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<mods:title id="6B5D830DFB3DBDF60CA77F686AAF037D">Bombardier beetles of the genus Pheropsophus Solier 1833 (Carabidae: Brachininae: Brachinini) from Indian subcontinent</mods:title>
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<emphasis id="B926EA86FFF577534AFEFC7934D9FDDD" bold="true" box="[151,440,980,1007]" italics="true" pageId="13" pageNumber="78">Pheropsophus occipitalis</emphasis>
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<bibRefCitation id="EFC34B65FFF577534B9CFC7937A6FDDD" author="Macleay, W. S." box="[501,711,980,1007]" pageId="13" pageNumber="78" refId="ref20195" refString="Macleay, W. S. (1825) Number I. of Annulosa Javanica, or an attempt to illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfield, M. D. F. L. &amp; G. S. and deposited by him in the museum of the honourable East-India Company. Kingsbury, Parbury, and Allen, London, xii + 50 pp., 1 pls." type="book" year="1825">MACLEAY, 1825</bibRefCitation>
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(
<figureCitation id="13692A11FFF577534AC9FC54359DFA21" box="[160,252,1017,1043]" captionStart="FIGURE 2" captionStartId="11.[151,250,1867,1892]" captionTargetBox="[220,1413,191,1818]" captionTargetId="figure@11.[204,1425,179,1827]" captionTargetPageId="11" captionText="FIGURE 2. Dorsal habitus of Pheropsophus from India (cont.)A) Pheropsophus lissoderus (Holotype, Tibet, MNHN), B) P. nanodes (Verified specimen, S.E.Tibet, BMNH), C) P. picicollis (Syntype, Siam (now Thailand) MNHN), D) P. nigricollis (Holotype, Bangalore, BMNH), E) P. occipitalis (Verified specimen, China, BMNH), F) P. prophylax (Holotype, Manipur, BMNH), G) P. scythropus (Holotype, Gopaldhara, BMNH), H) P. scythropus dissolutus (Laos, BMNH)" figureDoi="http://doi.org/10.5281/zenodo.3993615" httpUri="https://zenodo.org/record/3993615/files/figure.png" pageId="13" pageNumber="78">Fig. 2 E</figureCitation>
,
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<taxonomicName id="4C524D17FFF577534ACCFB90377EFA67" ID-CoL="MT63" authority="DEJEAN, 1825" authorityName="DEJEAN" authorityYear="1825" box="[165,543,1084,1109]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="B926EA86FFF577534ACCFB90341BFA67" box="[165,378,1085,1109]" italics="true" pageId="13" pageNumber="78">Brachinus ambiguus</emphasis>
<bibRefCitation id="EFC34B65FFF577534BE9FB90377EFA67" author="Dejean, P. F. M. A." box="[384,543,1084,1109]" pageId="13" pageNumber="78" refId="ref18945" refString="Dejean, P. F. M. A. (1825) Species general des coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp." type="book" year="1825">DEJEAN, 1825</bibRefCitation>
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<emphasis id="B926EA86FFF577534AFEFBF135C4FA46" box="[151,165,1116,1140]" italics="true" pageId="13" pageNumber="78">=</emphasis>
<taxonomicName id="4C524D17FFF577534ACCFBF13742FA46" ID-CoL="68QDR" authority="DEJEAN, 1825" authorityName="DEJEAN" authorityYear="1825" box="[165,547,1115,1140]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="fuscicollis">
<emphasis id="B926EA86FFF577534ACCFBF1341FFA46" box="[165,382,1116,1140]" italics="true" pageId="13" pageNumber="78">Brachinus fuscicollis</emphasis>
<bibRefCitation id="EFC34B65FFF577534BEDFBF13742FA46" author="Dejean, P. F. M. A." box="[388,547,1115,1140]" pageId="13" pageNumber="78" refId="ref18945" refString="Dejean, P. F. M. A. (1825) Species general des coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp." type="book" year="1825">DEJEAN, 1825</bibRefCitation>
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<emphasis id="B926EA86FFF577534AFEFBD735C4FAA0" box="[151,165,1146,1170]" italics="true" pageId="13" pageNumber="78">=</emphasis>
<taxonomicName id="4C524D17FFF577534ACCFBD7374CFAA1" ID-CoL="MTBK" authority="DEJEAN, 1825" authorityName="DEJEAN" authorityYear="1825" box="[165,557,1146,1171]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="B926EA86FFF577534ACCFBD734E9FAA0" box="[165,392,1146,1170]" italics="true" pageId="13" pageNumber="78">Brachinus interruptus</emphasis>
<bibRefCitation id="EFC34B65FFF577534BE7FBD6374CFAA1" author="Dejean, P. F. M. A." box="[398,557,1146,1171]" pageId="13" pageNumber="78" refId="ref18945" refString="Dejean, P. F. M. A. (1825) Species general des coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp." type="book" year="1825">DEJEAN, 1825</bibRefCitation>
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<emphasis id="B926EA86FFF577534AFEFB3435C4FA83" box="[151,165,1177,1201]" italics="true" pageId="13" pageNumber="78">=</emphasis>
<taxonomicName id="4C524D17FFF577534ACCFB3437D7FA83" ID-CoL="76WJP" authority="CHAUDOIR, 1843" authorityName="CHAUDOIR" authorityYear="1843" box="[165,694,1177,1201]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="quadripustulatus">
<emphasis id="B926EA86FFF577534ACCFB34348AFA83" box="[165,491,1177,1201]" italics="true" pageId="13" pageNumber="78">Pheropsophus quadripustulatus</emphasis>
<bibRefCitation id="EFC34B65FFF577534B98FB3437D7FA83" author="Chaudoir, M." box="[497,694,1177,1201]" pageId="13" pageNumber="78" pagination="671 - 791" refId="ref18827" refString="Chaudoir, M. (1843) Carabiques nouveaux. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 671 - 791." type="journal article" year="1843">CHAUDOIR, 1843</bibRefCitation>
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<emphasis id="B926EA86FFF577534AFEFB153435FAE2" box="[151,340,1208,1232]" italics="true" pageId="13" pageNumber="78">Aptinus occipitalis</emphasis>
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<bibRefCitation id="EFC34B65FFF577534BDBFB7B37E5FADD" author="Saha, S. K. &amp; Mukherjee, A. K. &amp; Sengupta, T." box="[434,644,1238,1263]" pageId="13" pageNumber="78" refId="ref20668" refString="Saha, S. K., Mukherjee, A. K. &amp; Sengupta, T. (1992) Records of the Zoological Survey of India. Occasional paper No. 144. Carabidae (Coleotera: Insecta) of Calcutta. A. K. Chatterjee at Jnanodaya Press, Calcutta, 63 pp." type="book" year="1992">
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<emphasis id="B926EA86FFF577534B80FB7A377DFADC" box="[489,540,1238,1262]" italics="true" pageId="13" pageNumber="78">et al.</emphasis>
, 1992: 50
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<treatmentCitationGroup id="AB4211BAFFF577534AFEFB5834F5FB1E" pageId="13" pageNumber="78">
<treatmentCitation id="0AF31085FFF577534AFEFB58374BFB3F" author="Dejean, P. F. M. A." box="[151,554,1268,1293]" page="304" pageId="13" pageNumber="78" year="1825">
<taxonomicName id="4C524D17FFF577534AFEFB58374BFB3F" ID-CoL="MT63" authority="Dejean, 1825: 304" authorityName="DEJEAN" authorityPageNumber="304" authorityYear="1825" box="[151,554,1268,1293]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="ambiguus">
<emphasis id="B926EA86FFF577534AFEFB583409FB3F" box="[151,360,1269,1293]" italics="true" pageId="13" pageNumber="78">Brachinus ambiguus</emphasis>
<bibRefCitation id="EFC34B65FFF577534B07FB59374BFB3F" author="Dejean, P. F. M. A." box="[366,554,1268,1293]" pageId="13" pageNumber="78" refId="ref18945" refString="Dejean, P. F. M. A. (1825) Species general des coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp." type="book" year="1825">Dejean, 1825: 304</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753485DFB593798FB3F" author="Chaudoir, M." box="[564,761,1268,1293]" page="37" pageId="13" pageNumber="78" year="1876">
<bibRefCitation id="EFC34B65FFF57753485DFB593798FB3F" author="Chaudoir, M." box="[564,761,1268,1293]" pageId="13" pageNumber="78" pagination="11 - 104" refId="ref18920" refString="Chaudoir, M. (1876) Monographie de Brachynides. Annales de la Societe entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">Chaudoir, 1876: 37</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753496AFB5B36DCFB3F" author="Bates, H. W." box="[771,957,1268,1293]" page="391" pageId="13" pageNumber="78" year="1892">
<bibRefCitation id="EFC34B65FFF57753496AFB5B36DCFB3F" author="Bates, H. W." box="[771,957,1268,1293]" pageId="13" pageNumber="78" pagination="267 - 428" refId="ref18669" refString="Bates, H. W. (1892 b) List of Carabidae. Viaggio Di Leonardo Fea in Birmania e Regioni Vicine. Annali Del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">Bates, 1892b: 391</bibRefCitation>
</treatmentCitation>
; Heyne-Tasch, 1908: 14;
<treatmentCitation id="0AF31085FFF577534ED4FB5930F9FB3F" author="Andrewes, H. E." box="[1213,1432,1268,1293]" page="275" pageId="13" pageNumber="78" year="1930">
<bibRefCitation id="EFC34B65FFF577534ED4FB5930F9FB3F" author="Andrewes, H. E." box="[1213,1432,1268,1293]" pageId="13" pageNumber="78" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes, 1930: 275</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF577534AAEFABE34F1FB1E" author="Jedlicka, A." box="[199,400,1299,1324]" page="533" pageId="13" pageNumber="78" year="1963">
<bibRefCitation id="EFC34B65FFF577534AAEFABE34F1FB1E" author="Jedlicka, A." box="[199,400,1299,1324]" pageId="13" pageNumber="78" pagination="269 - 579" refId="ref19830" refString="Jedlicka, A. (1963) Monographie der Truncatipennen aus Ostasien. Lebiinae-Odacanthinae-Brachyninae (Coleoptera: Carabi- dae). Entomologisсhe Abhandlungen und Berichte aus dem Staatliches Museum fur Tierkunde in Dresden, 28 (7), 269 - 579. [1962 - 1964]" type="journal article" year="1963">Jedlička, 1963: 533</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3694FFF577534AFEFA9F34BBFB5B" blockId="13.[151,1437,1084,1447]" pageId="13" pageNumber="78">
<treatmentCitationGroup id="AB4211BAFFF577534AFEFA9F34BBFB5B" pageId="13" pageNumber="78">
<treatmentCitation id="0AF31085FFF577534AFEFA9F374DFB79" author="Dejean, P. F. M. A." box="[151,556,1330,1355]" page="306" pageId="13" pageNumber="78" year="1825">
<taxonomicName id="4C524D17FFF577534AFEFA9F374DFB79" ID-CoL="68QDR" authority="Dejean, 1825: 306" authorityName="DEJEAN" authorityPageNumber="306" authorityYear="1825" box="[151,556,1330,1355]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="fuscicollis">
<emphasis id="B926EA86FFF577534AFEFA9F340DFB78" box="[151,364,1330,1354]" italics="true" pageId="13" pageNumber="78">Brachinus fuscicollis</emphasis>
<bibRefCitation id="EFC34B65FFF577534B18FA9F374DFB79" author="Dejean, P. F. M. A." box="[369,556,1330,1355]" pageId="13" pageNumber="78" refId="ref18945" refString="Dejean, P. F. M. A. (1825) Species general des coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp." type="book" year="1825">Dejean, 1825: 306</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753485CFA9E378BFB79" author="Putzeys, J. A. A. H." box="[565,746,1330,1355]" page="45" pageId="13" pageNumber="78" year="1875">
<bibRefCitation id="EFC34B65FFF57753485CFA9E378BFB79" author="Putzeys, J. A. A. H." box="[565,746,1330,1355]" pageId="13" pageNumber="78" pagination="45 - 55" refId="ref20551" refString="Putzeys, J. A. A. H. (1875) Notes sur les Carabiques recueillis par M. Jean Van Volxem a Ceylan, a Manille, en Chine et au Japon (1873 - 1874). Annales de la Societe Entomologique de Belgique, 18, 45 - 55." type="journal article" year="1875">Putzeys, 1875: 45</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753489DFA9F36D9FB79" author="Chaudoir, M." box="[756,952,1330,1355]" page="37" pageId="13" pageNumber="78" year="1876">
<bibRefCitation id="EFC34B65FFF57753489DFA9F36D9FB79" author="Chaudoir, M." box="[756,952,1330,1355]" pageId="13" pageNumber="78" pagination="11 - 104" refId="ref18920" refString="Chaudoir, M. (1876) Monographie de Brachynides. Annales de la Societe entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">Chaudoir, 1876: 37</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF5775349A8FA9E310CFB79" author="Bates, H. W." box="[961,1133,1330,1355]" page="200" pageId="13" pageNumber="78" year="1886">
<bibRefCitation id="EFC34B65FFF5775349A8FA9E310CFB79" author="Bates, H. W." box="[961,1133,1330,1355]" pageId="13" pageNumber="78" pagination="199 - 212" refId="ref18382" refString="Bates, H. W. (1886) On the Geodephagous Coleoptera collected by George Lewis in Ceylon. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 5 (17), 199 - 212. https: // doi. org / 10.1080 / 00222938609460134" type="journal article" year="1886">Bates, 1886: 200</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF577534E1EFA9F318AFB79" author="Bates, H. W." box="[1143,1259,1330,1355]" page="391" pageId="13" pageNumber="78" year="1892">
<bibRefCitation id="EFC34B65FFF577534E1EFA9F318AFB79" author="Bates, H. W." box="[1143,1259,1330,1355]" pageId="13" pageNumber="78" pagination="267 - 428" refId="ref18669" refString="Bates, H. W. (1892 b) List of Carabidae. Viaggio Di Leonardo Fea in Birmania e Regioni Vicine. Annali Del Museo Civico di Storia Naturale di Genova, 12 (2), 267 - 428." type="journal article" year="1892">1892b: 391</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF577534E9CFA9F3591FB5B" author="Bouchard, J." page="173" pageId="13" pageNumber="78" year="1903">
<bibRefCitation id="EFC34B65FFF577534E9CFA9F3591FB5B" author="Bouchard, J." pageId="13" pageNumber="78" pagination="169 - 176" refId="ref18783" refString="Bouchard, J. (1903) Insectes recueillis par M. le Professeur Dr. Forster a Borneo, Java et Sumatra (Palembang), Coleopteres Carabiques. Annales de la Societe entomologique de France, 72, 169 - 176." type="journal article" year="1903">Bouchard, 1903: 173</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF577534A90FAFD34B7FB5B" author="Andrewes, H. E." box="[249,470,1360,1385]" page="275" pageId="13" pageNumber="78" year="1930">
<bibRefCitation id="EFC34B65FFF577534A90FAFD34B7FB5B" author="Andrewes, H. E." box="[249,470,1360,1385]" pageId="13" pageNumber="78" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes, 1930: 275</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3694FFF577534AFEFADD3695FBBA" blockId="13.[151,1437,1084,1447]" box="[151,1012,1391,1416]" pageId="13" pageNumber="78">
<treatmentCitationGroup id="AB4211BAFFF577534AFEFADD3695FBBA" box="[151,1012,1391,1416]" pageId="13" pageNumber="78">
<treatmentCitation id="0AF31085FFF577534AFEFADD375BFBBA" author="Dejean, P. F. M. A." box="[151,570,1391,1416]" page="306" pageId="13" pageNumber="78" year="1825">
<taxonomicName id="4C524D17FFF577534AFEFADD375BFBBA" ID-CoL="MTBK" authority="Dejean, 1825: 306" authorityName="DEJEAN" authorityPageNumber="306" authorityYear="1825" box="[151,570,1391,1416]" class="Insecta" family="Carabidae" genus="Brachinus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="B926EA86FFF577534AFEFADD3416FBBA" box="[151,375,1392,1416]" italics="true" pageId="13" pageNumber="78">Brachinus interruptus</emphasis>
<bibRefCitation id="EFC34B65FFF577534B14FAC2375BFBBA" author="Dejean, P. F. M. A." box="[381,570,1391,1416]" pageId="13" pageNumber="78" refId="ref18945" refString="Dejean, P. F. M. A. (1825) Species general des coleopteres de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp." type="book" year="1825">Dejean, 1825: 306</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753482DFAC2366BFBBA" author="Chaudoir, M." box="[580,778,1391,1416]" page="37" pageId="13" pageNumber="78" year="1876">
<bibRefCitation id="EFC34B65FFF57753482DFAC2366BFBBA" author="Chaudoir, M." box="[580,778,1391,1416]" pageId="13" pageNumber="78" pagination="11 - 104" refId="ref18920" refString="Chaudoir, M. (1876) Monographie de Brachynides. Annales de la Societe entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">Chaudoir, 1876: 37</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753497DFAC23691FBBA" author="Andrewes, H. E." box="[788,1008,1391,1416]" page="275" pageId="13" pageNumber="78" year="1930">
<bibRefCitation id="EFC34B65FFF57753497DFAC23691FBBA" author="Andrewes, H. E." box="[788,1008,1391,1416]" pageId="13" pageNumber="78" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes, 1930: 275</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3694FFF577534AFEFA233162FB95" blockId="13.[151,1437,1084,1447]" box="[151,1027,1422,1447]" pageId="13" pageNumber="78">
<treatmentCitationGroup id="AB4211BAFFF577534AFEFA233162FB95" box="[151,1027,1422,1447]" pageId="13" pageNumber="78">
<treatmentCitation id="0AF31085FFF577534AFEFA2337D2FB95" author="Chaudoir, M." box="[151,691,1422,1447]" page="706" pageId="13" pageNumber="78" year="1843">
<taxonomicName id="4C524D17FFF577534AFEFA2337D2FB95" ID-CoL="76WJP" authority="Chaudoir, 1843: 706" authorityName="CHAUDOIR" authorityPageNumber="706" authorityYear="1843" box="[151,691,1422,1447]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="78" phylum="Arthropoda" rank="species" species="quadripustulatus">
<emphasis id="B926EA86FFF577534AFEFA2334BBFB94" box="[151,474,1422,1446]" italics="true" pageId="13" pageNumber="78">Pheropsophus quadripustulatus</emphasis>
<bibRefCitation id="EFC34B65FFF577534B89FA2337D2FB95" author="Chaudoir, M." box="[480,691,1422,1447]" pageId="13" pageNumber="78" pagination="671 - 791" refId="ref18827" refString="Chaudoir, M. (1843) Carabiques nouveaux. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 16, 671 - 791." type="journal article" year="1843">Chaudoir, 1843: 706</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF5775348D4FA233678FB95" author="Chaudoir, M." box="[701,793,1422,1447]" page="37" pageId="13" pageNumber="78" year="1876">
<bibRefCitation id="EFC34B65FFF5775348D4FA233678FB95" author="Chaudoir, M." box="[701,793,1422,1447]" pageId="13" pageNumber="78" pagination="11 - 104" refId="ref18920" refString="Chaudoir, M. (1876) Monographie de Brachynides. Annales de la Societe entomologique de Belgique, 19, 11 - 104." type="journal article" year="1876">1876: 37</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="0AF31085FFF57753494BFA23369EFB95" author="Andrewes, H. E." box="[802,1023,1422,1447]" page="275" pageId="13" pageNumber="78" year="1930">
<bibRefCitation id="EFC34B65FFF57753494BFA23369EFB95" author="Andrewes, H. E." box="[802,1023,1422,1447]" pageId="13" pageNumber="78" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes, 1930: 275</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BED3694FFF577534AFEFA79301BF805" blockId="13.[151,1437,1492,2023]" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AFEFA793487FBDD" bold="true" box="[151,486,1492,1519]" pageId="13" pageNumber="78">Specimens examined (n=22)</emphasis>
:
<materialsCitation id="3B3A3CC9FFF577534B9FFA783696FBDD" ID-GBIF-Occurrence="2846550304" box="[502,1015,1493,1519]" collectingDate="1973-08-11" collectionCode="BMNH" country="China" location="China" pageId="13" pageNumber="78" specimenCount="1">
<specimenCount id="9D54FD1DFFF577534B9FFA78374FFBDD" box="[502,558,1493,1519]" pageId="13" pageNumber="78" type="generic">1 ex.</specimenCount>
<collectingCountry id="F3457604FFF577534829FA7837E9FBDD" box="[576,648,1493,1519]" name="China" pageId="13" pageNumber="78">China</collectingCountry>
,
<date id="FFEC1054FFF5775348FDFA783649FBDD" box="[660,808,1493,1519]" pageId="13" pageNumber="78" value="1973-08-11">
<collectingDate id="EFA8E9BCFFF5775348FDFA783649FBDD" box="[660,808,1493,1519]" pageId="13" pageNumber="78" value="1973-08-11">11.VIII.1973</collectingDate>
</date>
, Yu.P,” in
<collectionCode id="ED43AE51FFF5775349F6FA783696FBDD" box="[927,1015,1493,1519]" pageId="13" pageNumber="78">BMNH</collectionCode>
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF577534E6AFA7830E3FBDD" ID-GBIF-Occurrence="2846550310" box="[1027,1410,1493,1519]" collectorName="Coimbatore" country="India" location="India" pageId="13" pageNumber="78" specimenCount="3">
<specimenCount id="9D54FD1DFFF577534E6AFA783126FBDD" box="[1027,1095,1493,1519]" pageId="13" pageNumber="78" type="generic">3 exs.</specimenCount>
,
<collectingCountry id="F3457604FFF577534E3DFA7831F3FBDD" box="[1108,1170,1493,1519]" name="India" pageId="13" pageNumber="78">India</collectingCountry>
:
<collectorName id="26A75342FFF577534EF4FA783047FBDD" box="[1181,1318,1493,1519]" pageId="13" pageNumber="78">Coimbatore</collectorName>
: TNAU
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF577534FE7FA78310FF821" ID-GBIF-Occurrence="2846550366" collectingDate="2016-02-28" collectorName="S. V. Akhil" country="India" county="Nilambur" location="Hand" municipality="Nedumkayam" pageId="13" pageNumber="78" specimenCount="7" stateProvince="Kerala">
<specimenCount id="9D54FD1DFFF577534FE7FA7835A7F821" pageId="13" pageNumber="78" type="generic">7 exs.</specimenCount>
,
<collectingCountry id="F3457604FFF577534AB8FA54346EF821" box="[209,271,1529,1555]" name="India" pageId="13" pageNumber="78">India</collectingCountry>
:
<collectingRegion id="4996F876FFF577534B73FA543408F821" box="[282,361,1529,1555]" country="India" name="Kerala" pageId="13" pageNumber="78">Kerala</collectingRegion>
:
<collectingCounty id="628C4E18FFF577534B1AFA543482F821" box="[371,483,1529,1555]" pageId="13" pageNumber="78">Nilambur</collectingCounty>
:
<collectingMunicipality id="6B89ACEEFFF577534B84FA5437ECF821" box="[493,653,1529,1555]" pageId="13" pageNumber="78">Nedumkayam</collectingMunicipality>
,
<location id="8E8D604FFFF5775348F1FA5437B4F821" LSID="urn:lsid:plazi:treatment:03FB8782FFF577504AFEFC79374CFF9C:8E8D604FFFF5775348F1FA5437B4F821" box="[664,725,1529,1555]" country="India" county="Nilambur" municipality="Nedumkayam" name="Hand" pageId="13" pageNumber="78" stateProvince="Kerala">Hand</location>
picking,
<date id="FFEC1054FFF577534954FA5436D5F821" box="[829,948,1529,1555]" pageId="13" pageNumber="78" value="2016-02-28">
<collectingDate id="EFA8E9BCFFF577534954FA5436D5F821" box="[829,948,1529,1555]" pageId="13" pageNumber="78" value="2016-02-28">28.II.2016</collectingDate>
</date>
, coll.
<collectorName id="26A75342FFF577534991FA54310FF821" box="[1016,1134,1529,1555]" pageId="13" pageNumber="78">S.V. Akhil</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF577534E13FA54373BF805" ID-GBIF-Occurrence="2846550373" collectingDate="2017-09-13" collectorName="K. Ashly" country="India" location="Kalpetta" pageId="13" pageNumber="78" specimenCount="8" stateProvince="Kerala">
<specimenCount id="9D54FD1DFFF577534E13FA5431DCF821" box="[1146,1213,1529,1555]" pageId="13" pageNumber="78" type="generic">8 exs.</specimenCount>
,
<collectingCountry id="F3457604FFF577534EA0FA543066F821" box="[1225,1287,1529,1555]" name="India" pageId="13" pageNumber="78">India</collectingCountry>
:
<collectingRegion id="4996F876FFF577534F78FA543001F821" box="[1297,1376,1529,1555]" country="India" name="Kerala" pageId="13" pageNumber="78">Kerala</collectingRegion>
:
<location id="8E8D604FFFF577534F03FA5435B1F805" LSID="urn:lsid:plazi:treatment:03FB8782FFF577504AFEFC79374CFF9C:8E8D604FFFF577534F03FA5435B1F805" country="India" name="Kalpetta" pageId="13" pageNumber="78" stateProvince="Kerala">Kalpetta</location>
,
<collectingMethod id="52134E83FFF577534AB5F9B03442F805" box="[220,291,1565,1591]" pageId="13" pageNumber="78">Pitfall</collectingMethod>
,
<date id="FFEC1054FFF577534B47F9B034D1F805" box="[302,432,1565,1591]" pageId="13" pageNumber="78" value="2017-09-13">
<collectingDate id="EFA8E9BCFFF577534B47F9B034D1F805" box="[302,432,1565,1591]" pageId="13" pageNumber="78" value="2017-09-13">13.IX.2017</collectingDate>
</date>
, coll.
<collectorName id="26A75342FFF577534B9DF9B0373BF805" box="[500,602,1565,1591]" pageId="13" pageNumber="78">K. Ashly</collectorName>
</materialsCitation>
;
<materialsCitation id="3B3A3CC9FFF57753480FF9B0301BF805" ID-GBIF-Occurrence="2846550338" box="[614,1402,1565,1591]" collectingDate="2017-11-27" collectorName="M. Divya" country="India" location="Malampuzha" pageId="13" pageNumber="78" specimenCount="3" stateProvince="Kerala">
<specimenCount id="9D54FD1DFFF57753480FF9B037CBF805" box="[614,682,1565,1591]" pageId="13" pageNumber="78" type="generic">3 exs.</specimenCount>
,
<collectingCountry id="F3457604FFF5775348DCF9B03792F805" box="[693,755,1565,1591]" name="India" pageId="13" pageNumber="78">India</collectingCountry>
:
<collectingRegion id="4996F876FFF577534897F9B0362CF805" box="[766,845,1565,1591]" country="India" name="Kerala" pageId="13" pageNumber="78">Kerala</collectingRegion>
:
<location id="8E8D604FFFF57753493EF9B0368DF805" LSID="urn:lsid:plazi:treatment:03FB8782FFF577504AFEFC79374CFF9C:8E8D604FFFF57753493EF9B0368DF805" box="[855,1004,1565,1591]" country="India" name="Malampuzha" pageId="13" pageNumber="78" stateProvince="Kerala">Malampuzha</location>
,
<collectingMethod id="52134E83FFF57753499EF9B0315FF805" box="[1015,1086,1565,1591]" pageId="13" pageNumber="78">Pitfall</collectingMethod>
,
<date id="FFEC1054FFF577534E23F9B031AAF805" box="[1098,1227,1565,1591]" pageId="13" pageNumber="78" value="2017-11-27">
<collectingDate id="EFA8E9BCFFF577534E23F9B031AAF805" box="[1098,1227,1565,1591]" pageId="13" pageNumber="78" value="2017-11-27">27.XI.2017</collectingDate>
</date>
, coll.
<collectorName id="26A75342FFF577534F66F9B0301BF805" box="[1295,1402,1565,1591]" pageId="13" pageNumber="78">M. Divya</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFF577504AAEF9ED3612FF71" lastPageId="14" lastPageNumber="79" pageId="13" pageNumber="78" type="description">
<paragraph id="8BED3694FFF577534AAEF9ED3700F869" blockId="13.[151,1437,1492,2023]" box="[199,609,1600,1627]" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AAEF9ED3438F869" bold="true" box="[199,345,1600,1627]" pageId="13" pageNumber="78">Description.</emphasis>
Length: 15.019.0 mm
</paragraph>
<paragraph id="8BED3694FFF577534AAEF9C93492F8D9" blockId="13.[151,1437,1492,2023]" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AAEF9C93445F84D" bold="true" box="[199,292,1636,1663]" pageId="13" pageNumber="78">Colour:</emphasis>
Head yellow with black spots on vertex, anterior and posterior black spot connected by middle narrow black line; antennal segment 12 reddish yellow and rest reddish brown; pronotum dark black, with two lateral yellow spots, in dumbbell shaped black background; elytra black with humeral spots; legs reddish yellow with hind femur with black spot on apex.
</paragraph>
<paragraph id="8BED3694FFF577534AAEF958364DF93D" blockId="13.[151,1437,1492,2023]" box="[199,812,1781,1807]" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AAEF9583471F93D" bold="true" box="[199,272,1781,1807]" pageId="13" pageNumber="78">Head:</emphasis>
Eyes protruding; pronounced wrinkles on neck.
</paragraph>
<paragraph id="8BED3694FFF577534AAEF8B4301AF965" blockId="13.[151,1437,1492,2023]" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AAEF8B43428F901" bold="true" box="[199,329,1817,1843]" pageId="13" pageNumber="78">Pronotum:</emphasis>
Glabrous, smooth, shiny; anterior margin curved inwards; disc laterally with anterior margin convex, narrow posteriorly with posterior 1/3
<superScript id="7C279BDCFFF57753485CF8913722F978" attach="left" box="[565,579,1852,1866]" fontSize="6" pageId="13" pageNumber="78">rd</superScript>
of lateral margin diverging rather than parallel; posterior margin straight.
</paragraph>
<paragraph id="8BED3694FFF577534AAEF8CC31A8F9F1" blockId="13.[151,1437,1492,2023]" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AAEF8CC347CF949" bold="true" box="[199,285,1889,1915]" pageId="13" pageNumber="78">Elytra:</emphasis>
Subparallel rather than convex, narrowed toward base; humeral spot large, extending somewhat posteriorly; median band narrow; lateral edge with yellow margin starting from the humeral spot and joins the prominent yellow apical band; disc matt and finely pubescent; narrow, deep striations with wider intervals.
</paragraph>
<paragraph id="8BED3694FFF577504AAEF8613612FF71" blockId="13.[151,1437,1492,2023]" lastBlockId="14.[151,1437,152,431]" lastPageId="14" lastPageNumber="79" pageId="13" pageNumber="78">
<emphasis id="B926EA86FFF577534AAEF8613764F9D5" bold="true" box="[199,517,1996,2023]" pageId="13" pageNumber="78">Geographical distribution.</emphasis>
<collectingCountry id="F3457604FFF577534865F8603701F9D5" box="[524,608,1997,2023]" name="India" pageId="13" pageNumber="78">INDIA</collectingCountry>
:
<collectingRegion id="4996F876FFF577534803F86037AFF9D5" box="[618,718,1997,2023]" country="India" name="Manipur" pageId="13" pageNumber="78">Manipur</collectingRegion>
;
<collectingRegion id="4996F876FFF5775348B1F8603649F9D5" box="[728,808,1997,2023]" country="India" name="Assam" pageId="13" pageNumber="78">Assam</collectingRegion>
; Mizoram-Tripura: Lushai Hills; Bengal: Kolkata (
<bibRefCitation id="EFC34B65FFF577504F0DF8603472FE81" author="Saha, S. K. &amp; Mukherjee, A. K. &amp; Sengupta, T." lastPageId="14" lastPageNumber="79" pageId="13" pageNumber="78" refId="ref20668" refString="Saha, S. K., Mukherjee, A. K. &amp; Sengupta, T. (1992) Records of the Zoological Survey of India. Occasional paper No. 144. Carabidae (Coleotera: Insecta) of Calcutta. A. K. Chatterjee at Jnanodaya Press, Calcutta, 63 pp." type="book" year="1992">
Saha
<emphasis id="B926EA86FFF677504AFEFF3435B1FE80" box="[151,208,152,178]" italics="true" pageId="14" pageNumber="79">et al.</emphasis>
1992
</bibRefCitation>
);
<collectingRegion id="4996F876FFF677504B41FF35340BFE80" box="[296,362,152,178]" country="India" name="Bihar" pageId="14" pageNumber="79">Bihar</collectingRegion>
;
<collectingRegion id="4996F876FFF677504B1CFF3534A9FE80" box="[373,456,152,178]" country="India" name="Odisha" pageId="14" pageNumber="79">Odisha</collectingRegion>
;
<collectingRegion id="4996F876FFF677504BBAFF35373DFE80" box="[467,604,152,178]" country="India" name="Tamil Nadu" pageId="14" pageNumber="79">Tamil Nadu</collectingRegion>
: Coimbatore (New record);
<collectingRegion id="4996F876FFF6775049F0FF353689FE80" box="[921,1000,152,178]" country="India" name="Kerala" pageId="14" pageNumber="79">Kerala</collectingRegion>
: Malampuzha (New record), Nilambur (New record), Thamarassery (New record), Kalpetta (New record); Andaman Is (
<bibRefCitation id="EFC34B65FFF677504E51FF113190FEE5" author="Andrewes, H. E." box="[1080,1265,188,215]" pageId="14" pageNumber="79" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes 1930</bibRefCitation>
);
<collectingCountry id="F3457604FFF677504F6EFF1030FDFEE5" box="[1287,1436,189,215]" name="Sri Lanka" pageId="14" pageNumber="79">SRI LANKA</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF677504AF6FF4D3438FEC9" author="Andrewes, H. E." box="[159,345,224,251]" pageId="14" pageNumber="79" pagination="119 - 217" refId="ref17745" refString="Andrewes, H. E. (1919) On the types of Oriental Carabidae in the British Museum and in the Hope Department of the Oxford University Museum. Transactions of Entomological Society of London, 1919, 119 - 217. https: // doi. org / 10.1111 / j. 1365 - 2311.1919. tb 00006. x" type="journal article" year="1919">Andrewes 1919</bibRefCitation>
);
<collectingCountry id="F3457604FFF677504B18FF4C3762FEC9" box="[369,515,225,251]" name="Myanmar" pageId="14" pageNumber="79">MYANMAR</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF67750487CFF4D37AEFEC9" author="Andrewes, H. E." box="[533,719,224,251]" pageId="14" pageNumber="79" pagination="119 - 217" refId="ref17745" refString="Andrewes, H. E. (1919) On the types of Oriental Carabidae in the British Museum and in the Hope Department of the Oxford University Museum. Transactions of Entomological Society of London, 1919, 119 - 217. https: // doi. org / 10.1111 / j. 1365 - 2311.1919. tb 00006. x" type="journal article" year="1919">Andrewes 1919</bibRefCitation>
);
<collectingCountry id="F3457604FFF67750488EFF4C3616FEC9" box="[743,887,225,251]" name="Malaysia" pageId="14" pageNumber="79">MALAYSIA</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF6775049E0FF4D3122FEC9" author="Andrewes, H. E." box="[905,1091,224,251]" pageId="14" pageNumber="79" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes 1930</bibRefCitation>
);
<collectingCountry id="F3457604FFF677504E32FF4C3199FEC9" box="[1115,1272,225,251]" name="Indonesia" pageId="14" pageNumber="79">INDONESIA</collectingCountry>
:
<collectingRegion id="4996F876FFF677504F6FFF4C305BFEC9" box="[1286,1338,225,251]" country="Indonesia" name="Jawa Timur" pageId="14" pageNumber="79">Java</collectingRegion>
, Borneo (
<bibRefCitation id="EFC34B65FFF677504AF6FEA93424FF2D" author="Macleay, W. S." box="[159,325,260,287]" pageId="14" pageNumber="79" refId="ref20195" refString="Macleay, W. S. (1825) Number I. of Annulosa Javanica, or an attempt to illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfield, M. D. F. L. &amp; G. S. and deposited by him in the museum of the honourable East-India Company. Kingsbury, Parbury, and Allen, London, xii + 50 pp., 1 pls." type="book" year="1825">Macleay 1825</bibRefCitation>
); Malay Peninsula and Archipelago (
<bibRefCitation id="EFC34B65FFF677504880FEA936C0FF2D" author="Andrewes, H. E." box="[745,929,260,287]" pageId="14" pageNumber="79" pagination="119 - 217" refId="ref17745" refString="Andrewes, H. E. (1919) On the types of Oriental Carabidae in the British Museum and in the Hope Department of the Oxford University Museum. Transactions of Entomological Society of London, 1919, 119 - 217. https: // doi. org / 10.1111 / j. 1365 - 2311.1919. tb 00006. x" type="journal article" year="1919">Andrewes 1919</bibRefCitation>
);
<collectingCountry id="F3457604FFF6775049DFFEA8316CFF2D" box="[950,1037,261,287]" name="China" pageId="14" pageNumber="79">CHINA</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF677504E74FEA931C1FF2D" author="Habu, A." box="[1053,1184,260,287]" pageId="14" pageNumber="79" pagination="101 - 173" refId="ref19513" refString="Habu, A. (1984) Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (IV) (Coleoptera, Carabidae). The Entomological Review of Japan, 39, 101 - 173." type="journal article" year="1984">Habu 1984</bibRefCitation>
);
<collectingCountry id="F3457604FFF677504EDCFEA8303EFF2D" box="[1205,1375,261,287]" name="Hong Kong" pageId="14" pageNumber="79">HONG KONG</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF677504F06FEA83448FF71" author="Andrewes, H. E." pageId="14" pageNumber="79" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes 1930</bibRefCitation>
);
<collectingCountry id="F3457604FFF677504B54FE8434C8FF71" box="[317,425,297,323]" name="Taiwan" pageId="14" pageNumber="79">TAIWAN</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF677504BD1FE85370FFF71" author="Andrewes, H. E." box="[440,622,296,323]" pageId="14" pageNumber="79" refId="ref18200" refString="Andrewes, H. E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp." type="book" year="1930">Andrewes 1930</bibRefCitation>
);
<collectingCountry id="F3457604FFF6775048EBFE8437B6FF71" box="[642,727,297,323]" name="Japan" pageId="14" pageNumber="79">JAPAN</collectingCountry>
(
<bibRefCitation id="EFC34B65FFF67750488CFE853606FF71" author="Habu, A." box="[741,871,296,323]" pageId="14" pageNumber="79" pagination="101 - 173" refId="ref19513" refString="Habu, A. (1984) Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (IV) (Coleoptera, Carabidae). The Entomological Review of Japan, 39, 101 - 173." type="journal article" year="1984">Habu 1984</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C348651FFFF677504AAEFEE0374CFF9C" pageId="14" pageNumber="79" type="discussion">
<paragraph id="8BED3694FFF677504AAEFEE0374CFF9C" blockId="14.[151,1437,152,431]" pageId="14" pageNumber="79">
<emphasis id="B926EA86FFF677504AAEFEE0345AFF55" bold="true" box="[199,315,333,359]" pageId="14" pageNumber="79">Remarks.</emphasis>
Very similar to
<taxonomicName id="4C524D17FFF677504B97FEE037C7FF54" baseAuthorityName="Linnaeus" baseAuthorityYear="1771" box="[510,678,332,358]" class="Insecta" family="Carabidae" genus="Pheropsophus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="79" phylum="Arthropoda" rank="species" species="bimaculatus">
<emphasis id="B926EA86FFF677504B97FEE037C7FF54" box="[510,678,332,358]" italics="true" pageId="14" pageNumber="79">P. bimaculatus</emphasis>
</taxonomicName>
but differs by the presence of three spots (anterior and posterior rounded, middle spot narrow connecting the two) on vertex, pronotum with larger spots reaching the lateral margin and narrow median transverse band.
</paragraph>
</subSubSection>
</treatment>
</document>