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<document id="6BD8FB8FA75DBBDA301AB059810CD775" ID-DOI="10.11646/zootaxa.3860.1.2" ID-GBIF-Dataset="a26eaf5f-4eba-45ad-8806-dd7288d78493" ID-ISSN="1175-5326" ID-Zenodo-Dep="287060" ID-ZooBank="B9713C6D-BC3B-4EF2-8495-8BDCD535A80D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460903601137" checkinUser="plazi" docAuthor="Nakthong, La-Au, Vitheepradit, Akekawat &amp; Sites, Robert W." docDate="2014" docId="03DF879BFFC3FFD909C9998F48DF35B9" docLanguage="en" docName="zt03860p063.pdf" docOrigin="Zootaxa 3860 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amemboa (Amemboa) riparia Polhemus and Andersen" docType="treatment" docVersion="5" lastPageNumber="56" masterDocId="FFE6FFE3FFCAFFD0095E9B004B3B3256" masterDocTitle="Key to the species of Eotrechinae (Hemiptera: Heteroptera: Gerridae) of Thailand and review of the fauna of the Phetchabun Mountain Range" masterLastPageNumber="63" masterPageNumber="47" pageNumber="56" updateTime="1698360479435" updateUser="plazi">
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<mods:title id="5E49F841C0343370ABA719DC7D56EAEA">Key to the species of Eotrechinae (Hemiptera: Heteroptera: Gerridae) of Thailand and review of the fauna of the Phetchabun Mountain Range</mods:title>
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<mods:namePart id="03FE6F8FF7B3A334CE50E929CDAA82EB">Nakthong, La-Au</mods:namePart>
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<mods:namePart id="5DDC06A5269AFBB7DF78C2C46113DCAE">Vitheepradit, Akekawat</mods:namePart>
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<mods:namePart id="879CD072142AE43926A75C2267324EF3">Sites, Robert W.</mods:namePart>
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<mods:title id="FD4923EB9C42BE08014CB0829C9F24DF">Zootaxa</mods:title>
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<mods:date id="7E70D1D16A31183DA7D946BC5B3D7567">2014</mods:date>
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<treatment id="03DF879BFFC3FFD909C9998F48DF35B9" ID-DOI="http://doi.org/10.5281/zenodo.6137521" ID-GBIF-Taxon="127669792" ID-Zenodo-Dep="6137521" LSID="urn:lsid:plazi:treatment:03DF879BFFC3FFD909C9998F48DF35B9" httpUri="http://treatment.plazi.org/id/03DF879BFFC3FFD909C9998F48DF35B9" lastPageNumber="56" pageId="9" pageNumber="56">
<subSubSection id="C36C6506FFC3FFD909C9998F4A20309D" pageId="9" pageNumber="56" type="nomenclature">
<paragraph id="8BC9368DFFC3FFD909C9998F480330FC" blockId="9.[151,824,655,715]" box="[151,824,655,682]" pageId="9" pageNumber="56">
<heading id="D08181E1FFC3FFD909C9998F480330FC" bold="true" box="[151,824,655,682]" fontSize="11" level="1" pageId="9" pageNumber="56" reason="1">
<taxonomicName id="4C764D0EFFC3FFD909C9998F480330FC" ID-CoL="CQYJ" authority="Polhemus and Andersen" authorityName="Polhemus and Andersen" box="[151,824,655,682]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia" subGenus="Amemboa">
<emphasis id="B902EA9FFFC3FFD909C9998F480330FC" bold="true" box="[151,824,655,682]" pageId="9" pageNumber="56">
<emphasis id="B902EA9FFFC3FFD909C9998F493B30FF" bold="true" box="[151,512,655,681]" italics="true" pageId="9" pageNumber="56">Amemboa (Amemboa) riparia</emphasis>
Polhemus and Andersen
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC9368DFFC3FFD909C999B24A20309D" blockId="9.[151,824,655,715]" box="[151,283,690,715]" pageId="9" pageNumber="56">
<figureCitation id="134D2A08FFC3FFD909C999B24BC8309D" box="[151,243,690,715]" captionStart="FIGURES 8 21" captionStartId="6.[151,264,1668,1691]" captionTargetBox="[151,1436,229,1647]" captionTargetId="figure@6.[151,1436,229,1647]" captionTargetPageId="6" captionText="FIGURES 8 21. Left forelegs of male Amemboa. (8) A. aquafrigida, (9) A. armata, (10) A. brevifasciata, (11) A. cristata, (12) A. heissi, (13) A. incurvata, (14) A. javanica, (15) A. laotica, (16) A. prostata, (17) A. riparia, (18) A. schwendingeri, (19) A. speciosa, (20) A. perlata, (21) A. velaris." httpUri="https://zenodo.org/record/287064/files/figure.png" pageId="9" pageNumber="56">Figs. 17</figureCitation>
,
<figureCitation id="134D2A08FFC3FFD909A199B34A20309D" box="[255,283,691,715]" captionStart="FIGURES 22 35" captionStartId="8.[151,264,1910,1933]" captionTargetBox="[152,1434,193,1888]" captionTargetId="figure@8.[152,1435,193,1889]" captionTargetPageId="8" captionText="FIGURES 22 35. Terminal abdominal segments of male Amemboa. (22) A. aquafrigida, (23) A. armata, (24) A. brevifasciata, (25) A. cristata, (26) A. heissi, (27) A. incurvata, (28) A. javanica, (29) A. laotica, (30) A. prostata, (31) A. riparia, (32) A. schwendingeri, (33) A. speciosa, (34) A. perlata, (35) A. velaris." httpUri="https://zenodo.org/record/287065/files/figure.png" pageId="9" pageNumber="56">31</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36C6506FFC3FFD909C999FA48D23118" pageId="9" pageNumber="56" type="reference_group">
<paragraph id="8BC9368DFFC3FFD909C999FA48EF3146" blockId="9.[151,1331,762,846]" box="[151,980,762,784]" pageId="9" pageNumber="56">
<taxonomicName id="4C764D0EFFC3FFD909C999FA49C43146" ID-CoL="CQYJ" authority="Polhemus and Andersen 1984" authorityName="Polhemus and Andersen" authorityYear="1984" box="[151,767,762,784]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia" subGenus="Amemboa">
<emphasis id="B902EA9FFFC3FFD909C999FA4AFC3146" box="[151,455,762,784]" italics="true" pageId="9" pageNumber="56">Amemboa (Amemboa) riparia</emphasis>
<bibRefCitation id="EFE74B7CFFC3FFD9089399FA49C43146" author="Polhemus" box="[461,767,762,784]" pageId="9" pageNumber="56" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus and Andersen 1984</bibRefCitation>
</taxonomicName>
: Steenstrupia 9092.
</paragraph>
<paragraph id="8BC9368DFFC3FFD909C9981948D23118" blockId="9.[151,1331,762,846]" pageId="9" pageNumber="56">
<taxonomicName id="4C764D0EFFC3FFD909C998194AC03179" ID-CoL="CQY8" authority="Paiva" authorityName="Paiva" box="[151,507,793,815]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lyra" subGenus="Amemboa">
<emphasis id="B902EA9FFFC3FFD909C998194A933179" box="[151,424,793,815]" italics="true" pageId="9" pageNumber="56">Amemboa (Amemboa) lyra</emphasis>
(Paiva)
</taxonomicName>
:
<bibRefCitation id="EFE74B7CFFC3FFD90B5A981949DA3179" author="Zettel" box="[516,737,793,815]" pageId="9" pageNumber="56" refString="Zettel, H. &amp; Chen, P. - P. (1997) Three new taxa of Amemboa ESAKI, 1925, from Thailand and Vietnam. Annales Historico- Naturales Musei Nationalis Hungarici, 89, 93 - 101." type="journal article" year="1997">Zettel and Chen 1997</bibRefCitation>
, Ann. Hist.-Natur. Mus. Nat. Hung. 156157 (synonymy).
<taxonomicName id="4C764D0EFFC3FFD909C9983849E83118" ID-CoL="CQYJ" authority="Polhemus and Tran 2012" authorityName="Polhemus and Tran" authorityYear="2012" box="[151,723,824,846]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia" subGenus="Amemboa">
<emphasis id="B902EA9FFFC3FFD909C998384AFC3118" box="[151,455,824,846]" italics="true" pageId="9" pageNumber="56">Amemboa (Amemboa) riparia</emphasis>
: Polhemus and Tran 2012
</taxonomicName>
, Zootaxa 5557 (stat. nov.)
</paragraph>
</subSubSection>
<subSubSection id="C36C6506FFC3FFD909C9987A4A4D3610" pageId="9" pageNumber="56" type="diagnosis">
<paragraph id="8BC9368DFFC3FFD909C9987A4A4D3610" blockId="9.[151,1437,890,2031]" pageId="9" pageNumber="56">
<emphasis id="B902EA9FFFC3FFD909C9987A4A2A31C5" bold="true" box="[151,273,890,915]" pageId="9" pageNumber="56">Diagnosis.</emphasis>
Males of
<taxonomicName id="4C764D0EFFC3FFD908DD987B4ACA31C5" box="[387,497,891,915]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD908DD987B4AAF31C4" box="[387,404,891,914]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD908FC987C4ACA31C5" box="[418,497,892,915]" italics="true" pageId="9" pageNumber="56">riparia</emphasis>
</taxonomicName>
have the pygophore with a pair of small posterolateral lobes and a digitate tip (
<figureCitation id="134D2A08FFC3FFD90C31987A4B8D31E1" captionStart="FIGURES 22 35" captionStartId="8.[151,264,1910,1933]" captionTargetBox="[152,1434,193,1888]" captionTargetId="figure@8.[152,1435,193,1889]" captionTargetPageId="8" captionText="FIGURES 22 35. Terminal abdominal segments of male Amemboa. (22) A. aquafrigida, (23) A. armata, (24) A. brevifasciata, (25) A. cristata, (26) A. heissi, (27) A. incurvata, (28) A. javanica, (29) A. laotica, (30) A. prostata, (31) A. riparia, (32) A. schwendingeri, (33) A. speciosa, (34) A. perlata, (35) A. velaris." httpUri="https://zenodo.org/record/287065/files/figure.png" pageId="9" pageNumber="56">Fig. 31</figureCitation>
). Further, the male profemur has two distinct hair patches (
<figureCitation id="134D2A08FFC3FFD90A12989E48A531E1" box="[844,926,926,951]" captionStart="FIGURES 8 21" captionStartId="6.[151,264,1668,1691]" captionTargetBox="[151,1436,229,1647]" captionTargetId="figure@6.[151,1436,229,1647]" captionTargetPageId="6" captionText="FIGURES 8 21. Left forelegs of male Amemboa. (8) A. aquafrigida, (9) A. armata, (10) A. brevifasciata, (11) A. cristata, (12) A. heissi, (13) A. incurvata, (14) A. javanica, (15) A. laotica, (16) A. prostata, (17) A. riparia, (18) A. schwendingeri, (19) A. speciosa, (20) A. perlata, (21) A. velaris." httpUri="https://zenodo.org/record/287064/files/figure.png" pageId="9" pageNumber="56">Fig. 17</figureCitation>
), whereas
<taxonomicName id="4C764D0EFFC3FFD90D48989F4FAD31E0" box="[1046,1174,927,950]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="javanica">
<emphasis id="B902EA9FFFC3FFD90D48989F4F1C31E0" box="[1046,1063,927,950]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD90D6B989F4FAD31E0" box="[1077,1174,927,950]" italics="true" pageId="9" pageNumber="56">javanica</emphasis>
</taxonomicName>
has a single hair patch, and
<taxonomicName id="4C764D0EFFC3FFD9099998C34A57318D" box="[199,364,962,987]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="aquafrigida">
<emphasis id="B902EA9FFFC3FFD9099998C34BE3318C" box="[199,216,963,986]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD909B998C24A57318D" box="[231,364,962,987]" italics="true" pageId="9" pageNumber="56">aquafrigida</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D0EFFC3FFD908FD98C3491F318D" box="[419,548,963,987]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="speciosa">
<emphasis id="B902EA9FFFC3FFD908FD98C34A8F318C" box="[419,436,963,986]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD9089D98C4491F318D" box="[451,548,964,987]" italics="true" pageId="9" pageNumber="56">speciosa</emphasis>
</taxonomicName>
each have three hair patches. Males of
<taxonomicName id="4C764D0EFFC3FFD90ABD98C34F69318D" box="[995,1106,963,987]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90ABD98C348CF318C" box="[995,1012,963,986]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD90D5D98C44F69318D" box="[1027,1106,964,987]" italics="true" pageId="9" pageNumber="56">riparia</emphasis>
</taxonomicName>
have a proximally incrassate profemur, whereas
<taxonomicName id="4C764D0EFFC3FFD9082C98E7491F31A9" box="[370,548,998,1023]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="brevifasciata">
<emphasis id="B902EA9FFFC3FFD9082C98E74AB831A8" box="[370,387,999,1022]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD908CD98E6491F31A9" box="[403,548,998,1023]" italics="true" pageId="9" pageNumber="56">brevifasciata</emphasis>
</taxonomicName>
has an unmodified proximal half of the profemur. Males of
<taxonomicName id="4C764D0EFFC3FFD90D8798E74E7131A8" box="[1241,1354,999,1022]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90D8798E74FD131A8" box="[1241,1258,999,1022]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD90DA498E74E7131A8" box="[1274,1354,999,1022]" italics="true" pageId="9" pageNumber="56">riparia</emphasis>
</taxonomicName>
have a well-developed profemoral tumescence beyond the proximal hair patch, whereas those of
<taxonomicName id="4C764D0EFFC3FFD90DD59F0B4E3B3675" box="[1163,1280,1035,1059]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="cristata">
<emphasis id="B902EA9FFFC3FFD90DD59F0B4FA73674" box="[1163,1180,1035,1058]" italics="true" pageId="9" pageNumber="56">A</emphasis>
.
<emphasis id="B902EA9FFFC3FFD90DF49F0C4E3B3675" box="[1194,1280,1036,1059]" italics="true" pageId="9" pageNumber="56">cristata</emphasis>
</taxonomicName>
lack the midventral tumescence.
</paragraph>
</subSubSection>
<subSubSection id="C36C6506FFC3FFD909999F524E1834D1" pageId="9" pageNumber="56" type="discussion">
<paragraph id="8BC9368DFFC3FFD909999F524E1834D1" blockId="9.[151,1437,890,2031]" pageId="9" pageNumber="56">
<emphasis id="B902EA9FFFC3FFD909999F524A7E363D" bold="true" box="[199,325,1106,1131]" pageId="9" pageNumber="56">Discussion</emphasis>
.
<bibRefCitation id="EFE74B7CFFC3FFD9080B9F524991363D" author="Polhemus" box="[341,682,1106,1131]" pageId="9" pageNumber="56" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen (1984)</bibRefCitation>
described
<taxonomicName id="4C764D0EFFC3FFD90A799F5348AC363D" box="[807,919,1107,1131]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90A799F5348AC363D" box="[807,919,1107,1131]" italics="true" pageId="9" pageNumber="56">A. riparia</emphasis>
</taxonomicName>
based on specimens from Chiang Mai. They described the similarities between
<taxonomicName id="4C764D0EFFC3FFD90B7B9F7749AE36D8" box="[549,661,1143,1166]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90B7B9F7749AE36D8" box="[549,661,1143,1166]" italics="true" pageId="9" pageNumber="56">A. riparia</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D0EFFC3FFD90B8C9F77484536D9" authority="Paiva" authorityName="Paiva" box="[722,894,1142,1167]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lyra">
<emphasis id="B902EA9FFFC3FFD90B8C9F77481A36D9" box="[722,801,1142,1167]" italics="true" pageId="9" pageNumber="56">A. lyra</emphasis>
(Paiva)
</taxonomicName>
, and distinguished them by the number of hair patches on the profemur in males. More specifically, males of
<taxonomicName id="4C764D0EFFC3FFD90A0C9F9B48A536E5" box="[850,926,1178,1203]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lyra">
<emphasis id="B902EA9FFFC3FFD90A0C9F9B48A536E5" box="[850,926,1178,1203]" italics="true" pageId="9" pageNumber="56">A. lyra</emphasis>
</taxonomicName>
have the profemur with a small middle patch on a tumescence, whereas males of
<taxonomicName id="4C764D0EFFC3FFD90B759FBF49A13680" box="[555,666,1215,1238]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90B759FBF49A13680" box="[555,666,1215,1238]" italics="true" pageId="9" pageNumber="56">A. riparia</emphasis>
</taxonomicName>
lack the middle patch. Later,
<bibRefCitation id="EFE74B7CFFC3FFD90AB59FBE4FE43681" author="Zettel" box="[1003,1247,1214,1239]" pageId="9" pageNumber="56" refString="Zettel, H. &amp; Chen, P. - P. (1997) Three new taxa of Amemboa ESAKI, 1925, from Thailand and Vietnam. Annales Historico- Naturales Musei Nationalis Hungarici, 89, 93 - 101." type="journal article" year="1997">Zettel &amp; Chen (1997)</bibRefCitation>
synonymized
<taxonomicName id="4C764D0EFFC3FFD90CDA9FBF4BDD36AD" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90CDA9FBF4BDD36AD" italics="true" pageId="9" pageNumber="56">A. riparia</emphasis>
</taxonomicName>
with
<taxonomicName id="4C764D0EFFC3FFD908729FE34A4036AD" box="[300,379,1250,1275]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lyra">
<emphasis id="B902EA9FFFC3FFD908729FE34A4036AD" box="[300,379,1250,1275]" italics="true" pageId="9" pageNumber="56">A. lyra</emphasis>
</taxonomicName>
because the hair patches vary within and among populations in both species, and the male genitalia of both species are not clearly distinguishable. More recently, Polhemus &amp; Tran (2012) reestablished
<taxonomicName id="4C764D0EFFC3FFD90CDA9E074BDD3715" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90CDA9E074BDD3715" italics="true" pageId="9" pageNumber="56">A. riparia</emphasis>
</taxonomicName>
and stated that it can be distinguished from
<taxonomicName id="4C764D0EFFC3FFD90BB49E2B48023715" box="[746,825,1322,1347]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="lyra">
<emphasis id="B902EA9FFFC3FFD90BB49E2B48023715" box="[746,825,1322,1347]" italics="true" pageId="9" pageNumber="56">A. lyra</emphasis>
</taxonomicName>
by the structure of the male fore leg and pygophore. Moreover, the presence of a middle hair patch on the fore leg in
<taxonomicName id="4C764D0EFFC3FFD90ACD9E4F4F3E3730" box="[915,1029,1359,1382]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="riparia">
<emphasis id="B902EA9FFFC3FFD90ACD9E4F4F3E3730" box="[915,1029,1359,1382]" italics="true" pageId="9" pageNumber="56">A. riparia</emphasis>
</taxonomicName>
is variable and can occur at a low frequency (Polhemus &amp; Tran 2012). This species has been recorded from
<collectingCountry id="F361761DFFC3FFD90AB79E734F1937DC" box="[1001,1058,1395,1418]" name="Laos" pageId="9" pageNumber="56">Laos</collectingCountry>
, peninsular
<collectingCountry id="F361761DFFC3FFD90DF39E724E2D37DD" box="[1197,1302,1394,1419]" name="Malaysia" pageId="9" pageNumber="56">Malaysia</collectingCountry>
,
<collectingCountry id="F361761DFFC3FFD90C7D9E724EA337DD" box="[1315,1432,1394,1419]" name="Singapore" pageId="9" pageNumber="56">Singapore</collectingCountry>
,
<collectingCountry id="F361761DFFC3FFD909C99E964BC637F9" box="[151,253,1430,1455]" name="Thailand" pageId="9" pageNumber="56">Thailand</collectingCountry>
, and
<collectingCountry id="F361761DFFC3FFD9081E9E964AA437F9" box="[320,415,1430,1455]" name="Vietnam" pageId="9" pageNumber="56">Vietnam</collectingCountry>
(
<bibRefCitation id="EFE74B7CFFC3FFD908EC9E96483B37F9" author="Polhemus" box="[434,768,1430,1455]" pageId="9" pageNumber="56" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen 1984</bibRefCitation>
,
<bibRefCitation id="EFE74B7CFFC3FFD90A4E9E9648DE37F9" author="Cheng" box="[784,997,1430,1455]" pageId="9" pageNumber="56" refString="Cheng, L., Yang, C. M., Li, D. &amp; Liu, H. (2006) Aquatic Heteroptera (Insecta: Gerromorpha and Nepomorpha) from Xishuangbanna, Yunnan, China. Raffles Bulletin of Zoology, 54, 203 - 214." type="journal article" year="2006">Cheng et al. 2006</bibRefCitation>
, Zettel et al. 2007). In
<collectingCountry id="F361761DFFC3FFD90DA09E964E5F37F9" box="[1278,1380,1430,1455]" name="Thailand" pageId="9" pageNumber="56">Thailand</collectingCountry>
, this species has been collected previously from Chiang Mai, Mae Hong Son, Nan, Phetchabun, Phrae, Songkhla, and Tak provinces (
<bibRefCitation id="EFE74B7CFFC3FFD908189EDE49B737A1" author="Polhemus" box="[326,652,1502,1527]" pageId="9" pageNumber="56" refString="Polhemus, J. T. &amp; Andersen, N. M. (1984) A revision of Amemboa Esaki with notes on the phylogeny and ecological evolution of eotrechine water striders (Insecta, Hemiptera, Gerridae). Steenstrupia, 10, 65 - 111." type="journal article" year="1984">Polhemus &amp; Andersen 1984</bibRefCitation>
,
<bibRefCitation id="EFE74B7CFFC3FFD90BC69EDE484437A1" author="Zettel" box="[664,895,1502,1527]" pageId="9" pageNumber="56" refString="Zettel, H. &amp; Chen, P. - P. (1997) Three new taxa of Amemboa ESAKI, 1925, from Thailand and Vietnam. Annales Historico- Naturales Musei Nationalis Hungarici, 89, 93 - 101." type="journal article" year="1997">Zettel &amp; Chen 1997</bibRefCitation>
,
<bibRefCitation id="EFE74B7CFFC3FFD90AD29EDE4F6737A1" author="Vitheepradit" box="[908,1116,1502,1527]" pageId="9" pageNumber="56" refString="Vitheepradit, A. (2000) The aquatic and semiaquatic Heteroptera of the Phu Pan and Phetchabun mountain ranges of Thailand. M. S. Thesis. University of Missouri, Columbia, Missouri, USA. [total page number not cited]" type="book" year="2000">Vitheepradit 2000</bibRefCitation>
). This species was collected throughout the Phetchabun Mountain Range and in various
<typeStatus id="54CD882FFFC3FFD90A1F9D034847344D" box="[833,892,1539,1563]" pageId="9" pageNumber="56">types</typeStatus>
of habitats (e.g., rock pools, leaf packs, muddy tire ruts, sandy stream margins). Also, we have observed variation in presence or absence of the small middle hair patches on male profemora in specimens collected from Heuy Prom Laeng, Nam Nao NP, Chaiyaphum Province. This species occurred synoptically with
<taxonomicName id="4C764D0EFFC3FFD90B079D6F49C734D1" box="[601,764,1646,1671]" class="Insecta" family="Gerridae" genus="Amemboa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="56" phylum="Arthropoda" rank="species" species="aquafrigida">
<emphasis id="B902EA9FFFC3FFD90B079D6F49C734D1" box="[601,764,1646,1671]" italics="true" pageId="9" pageNumber="56">A. aquafrigida</emphasis>
</taxonomicName>
in rock pools in Loei and Phitsanulok provinces.
</paragraph>
</subSubSection>
<subSubSection id="C36C6506FFC3FFD909999D9248DF35B9" pageId="9" pageNumber="56" type="materials_examined">
<paragraph id="8BC9368DFFC3FFD909999D9248DF35B9" blockId="9.[151,1437,890,2031]" pageId="9" pageNumber="56">
<emphasis id="B902EA9FFFC3FFD909999D924A8934FD" bold="true" box="[199,434,1682,1707]" pageId="9" pageNumber="56">Material examined.</emphasis>
<collectingCountry id="F361761DFFC3FFD908E59D93496B34FD" box="[443,592,1683,1707]" name="Thailand" pageId="9" pageNumber="56">THAILAND</collectingCountry>
:
<emphasis id="B902EA9FFFC3FFD90B039D92483F34FD" bold="true" box="[605,772,1682,1707]" pageId="9" pageNumber="56">Loei Province</emphasis>
: Amphur Phu Rua, Plan Ba Waterfall,
<geoCoordinate id="EE42504AFFC3FFD90D969D934E1F34FD" box="[1224,1316,1683,1707]" direction="north" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="17.383333">17°23'N</geoCoordinate>
<geoCoordinate id="EE42504AFFC3FFD90C739D934EAC34FD" box="[1325,1431,1683,1707]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="101.36667">101°22'E</geoCoordinate>
,
<quantity id="4C8E9B68FFC3FFD909C99DB64BDB3498" box="[151,224,1718,1743]" metricMagnitude="2" metricUnit="m" metricValue="6.49" pageId="9" pageNumber="56" unit="m" value="649.0">649 m</quantity>
,
<date id="FFC8104DFFC3FFD909B39DB74A533499" box="[237,360,1719,1743]" pageId="9" pageNumber="56" value="2004-05-10">10-V-2004</date>
, A. Vitheepradit and T. Prommi, L-675 (3 winged males, 1 wingless male, 2 winged females, 1 wingless female);
<emphasis id="B902EA9FFFC3FFD908389DDA494834A5" bold="true" box="[358,627,1754,1779]" pageId="9" pageNumber="56">Chaiyaphum Province</emphasis>
: Phu Kiew WS, Lam Nam Prom,
<geoCoordinate id="EE42504AFFC3FFD90AA79DDA4F6D34A5" box="[1017,1110,1754,1779]" direction="north" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="16.45">16°27'N</geoCoordinate>
<geoCoordinate id="EE42504AFFC3FFD90D009DDB4FF334A5" box="[1118,1224,1755,1779]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="101.65">101°39'E</geoCoordinate>
,
<quantity id="4C8E9B68FFC3FFD90D8A9DDA4E2534A4" box="[1236,1310,1754,1779]" metricMagnitude="2" metricUnit="m" metricValue="5.89" pageId="9" pageNumber="56" unit="m" value="589.0">589 m</quantity>
,
<date id="FFC8104DFFC3FFD90C759DDB4EA334A5" box="[1323,1432,1755,1779]" pageId="9" pageNumber="56" value="2004-05-02">2-V-2004</date>
, A. Vitheepradit, L-656 (1 winged male); Nam Nao NP, Heuy Prom Laeng,
<geoCoordinate id="EE42504AFFC3FFD90D5E9DFE4F663541" box="[1024,1117,1790,1815]" direction="north" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="16.633333">16°38'N</geoCoordinate>
101°034'E,
<quantity id="4C8E9B68FFC3FFD90DB29DFF4E033540" box="[1260,1336,1791,1815]" metricMagnitude="2" metricUnit="m" metricValue="8.02" pageId="9" pageNumber="56" unit="m" value="802.0">802 m</quantity>
,
<date id="FFC8104DFFC3FFD90C199DFF4BE9356D" pageId="9" pageNumber="56" value="2012-12-20">20-XII- 2012</date>
, KU team, C-239 (1 winged male);
<emphasis id="B902EA9FFFC3FFD90BCF9C2248AC356D" bold="true" box="[657,919,1826,1851]" pageId="9" pageNumber="56">Phetchabun Province</emphasis>
: Khao Kho NP, Lam Nam Khek,
<geoCoordinate id="EE42504AFFC3FFD90C619C224EA7356D" box="[1343,1436,1826,1851]" direction="north" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="16.566668">16°34'N</geoCoordinate>
<geoCoordinate id="EE42504AFFC3FFD909C99C464A3B3509" box="[151,256,1862,1887]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="100.96667">100°58'E</geoCoordinate>
,
<quantity id="4C8E9B68FFC3FFD9084F9C464A653508" box="[273,350,1862,1887]" metricMagnitude="2" metricUnit="m" metricValue="7.6" pageId="9" pageNumber="56" unit="m" value="760.0">760 m</quantity>
,
<date id="FFC8104DFFC3FFD908309C4749333509" box="[366,520,1863,1887]" pageId="9" pageNumber="56" value="2012-08-20">20-VIII-2012</date>
, KU team, C-178 (1 winged male, 1 winged female);
<emphasis id="B902EA9FFFC3FFD90DCE9C464EAF3509" bold="true" box="[1168,1428,1862,1887]" pageId="9" pageNumber="56">Phitsanulok Province</emphasis>
: Thung Salaeng NP, Huai Tub Dah Mi,
<geoCoordinate id="EE42504AFFC3FFD90B099C6A498835D5" box="[599,691,1898,1923]" direction="north" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="16.833334">16°50'N</geoCoordinate>
<geoCoordinate id="EE42504AFFC3FFD90BE39C6A481D35D5" box="[701,806,1898,1923]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="100.85">100°51'E</geoCoordinate>
,
<quantity id="4C8E9B68FFC3FFD90A6A9C6B484435D4" box="[820,895,1899,1923]" metricMagnitude="2" metricUnit="m" metricValue="4.4" pageId="9" pageNumber="56" unit="m" value="440.0">440 m</quantity>
,
<date id="FFC8104DFFC3FFD90AD29C6B4F3F35D5" box="[908,1028,1899,1923]" pageId="9" pageNumber="56" value="2002-03-08">8-III-2002</date>
, R.W. Sites, A. Vitheepradit and K. Kirawanich, L-274 (3 winged males, 4 winged females); same locality,
<date id="FFC8104DFFC3FFD90AE89C8F4F0A35F1" box="[950,1073,1935,1959]" pageId="9" pageNumber="56" value="2004-05-13">13-V-2004</date>
, A. Vitheepradit and T. Prommi, L-683 (9 winged males, 1 wingless male, 7 winged females); Thung Salaeng NP, Kaeng So Pha,
<geoCoordinate id="EE42504AFFC3FFD90D959CB24E1C359D" box="[1227,1319,1970,1995]" direction="north" orientation="latitude" pageId="9" pageNumber="56" precision="925" value="16.833334">16°50'N</geoCoordinate>
<geoCoordinate id="EE42504AFFC3FFD90C709CB24EAC359D" box="[1326,1431,1970,1995]" direction="east" orientation="longitude" pageId="9" pageNumber="56" precision="925" value="101.85">101°51'E</geoCoordinate>
,
<quantity id="4C8E9B68FFC3FFD909C99CD74BDB35B8" box="[151,224,2007,2031]" metricMagnitude="2" metricUnit="m" metricValue="3.97" pageId="9" pageNumber="56" unit="m" value="397.0">397 m</quantity>
,
<date id="FFC8104DFFC3FFD909B29CD74A5D35B9" box="[236,358,2007,2031]" pageId="9" pageNumber="56" value="2004-05-11">11-V-2004</date>
, A. Vitheepradit and T. Prommi, L-679 (1 winged male).
</paragraph>
</subSubSection>
</treatment>
</document>