13, subgenital plate of adult ♀ (Haute-Savoie); A.14, basal section of cerci of nymph (Staffora); A.15, basal section of cerci of nymph (Gryonne valley)." figureDoi="http://doi.org/10.5281/zenodo.12763180" httpUri="https://zenodo.org/record/12763180/files/figure.png" pageId="8" pageNumber="229">Fig. E.10</figureCitation>
). Yellow markings on tergites III to
<collectionCode id="A694AE0E3151A85AFC5FFB3EFBF0FB68" box="[1015,1034,1173,1196]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="229" type="Museum">V</collectionCode>
dot-like, medially separated by a dark stripe (
<figureCitation id="58BE2A4E3151A85AFC24FB1FFC0FFB08" box="[908,1013,1204,1228]" captionStart="Plate XIV" captionStartId="27.[136,182,679,698]" captionTargetBox="[283,1313,166,646]" captionTargetId="figure-4@27.[241,1313,164,647]" captionTargetPageId="27" captionText="Plate XIV, Figs F.1–F.2 – Type F (putative Perla bipunctata from the foothills of the French Pyrenees). F.1, illustrations of Despax (1942, his Figs 11, 12. Fig. 11 = adult ♂b with everted aedeagus, lateral view; Fig. 12 = adult ♂b, sclerotized apex of aedeagus); F.2, illustrations of Despax (1951, his Fig. 86, D, E, F. D = adult ♂b, mesal field with sensilla; E = adult ♂b, mesal field; F = adult ♂b, hemitergite)." figureDoi="http://doi.org/10.5281/zenodo.12763205" httpUri="https://zenodo.org/record/12763205/files/figure.png" pageId="8" pageNumber="229">Fig. E.12</figureCitation>
). Dorsal faces of femora of middle and hind legs with an indistinct dark area connected to the dorsal edge of the femora (
<figureCitation id="58BE2A4E3151A85AFBE2FB5FFB49FAC8" box="[1098,1203,1268,1292]" captionStart="Plate IX" captionStartId="22.[151,197,1447,1466]" captionTargetBox="[151,1451,166,1414]" captionTargetId="figure-10@22.[151,1454,163,1414]" captionTargetPageId="22" captionText="Plate IX, Figs C.13–C.18 – Type C (nymphs, Nure, Rivers Aare and Emme). C.13, head and pronotum of nymph (Nure); C.14, mediodorsal setation on tergites, lateral view (Nure); C.15, paraprocts, ventral view (Aare); C.16, head and pronotum of nymph (Aare); C.17, oval markings on femora (Emme); C.18, sclerotized apex of aedeagus of Perla bipunctata, from Aubert 1949: 225, his Fig. 5." figureDoi="http://doi.org/10.5281/zenodo.12763195" httpUri="https://zenodo.org/record/12763195/files/figure.png" pageId="8" pageNumber="229">Fig. E.13</figureCitation>
). Cerci with separated bunches of basal mediodorsal setae (Fig. E.14). Tergites with long mediodorsal flexible setae, in lateral view. Paraprocts with conical apex (Fig. E.15).
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDFADFFAADF9C8" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDFADFFB84FA48" bold="true" box="[869,1150,1396,1420]" pageId="8" pageNumber="229">Morphological affinities.</emphasis>
Nymphs of
<taxonomicName id="07854D483151A85AFABDFADFFC9BFA68" authorityName="Reding" authorityYear="2023" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="pyrenaica" status="sp. nov.">
<emphasis id="F2F1EAD93151A85AFABDFADFFC9BFA68" italics="true" pageId="8" pageNumber="229">Perla pyrenaica</emphasis>
</taxonomicName>
<emphasis id="F2F1EAD93151A85AFCC1FA3FFC52FA68" bold="true" box="[873,936,1428,1452]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="E9C257A23151A85AFCC1FA3FFC52FA68" box="[873,936,1428,1452]" pageId="8" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
are characterized by the presence of separated bunches of basal mediodorsal setae on their cerci (Fig. E.14), whereas
<taxonomicName id="07854D483151A85AFC47FA7EFBA4FA28" authorityName=", sensu Sivec &amp; Stark" authorityYear="2002" box="[1007,1118,1492,1516]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="F2F1EAD93151A85AFC47FA7EFBA4FA28" box="[1007,1118,1492,1516]" italics="true" pageId="8" pageNumber="229">P. grandis</emphasis>
</taxonomicName>
nymphs exhibit a continuous row of long, mediodorsal setae (Figs. A.14, A.15).
</paragraph>
</subSubSection>
<subSubSection id="889F65403151A85AFCCDF9BFFBA6F928" pageId="8" pageNumber="229" type="distribution">
<paragraph id="C03A36CB3151A85AFCCDF9BFFAABF9A8" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDF9BFFC0AF9E8" bold="true" box="[869,1008,1556,1580]" pageId="8" pageNumber="229">Distribution</emphasis>
. This species is known from high altitude brooks and springs on both slopes of the Pyrenees. The material is listed in the Appendix 1, under
<typeStatus id="1F3E88693151A85AFAADF9FEFAC9F9A9" box="[1285,1331,1621,1645]" pageId="8" pageNumber="229">type</typeStatus>
E.
</paragraph>
<paragraph id="C03A36CB3151A85AFCCDF9DFFBA6F928" blockId="8.[832,1452,180,1772]" pageId="8" pageNumber="229">
<emphasis id="F2F1EAD93151A85AFCCDF9DFFBB9F948" bold="true" box="[869,1091,1652,1676]" pageId="8" pageNumber="229">Derivation nominis</emphasis>
. This species is named after the region which it inhabits. The adjective
<taxonomicName id="07854D483151A85AFB41F93EFAA9F968" authorityName="Reding" authorityYear="2023" box="[1257,1363,1685,1708]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="pyrenaica">
<emphasis id="F2F1EAD93151A85AFB41F93EFAA9F968" box="[1257,1363,1685,1708]" italics="true" pageId="8" pageNumber="229">pyrenaica</emphasis>
</taxonomicName>
is to be treated as a feminine Latin adjective in the nominative case combined with
<taxonomicName id="07854D483151A85AFBB2F97FFBAFF928" authorityName="Geoffroy" authorityYear="1762" box="[1050,1109,1748,1772]" class="Insecta" family="Perlidae" genus="Perla" kingdom="Animalia" order="Plecoptera" pageId="8" pageNumber="229" phylum="Arthropoda" rank="genus">
<emphasis id="F2F1EAD93151A85AFBB2F97FFBAFF928" box="[1050,1109,1748,1772]" italics="true" pageId="8" pageNumber="229">Perla</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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