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<document id="420715AC177FF7A27BC745623F78D7EF" ID-DOI="10.5852/ejt.2018.459" ID-GBIF-Dataset="3cb35a49-8d51-490f-b2a1-4706ffca1686" ID-ISSN="2118-9773" ID-Zenodo-Dep="1468694" ID-ZooBank="urn:lsid:zoobank.org:pub:7F198F1E-A4F6-457E-9E03-667D709C35DD" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="guilherme" IM.treatments_approvedBy="guilherme" checkinTime="1540214293721" checkinUser="plazi" docAuthor="Reshchikov, Alexey, Sääksjärvi, Ilari E. &amp; Pollet, Marc" docDate="2018" docId="03E387CEFFE1CB75F768FC0FFA83F862" docLanguage="en" docName="ejt-459_reshchikov_sààksjàrvi_pollett.pdf.imf" docOrigin="European Journal of Taxonomy 459" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nanium Townes 1967" docType="treatment" docVersion="2" lastPageNumber="4" masterDocId="FFDAFFB6FFE3CB76F501FFEFFFF4FF89" masterDocTitle="Review of the New World genus Nanium Townes, 1967 (Hymenoptera: Ichneumonidae: Ctenopelmatinae), with two new species from the Neotropical region" masterLastPageNumber="18" masterPageNumber="1" pageNumber="3" updateTime="1736275312465" updateUser="ExternalLinkService">
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<mods:title id="7AE441697F82F03D40067B2C0CE42760">Review of the New World genus Nanium Townes, 1967 (Hymenoptera: Ichneumonidae: Ctenopelmatinae), with two new species from the Neotropical region</mods:title>
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<heading id="D0BD81B4FFE1CB74F768FC0FFC3FFC73" box="[617,971,992,1018]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
Genus 
<taxonomicName id="4C4A4D5BFFE1CB74F7BAFC0FFC3FFC73" ID-CoL="63CGB" ID-ENA="494892" authority="Townes, 1967" authorityName="Townes" authorityYear="1967" box="[699,971,992,1018]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEACAFFE1CB74F7BAFC0FFCEAFC73" bold="true" box="[699,798,992,1018]" italics="true" pageId="2" pageNumber="3">Nanium</emphasis>
<bibRefCitation id="EFDB4B29FFE1CB74F627FC0FFC3FFC73" box="[806,971,992,1018]" pageId="2" pageNumber="3" refId="ref5767" refString="Townes H. K. 1967. Genus Nanium Townes, n. genus. In: Krombein K. V. &amp; Burks B. D., Hymenoptera of America North of Mexico - Synoptic Catalog 2 (2 nd suppl.): 174 - 175. U. S. Dept of Agriculture Washington. https: // doi. org / 10.5962 / bhl. title. 4124" type="book chapter">Townes, 1967</bibRefCitation>
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<taxonomicName id="4C4A4D5BFFE1CB74F5BCFBCCFDF5FBB4" ID-CoL="63CGB" authority="Townes, 1967: 174" authorityName="Townes" authorityPageNumber="174" authorityYear="1967" box="[189,513,1058,1085]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEACAFFE1CB74F5BCFBCCFEEEFBB5" box="[189,282,1059,1084]" italics="true" pageId="2" pageNumber="3">Nanium</emphasis>
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: 174
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–175.
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<emphasis id="B93EEACAFFE1CB74F5BCFB8BFEACFBF7" bold="true" box="[189,344,1124,1150]" pageId="2" pageNumber="3">Type species</emphasis>
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<taxonomicName id="4C4A4D5BFFE1CB74F5BCFB7CFDB7FB24" authority="Cresson, 1864" authorityName="Cresson" authorityYear="1864" box="[189,579,1171,1197]" class="Insecta" family="Ichneumonidae" genus="Tryphon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="capitatus" status="by original designation">
<emphasis id="B93EEACAFFE1CB74F5BCFB7CFE66FB24" box="[189,402,1171,1197]" italics="true" pageId="2" pageNumber="3">Tryphon capitatus</emphasis>
<bibRefCitation id="EFDB4B29FFE1CB74F498FB7CFDB7FB24" box="[409,579,1171,1197]" pageId="2" pageNumber="3" refId="ref5533" refString="Cresson E. T. 1864. Descriptions of North American Hymenoptera in the collection of the Entomological Society of Philadelphia. Proceedings of the Entomological Society of Philadelphia 3: 257 - 321." type="journal article">Cresson, 1864</bibRefCitation>
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, 
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.
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<emphasis id="B93EEACAFFE1CB74F5BCFB3AFEC2FB66" bold="true" box="[189,310,1237,1263]" pageId="2" pageNumber="3">Diagnosis</emphasis>
(Updated from 
<bibRefCitation id="EFDB4B29FFE1CB74F4F0FB39FD7BFB66" box="[497,655,1237,1264]" pageId="2" pageNumber="3" refId="ref5840" refString="Townes H. K. 1970. The genera of Ichneumonidae, Part 3. Memoirs of the American Entomological Institute 13: 307." type="journal article">Townes 1970</bibRefCitation>
)
</paragraph>
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Generally small-bodied species (
<quantity id="4CB29B3DFFE1CB74F751FAEBFD49FA96" box="[592,701,1284,1311]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="6.0" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="4.5" valueMax="6.0" valueMin="3.0">3–6 mm</quantity>
), similar to the smaller species of 
<taxonomicName id="4C4A4D5BFFE1CB74F178FAEBFAFDFA97" authorityName="Forster" authorityYear="1869" box="[1145,1289,1284,1310]" class="Insecta" family="Ichneumonidae" genus="Lathrolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEACAFFE1CB74F178FAEBFAFDFA97" box="[1145,1289,1284,1310]" italics="true" pageId="2" pageNumber="3">Lathrolestes</emphasis>
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. Can be distinguished from the latter by the absence of propodeal carinae, except occasional traces of median longitudinal carinae posteriorly, and by the granulate sculpture on the propodeum and T1. In contrast with other genera of 
<taxonomicName id="4C4A4D5BFFE1CB74F4B7FA9FFDDAFA03" authorityName="Thomson" authorityYear="1883" box="[438,558,1392,1418]" class="Insecta" family="Ichneumonidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="tribe" tribe="Perilissini">Perilissini</taxonomicName>
, the epicnemial carina in 
<taxonomicName id="4C4A4D5BFFE1CB74F661FA9EFC49FA03" authorityName="Townes" authorityYear="1967" box="[864,957,1393,1418]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEACAFFE1CB74F661FA9EFC49FA03" box="[864,957,1393,1418]" italics="true" pageId="2" pageNumber="3">Nanium</emphasis>
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extends to the anterior margin of the mesopleuron, the pterostigma is short and the marginal cell is also short and deep. The genus can further be characterized as follows: clypeus with ventral margin thick and bluntly rounded, thinner laterally, without small lateral tooth or projection. Ventral margin of clypeus convex. Epistomal sulcus usually distinctly present. Clypeus in profile not projecting or only very weakly angled outwardly from face. Malar space distinct, from very short (0.2 × basal width of mandible) to moderately long (0.3–0.5 × basal width of mandible). Diameter of lateral ocellus much shorter than the distance between ocellus and eye. Maxillary palp moderately long, about equal to head height. Female antenna ranging from as long as to slightly, but distinctly, longer than body. Occipital carina dorsally complete, its lower part joining hypostomal carina distinctly above base of mandible. Dorsal end of epicnemial carina extending to anterior margin of mesopleuron. Notaulus varying from nearly absent to distinctly impressed on anterior declivity, extending posteriorly to level of tegula as a weak groove. Groove present between propodeum and metanotum indistinct in lateral view. Propodeal carinae usually completely absent (occasionally traces of median longitudinal carinae). Apex of mid tibia somewhat evenly rounded, with at most a small, weak tooth (similar to several species of genus 
<taxonomicName id="4C4A4D5BFFE1CB74F65AF88AFB8AF809" authority="Forster, 1855" authorityName="Forster" authorityYear="1855" box="[859,1150,1893,1920]" class="Insecta" family="Ichneumonidae" genus="Perilissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEACAFFE1CB74F65AF88AFC3BF8F6" box="[859,975,1893,1919]" italics="true" pageId="2" pageNumber="3">Perilissus</emphasis>
Förster, 1855
</taxonomicName>
). Apex of hind tibia with poorly developed comb, with setae widely spaced, not dense. Posterior hind tibial spur at least 7 × longer than maximum width at base. Claws varying from completely pectinate to pectinate only in basal half. Fore wing nearly always with vestigial areolet. Pterostigma short, broad, with 
<emphasis id="B93EEACAFFE1CB74F1D6F83DFB00F862" box="[1239,1268,2002,2027]" italics="true" pageId="2" pageNumber="3">Rs</emphasis>
+2 
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<emphasis id="B93EEACAFFE1CB74F015F83DFAEBF862" box="[1300,1311,2002,2027]" italics="true" pageId="2" pageNumber="3">r</emphasis>
arising
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near middle of pterostigma. Hind wing with first abscissa of 
<emphasis id="B93EEACAFFE0CB75F689FEE3FC54FEAC" box="[904,928,268,293]" italics="true" pageId="3" pageNumber="4">M</emphasis>
+ 
<emphasis id="B93EEACAFFE0CB75F6B0FEE3FC27FEAF" box="[945,979,268,294]" italics="true" pageId="3" pageNumber="4">Cu</emphasis>
1 strongly bowed; distal abscissa of 
<emphasis id="B93EEACAFFE0CB75F5BCFED7FF2BFEDB" box="[189,223,312,338]" italics="true" pageId="3" pageNumber="4">Cu</emphasis>
l not always discernible but when visible, joining cu-a closer to l 
<emphasis id="B93EEACAFFE0CB75F6DCFED7FC1BFEDB" box="[989,1007,312,338]" italics="true" pageId="3" pageNumber="4">A</emphasis>
than to 
<emphasis id="B93EEACAFFE0CB75F151FED7FB9CFED8" box="[1104,1128,312,337]" italics="true" pageId="3" pageNumber="4">M</emphasis>
; distal abscissae of 1 
<emphasis id="B93EEACAFFE0CB75F064FED7FA82FEDB" box="[1381,1398,312,338]" italics="true" pageId="3" pageNumber="4">A</emphasis>
and 
<emphasis id="B93EEACAFFE0CB75F5EEFE8BFEE5FEF7" box="[239,273,356,382]" italics="true" pageId="3" pageNumber="4">Cu</emphasis>
l spectral, vestigial or absent. T1 varying from moderately long and slender to shorter, distinctly broadening posteriorly and flattened dorsally. Dorsal carinae absent. Basal depression at dorsal tendon attachment absent. Dorsal-lateral carina absent or nearly so between spiracle and apex of T1. Glymmae on each side meeting on the midline posterior to dorsal tendon attachment, large, deep, and separated at midline by translucent partition. T2 with thyridium absent. Ovipositor straight, short, with very broad, deep, subapical depression. Ovipositor sheath from narrowly rectangular to weakly expanding distally. Male parameres broad, rectangular, never strongly narrowed posteriorly. Aedeagus rounded and weakly clubbed distally.
</paragraph>
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<heading id="D0BD81B4FFE0CB75F5BCFD0DFD3AFD75" bold="true" box="[189,718,738,764]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="B93EEACAFFE0CB75F5BCFD0DFD3AFD75" bold="true" box="[189,718,738,764]" pageId="3" pageNumber="4">
Identification key to the species of 
<taxonomicName id="4C4A4D5BFFE0CB75F76AFD0DFD3AFD75" authorityName="Townes" authorityYear="1967" box="[619,718,738,764]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B93EEACAFFE0CB75F76AFD0DFD3AFD75" bold="true" box="[619,718,738,764]" italics="true" pageId="3" pageNumber="4">Nanium</emphasis>
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</emphasis>
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1. Face distinctly broader than high (
<figureCitation id="13712A5DFFE0CB75F797FCF6FD08FCBA" box="[662,764,793,819]" captionStart="Fig. 2" captionStartId="5.[189,236,1948,1977]" captionTargetBox="[213,1375,263,1907]" captionTargetId="figure@5.[213,1375,263,1907]" captionText="Fig. 2. Nanium capitatum Townes, 1967, lectotype, Ƌ (ANSP). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468698" httpUri="https://zenodo.org/record/1468698/files/figure.png" pageId="3" pageNumber="4">Figs 2A</figureCitation>
, 
<figureCitation id="13712A5DFFE0CB75F60FFCF6FCC0FCBA" box="[782,820,793,819]" captionStart="Fig. 3" captionStartId="6.[189,236,1948,1977]" captionTargetBox="[218,1379,263,1905]" captionTargetId="figure@6.[218,1379,263,1905]" captionText="Fig. 3. Nanium capitatum Townes, 1967, ♀ (MZLU-HYM 00020508). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468700" httpUri="https://zenodo.org/record/1468700/files/figure.png" pageId="3" pageNumber="4">3A</figureCitation>
, 5A, 6A)...........................................................2
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− Face at most about as broad as high (
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) ................................................................................. 5
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</paragraph>
<keyStep id="30BE2E7DFFE0CB75F5BCFC60FA83FBD3" pageId="3" pageNumber="4">
<paragraph id="8BF536D8FFE0CB75F5BCFC60FA83FC5F" blockId="3.[189,1399,911,1114]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5BCFC60FA83FC5F" pageId="3" pageNumber="4">
2. Mesosoma with yellow stripes anteriorly on mesoscutum (
<figureCitation id="13712A5DFFE0CB75F6B6FC60FBEDFC23" box="[951,1049,911,938]" captionStart="Fig. 5" captionStartId="8.[189,236,1948,1977]" captionTargetBox="[213,1374,265,1907]" captionTargetId="figure@8.[213,1374,265,1907]" captionText="Fig. 5. Nanium medianum Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (EC/2009-37/ MP&amp;ADB-013 (RBINS)). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468704" httpUri="https://zenodo.org/record/1468704/files/figure.png" pageId="3" pageNumber="4">Fig. 5C</figureCitation>
). Metasoma with all tergites extensively pale medially (
<figureCitation id="13712A5DFFE0CB75F727FC54FD8BFC5F" box="[550,639,955,982]" captionStart="Fig. 5" captionStartId="8.[189,236,1948,1977]" captionTargetBox="[213,1374,265,1907]" captionTargetId="figure@8.[213,1374,265,1907]" captionText="Fig. 5. Nanium medianum Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (EC/2009-37/ MP&amp;ADB-013 (RBINS)). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468704" httpUri="https://zenodo.org/record/1468704/files/figure.png" pageId="3" pageNumber="4">Fig. 5B</figureCitation>
) ....................... 
<taxonomicName id="4C4A4D5BFFE0CB75F63EFC53FAE1FC5C" authority="Reshchikov &amp; Saaksjarvi" authorityName="Reshchikov &amp; Sääksjärvi &amp; Pollet" authorityYear="2018" box="[831,1301,955,982]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="medianum" status="sp. nov.">
<emphasis id="B93EEACAFFE0CB75F63EFC53FC10FC5F" bold="true" box="[831,996,955,982]" italics="true" pageId="3" pageNumber="4">N. medianum</emphasis>
Reshchikov &amp; Sääksjärvi
</taxonomicName>
<taxonomicNameLabel id="A20D57B1FFE0CB75F01DFC53FA83FC5F" box="[1308,1399,956,982]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BF536D8FFE0CB75F5C0FC07FA83FBD3" blockId="3.[189,1399,911,1114]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5C0FC07FA83FBD3" pageId="3" pageNumber="4">
− Mesosoma black dorsally and yellowish ventrally, or entirely black (
<figureCitation id="13712A5DFFE0CB75F6FCFC08FBAFFB8B" box="[1021,1115,999,1026]" captionStart="Fig. 2" captionStartId="5.[189,236,1948,1977]" captionTargetBox="[213,1375,263,1907]" captionTargetId="figure@5.[213,1375,263,1907]" captionText="Fig. 2. Nanium capitatum Townes, 1967, lectotype, Ƌ (ANSP). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468698" httpUri="https://zenodo.org/record/1468698/files/figure.png" pageId="3" pageNumber="4">Figs 2B</figureCitation>
, 
<figureCitation id="13712A5DFFE0CB75F164FC07FB7EFB8B" box="[1125,1162,1000,1026]" captionStart="Fig. 3" captionStartId="6.[189,236,1948,1977]" captionTargetBox="[218,1379,263,1905]" captionTargetId="figure@6.[218,1379,263,1905]" captionText="Fig. 3. Nanium capitatum Townes, 1967, ♀ (MZLU-HYM 00020508). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468700" httpUri="https://zenodo.org/record/1468700/files/figure.png" pageId="3" pageNumber="4">3C</figureCitation>
, 6C, 7C). Metasoma entirely black, with only T3 extensively pale, or with tergites black with hind margins narrowly yellow (
<figureCitation id="13712A5DFFE0CB75F44BFBD0FE5EFBD3" box="[330,426,1087,1114]" captionStart="Fig. 2" captionStartId="5.[189,236,1948,1977]" captionTargetBox="[213,1375,263,1907]" captionTargetId="figure@5.[213,1375,263,1907]" captionText="Fig. 2. Nanium capitatum Townes, 1967, lectotype, Ƌ (ANSP). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468698" httpUri="https://zenodo.org/record/1468698/files/figure.png" pageId="3" pageNumber="4">Figs 2B</figureCitation>
, 
<figureCitation id="13712A5DFFE0CB75F4B7FBAFFE2FFBD3" box="[438,475,1088,1114]" captionStart="Fig. 3" captionStartId="6.[189,236,1948,1977]" captionTargetBox="[218,1379,263,1905]" captionTargetId="figure@6.[218,1379,263,1905]" captionText="Fig. 3. Nanium capitatum Townes, 1967, ♀ (MZLU-HYM 00020508). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468700" httpUri="https://zenodo.org/record/1468700/files/figure.png" pageId="3" pageNumber="4">3B</figureCitation>
, 6B, 7B) ........................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BE2E7DFFE0CB75F5BCFB65FA83FB75" pageId="3" pageNumber="4">
<paragraph id="8BF536D8FFE0CB75F5BCFB65FA83FB59" blockId="3.[189,1399,1162,1282]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5BCFB65FA83FB59" pageId="3" pageNumber="4">
3. Metasoma with T3 extensively pale (
<figureCitation id="13712A5DFFE0CB75F797FB65FD02FB2D" box="[662,758,1162,1188]" captionStart="Fig. 2" captionStartId="5.[189,236,1948,1977]" captionTargetBox="[213,1375,263,1907]" captionTargetId="figure@5.[213,1375,263,1907]" captionText="Fig. 2. Nanium capitatum Townes, 1967, lectotype, Ƌ (ANSP). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468698" httpUri="https://zenodo.org/record/1468698/files/figure.png" pageId="3" pageNumber="4">Figs 2B</figureCitation>
, 
<figureCitation id="13712A5DFFE0CB75F603FB65FCD3FB2D" box="[770,807,1162,1188]" captionStart="Fig. 3" captionStartId="6.[189,236,1948,1977]" captionTargetBox="[218,1379,263,1905]" captionTargetId="figure@6.[218,1379,263,1905]" captionText="Fig. 3. Nanium capitatum Townes, 1967, ♀ (MZLU-HYM 00020508). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468700" httpUri="https://zenodo.org/record/1468700/files/figure.png" pageId="3" pageNumber="4">3B</figureCitation>
), and T2–4 uniformly dark (
<figureCitation id="13712A5DFFE0CB75F163FB65FB48FB2D" box="[1122,1212,1162,1188]" captionStart="Fig. 3" captionStartId="6.[189,236,1948,1977]" captionTargetBox="[218,1379,263,1905]" captionTargetId="figure@6.[218,1379,263,1905]" captionText="Fig. 3. Nanium capitatum Townes, 1967, ♀ (MZLU-HYM 00020508). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468700" httpUri="https://zenodo.org/record/1468700/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
) .................... .. .................................................................................................................. 
<taxonomicName id="4C4A4D5BFFE0CB75F12FFB59FA83FB59" authority="Townes, 1967" authorityName="Townes" authorityYear="1967" box="[1070,1399,1206,1232]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="capitatum">
<emphasis id="B93EEACAFFE0CB75F12FFB59FB3FFB59" bold="true" box="[1070,1227,1206,1232]" italics="true" pageId="3" pageNumber="4">N. capitatum</emphasis>
<bibRefCitation id="EFDB4B29FFE0CB75F1D3FB59FA83FB59" box="[1234,1399,1206,1232]" pageId="3" pageNumber="4" refId="ref5767" refString="Townes H. K. 1967. Genus Nanium Townes, n. genus. In: Krombein K. V. &amp; Burks B. D., Hymenoptera of America North of Mexico - Synoptic Catalog 2 (2 nd suppl.): 174 - 175. U. S. Dept of Agriculture Washington. https: // doi. org / 10.5962 / bhl. title. 4124" type="book chapter">Townes, 1967</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF536D8FFE0CB75F5C0FB0DFA83FB75" blockId="3.[189,1399,1162,1282]" box="[193,1399,1250,1282]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5C0FB0DFA83FB75" box="[193,1399,1250,1282]" pageId="3" pageNumber="4">
− Metasoma with T3 concolorous with T2–4 (
<figureCitation id="13712A5DFFE0CB75F7D4FB0DFCC2FB75" box="[725,822,1250,1276]" captionStart="Fig. 6" captionStartId="9.[189,236,1948,1977]" captionTargetBox="[189,1398,356,1906]" captionTargetId="figure@9.[189,1398,356,1906]" captionText="Fig. 6. Nanium nogueri Gauld, 1997, holotype, ♀ (BMNH). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468706" httpUri="https://zenodo.org/record/1468706/files/figure.png" pageId="3" pageNumber="4">Figs 6B</figureCitation>
, 
<figureCitation id="13712A5DFFE0CB75F643FB0DFC91FB75" box="[834,869,1250,1276]" captionStart="Fig. 7" captionStartId="10.[190,237,1948,1977]" captionTargetBox="[236,1399,343,1907]" captionTargetId="figure@10.[190,1399,343,1907]" captionText="Fig. 7. Nanium oriasi Gauld, 1997, holotype, ♀ (BMNH). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468708" httpUri="https://zenodo.org/record/1468708/files/figure.png" pageId="3" pageNumber="4">7B</figureCitation>
) ................................................................... 4
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BE2E7DFFE0CB75F5BCFAC3FA83FA43" pageId="3" pageNumber="4">
<paragraph id="8BF536D8FFE0CB75F5BCFAC3FA83FAFB" blockId="3.[189,1399,1324,1483]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5BCFAC3FA83FAFB" pageId="3" pageNumber="4">
4. Mesoscutum strongly polished and rather smooth. T1 about 1.5 × as long as broad posteriorly (
<figureCitation id="13712A5DFFE0CB75F5F2FAB7FEB9FAFA" box="[243,333,1368,1395]" captionStart="Fig. 6" captionStartId="9.[189,236,1948,1977]" captionTargetBox="[189,1398,356,1906]" captionTargetId="figure@9.[189,1398,356,1906]" captionText="Fig. 6. Nanium nogueri Gauld, 1997, holotype, ♀ (BMNH). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468706" httpUri="https://zenodo.org/record/1468706/files/figure.png" pageId="3" pageNumber="4">Fig. 6B</figureCitation>
) ........................................................................................................ 
<taxonomicName id="4C4A4D5BFFE0CB75F15DFAB7FA83FAFB" authority="Gauld, 1997" authorityName="Gauld" authorityYear="1997" box="[1116,1399,1368,1394]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="nogueri">
<emphasis id="B93EEACAFFE0CB75F15DFAB7FB2BFAFB" bold="true" box="[1116,1247,1368,1394]" italics="true" pageId="3" pageNumber="4">N. nogueri</emphasis>
<bibRefCitation id="EFDB4B29FFE0CB75F1E4FAB7FA83FAFB" box="[1253,1399,1368,1394]" pageId="3" pageNumber="4" refId="ref5568" refString="Gauld I. D. 1997. The Ichneumonidae of Costa Rica, 2. Memoirs of the American Entomological Institute 57: 1 - 485." type="journal article">Gauld, 1997</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF536D8FFE0CB75F5C0FA6AFA83FA43" blockId="3.[189,1399,1324,1483]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5C0FA6AFA83FA43" pageId="3" pageNumber="4">
− Mesoscutum granulate. T1 about 2.3 × as long as broad posteriorly (
<figureCitation id="13712A5DFFE0CB75F104FA6BFBABFA16" box="[1029,1119,1412,1439]" captionStart="Fig. 7" captionStartId="10.[190,237,1948,1977]" captionTargetBox="[236,1399,343,1907]" captionTargetId="figure@10.[190,1399,343,1907]" captionText="Fig. 7. Nanium oriasi Gauld, 1997, holotype, ♀ (BMNH). A. Face. B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468708" httpUri="https://zenodo.org/record/1468708/files/figure.png" pageId="3" pageNumber="4">Fig. 7B</figureCitation>
) .................................. .......................................................................................................................... 
<taxonomicName id="4C4A4D5BFFE0CB75F177FA5FFA83FA43" authority="Gauld, 1997" authorityName="Gauld" authorityYear="1997" box="[1142,1399,1456,1482]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oriasi">
<emphasis id="B93EEACAFFE0CB75F177FA5FFB2AFA43" bold="true" box="[1142,1246,1456,1482]" italics="true" pageId="3" pageNumber="4">N. oriasi</emphasis>
<bibRefCitation id="EFDB4B29FFE0CB75F1E4FA5FFA83FA43" box="[1253,1399,1456,1482]" pageId="3" pageNumber="4" refId="ref5568" refString="Gauld I. D. 1997. The Ichneumonidae of Costa Rica, 2. Memoirs of the American Entomological Institute 57: 1 - 485." type="journal article">Gauld, 1997</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BE2E7DFFE0CB75F5BCFA15FA83F911" pageId="3" pageNumber="4">
<paragraph id="8BF536D8FFE0CB75F5BCFA15FA83F9C9" blockId="3.[189,1399,1530,1689]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5BCFA15FA83F9C9" pageId="3" pageNumber="4">
5. Pleural carina distinct. T1 less than 2.4 × as long as broad posteriorly (
<figureCitation id="13712A5DFFE0CB75F12BFA15FB7CF99D" box="[1066,1160,1530,1556]" captionStart="Fig. 1" captionStartId="4.[189,236,1948,1977]" captionTargetBox="[234,1354,265,1906]" captionTargetId="figure@4.[234,1354,265,1908]" captionText="Fig. 1. Nanium atitlanensis Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (LLAMA#Ma-B-09-2-01 (ZMUT)). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468696" httpUri="https://zenodo.org/record/1468696/files/figure.png" pageId="3" pageNumber="4">Fig. 1A</figureCitation>
); hind coxa, at least partly, yellowish (
<figureCitation id="13712A5DFFE0CB75F4BFF9C9FDE2F9C9" box="[446,534,1574,1600]" captionStart="Fig. 1" captionStartId="4.[189,236,1948,1977]" captionTargetBox="[234,1354,265,1906]" captionTargetId="figure@4.[234,1354,265,1908]" captionText="Fig. 1. Nanium atitlanensis Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (LLAMA#Ma-B-09-2-01 (ZMUT)). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468696" httpUri="https://zenodo.org/record/1468696/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
) ................................................................................................................ 6
</keyLead>
</paragraph>
<paragraph id="8BF536D8FFE0CB75F5C0F9BCFA83F911" blockId="3.[189,1399,1530,1689]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5C0F9BCFA83F911" pageId="3" pageNumber="4">
− Pleural carina absent. Metasoma with T1 at least 2.8 × as long as broad posteriorly; hind coxa nearly entirely black .............................................................................................. 
<taxonomicName id="4C4A4D5BFFE0CB75F14CF990FA83F911" authority="Gauld, 1997" authorityName="Gauld" authorityYear="1997" box="[1101,1399,1662,1689]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="mairenai">
<emphasis id="B93EEACAFFE0CB75F14CF990FB2AF911" bold="true" box="[1101,1246,1662,1689]" italics="true" pageId="3" pageNumber="4">N. mairenai</emphasis>
<bibRefCitation id="EFDB4B29FFE0CB75F1E4F991FA83F911" box="[1253,1399,1662,1688]" pageId="3" pageNumber="4" refId="ref5568" refString="Gauld I. D. 1997. The Ichneumonidae of Costa Rica, 2. Memoirs of the American Entomological Institute 57: 1 - 485." type="journal article">Gauld, 1997</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30BE2E7DFFE0CB75F5BCF927FA83F862" pageId="3" pageNumber="4">
<paragraph id="8BF536D8FFE0CB75F5BCF927FA83F8B2" blockId="3.[189,1399,1736,2027]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5BCF927FA83F8B2" pageId="3" pageNumber="4">
6. Antenna with 26 flagellomeres. Hind tibia 9 × as long as broad apically. Ovipositor sheath diamond-shaped. Propleuron yellowish. T3 yellowish with dark anterior part. Hind coxa entirely pale yellow. Hypopygium pale yellow (
<figureCitation id="13712A5DFFE0CB75F724F8CEFD9FF8B2" box="[549,619,1825,1851]" captionStart="Fig. 4" captionStartId="7.[189,236,1948,1977]" captionTargetBox="[190,1397,287,1906]" captionTargetId="figure@7.[190,1397,287,1906]" captionText="Fig. 4. Nanium huberthi Gauld, 1997, holotype, ♀, habitus (MNCR)." figureDoi="http://doi.org/10.5281/zenodo.1468702" httpUri="https://zenodo.org/record/1468702/files/figure.png" pageId="3" pageNumber="4">Fig. 4</figureCitation>
) ................................................................ 
<taxonomicName id="4C4A4D5BFFE0CB75F153F8CEFA83F8B2" authority="Gauld, 1997" authorityName="Gauld" authorityYear="1997" box="[1106,1399,1824,1851]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="huberthi">
<emphasis id="B93EEACAFFE0CB75F153F8CEFB2AF8B2" bold="true" box="[1106,1246,1824,1851]" italics="true" pageId="3" pageNumber="4">N. huberthi</emphasis>
<bibRefCitation id="EFDB4B29FFE0CB75F1E4F8CEFA83F8B2" box="[1253,1399,1825,1851]" pageId="3" pageNumber="4" refId="ref5568" refString="Gauld I. D. 1997. The Ichneumonidae of Costa Rica, 2. Memoirs of the American Entomological Institute 57: 1 - 485." type="journal article">Gauld, 1997</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BF536D8FFE0CB75F5C0F8A2FA83F862" blockId="3.[189,1399,1736,2027]" pageId="3" pageNumber="4">
<keyLead id="30BB95EDFFE0CB75F5C0F8A2FA83F862" pageId="3" pageNumber="4">
− Antenna with 28 flagellomeres. Hind tibia 7 × as long as broad apically. Ovipositor sheath gradually expanding towards apex. Propleuron dark brown (
<figureCitation id="13712A5DFFE0CB75F645F896FC56F81A" box="[836,930,1913,1939]" captionStart="Fig. 1" captionStartId="4.[189,236,1948,1977]" captionTargetBox="[234,1354,265,1906]" captionTargetId="figure@4.[234,1354,265,1908]" captionText="Fig. 1. Nanium atitlanensis Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (LLAMA#Ma-B-09-2-01 (ZMUT)). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468696" httpUri="https://zenodo.org/record/1468696/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
). T3 entirely yellowish (
<figureCitation id="13712A5DFFE0CB75F1C7F896FAD7F81A" box="[1222,1315,1913,1939]" captionStart="Fig. 1" captionStartId="4.[189,236,1948,1977]" captionTargetBox="[234,1354,265,1906]" captionTargetId="figure@4.[234,1354,265,1908]" captionText="Fig. 1. Nanium atitlanensis Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (LLAMA#Ma-B-09-2-01 (ZMUT)). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468696" httpUri="https://zenodo.org/record/1468696/files/figure.png" pageId="3" pageNumber="4">Fig. 1B</figureCitation>
). Hind coxa dark brown with only its basodorsal part pale yellow (
<figureCitation id="13712A5DFFE0CB75F6CDF84AFBDFF836" box="[972,1067,1957,1983]" captionStart="Fig. 1" captionStartId="4.[189,236,1948,1977]" captionTargetBox="[234,1354,265,1906]" captionTargetId="figure@4.[234,1354,265,1908]" captionText="Fig. 1. Nanium atitlanensis Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (LLAMA#Ma-B-09-2-01 (ZMUT)). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468696" httpUri="https://zenodo.org/record/1468696/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
). Hypopygium dark brown (
<figureCitation id="13712A5DFFE0CB75F5F2F83EFEB8F862" box="[243,332,2001,2027]" captionStart="Fig. 1" captionStartId="4.[189,236,1948,1977]" captionTargetBox="[234,1354,265,1906]" captionTargetId="figure@4.[234,1354,265,1908]" captionText="Fig. 1. Nanium atitlanensis Reshchikov &amp; Sääksjärvi sp. nov., holotype, ♀ (LLAMA#Ma-B-09-2-01 (ZMUT)). A. Face (frontal view). B. Metasoma (dorsal view). C. Habitus." figureDoi="http://doi.org/10.5281/zenodo.1468696" httpUri="https://zenodo.org/record/1468696/files/figure.png" pageId="3" pageNumber="4">Fig. 1C</figureCitation>
) ................................................................ 
<taxonomicName id="4C4A4D5BFFE0CB75F634F83EFAE1F862" authority="Reshchikov &amp; Saaksjarvi" authorityName="Reshchikov &amp; Sääksjärvi &amp; Pollet" authorityYear="2018" box="[821,1301,2000,2027]" class="Insecta" family="Ichneumonidae" genus="Nanium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="atitlanensis" status="sp. nov.">
<emphasis id="B93EEACAFFE0CB75F634F83EFC10F862" bold="true" box="[821,996,2000,2027]" italics="true" pageId="3" pageNumber="4">N. atitlanensis</emphasis>
Reshchikov &amp; Sääksjärvi
</taxonomicName>
<taxonomicNameLabel id="A20D57B1FFE0CB75F01DF83EFA83F862" box="[1308,1399,2001,2027]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>