treatments-xml/data/03/DF/87/03DF87F1FFF6DB3DFF39FDB2FE70FDF6.xml

<document id="328B0FBEA0ACB9793425F85C6ADAF88A" ID-DOI="10.11646/phytotaxa.658.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217417" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722615130224" checkinUser="felipe" docAuthor="Averyanov, Leonid V., Nguyen, Van Canh, Vuong, Truong Ba, Nuraliev, Maxim S., Nguyen, Khang Sinh, Maisak, Tatiana V., Yudova, Daria A., Saidov, Nikita T., Nguyen, Cuong Huu &amp; Nong, Duy Van" docDate="2024" docId="03DF87F1FFF6DB3DFF39FDB2FE70FDF6" docLanguage="en" docName="phytotaxa.658.1.1.pdf" docOrigin="Phytotaxa 658 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.658.1.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Listera anthropophora Aver. &amp; V. C. Nguyen, sp. nov." docType="treatment" docVersion="3" lastPageNumber="4" masterDocId="FFE6FF89FFF4DB3EFFB1FFA6FFAAFFB3" masterDocTitle="New orchids in the flora of Vietnam VIII (Orchidaceae: Epidendroideae, tribes Malaxideae, Neottieae and Podochileae)" masterLastPageNumber="49" masterPageNumber="1" pageNumber="3" updateTime="1722811287782" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5E820E5E540D6E5777CA0872E3F73712">New orchids in the flora of Vietnam VIII (Orchidaceae: Epidendroideae, tribes Malaxideae, Neottieae and Podochileae)</mods:title>
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<mods:namePart id="4944183CD579914624B61E4EC4C478B6">Averyanov, Leonid V.</mods:namePart>
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<mods:namePart id="E3FB0F2C81B045BF555E91807C33B09D">Nguyen, Van Canh</mods:namePart>
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<mods:affiliation id="1D0F54DC5B4B527E6B7FF5C64E9ECA02">Institute of Applied Technology, Thu Dau Mot University, No. 6, Tran Van On Street, Phu Hoa Ward, Thu Dau Mot City, Binh Duong Province, Vietnam</mods:affiliation>
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<mods:namePart id="A9F4CA6C729200071443F682395EB382">Vuong, Truong Ba</mods:namePart>
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<mods:affiliation id="626F471D7C50D26BBBE2D5AC490D3B6D">Institute of Tropical Biology, Vietnam Academy of Science and Technology, 85, Tran Quoc Toan Street, District 3, Ho Chi Minh City, Vietnam</mods:affiliation>
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<mods:namePart id="AD2333790F63303465E65FD9B1860B53">Nuraliev, Maxim S.</mods:namePart>
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<mods:affiliation id="474E46FCD860A186ACDF059A338C50BD">Department of Higher Plants, Biological Faculty, M. V. Lomonosov Moscow State University, 1, 12, Leninskie Gory, 119234, Moscow, Russia &amp; Joint Russian-Vietnamese Tropical Scientific and Technological Center, 63, Nguyen Van Huyen Street, Cau Giay District, 122000, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="ACF48621CB622B126B6D9962AEAD82D7">Nguyen, Khang Sinh</mods:namePart>
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<mods:namePart id="D75049904E9C9156722DF3877F6740AB">Maisak, Tatiana V.</mods:namePart>
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<mods:affiliation id="7479F6EEB0A5C39AD13C3B028EC2A41E">Komarov Botanical Institute of the Russian Academy of Sciences, 2, Prof. Popov Street, 197022, St. Petersburg, Russia</mods:affiliation>
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<mods:namePart id="FFEA5799DE0F9AAE9DC85930B5377753">Yudova, Daria A.</mods:namePart>
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<mods:namePart id="67E804F6E583B3C10ADC697E2AB916F0">Saidov, Nikita T.</mods:namePart>
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<mods:namePart id="D058388ABF952BACAC8D7DCEE8F1CE6A">Nguyen, Cuong Huu</mods:namePart>
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<mods:namePart id="AA0312EA4ACAD2A783584B9DD98BE334">Nong, Duy Van</mods:namePart>
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<mods:affiliation id="6F082FA20D308DD159F55B08B00A9CB5">Tay Nguyen Institute for Scientific Research, Vietnam Academy of Science and Technology, Da Lat, Vietnam</mods:affiliation>
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<mods:date id="45ACA7E55BC7AE01646EEFEB7C24F97F">2024</mods:date>
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<taxonomicName id="4C764D64FFF6DB3CFF39FDB2FDD5FD9D" authority="Aver. &amp; V.C.Nguyen" authorityName="Aver. &amp; V. C. Nguyen" box="[136,639,532,559]" class="Liliopsida" family="Orchidaceae" genus="Listera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="anthropophora" status="sp. nov.">
<emphasis id="B902EAF5FFF6DB3CFF39FDB2FE24FD9C" bold="true" box="[136,398,532,559]" italics="true" pageId="2" pageNumber="3">Listera anthropophora</emphasis>
Aver. &amp; V.C.Nguyen
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, 
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(
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&amp; 
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)
</paragraph>
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<emphasis id="B902EAF5FFF6DB3CFF39FD25FF59FD2A" bold="true" box="[136,243,643,665]" pageId="2" pageNumber="3">Diagnosis:</emphasis>
—The new species differs from the closest 
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<emphasis id="B902EAF5FFF6DB3CFD20FD22FCF8FD2A" box="[657,850,644,665]" italics="true" pageId="2" pageNumber="3">Listera yunnanensis</emphasis>
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in glabrous peduncle, hispid rachis, glabrous pedicels, broader sepals (
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wide) and petals (
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wide), entirely glabrous lip, side lobes of lip erect, spathulate, broadening towards apex, and column with anther prominently bent back at 90°.
</paragraph>
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<emphasis id="B902EAF5FFF6DB3CFF39FD56FF68FCB6" bold="true" box="[136,194,752,773]" pageId="2" pageNumber="3">Type:</emphasis>
— 
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. 
<collectingRegion id="49B2F805FFF6DB3CFEE7FD56FDA9FCB5" box="[342,515,752,774]" country="Vietnam" name="Lao Cai" pageId="2" pageNumber="3">Lao Cai Province</collectingRegion>
: Bat Xat District, evergreen broad-leaved forest at elevation of about 
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a.s.l., living plants collected by unknown collector in 
<date id="FFC81027FFF6DB3CFDA3FCB2FD32FC9A" box="[530,664,788,810]" pageId="2" pageNumber="3" value="2023-10">October 2023</date>
, type herbarium specimen prepared on 
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, 
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Nguyen Van Canh, 
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. Averyanov, 
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. Maisak, 
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2439
</emphasis>
(
<typeStatus id="54CD8845FFF6DB3CFE44FC9EFDE1FCFD" box="[501,587,824,846]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
<collectionCode id="ED67AE22FFF6DB3CFDE2FC9EFD3DFCFD" box="[595,663,824,846]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="3">LE LE</collectionCode>
01253791! https://en.herbariumle.ru/?t=occ&amp;id=212990, photos of plant used for preparation of the type herbarium specimen 
<collectionCode id="ED67AE22FFF6DB3CFDEFFCFAFD36FCC1" box="[606,668,860,882]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="3">LE LE</collectionCode>
01124550 https://en.herbariumle.ru/?t=occ&amp;id=207739, drawing 
<collectionCode id="ED67AE22FFF6DB3CFABCFCFAFAE6FCC1" box="[1293,1356,860,882]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="2" pageNumber="3">LE LE</collectionCode>
01124776 https://en.herbariumle.ru/?t=occ&amp;id=213009).
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFF6DB3CFF39FC62FC42F9F1" pageId="2" pageNumber="3" type="description">
<paragraph id="8BC936E7FFF6DB3CFF39FC62FC42F9F1" blockId="2.[136,1452,964,2071]" pageId="2" pageNumber="3">
<emphasis id="B902EAF5FFF6DB3CFF39FC62FEDCFC6D" bold="true" box="[136,374,964,991]" pageId="2" pageNumber="3">Description:—Herb</emphasis>
terrestrial, autotrophic. 
<emphasis id="B902EAF5FFF6DB3CFD3EFC62FD53FC6D" bold="true" box="[655,761,964,990]" pageId="2" pageNumber="3">Rhizome</emphasis>
underground, thin, light brownish, glabrous, bearing a few distant, wiry, white roots. 
<emphasis id="B902EAF5FFF6DB3CFE1EFC4EFE46FBB1" bold="true" box="[431,492,1000,1026]" pageId="2" pageNumber="3">Stem</emphasis>
erect, light greenish to almost white, terete, 
<quantity id="4C8E9B02FFF6DB3CFC56FC4EFBFDFBB0" box="[999,1111,1000,1027]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="2" pageNumber="3" unit="cm" value="29.0" valueMax="32.0" valueMin="26.0">26–32 cm</quantity>
tall (including inflorescence), 
<quantity id="4C8E9B02FFF6DB3CFF39FBAAFF50FB94" box="[136,250,1036,1063]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="3" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
in diameter, glabrous, with 3–4 scarious thin tubular sheaths at base. 
<emphasis id="B902EAF5FFF6DB3CFBB2FBAAFBFFFB95" bold="true" box="[1027,1109,1036,1062]" pageId="2" pageNumber="3">Leaves</emphasis>
2, arranged at middle of stem, pseudo-opposite, sessile, grassy green adaxially, pale green to almost white abaxially, broadly ovate, 
<quantity id="4C8E9B02FFF6DB3CFB5EFB96FAC6FBF8" box="[1263,1388,1072,1099]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="2" pageNumber="3" unit="cm" value="5.0" valueMax="5.5" valueMin="4.5">4.5–5.5 cm</quantity>
long, 
<quantity id="4C8E9B02FFF6DB3CFF39FBF2FF45FBDC" box="[136,239,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="2" pageNumber="3" unit="cm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 cm</quantity>
wide, glabrous, base slightly cordate, apex acute or apiculate. 
<emphasis id="B902EAF5FFF6DB3CFC11FBF2FBA7FBDD" bold="true" box="[928,1037,1108,1134]" pageId="2" pageNumber="3">Peduncle</emphasis>
<quantity id="4C8E9B02FFF6DB3CFBA3FBF2FBCEFBDC" box="[1042,1124,1108,1135]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="2" pageNumber="3" unit="cm" value="7.5" valueMax="8.0" valueMin="7.0">7–8 cm</quantity>
long, glabrous, with 2 distant sterile bracts; bracts narrowly triangular, 
<quantity id="4C8E9B02FFF6DB3CFDD1FBDEFD67FB20" box="[608,717,1144,1171]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" metricValueMax="1.6" metricValueMin="0.5" pageId="2" pageNumber="3" unit="mm" value="10.5" valueMax="16.0" valueMin="5.0">5–16 mm</quantity>
long, glabrous, apex acuminate. 
<emphasis id="B902EAF5FFF6DB3CFBFBFBDEFB42FB21" bold="true" box="[1098,1256,1144,1170]" pageId="2" pageNumber="3">Inflorescence</emphasis>
a raceme; rachis 
<quantity id="4C8E9B02FFF6DB3CFF39FB3AFF77FB04" box="[136,221,1180,1207]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="2" pageNumber="3" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 cm</quantity>
long, hispid, laxly 10–15-flowered; floral bracts narrowly triangular, as long as pedicels or slightly shorter, glabrous. 
<emphasis id="B902EAF5FFF6DB3CFF49FB66FEE4FB69" bold="true" box="[248,334,1216,1242]" pageId="2" pageNumber="3">Pedicel</emphasis>
<quantity id="4C8E9B02FFF6DB3CFEE7FB66FE60FB68" box="[342,458,1216,1243]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.5" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="4.25" valueMax="4.5" valueMin="4.0">4–4.5 mm</quantity>
long, glabrous, prominently bent upwards near the apex. 
<emphasis id="B902EAF5FFF6DB3CFBECFB66FB03FB69" bold="true" box="[1117,1193,1216,1242]" pageId="2" pageNumber="3">Ovary</emphasis>
fusiform, about 
<quantity id="4C8E9B02FFF6DB3CFAD8FB66FA06FB68" box="[1385,1452,1216,1243]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="3" unit="mm" value="3.0">3 mm</quantity>
long, glabrous. 
<emphasis id="B902EAF5FFF6DB3CFE88FB42FE32FB4D" bold="true" box="[313,408,1252,1278]" pageId="2" pageNumber="3">Flowers</emphasis>
widely opening, glabrous; sepals and petals spreading, indistinctly 1-veined. 
<emphasis id="B902EAF5FFF6DB3CFABBFB42FA06FB4D" bold="true" box="[1290,1452,1252,1278]" pageId="2" pageNumber="3">Median sepal</emphasis>
narrowly ovate, 
<quantity id="4C8E9B02FFF6DB3CFE8EFAAEFE1BFA90" box="[319,433,1288,1315]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="3" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
long, 
<quantity id="4C8E9B02FFF6DB3CFE46FAAEFDD4FA90" box="[503,638,1288,1315]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="3" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
wide, blunt to obtuse at apex. 
<emphasis id="B902EAF5FFF6DB3CFC64FAAEFBD6FA91" bold="true" box="[981,1148,1288,1314]" pageId="2" pageNumber="3">Lateral sepals</emphasis>
narrowly ovate, somewhat oblique, 
<quantity id="4C8E9B02FFF6DB3CFF5BFA8AFEF1FAF4" box="[234,347,1324,1351]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" metricValueMax="3.2" metricValueMin="3.0" pageId="2" pageNumber="3" unit="mm" value="3.1" valueMax="3.2" valueMin="3.0">3–3.2 mm</quantity>
long, 
<quantity id="4C8E9B02FFF6DB3CFE10FA8AFD82FAF4" box="[417,552,1324,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="3" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
wide, obtuse. 
<emphasis id="B902EAF5FFF6DB3CFD78FA8AFCBBFAF5" bold="true" box="[713,785,1324,1350]" pageId="2" pageNumber="3">Petals</emphasis>
oblong, slightly oblique, as long as sepals, 
<quantity id="4C8E9B02FFF6DB3CFB47FA8AFAC2FAF4" box="[1270,1384,1324,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="2" pageNumber="3" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
wide, blunt to roundish at apex. 
<emphasis id="B902EAF5FFF6DB3CFE1DFAF6FE7CFAD9" bold="true" box="[428,470,1360,1386]" pageId="2" pageNumber="3">Lip</emphasis>
narrowly obovate in outline, 
<quantity id="4C8E9B02FFF6DB3CFC91FAF6FC0DFAD8" box="[800,935,1360,1387]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="8.5" metricValueMin="7.5" pageId="2" pageNumber="3" unit="mm" value="8.0" valueMax="8.5" valueMin="7.5">7.5–8.5 mm</quantity>
long, 
<quantity id="4C8E9B02FFF6DB3CFC5CFAF6FBF4FAD8" box="[1005,1118,1360,1387]" metricMagnitude="-3" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="2" pageNumber="3" unit="mm" value="2.75" valueMax="3.0" valueMin="2.5">2.5–3 mm</quantity>
wide, at base with side lobes, median lobe apex deeply 2-lobuled; side lobes directed towards lip base, erect, spathulate, broadened towards apex, 
<quantity id="4C8E9B02FFF6DB3CFF39FA3EFEBAFA00" box="[136,272,1432,1459]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="3" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
long, 
<quantity id="4C8E9B02FFF6DB3CFEE8FA3EFE4BFA00" box="[345,481,1432,1459]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
wide; lobules of median lobe diverging at a sharp angle, narrowly triangular, slightly falcate inwards, acuminate to shortly caudate, 
<quantity id="4C8E9B02FFF6DB3CFD22FA1AFCB1FA64" box="[659,795,1468,1495]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.2" metricValueMin="3.8" pageId="2" pageNumber="3" unit="mm" value="4.0" valueMax="4.2" valueMin="3.8">3.8–4.2 mm</quantity>
long; disk fleshy, at center with 2 short, low, fleshy keels, near base with 2 lateral fold-like teeth placed more proximally than side lobes. 
<emphasis id="B902EAF5FFF6DB3CFB85FA46FB3EFA49" bold="true" box="[1076,1172,1504,1530]" pageId="2" pageNumber="3">Column</emphasis>
<quantity id="4C8E9B02FFF6DB3CFB11FA46FABCFA48" box="[1184,1302,1504,1531]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="2" pageNumber="3" unit="mm" value="0.9" valueMax="1.0" valueMin="0.8">0.8–1 mm</quantity>
tall on stalk 
<quantity id="4C8E9B02FFF6DB3CFF39F9A2FEA4F9AC" box="[136,270,1540,1567]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
long; stigma orbicular; rostellum insignificant; anther narrowly ovoid, 
<quantity id="4C8E9B02FFF6DB3CFB97F9A2FB06F9AC" box="[1062,1196,1540,1567]" metricMagnitude="-4" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="2" pageNumber="3" unit="mm" value="0.45" valueMax="0.5" valueMin="0.4">0.4–0.5 mm</quantity>
long, bent back at 90°. 
<emphasis id="B902EAF5FFF6DB3CFF39F98EFF42F9F1" bold="true" box="[136,232,1576,1602]" pageId="2" pageNumber="3">Capsule</emphasis>
narrowly obovoid, 6-ribbed, 
<quantity id="4C8E9B02FFF6DB3CFD83F98EFD0EF9F0" box="[562,676,1576,1603]" metricMagnitude="-3" metricUnit="m" metricValue="6.75" metricValueMax="7.0" metricValueMin="6.5" pageId="2" pageNumber="3" unit="mm" value="6.75" valueMax="7.0" valueMin="6.5">6.5–7 mm</quantity>
long, 
<quantity id="4C8E9B02FFF6DB3CFD5AF98EFCF7F9F0" box="[747,861,1576,1603]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="2" pageNumber="3" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 mm</quantity>
in diameter.
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFF6DB3CFF0CF9EAFBD4F91D" pageId="2" pageNumber="3" type="etymology">
<paragraph id="8BC936E7FFF6DB3CFF0CF9EAFB20F9D5" blockId="2.[136,1452,964,2071]" box="[189,1162,1612,1639]" pageId="2" pageNumber="3">
<emphasis id="B902EAF5FFF6DB3CFF0CF9EAFEE2F9D5" bold="true" box="[189,328,1612,1638]" pageId="2" pageNumber="3">Etymology:</emphasis>
—The species name refers to the anthropomorphic shape of the flower lip.
</paragraph>
<paragraph id="8BC936E7FFF6DB3CFF0CF9D6FBD4F91D" blockId="2.[136,1452,964,2071]" pageId="2" pageNumber="3">
<emphasis id="B902EAF5FFF6DB3CFF0CF9D6FE72F939" bold="true" box="[189,472,1648,1674]" pageId="2" pageNumber="3">Habitat and phenology:</emphasis>
—Terrestrial rhizomatous herb. Evergreen broad-leaved forest on granite at elevation of about 
<quantity id="4C8E9B02FFF6DB3CFF7CF932FE88F91C" box="[205,290,1684,1711]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="3" unit="m" value="2500.0">2500 m</quantity>
a.s.l., on wet mossy ground. Very rare. Flowers likely in August–September.
</paragraph>
</subSubSection>
<subSubSection id="C36C656CFFF6DB3DFF0CF91EFE70FDF6" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="distribution">
<paragraph id="8BC936E7FFF6DB3CFF0CF91EFB9FF961" blockId="2.[136,1452,964,2071]" box="[189,1077,1720,1747]" pageId="2" pageNumber="3">
<emphasis id="B902EAF5FFF6DB3CFF0CF91EFEF3F961" bold="true" box="[189,345,1720,1746]" pageId="2" pageNumber="3">Distribution:</emphasis>
—Endemic to NW 
<collectingCountry id="F3617677FFF6DB3CFD80F91EFD3AF961" box="[561,656,1720,1746]" name="Vietnam" pageId="2" pageNumber="3">Vietnam</collectingCountry>
(
<collectingRegion id="49B2F805FFF6DB3CFD2FF91EFCC2F961" box="[670,872,1720,1746]" country="Vietnam" name="Lao Cai" pageId="2" pageNumber="3">Lao Cai Province</collectingRegion>
: Bat Xat District).
</paragraph>
<paragraph id="8BC936E7FFF6DB3DFF0CF97AFCEAFF6F" blockId="2.[136,1452,964,2071]" lastBlockId="3.[136,1452,158,581]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="B902EAF5FFF6DB3CFF0CF97AFE18F945" bold="true" box="[189,434,1756,1782]" pageId="2" pageNumber="3">Conservation status:</emphasis>
— 
<taxonomicName id="4C764D64FFF6DB3CFE7FF97BFD61F945" box="[462,715,1756,1782]" class="Liliopsida" family="Orchidaceae" genus="Listera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="anthropophora">
<emphasis id="B902EAF5FFF6DB3CFE7FF97BFD61F945" box="[462,715,1756,1782]" italics="true" pageId="2" pageNumber="3">Listera anthropophora</emphasis>
</taxonomicName>
is known from a single location. Taking into consideration the very small area of its expected occurrence and extensive selected logging in this area, we estimate the conservation status of this species as globally Critically Endangered (CR) following the formal criteria proposed by the 
<bibRefCitation id="EFE74B16FFF6DB3CFB0FF882FAF2F88D" author="IUCN Standards &amp; Petitions Committee" box="[1214,1368,1828,1854]" pageId="2" pageNumber="3" refId="ref31315" refString="IUCN Standards and Petitions Committee (2024) Guidelines for Using the IUCN Red List Categories and Criteria. Version 15.1. Prepared by the Standards and Petitions Committee. Available from: https: // nc. iucnredlist. org / redlist / content / attachment _ files / RedListGuidelines. pdf (accessed 20 February 2024)." type="book" year="2024">IUCN (2024)</bibRefCitation>
: A2a,b; B1a,b(i–iii)+2a,b(i–iii); C1+2(i,ii). In details, we estimate (within the statements employed by the IUCN):—expected population reduction for 10 years or 3 generation ≥ 80% is not reversible (A2), based on direct observation (a) and observed decline of the area of occupancy, extent of occurrence and habitat quality (b);—the area of occurrence &lt;
<quantity id="4C8E9B02FFF6DB3CFA30F836FF06F863" metricMagnitude="5" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="km" value="100.0">100 km</quantity>
² (B1) and the area of occupancy &lt;
<quantity id="4C8E9B02FFF6DB3CFD8CF812FD2EF863" box="[573,644,1972,2000]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="3" unit="km" value="10.0">10 km</quantity>
² (B2), with 1 known population (a) and continuing observed decline of extent of occurrence (bi); the area of occupancy (bii); area, quality of habitat (biii);—the number of mature individuals &lt;250, estimated or projected continuing decline in 25% in 3 years or 1 generation (C1), and observed, estimated, projected or inferred continuing decline when the number of mature individuals in each subpopulation (C2(i)) ≤ 50 and 90–100% of mature individuals in one subpopulation (C2(ii)).
</paragraph>
<paragraph id="8BC936E7FFF7DB3DFF0CFF40FE70FDF6" blockId="3.[136,1452,158,581]" pageId="3" pageNumber="4">
<emphasis id="B902EAF5FFF7DB3DFF0CFF40FEA3FEB3" bold="true" box="[189,265,230,256]" pageId="3" pageNumber="4">Notes:</emphasis>
— 
<taxonomicName id="4C764D64FFF7DB3DFE97FF41FD8CFEB3" box="[294,550,230,256]" class="Liliopsida" family="Orchidaceae" genus="Listera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="anthropophora">
<emphasis id="B902EAF5FFF7DB3DFE97FF41FD8CFEB3" box="[294,550,230,256]" italics="true" pageId="3" pageNumber="4">Listera anthropophora</emphasis>
</taxonomicName>
is morphologically very close to 
<taxonomicName id="4C764D64FFF7DB3DFC10FF41FAD6FEB2" authority="S. C. Chen (1981: 759" authorityName="S. C. Chen" authorityYear="1981" box="[929,1404,230,257]" class="Liliopsida" family="Orchidaceae" genus="Listera" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="4" phylum="Tracheophyta" rank="species" species="yunnanensis">
<emphasis id="B902EAF5FFF7DB3DFC10FF41FB28FEB3" box="[929,1154,231,256]" italics="true" pageId="3" pageNumber="4">Listera yunnanensis</emphasis>
S.C. 
<bibRefCitation id="EFE74B16FFF7DB3DFB05FF40FAD6FEB2" author="Chen, S. - C." box="[1204,1404,230,257]" pageId="3" pageNumber="4" pagination="759 - 762" refId="ref29743" refString="Chen, S. - C. (1981) Three new species of Listera (Orchidaceae) from China. Kew Bulletin 35 (4): 759 - 762. https: // doi. org / 10.2307 / 4110171" type="journal article" year="1981">Chen (1981: 759</bibRefCitation>
</taxonomicName>
, see also 
<bibRefCitation id="EFE74B16FFF7DB3DFF0FFEACFE2AFE97" author="Chen, S. C. &amp; Liu, Z. J. &amp; Zhu, G. H. &amp; Lang, K. Y. &amp; Ji, Z. H. &amp; Luo, Y. B. &amp; Jin, X. H. &amp; Cribb, P. J. &amp; Wood, J. J. &amp; Gale, S. W. &amp; Ormerod, P. &amp; Vermeulen, J. J. &amp; Wood, H. P. &amp; Clayton, D. &amp; Bell, A." box="[190,384,266,292]" pageId="3" pageNumber="4" pagination="211 - 235" refId="ref29820" refString="Chen, S. C., Liu, Z. J., Zhu, G. H., Lang, K. Y., Ji, Z. H., Luo, Y. B., Jin, X. H., Cribb, P. J., Wood, J. J., Gale, S. W., Ormerod, P., Vermeulen, J. J., Wood, H. P., Clayton, D. &amp; Bell, A. (2009) Orchidaceae. In: Wu, Z. Y., Raven, P. H. &amp; Hong, D. (Eds.) Flora of China, vol. 25. Science Press, Beijing &amp; Missouri Botanical Garden Press, St. Louis, pp. 211 - 235." type="book chapter" year="2009">
Chen 
<emphasis id="B902EAF5FFF7DB3DFEB3FEADFE96FE97" box="[258,316,266,292]" italics="true" pageId="3" pageNumber="4">et al.</emphasis>
2009
</bibRefCitation>
), but distinctly differs from the latter species in shorter stem 
<quantity id="4C8E9B02FFF7DB3DFB8CFEACFB04FE96" box="[1085,1198,266,293]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" metricValueMax="3.2" metricValueMin="2.6" pageId="3" pageNumber="4" unit="cm" value="29.0" valueMax="32.0" valueMin="26.0">26–32 cm</quantity>
(vs. 
<quantity id="4C8E9B02FFF7DB3DFB59FEACFADAFE97" box="[1256,1392,266,293]" metricMagnitude="-1" metricUnit="m" metricValue="4.225" metricValueMax="4.75" metricValueMin="3.7" pageId="3" pageNumber="4" unit="cm" value="42.25" valueMax="47.5" valueMin="37.0">37–47.5 cm</quantity>
) tall, glabrous (vs. pubescent) peduncle, rachis 
<quantity id="4C8E9B02FFF7DB3DFDECFE88FD1AFEFA" box="[605,688,302,329]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="3" pageNumber="4" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 cm</quantity>
long, hispid, 10–15-flowered (vs. rachis 
<quantity id="4C8E9B02FFF7DB3DFBCFFE88FB47FEFA" box="[1150,1261,302,329]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="3" pageNumber="4" unit="cm" value="15.0" valueMax="16.0" valueMin="14.0">14–16 cm</quantity>
long, pubescent, 24–35-flowered), glabrous (vs. sparsely pubescent) pedicel, broader sepals (
<quantity id="4C8E9B02FFF7DB3DFC49FEF4FB2EFEDE" box="[1016,1156,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="3" pageNumber="4" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
vs. 
<quantity id="4C8E9B02FFF7DB3DFB08FEF4FA84FEDE" box="[1209,1326,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.3" metricValueMin="1.0" pageId="3" pageNumber="4" unit="mm" value="1.15" valueMax="1.3" valueMin="1.0">1–1.3 mm</quantity>
wide) and petals (
<quantity id="4C8E9B02FFF7DB3DFF68FED0FEE6FE22" box="[217,332,374,401]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="3" pageNumber="4" unit="mm" value="1.1" valueMax="1.2" valueMin="1.0">1–1.2 mm</quantity>
vs. 
<quantity id="4C8E9B02FFF7DB3DFEC8FED0FDABFE22" box="[377,513,374,401]" metricMagnitude="-4" metricUnit="m" metricValue="5.5" metricValueMax="6.0" metricValueMin="5.0" pageId="3" pageNumber="4" unit="mm" value="0.55" valueMax="0.6" valueMin="0.5">0.5–0.6 mm</quantity>
wide), entirely glabrous lip (vs. lip minutely ciliate along the margin), side lobes of lip erect, spathulate, broadening to the apex, 
<quantity id="4C8E9B02FFF7DB3DFD31FE3CFCADFE06" box="[640,775,410,437]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="3" pageNumber="4" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.4–1.6 mm</quantity>
long, 
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wide (vs. side lobes horizontally or down directed, ovate-oblong, tapering to the apex, about 
<quantity id="4C8E9B02FFF7DB3DFD0DFE18FCBBFE6A" box="[700,785,446,473]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="3" pageNumber="4" unit="mm" value="1.3">1.3 mm</quantity>
long, 
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wide), disk with 2 short, low, fleshy keels (vs. disk with 2 swollen, suborbicular lamellae), and anther prominently bent back at 90° (vs. anther erect to suberect). The prominent erect spathulate side lobes of lip broadened towards the apex is a unique feature that is unknown from all the other species of the genus.
</paragraph>
</subSubSection>
</treatment>
</document>