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<document id="3F02E3DDB63107EE2C42BF2D1B4B88A9" ID-DOI="10.5281/zenodo.13229251" ID-ISSN="2345-7600" ID-Zenodo-Dep="13229251" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722894128278" checkinUser="felipe" docAuthor="Khaironizam, M. Z. &amp; Norma-Rashid, Y." docDate="2003" docId="03FB87EF206EFFEED14E5E93FC03FC4A" docLanguage="en" docName="RafflesBZool.51.1.97-100.pdf" docOrigin="Raffles Bulletin of Zoology 51 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Periophthalmodon septemradiatus" docType="treatment" docVersion="2" lastPageNumber="98" masterDocId="FFC2FF97206EFFECD2305A3AFF82FF82" masterDocTitle="First Record Of The Mudskipper, Periophthalmodon Septemradiatus (Hamilton) (Teleostei: Gobiidae) From Peninsular Malaysia" masterLastPageNumber="100" masterPageNumber="97" pageNumber="97" updateTime="1722902215981" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="8B9982E71E478FE51031E8E88C64579C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="02EC603FB4B9CC3B2A09BC54D3D988AF">
<mods:title id="E823CC2D5B99292ED9C0B5C8A82EB039">First Record Of The Mudskipper, Periophthalmodon Septemradiatus (Hamilton) (Teleostei: Gobiidae) From Peninsular Malaysia</mods:title>
</mods:titleInfo>
<mods:name id="FC119D15D99985FBCE452F2318968B59" type="personal">
<mods:role id="D00E83B14B04050468C6C117CB7F10FC">
<mods:roleTerm id="52EF933BE4B7877393C3D5A1C94FC7CE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="46867DB656F9099AD3AB8455DD3C6FFA">Khaironizam, M. Z.</mods:namePart>
</mods:name>
<mods:name id="CCA91FB7384CC2779AE160AF149450D1" type="personal">
<mods:role id="82AD3A5602A5FE2F7769198BEAECA3AC">
<mods:roleTerm id="B2C0A54357AA286484ABA8723FFEA6AC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAA1D61472CB60B0A800566B4A34DB26">Norma-Rashid, Y.</mods:namePart>
</mods:name>
<mods:typeOfResource id="67995542EA6F40B182F50B6C5722D4E9">text</mods:typeOfResource>
<mods:relatedItem id="6FBB9C7923823071B27B34A95177C89C" type="host">
<mods:titleInfo id="60C0E52C2511C311E6780D52B21FB8E9">
<mods:title id="D518A190CBD81AFF99683C84B89BBB3D">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="7FF5B56BB7D6422DB3FB8A8784D64E41">
<mods:date id="296C0764B05BB7FF34E9E684F2EF2561">2003</mods:date>
<mods:detail id="03FCAEB2A52933EF0973155FE7C41DFA" type="volume">
<mods:number id="9D0AD46BE584C1239F33296C13394C10">51</mods:number>
</mods:detail>
<mods:detail id="8A2A5AC0A05FE4625C3852050B812D9A" type="issue">
<mods:number id="9D3D7FE94302733616A3A9610AE71CD2">1</mods:number>
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<mods:start id="9DE1994708F38538BE4A3D45555D66DF">97</mods:start>
<mods:end id="720BB4523D58A36E6FF0D110F7B3D232">100</mods:end>
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<mods:classification id="BA3D2F5877F56293E608BF446077F49D">journal article</mods:classification>
<mods:identifier id="1432DDF8D8579AB75224422A523D49B5" type="DOI">10.5281/zenodo.13229251</mods:identifier>
<mods:identifier id="8627DC97DD28955CC3215D98431B4272" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="C1EA54657BE2C1D8EF974141B339731A" type="Zenodo-Dep">13229251</mods:identifier>
<mods:identifier id="A8649D5DD3068DAFC25DE07D9300CFE9" type="ZooBank">B924E4BA-A5AB-4C4F-87E9-2166400EAA93</mods:identifier>
</mods:mods>
<treatment id="03FB87EF206EFFEED14E5E93FC03FC4A" LSID="urn:lsid:plazi:treatment:03FB87EF206EFFEED14E5E93FC03FC4A" httpUri="http://treatment.plazi.org/id/03FB87EF206EFFEED14E5E93FC03FC4A" lastPageId="2" lastPageNumber="98" pageId="0" pageNumber="97">
<subSubSection id="C3486572206EFFECD14E5E93FA05FB43" box="[894,1415,1193,1217]" pageId="0" pageNumber="97" type="nomenclature">
<paragraph id="8BED36F9206EFFECD14E5E93FA05FB43" blockId="0.[894,1415,1193,1217]" box="[894,1415,1193,1217]" pageId="0" pageNumber="97">
<heading id="D0A58195206EFFECD14E5E93FA05FB43" bold="true" box="[894,1415,1193,1217]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="97" reason="2">
<taxonomicName id="4C524D7A206EFFECD14E5E93FA05FB43" ID-CoL="4F2SW" authority="(Hamilton)" baseAuthorityName="Hamilton" box="[894,1415,1193,1217]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="0" pageNumber="97" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206EFFECD14E5E93FA05FB43" bold="true" box="[894,1415,1193,1217]" pageId="0" pageNumber="97">
<emphasis id="B926EAEB206EFFECD14E5E93FB7DFB43" bold="true" box="[894,1279,1193,1217]" italics="true" pageId="0" pageNumber="97">Periophthalmodon septemradiatus</emphasis>
(Hamilton)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3486572206EFFECD1735ED3FACCFA0C" pageId="0" pageNumber="97" type="materials_examined">
<paragraph id="8BED36F9206EFFECD1735ED3FACCFA0C" blockId="0.[835,1475,1257,1422]" pageId="0" pageNumber="97">
<emphasis id="B926EAEB206EFFECD1735ED3FB96FB7C" bold="true" box="[835,1044,1257,1278]" italics="true" pageId="0" pageNumber="97">Material examined. –</emphasis>
<materialsCitation id="3B3A3CA4206EFFECD6295ED3FA83FAB5" collectingDate="2000-03-06" collectorName="M. Z. Khaironizam &amp; Y. Norma-Rashid" country="Malaysia" location="Selangor" pageId="0" pageNumber="97" specimenCode="KMZ-NYR000306" specimenCount="20" stateProvince="Selangor">
<specimenCount id="9D54FD70206EFFECD6295ED3FBD7FB7C" box="[1049,1109,1257,1278]" count="20" pageId="0" pageNumber="97" type="generic">20 ex.</specimenCount>
,
<specimenCode id="DBF49E82206EFFECD66D5ED3FA94FB7C" box="[1117,1302,1257,1278]" collectionCode="KMZ-NYR" pageId="0" pageNumber="97">KMZ-NYR000306</specimenCode>
KK (2),
<collectingCountry id="F3457669206EFFECD7535ED3FA3DFB7C" box="[1379,1471,1257,1278]" name="Malaysia" pageId="0" pageNumber="97">Malaysia</collectingCountry>
:
<collectingRegion id="4996F81B206EFFECD1735F3FFC23FA98" box="[835,929,1285,1306]" country="Malaysia" name="Selangor" pageId="0" pageNumber="97">Selangor</collectingRegion>
,
<collectingRegion id="4996F81B206EFFECD1805F3FFB89FA98" box="[944,1035,1285,1306]" country="Malaysia" name="Selangor" pageId="0" pageNumber="97">Selangor</collectingRegion>
River at Kampung Kuantan, coll.
<collectorName id="26A7532F206EFFECD7B65F3FFC42FAB5" pageId="0" pageNumber="97">M. Z. Khaironizam</collectorName>
&amp;
<collectorName id="26A7532F206EFFECD1EC5F18FB05FAB5" box="[988,1159,1314,1335]" pageId="0" pageNumber="97">Y. Norma-Rashid</collectorName>
,
<date id="FFEC1039206EFFECD6BF5F18FA83FAB5" box="[1167,1281,1314,1335]" pageId="0" pageNumber="97" value="2000-03-06">
<collectingDate id="EFA8E9D1206EFFECD6BF5F18FA83FAB5" box="[1167,1281,1314,1335]" pageId="0" pageNumber="97" value="2000-03-06">6 Mar.2000</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3A3CA4206EFFECD7395F18FBA3FAF3" collectingDate="1997-09-08" collectionCode="ZRC" collectorName="Y. Y. Goh" country="Malaysia" location="Muar River" pageId="0" pageNumber="97" specimenCode="ZRC 41648" specimenCount="10" stateProvince="Johor">
<specimenCount id="9D54FD70206EFFECD7395F18FAC7FAB5" box="[1289,1349,1314,1335]" count="10" pageId="0" pageNumber="97" type="generic">10 ex.</specimenCount>
,
<specimenCode id="DBF49E82206EFFECD77D5F18FA3CFAB5" box="[1357,1470,1314,1335]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="97">ZRC 41648</specimenCode>
,
<collectingCountry id="F3457669206EFFECD1735F05FC22FAD6" box="[835,928,1343,1364]" name="Malaysia" pageId="0" pageNumber="97">Malaysia</collectingCountry>
:
<collectingRegion id="4996F81B206EFFECD19B5F05FC66FAD6" box="[939,996,1343,1364]" country="Malaysia" name="Johor" pageId="0" pageNumber="97">Johor</collectingRegion>
, Muar,
<location id="8E8D6022206EFFECD6015F05FB26FAD6" LSID="urn:lsid:plazi:treatment:03FB87EF206EFFEED14E5E93FC03FC4A:8E8D6022206EFFECD6015F05FB26FAD6" box="[1073,1188,1343,1364]" country="Malaysia" name="Muar River" pageId="0" pageNumber="97" stateProvince="Johor">Muar River</location>
below Tanjung Selabu, coll.
<collectorName id="26A7532F206EFFECD1735F66FC2BFAF3" box="[835,937,1372,1393]" pageId="0" pageNumber="97">Y. Y. Goh</collectorName>
,
<date id="FFEC1039206EFFECD1825F66FBA3FAF3" box="[946,1057,1372,1393]" pageId="0" pageNumber="97" value="1997-09-08">
<collectingDate id="EFA8E9D1206EFFECD1825F66FBA3FAF3" box="[946,1057,1372,1393]" pageId="0" pageNumber="97" value="1997-09-08">8 Sep.1997</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B3A3CA4206EFFECD61A5F66FAC8FA0C" collectingDate="1996-11-18" collectionCode="ZRC" collectorName="Y. Y. Goh" country="Malaysia" location="Muar River" pageId="0" pageNumber="97" specimenCode="ZRC 41172" specimenCount="6" stateProvince="Johor">
<specimenCount id="9D54FD70206EFFECD61A5F66FBD9FAF3" box="[1066,1115,1372,1393]" count="6" pageId="0" pageNumber="97" type="generic">6 ex.</specimenCount>
,
<specimenCode id="DBF49E82206EFFECD6545F66FB55FAF3" box="[1124,1239,1372,1393]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="0" pageNumber="97">ZRC 41172</specimenCode>
,
<collectingCountry id="F3457669206EFFECD6D05F66FABFFAF3" box="[1248,1341,1372,1393]" name="Malaysia" pageId="0" pageNumber="97">Malaysia</collectingCountry>
:
<collectingRegion id="4996F81B206EFFECD7765F66FAFDFAF3" box="[1350,1407,1372,1393]" country="Malaysia" name="Johor" pageId="0" pageNumber="97">Johor</collectingRegion>
, Muar,
<location id="8E8D6022206EFFECD1735F43FC34FA0C" LSID="urn:lsid:plazi:treatment:03FB87EF206EFFEED14E5E93FC03FC4A:8E8D6022206EFFECD1735F43FC34FA0C" box="[835,950,1401,1422]" country="Malaysia" name="Muar River" pageId="0" pageNumber="97" stateProvince="Johor">Muar River</location>
at Jorak, coll.
<collectorName id="26A7532F206EFFECD67E5F43FB3BFA0C" box="[1102,1209,1401,1422]" pageId="0" pageNumber="97">Y. Y. Goh</collectorName>
,
<date id="FFEC1039206EFFECD6F55F43FAC8FA0C" box="[1221,1354,1401,1422]" pageId="0" pageNumber="97" value="1996-11-18">
<collectingDate id="EFA8E9D1206EFFECD6F55F43FAC8FA0C" box="[1221,1354,1401,1422]" pageId="0" pageNumber="97" value="1996-11-18">18 Nov.1996</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3486572206EFFEDD1735F8CFD0AFD1A" lastPageId="1" lastPageNumber="98" pageId="0" pageNumber="97" type="diagnosis">
<paragraph id="8BED36F9206EFFECD1735F8CFBE5F8EF" blockId="0.[835,1475,1462,1901]" pageId="0" pageNumber="97">
<emphasis id="B926EAEB206EFFECD1735F8CFC54FA4C" bold="true" box="[835,982,1462,1486]" italics="true" pageId="0" pageNumber="97">Diagnosis. –</emphasis>
No pelvic frenum, pelvic fins completely separated. Snout scaleless, isthmus completely scaled. Scales on entire body large and cycloid. Teeth of upper jaw in two rows: anterior teeth of outer jaw small caninoid and smaller in the inner row. First dorsal fins (D1) in the female poorly developed possessing 3-5 short spines, and in some specimens are rudimentary. In the male, the first dorsal fin is of moderate height with a concave margin and an elongated first spine with 12 unbranched spines (
<figureCitation id="13692A7C206EFFECD6C25C8CFAD1F94F" box="[1266,1363,1718,1741]" captionStart="Fig" captionStartId="2.[136,172,2037,2058]" captionTargetBox="[292,1328,1307,2005]" captionTargetId="figure-7@2.[289,1330,1306,2020]" captionTargetPageId="2" captionText="Fig. 2. Sexual dimorphism on Periophthalmodon septemradiatus [KMZ-NYR000306KK (2)]. a) Male: the first dorsal fin is of moderate height with an elongated spine and contiguous with the second dorsal fins. b) Female: the first dorsal fin is poorly developed and not contiguous with the second dorsal fins." figureDoi="http://doi.org/10.5281/zenodo.13229255" httpUri="https://zenodo.org/record/13229255/files/figure.png" pageId="0" pageNumber="97">Figure 2</figureCitation>
). Second dorsal fin (D2) with 12-14 spines (the first two unbranched). The first and second dorsal fins are contiguous. Pectoral and caudal fin spines are also branched. Table 1 shows the meristic and morphometric data of the specimens under KMZ-NYR000306KK (2).
</paragraph>
<paragraph id="8BED36F9206EFFEDD1735DACFDE4FE3A" blockId="0.[835,1474,1942,2093]" lastBlockId="1.[113,752,193,440]" lastPageId="1" lastPageNumber="98" pageId="0" pageNumber="97">
<emphasis id="B926EAEB206EFFECD1735DACFBB3F82C" bold="true" box="[835,1073,1942,1966]" italics="true" pageId="0" pageNumber="97">Colour description. –</emphasis>
Freshly dead specimens have dusky grey heads and, brown on dorsum, white on ventrum and numerous pale red and pale blue spots on snout, opercle and trunk. A dusky longitudinal stripe on the predorsum starts from eye and terminates under the end of D2 or just before the D
<quantity id="4CAA9B1C206FFFEDD2835AFBFF60FF5A" box="[179,226,193,216]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="98" unit="in" value="2.0">2 in</quantity>
certain specimens. Eight black, saddle-like blotches occur dorsally: first anterior to D1, second across base of D1, third between D1 and D2, fourth, fifth and sixth across base of D2, seventh between D2 and caudal fins and eighth anterior to caudal fin. Dorsal fins: D1 black with reddish margin and D2 dusky with a red margin. Caudal and pectoral fins greyish and numerous red and blue speckles on pectoral fins. Anal and pelvic fins dusky.
</paragraph>
<paragraph id="8BED36F9206FFFEDD2415BDBFD0AFD1A" blockId="1.[113,753,481,664]" pageId="1" pageNumber="98">Preserved specimens are dark brown on body and white on ventrum. Black spots on body and the dusky stripe can only be seen in some specimens. The black blotches on dorsum also can be seen in preserved specimens. D1 black with a transparent margin, D2 dusky with dark brown stripes. Caudal, pectoral, anal and pelvic fin rays dusky.</paragraph>
</subSubSection>
<subSubSection id="C3486572206FFFEDD24158FBFAAEFD7A" pageId="1" pageNumber="98" type="discussion">
<paragraph id="8BED36F9206FFFEDD24158FBFD41FCFA" blockId="1.[113,753,705,888]" pageId="1" pageNumber="98">
<emphasis id="B926EAEB206FFFEDD24158FBFF6FFD5B" bold="true" box="[113,237,705,729]" italics="true" pageId="1" pageNumber="98">Remarks. –</emphasis>
Murdy (1989) recognised three species from the genus
<taxonomicName id="4C524D7A206FFFEDD28558DBFEFFFD7A" authorityName="Bleeker" authorityYear="1874" box="[181,381,737,760]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="B926EAEB206FFFEDD28558DBFEFFFD7A" box="[181,381,737,760]" italics="true" pageId="1" pageNumber="98">Periophthalmodon</emphasis>
</taxonomicName>
. They are distributed from Ganges Delta eastward to
<collectingCountry id="F3457669206FFFEDD361593BFE47FC9A" box="[337,453,769,792]" name="Indonesia" pageId="1" pageNumber="98">Indonesia</collectingCountry>
,
<collectingCountry id="F3457669206FFFEDD3E6593BFD39FC9A" box="[470,699,769,792]" name="Papua New Guinea" pageId="1" pageNumber="98">Papua New Guinea</collectingCountry>
and
<collectingCountry id="F3457669206FFFEDD241591BFF54FCBA" box="[113,214,801,824]" name="Australia" pageId="1" pageNumber="98">Australia</collectingCountry>
, but only one species,
<taxonomicName id="4C524D7A206FFFEDD3F2591BFDD5FCBA" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[450,599,801,824]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="schlosseri">
<emphasis id="B926EAEB206FFFEDD3F2591BFDD5FCBA" box="[450,599,801,824]" italics="true" pageId="1" pageNumber="98">Pn. schlosseri</emphasis>
</taxonomicName>
, was recorded from Peninsular
<collectingCountry id="F3457669206FFFEDD319597BFE0EFCDA" box="[297,396,833,856]" name="Malaysia" pageId="1" pageNumber="98">Malaysia</collectingCountry>
(
<bibRefCitation id="EFC34B08206FFFEDD3AC597BFDADFCDA" author="Cantor, T" box="[412,559,833,856]" pageId="1" pageNumber="98" pagination="1 - 1443" refId="ref1885" refString="Cantor, T. E., 1849. Catalogue of Malayan fishes. Journal of the Asiatic Society of Bengal, 18 (2): 1 - 1443." type="journal article" year="1849">Cantor, 1849</bibRefCitation>
;
<bibRefCitation id="EFC34B08206FFFEDD00C597BFD6EFCDA" author="Koumans, F" box="[572,748,833,856]" pageId="1" pageNumber="98" refId="ref1985" refString="Koumans, F. P., 1953. The fishes of the Indo-Australian Archipelago X. Leiden, E. J. Brill. xiii + 423 pp." type="book" year="1953">Koumans, 1953</bibRefCitation>
;
<bibRefCitation id="EFC34B08206FFFEDD241595BFF75FCFA" author="Berry, A" box="[113,247,865,888]" pageId="1" pageNumber="98" pagination="135 - 162" refId="ref1857" refString="Berry, A. J., 1972. The natural history of West Malaysian mangrove faunas. Malayan Nature Journal, 25: 135 - 162." type="journal article" year="1972">Berry, 1972</bibRefCitation>
;
<bibRefCitation id="EFC34B08206FFFEDD334595BFE17FCFA" author="Murdy, E" box="[260,405,865,888]" pageId="1" pageNumber="98" pagination="20 - 23" refId="ref2051" refString="Murdy, E. O., 1986. The lords of the Mudflats. Freshwater and Marine Aquarium, 9: 20 - 23." type="journal article" year="1986">Murdy, 1986</bibRefCitation>
, 1989;
<bibRefCitation id="EFC34B08206FFFEDD3D9595BFD35FCFA" author="Takita, T" box="[489,695,865,888]" pageId="1" pageNumber="98" pagination="131 - 138" refId="ref2159" refString="Takita, T., Agusnimar &amp; A. B. Ali, 1999. Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca. Ichthyological Research, 46 (2): 131 - 138." type="journal article" year="1999">Takita et al., 1999</bibRefCitation>
).
</paragraph>
<paragraph id="8BED36F9206FFFEDD241599BFDC5FBFA" blockId="1.[113,753,929,1144]" pageId="1" pageNumber="98">
The genus
<taxonomicName id="4C524D7A206FFFEDD2CC599BFE59FC3A" authorityName="Bleeker" authorityYear="1874" box="[252,475,929,952]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="B926EAEB206FFFEDD2CC599BFE59FC3A" box="[252,475,929,952]" italics="true" pageId="1" pageNumber="98">Periophthalmodon</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="4C524D7A206FFFEDD24159FBFE98FC5A" authorityName="Bloch &amp; J.G.Schneider" authorityYear="1801" box="[113,282,961,984]" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="genus">
<emphasis id="B926EAEB206FFFEDD24159FBFE98FC5A" box="[113,282,961,984]" italics="true" pageId="1" pageNumber="98">Periophthalmus</emphasis>
</taxonomicName>
by having a black stripe coursing posteriorly from the eye to the caudal peduncle, and by having two rows instead of one row of teeth in the upper jaw. These characters differentiate
<taxonomicName id="4C524D7A206FFFEDD3305E1BFE56FBBA" box="[256,468,1057,1080]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206FFFEDD3305E1BFE56FBBA" box="[256,468,1057,1080]" italics="true" pageId="1" pageNumber="98">Pn. septemradiatus</emphasis>
</taxonomicName>
from the similar looking
<emphasis id="B926EAEB206FFFEDD2415E7BFEF4FBDA" box="[113,374,1089,1112]" italics="true" pageId="1" pageNumber="98">
<taxonomicName id="4C524D7A206FFFEDD2415E7BFEF0FBDA" authorityName="Eggert" authorityYear="1935" box="[113,370,1089,1112]" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="weberi">Periophthalmus weberi</taxonomicName>
,
</emphasis>
which is known only from
<collectingCountry id="F3457669206FFFEDD09F5E7BFF79FBFA" name="Papua New Guinea" pageId="1" pageNumber="98">Papua New Guinea</collectingCountry>
and
<collectingCountry id="F3457669206FFFEDD3035E5BFE14FBFA" box="[307,406,1121,1144]" name="Australia" pageId="1" pageNumber="98">Australia</collectingCountry>
(Murdy, 1989).
</paragraph>
<paragraph id="8BED36F9206FFFEDD11D5AFBFA2AFE7A" blockId="1.[813,1452,193,504]" pageId="1" pageNumber="98">
The separated pelvic fins of
<taxonomicName id="4C524D7A206FFFEDD6665AFBFAA1FF5A" box="[1110,1315,193,216]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206FFFEDD6665AFBFAA1FF5A" box="[1110,1315,193,216]" italics="true" pageId="1" pageNumber="98">Pn. septemradiatus</emphasis>
</taxonomicName>
differentiate it from its congeners.
<taxonomicName id="4C524D7A206FFFEDD62A5ADBFB33FF7A" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[1050,1201,225,248]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="schlosseri">
<emphasis id="B926EAEB206FFFEDD62A5ADBFB33FF7A" box="[1050,1201,225,248]" italics="true" pageId="1" pageNumber="98">Pn. schlosseri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D7A206FFFEDD6D35ADBFAF1FF7A" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[1251,1395,225,248]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="freycineti">
<emphasis id="B926EAEB206FFFEDD6D35ADBFAF1FF7A" box="[1251,1395,225,248]" italics="true" pageId="1" pageNumber="98">Pn. freycineti</emphasis>
</taxonomicName>
have a strong frenum across their pelvic fins which are totally united to form a round disk. Unlike its congeners,
<taxonomicName id="4C524D7A206FFFEDD7B65B1BFC52FEDA" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206FFFEDD7B65B1BFC52FEDA" italics="true" pageId="1" pageNumber="98">Pn. septemradiatus</emphasis>
</taxonomicName>
is also sexually dimorphic. In mature males, the dorsal fins are contiguous, and the first dorsal fin is very reduced on females (
<figureCitation id="13692A7C206FFFEDD6135BBBFB09FE1A" box="[1059,1163,385,408]" captionStart="Fig" captionStartId="2.[136,172,2037,2058]" captionTargetBox="[292,1328,1307,2005]" captionTargetId="figure-7@2.[289,1330,1306,2020]" captionTargetPageId="2" captionText="Fig. 2. Sexual dimorphism on Periophthalmodon septemradiatus [KMZ-NYR000306KK (2)]. a) Male: the first dorsal fin is of moderate height with an elongated spine and contiguous with the second dorsal fins. b) Female: the first dorsal fin is poorly developed and not contiguous with the second dorsal fins." figureDoi="http://doi.org/10.5281/zenodo.13229255" httpUri="https://zenodo.org/record/13229255/files/figure.png" pageId="1" pageNumber="98">Figure 2</figureCitation>
). In addition, adult
<taxonomicName id="4C524D7A206FFFEDD7B65BBBFC62FE3A" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206FFFEDD7B65BBBFC62FE3A" italics="true" pageId="1" pageNumber="98">Pn. septemradiatus</emphasis>
</taxonomicName>
is smaller (SL: about 40.0-60.0 mm) compared to
<taxonomicName id="4C524D7A206FFFEDD1865BFBFBC9FE5A" baseAuthorityName="Pallas" baseAuthorityYear="1770" box="[950,1099,449,472]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="schlosseri">
<emphasis id="B926EAEB206FFFEDD1865BFBFBC9FE5A" box="[950,1099,449,472]" italics="true" pageId="1" pageNumber="98">Pn. schlosseri</emphasis>
</taxonomicName>
(SL: about 100.0-300.0 mm) and
<taxonomicName id="4C524D7A206FFFEDD11D5BDBFC42FE7A" baseAuthorityName="Valenciennes" baseAuthorityYear="1837" box="[813,960,481,504]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="freycineti">
<emphasis id="B926EAEB206FFFEDD11D5BDBFC42FE7A" box="[813,960,481,504]" italics="true" pageId="1" pageNumber="98">Pn. freycineti</emphasis>
</taxonomicName>
(SL: about 80.0-200.0 mm) (Murdy, 1989).
</paragraph>
<paragraph id="8BED36F9206FFFEDD11D581BFAAEFD7A" blockId="1.[813,1452,545,760]" pageId="1" pageNumber="98">
<taxonomicName id="4C524D7A206FFFEDD11D581BFB27FDBA" box="[813,1189,545,568]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206FFFEDD11D581BFB27FDBA" box="[813,1189,545,568]" italics="true" pageId="1" pageNumber="98">Periophthalmodon septemradiatus</emphasis>
</taxonomicName>
was found utilising the habitat that is far from the sea. Although under tidal influence, the water there was almost fresh, with a salinity that ranged between 1 and 3 ppt. In contrast,
<emphasis id="B926EAEB206FFFEDD71858BBFA27FD1A" box="[1320,1445,641,664]" italics="true" pageId="1" pageNumber="98">B. boddarti</emphasis>
,
<taxonomicName id="4C524D7A206FFFEDD11D589BFC60FD3A" authorityName="Bleeker" authorityYear="1853" box="[813,994,673,696]" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="chrysospilos">
<emphasis id="B926EAEB206FFFEDD11D589BFC60FD3A" box="[813,994,673,696]" italics="true" pageId="1" pageNumber="98">Ps. chrysospilos</emphasis>
</taxonomicName>
and most other mudskippers live in the littoral zone on the foreshore where the salinity ranges from 30-34 ppt. (Chew &amp; Ip, 1990;
<bibRefCitation id="EFC34B08206FFFEDD6B158DBFAA2FD7A" author="Clayton, D" box="[1153,1312,737,760]" pageId="1" pageNumber="98" pagination="507 - 577" refId="ref1961" refString="Clayton, D. A., 1993. Mudskippers. Oceanography and Marine Biology Annual Review, 31: 507 - 577." type="journal article" year="1993">Clayton, 1993</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3486572206FFFEED11D591BFC03FC4A" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98" type="biology_ecology">
<paragraph id="8BED36F9206FFFEED11D591BFC03FC4A" blockId="1.[813,1452,801,1144]" lastBlockId="2.[835,1474,849,968]" lastPageId="2" lastPageNumber="99" pageId="1" pageNumber="98">
<emphasis id="B926EAEB206FFFEDD11D591BFC3BFCBB" bold="true" box="[813,953,801,825]" italics="true" pageId="1" pageNumber="98">Ecology. –</emphasis>
The specimens of
<taxonomicName id="4C524D7A206FFFEDD6F9591BFC4FFCDA" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="98" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206FFFEDD6F9591BFC4FFCDA" italics="true" pageId="1" pageNumber="98">Periophthalmodon septemradiatus</emphasis>
</taxonomicName>
examined by us were obtained from the banks of the
<collectingRegion id="4996F81B206FFFEDD142595BFC50FCFA" box="[882,978,865,888]" country="Malaysia" name="Selangor" pageId="1" pageNumber="98">Selangor</collectingRegion>
River at Kampung Kuantan (
<geoCoordinate id="EE66503E206FFFEDD724595BFAEFFCFA" box="[1300,1389,865,888]" degrees="3" direction="north" minutes="11" orientation="latitude" pageId="1" pageNumber="98" precision="925" value="3.1833334">3 11’N</geoCoordinate>
,
<geoCoordinate id="EE66503E206FFFEDD749595BFCE6FC1A" degrees="101" direction="east" minutes="18" orientation="longitude" pageId="1" pageNumber="98" precision="925" value="101.3">101 18’E</geoCoordinate>
). This study area (
<figureCitation id="13692A7C206FFFEDD60D59BBFB18FC1A" box="[1085,1178,897,920]" captionStart="Fig" captionStartId="1.[114,150,2065,2086]" captionTargetBox="[211,1354,1207,2026]" captionTargetId="figure-255@1.[211,1357,1201,2030]" captionTargetPageId="1" captionText="Fig. 1. Map of Selangor River at Kampung Kuantan, Selangor, showing known localities where Periophthalmodon septemradiatus were collected (triangles). Inset: map of Peninsular Malaysia with an additional locality (round spot) in Johor." figureDoi="http://doi.org/10.5281/zenodo.13229253" httpUri="https://zenodo.org/record/13229253/files/figure.png" pageId="1" pageNumber="98">Figure 1</figureCitation>
) is located about
<quantity id="4CAA9B1C206FFFEDD75559BBFA29FC1A" box="[1381,1451,897,920]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="98" unit="km" value="15.0">15 km</quantity>
from the river mouth at Kuala
<collectingRegion id="4996F81B206FFFEDD6B2599BFB67FC3A" box="[1154,1253,929,952]" country="Malaysia" name="Selangor" pageId="1" pageNumber="98">Selangor</collectingRegion>
, and about
<quantity id="4CAA9B1C206FFFEDD756599BFA29FC3A" box="[1382,1451,929,952]" metricMagnitude="4" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="98" unit="km" value="64.0">64 km</quantity>
from northwest
<collectingRegion id="4996F81B206FFFEDD1ED59FBFB02FC5A" box="[989,1152,961,984]" country="Malaysia" name="Kuala Lumpur" pageId="1" pageNumber="98">Kuala Lumpur</collectingRegion>
. It was largely utilised for fishing, touristic and residential accommodation. A single species of mangrove,
<taxonomicName id="4C524D7A206FFFEDD62A5E3BFA85FB9A" box="[1050,1287,1025,1048]" class="Cephalopoda" family="Hoplitidae" genus="Sonneratia" kingdom="Animalia" pageId="1" pageNumber="98" phylum="Mollusca" rank="species" species="caseolaris">
<emphasis id="B926EAEB206FFFEDD62A5E3BFA85FB9A" box="[1050,1287,1025,1048]" italics="true" pageId="1" pageNumber="98">Sonneratia caseolaris</emphasis>
</taxonomicName>
was dominant. It formed a 10 metre wide belt above the mud banks, and ran parallel to the bund. Mature trees have brownish-grey bark and protruding pneumatophores that grow as high as
<quantity id="4CAA9B1C206CFFEED2B8596BFE88FCEA" box="[136,266,849,872]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="2" pageNumber="99" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10 to 15 cm</quantity>
. The mangrove forest was relatively open with slightly over 15 plants per 6 square metres. There were also some small tributaries draining through stands of
<taxonomicName id="4C524D7A206CFFEED0FD59ABFF67FC4A" class="Liliopsida" family="Arecaceae" genus="Nypa" kingdom="Plantae" order="Arecales" pageId="2" pageNumber="99" phylum="Tracheophyta" rank="species" species="fruticans">
<emphasis id="B926EAEB206CFFEED0FD59ABFF67FC4A" italics="true" pageId="2" pageNumber="99">Nypa fruticans</emphasis>
</taxonomicName>
and village orchard. The mud-banks were gradually sloping and the substrate was extremely soft with high water content.
<taxonomicName id="4C524D7A206CFFEED2C059CBFDEAFB8A" box="[240,616,1009,1032]" family="Gobiidae" genus="Periophthalmodon" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="99" phylum="Chordata" rank="species" species="septemradiatus">
<emphasis id="B926EAEB206CFFEED2C059CBFDEAFB8A" box="[240,616,1009,1032]" italics="true" pageId="2" pageNumber="99">Periophthalmodon septemradiatus</emphasis>
</taxonomicName>
was found on the mud banks and in the small tributaries along the upper reaches of the estuary. It was observed living sympatrically with the mudskippers
<taxonomicName id="4C524D7A206CFFEED3B05E6BFD54FBEA" baseAuthorityName="Hamilton" baseAuthorityYear="1822" box="[384,726,1105,1128]" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="99" phylum="Chordata" rank="species" species="novemradiatus">
<emphasis id="B926EAEB206CFFEED3B05E6BFD54FBEA" box="[384,726,1105,1128]" italics="true" pageId="2" pageNumber="99">Periophthalmus novemradiatus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D7A206CFFEED2B85E4AFE88FB05" authorityName="Eggert" authorityYear="1935" box="[136,266,1136,1159]" family="Gobiidae" genus="Periophthalmus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="99" phylum="Chordata" rank="species" species="gracilis">
<emphasis id="B926EAEB206CFFEED2B85E4AFE88FB05" box="[136,266,1136,1159]" italics="true" pageId="2" pageNumber="99">Ps. gracilis</emphasis>
</taxonomicName>
. Along the upper reaches of the estuary at Kampung Kuantan, the water was fresh (0 ppt) at low tide. During extreme high tides, the salinities recorded in the river ranged from 1 to 3 ppt.
<bibRefCitation id="EFC34B08206CFFEED3B05EEAFDC5FB65" author="Takita, T" box="[384,583,1232,1255]" pageId="2" pageNumber="99" pagination="131 - 138" refId="ref2159" refString="Takita, T., Agusnimar &amp; A. B. Ali, 1999. Distribution and habitat requirements of oxudercine gobies (Gobiidae: Oxudercinae) along the Straits of Malacca. Ichthyological Research, 46 (2): 131 - 138." type="journal article" year="1999">Takita et al. (1999)</bibRefCitation>
found this species in a small stream at Tebing Tinggi,
<collectingRegion id="4996F81B206CFFEED69F596BFA8BFCEA" box="[1199,1289,849,872]" country="Indonesia" name="Sumatera Utara" pageId="2" pageNumber="99">Sumatra</collectingRegion>
, in a place similar to the present study area, but they did not report on the water salinity or the presence of sympatric mudskipper species there.
</paragraph>
</subSubSection>
</treatment>
</document>
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