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<document id="0C9A18C7C77ADBBBA3802B1805FC66AE" ID-DOI="10.5281/zenodo.207370" ID-GBIF-Dataset="a7ff0ca7-8f31-4e46-8f00-fe8be5a64b18" ID-ISSN="1175-5326" ID-Zenodo-Dep="207370" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460328686883" checkinUser="plazi" docAuthor="Liu, Xingyue, Hayashi, Fumio &amp; Yang, Ding" docDate="2011" docId="03FB87B2FF87FF9FFF4C8BF1248BF8D4" docLanguage="en" docName="zt02981p035.pdf" docOrigin="Zootaxa 2981" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ctenochauliodes khasianus" docType="treatment" docVersion="9" lastPageNumber="27" masterDocId="FFC2FFCAFF85FF9BFFDB882B255CFFBF" masterDocTitle="Taxonomic notes and updated phylogeny of the fishfly genus Ctenochauliodes van der Weele (Megaloptera: Corydalidae)" masterLastPageNumber="35" masterPageNumber="23" pageNumber="25" updateTime="1698258261109" updateUser="plazi">
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<mods:title id="A188D12AD1D1B92DECDBF534B52ECB2E">Taxonomic notes and updated phylogeny of the fishfly genus Ctenochauliodes van der Weele (Megaloptera: Corydalidae)</mods:title>
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<mods:namePart id="093F72F3B5F3B1820C0781CE0E106326">Liu, Xingyue</mods:namePart>
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<mods:namePart id="320757AF9D516DCDFB163F82A9BC4548">Hayashi, Fumio</mods:namePart>
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<treatment id="03FB87B2FF87FF9FFF4C8BF1248BF8D4" ID-DOI="http://doi.org/10.5281/zenodo.5689718" ID-GBIF-Taxon="119404193" ID-Zenodo-Dep="5689718" LSID="urn:lsid:plazi:treatment:03FB87B2FF87FF9FFF4C8BF1248BF8D4" httpUri="http://treatment.plazi.org/id/03FB87B2FF87FF9FFF4C8BF1248BF8D4" lastPageId="4" lastPageNumber="27" pageId="2" pageNumber="25">
<subSubSection id="C348652FFF87FF99FF4C8BF12159FBE4" pageId="2" pageNumber="25" type="nomenclature">
<paragraph id="8BED36A4FF87FF99FF4C8BF12618FC4B" blockId="2.[151,836,986,1045]" box="[151,836,986,1012]" pageId="2" pageNumber="25">
<heading id="D0A581C8FF87FF99FF4C8BF12618FC4B" bold="true" box="[151,836,986,1012]" fontSize="11" level="1" pageId="2" pageNumber="25" reason="1">
<emphasis id="B926EAB6FF87FF99FF4C8BF12618FC4B" bold="true" box="[151,836,986,1012]" pageId="2" pageNumber="25">
<taxonomicName id="4C524D27FF87FF99FF4C8BF124B8FC4B" ID-CoL="325DN" box="[151,484,986,1012]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="B926EAB6FF87FF99FF4C8BF124B8FC4B" bold="true" box="[151,484,986,1012]" italics="true" pageId="2" pageNumber="25">Ctenochauliodes khasianus</emphasis>
</taxonomicName>
(van der Weele), comb. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36A4FF87FF99FF4C8BD62404FBAA" blockId="2.[151,836,986,1045]" box="[151,344,1021,1045]" pageId="2" pageNumber="25">
(
<figureCitation id="13692A21FF87FF99FF458BD62455FBAA" box="[158,265,1021,1045]" captionStart="FIGURES 1 3" captionStartId="3.[151,264,1949,1972]" captionTargetBox="[190,1389,204,1925]" captionTargetId="figure@3.[190,1397,179,1941]" captionTargetPageId="3" captionText="FIGURES 1 3. Habitus images of Ctenochauliodes spp. 1. C. khasianus (van der Weele), holotype male; 2. C. nigrovenosus (van der Weele), lectotype female; 3. C. nigrovenosus (van der Weele), lectotype male of C. friedrichi Navás. Scale bars = 5.0 mm." httpUri="https://zenodo.org/record/207371/files/figure.png" pageId="2" pageNumber="25">Figures 1</figureCitation>
,
<figureCitation id="13692A21FF87FF99FECC8BD6240CFBAA" box="[279,336,1021,1045]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1449,1472]" captionTargetBox="[180,1390,207,1408]" captionTargetId="figure@5.[177,1410,193,1428]" captionTargetPageId="5" captionText="FIGURES 6 10. Ctenochauliodes khasianus (van der Weele). 6. Male genitalia, lateral view; 7. Male tenth tergum, dorsal view; 8. Male tenth tergum, ventral view; 9. Male tenth gonocoxite, ventral view; 10. Sclerite beneath anus, ventral view. Gx 10: tenth gonocoxite; S 9: ninth sternum; sa: sclerite beneath anus; T 9 10: ninth to tenth tergum. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/207373/files/figure.png" pageId="2" pageNumber="25">610</figureCitation>
)
</paragraph>
<paragraph id="8BED36A4FF87FF99FF4C8C6E2159FBE4" blockId="2.[151,1029,1093,1115]" box="[151,1029,1093,1115]" pageId="2" pageNumber="25">
<treatmentCitationGroup id="AB42118AFF87FF99FF4C8C6E2159FBE4" box="[151,1029,1093,1115]" pageId="2" pageNumber="25">
<taxonomicName id="4C524D27FF87FF99FF4C8C6E24C5FBE4" ID-CoL="46CP9" box="[151,409,1093,1115]" class="Insecta" family="Corydalidae" genus="Neochauliodes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="B926EAB6FF87FF99FF4C8C6E24C5FBE4" box="[151,409,1093,1115]" italics="true" pageId="2" pageNumber="25">Neochauliodes khasianus</emphasis>
</taxonomicName>
van der
<treatmentCitation id="0AF310B5FF87FF99FE2B8C6E27F9FBE4" author="Weele" box="[496,677,1093,1115]" page="259" pageId="2" pageNumber="25" year="1909">
<bibRefCitation id="EFC34B55FF87FF99FE2B8C6E272EFBE4" author="Weele" box="[496,626,1093,1115]" pageId="2" pageNumber="25" refString="Weele, H. W. van der (1909) New genera and species of Megaloptera Latr. Notes from the Leyden Museum, 30, 249 - 264." type="journal article" year="1909">Weele, 1909</bibRefCitation>
: 259
</treatmentCitation>
.
<typeStatus id="54E98806FF87FF99FD758C6D27BCFBE4" box="[686,736,1094,1115]" pageId="2" pageNumber="25">Type</typeStatus>
locality:
<collectingCountry id="F3457634FF87FF99FC9B8C6E2628FBE4" box="[832,884,1093,1115]" name="India" pageId="2" pageNumber="25">India</collectingCountry>
(Khasi Hills).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C348652FFF87FF99FF4C8CAC27D3FAB3" pageId="2" pageNumber="25" type="diagnosis">
<paragraph id="8BED36A4FF87FF99FF4C8CAC27D3FAB3" blockId="2.[151,1436,1159,2015]" pageId="2" pageNumber="25">
<emphasis id="B926EAB6FF87FF99FF4C8CAC244DFB1F" bold="true" box="[151,273,1159,1184]" pageId="2" pageNumber="25">Diagnosis.</emphasis>
The forewings of this species are densely marked by numerous small brownish spots. The male tenth tergum is twisted apically forming a triangular, somewhat flattened lobe. The male tenth gonocoxite is composed of a pair of bluntly tapered lobes that form nearly equal in length and subtriangular on posterior half of right lobe. Lateral lobes beneath anus feebly developed.
</paragraph>
</subSubSection>
<subSubSection id="C348652FFF87FF99FF1E8D3C24C7F8CD" pageId="2" pageNumber="25" type="description">
<paragraph id="8BED36A4FF87FF99FF1E8D3C21E0FA8F" blockId="2.[151,1436,1159,2015]" box="[197,1212,1303,1328]" pageId="2" pageNumber="25">
<emphasis id="B926EAB6FF87FF99FF1E8D3C240BFA8F" bold="true" box="[197,343,1303,1328]" pageId="2" pageNumber="25">Description.</emphasis>
Male. Body length
<quantity id="4CAA9B41FF87FF99FDEE8D3C27DBFA8F" box="[565,647,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="25" unit="mm" value="10.0">10 mm</quantity>
; forewing length
<quantity id="4CAA9B41FF87FF99FC908D3C26C1FA8F" box="[843,925,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="25" unit="mm" value="22.0">22 mm</quantity>
, hindwing length
<quantity id="4CAA9B41FF87FF99FBBD8D3C21E5FA8F" box="[1126,1209,1303,1328]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="25" unit="mm" value="20.0">20 mm</quantity>
.
</paragraph>
<paragraph id="8BED36A4FF87FF99FF1E8D17266AFAC8" blockId="2.[151,1436,1159,2015]" pageId="2" pageNumber="25">Head blackish brown with frons black. Compound eyes brown, ocelli yellow. Antenna blackish brown. Mouthparts blackish brown, mandibles with apices reddish brown.</paragraph>
<paragraph id="8BED36A4FF87FF99FF1E8DAF219FF993" blockId="2.[151,1436,1159,2015]" pageId="2" pageNumber="25">Thorax blackish brown. Legs brown, with short dense yellowish setae; tarsal claws reddish brown. Wings hyaline, slightly tinged pale smoky brown, with a brownish marking over each black nygmata. Forewings with proximal half of costal area pale brown and brownish markings on pterostigmatic areas, and with numerous indistinct small brownish spots throughout. Hindwings much paler than forewings, immaculate except for black nygmata. Rs four to five-branched; three crossveins between R1 and Rs; M two-branched; 1A two-branched.</paragraph>
<paragraph id="8BED36A4FF87FF99FF1E8E1124F2F8F0" blockId="2.[151,1436,1159,2015]" pageId="2" pageNumber="25">
Abdomen blackish brown with venter much paler. Tenth tergum (
<figureCitation id="13692A21FF87FF99FC408E112159F9ED" box="[923,1029,1594,1618]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1449,1472]" captionTargetBox="[180,1390,207,1408]" captionTargetId="figure@5.[177,1410,193,1428]" captionTargetPageId="5" captionText="FIGURES 6 10. Ctenochauliodes khasianus (van der Weele). 6. Male genitalia, lateral view; 7. Male tenth tergum, dorsal view; 8. Male tenth tergum, ventral view; 9. Male tenth gonocoxite, ventral view; 10. Sclerite beneath anus, ventral view. Gx 10: tenth gonocoxite; S 9: ninth sternum; sa: sclerite beneath anus; T 9 10: ninth to tenth tergum. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/207373/files/figure.png" pageId="2" pageNumber="25">Figs. 68</figureCitation>
) claviform, with apex distinctly narrowed and strongly twisted anteroventrad. Tenth gonocoxite (
<figureCitation id="13692A21FF87FF99FC918E7426EAF9C8" box="[842,950,1631,1655]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1449,1472]" captionTargetBox="[180,1390,207,1408]" captionTargetId="figure@5.[177,1410,193,1428]" captionTargetPageId="5" captionText="FIGURES 6 10. Ctenochauliodes khasianus (van der Weele). 6. Male genitalia, lateral view; 7. Male tenth tergum, dorsal view; 8. Male tenth tergum, ventral view; 9. Male tenth gonocoxite, ventral view; 10. Sclerite beneath anus, ventral view. Gx 10: tenth gonocoxite; S 9: ninth sternum; sa: sclerite beneath anus; T 9 10: ninth to tenth tergum. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/207373/files/figure.png" pageId="2" pageNumber="25">Figs. 6, 9</figureCitation>
) strongly sclerotized, median plate divided into a pair of asymmetric lobes by a narrow incision; in lateral view median plate distinctly broadened and rounded dorsad and bearing short setae distally; in ventral view lateral arms suboval, median plate concave anteriorly forming a triangular emargination; lobes formed by narrow cleft nearly equal in length but asymmetrical, left lobe slightly incurved on medial surface and rounded apically, right lobe wider, posterior half strongly narrowed into a subtriangular apex. Sclerite ventrally to anus (
<figureCitation id="13692A21FF87FF99FD798F392640F895" box="[674,796,1810,1834]" captionStart="FIGURES 6 10" captionStartId="5.[151,264,1449,1472]" captionTargetBox="[180,1390,207,1408]" captionTargetId="figure@5.[177,1410,193,1428]" captionTargetPageId="5" captionText="FIGURES 6 10. Ctenochauliodes khasianus (van der Weele). 6. Male genitalia, lateral view; 7. Male tenth tergum, dorsal view; 8. Male tenth tergum, ventral view; 9. Male tenth gonocoxite, ventral view; 10. Sclerite beneath anus, ventral view. Gx 10: tenth gonocoxite; S 9: ninth sternum; sa: sclerite beneath anus; T 9 10: ninth to tenth tergum. Scale bars = 1.0 mm." httpUri="https://zenodo.org/record/207373/files/figure.png" pageId="2" pageNumber="25">Figs. 6, 10</figureCitation>
) indistinct except for tuberculate lateral lobes which are posterolaterally directed.
</paragraph>
<paragraph id="8BED36A4FF87FF99FF1E8F7124C7F8CD" blockId="2.[151,1436,1159,2015]" box="[197,411,1882,1906]" pageId="2" pageNumber="25">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="C348652FFF87FF99FF1E8F54249AF860" pageId="2" pageNumber="25" type="materials_examined">
<paragraph id="8BED36A4FF87FF99FF1E8F54249AF860" blockId="2.[151,1436,1159,2015]" pageId="2" pageNumber="25">
<emphasis id="B926EAB6FF87FF99FF1E8F542426F828" bold="true" box="[197,378,1918,1943]" pageId="2" pageNumber="25">
<typeStatus id="54E98806FF87FF99FF1E8F54245CF828" box="[197,256,1919,1943]" pageId="2" pageNumber="25">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54E98806FF87FF99FE538F5424ACF828" box="[392,496,1919,1943]" pageId="2" pageNumber="25" type="holotype">Holotype</typeStatus>
3, “Khasis [= Khasi Hills,
<geoCoordinate id="EE665063FF87FF99FC888F5426F3F828" box="[851,943,1919,1943]" direction="east" orientation="longitude" pageId="2" pageNumber="25" precision="925" value="91.63333">91°38'E</geoCoordinate>
,
<geoCoordinate id="EE665063FF87FF99FC1A8F54217CF828" box="[961,1056,1919,1943]" direction="north" orientation="latitude" pageId="2" pageNumber="25" precision="925" value="25.583334">25°35'N</geoCoordinate>
, in Meghalaya State]. Nat[ive]. Coll[ector]./Cat[alog] No.1./3/
<taxonomicName id="4C524D27FF87FF99FE228F8826F8F805" authority="Type 1908" authorityName="Type" authorityYear="1908" box="[505,932,1954,1979]" class="Insecta" family="Corydalidae" genus="Neochauliodes" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="25" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="B926EAB6FF87FF99FE228F882640F804" box="[505,796,1955,1979]" italics="true" pageId="2" pageNumber="25">Neochauliodes khasianus</emphasis>
<typeStatus id="54E98806FF87FF99FCF38F88263CF804" box="[808,864,1955,1979]" pageId="2" pageNumber="25">Type</typeStatus>
1908
</taxonomicName>
Weele/
<typeStatus id="54E98806FF87FF99FBD98F892160F805" box="[1026,1084,1954,1978]" pageId="2" pageNumber="25">Type</typeStatus>
/Coll[ection]. v[an].d[er].Weele leg. 1907 No.6.” (
<collectionCode id="ED43AE61FF87FF99FE848FEC24E6F860" LSID="urn:lsid:biocol.org:col:34992" box="[351,442,1991,2015]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="2" pageNumber="25">RMNH</collectionCode>
).
</paragraph>
</subSubSection>
<caption id="DF2D662CFF86FF98FF4C8FB62599F84E" httpUri="https://zenodo.org/record/207371/files/figure.png" pageId="3" pageNumber="26" targetBox="[190,1389,204,1925]" targetPageId="3">
<paragraph id="8BED36A4FF86FF98FF4C8FB62599F84E" blockId="3.[151,1436,1949,2033]" pageId="3" pageNumber="26">
<emphasis id="B926EAB6FF86FF98FF4C8FB62468F80C" bold="true" box="[151,308,1949,1972]" pageId="3" pageNumber="26">FIGURES 13</emphasis>
. Habitus images of
<taxonomicName id="4C524D27FF86FF98FDDF8FB527F2F80B" box="[516,686,1950,1972]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB6FF86FF98FDDF8FB527F2F80B" box="[516,686,1950,1972]" italics="true" pageId="3" pageNumber="26">Ctenochauliodes</emphasis>
</taxonomicName>
spp. 1.
<taxonomicName id="4C524D27FF86FF98FCDA8FB426D9F80B" box="[769,901,1950,1972]" class="Insecta" family="Corydalidae" genus="Neochauliodes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="B926EAB6FF86FF98FCDA8FB426D9F80B" box="[769,901,1950,1972]" italics="true" pageId="3" pageNumber="26">C. khasianus</emphasis>
</taxonomicName>
(van der Weele), holotype male; 2.
<taxonomicName id="4C524D27FF86FF98FB2C8FB420C7F80B" box="[1271,1435,1951,1972]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="nigrovenosus">
<emphasis id="B926EAB6FF86FF98FB2C8FB420C7F80B" box="[1271,1435,1951,1972]" italics="true" pageId="3" pageNumber="26">C. nigrovenosus</emphasis>
</taxonomicName>
(van der Weele), lectotype female; 3.
<taxonomicName id="4C524D27FF86FF98FDCD8F9627E6F86D" box="[534,698,1981,2002]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="nigrovenosus">
<emphasis id="B926EAB6FF86FF98FDCD8F9627E6F86D" box="[534,698,1981,2002]" italics="true" pageId="3" pageNumber="26">C. nigrovenosus</emphasis>
</taxonomicName>
(van der Weele), lectotype male of
<taxonomicName id="4C524D27FF86FF98FBF38F96206CF86D" authority="Navas. Scale" authorityName="Navas. Scale" box="[1064,1328,1980,2002]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="26" phylum="Arthropoda" rank="species" species="friedrichi">
<emphasis id="B926EAB6FF86FF98FBF38F9621FAF86D" box="[1064,1190,1980,2002]" italics="true" pageId="3" pageNumber="26">C. friedrichi</emphasis>
Navás. Scale
</taxonomicName>
bars = 5.0 mm.
</paragraph>
</caption>
<caption id="DF2D662CFF81FF9FFF4C8D4E2629FA25" httpUri="https://zenodo.org/record/207372/files/figure.png" pageId="4" pageNumber="27" targetBox="[157,1436,201,1334]" targetPageId="4">
<paragraph id="8BED36A4FF81FF9FFF4C8D4E2629FA25" blockId="4.[151,1436,1381,1435]" pageId="4" pageNumber="27">
<emphasis id="B926EAB6FF81FF9FFF4C8D4E2468FAC4" bold="true" box="[151,308,1381,1404]" pageId="4" pageNumber="27">FIGURES 45</emphasis>
. Habitus images of
<taxonomicName id="4C524D27FF81FF9FFDDE8D4D27F3FAC3" box="[517,687,1382,1404]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB6FF81FF9FFDDE8D4D27F3FAC3" box="[517,687,1382,1404]" italics="true" pageId="4" pageNumber="27">Ctenochauliodes</emphasis>
</taxonomicName>
spp. 4.
<taxonomicName id="4C524D27FF81FF9FFCD88D4C26F4FAC3" box="[771,936,1383,1404]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="nigrovenosus">
<emphasis id="B926EAB6FF81FF9FFCD88D4C26F4FAC3" box="[771,936,1383,1404]" italics="true" pageId="4" pageNumber="27">C. nigrovenosus</emphasis>
</taxonomicName>
(van der Weele), holotype male of
<taxonomicName id="4C524D27FF81FF9FFACC8D4C25A5FA25" authority="Kimmins" authorityName="Kimmins" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="forcipatus">
<emphasis id="B926EAB6FF81FF9FFACC8D4C20C0FAC3" box="[1303,1436,1382,1404]" italics="true" pageId="4" pageNumber="27">C. forcipatus</emphasis>
Kimmins
</taxonomicName>
; 5.
<taxonomicName id="4C524D27FF81FF9FFEC78DAE24C1FA24" box="[284,413,1413,1435]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="stigmosus" status="sp. nov.">
<emphasis id="B926EAB6FF81FF9FFEC78DAE24C1FA24" box="[284,413,1413,1435]" italics="true" pageId="4" pageNumber="27">C. stigmosus</emphasis>
</taxonomicName>
<emphasis id="B926EAB6FF81FF9FFE788DAE24AFFA25" bold="true" box="[419,499,1413,1434]" pageId="4" pageNumber="27">
<taxonomicNameLabel id="A21557CDFF81FF9FFE788DAE24AFFA25" box="[419,499,1413,1434]" pageId="4" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. Scale bars = 5.0 mm.
</paragraph>
</caption>
<subSubSection id="C348652FFF81FF9FFF1E8DED2760FA60" box="[197,572,1478,1503]" pageId="4" pageNumber="27" type="distribution">
<paragraph id="8BED36A4FF81FF9FFF1E8DED2760FA60" blockId="4.[151,1436,1478,1899]" box="[197,572,1478,1503]" pageId="4" pageNumber="27">
<emphasis id="B926EAB6FF81FF9FFF1E8DED2402FA60" bold="true" box="[197,350,1478,1503]" pageId="4" pageNumber="27">Distribution.</emphasis>
<collectingCountry id="F3457634FF81FF9FFEBE8DEC24C2FA60" box="[357,414,1479,1503]" name="India" pageId="4" pageNumber="27">India</collectingCountry>
(Meghalaya).
</paragraph>
</subSubSection>
<subSubSection id="C348652FFF81FF9FFF1E8DC7248BF8D4" pageId="4" pageNumber="27" type="discussion">
<paragraph id="8BED36A4FF81FF9FFF1E8DC7248BF8D4" blockId="4.[151,1436,1478,1899]" pageId="4" pageNumber="27">
<emphasis id="B926EAB6FF81FF9FFF1E8DC72465F9BB" bold="true" box="[197,313,1516,1540]" pageId="4" pageNumber="27">Remarks.</emphasis>
The taxonomic status of this species has remained obscure for more than a century. The male
<typeStatus id="54E98806FF81FF9FFABA8DC0259BF997" pageId="4" pageNumber="27" type="holotype">holotype</typeStatus>
of this species was first identified by van der
<bibRefCitation id="EFC34B55FF81FF9FFD158E242639F998" author="Weele" box="[718,869,1551,1575]" pageId="4" pageNumber="27" refString="Weele, H. W. van der (1907) Notizen uber Sialiden und Beschreibung einiger neuen Arten. Notes from the Leyden Museum, 28, 227 - 264." type="journal article" year="1907">Weele (1907)</bibRefCitation>
as
<taxonomicName id="4C524D27FF81FF9FFC508E242065F998" authority="McLachlan, 1867" authorityName="McLachlan" authorityYear="1867" box="[907,1337,1551,1575]" class="Insecta" family="Corydalidae" genus="Chauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="pusillus">
<emphasis id="B926EAB6FF81FF9FFC508E242135F998" box="[907,1129,1551,1575]" italics="true" pageId="4" pageNumber="27">Chauliodes pusillus</emphasis>
<bibRefCitation id="EFC34B55FF81FF9FFBAA8E242065F998" author="McLachlan" box="[1137,1337,1551,1575]" pageId="4" pageNumber="27" refString="McLachlan, R. (1867) New genera and species, etc., of Neuropterous insects; and a revision of Mr F. Walker's British Museum Catalogue, part ii (1853), as far as the end of the genus Myrmeleon. Journal of the Linnean Society (Zoology), 9, 230 - 281." type="journal article" year="1867">McLachlan, 1867</bibRefCitation>
</taxonomicName>
, but was moved to the South African genus
<taxonomicName id="4C524D27FF81FF9FFDC28E182791F9F4" box="[537,717,1587,1611]" class="Insecta" family="Corydalidae" genus="Platychauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB6FF81FF9FFDC28E182791F9F4" box="[537,717,1587,1611]" italics="true" pageId="4" pageNumber="27">Platychauliodes</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC34B55FF81FF9FFD2D8E1826F9F9F4" author="Barnard" box="[758,933,1587,1611]" pageId="4" pageNumber="27" refString="Barnard, K. H. (1931) The Cape alder-flies (Neuroptera, Megaloptera). Transactions of the Royal Society of South Africa, 19, 169 - 184." type="journal article" year="1931">Barnard (1931)</bibRefCitation>
. Later, van der
<bibRefCitation id="EFC34B55FF81FF9FFB948E1821B9F9F4" author="Weele" box="[1103,1253,1587,1611]" pageId="4" pageNumber="27" refString="Weele, H. W. van der (1909) New genera and species of Megaloptera Latr. Notes from the Leyden Museum, 30, 249 - 264." type="journal article" year="1909">Weele (1909)</bibRefCitation>
used this Indian specimen as the
<typeStatus id="54E98806FF81FF9FFE8F8E7C24EAF9D0" box="[340,438,1623,1647]" pageId="4" pageNumber="27" type="holotype">holotype</typeStatus>
for a new species,
<taxonomicName id="4C524D27FF81FF9FFD4D8E7C26EAF9D0" box="[662,950,1623,1647]" class="Insecta" family="Corydalidae" genus="Neochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="khasianus">
<emphasis id="B926EAB6FF81FF9FFD4D8E7C26EAF9D0" box="[662,950,1623,1647]" italics="true" pageId="4" pageNumber="27">Neochauliodes khasianus</emphasis>
</taxonomicName>
van der Weele. However, we realized that this species resembles species of
<taxonomicName id="4C524D27FF81FF9FFDD18E50279AF92C" box="[522,710,1659,1683]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB6FF81FF9FFDD18E50279AF92C" box="[522,710,1659,1683]" italics="true" pageId="4" pageNumber="27">Ctenochauliodes</emphasis>
</taxonomicName>
with its narrowed wings and strongly pectinate male antennae as shown in the habitus photo of the
<typeStatus id="54E98806FF81FF9FFDF18EB427D0F908" box="[554,652,1695,1719]" pageId="4" pageNumber="27" type="holotype">holotype</typeStatus>
(van der
<bibRefCitation id="EFC34B55FF81FF9FFCDA8EB426D0F908" author="Weele" box="[769,908,1695,1719]" pageId="4" pageNumber="27" refString="Weele, H. W. van der (1907) Notizen uber Sialiden und Beschreibung einiger neuen Arten. Notes from the Leyden Museum, 28, 227 - 264." type="journal article" year="1907">Weele 1907</bibRefCitation>
: Plate 4,
<figureCitation id="13692A21FF81FF9FFC278EB4211FF908" box="[1020,1091,1695,1719]" captionStart="FIGURES 4 5" captionStartId="4.[151,264,1381,1404]" captionTargetBox="[157,1436,201,1334]" captionTargetId="figure@4.[151,1436,193,1360]" captionTargetPageId="4" captionText="FIGURES 4 5. Habitus images of Ctenochauliodes spp. 4. C. nigrovenosus (van der Weele), holotype male of C. forcipatus Kimmins; 5. C. stigmosus sp. nov., holotype male. Scale bars = 5.0 mm." httpUri="https://zenodo.org/record/207372/files/figure.png" pageId="4" pageNumber="27">Fig. 4</figureCitation>
). Recently, we confirmed our assumption that this species truly belongs to
<taxonomicName id="4C524D27FF81FF9FFD518EE8261AF964" box="[650,838,1731,1755]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB6FF81FF9FFD518EE8261AF964" box="[650,838,1731,1755]" italics="true" pageId="4" pageNumber="27">Ctenochauliodes</emphasis>
</taxonomicName>
when we re-examined the
<typeStatus id="54E98806FF81FF9FFBA98EE82188F964" box="[1138,1236,1731,1755]" pageId="4" pageNumber="27" type="holotype">holotype</typeStatus>
at the RMNH. Its similarly shaped male tenth gonocoxite appears to place it in close relationships with
<taxonomicName id="4C524D27FF81FF9FFB868EC32048F940" box="[1117,1300,1768,1791]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="nigrovenosus">
<emphasis id="B926EAB6FF81FF9FFB868EC32048F940" box="[1117,1300,1768,1791]" italics="true" pageId="4" pageNumber="27">C. nigrovenosus</emphasis>
</taxonomicName>
from
<collectingCountry id="F3457634FF81FF9FFA828ECC20C0F940" box="[1369,1436,1767,1791]" name="China" pageId="4" pageNumber="27">China</collectingCountry>
and
<collectingCountry id="F3457634FF81FF9FFF1C8F202476F89C" box="[199,298,1803,1827]" name="Vietnam" pageId="4" pageNumber="27">Vietnam</collectingCountry>
. However, it can be distinguished from the latter species by the male tenth gonocoxite with the right lobe being rapidly narrowed at its apex. In
<taxonomicName id="4C524D27FF81FF9FFD518F1B261CF8F8" box="[650,832,1840,1863]" class="Insecta" family="Corydalidae" genus="Ctenochauliodes" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="27" phylum="Arthropoda" rank="species" species="nigrovenosus">
<emphasis id="B926EAB6FF81FF9FFD518F1B261CF8F8" box="[650,832,1840,1863]" italics="true" pageId="4" pageNumber="27">C. nigrovenosus</emphasis>
</taxonomicName>
, the right lobe of the male tenth gonocoxite is more gradually narrowed apically.
</paragraph>
</subSubSection>
</treatment>
</document>