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<document id="E42261ABE63DBF85F907E0E2A414A44D" ID-CLB-Dataset="298072" ID-DOI="10.5252/geodiversitas2024v46a7" ID-GBIF-Dataset="0ac5cd8a-73cd-4cd0-9171-3a8111f02165" ID-ISSN="1638-9395" ID-Zenodo-Dep="11408605" ID-ZooBank="urn:lsid:zoobank.org:pub:7D6B75B9-71D3-48B6-8217-261B91381E99" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.tables_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1717238347288" checkinUser="plazi" docAuthor="Allaire, Ninon, Rustán, Juan J., Korn, Dieter, Vaccari, Emilio, Ezpeleta, Miguel &amp; Balseiro, Diego" docDate="2024" docId="03DF879650081208FF69CE403A1BF853" docLanguage="en" docName="geodiversitas.2024.46.7.pdf" docOrigin="Geodiversitas 46 (7)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2024v46a7.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Epitornoceras baldisi" docType="treatment" docVersion="5" lastPageNumber="337" masterDocId="FFE6FFEE50011204FF96C8123E68FFB8" masterDocTitle="The late Givetian (Middle Devonian) ammonoid Epitornoceras Frech, 1902 from Argentina (southwestern Gondwana)" masterLastPageNumber="341" masterPageNumber="327" pageNumber="334" updateTime="1717509125305" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="3826A2B09C516EE5A6A5BD9FB8293EF2">The late Givetian (Middle Devonian) ammonoid Epitornoceras Frech, 1902 from Argentina (southwestern Gondwana)</mods:title>
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<mods:namePart id="2316A2933165251F3066943A956D1C97">Allaire, Ninon</mods:namePart>
<mods:affiliation id="0E7589B5CA3FE01EFB5C0B4565188527">Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Centro de Investigaciones en Ciencias de la Tierra (CICTERRA), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) ninon. allaire @ gmail. com (corresponding author)</mods:affiliation>
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<mods:namePart id="6DF93B448054B20F16EBC77EC91F3414">Rustán, Juan J.</mods:namePart>
<mods:affiliation id="B4278A2B966671534A557DD4EF1B8C97">Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Centro de Investigaciones en Ciencias de la Tierra (CICTERRA), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Universidad Nacional de La Rioja, Avenida Luis M. de la Fuente s / n, F 5300, La Rioja (Argentina) juanjorustan @ gmail. com</mods:affiliation>
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<mods:namePart id="EAD302B6F4A74102033045EF70CB649B">Vaccari, Emilio</mods:namePart>
<mods:affiliation id="E33B48A2BA8F07B10B4D5295C7C2C1A1">Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Centro de Investigaciones en Ciencias de la Tierra (CICTERRA), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Universidad Nacional de La Rioja, Avenida Luis M. de la Fuente s / n, F 5300, La Rioja (Argentina)</mods:affiliation>
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<mods:namePart id="DCCAFC9545BBE3E46782E8189DA1708A">Ezpeleta, Miguel</mods:namePart>
<mods:affiliation id="3349BD020125D2F4ADAEC36956A9F89E">Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Centro de Investigaciones en Ciencias de la Tierra (CICTERRA), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Universidad Nacional de La Rioja, Avenida Luis M. de la Fuente s / n, F 5300, La Rioja (Argentina) evaccari @ unc. edu. ar miguelezpeleta @ gmail. com</mods:affiliation>
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<mods:affiliation id="98B87148D2C60F8EFA8B688F6997C726">Facultad de Ciencias Exactas, Físicas y Naturales, Universidad Nacional de Córdoba, Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) and Centro de Investigaciones en Ciencias de la Tierra (CICTERRA), Consejo Nacional de Investigaciones Científicas y Técnicas (CONICET), Avenida Vélez Sarsfield 1611, CP X 5016 GCA Córdoba (Argentina) balsedie @ gmail. com</mods:affiliation>
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<treatment id="03DF879650081208FF69CE403A1BF853" ID-DOI="http://doi.org/10.5281/zenodo.11473440" ID-GBIF-Taxon="232131763" ID-Zenodo-Dep="11473440" LSID="urn:lsid:plazi:treatment:03DF879650081208FF69CE403A1BF853" httpUri="http://treatment.plazi.org/id/03DF879650081208FF69CE403A1BF853" lastPageId="12" lastPageNumber="337" pageId="9" pageNumber="334">
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<taxonomicName id="4C764D035008120DFF69CE403CE4F9D4" authority="(Leanza, 1968)" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[255,652,1618,1645]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" order="Goniatitida" pageId="9" pageNumber="334" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA925008120DFF69CE403F8AF9D5" bold="true" box="[255,482,1618,1645]" italics="true" pageId="9" pageNumber="334">Epitornoceras baldisi</emphasis>
(
<bibRefCitation id="EFE74B715008120DFE67CE403CECF9D4" author="LEANZA A. F." box="[497,644,1618,1644]" pageId="9" pageNumber="334" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza, 1968</bibRefCitation>
)
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<paragraph id="8BC936805008120DFE18CE603F95F934" blockId="9.[398,509,1650,1677]" box="[398,509,1650,1677]" pageId="9" pageNumber="334">
(
<figureCitation id="134D2A055008120DFE01CE603FB0F934" box="[407,472,1650,1676]" captionStart="FIG" captionStartId="9.[133,143,615,632]" captionTargetBox="[279,1308,194,590]" captionTargetId="figure-697@9.[279,1121,194,590]" captionTargetPageId="9" captionText="FIG. 6. — Historical material of Epitornoceras baldisi (Leanza, 1968), identified as Tornoceras baldisi Leanza, 1968 by Leanza (1968), from the Chavela Creek (“Quebrada de la Chavela”),Chavela Member, Chigua Formation (San Juan Province, Argentina): A, B, holotype CPUBA 8088 (part and counterpart showing the external mold of the conch); C, additional specimen CPUBA 8089. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.11408615" httpUri="https://zenodo.org/record/11408615/files/figure.png" pageId="9" pageNumber="334">Figs 6</figureCitation>
;
<figureCitation id="134D2A055008120DFE70CE613F9CF935" box="[486,500,1651,1677]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">7</figureCitation>
)
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<taxonomicName id="4C764D035008120DFF12CEB83FDCF97A" ID-CoL="57G3S" authority="Leanza, 1968" authorityName="Leanza" authorityYear="1968" box="[132,436,1706,1730]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" order="Goniatitida" pageId="9" pageNumber="334" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA925008120DFF12CEB83F43F97A" box="[132,299,1706,1730]" italics="true" pageId="9" pageNumber="334">Tornoceras baldisi</emphasis>
<bibRefCitation id="EFE74B715008120DFEA7CEB83FDCF97A" author="LEANZA A. F." box="[305,436,1706,1730]" pageId="9" pageNumber="334" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza, 1968</bibRefCitation>
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: figs 1-4.
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<paragraph id="8BC936805008120DFF12CECA3D6FF948" blockId="9.[132,775,1752,1776]" box="[132,775,1752,1776]" pageId="9" pageNumber="334">
non
<taxonomicName id="4C764D035008120DFF26CECB3C00F948" ID-CoL="57G3S" authority="House, 1978" authorityName="House" authorityYear="1978" box="[176,616,1752,1776]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" isUncertain="true" kingdom="Animalia" order="Goniatitida" pageId="9" pageNumber="334" phylum="Mollusca" rank="species" species="baldisi" subGenus="Tornoceras">
<emphasis id="B902EA925008120DFF26CECB3F7AF948" box="[176,274,1753,1776]" italics="true" pageId="9" pageNumber="334">Tornoceras</emphasis>
(
<emphasis id="B902EA925008120DFE89CECB3FE9F948" box="[287,385,1753,1776]" italics="true" pageId="9" pageNumber="334">Tornoceras</emphasis>
) cf.
<emphasis id="B902EA925008120DFE3ECECA3F8EF948" box="[424,486,1752,1776]" italics="true" pageId="9" pageNumber="334">baldisi</emphasis>
<bibRefCitation id="EFE74B715008120DFE7ACECA3C00F948" author="HOUSE M. R." box="[492,616,1752,1776]" pageId="9" pageNumber="334" pagination="1 - 70" refId="ref10787" refString="HOUSE M. R. 1978. - Devonian Ammonoids from the Appalachians and their bearing on international zonation and correlation. Special Papers in Palaeontology 21: 1 - 70." type="journal article" year="1978">House, 1978</bibRefCitation>
</taxonomicName>
: pl. 6, figs 9, 10.
</paragraph>
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. —
<materialsCitation id="3B1E3CDD5008120DFEADCF143FC5F8EB" ID-GBIF-Occurrence="4872739301" collectionCode="CPUBA" country="Argentina" location="Argentina" pageId="9" pageNumber="334" specimenCode="CPUBA 8088" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54CD88225008120DFEADCF143FF3F8A6" box="[315,411,1798,1822]" pageId="9" pageNumber="334" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="F36176105008120DFE35CF153C6EF8A7" box="[419,518,1799,1823]" name="Argentina" pageId="9" pageNumber="334">Argentina</collectingCountry>
</emphasis>
• Specimen
<specimenCode id="DBD09EFB5008120DFDEBCF143D6DF8A6" box="[637,773,1798,1822]" collectionCode="CPUBA" pageId="9" pageNumber="334">CPUBA 8088</specimenCode>
designed by
<bibRefCitation id="EFE74B715008120DFF6ECF333FEBF880" author="LEANZA A. F." box="[248,387,1824,1848]" pageId="9" pageNumber="334" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza (1968)</bibRefCitation>
, stored at the University of Buenos Aires (re-illustrated here, see
<figureCitation id="134D2A055008120DFEF1CF293FF7F8EB" box="[359,415,1851,1875]" captionStart="FIG" captionStartId="9.[133,143,615,632]" captionTargetBox="[279,1308,194,590]" captionTargetId="figure-697@9.[279,1121,194,590]" captionTargetPageId="9" captionText="FIG. 6. — Historical material of Epitornoceras baldisi (Leanza, 1968), identified as Tornoceras baldisi Leanza, 1968 by Leanza (1968), from the Chavela Creek (“Quebrada de la Chavela”),Chavela Member, Chigua Formation (San Juan Province, Argentina): A, B, holotype CPUBA 8088 (part and counterpart showing the external mold of the conch); C, additional specimen CPUBA 8089. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.11408615" httpUri="https://zenodo.org/record/11408615/files/figure.png" pageId="9" pageNumber="334">Fig. 6</figureCitation>
).
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</paragraph>
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<subSubSection id="C36C650B5008120DFF12CF7B3B24FC91" pageId="9" pageNumber="334" type="diagnosis">
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. —
<taxonomicName id="4C764D035008120DFE86CF783FE1F839" authorityName="Frech" authorityYear="1902" box="[272,393,1898,1921]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="9" pageNumber="334" phylum="Mollusca" rank="genus">
<emphasis id="B902EA925008120DFE86CF783FE1F839" box="[272,393,1898,1921]" italics="true" pageId="9" pageNumber="334">Epitornoceras</emphasis>
</taxonomicName>
reaching at least
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conch diameter. Extremely discoidal and involute adult stage (CWI
<emphasis id="B902EA925008120DFDC9CF963C06F823" box="[607,622,1924,1947]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
0.25; umbilicus closed at
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dm); whorl profile strongly compressed (WWI
<emphasis id="B902EA925008120DFF10CFAB3EFEF868" box="[134,150,1977,2000]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
0.40 at
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dm). Flanks weakly convex, convergent, umbilical margin continuously rounded. Without any ventrolateral grooves. Suture line with broadly rounded, nearly symmetric adventive lobe with convergent flanks; adventive lobe wider than deep.
</paragraph>
</subSubSection>
<subSubSection id="C36C650B5008120DFCBBCB533B31FB9A" pageId="9" pageNumber="334" type="materials_examined">
<paragraph id="8BC936805008120DFCBBCB533BC1FC36" blockId="9.[813,1457,833,910]" pageId="9" pageNumber="334">
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<collectionCode id="ED67AE455008120DFCF3CB543DA8FCEF" box="[869,960,838,855]" pageId="9" pageNumber="334">LOCALITY</collectionCode>
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<collectionCode id="ED67AE455008120DFC51CB543D99FCEF" box="[967,1009,838,855]" pageId="9" pageNumber="334">AND</collectionCode>
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. — Uppermost part of the Chavela Member, uppermost part of the Chigua Formation, Chinguillos Group, north of Angualasto town,
<collectingRegion id="49B2F8625008120DFB12CB643B5FFC36" box="[1156,1335,886,910]" country="Argentina" name="San Juan" pageId="9" pageNumber="334">San Juan province</collectingRegion>
,
<collectingCountry id="F36176105008120DFAD7CB643BCDFC36" box="[1345,1445,886,910]" name="Argentina" pageId="9" pageNumber="334">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="8BC936805008120DFCBBCBB73D83FC4A" blockId="9.[812,1455,933,1010]" pageId="9" pageNumber="334">
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. — The species is known only from the
<typeStatus id="54CD88225008120DFAAFCBB43B0BFC06" box="[1337,1379,934,958]" pageId="9" pageNumber="334">type</typeStatus>
locality (Chavela Member, Chigua Formation,
<collectingRegion id="49B2F8625008120DFB3ACBD23B09FC60" box="[1196,1377,960,984]" country="Argentina" name="San Juan" pageId="9" pageNumber="334">San Juan province</collectingRegion>
,
<collectingCountry id="F36176105008120DFAFACBD23D3EFC4A" name="Argentina" pageId="9" pageNumber="334">Argentina</collectingCountry>
), late Givetian.
</paragraph>
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. — Ten specimens, INGEO-PI-2002 to 2011.
</paragraph>
</subSubSection>
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</paragraph>
<paragraph id="8BC936805008120DFCBBCC663A63F934" blockId="9.[810,1457,1109,2028]" pageId="9" pageNumber="334">
The conch is extremely discoidal (CWI
<emphasis id="B902EA925008120DFB54CC673ABAFB36" box="[1218,1234,1141,1166]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
0.25) with a strongly compressed whorl section (WWI
<emphasis id="B902EA925008120DFB11CC873AFFFB16" box="[1159,1175,1173,1198]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
0.40) in the subadult stage (
<quantity id="4C8E9B655008120DFCA4CCA63DCAFB76" box="[818,930,1204,1231]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="9" pageNumber="334" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">30-50 mm</quantity>
dm); its umbilicus is completely closed (specimens INGEO-PI-2011, INGEO-PI-2007 and INGEO-PI-2004;
<figureCitation id="134D2A055008120DFCBACCE13D02FAB6" box="[812,874,1267,1294]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">Fig. 7</figureCitation>
A-C, I). The flanks are weakly convex and convergent, and the venter is narrowly rounded; the umbilical margin is continuously rounded. At a smaller diameter (
<quantity id="4C8E9B655008120DFAADCD213BD9FAF6" box="[1339,1457,1331,1358]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="9" pageNumber="334" unit="mm" value="25.0" valueMax="30.0" valueMin="20.0">20-30 mm</quantity>
dm), the conch is also discoidal, but has a slightly opened umbilicus (specimens INGEO-PI-2005 and INGEO-PI-2002;
<figureCitation id="134D2A055008120DFCBCCD813D0FFA16" box="[810,871,1427,1454]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">Fig. 7</figureCitation>
F-H). At about
<quantity id="4C8E9B655008120DFB99CD813A35FA16" box="[1039,1117,1427,1454]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="9" pageNumber="334" unit="mm" value="20.0">20 mm</quantity>
dm, the coiling rate is very high (WER
<emphasis id="B902EA925008120DFCE0CDA63DEEFA75" box="[886,902,1460,1485]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
2.35). Based on specimen INGEO-PI-2003 (
<figureCitation id="134D2A055008120DFACFCDA13D28FA55" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">Fig. 7Q, L</figureCitation>
), the morphology of the earlier whorls can be investigated by the use of latex casts: around
<quantity id="4C8E9B655008120DFB12CDE03A9FF9B5" box="[1156,1271,1522,1549]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="9" pageNumber="334" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10-15 mm</quantity>
dm, the conch is thinly discoidal (CWI
<emphasis id="B902EA925008120DFB83CE013A4EF994" box="[1045,1062,1555,1580]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
0.35-0.40) with a weakly compressed whorl section (WWI
<emphasis id="B902EA925008120DFB83CE213A4EF9F4" box="[1045,1062,1587,1612]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
0.50-0.60) and a high whorl expansion rate (WER
<emphasis id="B902EA925008120DFC4CCE413D83F9D4" box="[986,1003,1619,1644]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
2.10). Surprisingly, the umbilicus appears to be entirely closed.
</paragraph>
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In most specimens, partially preserved suture lines or septal surfaces are visible. The entire suture line can be reconstructed for specimen INGEO-PI-2007 (
<figureCitation id="134D2A055008120DFB13CEC03A9CF954" box="[1157,1268,1746,1772]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">Fig. 7C, E</figureCitation>
); at wh
<emphasis id="B902EA925008120DFAD2CEC13B3CF954" box="[1348,1364,1747,1772]" italics="true" pageId="9" pageNumber="334">c.</emphasis>
<quantity id="4C8E9B655008120DFACDCEC03BC3F954" box="[1371,1451,1746,1772]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="9" pageNumber="334" unit="mm" value="18.0">18 mm</quantity>
, it consists of a narrow and deep external lobe, a narrow and subacute E/A saddle, a broadly rounded, wide and deep nearly symmetric adventive lobe with convergent flanks, followed by a similarly broadly rounded and deep but wider saddle. Suture lines are also relatively well preserved in specimen INGEO- PI-2004 (
<figureCitation id="134D2A055008120DFC0FCF833A63F813" box="[921,1035,1937,1964]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">Fig. 7D, I</figureCitation>
); the shape of the adventive lobe and the following saddle is similar to specimen INGEO-PI-2007 (
<figureCitation id="134D2A055008120DFCA0CFC33DE0F853" box="[822,904,2001,2028]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="9" pageNumber="334">Fig. 7E</figureCitation>
); the external lobe and the ventrolateral saddle are not visible. Interestingly, the larger specimens show a septal thickening when comparing with smaller ones (i.e., increase in thickness of the septal wall when it meets the shell; compare the wideness of the suture lines of specimens INGEO- PI-2007 and INGEO-PI-2004,
<figureCitation id="134D2A05500B120EFE7DCDC13C30FA55" box="[491,600,1491,1517]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="10" pageNumber="335">Fig. 7C, I</figureCitation>
, with specimen INGEO-PI-2002,
<figureCitation id="134D2A05500B120EFEC7CDE13FC1F9B5" box="[337,425,1523,1549]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="10" pageNumber="335">Fig. 7G</figureCitation>
). This trend of increasing septal thickness toward adulthood (seeKlug
<emphasis id="B902EA92500B120EFD9ACE013C54F994" box="[524,572,1554,1580]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
2015b) was already documented in several ammonoids (
<bibRefCitation id="EFE74B71500B120EFD92CE213CA3F9F5" author="WESTERMANN G. E. G." box="[516,715,1587,1613]" pageId="10" pageNumber="335" pagination="1 - 39" refId="ref12309" refString="WESTERMANN G. E. G. 1971. - Form, structure and function of shell and siphuncle in coiled Mesozoic ammonoids. Royal Ontario Museum, Life Science Contributions 78: 1 - 39. https: // www. biodiversitylibrary. org / page / 36138916" type="journal article" year="1971">Westermann 1971</bibRefCitation>
) as in modern nautilids (
<bibRefCitation id="EFE74B71500B120EFED8CE403C53F9D4" author="COLLINS D. &amp; WARD P. D." box="[334,571,1618,1644]" pageId="10" pageNumber="335" pagination="421 - 432" refId="ref10107" refString="COLLINS D. &amp; WARD P. D. 1987. - Adolescent Growth and Maturity in Nautilus, in SAUNDERS W. B. &amp; LANDMAN N. H. (eds), Nautilus: The Biology and Paleobiology of a Living Fossil, Reprint with additions. Plenum Press (Topics in Geobiology; 6), New York: 421 - 432. https: // doi. org / 10.1007 / 978 - 90 - 481 - 3299 - 7 _ 29" type="book chapter" year="1987">Collins &amp; Ward 1987</bibRefCitation>
).
</paragraph>
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. 7. — New specimens of
<taxonomicName id="4C764D03500B120EFEEECCA43CCEFB7F" authority="(Leanza, 1968)" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[376,678,1206,1223]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500B120EFEEECCA43C4EFB7F" box="[376,550,1206,1223]" italics="true" pageId="10" pageNumber="335">Epitornoceras baldisi</emphasis>
(
<bibRefCitation id="EFE74B71500B120EFDA7CCA43CCAFB7F" author="LEANZA A. F." box="[561,674,1206,1223]" pageId="10" pageNumber="335" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza, 1968</bibRefCitation>
)
</taxonomicName>
from the type area of the Chigua Formation (San Juan Province, Argentina):
<emphasis id="B902EA92500B120EFAB8CCA43B53FB7F" bold="true" box="[1326,1339,1206,1223]" pageId="10" pageNumber="335">A</emphasis>
,
<emphasis id="B902EA92500B120EFAD0CCA43B3BFB7F" bold="true" box="[1350,1363,1206,1223]" pageId="10" pageNumber="335">B</emphasis>
, specimen INGEO-PI-2011;
<emphasis id="B902EA92500B120EFE85CCDF3F49FB66" bold="true" box="[275,289,1229,1246]" pageId="10" pageNumber="335">C</emphasis>
, specimen INGEO-PI-2007;
<emphasis id="B902EA92500B120EFD84CCDF3C48FB66" bold="true" box="[530,544,1229,1246]" pageId="10" pageNumber="335">D</emphasis>
, suture line of specimen INGEO-PI- 2004 at wh
<emphasis id="B902EA92500B120EFC2BCCDF3DA5FB66" box="[957,973,1229,1246]" italics="true" pageId="10" pageNumber="335">c.</emphasis>
33 mm;
<emphasis id="B902EA92500B120EFB8FCCDF3A4DFB66" bold="true" box="[1049,1061,1229,1246]" pageId="10" pageNumber="335">E</emphasis>
, reconstructed suture line based on specimen INGEO-PI-2007 at wh
<emphasis id="B902EA92500B120EFEA9CCF13F38FB4C" box="[319,336,1251,1268]" italics="true" pageId="10" pageNumber="335">c.</emphasis>
18 mm;
<emphasis id="B902EA92500B120EFE0ECCF13FCBFB4C" bold="true" box="[408,419,1251,1268]" pageId="10" pageNumber="335">F</emphasis>
, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch);
<emphasis id="B902EA92500B120EFBC6CCF13A36FB4C" bold="true" box="[1104,1118,1251,1268]" pageId="10" pageNumber="335">G</emphasis>
,
<emphasis id="B902EA92500B120EFBFECCF13A1EFB4C" bold="true" box="[1128,1142,1251,1268]" pageId="10" pageNumber="335">H</emphasis>
, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch);
<emphasis id="B902EA92500B120EFDA9CCE83C2DFAB3" bold="true" box="[575,581,1274,1291]" pageId="10" pageNumber="335">I</emphasis>
, specimen INGEO-PI-2004;
<emphasis id="B902EA92500B120EFCA7CCE83D53FAB3" bold="true" box="[817,827,1274,1291]" pageId="10" pageNumber="335">J</emphasis>
, specimen INGEO-PI-2006;
<emphasis id="B902EA92500B120EFBBECCE83A5DFAB3" bold="true" box="[1064,1077,1274,1291]" pageId="10" pageNumber="335">K</emphasis>
,
<emphasis id="B902EA92500B120EFBD6CCE83A23FAB3" bold="true" box="[1088,1099,1274,1291]" pageId="10" pageNumber="335">L</emphasis>
, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K).
</paragraph>
</caption>
<paragraph id="8BC93680500B120EFF0DCE603C6BF974" blockId="10.[130,776,1363,1741]" pageId="10" pageNumber="335">
Although cracked, the shell is partially preserved in
<specimenCount id="9D70FD09500B120EFD49CE613E83F914" count="1" pageId="10" pageNumber="335" type="generic">one specimen</specimenCount>
(INGEO-PI-2011,
<figureCitation id="134D2A05500B120EFE50CE803C71F914" box="[454,537,1682,1709]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="10" pageNumber="335">Fig. 7A</figureCitation>
); it exhibits iridescent colors. Growth lines are not visible.
</paragraph>
</subSubSection>
<subSubSection id="C36C650B500B1208FF12CEE03A1BF853" lastPageId="12" lastPageNumber="337" pageId="10" pageNumber="335" type="discussion">
<paragraph id="8BC93680500B120EFF12CEE03E80F8B2" blockId="10.[131,776,1778,2028]" box="[132,232,1778,1804]" pageId="10" pageNumber="335">
<smallCapsWord id="8D2FA05C500B120EFF12CEE03E80F8B2" baselines="1797,1797" box="[132,232,1778,1804]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Remarks" pageId="10" pageNumber="335">REMARKS</smallCapsWord>
</paragraph>
<paragraph id="8BC93680500B120EFF12CF003AC2FA36" blockId="10.[131,776,1778,2028]" lastBlockId="10.[810,1457,1363,2028]" pageId="10" pageNumber="335">
This material fits well with the description of
<bibRefCitation id="EFE74B71500B120EFDFCCF003F76F8F4" author="KORN D. &amp; BOCKWINKEL J." pageId="10" pageNumber="335" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel (2022)</bibRefCitation>
defining the genus
<taxonomicName id="4C764D03500B120EFE62CF203D6FF8F4" authority="Frech, 1902" authorityName="Frech" authorityYear="1902" box="[500,775,1841,1868]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500B120EFE62CF203C16F8F3" box="[500,638,1842,1867]" italics="true" pageId="10" pageNumber="335">Epitornoceras</emphasis>
<bibRefCitation id="EFE74B71500B120EFD13CF233D6FF8F4" author="FRECH F." box="[645,775,1841,1868]" pageId="10" pageNumber="335" pagination="27 - 112" refId="ref10386" refString="FRECH F. 1902. - Uber devonische Ammoneen. Beitrage zur Palaontologie Osterreich-Ungarns und des Orients. Wien and Leipzig 14: 27 - 112 (in German)." type="journal article" year="1902">Frech, 1902</bibRefCitation>
</taxonomicName>
with a compressed and involute conch, a completely closed umbilicus in the pre-adult stage but slightly opened in the juvenile stage, sutural formula E A L I with broadly rounded adventive lobe, narrow and subacute ventrolateral saddle.
<taxonomicName id="4C764D03500B120EFF12CFC03F66F853" authorityName="Frech" authorityYear="1902" box="[132,270,2002,2027]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500B120EFF12CFC03F66F853" box="[132,270,2002,2027]" italics="true" pageId="10" pageNumber="335">Epitornoceras</emphasis>
</taxonomicName>
differs from the other genera of the subfamily
<taxonomicName id="4C764D03500B120EFCBBCD413DA7FAD5" authorityName="von Arthaber" authorityYear="1911" box="[813,975,1363,1389]" class="Cephalopoda" family="Tornoceratidae" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="subFamily" subFamily="Tornoceratinae">Tornoceratinae</taxonomicName>
by its very narrow and subacute ventrolateral saddle (
<bibRefCitation id="EFE74B71500B120EFCE9CD663AF3FA35" author="KORN D. &amp; BOCKWINKEL J." box="[895,1179,1395,1422]" pageId="10" pageNumber="335" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
).
</paragraph>
<paragraph id="8BC93680500B120EFCD5CD863BC1F894" blockId="10.[810,1457,1363,2028]" pageId="10" pageNumber="335">
Four other species of
<taxonomicName id="4C764D03500B120EFBA1CD863AABFA15" authorityName="Frech" authorityYear="1902" box="[1079,1219,1428,1453]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500B120EFBA1CD863AABFA15" box="[1079,1219,1428,1453]" italics="true" pageId="10" pageNumber="335">Epitornoceras</emphasis>
</taxonomicName>
have been described:
<taxonomicName id="4C764D03500B120EFCB9CDA63A8DFA75" authority="(Frech, 1888)" baseAuthorityName="Frech" baseAuthorityYear="1888" box="[815,1253,1459,1485]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="mithracoides">
<emphasis id="B902EA92500B120EFCB9CDA63A2EFA75" box="[815,1094,1459,1485]" italics="true" pageId="10" pageNumber="335">Epitornoceras mithracoides</emphasis>
(Frech, 1888)
</taxonomicName>
from the Rhenish Mountains of
<collectingCountry id="F3617610500B120EFC2BCDC13A76FA55" box="[957,1054,1491,1517]" name="Germany" pageId="10" pageNumber="335">Germany</collectingCountry>
(
<bibRefCitation id="EFE74B71500B120EFBBCCDC13B67FA55" author="BOCKWINKEL J. &amp; KORN D. &amp; EBBIGHAUSEN V. &amp; GRAF S." box="[1066,1295,1491,1517]" pageId="10" pageNumber="335" pagination="257 - 274" refId="ref9432" refString="BOCKWINKEL J., KORN D., EBBIGHAUSEN V. &amp; GRAF S. 2013. - Late Givetian ammonoids from Hagen-Herbeck, Donnerkuhle quarry (Devonian, Rhenish Mountains). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 270 (3): 257 - 274. https: // doi. org / 10.1127 / 0077 - 7749 / 2013 / 0369" type="journal article" year="2013">
Bockwinkel
<emphasis id="B902EA92500B120EFB31CDC63ABFFA55" box="[1191,1239,1491,1517]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFE74B71500B120EFA8DCDC13DC9F9B5" author="BOCKWINKEL J. &amp; KORN D." pageId="10" pageNumber="335" pagination="351 - 363" refId="ref9339" refString="BOCKWINKEL J. &amp; KORN D. 2015. - Late Givetian ammonoids from Oberberge (Middle Devonian; Rhenish Mountains). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 278 (3): 351 - 363. https: // doi. org / 10.1127 / njgpa / 2015 / 0534" type="journal article" year="2015">Bockwinkel &amp; Korn 2015</bibRefCitation>
;
<bibRefCitation id="EFE74B71500B120EFC24CDE13AA7F9B5" author="KORN D. &amp; BOCKWINKEL J." box="[946,1231,1523,1549]" pageId="10" pageNumber="335" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
),
<taxonomicName id="4C764D03500B120EFB73CDE13A89F995" authority="Korn &amp; Bockwinkel, 2022" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="transmediterraneum">
<emphasis id="B902EA92500B120EFB73CDE13DAEF994" italics="true" pageId="10" pageNumber="335">Epitornoceras transmediterraneum</emphasis>
<bibRefCitation id="EFE74B71500B120EFC5DCE013A89F995" author="KORN D. &amp; BOCKWINKEL J." box="[971,1249,1554,1581]" pageId="10" pageNumber="335" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel, 2022</bibRefCitation>
</taxonomicName>
from the Anti-Atlas of
<collectingCountry id="F3617610500B120EFCD1CE213DCFF9F5" box="[839,935,1587,1613]" name="Morocco" pageId="10" pageNumber="335">Morocco</collectingCountry>
(
<bibRefCitation id="EFE74B71500B120EFC2FCE203AC7F9F5" author="BOCKWINKEL J. &amp; BECKER R. T. &amp; EBBIGHAUSEN V." box="[953,1199,1586,1613]" pageId="10" pageNumber="335" pagination="61 - 128" refId="ref9393" refString="BOCKWINKEL J., BECKER R. T. &amp; EBBIGHAUSEN V. 2009. - Upper Givetian ammonoids from Dar Kaoua (Tafilalt, SE Anti-Atlas, Morocco). Berliner palaobiologische Abhandlungen 10: 61 - 128" type="journal article" year="2009">
Bockwinkel
<emphasis id="B902EA92500B120EFBA8CE213A19F9F4" box="[1086,1137,1586,1612]" italics="true" pageId="10" pageNumber="335">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="EFE74B71500B120EFB2BCE213A9CF9F5" author="BOCKWINKEL J. &amp; KORN D. &amp; EBBIGHAUSEN V. &amp; GRAF S." box="[1213,1268,1587,1613]" pageId="10" pageNumber="335" pagination="257 - 274" refId="ref9432" refString="BOCKWINKEL J., KORN D., EBBIGHAUSEN V. &amp; GRAF S. 2013. - Late Givetian ammonoids from Hagen-Herbeck, Donnerkuhle quarry (Devonian, Rhenish Mountains). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 270 (3): 257 - 274. https: // doi. org / 10.1127 / 0077 - 7749 / 2013 / 0369" type="journal article" year="2013">2013</bibRefCitation>
,
<bibRefCitation id="EFE74B71500B120EFA95CE213B52F9F5" author="BOCKWINKEL J. &amp; BECKER R. T. &amp; ABOUSSALAM Z. S." box="[1283,1338,1587,1613]" pageId="10" pageNumber="335" pagination="307 - 354" refId="ref9559" refString="BOCKWINKEL J., BECKER R. T. &amp; ABOUSSALAM Z. S. 2017. - Ammonoids from the late Givetian Taouzites Bed of Ouidane Chebbi (eastern Tafilalt, SE Morocco). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 284 (3): 307 - 354. https: // doi. org / 10.1127 / njgpa / 2017 / 0664" type="journal article" year="2017">2017</bibRefCitation>
;
<bibRefCitation id="EFE74B71500B120EFADECE203D80F9D5" author="BOCKWINKEL J. &amp; KORN D." pageId="10" pageNumber="335" pagination="351 - 363" refId="ref9339" refString="BOCKWINKEL J. &amp; KORN D. 2015. - Late Givetian ammonoids from Oberberge (Middle Devonian; Rhenish Mountains). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 278 (3): 351 - 363. https: // doi. org / 10.1127 / njgpa / 2015 / 0534" type="journal article" year="2015">Bockwinkel &amp; Korn 2015</bibRefCitation>
;
<bibRefCitation id="EFE74B71500B120EFC60CE413B7AF9D4" author="KORN D. &amp; BOCKWINKEL J." box="[1014,1298,1618,1645]" pageId="10" pageNumber="335" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
),
<taxonomicName id="4C764D03500B120EFAB1CE413DF1F934" baseAuthorityName="Hall" baseAuthorityYear="1876" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500B120EFAB1CE413DF1F934" italics="true" pageId="10" pageNumber="335">Epitornoceras peracutum</emphasis>
</taxonomicName>
(Hall, 1876;
<typeStatus id="54CD8822500B120EFBB9CE603AE5F934" box="[1071,1165,1650,1676]" pageId="10" pageNumber="335" type="holotype">holotype</typeStatus>
described by
<bibRefCitation id="EFE74B71500B120EFAB0CE603BC1F934" author="MILLER A. K." box="[1318,1449,1650,1676]" pageId="10" pageNumber="335" pagination="1 - 262" refId="ref11505" refString="MILLER A. K. 1938. - Devonian Ammonoids of America. Geological Society American Special Paper 14: 1 - 262. https: // doi. org / 10.1130 / SPE 14" type="journal article" year="1938">Miller 1938</bibRefCitation>
) and
<taxonomicName id="4C764D03500B120EFCCDCE813BC7F914" authority="Kirchgasser, Baird &amp; Brett, 2020" authorityName="Kirchgasser, Baird &amp; Brett" authorityYear="2020" box="[859,1455,1682,1709]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="dennisoni">
<emphasis id="B902EA92500B120EFCCDCE813A24F914" box="[859,1100,1682,1708]" italics="true" pageId="10" pageNumber="335">Epitornoceras dennisoni</emphasis>
<bibRefCitation id="EFE74B71500B120EFBC4CE803BC7F914" author="KIRCHGASSER W. T. &amp; BAIRD G. C. &amp; BRETT C. E." box="[1106,1455,1682,1709]" pageId="10" pageNumber="335" refId="ref10915" refString="KIRCHGASSER W. T., BAIRD G. C. &amp; BRETT C. E. 2020. - Ammonoids from the Tully Formation and Harrell Shale in Pennsylvania: markers of sea-level highstands in the Middle to Late Devonian transition, Appalachian foreland basin, eastern North America, in AVARY K. L., HASSON K. O. &amp; DIECCHIO R. J. (eds), The Appalachian Geology of John M. Dennison: Rocks, People, and a Few Good Restaurants along the Way. Vol. 545. Geological Society of America, Boulder. https: // doi. org / 10.1130 / 2020.2545 (10)" type="book" year="2020">Kirchgasser, Baird &amp; Brett, 2020</bibRefCitation>
</taxonomicName>
reported from North America (seeTable 1 for a comparison of the morphological characters of the different species).
<bibRefCitation id="EFE74B71500B120EFAFCCEC03D02F8B5" author="HOUSE M. R." pageId="10" pageNumber="335" pagination="79 - 130" refId="ref10725" refString="HOUSE M. R. 1965. - A study in the tornoceratidae: the succession of tornoceras and related genera in the North American devonian. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 250 (763): 79 - 130. https: // doi. org / 10.1098 / rstb. 1965.0021" type="journal article" year="1965">House (1965</bibRefCitation>
,
<bibRefCitation id="EFE74B71500B120EFCE3CEE03DC3F8B4" author="HOUSE M. R." box="[885,939,1778,1804]" pageId="10" pageNumber="335" pagination="1 - 70" refId="ref10787" refString="HOUSE M. R. 1978. - Devonian Ammonoids from the Appalachians and their bearing on international zonation and correlation. Special Papers in Palaeontology 21: 1 - 70." type="journal article" year="1978">1978</bibRefCitation>
)also identified forms showing affinities with
<taxonomicName id="4C764D03500B120EFAFECEE13D8EF893" baseAuthorityName="Hall" baseAuthorityYear="1876" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500B120EFAFECEE13D8EF893" italics="true" pageId="10" pageNumber="335">Epitornoceras peracutum</emphasis>
</taxonomicName>
(refigured in
<bibRefCitation id="EFE74B71500B120EFBEECF003BF3F894" author="HOUSE M. R. &amp; KIRCHGASSER W. T." box="[1144,1435,1810,1836]" pageId="10" pageNumber="335" pagination="1 - 288" refId="ref10819" refString="HOUSE M. R. &amp; KIRCHGASSER W. T. 2008. - Late Devonian Goniatites (Cephalopoda, Ammonoidea) from New York State. Bulletins of American Paleontology 374: 1 - 288." type="journal article" year="2008">House &amp; Kirchgasser 2008</bibRefCitation>
).
</paragraph>
<paragraph id="8BC93680500B120EFCD5CF203B48F854" blockId="10.[810,1457,1363,2028]" pageId="10" pageNumber="335">
As a general observation, the conch of
<taxonomicName id="4C764D03500B120EFB4CCF203B54F8F3" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[1242,1340,1841,1867]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500B120EFB4CCF203B54F8F3" box="[1242,1340,1841,1867]" italics="true" pageId="10" pageNumber="335">E. baldisi</emphasis>
</taxonomicName>
appears to be more slender (i.e., extremely discoidal, see
<tableCitation id="C6F4033B500B120EFA85CF433B0BF8D4" box="[1299,1379,1873,1900]" captionStart="TABLE" captionStartId="11.[134,143,220,237]" captionTargetPageId="11" captionText="TABLE 1. — Comparison of the morphological characteristics of the different species belonging to the genus Epitornoceras Frech,1902.Abbreviations:CWI, conch width index; UWI, umbilical width index; WER, whorl expansion rate; WWI, whorl width index." httpUri="http://table.plazi.org/id/DF096608500A120FFF10C8CE3DFBFEBB" pageId="10" pageNumber="335" tableUuid="DF096608500A120FFF10C8CE3DFBFEBB">Table 1</tableCitation>
) at the adult stage than in all other
<taxonomicName id="4C764D03500B120EFBDECF603B75F833" box="[1096,1309,1905,1931]" pageId="10" pageNumber="335">
<emphasis id="B902EA92500B120EFBDECF603AA7F833" box="[1096,1231,1906,1931]" italics="true" pageId="10" pageNumber="335">Epitornoceras</emphasis>
species
</taxonomicName>
(by excluding
<taxonomicName id="4C764D03500B120EFCBDCF803DD9F813" baseAuthorityName="Hall" baseAuthorityYear="1876" box="[811,945,1938,1963]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="10" pageNumber="335" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500B120EFCBDCF803DD9F813" box="[811,945,1938,1963]" italics="true" pageId="10" pageNumber="335">E. peracutum</emphasis>
</taxonomicName>
from the comparison, for which the conch shape is not well known; the description of this species is almost only based on its suture line, see
<bibRefCitation id="EFE74B71500B120EFB18CFC33B79F853" author="MILLER A. K." box="[1166,1297,2001,2027]" pageId="10" pageNumber="335" pagination="1 - 262" refId="ref11505" refString="MILLER A. K. 1938. - Devonian Ammonoids of America. Geological Society American Special Paper 14: 1 - 262. https: // doi. org / 10.1130 / SPE 14" type="journal article" year="1938">Miller 1938</bibRefCitation>
).
</paragraph>
<caption id="DF096608500A120FFF10C8CE3DFBFEBB" ID-Table-UUID="DF096608500A120FFF10C8CE3DFBFEBB" httpUri="http://table.plazi.org/id/DF096608500A120FFF10C8CE3DFBFEBB" pageId="11" pageNumber="336" startId="11.[134,143,220,237]" targetBox="[134,1448,317,911]" targetIsTable="true" targetPageId="11" targetType="table">
<paragraph id="8BC93680500A120FFF10C8CE3DFBFEBB" blockId="11.[132,1455,220,259]" pageId="11" pageNumber="336">
<smallCapsWord id="8D2FA05C500A120FFF10C8CE3EDAFF54" baselines="233,233" box="[134,178,220,237]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Table" pageId="11" pageNumber="336">TABLE</smallCapsWord>
1. — Comparison of the morphological characteristics of the different species belonging to the genus
<taxonomicName id="4C764D03500A120FFC67C8CE3AA4FF55" authority="Frech, 1902" authorityName="Frech" authorityYear="1902" box="[1009,1228,220,237]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFC67C8CE3A0CFF55" box="[1009,1124,220,237]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
<bibRefCitation id="EFE74B71500A120FFBFEC8CE3AA4FF55" author="FRECH F." box="[1128,1228,220,237]" pageId="11" pageNumber="336" pagination="27 - 112" refId="ref10386" refString="FRECH F. 1902. - Uber devonische Ammoneen. Beitrage zur Palaontologie Osterreich-Ungarns und des Orients. Wien and Leipzig 14: 27 - 112 (in German)." type="journal article" year="1902">Frech,1902</bibRefCitation>
</taxonomicName>
.Abbreviations:
<emphasis id="B902EA92500A120FFAD8C8CE3B1BFF55" bold="true" box="[1358,1395,220,237]" pageId="11" pageNumber="336">CWI</emphasis>
, conch width index;
<emphasis id="B902EA92500A120FFF79C8E03F7CFEBB" bold="true" box="[239,276,242,259]" pageId="11" pageNumber="336">UWI</emphasis>
, umbilical width index;
<emphasis id="B902EA92500A120FFE4EC8E03C6BFEBB" bold="true" box="[472,515,242,259]" pageId="11" pageNumber="336">WER</emphasis>
, whorl expansion rate;
<emphasis id="B902EA92500A120FFD50C8E03C87FEBB" bold="true" box="[710,751,242,259]" pageId="11" pageNumber="336">WWI</emphasis>
, whorl width index.
</paragraph>
</caption>
<paragraph id="8BC93680500A120FFF34C92D3B5DFC37" pageId="11" pageNumber="336">
<table id="F976C420500AEDFBFF10C92F3BC0FC37" box="[134,1448,317,911]" gridcols="6" gridrows="23" pageId="11" pageNumber="336">
<tr id="354634C2500AEDFBFF10C92F3BC0FEE9" box="[134,1448,317,337]" gridrow="0" pageId="11" pageNumber="336" rowspan-1="1" rowspan-2="1" rowspan-5="1">
<th id="76975DBE500AEDFBFF10C92F3F57FEE9" box="[134,319,317,337]" gridcol="0" gridrow="0" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFF34C92D3F5CFEE9" bold="true" box="[162,308,319,337]" pageId="11" pageNumber="336">Morphological</emphasis>
</th>
<th id="76975DBE500AEDFBFC9EC92F3D89FEE9" box="[776,993,317,337]" gridcol="3" gridrow="0" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFCA7C92F3DA8FEE9" authorityName="Frech" authorityYear="1902" box="[817,960,317,337]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFCA7C92F3DA8FEE9" bold="true" box="[817,960,317,337]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
</th>
<th id="76975DBE500AEDFBFC7DC92F3AADFEE9" box="[1003,1221,317,337]" gridcol="4" gridrow="0" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFB82C92F3ACBFEE9" authorityName="Frech" authorityYear="1902" box="[1044,1187,317,337]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFB82C92F3ACBFEE9" bold="true" box="[1044,1187,317,337]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="354634C2500AEDFBFF10C9453BC0FED3" box="[134,1448,343,363]" gridrow="1" pageId="11" pageNumber="336" rowspan-1="1">
<th id="76975DBE500AEDFBFF10C9453F57FED3" box="[134,319,343,363]" gridcol="0" gridrow="1" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFF3EC94B3F47FED3" bold="true" box="[168,303,345,363]" pageId="11" pageNumber="336">characters at</emphasis>
</th>
<td id="76975DBE500AEDFBFDBBC9453D6FFED3" box="[557,775,343,363]" gridcol="2" gridrow="1" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFDC4C9453C89FED3" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[594,737,343,363]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="335" phylum="Mollusca" rank="species" species="transmediterraneum">
<emphasis id="B902EA92500A120FFDC4C9453C89FED3" bold="true" box="[594,737,343,363]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
</td>
<td id="76975DBE500AEDFBFC9EC9453D89FED3" box="[776,993,343,363]" gridcol="3" gridrow="1" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFC86C9453D89FED3" bold="true" box="[784,993,343,363]" italics="true" pageId="11" pageNumber="336">transmediterraneum</emphasis>
</td>
<td id="76975DBE500AEDFBFC7DC9453AADFED3" box="[1003,1221,343,363]" gridcol="4" gridrow="1" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFBBCC9453AE5FED3" authorityName="Kirchgasser, Baird &amp; Brett" authorityYear="2020" box="[1066,1165,343,363]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="dennisoni">
<emphasis id="B902EA92500A120FFBBCC9453AE5FED3" bold="true" box="[1066,1165,343,363]" italics="true" pageId="11" pageNumber="336">dennisoni</emphasis>
</taxonomicName>
</td>
<td id="76975DBE500AEDFBFB58C9453BC0FED3" box="[1230,1448,343,363]" gridcol="5" gridrow="1" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFB61C9453BEEFED3" authorityName="Frech" authorityYear="1902" box="[1271,1414,343,363]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFB61C9453BEEFED3" bold="true" box="[1271,1414,343,363]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="354634C2500AEDFBFF10C9633BC0FE3D" box="[134,1448,369,389]" gridrow="2" pageId="11" pageNumber="336">
<th id="76975DBE500AEDFBFF10C9633F57FE3D" box="[134,319,369,389]" gridcol="0" gridrow="2" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFF09C9613F5EFE3C" bold="true" box="[159,310,371,389]" pageId="11" pageNumber="336">the adult stage</emphasis>
</th>
<td id="76975DBE500AEDFBFEC4C9633C44FE3D" box="[338,556,369,389]" gridcol="1" gridrow="2" pageId="11" pageNumber="336">Epitornoceras baldisi</td>
<td id="76975DBE500AEDFBFDBBC9633D6FFE3D" box="[557,775,369,389]" gridcol="2" gridrow="2" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFDCEC9633CB5FE3D" box="[600,733,369,389]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="subSpecies" species="baldisi" subSpecies="mithracoides">
<emphasis id="B902EA92500A120FFDCEC9633CB5FE3D" bold="true" box="[600,733,369,389]" italics="true" pageId="11" pageNumber="336">mithracoides</emphasis>
</taxonomicName>
</td>
<td id="76975DBE500AEDFBFC9EC9633D89FE3D" box="[776,993,369,389]" gridcol="3" gridrow="2" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFC83C9613DB5FE3D" bold="true" box="[789,989,371,389]" pageId="11" pageNumber="336">Korn &amp; Bockwinkel,</emphasis>
</td>
<td id="76975DBE500AEDFBFC7DC9633AADFE3D" box="[1003,1221,369,389]" gridcol="4" gridrow="2" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFB8DC9613AF4FE3D" bold="true" box="[1051,1180,371,389]" pageId="11" pageNumber="336">Kirchgasser,</emphasis>
</td>
<td id="76975DBE500AEDFBFB58C9633BC0FE3D" box="[1230,1448,369,389]" gridcol="5" gridrow="2" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFA91C9633B1EFE3D" baseAuthorityName="Hall" baseAuthorityYear="1876" box="[1287,1398,369,389]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500A120FFA91C9633B1EFE3D" bold="true" box="[1287,1398,369,389]" italics="true" pageId="11" pageNumber="336">peracutum</emphasis>
</taxonomicName>
</td>
</tr>
<tr id="354634C2500AEDFBFF10C9983BC0FE26" box="[134,1448,394,414]" gridrow="3" pageId="11" pageNumber="336">
<th id="76975DBE500AEDFBFF10C9983F57FE26" box="[134,319,394,414]" gridcol="0" gridrow="3" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFF0EC99E3F57FE26" bold="true" box="[152,319,394,414]" pageId="11" pageNumber="336">
(
<emphasis id="B902EA92500A120FFF08C9983EC6FE26" bold="true" box="[158,174,394,414]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
30-50 mm dm)
</emphasis>
</th>
<td id="76975DBE500AEDFBFEC4C9983C44FE26" box="[338,556,394,414]" gridcol="1" gridrow="3" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFEE0C99E3C6FFE26" bold="true" box="[374,519,396,414]" pageId="11" pageNumber="336">
(
<bibRefCitation id="EFE74B71500A120FFEEDC99E3C6BFE26" author="LEANZA A. F." box="[379,515,396,414]" pageId="11" pageNumber="336" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza, 1968</bibRefCitation>
)
</emphasis>
</td>
<td id="76975DBE500AEDFBFDBBC9983D6FFE26" box="[557,775,394,414]" gridcol="2" gridrow="3" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFDCFC99E3CB3FE26" bold="true" box="[601,731,396,414]" pageId="11" pageNumber="336">(Frech, 1888)</emphasis>
</td>
<td id="76975DBE500AEDFBFC9EC9983D89FE26" box="[776,993,394,414]" gridcol="3" gridrow="3" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFCF7C99E3DF8FE26" bold="true" box="[865,912,396,414]" pageId="11" pageNumber="336">2022</emphasis>
</td>
<td id="76975DBE500AEDFBFC7DC9983AADFE26" box="[1003,1221,394,414]" gridcol="4" gridrow="3" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFC6AC99E3AD3FE26" bold="true" box="[1020,1211,396,414]" pageId="11" pageNumber="336">Baird &amp; Brett, 2020</emphasis>
</td>
<td id="76975DBE500AEDFBFB58C9983BC0FE26" box="[1230,1448,394,414]" gridcol="5" gridrow="3" pageId="11" pageNumber="336">
<emphasis id="B902EA92500A120FFA91C99E3B1EFE26" bold="true" box="[1287,1398,396,414]" pageId="11" pageNumber="336">(Hall, 1876)</emphasis>
</td>
</tr>
<tr id="354634C2500AEDFBFF10C9BC3BC0FE7A" box="[134,1448,430,450]" gridrow="4" pageId="11" pageNumber="336">
<th id="76975DBE500AEDFBFF10C9BC3F57FE7A" box="[134,319,430,450]" gridcol="0" gridrow="4" pageId="11" pageNumber="336">Conch geometry</th>
<td id="76975DBE500AEDFBFEC4C9BC3C44FE7A" box="[338,556,430,450]" gridcol="1" gridrow="4" pageId="11" pageNumber="336">Extremely discoidal</td>
<td id="76975DBE500AEDFBFDBBC9BC3D6FFE7A" box="[557,775,430,450]" gridcol="2" gridrow="4" pageId="11" pageNumber="336">Thinly discoidal</td>
<td id="76975DBE500AEDFBFC9EC9BC3D89FE7A" box="[776,993,430,450]" gridcol="3" gridrow="4" pageId="11" pageNumber="336">Thinly discoidal</td>
<td id="76975DBE500AEDFBFC7DC9BC3AADFE7A" box="[1003,1221,430,450]" gridcol="4" gridrow="4" pageId="11" pageNumber="336">Thinly discoidal</td>
<td id="76975DBE500AEDFBFB58C9BC3BC0FE7A" box="[1230,1448,430,450]" gridcol="5" gridrow="4" pageId="11" pageNumber="336">Conch probably</td>
</tr>
<tr id="354634C2500AEDFBFF10C9D53BC0FE63" box="[134,1448,455,475]" gridrow="5" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4C9D53C44FE63" box="[338,556,455,475]" gridcol="1" gridrow="5" pageId="11" pageNumber="336">
(CWI
<emphasis id="B902EA92500A120FFE03C9D53FCFFE63" box="[405,423,455,475]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.25)
</td>
<td id="76975DBE500AEDFBFDBBC9D53D6FFE63" box="[557,775,455,475]" gridcol="2" gridrow="5" pageId="11" pageNumber="336">
(CWI
<emphasis id="B902EA92500A120FFDE6C9D53CEAFE63" box="[624,642,455,475]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.35)
</td>
<td id="76975DBE500AEDFBFC9EC9D53D89FE63" box="[776,993,455,475]" gridcol="3" gridrow="5" pageId="11" pageNumber="336">
(CWI
<emphasis id="B902EA92500A120FFCDAC9D53D35FE63" box="[844,861,455,475]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.35)
</td>
<td id="76975DBE500AEDFBFC7DC9D53AADFE63" box="[1003,1221,455,475]" gridcol="4" gridrow="5" pageId="11" pageNumber="336">
(CWI
<emphasis id="B902EA92500A120FFBB8C9D53A28FE63" box="[1070,1088,455,475]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.35)
</td>
<td id="76975DBE500AEDFBFB58C9D53BC0FE63" box="[1230,1448,455,475]" gridcol="5" gridrow="5" pageId="11" pageNumber="336">discoidal</td>
</tr>
<tr id="354634C2500AEDFBFF10C9CD3BC0FE4B" box="[134,1448,479,499]" gridrow="6" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4C9CD3C44FE4B" box="[338,556,479,499]" gridcol="1" gridrow="6" pageId="11" pageNumber="336">Whorl profile strongly</td>
<td id="76975DBE500AEDFBFDBBC9CD3D6FFE4B" box="[557,775,479,499]" gridcol="2" gridrow="6" pageId="11" pageNumber="336">Whorl profile</td>
<td id="76975DBE500AEDFBFC9EC9CD3D89FE4B" box="[776,993,479,499]" gridcol="3" gridrow="6" pageId="11" pageNumber="336">Whorl profile</td>
<td id="76975DBE500AEDFBFC7DC9CD3AADFE4B" box="[1003,1221,479,499]" gridcol="4" gridrow="6" pageId="11" pageNumber="336">Whorl profile</td>
<td id="76975DBE500AEDFBFB58C9CD3BC0FE4B" box="[1230,1448,479,499]" gridcol="5" gridrow="6" pageId="11" pageNumber="336">Whorl profile shape</td>
</tr>
<tr id="354634C2500AEDFBFF10C9EB3BC0FDB5" box="[134,1448,505,525]" gridrow="7" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4C9EB3C44FDB5" box="[338,556,505,525]" gridcol="1" gridrow="7" pageId="11" pageNumber="336">compressed</td>
<td id="76975DBE500AEDFBFDBBC9EB3D6FFDB5" box="[557,775,505,525]" gridcol="2" gridrow="7" pageId="11" pageNumber="336">compressed</td>
<td id="76975DBE500AEDFBFC9EC9EB3D89FDB5" box="[776,993,505,525]" gridcol="3" gridrow="7" pageId="11" pageNumber="336">compressed</td>
<td id="76975DBE500AEDFBFC7DC9EB3AADFDB5" box="[1003,1221,505,525]" gridcol="4" gridrow="7" pageId="11" pageNumber="336">compressed</td>
<td id="76975DBE500AEDFBFB58C9EB3BC0FDB5" box="[1230,1448,505,525]" gridcol="5" gridrow="7" pageId="11" pageNumber="336">unknown</td>
</tr>
<tr id="354634C2500AEDFBFF10CA013BC0FD9F" box="[134,1448,531,551]" gridrow="8" pageId="11" pageNumber="336" rowspan-0="1" rowspan-5="1">
<td id="76975DBE500AEDFBFEC4CA013C44FD9F" box="[338,556,531,551]" gridcol="1" gridrow="8" pageId="11" pageNumber="336">
(WWI
<emphasis id="B902EA92500A120FFE0FCA013FC3FD9F" box="[409,427,531,551]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.40)
</td>
<td id="76975DBE500AEDFBFDBBCA013D6FFD9F" box="[557,775,531,551]" gridcol="2" gridrow="8" pageId="11" pageNumber="336">
(WWI
<emphasis id="B902EA92500A120FFDE3CA013CEEFD9F" box="[629,646,531,551]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.65)
</td>
<td id="76975DBE500AEDFBFC9ECA013D89FD9F" box="[776,993,531,551]" gridcol="3" gridrow="8" pageId="11" pageNumber="336">
(WWI
<emphasis id="B902EA92500A120FFCC6CA013D0AFD9F" box="[848,866,531,551]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.65)
</td>
<td id="76975DBE500AEDFBFC7DCA013AADFD9F" box="[1003,1221,531,551]" gridcol="4" gridrow="8" pageId="11" pageNumber="336">
(WWI
<emphasis id="B902EA92500A120FFBA5CA013A2CFD9F" box="[1075,1092,531,551]" italics="true" pageId="11" pageNumber="336">c.</emphasis>
0.65)
</td>
</tr>
<tr id="354634C2500AEDFBFF10CA393BC0FD87" box="[134,1448,555,575]" gridrow="9" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CA393C44FD87" box="[338,556,555,575]" gridcol="1" gridrow="9" pageId="11" pageNumber="336">Umbilicus closed at</td>
<td id="76975DBE500AEDFBFDBBCA393D6FFD87" box="[557,775,555,575]" gridcol="2" gridrow="9" pageId="11" pageNumber="336">Umbilicus closed at</td>
<td id="76975DBE500AEDFBFC9ECA393D89FD87" box="[776,993,555,575]" gridcol="3" gridrow="9" pageId="11" pageNumber="336">Umbilicus closed at</td>
<td id="76975DBE500AEDFBFC7DCA393AADFD87" box="[1003,1221,555,575]" gridcol="4" gridrow="9" pageId="11" pageNumber="336">Diameter of umbilical</td>
<td id="76975DBE500AEDFBFB58CA393BC0FD87" box="[1230,1448,555,575]" gridcol="5" gridrow="9" pageId="11" pageNumber="336">Diameter of umbilical</td>
</tr>
<tr id="354634C2500AEDFBFF10CA563BC0FDE0" box="[134,1448,580,600]" gridrow="10" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CA563C44FDE0" box="[338,556,580,600]" gridcol="1" gridrow="10" pageId="11" pageNumber="336">30 mm dm</td>
<td id="76975DBE500AEDFBFDBBCA563D6FFDE0" box="[557,775,580,600]" gridcol="2" gridrow="10" pageId="11" pageNumber="336">50 mm dm</td>
<td id="76975DBE500AEDFBFC9ECA563D89FDE0" box="[776,993,580,600]" gridcol="3" gridrow="10" pageId="11" pageNumber="336">50 mm dm</td>
<td id="76975DBE500AEDFBFC7DCA563AADFDE0" box="[1003,1221,580,600]" gridcol="4" gridrow="10" pageId="11" pageNumber="336">closure unknown</td>
<td id="76975DBE500AEDFBFB58CA563BC0FDE0" box="[1230,1448,580,600]" gridcol="5" gridrow="10" pageId="11" pageNumber="336">closure unknown</td>
</tr>
<tr id="354634C2500AEDFBFF10CA4E3BC0FDC8" box="[134,1448,604,624]" gridrow="11" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CA4E3C44FDC8" box="[338,556,604,624]" gridcol="1" gridrow="11" pageId="11" pageNumber="336">Umbilical margin</td>
<td id="76975DBE500AEDFBFDBBCA4E3D6FFDC8" box="[557,775,604,624]" gridcol="2" gridrow="11" pageId="11" pageNumber="336">Umbilical margin</td>
<td id="76975DBE500AEDFBFC9ECA4E3D89FDC8" box="[776,993,604,624]" gridcol="3" gridrow="11" pageId="11" pageNumber="336">Umbilical margin</td>
<td id="76975DBE500AEDFBFC7DCA4E3AADFDC8" box="[1003,1221,604,624]" gridcol="4" gridrow="11" pageId="11" pageNumber="336">Umbilical margin</td>
<td id="76975DBE500AEDFBFB58CA4E3BC0FDC8" box="[1230,1448,604,624]" gridcol="5" gridrow="11" pageId="11" pageNumber="336">Umbilical margin</td>
</tr>
<tr id="354634C2500AEDFBFF10CA643BC0FD32" box="[134,1448,630,650]" gridrow="12" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CA643C44FD32" box="[338,556,630,650]" gridcol="1" gridrow="12" pageId="11" pageNumber="336">continuously rounded</td>
<td id="76975DBE500AEDFBFDBBCA643D6FFD32" box="[557,775,630,650]" gridcol="2" gridrow="12" pageId="11" pageNumber="336">continuously rounded</td>
<td id="76975DBE500AEDFBFC9ECA643D89FD32" box="[776,993,630,650]" gridcol="3" gridrow="12" pageId="11" pageNumber="336">obliquely flattened</td>
<td id="76975DBE500AEDFBFC7DCA643AADFD32" box="[1003,1221,630,650]" gridcol="4" gridrow="12" pageId="11" pageNumber="336">continuously rounded</td>
<td id="76975DBE500AEDFBFB58CA643BC0FD32" box="[1230,1448,630,650]" gridcol="5" gridrow="12" pageId="11" pageNumber="336">continuously rounded</td>
</tr>
<tr id="354634C2500AEDFBFF10CA823BC0FD1C" box="[134,1448,656,676]" gridrow="13" pageId="11" pageNumber="336" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-4="1" rowspan-5="1">
<td id="76975DBE500AEDFBFC9ECA823D89FD1C" box="[776,993,656,676]" gridcol="3" gridrow="13" pageId="11" pageNumber="336">(funnel-shaped)</td>
</tr>
<tr id="354634C2500AEDFBFF10CABA3BC0FD04" box="[134,1448,680,700]" gridrow="14" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CABA3D89FD04" box="[338,993,680,700]" colspan="3" colspanRight="2" gridcol="1" gridrow="14" pageId="11" pageNumber="336">Flanks weakly convex, Flanks weakly convex, Flanks nearly parallel</td>
<td id="76975DBE500AEDFBFC7DCABA3AADFD04" box="[1003,1221,680,700]" gridcol="4" gridrow="14" pageId="11" pageNumber="336">Flanks weakly convex,</td>
<td id="76975DBE500AEDFBFB58CABA3BC0FD04" box="[1230,1448,680,700]" gridcol="5" gridrow="14" pageId="11" pageNumber="336">Flanks weakly convex</td>
</tr>
<tr id="354634C2500AEDFBFF10CAD33BC0FD6D" box="[134,1448,705,725]" gridrow="15" pageId="11" pageNumber="336" rowspan-0="1" rowspan-3="1" rowspan-5="1">
<td id="76975DBE500AEDFBFEC4CAD33C44FD6D" box="[338,556,705,725]" gridcol="1" gridrow="15" pageId="11" pageNumber="336">convergent</td>
<td id="76975DBE500AEDFBFDBBCAD33D6FFD6D" box="[557,775,705,725]" gridcol="2" gridrow="15" pageId="11" pageNumber="336">convergent</td>
<td id="76975DBE500AEDFBFC7DCAD33AADFD6D" box="[1003,1221,705,725]" gridcol="4" gridrow="15" pageId="11" pageNumber="336">convergent</td>
</tr>
<tr id="354634C2500AEDFBFF10CAF73BC0FD41" box="[134,1448,741,761]" gridrow="16" pageId="11" pageNumber="336">
<th id="76975DBE500AEDFBFF10CAF73F57FD41" box="[134,319,741,761]" gridcol="0" gridrow="16" pageId="11" pageNumber="336">Suture line</th>
<td id="76975DBE500AEDFBFEC4CAF73C44FD41" box="[338,556,741,761]" gridcol="1" gridrow="16" pageId="11" pageNumber="336">Narrow E lobe</td>
<td id="76975DBE500AEDFBFDBBCAF73D6FFD41" box="[557,775,741,761]" gridcol="2" gridrow="16" pageId="11" pageNumber="336">V-shaped E lobe</td>
<td id="76975DBE500AEDFBFC9ECAF73D89FD41" box="[776,993,741,761]" gridcol="3" gridrow="16" pageId="11" pageNumber="336">V-shaped E lobe</td>
<td id="76975DBE500AEDFBFC7DCAF73AADFD41" box="[1003,1221,741,761]" gridcol="4" gridrow="16" pageId="11" pageNumber="336">Wide E lobe with</td>
<td id="76975DBE500AEDFBFB58CAF73BC0FD41" box="[1230,1448,741,761]" gridcol="5" gridrow="16" pageId="11" pageNumber="336">Wide E lobe with</td>
</tr>
<tr id="354634C2500AEDFBFF10CAEC3BC0FCAA" box="[134,1448,766,786]" gridrow="17" pageId="11" pageNumber="336" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="76975DBE500AEDFBFC7DCAEC3AADFCAA" box="[1003,1221,766,786]" gridcol="4" gridrow="17" pageId="11" pageNumber="336">convex flanks</td>
<td id="76975DBE500AEDFBFB58CAEC3BC0FCAA" box="[1230,1448,766,786]" gridcol="5" gridrow="17" pageId="11" pageNumber="336">concave flanks</td>
</tr>
<tr id="354634C2500AEDFBFF10CB043BC0FC92" box="[134,1448,790,810]" gridrow="18" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CB043C44FC92" box="[338,556,790,810]" gridcol="1" gridrow="18" pageId="11" pageNumber="336">Nearly symmetric</td>
<td id="76975DBE500AEDFBFDBBCB043D6FFC92" box="[557,775,790,810]" gridcol="2" gridrow="18" pageId="11" pageNumber="336">Asymmetric A lobe,</td>
<td id="76975DBE500AEDFBFC9ECB043D89FC92" box="[776,993,790,810]" gridcol="3" gridrow="18" pageId="11" pageNumber="336">Nearly symmetric</td>
<td id="76975DBE500AEDFBFC7DCB043AADFC92" box="[1003,1221,790,810]" gridcol="4" gridrow="18" pageId="11" pageNumber="336">Asymmetric A lobe,</td>
<td id="76975DBE500AEDFBFB58CB043BC0FC92" box="[1230,1448,790,810]" gridcol="5" gridrow="18" pageId="11" pageNumber="336">Asymmetric A lobe,</td>
</tr>
<tr id="354634C2500AEDFBFF10CB223BC0FCFC" box="[134,1448,816,836]" gridrow="19" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CB223C44FCFC" box="[338,556,816,836]" gridcol="1" gridrow="19" pageId="11" pageNumber="336">A lobe, wider than</td>
<td id="76975DBE500AEDFBFDBBCB223D6FFCFC" box="[557,775,816,836]" gridcol="2" gridrow="19" pageId="11" pageNumber="336">as deep as wide,</td>
<td id="76975DBE500AEDFBFC9ECB223D89FCFC" box="[776,993,816,836]" gridcol="3" gridrow="19" pageId="11" pageNumber="336">A lobe, deeper than</td>
<td id="76975DBE500AEDFBFC7DCB223AADFCFC" box="[1003,1221,816,836]" gridcol="4" gridrow="19" pageId="11" pageNumber="336">slightly wider than</td>
<td id="76975DBE500AEDFBFB58CB223BC0FCFC" box="[1230,1448,816,836]" gridcol="5" gridrow="19" pageId="11" pageNumber="336">very deep, deeper</td>
</tr>
<tr id="354634C2500AEDFBFF10CB583BC0FCE6" box="[134,1448,842,862]" gridrow="20" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CB583C44FCE6" box="[338,556,842,862]" gridcol="1" gridrow="20" pageId="11" pageNumber="336">deep</td>
<td id="76975DBE500AEDFBFDBBCB583D6FFCE6" box="[557,775,842,862]" gridcol="2" gridrow="20" pageId="11" pageNumber="336">convergent flanks</td>
<td id="76975DBE500AEDFBFC9ECB583D89FCE6" box="[776,993,842,862]" gridcol="3" gridrow="20" pageId="11" pageNumber="336">wide, parallel flanks</td>
<td id="76975DBE500AEDFBFC7DCB583AADFCE6" box="[1003,1221,842,862]" gridcol="4" gridrow="20" pageId="11" pageNumber="336">deep</td>
<td id="76975DBE500AEDFBFB58CB583BC0FCE6" box="[1230,1448,842,862]" gridcol="5" gridrow="20" pageId="11" pageNumber="336">than wide</td>
</tr>
<tr id="354634C2500AEDFBFF10CB703BC0FCCE" box="[134,1448,866,886]" gridrow="21" pageId="11" pageNumber="336" rowspan-0="1">
<td id="76975DBE500AEDFBFEC4CB703C44FCCE" box="[338,556,866,886]" gridcol="1" gridrow="21" pageId="11" pageNumber="336"></td>
<td id="76975DBE500AEDFBFDBBCB703D6FFCCE" box="[557,775,866,886]" gridcol="2" gridrow="21" pageId="11" pageNumber="336"></td>
<td id="76975DBE500AEDFBFC9ECB703D89FCCE" box="[776,993,866,886]" gridcol="3" gridrow="21" pageId="11" pageNumber="336"></td>
<td id="76975DBE500AEDFBFC7DCB703AADFCCE" box="[1003,1221,866,886]" gridcol="4" gridrow="21" pageId="11" pageNumber="336">Flat-topped</td>
<td id="76975DBE500AEDFBFB58CB703BC0FCCE" box="[1230,1448,866,886]" gridcol="5" gridrow="21" pageId="11" pageNumber="336">Dorsolateral saddle</td>
</tr>
<tr id="354634C2500AEDFBFF10CB693BC0FC37" box="[134,1448,891,911]" gridrow="22" pageId="11" pageNumber="336" rowspan-0="1" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<td id="76975DBE500AEDFBFC7DCB693AADFC37" box="[1003,1221,891,911]" gridcol="4" gridrow="22" pageId="11" pageNumber="336">dorsolateral saddle</td>
<td id="76975DBE500AEDFBFB58CB693BC0FC37" box="[1230,1448,891,911]" gridcol="5" gridrow="22" pageId="11" pageNumber="336">less wide</td>
</tr>
</table>
</paragraph>
<paragraph id="8BC93680500A120FFF0DCBC73F21FA35" blockId="11.[130,777,980,2027]" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFF0DCBC73F19FC56" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[155,369,980,1006]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFF0DCBC73F19FC56" box="[155,369,980,1006]" italics="true" pageId="11" pageNumber="336">Epitornoceras baldisi</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C764D03500A120FFE6ACBC73C82FC56" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[508,746,980,1006]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="transmediterraneum">
<emphasis id="B902EA92500A120FFE6ACBC73C82FC56" box="[508,746,980,1006]" italics="true" pageId="11" pageNumber="336">E. transmediterraneum</emphasis>
</taxonomicName>
in the shape of the umbilicus: the umbilical wall is obliquely flattened caused by an umbilical shell thickening in
<taxonomicName id="4C764D03500A120FFD38CC073F42FBF6" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="transmediterraneum">
<emphasis id="B902EA92500A120FFD38CC073F42FBF6" italics="true" pageId="11" pageNumber="336">E. transmediterraneum</emphasis>
</taxonomicName>
(i.e., funnel-shaped umbilicus), while in
<taxonomicName id="4C764D03500A120FFF10CC473E82FBD6" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[134,234,1108,1134]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFF10CC473E82FBD6" box="[134,234,1108,1134]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
the umbilical margin is continuously rounded (as in all others species, see
<tableCitation id="C6F4033B500A120FFE11CC663FBFFB36" box="[391,471,1140,1166]" captionStart="TABLE" captionStartId="11.[134,143,220,237]" captionTargetPageId="11" captionText="TABLE 1. — Comparison of the morphological characteristics of the different species belonging to the genus Epitornoceras Frech,1902.Abbreviations:CWI, conch width index; UWI, umbilical width index; WER, whorl expansion rate; WWI, whorl width index." httpUri="http://table.plazi.org/id/DF096608500A120FFF10C8CE3DFBFEBB" pageId="11" pageNumber="336" tableUuid="DF096608500A120FFF10C8CE3DFBFEBB">Table 1</tableCitation>
). The two species also differ when comparing the suture line. The suture line of
<taxonomicName id="4C764D03500A120FFD32CC873D6EFB16" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[676,774,1172,1198]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFD32CC873D6EFB16" box="[676,774,1172,1198]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
possesses a narrower external lobe, in
<taxonomicName id="4C764D03500A120FFD8FCCA73D6FFB76" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[537,775,1204,1230]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="transmediterraneum">
<emphasis id="B902EA92500A120FFD8FCCA73D6FFB76" box="[537,775,1204,1230]" italics="true" pageId="11" pageNumber="336">E. transmediterraneum</emphasis>
</taxonomicName>
the E lobe is V-shaped and wider. Furthermore, the A lobe is considerably deeper in
<taxonomicName id="4C764D03500A120FFE3FCCE63CC8FAB5" authorityName="Korn &amp; Bockwinkel" authorityYear="2022" box="[425,672,1267,1293]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="transmediterraneum">
<emphasis id="B902EA92500A120FFE3FCCE63CC8FAB5" box="[425,672,1267,1293]" italics="true" pageId="11" pageNumber="336">E. transmediterraneum</emphasis>
</taxonomicName>
, and the flanks are nearly parallel (
<bibRefCitation id="EFE74B71500A120FFE00CD063CD9FA96" author="KORN D. &amp; BOCKWINKEL J." box="[406,689,1299,1326]" pageId="11" pageNumber="336" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
). In the latter, the A lobe is deeper than wide (
<bibRefCitation id="EFE74B71500A120FFDBECD263ED6FAD5" author="KORN D. &amp; BOCKWINKEL J." pageId="11" pageNumber="336" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
), while in
<taxonomicName id="4C764D03500A120FFEAFCD463FF6FAD5" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[313,414,1363,1389]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFEAFCD463FF6FAD5" box="[313,414,1363,1389]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
the A lobe is slightly wider than deep (
<figureCitation id="134D2A05500A120FFF51CD613F55FA35" box="[199,317,1395,1422]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="11" pageNumber="336">Fig. 7D, E</figureCitation>
).
</paragraph>
<paragraph id="8BC93680500A120FFF0DCD863F3BF9D4" blockId="11.[130,777,980,2027]" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFF0DCD863F19FA15" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[155,369,1427,1453]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFF0DCD863F19FA15" box="[155,369,1427,1453]" italics="true" pageId="11" pageNumber="336">Epitornoceras baldisi</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D03500A120FFE30CD863C2DFA15" ID-CoL="6G8RQ" baseAuthorityName="Frech" baseAuthorityYear="1888" box="[422,581,1427,1453]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="mithracoides">
<emphasis id="B902EA92500A120FFE30CD863C2DFA15" box="[422,581,1427,1453]" italics="true" pageId="11" pageNumber="336">E. mithracoides</emphasis>
</taxonomicName>
are close in terms of conch shape (seeTable 1). However, differences can be seen in the suture line: in
<taxonomicName id="4C764D03500A120FFECDCDC63F90FA55" ID-CoL="6G8RQ" baseAuthorityName="Frech" baseAuthorityYear="1888" box="[347,504,1491,1517]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="mithracoides">
<emphasis id="B902EA92500A120FFECDCDC63F90FA55" box="[347,504,1491,1517]" italics="true" pageId="11" pageNumber="336">E. mithracoides</emphasis>
</taxonomicName>
, the A lobe is asymmetric and as wide as it is deep (
<bibRefCitation id="EFE74B71500A120FFE0FCDE13CDEF9B5" author="KORN D. &amp; BOCKWINKEL J." box="[409,694,1523,1549]" pageId="11" pageNumber="336" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
), while in
<taxonomicName id="4C764D03500A120FFF37CE013F6BF994" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[161,259,1554,1580]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFF37CE013F6BF994" box="[161,259,1554,1580]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
, the A lobe is nearly symmetric (and wider than deep), furthermore the L saddle is deeper and considerably wider (
<figureCitation id="134D2A05500A120FFF47CE403F2EF9D4" box="[209,326,1618,1645]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="11" pageNumber="336">Fig. 7D, E</figureCitation>
).
</paragraph>
<paragraph id="8BC93680500A120FFF0DCE613FF2F834" blockId="11.[130,777,980,2027]" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFF0DCE613F18F934" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[155,368,1650,1676]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFF0DCE613F18F934" box="[155,368,1650,1676]" italics="true" pageId="11" pageNumber="336">Epitornoceras baldisi</emphasis>
</taxonomicName>
mainly differs from the North American species (
<taxonomicName id="4C764D03500A120FFE85CE813FCDF914" ID-CoL="6G8S2" baseAuthorityName="Hall" baseAuthorityYear="1876" box="[275,421,1683,1708]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500A120FFE85CE813FCDF914" box="[275,421,1683,1708]" italics="true" pageId="11" pageNumber="336">E. peracutum</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D03500A120FFE74CE813C04F914" authorityName="Kirchgasser, Baird &amp; Brett" authorityYear="2020" box="[482,620,1682,1708]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="dennisoni">
<emphasis id="B902EA92500A120FFE74CE813C04F914" box="[482,620,1682,1708]" italics="true" pageId="11" pageNumber="336">E. dennisoni</emphasis>
</taxonomicName>
) in its suture line (see
<tableCitation id="C6F4033B500A120FFF79CEA03F2CF974" box="[239,324,1714,1740]" captionStart="TABLE" captionStartId="11.[134,143,220,237]" captionTargetPageId="11" captionText="TABLE 1. — Comparison of the morphological characteristics of the different species belonging to the genus Epitornoceras Frech,1902.Abbreviations:CWI, conch width index; UWI, umbilical width index; WER, whorl expansion rate; WWI, whorl width index." httpUri="http://table.plazi.org/id/DF096608500A120FFF10C8CE3DFBFEBB" pageId="11" pageNumber="336" tableUuid="DF096608500A120FFF10C8CE3DFBFEBB">Table 1</tableCitation>
).
<taxonomicName id="4C764D03500A120FFEC9CEA13C19F974" ID-CoL="6G8S2" baseAuthorityName="Hall" baseAuthorityYear="1876" box="[351,625,1715,1740]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500A120FFEC9CEA13C19F974" box="[351,625,1715,1740]" italics="true" pageId="11" pageNumber="336">Epitornoceras peracutum</emphasis>
</taxonomicName>
possesses an E lobe much wider (with concave flanks), and an A lobe much deeper and more asymmetric, furthermore, the dorsolateral saddle is comparatively less wide (
<bibRefCitation id="EFE74B71500A120FFDF8CF003C90F894" author="MILLER A. K." box="[622,760,1810,1836]" pageId="11" pageNumber="336" pagination="1 - 262" refId="ref11505" refString="MILLER A. K. 1938. - Devonian Ammonoids of America. Geological Society American Special Paper 14: 1 - 262. https: // doi. org / 10.1130 / SPE 14" type="journal article" year="1938">Miller 1938</bibRefCitation>
).
<taxonomicName id="4C764D03500A120FFF12CF203F15F8F3" authorityName="Kirchgasser, Baird &amp; Brett" authorityYear="2020" box="[132,381,1841,1867]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="dennisoni">
<emphasis id="B902EA92500A120FFF12CF203F15F8F3" box="[132,381,1841,1867]" italics="true" pageId="11" pageNumber="336">Epitornoceras dennisoni</emphasis>
</taxonomicName>
also possesses an E lobe much wider (with convex flanks), and a flat-topped dorsolateral saddle (
<bibRefCitation id="EFE74B71500A120FFF18CF633FE3F833" author="KIRCHGASSER W. T. &amp; BAIRD G. C. &amp; BRETT C. E." box="[142,395,1905,1931]" pageId="11" pageNumber="336" refId="ref10915" refString="KIRCHGASSER W. T., BAIRD G. C. &amp; BRETT C. E. 2020. - Ammonoids from the Tully Formation and Harrell Shale in Pennsylvania: markers of sea-level highstands in the Middle to Late Devonian transition, Appalachian foreland basin, eastern North America, in AVARY K. L., HASSON K. O. &amp; DIECCHIO R. J. (eds), The Appalachian Geology of John M. Dennison: Rocks, People, and a Few Good Restaurants along the Way. Vol. 545. Geological Society of America, Boulder. https: // doi. org / 10.1130 / 2020.2545 (10)" type="book" year="2020">
Kirchgasser
<emphasis id="B902EA92500A120FFE82CF603F21F833" box="[276,329,1905,1931]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
2020
</bibRefCitation>
).
</paragraph>
<paragraph id="8BC93680500A120FFF0DCF833A36FA15" blockId="11.[130,777,980,2027]" lastBlockId="11.[811,1457,980,2028]" pageId="11" pageNumber="336">
The type material of
<taxonomicName id="4C764D03500A120FFE1ECF803F87F813" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[392,495,1937,1963]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFE1ECF803F87F813" box="[392,495,1937,1963]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
was originally attributed to the genus
<taxonomicName id="4C764D03500A120FFE9BCFA03F15F873" ID-CoL="63WJJ" authorityName="Hyatt" authorityYear="1884" box="[269,381,1970,1995]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFE9BCFA03F15F873" box="[269,381,1970,1995]" italics="true" pageId="11" pageNumber="336">Tornoceras</emphasis>
</taxonomicName>
.
<specimenCount id="9D70FD09500A120FFE1CCFA03C58F873" box="[394,560,1969,1996]" count="2" pageId="11" pageNumber="336" type="generic">Two specimens</specimenCount>
were described: the
<typeStatus id="54CD8822500A120FFF15CFC33E8DF853" box="[131,229,2001,2027]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
CPUBA 8088 and CPUBA 8089, re-illustrated here inFig. 6. The conch of the
<typeStatus id="54CD8822500A120FFB14CBC63A8BFC56" box="[1154,1251,980,1006]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
is involute with an entirely closed umbilicus (
<figureCitation id="134D2A05500A120FFBD3CBE63ADCFBB7" box="[1093,1204,1012,1039]" captionStart="FIG" captionStartId="9.[133,143,615,632]" captionTargetBox="[279,1308,194,590]" captionTargetId="figure-697@9.[279,1121,194,590]" captionTargetPageId="9" captionText="FIG. 6. — Historical material of Epitornoceras baldisi (Leanza, 1968), identified as Tornoceras baldisi Leanza, 1968 by Leanza (1968), from the Chavela Creek (“Quebrada de la Chavela”),Chavela Member, Chigua Formation (San Juan Province, Argentina): A, B, holotype CPUBA 8088 (part and counterpart showing the external mold of the conch); C, additional specimen CPUBA 8089. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.11408615" httpUri="https://zenodo.org/record/11408615/files/figure.png" pageId="11" pageNumber="336">Fig. 6A, B</figureCitation>
); suture lines are visible (see
<bibRefCitation id="EFE74B71500A120FFCC9CC073D86FB96" author="LEANZA A. F." box="[863,1006,1044,1071]" pageId="11" pageNumber="336" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza 1968</bibRefCitation>
: fig. 4) and their shape is characteristic of the genus
<taxonomicName id="4C764D03500A120FFC53CC273A82FBF6" ID-CoL="4DGY" authority="Frech, 1902" authorityName="Frech" authorityYear="1902" box="[965,1258,1076,1102]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFC53CC273A3DFBF6" box="[965,1109,1077,1102]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
<bibRefCitation id="EFE74B71500A120FFBCBCC263A82FBF6" author="FRECH F." box="[1117,1258,1076,1102]" pageId="11" pageNumber="336" pagination="27 - 112" refId="ref10386" refString="FRECH F. 1902. - Uber devonische Ammoneen. Beitrage zur Palaontologie Osterreich-Ungarns und des Orients. Wien and Leipzig 14: 27 - 112 (in German)." type="journal article" year="1902">Frech, 1902</bibRefCitation>
</taxonomicName>
. The counterpart of the specimen shows the external mold with imprints of the growth lines (
<figureCitation id="134D2A05500A120FFC79CC663A2BFB36" box="[1007,1091,1140,1166]" captionStart="FIG" captionStartId="9.[133,143,615,632]" captionTargetBox="[279,1308,194,590]" captionTargetId="figure-697@9.[279,1121,194,590]" captionTargetPageId="9" captionText="FIG. 6. — Historical material of Epitornoceras baldisi (Leanza, 1968), identified as Tornoceras baldisi Leanza, 1968 by Leanza (1968), from the Chavela Creek (“Quebrada de la Chavela”),Chavela Member, Chigua Formation (San Juan Province, Argentina): A, B, holotype CPUBA 8088 (part and counterpart showing the external mold of the conch); C, additional specimen CPUBA 8089. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.11408615" httpUri="https://zenodo.org/record/11408615/files/figure.png" pageId="11" pageNumber="336">Fig. 6B</figureCitation>
). As indicated by
<bibRefCitation id="EFE74B71500A120FFA9ECC673BCFFB36" author="LEANZA A. F." box="[1288,1447,1140,1167]" pageId="11" pageNumber="336" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza (1968)</bibRefCitation>
, they have a biconvex course (characteristic of the genus
<taxonomicName id="4C764D03500A120FFCB9CCA73DD2FB76" ID-CoL="4DGY" authorityName="Frech" authorityYear="1902" box="[815,954,1205,1230]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFCB9CCA73DD2FB76" box="[815,954,1205,1230]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFE74B71500A120FFC7DCCA63B61FB76" author="KORN D. &amp; BOCKWINKEL J." box="[1003,1289,1204,1230]" pageId="11" pageNumber="336" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
). The
<typeStatus id="54CD8822500A120FFAD8CCA63BC7FB76" box="[1358,1455,1204,1230]" pageId="11" pageNumber="336" type="holotype">holotype</typeStatus>
and the new specimens described herein share very similar characteristics and came from the same levels and localities, so they most likely correspond to the same species. The additional specimen of
<bibRefCitation id="EFE74B71500A120FFBC8CD263A95FAF5" author="LEANZA A. F." box="[1118,1277,1331,1358]" pageId="11" pageNumber="336" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza (1968)</bibRefCitation>
, CPUBA 8089, consists of a poorly preserved internal mold (
<figureCitation id="134D2A05500A120FFA81CD413B01FAD5" box="[1303,1385,1363,1389]" captionStart="FIG" captionStartId="9.[133,143,615,632]" captionTargetBox="[279,1308,194,590]" captionTargetId="figure-697@9.[279,1121,194,590]" captionTargetPageId="9" captionText="FIG. 6. — Historical material of Epitornoceras baldisi (Leanza, 1968), identified as Tornoceras baldisi Leanza, 1968 by Leanza (1968), from the Chavela Creek (“Quebrada de la Chavela”),Chavela Member, Chigua Formation (San Juan Province, Argentina): A, B, holotype CPUBA 8088 (part and counterpart showing the external mold of the conch); C, additional specimen CPUBA 8089. Scale bars: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.11408615" httpUri="https://zenodo.org/record/11408615/files/figure.png" pageId="11" pageNumber="336">Fig. 6C</figureCitation>
) of an involute conch; the internal whorl shows a slightly opened umbilicus (pre-adult stage).
</paragraph>
<paragraph id="8BC93680500A120FFCD5CDA13A9AF995" blockId="11.[811,1457,980,2028]" pageId="11" pageNumber="336">
The specimen from
<collectingCountry id="F3617610500A120FFBB7CDA13A05FA75" box="[1057,1133,1459,1485]" name="Bolivia" pageId="11" pageNumber="336">Bolivia</collectingCountry>
identified as
<taxonomicName id="4C764D03500A120FFA93CDA63D9AFA55" authorityName="House" authorityYear="1978" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi" subGenus="Tornoceras">
<emphasis id="B902EA92500A120FFA93CDA63B1AFA75" box="[1285,1394,1460,1485]" italics="true" pageId="11" pageNumber="336">Tornoceras</emphasis>
(
<emphasis id="B902EA92500A120FFA15CDA63D1EFA55" italics="true" pageId="11" pageNumber="336">Tornoceras</emphasis>
) cf.
<emphasis id="B902EA92500A120FFC3BCDC13D9AFA55" box="[941,1010,1491,1517]" italics="true" pageId="11" pageNumber="336">baldisi</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE74B71500A120FFB8BCDC13AC4FA55" author="HOUSE M. R." box="[1053,1196,1491,1517]" pageId="11" pageNumber="336" pagination="1 - 70" refId="ref10787" refString="HOUSE M. R. 1978. - Devonian Ammonoids from the Appalachians and their bearing on international zonation and correlation. Special Papers in Palaeontology 21: 1 - 70." type="journal article" year="1978">House (1978</bibRefCitation>
: pl. 6, figs 9, 10) has a much stouter conch than
<taxonomicName id="4C764D03500A120FFBDECDE13AC2F9B5" authorityName="House" authorityYear="1978" box="[1096,1194,1523,1549]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFBDECDE13AC2F9B5" box="[1096,1194,1523,1549]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
. Thus, it can be neither identified as
<taxonomicName id="4C764D03500A120FFC20CE013A71F994" authorityName="House" authorityYear="1978" box="[950,1049,1554,1580]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFC20CE013A71F994" box="[950,1049,1554,1580]" italics="true" pageId="11" pageNumber="336">E. baldisi</emphasis>
</taxonomicName>
nor as
<taxonomicName id="4C764D03500A120FFBF1CE013A83F994" authorityName="House" authorityYear="1978" box="[1127,1259,1554,1580]" class="Cephalopoda" family="Tornoceratidae" genus="Tornoceras" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500A120FFBF1CE013A17F994" box="[1127,1151,1555,1580]" italics="true" pageId="11" pageNumber="336">E.</emphasis>
cf.
<emphasis id="B902EA92500A120FFB30CE003A83F994" box="[1190,1259,1554,1580]" italics="true" pageId="11" pageNumber="336">baldisi</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC93680500A1208FCD5CE213FADF853" blockId="11.[811,1457,980,2028]" lastBlockId="12.[132,775,1873,2027]" lastPageId="12" lastPageNumber="337" pageId="11" pageNumber="336">
In
<collectingCountry id="F3617610500A120FFCF4CE213DAEF9F5" box="[866,966,1587,1613]" name="Morocco" pageId="11" pageNumber="336">Morocco</collectingCountry>
, the genus
<taxonomicName id="4C764D03500A120FFBD6CE213AA2F9F4" authorityName="Frech" authorityYear="1902" box="[1088,1226,1587,1612]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFBD6CE213AA2F9F4" box="[1088,1226,1587,1612]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
occurs from the topmost middle Givetian (
<taxonomicName id="4C764D03500A120FFBA6CE403BE0F9D4" authority="Zone" authorityName="Zone" box="[1072,1416,1618,1644]" class="Cephalopoda" family="Pharciceratidae" genus="Pharciceras" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="amplexum">
<emphasis id="B902EA92500A120FFBA6CE403ACCF9D4" box="[1072,1188,1618,1644]" italics="true" pageId="11" pageNumber="336">Pharciceras</emphasis>
aff.
<emphasis id="B902EA92500A120FFB4CCE403B2FF9D4" box="[1242,1351,1618,1644]" italics="true" pageId="11" pageNumber="336">amplexum</emphasis>
Zone
</taxonomicName>
) to the topmost late Givetian (
<taxonomicName id="4C764D03500A120FFBC1CE613B30F934" authority="Zone" authorityName="Zone" box="[1111,1368,1650,1676]" class="Cephalopoda" family="Petteroceratidae" genus="Petteroceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="errans">
<emphasis id="B902EA92500A120FFBC1CE613B7CF934" box="[1111,1300,1651,1676]" italics="true" pageId="11" pageNumber="336">Petteroceras errans</emphasis>
Zone
</taxonomicName>
) deposits (
<bibRefCitation id="EFE74B71500A120FFCC1CE803A39F914" author="BECKER R. T. &amp; HOUSE M. R." box="[855,1105,1682,1709]" pageId="11" pageNumber="336" pagination="27 - 36" refId="ref9190" refString="BECKER R. T. &amp; HOUSE M. R. 2000. - Late Givetian and Frasnian ammonoid succession at Bou Tchrafine (Anti-Atlas, Southern Morocco). Notes et Memoires du Service Geologique du Maroc 399: 27 - 36." type="journal article" year="2000">Becker &amp; House 2000</bibRefCitation>
;
<bibRefCitation id="EFE74B71500A120FFBF7CE803B08F914" author="BOCKWINKEL J. &amp; BECKER R. T. &amp; EBBIGHAUSEN V." box="[1121,1376,1682,1708]" pageId="11" pageNumber="336" pagination="61 - 128" refId="ref9393" refString="BOCKWINKEL J., BECKER R. T. &amp; EBBIGHAUSEN V. 2009. - Upper Givetian ammonoids from Dar Kaoua (Tafilalt, SE Anti-Atlas, Morocco). Berliner palaobiologische Abhandlungen 10: 61 - 128" type="journal article" year="2009">
Bockwinkel
<emphasis id="B902EA92500A120FFB7CCE813B77F914" box="[1258,1311,1682,1708]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
2009
</bibRefCitation>
,
<bibRefCitation id="EFE74B71500A120FFAE6CE803BC0F914" author="BOCKWINKEL J. &amp; KORN D. &amp; EBBIGHAUSEN V. &amp; GRAF S." box="[1392,1448,1682,1708]" pageId="11" pageNumber="336" pagination="257 - 274" refId="ref9432" refString="BOCKWINKEL J., KORN D., EBBIGHAUSEN V. &amp; GRAF S. 2013. - Late Givetian ammonoids from Hagen-Herbeck, Donnerkuhle quarry (Devonian, Rhenish Mountains). Neues Jahrbuch fur Geologie und Palaontologie - Abhandlungen 270 (3): 257 - 274. https: // doi. org / 10.1127 / 0077 - 7749 / 2013 / 0369" type="journal article" year="2013">2013</bibRefCitation>
, 2015, 2017;
<bibRefCitation id="EFE74B71500A120FFC20CEA03AB6F974" author="ABOUSSALAM Z. S. &amp; BECKER R. T." box="[950,1246,1714,1740]" pageId="11" pageNumber="336" pagination="136 - 164" refId="ref8499" refString="ABOUSSALAM Z. S. &amp; BECKER R. T. 2011. - The global Taghanic Biocrisis (Givetian) in the eastern Anti-Atlas, Morocco, Sea-level cyclicity, climate change, and bioevents in Middle Devonian marine and terrestrial environments. Palaeogeography, Palaeoclimatology, Palaeoecology 304 (1): 136 - 164. https: // doi. org / 10.1016 / j. palaeo. 2010.10.015" type="journal article" year="2011">Aboussalam &amp; Becker 2011</bibRefCitation>
;
<bibRefCitation id="EFE74B71500A120FFB7DCEA03BC0F974" author="BECKER R. T. &amp; ABOUSSALAM Z. S. &amp; HARTENFELS S. &amp; EL HASSANI A. &amp; FISHER T." box="[1259,1448,1714,1740]" pageId="11" pageNumber="336" pagination="61 - 76" refId="ref9234" refString="BECKER R. T., ABOUSSALAM Z. S., HARTENFELS S., EL HASSANI A. &amp; FISHER T. 2013. - The Givetian - Famennian at Oum el Jerane (Amessoui Syncline, southern Tafilalt). Document de l'Institut Scientifique, Rabat: 61 - 76." type="book chapter" year="2013">
Becker
<emphasis id="B902EA92500A120FFAAFCEA13B03F974" box="[1337,1387,1714,1740]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
2013
</bibRefCitation>
;
<bibRefCitation id="EFE74B71500A120FFCBBCEC03A52F954" author="ABOUSSALAM Z. S. &amp; BECKER R. T. &amp; MAYER O. &amp; GIBB M. A. &amp; HUNEKE H." box="[813,1082,1746,1772]" pageId="11" pageNumber="336" pagination="312 - 326" refId="ref8571" refString="ABOUSSALAM Z. S., BECKER R. T., MAYER O., GIBB M. A. &amp; HUNEKE H. 2018. - Emsian to middle Famennian bioevents and cephalopod faunas at Mdoura (western Tafilalt Platform). Munstersche Forschungen zur Geologie und Palaontologie 110: 312 - 326." type="journal article" year="2018">
Aboussalam
<emphasis id="B902EA92500A120FFC2ACEC13D9CF954" box="[956,1012,1746,1772]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
2018
</bibRefCitation>
). In
<collectingCountry id="F3617610500A120FFBE3CEC03AB7F954" box="[1141,1247,1746,1772]" name="Germany" pageId="11" pageNumber="336">Germany</collectingCountry>
,
<taxonomicName id="4C764D03500A120FFB79CEC13BE8F954" authorityName="Frech" authorityYear="1902" box="[1263,1408,1747,1772]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFB79CEC13BE8F954" box="[1263,1408,1747,1772]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
was reported in one late Givetian biozone (
<taxonomicName id="4C764D03500A120FFB7FCEE03DD9F894" authority="Zone" authorityName="Zone" class="Cephalopoda" family="Ponticeratidae" genus="Pseudoprobeloceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="pernai">
<emphasis id="B902EA92500A120FFB7FCEE03D07F893" italics="true" pageId="11" pageNumber="336">Pseudoprobeloceras pernai</emphasis>
Zone
</taxonomicName>
;
<bibRefCitation id="EFE74B71500A120FFC2FCF003ABAF894" author="KORN D. &amp; BOCKWINKEL J." box="[953,1234,1810,1836]" pageId="11" pageNumber="336" pagination="32 - 51" refId="ref11277" refString="KORN D. &amp; BOCKWINKEL J. 2022. - The tornoceratid ammonoids from the Roteisenstein Formation of Dillenburg (Cephalopoda, Ammonoidea). European Journal ofTaxonomy 806: 32 - 51. https: // doi. org / 10.5852 / ejt. 2022.806.1699" type="journal article" year="2022">Korn &amp; Bockwinkel 2022</bibRefCitation>
). In North America,
<emphasis id="B902EA92500A120FFCBDCF203D82F8F3" box="[811,1002,1842,1867]" italics="true" pageId="11" pageNumber="336">
<taxonomicName id="4C764D03500A120FFCBDCF203DD1F8F3" authorityName="Frech" authorityYear="1902" box="[811,953,1842,1867]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">Epitornoceras</taxonomicName>
was
</emphasis>
recognized in the lower part of the late Givetian (i.e.,
<taxonomicName id="4C764D03500A120FFC5DCF403AE9F8D3" baseAuthorityName="Hall" baseAuthorityYear="1876" box="[971,1153,1873,1899]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" isUncertain="true" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500A120FFC5DCF403D8BF8D3" box="[971,995,1874,1899]" italics="true" pageId="11" pageNumber="336">E.</emphasis>
aff.
<emphasis id="B902EA92500A120FFB82CF403AE9F8D3" box="[1044,1153,1874,1899]" italics="true" pageId="11" pageNumber="336">peracutum</emphasis>
</taxonomicName>
; upper part of the Geneseo shale,
<bibRefCitation id="EFE74B71500A120FFCE7CF603D9FF834" author="HOUSE M. R." box="[881,1015,1905,1932]" pageId="11" pageNumber="336" pagination="79 - 130" refId="ref10725" refString="HOUSE M. R. 1965. - A study in the tornoceratidae: the succession of tornoceras and related genera in the North American devonian. Philosophical Transactions of the Royal Society of London. Series B, Biological Sciences 250 (763): 79 - 130. https: // doi. org / 10.1098 / rstb. 1965.0021" type="journal article" year="1965">House 1965</bibRefCitation>
,
<bibRefCitation id="EFE74B71500A120FFB91CF633A28F833" author="HOUSE M. R." box="[1031,1088,1905,1931]" pageId="11" pageNumber="336" pagination="1 - 70" refId="ref10787" refString="HOUSE M. R. 1978. - Devonian Ammonoids from the Appalachians and their bearing on international zonation and correlation. Special Papers in Palaeontology 21: 1 - 70." type="journal article" year="1978">1978</bibRefCitation>
) and in early Frasnian rocks (i.e.,
<taxonomicName id="4C764D03500A120FFCB8CF803DC6F813" authorityName="Kirchgasser, Baird &amp; Brett" authorityYear="2020" box="[814,942,1937,1963]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="species" species="dennisoni">
<emphasis id="B902EA92500A120FFCB8CF803DC6F813" box="[814,942,1937,1963]" italics="true" pageId="11" pageNumber="336">E. dennisoni</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFE74B71500A120FFC2ACF833AD9F813" author="KIRCHGASSER W. T. &amp; BAIRD G. C. &amp; BRETT C. E." box="[956,1201,1937,1963]" pageId="11" pageNumber="336" refId="ref10915" refString="KIRCHGASSER W. T., BAIRD G. C. &amp; BRETT C. E. 2020. - Ammonoids from the Tully Formation and Harrell Shale in Pennsylvania: markers of sea-level highstands in the Middle to Late Devonian transition, Appalachian foreland basin, eastern North America, in AVARY K. L., HASSON K. O. &amp; DIECCHIO R. J. (eds), The Appalachian Geology of John M. Dennison: Rocks, People, and a Few Good Restaurants along the Way. Vol. 545. Geological Society of America, Boulder. https: // doi. org / 10.1130 / 2020.2545 (10)" type="book" year="2020">
Kirchgasser
<emphasis id="B902EA92500A120FFBA8CF803A1AF813" box="[1086,1138,1937,1963]" italics="true" pageId="11" pageNumber="336">et al.</emphasis>
2020
</bibRefCitation>
).
<taxonomicName id="4C764D03500A120FFB51CF803B39F813" authorityName="Frech" authorityYear="1902" box="[1223,1361,1938,1963]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="11" pageNumber="336" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500A120FFB51CF803B39F813" box="[1223,1361,1938,1963]" italics="true" pageId="11" pageNumber="336">Epitornoceras</emphasis>
</taxonomicName>
was also reported in early Famennian deposits from
<collectingCountry id="F3617610500A120FFB7BCFA33B22F873" box="[1261,1354,1969,1995]" name="Australia" pageId="11" pageNumber="336">Australia</collectingCountry>
(
<bibRefCitation id="EFE74B71500A120FFACECFA03D0FF854" author="PETERSEN M. S." pageId="11" pageNumber="336" pagination="1 - 55" refId="ref11698" refString="PETERSEN M. S. 1975. - Upper Devonian (Famennian) Ammonoids from the Canning Basin, Western Australia. The Paleontological Society Memoir 8 (5): 1 - 55. https: // www. jstor. org / stable / 1315492" type="journal article" year="1975">Petersen 1975</bibRefCitation>
).
<bibRefCitation id="EFE74B71500A120FFCE9CFC03A73F853" author="HOUSE M. R." box="[895,1051,2001,2028]" pageId="11" pageNumber="336" pagination="1 - 70" refId="ref10787" refString="HOUSE M. R. 1978. - Devonian Ammonoids from the Appalachians and their bearing on international zonation and correlation. Special Papers in Palaeontology 21: 1 - 70." type="journal article" year="1978">House (1978)</bibRefCitation>
, however, stated that the specimens identified there as
<taxonomicName id="4C764D03500D1208FED7CF403C53F8D3" baseAuthorityName="Hall" baseAuthorityYear="1876" box="[321,571,1874,1899]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="12" pageNumber="337" phylum="Mollusca" rank="species" species="peracutum">
<emphasis id="B902EA92500D1208FED7CF403C53F8D3" box="[321,571,1874,1899]" italics="true" pageId="12" pageNumber="337">Epitornoceras peracutum</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFE74B71500D1208FDF7CF403D6FF8D4" author="PETERSEN M. S." box="[609,775,1873,1900]" pageId="12" pageNumber="337" pagination="1 - 55" refId="ref11698" refString="PETERSEN M. S. 1975. - Upper Devonian (Famennian) Ammonoids from the Canning Basin, Western Australia. The Paleontological Society Memoir 8 (5): 1 - 55. https: // www. jstor. org / stable / 1315492" type="journal article" year="1975">Petersen (1975)</bibRefCitation>
must be assigned to a different genus. Therefore, the Argentinian
<taxonomicName id="4C764D03500D1208FF5DCF803F3DF813" authorityName="Frech" authorityYear="1902" box="[203,341,1938,1963]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="12" pageNumber="337" phylum="Mollusca" rank="genus">
<emphasis id="B902EA92500D1208FF5DCF803F3DF813" box="[203,341,1938,1963]" italics="true" pageId="12" pageNumber="337">Epitornoceras</emphasis>
</taxonomicName>
most likely indicates a late Givetian age (see also Discussion section below for more details about biostratigraphic correlations).
</paragraph>
<caption id="DF096608500D1208FF12CEE73C6DF8A5" ID-DOI="http://doi.org/10.5281/zenodo.11408619" ID-Zenodo-Dep="11408619" httpUri="https://zenodo.org/record/11408619/files/figure.png" pageId="12" pageNumber="337" startId="12.[132,143,1781,1798]" targetBox="[132,1456,211,1741]" targetPageId="12" targetType="figure">
<paragraph id="8BC93680500D1208FF12CEE73C6DF8A5" blockId="12.[132,1457,1781,1821]" pageId="12" pageNumber="337">
<smallCapsWord id="8D2FA05C500D1208FF12CEE73EF5F8BD" baselines="1794,1794" box="[132,157,1781,1798]" lowerCaseFontSize="5" mainFontSize="7" normCase="title" normString="Fig" pageId="12" pageNumber="337">FIG</smallCapsWord>
. 8. — Artistic reconstruction of the ammonoid
<taxonomicName id="4C764D03500D1208FDB7CEE73D23F8BE" authority="(Leanza, 1968)" baseAuthorityName="Leanza" baseAuthorityYear="1968" box="[545,843,1781,1798]" class="Cephalopoda" family="Tornoceratidae" genus="Epitornoceras" kingdom="Animalia" pageId="12" pageNumber="337" phylum="Mollusca" rank="species" species="baldisi">
<emphasis id="B902EA92500D1208FDB7CEE73CA6F8BE" box="[545,718,1781,1798]" italics="true" pageId="12" pageNumber="337">Epitornoceras baldisi</emphasis>
(
<bibRefCitation id="EFE74B71500D1208FD41CEE73D2FF8BE" author="LEANZA A. F." box="[727,839,1781,1798]" pageId="12" pageNumber="337" pagination="326 - 330" refId="ref11393" refString="LEANZA A. F. 1968. - Acerca del descubrimiento de Ammonoideos Devonicos en la Republica Argentina. Revista de la Asociacion Geologica Argentina 23 (4): 326 - 330 (in Spanish)." type="journal article" year="1968">Leanza, 1968</bibRefCitation>
)
</taxonomicName>
during the Middle Devonian. Paleoartist: H. Santiago Druetta (CICTERRA, Universidad Nacional de Córdoba, Argentina).
</paragraph>
</caption>
<paragraph id="8BC93680500D1208FCD5CF433A1BF853" blockId="12.[812,1457,1873,2028]" pageId="12" pageNumber="337">
The specimen whose shell shows iridescent colors (INGEO- PI-2011;
<figureCitation id="134D2A05500D1208FC0FCF633D86F833" box="[921,1006,1905,1932]" captionStart="FIG" captionStartId="10.[132,143,1206,1223]" captionTargetBox="[140,1453,220,1166]" captionTargetId="figure-531@10.[698,1267,200,746]" captionTargetPageId="10" captionText="FIG. 7. — New specimens of Epitornoceras baldisi (Leanza, 1968) from the type area of the Chigua Formation (San Juan Province, Argentina): A, B, specimen INGEO-PI-2011; C, specimen INGEO-PI-2007; D, suture line of specimen INGEO-PI- 2004 at wh c. 33 mm; E, reconstructed suture line based on specimen INGEO-PI-2007 at wh c. 18 mm; F, specimen INGEO-PI-2005 (counterpart showing the external mold of the conch); G, H, specimen INGEO-PI- 2002 (part and counterpart showing the external mold of the conch); I, specimen INGEO-PI-2004;J, specimen INGEO-PI-2006; K, L, specimen INGEO-PI-2003 (L corresponds to latex cast of internal whorl). Scale bars: 2 mm. The pictures of some specimens have been flipped (A, B, and K)." figureDoi="http://doi.org/10.5281/zenodo.11408617" httpUri="https://zenodo.org/record/11408617/files/figure.png" pageId="12" pageNumber="337">Fig. 7A</figureCitation>
) is the subject of further investigations. Chemical analyses and SEM observations will be performed to understand the origin of these colors, and the results will be presented in a future study.
</paragraph>
</subSubSection>
</treatment>
</document>