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<document id="7FD6FDEB56B28C6D40C8EA6F4A874635" ID-CLB-Dataset="22084" ID-DOI="10.5852/ejt.2018.446" ID-GBIF-Dataset="08fb092e-b3df-47c4-a1ad-8ea8ffd2457a" ID-Zenodo-Dep="3829902" ID-ZooBank="56E50F4E-6A7E-4CE6-963E-3B49AA7A03B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589575450896" checkinUser="valdenar" docAuthor="Schmidt, Joachim &amp; Faille, Arnaud" docDate="2018" docId="03DF87A4FF84FFE3FF5BF85FFAD95B4D" docLanguage="en" docName="ejt-446_schmidt_faille.pdf.imf" docOrigin="European Journal of Taxonomy 446" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Trechus Clairville 1806" docType="key" docVersion="6" lastPageNumber="79" masterDocId="FFE6FFDCFFCFFFADFFE6F953FFAE5C66" masterDocTitle="Revision of Trechus Clairville, 1806 of the Bale Mountains and adjacent volcanos, Ethiopia (Coleoptera, Carabidae, Trechini)" masterLastPageNumber="82" masterPageNumber="1" pageNumber="76" updateTime="1698839989887" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="EB8038B93182E59AE84979967DB4F9E4">Revision of Trechus Clairville, 1806 of the Bale Mountains and adjacent volcanos, Ethiopia (Coleoptera, Carabidae, Trechini)</mods:title>
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<mods:namePart id="E72E01E4D55FA68A8666379C24403C48">Schmidt, Joachim</mods:namePart>
<mods:affiliation id="E540C68B5F0B0D8754EDFE9FC0A109A1">University of Rostock, Institute of Biosciences, General and Systematic Zoology, Universitätsplatz 2, 18055 Rostock, Germany. University of Marburg, Fb. 17 - Biologie, Karl-von-Frisch-Strasse 8, 35043 Marburg, Germany. Institute of Evolutionary Biology (CSIC-Universitat Pompeu Fabra), Passeig Maritim de la Barceloneta 37, 08003 Barcelona, Spain. MECADEV - UMR 7179 MNHN / CNRS, Paris, France.</mods:affiliation>
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<mods:namePart id="9CC67C6D2F7FA1442F5C313D8E54A095">Faille, Arnaud</mods:namePart>
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<mods:title id="4BC58CA9FDE0C2688BD214C0FDEF11E5">European Journal of Taxonomy</mods:title>
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<mods:date id="C881398AEB96AE66D2FFE47D26B0C1F8">2018</mods:date>
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<subSection id="E2F92F59FF84FFE3FF5BF85FFAD95B4D" lastPageId="78" lastPageNumber="79" pageId="75" pageNumber="76" type="key">
<treatment id="03DF87A4FF84FFE3FF5BF85FFAD95B4D" ID-DOI="http://doi.org/10.5281/zenodo.3848495" ID-GBIF-Taxon="164165531" ID-Zenodo-Dep="3848495" LSID="urn:lsid:plazi:treatment:03DF87A4FF84FFE3FF5BF85FFAD95B4D" httpUri="http://treatment.plazi.org/id/03DF87A4FF84FFE3FF5BF85FFAD95B4D" lastPageId="78" lastPageNumber="79" pageId="75" pageNumber="76">
<subSubSection id="C36C6539FF84FFE6FF5BF85FFAA55D4C" box="[189,1291,267,299]" pageId="75" pageNumber="76" type="nomenclature">
<paragraph id="8BC936B2FF84FFE6FF5BF85FFAA55D4C" blockId="75.[189,1291,267,299]" box="[189,1291,267,299]" pageId="75" pageNumber="76">
<heading id="D08181DEFF84FFE6FF5BF85FFAA55D4C" bold="true" box="[189,1291,267,299]" fontSize="13" level="1" pageId="75" pageNumber="76" reason="1">
<emphasis id="B902EAA0FF84FFE6FF5BF85FFAA55D4C" bold="true" box="[189,1291,267,299]" pageId="75" pageNumber="76">
<emphasis id="B902EAA0FF84FFE6FF5BF85FFE705D4C" bold="true" box="[189,478,267,298]" italics="true" pageId="75" pageNumber="76">Key to the species of</emphasis>
<taxonomicName id="4C764D31FF84FFE6FE00F85FFDF25D4D" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[486,604,268,299]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">Trechus</taxonomicName>
<emphasis id="B902EAA0FF84FFE6FD83F858FAA55D4C" bold="true" box="[613,1291,267,298]" italics="true" pageId="75" pageNumber="76">
from the Arussi and Bale Provinces of 
<collectingCountry id="F3617622FF84FFE6FB76F858FAA55D4C" box="[1168,1291,267,298]" name="Ethiopia" pageId="75" pageNumber="76">Ethiopia</collectingCountry>
</emphasis>
</emphasis>
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</paragraph>
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<key id="FFE76183FF84FFE3FF5BF812FAD95B4D" lastPageId="78" lastPageNumber="79" pageId="75" pageNumber="76">
<paragraph id="8BC936B2FF84FFE6FF5BF812FAD85D18" blockId="75.[189,1399,321,383]" pageId="75" pageNumber="76">
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<keyLead id="30879587FF84FFE6FF5BF812FAD85D18" pageId="75" pageNumber="76">1. Males with only one basal protarsomere dilated .............................................................................. 2 – Males with two basal protarsomeres dilated ................................................................................... 10</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF84FFE6FF5BF8F5FAD95E4C" pageId="75" pageNumber="76">
<paragraph id="8BC936B2FF84FFE6FF5BF8F5FAD85D85" blockId="75.[189,1399,422,554]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BF8F5FAD85D85" pageId="75" pageNumber="76">2. Disc of elytra with at least 4–5 internal striae distinctly engraved. Antennae slenderer with antennomeres 7–9 1.6–1.8 times as long as wide. Species from southern slope of Bale Mts …3</keyLead>
</paragraph>
<paragraph id="8BC936B2FF84FFE6FF27F8BEFAD95E4C" blockId="75.[189,1399,422,554]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF27F8BEFAD95E4C" pageId="75" pageNumber="76">– Elytral discal striae very finely engraved, usually indistinct with first stria often absent. Antennae short with antennomeres 7–9 1.2–1.4 times as long as wide ............................................................ 4</keyLead>
</paragraph>
</keyStep>
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3. Each elytron with a single dorsal seta (both posterior discal seta on third stria and preapical seta are absent). Pronotum with laterobasal angles very obtuse, blunt at tip, and with laterobasal impressions indistinct; lateral gutter of pronotum not widened near base ................... 
<taxonomicName id="4C764D31FF84FFE6FBA6FBCBFABB5ED4" ID-CoL="8M78L" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1088,1301,663,690]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="transversicollis" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FBA6FBCBFABB5ED4" bold="true" box="[1088,1301,663,690]" italics="true" pageId="75" pageNumber="76">T. transversicollis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFBCBFAD95ED4" box="[1308,1399,664,690]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
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<keyLead id="30879587FF84FFE6FF27FBEFFAD95F7A" pageId="75" pageNumber="76">
– Two elytral discal setae present. Pronotum with laterobasal angles slightly obtuse, sharp at tip, and with laterobasal impressions broad; lateral gutter of pronotum distinctly widened towards base ......................................................................................................................... 
<taxonomicName id="4C764D31FF84FFE6FB40FA51FABB5F7A" ID-CoL="8MNFL" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1190,1301,770,796]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="mattisi" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB40FA51FABB5F7A" bold="true" box="[1190,1301,770,796]" italics="true" pageId="75" pageNumber="76">T. mattisi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFA51FAD95F7A" box="[1308,1399,770,796]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BC936B2FF84FFE6FF5BFA17FAD95FE4" blockId="75.[189,1399,835,968]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFA17FAD95FE4" pageId="75" pageNumber="76">4. Elytra with preapical seta absent. Species of Mt Enkuolo with body colour of mature specimens light brown ................................................................................................................................................ 5</keyLead>
</paragraph>
<paragraph id="8BC936B2FF84FFE6FF27FAD8FAD95FA1" blockId="75.[189,1399,835,968]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF27FAD8FAD95FA1" pageId="75" pageNumber="76">– Elytra with preapical seta present (situated at the apical anastomosis of second and third striae). Species of the Bale Mts; body colour of mature specimens usually brown to dark brown .............. 6</keyLead>
</paragraph>
</keyStep>
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<keyStep id="30822E17FF84FFE6FF5BFABCFAD95836" pageId="75" pageNumber="76">
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5. Each elytron with two dorsal setae on third stria in normal position ................................................. ........................................................................................ 
<taxonomicName id="4C764D31FF84FFE6FC9FFD40FAD9584A" ID-CoL="57ZMR" authority="Vigna Taglianti &amp; Magrini, 2010" authorityName="Vigna Taglianti &amp; Magrini" authorityYear="2010" box="[889,1399,1042,1069]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="gypaeti">
<emphasis id="B902EAA0FF84FFE6FC9FFD40FC43584A" bold="true" box="[889,1005,1042,1069]" italics="true" pageId="75" pageNumber="76">T. gypaeti</emphasis>
Vigna Taglianti &amp; Magrini, 2010
</taxonomicName>
– The posterior elytral dorsal seta is lacking ................................................... 
<taxonomicName id="4C764D31FF84FFE6FB80FD65FABB5836" ID-CoL="8MF2N" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1126,1301,1077,1104]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="minitrechus" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB80FD65FABB5836" bold="true" box="[1126,1301,1077,1104]" italics="true" pageId="75" pageNumber="76">T. minitrechus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFD65FAD95836" box="[1308,1399,1078,1104]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="30822E17FF84FFE6FF5BFD24FAD958BE" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFD24FAD958BE" pageId="75" pageNumber="76">
6. Apex of terminal lamella of aedeagal median lobe simply tapered or with a small globose apical capitulum .......................................................................................................................................... 7 – Median lobe of aedeagus with a distinct button-like apical capitulum (
<figureCitation id="134D2A37FF84FFE6FBC6FDEDFB7C58BE" box="[1056,1234,1214,1241]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Figs 45, 47, 48</figureCitation>
) ................. 8
</keyLead>
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</paragraph>
<paragraph id="8BC936B2FF84FFE6FF5BFC53FAD95906" blockId="75.[189,1399,1279,1376]" pageId="75" pageNumber="76">
<keyStep id="30822E17FF84FFE6FF5BFC53FAD95906" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFC53FAD95906" pageId="75" pageNumber="76">
7. Eyes slightly larger than tempora. Aedeagal median lobe stouter (
<figureCitation id="134D2A37FF84FFE6FC0FFDACFB96597F" box="[1001,1080,1279,1305]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Fig 43</figureCitation>
) .......... 
<taxonomicName id="4C764D31FF84FFE6FB72FC53FABA597C" ID-CoL="8MJ7S" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1172,1300,1279,1306]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="nanulus" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB72FC53FABA597C" bold="true" box="[1172,1300,1279,1306]" italics="true" pageId="75" pageNumber="76">T. nanulus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFC53FAD9597C" box="[1308,1399,1280,1306]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
– Eyes slightly smaller than tempora. Aedeagal median lobe more elongated (
<figureCitation id="134D2A37FF84FFE6FB87FC70FB1F595B" box="[1121,1201,1315,1341]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Fig 44</figureCitation>
) ........................ ...................................................................................................................................... 
<taxonomicName id="4C764D31FF84FFE6FB2FFC15FABB5906" ID-CoL="8MCQC" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1225,1301,1350,1376]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="rira" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB2FFC15FABB5906" bold="true" box="[1225,1301,1350,1376]" italics="true" pageId="75" pageNumber="76">T. rira</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFC15FAD95906" box="[1308,1399,1350,1376]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
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</paragraph>
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<keyStep id="30822E17FF84FFE6FF5BFCDBFAD9598E" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFCDBFAD9598E" pageId="75" pageNumber="76">
8. Aedeagal median lobe in lateral view with ventral margin sinusoidal (
<figureCitation id="134D2A37FF84FFE6FBFAFCD4FBC559C4" box="[1052,1131,1415,1442]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Fig 47</figureCitation>
) ................................. ...................................................................... 
<taxonomicName id="4C764D31FF84FFE6FD11FCF8FAD959A3" ID-CoL="8MBZG" authority="Magrini, Queinnec &amp; Vigna Taglianti, 2012" authorityName="Magrini, Queinnec &amp; Vigna Taglianti" authorityYear="2012" box="[759,1399,1450,1477]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="relictus">
<emphasis id="B902EAA0FF84FFE6FD11FCF8FCC159A3" bold="true" box="[759,879,1450,1477]" italics="true" pageId="75" pageNumber="76">T. relictus</emphasis>
<bibRefCitation id="EFE74B43FF84FFE6FC90FCF8FAD959A3" author="Magrini P. &amp; Queinnec E. &amp; Vigna Taglianti A." box="[886,1399,1451,1477]" pageId="75" pageNumber="76" pagination="19 - 32" refId="ref36764" refString="Magrini P., Queinnec E. &amp; Vigna Taglianti A. 2012. A new subgenus and two new species of Trechus from Ethiopia (Coleoptera, Carabidae). Fragmenta entomologica 44 (1): 19 - 32. https: // doi. org / 10.4081 / fe. 2012.30" type="journal article" year="2012">Magrini, Quéinnec &amp; Vigna Taglianti, 2012</bibRefCitation>
</taxonomicName>
– Aedeagal median lobe in lateral view with ventral margin concave or straight ............................... 9
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BC936B2FF84FFE6FF5BFF43FAD95A28" blockId="75.[189,1399,1552,1614]" pageId="75" pageNumber="76">
<keyStep id="30822E17FF84FFE6FF5BFF43FAD95A28" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFF43FAD95A28" pageId="75" pageNumber="76">
9. Aedeagal median lobe robust in view from laterad and dorsad (
<figureCitation id="134D2A37FF84FFE6FC28FF43FBFB5A4C" box="[974,1109,1552,1578]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Figs 48–49</figureCitation>
) .... 
<taxonomicName id="4C764D31FF84FFE6FB63FF43FABB5A4C" ID-CoL="8MKKX" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1157,1301,1552,1578]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="oppositus" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB63FF43FABB5A4C" bold="true" box="[1157,1301,1552,1578]" italics="true" pageId="75" pageNumber="76">T. oppositus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFF43FAD95A4C" box="[1308,1399,1552,1578]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
– Aedeagal median lobe slender in view from laterad and dorsad (
<figureCitation id="134D2A37FF84FFE6FC3FFF60FBCE5A28" box="[985,1120,1587,1614]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Figs 45–46</figureCitation>
) .. 
<taxonomicName id="4C764D31FF84FFE6FB98FF60FABB5A28" ID-CoL="8M8KW" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1150,1301,1587,1614]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="iridescens" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB98FF60FABB5A28" bold="true" box="[1150,1301,1587,1614]" italics="true" pageId="75" pageNumber="76">T. iridescens</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFF67FAD95A28" box="[1308,1399,1588,1614]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF84FFE6FF5BFF26FAD85B26" pageId="75" pageNumber="76">
<paragraph id="8BC936B2FF84FFE6FF5BFF26FADB5AB3" blockId="75.[189,1399,1652,1856]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFF26FADB5AB3" pageId="75" pageNumber="76">
10. Body length &lt;
<quantity id="4C8E9B57FF84FFE6FE78FF27FE545AE9" box="[414,506,1652,1679]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="75" pageNumber="76" unit="mm" value="3.5">3.5 mm</quantity>
and eyes rather small, tempora at least 0.7 times as long as eyes, first stria of elytra indistinctly impressed or absent, microsculpture of elytra with very finely engraved very transverse meshes ............................................................................................................................11
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF84FFE6FF27FF8CFAD85B26" blockId="75.[189,1399,1652,1856]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF27FF8CFAD85B26" pageId="75" pageNumber="76">
– Either body length ± 
<quantity id="4C8E9B57FF84FFE6FE3AFF8DFD995A9F" box="[476,567,1758,1785]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="75" pageNumber="76" unit="mm" value="3.5">3.5 mm</quantity>
, or eyes rather large, tempora at maximum 0.5 times as long as eyes, first stria of elytra distinctly impressed, microsculpture of elytra with distinctly engraved isodiametric or slightly transverse meshes .............................................................................................................. 15
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF84FFE6FF5BFE34FAD85B8D" pageId="75" pageNumber="76">
<paragraph id="8BC936B2FF84FFE6FF5BFE34FAD95BC3" blockId="75.[189,1399,1895,2027]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF5BFE34FAD95BC3" pageId="75" pageNumber="76">
11. Body length approximately 
<quantity id="4C8E9B57FF84FFE6FDD8FE34FD7C5BE7" box="[574,722,1895,1921]" metricMagnitude="-3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="75" pageNumber="76" unit="mm" value="3.25" valueMax="3.3" valueMin="3.2">3.2–3.3 mm</quantity>
. Aedeagal median lobe slenderer, with terminal lamella long, markedly bent upward at tip (
<figureCitation id="134D2A37FF84FFE6FD6EFED9FD705BC3" box="[648,734,1930,1957]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="75" pageNumber="76">Fig. 56</figureCitation>
) ............................................................ 
<taxonomicName id="4C764D31FF84FFE6FB4AFED9FABB5BC3" ID-CoL="8MG4M" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1196,1301,1930,1957]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="bombi" status="sp. nov.">
<emphasis id="B902EAA0FF84FFE6FB4AFED9FABB5BC3" bold="true" box="[1196,1301,1930,1957]" italics="true" pageId="75" pageNumber="76">T. bombi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF84FFE6FAFAFED8FAD95BC3" box="[1308,1399,1931,1957]" pageId="75" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF84FFE6FF27FEFDFAD85B8D" blockId="75.[189,1399,1895,2027]" pageId="75" pageNumber="76">
<keyLead id="30879587FF84FFE6FF27FEFDFAD85B8D" pageId="75" pageNumber="76">– Body length below 3.0 mm. Aedeagal median lobe rather stout, with terminal lamella moderately short, at most slightly bent upward at tip ........................................................................................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF83FFE1FF5BF858FAD85DD9" pageId="76" pageNumber="77">
<paragraph id="8BC936B2FF83FFE1FF5BF858FAD95D15" blockId="76.[189,1399,267,448]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BF858FAD95D15" pageId="76" pageNumber="77">
12. Endophallus very slightly sclerotized throughout, without suggestion of a copulatory piece even in view from laterad (
<figureCitation id="134D2A37FF83FFE1FE08F861FDE65D2B" box="[494,584,306,333]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="76" pageNumber="77">Fig. 53</figureCitation>
). Species from northern slope of western Bale Mts above the city of Adaba ....................................................................................................................... 
<taxonomicName id="4C764D31FF83FFE1FB48F80AFABB5D15" ID-CoL="8M9MN" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1198,1301,344,371]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="adaba" status="sp. nov.">
<emphasis id="B902EAA0FF83FFE1FB48F80AFABB5D15" bold="true" box="[1198,1301,344,371]" italics="true" pageId="76" pageNumber="77">T. adaba</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF83FFE1FAFAF80AFAD95D15" box="[1308,1399,345,371]" pageId="76" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF27F82CFAD85DD9" blockId="76.[189,1399,267,448]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF27F82CFAD85DD9" pageId="76" pageNumber="77">– Endophallus with a more markedly sclerotized part near dorsoapical ostium which is easily recognizable in lateral view ............................................................................................................ 13</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF83FFE1FF5BF8B9FAD85E11" pageId="76" pageNumber="77">
<paragraph id="8BC936B2FF83FFE1FF5BF8B9FAD95E4D" blockId="76.[189,1399,490,632]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BF8B9FAD95E4D" pageId="76" pageNumber="77">
13. Aedeagal median lobe in lateral view suggestively sinusoidal with ventral margin slightly convex in apical half (
<figureCitation id="134D2A37FF83FFE1FE93FB43FE635E4D" box="[373,461,528,555]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="76" pageNumber="77">Fig. 55</figureCitation>
). Species from southeastern slope of Bale Mts .................... 
<taxonomicName id="4C764D31FF83FFE1FB76FB42FABB5E4D" ID-CoL="8MHV3" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1168,1301,528,555]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="harenna" status="sp. nov.">
<emphasis id="B902EAA0FF83FFE1FB76FB42FABB5E4D" bold="true" box="[1168,1301,528,555]" italics="true" pageId="76" pageNumber="77">T. harenna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF83FFE1FAFAFB42FAD95E4D" box="[1308,1399,529,555]" pageId="76" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF27FB64FAD85E11" blockId="76.[189,1399,490,632]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF27FB64FAD85E11" pageId="76" pageNumber="77">– Median lobe in lateral view with ventral margin evenly curved throughout or straight in apical half .................................................................................................................................................. 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF83FFE1FF5BFBF1FAD95F1B" pageId="76" pageNumber="77">
<paragraph id="8BC936B2FF83FFE1FF5BFBF1FAD95F6F" blockId="76.[189,1399,674,893]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFBF1FAD95F6F" pageId="76" pageNumber="77">
14. Aedeagal median lobe markedly short (EL/AL = 3.96–4.16), in lateral view almost evenly bent throughout (
<figureCitation id="134D2A37FF83FFE1FE59FB9BFDB25E85" box="[447,540,712,739]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="76" pageNumber="77">Fig. 52</figureCitation>
). Species from northern slope of western Bale Mts above the city of Dodola ..................................................................................................................... 
<taxonomicName id="4C764D31FF83FFE1FB40FBBCFABA5F6F" ID-CoL="8MLN6" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1190,1300,750,777]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="dodola" status="sp. nov.">
<emphasis id="B902EAA0FF83FFE1FB40FBBCFABA5F6F" bold="true" box="[1190,1300,750,777]" italics="true" pageId="76" pageNumber="77">T. dodola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF83FFE1FAFAFBBCFAD95F6F" box="[1308,1399,751,777]" pageId="76" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF27FA45FAD95F1B" blockId="76.[189,1399,674,893]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF27FA45FAD95F1B" pageId="76" pageNumber="77">
– Aedeagal median lobe more elongated (EL/AL = 3.03–3.22), in lateral view with ventral margin straight in apical half (
<figureCitation id="134D2A37FF83FFE1FE14FA6FFDE45F30" box="[498,586,827,854]" captionStart="Figs 37–56" captionStartId="28.[189,241,1788,1814]" captionTargetBox="[273,1317,319,1746]" captionTargetId="figure@28.[271,1317,317,1746]" captionTargetPageId="28" captionText="Figs 37–56. Trechus spp., aedeagal median lobe, left lateral view (37, 39, 40, 42, 44–45, 47–48, 50, 52–53, 55–56) and dorsal view (38, 41, 43, 46, 49, 51, 54). 37–38. T. mattisi sp. nov., paratype. 39. T. transversicollis sp. nov., holotype. 40–41. T. minitrechus sp. nov., paratype. 42–43. T. nanulus sp. nov., paratype. 44. T. rira sp. nov., holotype. 45–46. T. iridescens sp. nov., holotype. 47. T. relictus Magrini, Quéinnec &amp; Vigna Taglianti, 2012, specimen from brook valley above Goba. 48–49. T. oppositus sp. nov., paratype. 50–51. T. harryi sp. nov., paratype. 52. T. dodola sp. nov., holotype. 53–54. T. adaba sp. nov., paratype. 55. T. harenna sp. nov., paratype. 56. T. bombi sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829924" httpUri="https://zenodo.org/record/3829924/files/figure.png" pageId="76" pageNumber="77">Fig. 50</figureCitation>
). Species from southern slope of eastern Bale Mts, near the village of Rira .......................................................................................................................... 
<taxonomicName id="4C764D31FF83FFE1FB4AFA30FABB5F1B" ID-CoL="8MFFV" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1196,1301,866,893]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="harryi" status="sp. nov.">
<emphasis id="B902EAA0FF83FFE1FB4AFA30FABB5F1B" bold="true" box="[1196,1301,866,893]" italics="true" pageId="76" pageNumber="77">T. harryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF83FFE1FAFAFA30FAD95F1B" box="[1308,1399,867,893]" pageId="76" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BC936B2FF83FFE1FF5BFAF4FAD85868" blockId="76.[189,1398,935,1038]" pageId="76" pageNumber="77">
<keyStep id="30822E17FF83FFE1FF5BFAF4FAD85868" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFAF4FAD85868" pageId="76" pageNumber="77">15. Each elytron with a single discal seta which inserts near the end of the anterior elytral quarter ... 16 – Each elytron with two discal setae, the anterior one near the end of the anterior elytral quarter, the posterior one near elytral middle .................................................................................................... 17</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF5BFD6BFAD958F9" blockId="76.[189,1399,1080,1184]" pageId="76" pageNumber="77">
<keyStep id="30822E17FF83FFE1FF5BFD6BFAD958F9" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFD6BFAD958F9" pageId="76" pageNumber="77">
16. Pronotum with laterobasal setae absent. Species from Mt Chillalo ............. 
<taxonomicName id="4C764D31FF83FFE1FBBDFD6AFAD95834" ID-CoL="7CNPB" authority="Jeannel, 1960" authorityName="Jeannel" authorityYear="1960" box="[1115,1399,1080,1107]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="patrizii">
<emphasis id="B902EAA0FF83FFE1FBBDFD6AFB605834" bold="true" box="[1115,1230,1080,1107]" italics="true" pageId="76" pageNumber="77">T. patrizii</emphasis>
<bibRefCitation id="EFE74B43FF83FFE1FB33FD6BFAD95834" author="Jeannel R." box="[1237,1399,1080,1106]" pageId="76" pageNumber="77" pagination="265 - 266" refId="ref36713" refString="Jeannel R. 1960. Un nouveau Trechus de l'Abyssinie. Revue francaise d'Entomologie 27 (4): 265 - 266." type="journal article" year="1960">Jeannel, 1960</bibRefCitation>
</taxonomicName>
– Pronotum with laterobasal setae present. Species from Mt Chillalo, Mt Enkuolo, and northern slope of Bale Mts ............................................. 
<taxonomicName id="4C764D31FF83FFE1FD2BFDD5FAD958F9" ID-CoL="57ZW9" authority="Magrini, Queinnec &amp; Vigna Taglianti, 2012" authorityName="Magrini, Queinnec &amp; Vigna Taglianti" authorityYear="2012" box="[717,1399,1157,1184]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="oromiensis">
<emphasis id="B902EAA0FF83FFE1FD2BFDD5FCC158F9" bold="true" box="[717,879,1157,1184]" italics="true" pageId="76" pageNumber="77">T. oromiensis</emphasis>
<bibRefCitation id="EFE74B43FF83FFE1FC90FDD6FAD958F9" author="Magrini P. &amp; Queinnec E. &amp; Vigna Taglianti A." box="[886,1399,1157,1183]" pageId="76" pageNumber="77" pagination="19 - 32" refId="ref36764" refString="Magrini P., Queinnec E. &amp; Vigna Taglianti A. 2012. A new subgenus and two new species of Trechus from Ethiopia (Coleoptera, Carabidae). Fragmenta entomologica 44 (1): 19 - 32. https: // doi. org / 10.4081 / fe. 2012.30" type="journal article" year="2012">Magrini, Quéinnec &amp; Vigna Taglianti, 2012</bibRefCitation>
</taxonomicName>
</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF5BFD99FAD8596D" blockId="76.[189,1398,1226,1291]" pageId="76" pageNumber="77">
<keyStep id="30822E17FF83FFE1FF5BFD99FAD8596D" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFD99FAD8596D" pageId="76" pageNumber="77">17. Eyes flat and very small, tempora as long as or longer than eyes .................................................. 18 – Eyes moderately large and convexly protruded laterally, tempora distinctly shorter than eyes ..... 19</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF83FFE1FF5BFC66FAD859A5" pageId="76" pageNumber="77">
<paragraph id="8BC936B2FF83FFE1FF5BFC66FAD95913" blockId="76.[189,1399,1333,1475]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFC66FAD95913" pageId="76" pageNumber="77">
18. Aedeagal median lobe markedly funnel-shaped, widened near apex (
<bibRefCitation id="EFE74B43FF83FFE1FBC3FC66FAA95929" author="Jeannel R." box="[1061,1287,1333,1359]" pageId="76" pageNumber="77" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel 1936: 209</bibRefCitation>
). Species from Mt Chillalo .............................................................................................. 
<taxonomicName id="4C764D31FF83FFE1FB96FC0FFAD95913" ID-CoL="5822G" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[1136,1399,1371,1398]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="scotti">
<emphasis id="B902EAA0FF83FFE1FB96FC0FFB605910" bold="true" box="[1136,1230,1372,1398]" italics="true" pageId="76" pageNumber="77">T. scotti</emphasis>
<bibRefCitation id="EFE74B43FF83FFE1FB33FC0FFAD95913" author="Jeannel R." box="[1237,1399,1371,1398]" pageId="76" pageNumber="77" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF27FCD0FAD859A5" blockId="76.[189,1399,1333,1475]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF27FCD0FAD859A5" pageId="76" pageNumber="77">
– Aedeagal median lobe simply narrowed towards apex (
<bibRefCitation id="EFE74B43FF83FFE1FC9EFCD1FBF559FB" author="Geginat G." box="[888,1115,1410,1437]" pageId="76" pageNumber="77" pagination="115 - 126" refId="ref36602" refString="Geginat G. 2008. Revision of the genus Cothresia Jeannel, 1964 (Coleoptera: Carabidae: Trechinae: Trechodina). Annals of the Transvaal Museum 45: 115 - 126." type="journal article" year="2008">Geginat 2008: 125</bibRefCitation>
). Species from the Bale Mts ............................................................................................... 
<taxonomicName id="4C764D31FF83FFE1FC07FCFAFAD859A5" ID-CoL="57ZH9" authority="(Basilewsky, 1974)" baseAuthorityName="Basilewsky" baseAuthorityYear="1974" box="[993,1398,1448,1475]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="clarkeianus">
<emphasis id="B902EAA0FF83FFE1FC07FCFAFB2359A5" bold="true" box="[993,1165,1448,1475]" italics="true" pageId="76" pageNumber="77">T. clarkeianus</emphasis>
(
<bibRefCitation id="EFE74B43FF83FFE1FB7AFCFBFAC059A5" author="Basilewsky P." box="[1180,1390,1448,1475]" pageId="76" pageNumber="77" pagination="148 - 158" refId="ref36378" refString="Basilewsky P. 1974. Les Placomotrechina d'Ethiopie (Col. Carabidae Trechinae). Revue de Zoologie africaine 88 (1): 148 - 158." type="journal article" year="1974">Basilewsky, 1974</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF83FFE1FF5BFCBEFAD85AC7" pageId="76" pageNumber="77">
<paragraph id="8BC936B2FF83FFE1FF5BFCBEFAD85A32" blockId="76.[189,1399,1517,1698]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFCBEFAD85A32" pageId="76" pageNumber="77">
19. Body length&gt; 
<quantity id="4C8E9B57FF83FFE1FE4CFCBEFDA75A6E" box="[426,521,1517,1544]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="76" pageNumber="77" unit="mm" value="5.1">5.1 mm</quantity>
(usually&gt; 
<quantity id="4C8E9B57FF83FFE1FD7BFCBEFD505A6E" box="[669,766,1517,1544]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="76" pageNumber="77" unit="mm" value="5.5">5.5 mm</quantity>
) and endophallus extensively sclerotized with total length of sclerotized portions more than half of length of aedeagal median lobe (
<bibRefCitation id="EFE74B43FF83FFE1FB72FF47FADF5A48" author="Geginat G." box="[1172,1393,1555,1582]" pageId="76" pageNumber="77" pagination="115 - 126" refId="ref36602" refString="Geginat G. 2008. Revision of the genus Cothresia Jeannel, 1964 (Coleoptera: Carabidae: Trechinae: Trechodina). Annals of the Transvaal Museum 45: 115 - 126." type="journal article" year="2008">Geginat 2008: 125</bibRefCitation>
, fig. 5A–C) ....................................................................................................................................... 20
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF27FF32FAD85AC7" blockId="76.[189,1399,1517,1698]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF27FF32FAD85AC7" pageId="76" pageNumber="77">
– Either body length &lt;
<quantity id="4C8E9B57FF83FFE1FE05FF33FDEE5A1D" box="[483,576,1632,1659]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="76" pageNumber="77" unit="mm" value="5.1">5.1 mm</quantity>
, or endophallus moderately sclerotized with total length of sclerotized portions approx. one third of length of aedeagal median lobe ....................................................... 22
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF83FFE1FF5BFF9FFAD85BC0" pageId="76" pageNumber="77">
<paragraph id="8BC936B2FF83FFE1FF5BFF9FFAD85B55" blockId="76.[189,1399,1740,1959]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF5BFF9FFAD85B55" pageId="76" pageNumber="77">
20. Colour of dorsal surface mid-brown, elytra silk-matt, not iridescent; elytral microsculpture with meshes larger, more deeply engraved; aedeagal median lobe markedly elongated (
<bibRefCitation id="EFE74B43FF83FFE1FB2CFFA1FEB65B55" author="Geginat G." pageId="76" pageNumber="77" pagination="115 - 126" refId="ref36602" refString="Geginat G. 2008. Revision of the genus Cothresia Jeannel, 1964 (Coleoptera: Carabidae: Trechinae: Trechodina). Annals of the Transvaal Museum 45: 115 - 126." type="journal article" year="2008">Geginat 2008: 125</bibRefCitation>
, fig. 5A). Species from the Bale Mts ................................... 
<taxonomicName id="4C764D31FF83FFE1FC30FE4AFAD85B55" ID-CoL="7CNPF" authority="(Basilewsky, 1974)" baseAuthorityName="Basilewsky" baseAuthorityYear="1974" box="[982,1398,1816,1843]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="rotundicollis">
<emphasis id="B902EAA0FF83FFE1FC30FE4AFB235B55" bold="true" box="[982,1165,1816,1843]" italics="true" pageId="76" pageNumber="77">T. rotundicollis</emphasis>
(
<bibRefCitation id="EFE74B43FF83FFE1FB7AFE4AFAC05B55" author="Basilewsky P." box="[1180,1390,1817,1843]" pageId="76" pageNumber="77" pagination="148 - 158" refId="ref36378" refString="Basilewsky P. 1974. Les Placomotrechina d'Ethiopie (Col. Carabidae Trechinae). Revue de Zoologie africaine 88 (1): 148 - 158." type="journal article" year="1974">Basilewsky, 1974</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF83FFE1FF27FE13FAD85BC0" blockId="76.[189,1399,1740,1959]" pageId="76" pageNumber="77">
<keyLead id="30879587FF83FFE1FF27FE13FAD85BC0" pageId="76" pageNumber="77">
– Colour of dorsal surface dark or blackish brown, elytra markedly shiny, iridescent; elytral microsculpture with meshes narrower, more slightly engraved; aedeagal median lobe markedly robust (
<bibRefCitation id="EFE74B43FF83FFE1FEA5FEDFFD8F5BC1" author="Geginat G." box="[323,545,1932,1959]" pageId="76" pageNumber="77" pagination="115 - 126" refId="ref36602" refString="Geginat G. 2008. Revision of the genus Cothresia Jeannel, 1964 (Coleoptera: Carabidae: Trechinae: Trechodina). Annals of the Transvaal Museum 45: 115 - 126." type="journal article" year="2008">Geginat 2008: 125</bibRefCitation>
, fig. 5B–C) ........................................................................................... 21
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF82FFE0FF5BF858FAD85DBE" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BF858FAD95D0A" blockId="77.[189,1399,267,472]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BF858FAD95D0A" pageId="77" pageNumber="78">
21. Elytra with striae 1–4 very finely impressed and intervals flat; aedeagal median lobe in lateral view with ventral margin slightly but distinctly convex (
<bibRefCitation id="EFE74B43FF82FFE0FCA8F87CFB9C5D2C" author="Geginat G." box="[846,1074,303,330]" pageId="77" pageNumber="78" pagination="115 - 126" refId="ref36602" refString="Geginat G. 2008. Revision of the genus Cothresia Jeannel, 1964 (Coleoptera: Carabidae: Trechinae: Trechodina). Annals of the Transvaal Museum 45: 115 - 126." type="journal article" year="2008">Geginat 2008: 125</bibRefCitation>
, fig. 5C). Species from Mt Chillalo ................................................................................................... 
<taxonomicName id="4C764D31FF82FFE0FBD4F800FAD95D0A" ID-CoL="57ZH2" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[1074,1399,338,365]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="chillalicus">
<emphasis id="B902EAA0FF82FFE0FBD4F800FB605D0B" bold="true" box="[1074,1230,338,365]" italics="true" pageId="77" pageNumber="78">T. chillalicus</emphasis>
<bibRefCitation id="EFE74B43FF82FFE0FB33F800FAD95D0A" author="Jeannel R." box="[1237,1399,338,365]" pageId="77" pageNumber="78" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27F824FAD85DBE" blockId="77.[189,1399,267,472]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27F824FAD85DBE" pageId="77" pageNumber="78">
– Elytra with striae 1–4 rather deeply impressed and intervals moderately convex; aedeagal median lobe in lateral view with ventral margin nearly straight (
<bibRefCitation id="EFE74B43FF82FFE0FC65F8C9FBCD5DD3" author="Geginat G." box="[899,1123,410,437]" pageId="77" pageNumber="78" pagination="115 - 126" refId="ref36602" refString="Geginat G. 2008. Revision of the genus Cothresia Jeannel, 1964 (Coleoptera: Carabidae: Trechinae: Trechodina). Annals of the Transvaal Museum 45: 115 - 126." type="journal article" year="2008">Geginat 2008: 125</bibRefCitation>
, fig. 5B). Species from the Bale Mts ..................................................................................... 
<taxonomicName id="4C764D31FF82FFE0FBE6F8EDFAD85DBE" ID-CoL="7CNYZ" authority="(Basilewsky, 1974)" baseAuthorityName="Basilewsky" baseAuthorityYear="1974" box="[1024,1398,445,472]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="baleensis">
<emphasis id="B902EAA0FF82FFE0FBE6F8EDFB235DBE" bold="true" box="[1024,1165,445,472]" italics="true" pageId="77" pageNumber="78">T. baleensis</emphasis>
(
<bibRefCitation id="EFE74B43FF82FFE0FB7AF8EEFAC05DBE" author="Basilewsky P." box="[1180,1390,445,472]" pageId="77" pageNumber="78" pagination="148 - 158" refId="ref36378" refString="Basilewsky P. 1974. Les Placomotrechina d'Ethiopie (Col. Carabidae Trechinae). Revue de Zoologie africaine 88 (1): 148 - 158." type="journal article" year="1974">Basilewsky, 1974</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BC936B2FF82FFE0FF5BFB53FAD85E5B" blockId="77.[189,1398,511,574]" pageId="77" pageNumber="78">
<keyStep id="30822E17FF82FFE0FF5BFB53FAD85E5B" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFB53FAD85E5B" pageId="77" pageNumber="78">22. Species from Mt Chillalo ................................................................................................................ 23 – Species from the Bale Mts .............................................................................................................. 24</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF82FFE0FF5BFB36FAD95F6B" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BFB36FAD95EA0" blockId="77.[189,1399,613,782]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFB36FAD95EA0" pageId="77" pageNumber="78">
23. Elytra shinier with meshes of microsculpture more finely engraved, narrower, more transverse; aedeagal median lobe with apex hook-like bent upward (
<bibRefCitation id="EFE74B43FF82FFE0FC65FBDBFBF35EC5" author="Jeannel R." box="[899,1117,648,675]" pageId="77" pageNumber="78" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel 1936: 207</bibRefCitation>
, fig. 2) ......................... ................................................................................................................... 
<taxonomicName id="4C764D31FF82FFE0FBD9FBFEFAD95EA0" ID-CoL="57ZLS" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[1087,1399,684,711]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="gallorites">
<emphasis id="B902EAA0FF82FFE0FBD9FBFEFB635EA1" bold="true" box="[1087,1229,684,711]" italics="true" pageId="77" pageNumber="78">T. gallorites</emphasis>
<bibRefCitation id="EFE74B43FF82FFE0FB33FBFFFAD95EA0" author="Jeannel R." box="[1237,1399,684,710]" pageId="77" pageNumber="78" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27FB83FAD95F6B" blockId="77.[189,1399,613,782]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27FB83FAD95F6B" pageId="77" pageNumber="78">
– Elytra silk-like matt with meshes of microsculpture more deeply engraved, larger, almost isodiametric; aedeagal median lobe with apex simple (
<bibRefCitation id="EFE74B43FF82FFE0FD21FBA0FC0C5F68" author="Jeannel R." box="[711,930,755,782]" pageId="77" pageNumber="78" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel 1936: 208</bibRefCitation>
, fig. 5) .... 
<taxonomicName id="4C764D31FF82FFE0FBC6FBA7FAD95F6B" ID-CoL="57ZHS" authority="Jeannel, 1936" authorityName="Jeannel" authorityYear="1936" box="[1056,1399,755,782]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="culminicola">
<emphasis id="B902EAA0FF82FFE0FBC6FBA7FB635F68" bold="true" box="[1056,1229,755,782]" italics="true" pageId="77" pageNumber="78">T. culminicola</emphasis>
<bibRefCitation id="EFE74B43FF82FFE0FB33FBA0FAD95F6B" author="Jeannel R." box="[1237,1399,755,781]" pageId="77" pageNumber="78" pagination="205 - 212" refId="ref36636" refString="Jeannel R. 1936. Sur des Trechinae recueillis par M. Hugh Scott en Abyssinie. Revue francaise d'Entomologie 2 (4) (1935): 205 - 212." type="journal article" year="1936">Jeannel, 1936</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF82FFE0FF5BFA66FAD85FB9" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BFA66FAD95FF1" blockId="77.[189,1399,821,991]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFA66FAD95FF1" pageId="77" pageNumber="78">
24. Pronotum with basal margin slightly but distinctly concave at outer quarters, and with laterobasal angles large, acute, posteriorly and laterally protruded (
<figureCitation id="134D2A37FF82FFE0FC8DFA0AFC6C5F15" box="[875,962,857,883]" captionStart="Figs 71–74" captionStartId="43.[189,241,1965,1991]" captionTargetBox="[295,1292,317,1923]" captionTargetId="figure@43.[295,1292,317,1923]" captionTargetPageId="43" captionText="Figs 71–74. Trechus spp., head, pronotum. 71. T. depressipennis sp. nov., holotype. 72. T. mekbibi sp. nov., paratype, ³. 73. T. hagenia sp. nov., paratype, ³. 74. T. colobus sp. nov., paratype, ³." figureDoi="http://doi.org/10.5281/zenodo.3829932" httpUri="https://zenodo.org/record/3829932/files/figure.png" pageId="77" pageNumber="78">Fig. 72</figureCitation>
). Species from the southern slope and the western part of the Bale Mts............................................................................ 
<taxonomicName id="4C764D31FF82FFE0FB71FA2EFABB5FF1" ID-CoL="8M8X2" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1175,1301,892,919]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="mekbibi" status="sp. nov.">
<emphasis id="B902EAA0FF82FFE0FB71FA2EFABB5FF1" bold="true" box="[1175,1301,892,919]" italics="true" pageId="77" pageNumber="78">T. mekbibi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF82FFE0FAFAFA2EFAD95FF1" box="[1308,1399,893,919]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27FAF2FAD85FB9" blockId="77.[189,1399,821,991]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27FAF2FAD85FB9" pageId="77" pageNumber="78">– Pronotum with basal margin straight or convexly rounded at outer quarters; laterobasal angles not protruded posteriad ......................................................................................................................... 25</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF82FFE0FF5BFD55FAD858C8" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BFD55FAD85801" blockId="77.[189,1399,1030,1199]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFD55FAD85801" pageId="77" pageNumber="78">
25. Pronotum with basal margin straight or very slightly bent anteriorly at outer quarters, and with laterobasal angles acute or rectangular or formed as a rather large, +/- sharp tooth (
<figureCitation id="134D2A37FF82FFE0FB3BFD7AFEA75801" captionStart="Figs 71–74" captionStartId="43.[189,241,1965,1991]" captionTargetBox="[295,1292,317,1923]" captionTargetId="figure@43.[295,1292,317,1923]" captionTargetPageId="43" captionText="Figs 71–74. Trechus spp., head, pronotum. 71. T. depressipennis sp. nov., holotype. 72. T. mekbibi sp. nov., paratype, ³. 73. T. hagenia sp. nov., paratype, ³. 74. T. colobus sp. nov., paratype, ³." figureDoi="http://doi.org/10.5281/zenodo.3829932" httpUri="https://zenodo.org/record/3829932/files/figure.png" pageId="77" pageNumber="78">Figs 71, 73– 74</figureCitation>
) ................................................................................................................................................... 26
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27FD22FAD858C8" blockId="77.[189,1399,1030,1199]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27FD22FAD858C8" pageId="77" pageNumber="78">
– Pronotum with basal margin distinctly bent anteriorly at outer quarters, and with laterobasal angles rounded or formed as a small blunt tooth (
<figureCitation id="134D2A37FF82FFE0FD3EFDC7FCF158C8" box="[728,863,1172,1198]" captionStart="Figs 83–86" captionStartId="47.[189,241,1966,1992]" captionTargetBox="[258,1330,317,1927]" captionTargetId="figure@47.[258,1330,317,1927]" captionTargetPageId="47" captionText="Figs 83–86. Trechus spp., head, pronotum. 83. T. wiersbowskyi sp. nov., paratype, ³. 84. T. grandipennis sp. nov., holotype. 85. T. sanettii sp. nov., paratype, ³. 86. T. abalkhasimi sp. nov., paratype, ³." figureDoi="http://doi.org/10.5281/zenodo.3829938" httpUri="https://zenodo.org/record/3829938/files/figure.png" pageId="77" pageNumber="78">Figs 83–86</figureCitation>
, 
<figureCitation id="134D2A37FF82FFE0FC8AFDC7FC1658C9" box="[876,952,1172,1199]" captionStart="Figs 95–98" captionStartId="55.[189,241,1930,1956]" captionTargetBox="[297,1289,317,1890]" captionTargetId="figure@55.[297,1289,317,1890]" captionTargetPageId="55" captionText="Figs 95–98. Trechus spp., head, pronotum. 95. T. fisehai sp. nov., paratype, ³. 96. T. angavoensis sp. nov., holotype. 97. T. batuensis Magrini &amp; Sciaky, 2006, ³ from Sanetti Plateau near Tulo Dimptu. 98. T. batuensis, ³ from Tegona Valley." figureDoi="http://doi.org/10.5281/zenodo.3829944" httpUri="https://zenodo.org/record/3829944/files/figure.png" pageId="77" pageNumber="78">95–98</figureCitation>
, 
<figureCitation id="134D2A37FF82FFE0FC22FDC7FB8558C8" box="[964,1067,1172,1198]" captionStart="Figs 107–110" captionStartId="65.[189,241,1965,1991]" captionTargetBox="[301,1287,317,1923]" captionTargetId="figure@65.[301,1287,317,1923]" captionTargetPageId="65" captionText="Figs 107–110. Trechus spp., head, pronotum. 107. T. haggei sp. nov., paratype, ³. 108. T. tragelaphus sp. nov., holotype. 109. T. nigrifemoralis, sp. nov., holotype. 110. T. balesilvestris, sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829950" httpUri="https://zenodo.org/record/3829950/files/figure.png" pageId="77" pageNumber="78">108–110</figureCitation>
) ...................................... 29
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF82FFE0FF5BFD85FAD85919" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BFD85FAD9595E" blockId="77.[189,1399,1238,1407]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFD85FAD9595E" pageId="77" pageNumber="78">
26. Pronotum with laterobasal angles comparatively large and acute (
<figureCitation id="134D2A37FF82FFE0FC3FFD85FB815896" box="[985,1071,1238,1264]" captionStart="Figs 71–74" captionStartId="43.[189,241,1965,1991]" captionTargetBox="[295,1292,317,1923]" captionTargetId="figure@43.[295,1292,317,1923]" captionTargetPageId="43" captionText="Figs 71–74. Trechus spp., head, pronotum. 71. T. depressipennis sp. nov., holotype. 72. T. mekbibi sp. nov., paratype, ³. 73. T. hagenia sp. nov., paratype, ³. 74. T. colobus sp. nov., paratype, ³." figureDoi="http://doi.org/10.5281/zenodo.3829932" httpUri="https://zenodo.org/record/3829932/files/figure.png" pageId="77" pageNumber="78">Fig. 71</figureCitation>
). Elytra markedly depressed on disc; meshes of elytral microsculpture rather large, as wide as long or very slightly transverse. Species from northern slope of Bale Mts .................................................. 
<taxonomicName id="4C764D31FF82FFE0FBA3FC4DFABB595E" ID-CoL="8LP5T" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1093,1301,1309,1336]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="depressipennis" status="sp. nov.">
<emphasis id="B902EAA0FF82FFE0FBA3FC4DFABB595E" bold="true" box="[1093,1301,1309,1336]" italics="true" pageId="77" pageNumber="78">T. depressipennis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF82FFE0FAFAFC4DFAD9595E" box="[1308,1399,1310,1336]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27FC11FAD85919" blockId="77.[189,1399,1238,1407]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27FC11FAD85919" pageId="77" pageNumber="78">– Laterobasal angles of pronotum smaller. Elytra more convex, at most slightly depressed on disc, with micro-meshes very narrow, distinctly transverse .................................................................... 27</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF82FFE0FF5BFCF4FAD85A29" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BFCF4FAD95983" blockId="77.[189,1399,1447,1616]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFCF4FAD95983" pageId="77" pageNumber="78">
27. Aedeagal median lobe in lateral view distinctly sinusoidal, markedly bent behind basal bulb and markedly convex in middle (
<figureCitation id="134D2A37FF82FFE0FDDAFC99FD0D5982" box="[572,675,1482,1508]" captionStart="Figs 115–125" captionStartId="70.[189,241,1860,1886]" captionTargetBox="[199,1387,318,1817]" captionTargetId="figure@70.[199,1389,317,1818]" captionTargetPageId="70" captionText="Figs 115–125. Trechus spp., aedeagal median lobe, left lateral view (115–116, 118, 120, 122, 124–125) and dorsal view (117, 119, 121, 123). 115. T. depressipennis sp. nov., holotype. 116–117. T. mekbibi sp. nov., paratypes. 118–119. T. hagenia sp. nov., paratypes. 120–121. T. colobus sp. nov., paratypes. 122–123. T. wiersbowskyi sp. nov., paratypes. 124. T. haggei sp. nov., paratype. 125. T. grandipennis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829954" httpUri="https://zenodo.org/record/3829954/files/figure.png" pageId="77" pageNumber="78">Fig. 120</figureCitation>
). Species from the western Bale Mts ........ 
<taxonomicName id="4C764D31FF82FFE0FB7CFC98FABB5983" ID-CoL="8MH68" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1178,1301,1482,1509]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="colobus" status="sp. nov.">
<emphasis id="B902EAA0FF82FFE0FB7CFC98FABB5983" bold="true" box="[1178,1301,1482,1509]" italics="true" pageId="77" pageNumber="78">T. colobus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF82FFE0FAFAFC98FAD95983" box="[1308,1399,1483,1509]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27FCBDFAD85A29" blockId="77.[189,1399,1447,1616]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27FCBDFAD85A29" pageId="77" pageNumber="78">
– Aedeagal median lobe in lateral view markedly elongated, slightly bent behind basal bulb and slightly convex or straight in middle (
<figureCitation id="134D2A37FF82FFE0FD4DFF42FCBF5A4A" box="[683,785,1553,1580]" captionStart="Figs 115–125" captionStartId="70.[189,241,1860,1886]" captionTargetBox="[199,1387,318,1817]" captionTargetId="figure@70.[199,1389,317,1818]" captionTargetPageId="70" captionText="Figs 115–125. Trechus spp., aedeagal median lobe, left lateral view (115–116, 118, 120, 122, 124–125) and dorsal view (117, 119, 121, 123). 115. T. depressipennis sp. nov., holotype. 116–117. T. mekbibi sp. nov., paratypes. 118–119. T. hagenia sp. nov., paratypes. 120–121. T. colobus sp. nov., paratypes. 122–123. T. wiersbowskyi sp. nov., paratypes. 124. T. haggei sp. nov., paratype. 125. T. grandipennis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829954" httpUri="https://zenodo.org/record/3829954/files/figure.png" pageId="77" pageNumber="78">Fig. 118</figureCitation>
; 
<bibRefCitation id="EFE74B43FF82FFE0FCF9FF42FBD95A4A" author="Magrini P. &amp; Sciaky R." box="[799,1143,1553,1580]" pageId="77" pageNumber="78" pagination="173 - 190" refId="ref36872" refString="Magrini P. &amp; Sciaky R. 2006. Cinque nuove specie di Trechus dell'Ethiopia (Coleoptera, Carabidae). Fragmenta entomologica 38 (2): 173 - 190. https: // doi. org / 10.4081 / fe. 2006.4" type="journal article" year="2006">Magrini &amp; Sciaky 2006: 185</bibRefCitation>
, fig. 9). Species from the northern and southern slopes of the eastern Bale Mts .............................................................. 28
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF82FFE0FF5BFF24FAD95B46" pageId="77" pageNumber="78">
<paragraph id="8BC936B2FF82FFE0FF5BFF24FAD95ABE" blockId="77.[189,1399,1655,1824]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFF24FAD95ABE" pageId="77" pageNumber="78">
28. Aedeagal median lobe in dorsal view distinctly widened before apex (
<bibRefCitation id="EFE74B43FF82FFE0FBFFFF24FADF5AF4" author="Magrini P. &amp; Sciaky R." box="[1049,1393,1655,1682]" pageId="77" pageNumber="78" pagination="173 - 190" refId="ref36872" refString="Magrini P. &amp; Sciaky R. 2006. Cinque nuove specie di Trechus dell'Ethiopia (Coleoptera, Carabidae). Fragmenta entomologica 38 (2): 173 - 190. https: // doi. org / 10.4081 / fe. 2006.4" type="journal article" year="2006">Magrini &amp; Sciaky, 2006: 185</bibRefCitation>
, fig. 10), and with endophallic copulatory piece short, triangular. Species from northeastern slope of Bale Mts ................................................................................. 
<taxonomicName id="4C764D31FF82FFE0FC53FFECFAD95ABE" ID-CoL="7CNZJ" authority="Magrini &amp; Sciaky, 2006" authorityName="Magrini &amp; Sciaky" authorityYear="2006" box="[949,1399,1726,1753]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="bastianinii">
<emphasis id="B902EAA0FF82FFE0FC53FFECFBFA5ABF" bold="true" box="[949,1108,1726,1753]" italics="true" pageId="77" pageNumber="78">T. bastianinii</emphasis>
<bibRefCitation id="EFE74B43FF82FFE0FBBDFFEDFAD95ABE" author="Magrini P. &amp; Sciaky R." box="[1115,1399,1726,1752]" pageId="77" pageNumber="78" pagination="173 - 190" refId="ref36872" refString="Magrini P. &amp; Sciaky R. 2006. Cinque nuove specie di Trechus dell'Ethiopia (Coleoptera, Carabidae). Fragmenta entomologica 38 (2): 173 - 190. https: // doi. org / 10.4081 / fe. 2006.4" type="journal article" year="2006">Magrini &amp; Sciaky, 2006</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF82FFE0FF27FFB0FAD95B46" blockId="77.[189,1399,1655,1824]" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF27FFB0FAD95B46" pageId="77" pageNumber="78">
– Aedeagal median lobe in dorsal view parallel before apex and with endophallic copulatory daggerlike (
<figureCitation id="134D2A37FF82FFE0FEC0FE55FE225B46" box="[294,396,1798,1824]" captionStart="Figs 115–125" captionStartId="70.[189,241,1860,1886]" captionTargetBox="[199,1387,318,1817]" captionTargetId="figure@70.[199,1389,317,1818]" captionTargetPageId="70" captionText="Figs 115–125. Trechus spp., aedeagal median lobe, left lateral view (115–116, 118, 120, 122, 124–125) and dorsal view (117, 119, 121, 123). 115. T. depressipennis sp. nov., holotype. 116–117. T. mekbibi sp. nov., paratypes. 118–119. T. hagenia sp. nov., paratypes. 120–121. T. colobus sp. nov., paratypes. 122–123. T. wiersbowskyi sp. nov., paratypes. 124. T. haggei sp. nov., paratype. 125. T. grandipennis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829954" httpUri="https://zenodo.org/record/3829954/files/figure.png" pageId="77" pageNumber="78">Fig. 119</figureCitation>
). Species from southeastern slope of Bale Mts .............................. 
<taxonomicName id="4C764D31FF82FFE0FB73FE55FABB5B46" ID-CoL="8MKXF" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1173,1301,1797,1824]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hagenia" status="sp. nov.">
<emphasis id="B902EAA0FF82FFE0FB73FE55FABB5B46" bold="true" box="[1173,1301,1797,1824]" italics="true" pageId="77" pageNumber="78">T. hagenia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF82FFE0FAFAFE55FAD95B46" box="[1308,1399,1798,1824]" pageId="77" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BC936B2FF82FFE0FF5BFE1BFAD85BCF" blockId="77.[189,1398,1863,1961]" pageId="77" pageNumber="78">
<keyStep id="30822E17FF82FFE0FF5BFE1BFAD85BCF" pageId="77" pageNumber="78">
<keyLead id="30879587FF82FFE0FF5BFE1BFAD85BCF" pageId="77" pageNumber="78">29. Microsculpture of elytra consists of rather large almost isodiametric meshes; elytra silk-shiny, not iridescent ......................................................................................................................................... 30 – Microsculpture with sculpticells narrow and transverse; elytra markedly shiny, iridescent .......... 32</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF5BF858FAD85D2F" blockId="78.[189,1399,267,329]" pageId="78" pageNumber="79">
<keyStep id="30822E17FF81FFE3FF5BF858FAD85D2F" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BF858FAD85D2F" pageId="78" pageNumber="79">
30. Elytra with pores of discal setae conspicuously dimple-like, deepened ................. 
<taxonomicName id="4C764D31FF81FFE3FB40F85FFABA5D40" ID-CoL="8M663" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1190,1300,267,294]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="haggei" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FB40F85FFABA5D40" bold="true" box="[1190,1300,267,294]" italics="true" pageId="78" pageNumber="79">T. haggei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAF85FFAD95D40" box="[1308,1399,268,294]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
– Elytral discal pores slightly impressed, normal for 
<taxonomicName id="4C764D31FF81FFE3FCDBF87CFC345D2F" ID-CoL="8KZB9" authorityName="Clairville" authorityYear="1806" box="[829,922,303,329]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="genus">
<emphasis id="B902EAA0FF81FFE3FCDBF87CFC345D2F" box="[829,922,303,329]" italics="true" pageId="78" pageNumber="79">Trechus</emphasis>
</taxonomicName>
........................................................... 31
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF81FFE3FF5BF823FAD95D92" pageId="78" pageNumber="79">
<paragraph id="8BC936B2FF81FFE3FF5BF823FAD95DC8" blockId="78.[189,1399,368,500]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BF823FAD95DC8" pageId="78" pageNumber="79">
31. Endophallus with sclerotized portions located in centre of aedeagal median lobe (
<figureCitation id="134D2A37FF81FFE3FB1FF823FEB95DCB" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Figs 126– 127</figureCitation>
) ........................................................................................................................ 
<taxonomicName id="4C764D31FF81FFE3FB79F8C7FABB5DCB" ID-CoL="8MB9Q" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1183,1301,403,430]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="sanettii" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FB79F8C7FABB5DCB" bold="true" box="[1183,1301,403,430]" italics="true" pageId="78" pageNumber="79">T. sanettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAF8C7FAD95DC8" box="[1308,1399,404,430]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF27F8E4FAD95D92" blockId="78.[189,1399,368,500]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF27F8E4FAD95D92" pageId="78" pageNumber="79">
– Endophallus with sclerotized portions located in anterior half of aedeagal median lobe (
<figureCitation id="134D2A37FF81FFE3FB19F8E5FEB95D92" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Figs 129– 130</figureCitation>
) .......................................................................................... 
<taxonomicName id="4C764D31FF81FFE3FC22F889FAD95D92" ID-CoL="7CNZH" authority="Magrini &amp; Sciaky, 2006" authorityName="Magrini &amp; Sciaky" authorityYear="2006" box="[964,1399,473,500]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="batuensis">
<emphasis id="B902EAA0FF81FFE3FC22F889FBFA5D92" bold="true" box="[964,1108,473,500]" italics="true" pageId="78" pageNumber="79">T. batuensis</emphasis>
<bibRefCitation id="EFE74B43FF81FFE3FBBDF889FAD95D92" author="Magrini P. &amp; Sciaky R." box="[1115,1399,474,500]" pageId="78" pageNumber="79" pagination="173 - 190" refId="ref36872" refString="Magrini P. &amp; Sciaky R. 2006. Cinque nuove specie di Trechus dell'Ethiopia (Coleoptera, Carabidae). Fragmenta entomologica 38 (2): 173 - 190. https: // doi. org / 10.4081 / fe. 2006.4" type="journal article" year="2006">Magrini &amp; Sciaky, 2006</bibRefCitation>
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</paragraph>
</keyStep>
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<keyStep id="30822E17FF81FFE3FF5BFB48FAD85E1A" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFB48FAD85E1A" pageId="78" pageNumber="79">32. Pronotum comparatively large, elytra slenderer oval (proportion EW/PW below 1.45) ................ 33 – Markedly broad oval elytra contrasting with a rather small pronotum (proportion EW/PW above 1.5) .................................................................................................................................................. 35</keyLead>
</keyStep>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF5BFBF0FAD85F62" blockId="78.[189,1399,675,772]" pageId="78" pageNumber="79">
<keyStep id="30822E17FF81FFE3FF5BFBF0FAD85F62" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFBF0FAD85F62" pageId="78" pageNumber="79">
33. Aedeagal median lobe slenderer, proportion EL/AL distinctly below 2.5 (
<figureCitation id="134D2A37FF81FFE3FBA0FBF0FB045EDB" box="[1094,1194,675,701]" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Fig. 137</figureCitation>
) ........................ ....................................................................................................................... 
<taxonomicName id="4C764D31FF81FFE3FBBAFB94FABB5E87" ID-CoL="8M7WG" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1116,1301,710,737]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="balesilvestris" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FBBAFB94FABB5E87" bold="true" box="[1116,1301,710,737]" italics="true" pageId="78" pageNumber="79">T. balesilvestris</emphasis>
</taxonomicName>
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– Proportion EL/AL distinctly above 2.5 ........................................................................................... 34
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF81FFE3FF5BFA78FAD95FC9" pageId="78" pageNumber="79">
<paragraph id="8BC936B2FF81FFE3FF5BFA78FAD95F0F" blockId="78.[189,1399,811,944]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFA78FAD95F0F" pageId="78" pageNumber="79">
34. Legs yellowish brown, femora not distinctly darkened; aedeagal median lobe in lateral view markedly widened near apical ostium (
<figureCitation id="134D2A37FF81FFE3FD54FA1DFCB85F0F" box="[690,790,846,873]" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Fig. 135</figureCitation>
) ........................................... 
<taxonomicName id="4C764D31FF81FFE3FB8EFA1CFABA5F0F" ID-CoL="8MLZL" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1128,1300,846,873]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="tragelaphus" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FB8EFA1CFABA5F0F" bold="true" box="[1128,1300,846,873]" italics="true" pageId="78" pageNumber="79">T. tragelaphus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFA1CFAD95F0F" box="[1308,1399,847,873]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF27FA21FAD95FC9" blockId="78.[189,1399,811,944]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF27FA21FAD95FC9" pageId="78" pageNumber="79">
– Basal three quarter of femora distinctly darkened; aedeagal median lobe in lateral view not widened near apical ostium (
<figureCitation id="134D2A37FF81FFE3FE37FAC6FD9B5FD6" box="[465,565,917,944]" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Fig. 136</figureCitation>
) ...................................................................... 
<taxonomicName id="4C764D31FF81FFE3FBACFAC6FABB5FC9" ID-CoL="8M5ST" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1098,1301,916,943]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="nigrifemoralis" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FBACFAC6FABB5FC9" bold="true" box="[1098,1301,916,943]" italics="true" pageId="78" pageNumber="79">T. nigrifemoralis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFAC6FAD95FC9" box="[1308,1399,917,943]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF81FFE3FF5BFA85FAD8583C" pageId="78" pageNumber="79">
<paragraph id="8BC936B2FF81FFE3FF5BFA85FAD95872" blockId="78.[189,1399,982,1115]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFA85FAD95872" pageId="78" pageNumber="79">
35. Aedeagal median lobe markedly elongated with middle portion tube-like, in lateral and dorsal views with side margins straight and parallel in middle (
<figureCitation id="134D2A37FF81FFE3FCD7FAAAFC7B5872" box="[817,981,1017,1044]" captionStart="Figs 115–125" captionStartId="70.[189,241,1860,1886]" captionTargetBox="[199,1387,318,1817]" captionTargetId="figure@70.[199,1389,317,1818]" captionTargetPageId="70" captionText="Figs 115–125. Trechus spp., aedeagal median lobe, left lateral view (115–116, 118, 120, 122, 124–125) and dorsal view (117, 119, 121, 123). 115. T. depressipennis sp. nov., holotype. 116–117. T. mekbibi sp. nov., paratypes. 118–119. T. hagenia sp. nov., paratypes. 120–121. T. colobus sp. nov., paratypes. 122–123. T. wiersbowskyi sp. nov., paratypes. 124. T. haggei sp. nov., paratype. 125. T. grandipennis sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829954" httpUri="https://zenodo.org/record/3829954/files/figure.png" pageId="78" pageNumber="79">Figs 122–123</figureCitation>
) .............. 
<taxonomicName id="4C764D31FF81FFE3FBB1FAA9FABB5872" ID-CoL="8MGSM" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1111,1301,1017,1044]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="wiersbowskyi" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FBB1FAA9FABB5872" bold="true" box="[1111,1301,1017,1044]" italics="true" pageId="78" pageNumber="79">T. wiersbowskyi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFAA9FAD95872" box="[1308,1399,1018,1044]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF27FD4EFAD8583C" blockId="78.[189,1399,982,1115]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF27FD4EFAD8583C" pageId="78" pageNumber="79">– Aedeagal median lobe not tube-like, in lateral view with side margins not parallel, and with dorsal margin distinctly convex ................................................................................................................. 36</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BC936B2FF81FFE3FF5BFDD2FAD858D9" blockId="78.[189,1399,1153,1215]" pageId="78" pageNumber="79">
<keyStep id="30822E17FF81FFE3FF5BFDD2FAD858D9" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFDD2FAD858D9" pageId="78" pageNumber="79">
36. Aedeagal median lobe larger with total length about 
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..................... 
<taxonomicName id="4C764D31FF81FFE3FBB1FDD1FABB58FA" ID-CoL="8MCCM" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1111,1301,1153,1180]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="grandipennis" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FBB1FDD1FABB58FA" bold="true" box="[1111,1301,1153,1180]" italics="true" pageId="78" pageNumber="79">T. grandipennis</emphasis>
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<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFDD1FAD958FA" box="[1308,1399,1154,1180]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
– Maximum length of aedeagal median lobe not above 
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.......................................................... 37
</keyLead>
</keyStep>
</paragraph>
<keyStep id="30822E17FF81FFE3FF5BFDB5FAD859B2" pageId="78" pageNumber="79">
<paragraph id="8BC936B2FF81FFE3FF5BFDB5FAD9590C" blockId="78.[189,1399,1254,1492]" pageId="78" pageNumber="79">
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37. Aedeagal median lobe robust in lateral and dorsal view, with large basal bulb contrasting with markedly small sagittal aileron; endophallus more markedly sclerotized, with sclerotized portions distinctly longer than one third of total length of aedeagal median lobe (
<bibRefCitation id="EFE74B43FF81FFE3FBC9FC7EFAEF5921" author="Magrini P. &amp; Queinnec E. &amp; Vigna Taglianti A." box="[1071,1345,1325,1351]" pageId="78" pageNumber="79" pagination="17 - 24" refId="ref36818" refString="Magrini P., Queinnec E. &amp; Vigna Taglianti A. 2013. A new species of Trechus from the Ethiopian Plateau (Coleoptera, Carabidae). Fragmenta entomologica 45 (1 - 2): 17 - 24. https: // doi. org / 10.4081 / fe. 2013.13" type="journal article" year="2013">
Magrini 
<emphasis id="B902EAA0FF81FFE3FB70FC7DFB7E5921" box="[1174,1232,1325,1351]" italics="true" pageId="78" pageNumber="79">et al.</emphasis>
2013: 22
</bibRefCitation>
, figs 2–4) ............................................................... 
<taxonomicName id="4C764D31FF81FFE3FD1FFC03FAD9590C" ID-CoL="57ZKQ" authority="Magrini, Queinnec &amp; Vigna Taglianti, 2013" authorityName="Magrini, Queinnec &amp; Vigna Taglianti" authorityYear="2013" box="[761,1399,1359,1386]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="ericalis">
<emphasis id="B902EAA0FF81FFE3FD1FFC03FCC1590C" bold="true" box="[761,879,1359,1386]" italics="true" pageId="78" pageNumber="79">T. ericalis</emphasis>
<bibRefCitation id="EFE74B43FF81FFE3FC90FC03FAD9590C" author="Magrini P. &amp; Queinnec E. &amp; Vigna Taglianti A." box="[886,1399,1360,1386]" pageId="78" pageNumber="79" pagination="17 - 24" refId="ref36818" refString="Magrini P., Queinnec E. &amp; Vigna Taglianti A. 2013. A new species of Trechus from the Ethiopian Plateau (Coleoptera, Carabidae). Fragmenta entomologica 45 (1 - 2): 17 - 24. https: // doi. org / 10.4081 / fe. 2013.13" type="journal article" year="2013">Magrini, Quéinnec &amp; Vigna Taglianti, 2013</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF27FC27FAD859B2" blockId="78.[189,1399,1254,1492]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF27FC27FAD859B2" pageId="78" pageNumber="79">– Aedeagal median lobe slenderer, with basal bulb and sagittal aileron average; endophallus less markedly sclerotized, with maximum length of sclerotized portions one third of total length of aedeagal median lobe ...................................................................................................................... 38</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF81FFE3FF5BFCA8FAD85A19" pageId="78" pageNumber="79">
<paragraph id="8BC936B2FF81FFE3FF5BFCA8FAD95A5F" blockId="78.[189,1399,1531,1663]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFCA8FAD95A5F" pageId="78" pageNumber="79">
38. Aedeagal median lobe less markedly bent in basal portion, straight from the end of the basal third towards apex (
<figureCitation id="134D2A37FF81FFE3FE70FF4DFE545A5E" box="[406,506,1566,1592]" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Fig. 128</figureCitation>
) ................................................................................. 
<taxonomicName id="4C764D31FF81FFE3FB85FF4CFABB5A5E" ID-CoL="8MMPD" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1123,1301,1566,1593]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="angavoensis" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FB85FF4CFABB5A5E" bold="true" box="[1123,1301,1566,1593]" italics="true" pageId="78" pageNumber="79">T. angavoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFF4CFAD95A5F" box="[1308,1399,1567,1593]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF27FF11FAD85A19" blockId="78.[189,1399,1531,1663]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF27FF11FAD85A19" pageId="78" pageNumber="79">– Aedeagal median lobe more markedly bent in basal portion, straight after beginning of the second third ................................................................................................................................................. 39</keyLead>
</paragraph>
</keyStep>
<keyStep id="30822E17FF81FFE3FF5BFFF5FAD95B4D" pageId="78" pageNumber="79">
<paragraph id="8BC936B2FF81FFE3FF5BFFF5FAD95A82" blockId="78.[189,1399,1702,1835]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF5BFFF5FAD95A82" pageId="78" pageNumber="79">
39. Aedeagal median lobe larger, total length 
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, with button-like apex more markedly bent upward at tip (
<figureCitation id="134D2A37FF81FFE3FE73FF9AFE575A82" box="[405,505,1737,1764]" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Fig. 131</figureCitation>
) ................................................................................. 
<taxonomicName id="4C764D31FF81FFE3FB85FF99FABB5A82" ID-CoL="8MDDM" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1123,1301,1737,1764]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="abalkhasimi" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FB85FF99FABB5A82" bold="true" box="[1123,1301,1737,1764]" italics="true" pageId="78" pageNumber="79">T. abalkhasimi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFF99FAD95A82" box="[1308,1399,1738,1764]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
<paragraph id="8BC936B2FF81FFE3FF27FFBEFAD95B4D" blockId="78.[189,1399,1702,1835]" pageId="78" pageNumber="79">
<keyLead id="30879587FF81FFE3FF27FFBEFAD95B4D" pageId="78" pageNumber="79">
– Aedeagal median lobe smaller, total length 
<quantity id="4C8E9B57FF81FFE3FCEFFFBEFC125B61" box="[777,956,1773,1799]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="78" pageNumber="79" unit="mm" value="0.75" valueMax="0.8" valueMin="0.7">0.70–0.80 mm</quantity>
, with button-like apex slightly bent upward at tip (
<figureCitation id="134D2A37FF81FFE3FE73FE43FE575B4C" box="[405,505,1808,1834]" captionStart="Figs 126–137" captionStartId="71.[189,241,1860,1886]" captionTargetBox="[220,1366,318,1804]" captionTargetId="figure@71.[220,1367,317,1810]" captionTargetPageId="71" captionText="Figs 126–137. Trechus spp., aedeagal median lobe, left lateral view (126, 128–129, 131, 133, 135–137) and dorsal view (127, 130, 132, 134). 126–127. T. sanettii sp. nov., paratypes. 128. T. angavoensis sp. nov., holotype. 129–130. T. batuensis Magrini &amp; Sciaky, 2006, specimens from Wasama Valley. 131–132. T. abalkhasimi sp. nov., paratypes. 133–134. T. fisehai sp. nov., paratypes. 135. T. tragelaphus sp. nov., holotype. 136. T. nigrifemoralis sp. nov., holotype. 137. T. balesilvestris sp. nov., holotype." figureDoi="http://doi.org/10.5281/zenodo.3829956" httpUri="https://zenodo.org/record/3829956/files/figure.png" pageId="78" pageNumber="79">Fig. 133</figureCitation>
) .......................................................................................... 
<taxonomicName id="4C764D31FF81FFE3FB41FE43FABB5B4D" ID-CoL="8MBMZ" authority="Schmidt &amp; Faille, 2018" authorityName="Schmidt &amp; Faille" authorityYear="2018" box="[1191,1301,1808,1835]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="78" pageNumber="79" phylum="Arthropoda" rank="species" species="fisehai" status="sp. nov.">
<emphasis id="B902EAA0FF81FFE3FB41FE43FABB5B4D" bold="true" box="[1191,1301,1808,1835]" italics="true" pageId="78" pageNumber="79">T. fisehai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23157DBFF81FFE3FAFAFE42FAD95B4D" box="[1308,1399,1809,1835]" pageId="78" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>