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<document id="0566149695C258E7A9DB4063F8381884" ID-DOI="10.1206/816.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5401238" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1630334480974" checkinUser="felipe" docAuthor="Herman, Lee H." docDate="2013" docId="03DF87947D2AD153FF02517EFDF20689" docLanguage="en" docName="B375.pdf" docOrigin="Bulletin of the American Museum of Natural History 2013 (375)" docSource="http://www.bioone.org/doi/abs/10.1206/816.1" docStyle="DocumentStyle:3507F1A1389FCD0DFF4CB4E0ACF2D98A.3:BulAmeMusNatHis.2011-.journal_article.1cover.type2" docStyleId="3507F1A1389FCD0DFF4CB4E0ACF2D98A" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type2" docStyleVersion="3" docTitle="Oedichirus brunneus Wendeler" docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="FFE6FFEC7D18D167FFB6536AFFC2023A" masterDocTitle="Revision Of The New World Species Of Oedichirus (Coleoptera: Staphylinidae: Paederinae: Pinophilini: Procirrina)" masterLastPageNumber="137" masterPageNumber="1" pageNumber="50" updateTime="1699232470662" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="871A40EA9710CE8ACC3071340A7A50BA" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="6851A4DBFDE95ADFB9C60E47F21CE940">Revision Of The New World Species Of Oedichirus (Coleoptera: Staphylinidae: Paederinae: Pinophilini: Procirrina)</mods:title>
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<mods:identifier id="F87DB1B0CF11EE9704E76B658DF801CE" type="DOI">10.1206/816.1</mods:identifier>
<mods:identifier id="1DDE8B18CD77BA9AE5AC9E3E823187E8" type="ISSN">0003-0090</mods:identifier>
<mods:identifier id="C1EC6607AF1B3C02576CC87458CEF8CA" type="Zenodo-Dep">5401238</mods:identifier>
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<treatment id="03DF87947D2AD153FF02517EFDF20689" LSID="urn:lsid:plazi:treatment:03DF87947D2AD153FF02517EFDF20689" httpUri="http://treatment.plazi.org/id/03DF87947D2AD153FF02517EFDF20689" lastPageId="52" lastPageNumber="52" pageId="50" pageNumber="50">
<subSubSection id="C36C65097D2AD155FF02517EFE200073" pageId="50" pageNumber="50" type="nomenclature">
<paragraph id="8BC936827D2AD155FF02517EFDD60011" blockId="50.[180,532,532,585]" box="[180,532,532,555]" pageId="50" pageNumber="50">
<heading id="D08181EE7D2AD155FF02517EFDD60011" box="[180,532,532,555]" centered="true" fontSize="9" level="2" pageId="50" pageNumber="50" reason="2">
<taxonomicName id="4C764D017D2AD155FF02517EFDD60011" ID-CoL="48VSB" authority="Wendeler" authorityName="Wendeler" box="[180,532,532,555]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FF02517EFE5E0011" box="[180,412,532,555]" italics="true" pageId="50" pageNumber="50">Oedichirus brunneus</emphasis>
Wendeler
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC936827D2AD155FF505159FE200073" blockId="50.[180,532,532,585]" box="[230,482,563,585]" pageId="50" pageNumber="50">
<heading id="D08181EE7D2AD155FF505159FE200073" box="[230,482,563,585]" centered="true" fontSize="9" level="2" pageId="50" pageNumber="50" reason="2">
<figureCitation id="134D2A077D2AD155FF505159FEA60072" box="[230,356,563,585]" captionStart="Figs" captionStartId="30.[116,164,1046,1065]" captionTargetBox="[96,1189,212,1016]" captionTargetId="figure-294@30.[96,1189,212,1016]" captionTargetPageId="30" captionText="Figs. 29–31. 29. Oedichirus brunneus. Segment IX, ventral, female. 30. Palaminus sp. Right elytron. 31. Palaminus sp. Abdominal tergum V (SEM)." figureDoi="http://doi.org/10.5281/zenodo.5401266" httpUri="https://zenodo.org/record/5401266/files/figure.png" pageId="50" pageNumber="50">Figures 29</figureCitation>
,
<figureCitation id="134D2A077D2AD155FEC45159FE530072" box="[370,401,563,584]" captionStart="Figs" captionStartId="37.[150,198,1630,1649]" captionTargetBox="[132,1222,211,1597]" captionTargetId="figure-1@37.[131,1224,210,1600]" captionTargetPageId="37" captionText="Figs. 39–40. Oedichirus species. Distribution. 39. Mesoamerica and the Caribbean. 40. South America. (Species described prior to present work preceded with *. Symbols ³ or ♀ signify species known only by the sex indicated. Open circle denotes multiple collecting sites.)" figureDoi="http://doi.org/10.5281/zenodo.5401272" httpUri="https://zenodo.org/record/5401272/files/figure.png" pageId="50" pageNumber="50">40</figureCitation>
,
<figureCitation id="134D2A077D2AD155FE285159FE200073" box="[414,482,563,585]" captionStart="Figs" captionStartId="51.[150,200,1060,1079]" captionTargetBox="[131,1224,212,1029]" captionTargetId="figure-244@51.[131,1224,212,1030]" captionTargetPageId="51" captionText="Figs. 68–69. Oedichirus brunneus. 68. Segment IX, female. 69. Segment IX, vulvar plate, enlarged (CLSM)." figureDoi="http://doi.org/10.5281/zenodo.5401286" httpUri="https://zenodo.org/record/5401286/files/figure.png" pageId="50" pageNumber="50">68–69</figureCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D2AD155FFEB5135FE680093" pageId="50" pageNumber="50" type="reference_group">
<paragraph id="8BC936827D2AD155FFEB5135FDD20049" blockId="50.[93,619,607,681]" box="[93,528,607,627]" pageId="50" pageNumber="50">
<treatmentCitationGroup id="AB6611AC7D2AD155FFEB5135FDD20049" box="[93,528,607,627]" pageId="50" pageNumber="50">
<treatmentCitation id="0AD710937D2AD155FFEB5135FDCF0049" author="Wendeler, H." box="[93,525,607,627]" page="185" pageId="50" pageNumber="50" year="1930">
<taxonomicName id="4C764D017D2AD155FFEB5135FDCF0049" ID-CoL="48VSB" authority="Wendeler, 1930: 185" authorityName="Wendeler" authorityPageNumber="185" authorityYear="1930" box="[93,525,607,627]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FFEB5135FEEE0049" box="[93,300,607,627]" italics="true" pageId="50" pageNumber="50">Oedichirus brunneus</emphasis>
<bibRefCitation id="EFE74B737D2AD155FE82510AFDCF0049" author="Wendeler, H." box="[308,525,608,627]" pageId="50" pageNumber="50" pagination="181 - 192" refId="ref78244" refString="Wendeler, H. 1930. Neue exotische Staphyliniden. Neue Beitrage zur systematischen Insektenkunde 4: 181 - 192." type="journal article" year="1930">Wendeler, 1930: 185</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC936827D2AD155FFCC5116FE680093" blockId="50.[93,619,607,681]" pageId="50" pageNumber="50">
—
<bibRefCitation id="EFE74B737D2AD155FF2A5116FE5000B5" author="Scheerpeltz, O." box="[156,402,636,655]" pageId="50" pageNumber="50" pagination="989 - 1500" refId="ref77791" refString="Scheerpeltz, O. 1933. Staphylinidae VII. In S. Schenkling (editor), Coleopterorum catalogus 6 (129): 989 - 1500. Berlin: Junk." type="journal article" year="1933">Scheerpeltz, 1933: 1217</bibRefCitation>
(catalog). —
<bibRefCitation id="EFE74B737D2AD155FD9E5116FEF30093" author="Blackwelder, R. E." pageId="50" pageNumber="50" pagination="1 - 188" refId="ref74496" refString="Blackwelder, R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 1. U. S. National Museum Bulletin 185: xii + 1 - 188." type="journal article" year="1944">Blackwelder, 1944: 131</bibRefCitation>
(checklist).
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D2AD155FFEB51A8FDEA04A9" pageId="50" pageNumber="50" type="materials_examined">
<paragraph id="8BC936827D2AD155FFEB51A8FF1F0672" blockId="50.[93,620,705,1683]" pageId="50" pageNumber="50">
TYPE MATERIAL: One
<typeStatus id="54CD88207D2AD155FEFD51A8FE5C00EF" box="[331,414,706,725]" pageId="50" pageNumber="50" type="syntype">syntype</typeStatus>
. One specimen was sent to me from the Museum für Naturkunde, Berlin, where the Wendeler collection is distributed in the main collection. The locality data on the label differ slightly from those published by Wendeler (see Remarks below), but his identification label is attached to the specimen. Although a red ‘‘Holotypus’’ label is one of five attached to the specimen, the
<typeStatus id="54CD88207D2AD155FE1950FCFDC80193" box="[431,522,918,937]" pageId="50" pageNumber="50" type="holotype">holotype</typeStatus>
selection was not published nor did Wendeler state or imply that he had studied only
<specimenCount id="9D70FD0B7D2AD155FE0A50A6FD8801E4" box="[444,586,971,991]" pageId="50" pageNumber="50" type="generic">one specimen</specimenCount>
, so the specimen is a
<typeStatus id="54CD88207D2AD155FE86508CFE4201C3" box="[304,384,998,1017]" pageId="50" pageNumber="50" type="syntype">syntype</typeStatus>
(
<bibRefCitation id="EFE74B737D2AD155FE39508CFDD201C3" author="ICZN &amp; International Commission on Zoological Nomenclature" box="[399,528,998,1017]" pageId="50" pageNumber="50" refId="ref76185" refString="ICZN (International Commission on Zoological Nomenclature). 1999. International code of zoological nomenclature. 4 th ed. London: International Trust for Zoological Nomenclature, xxix + 306 pp." type="book" year="1999">ICZN, 1999</bibRefCitation>
: Articles 73.1.1, 73.1.2, 73.2). To fix the identity of the species that specimen is herein designated as the
<typeStatus id="54CD88207D2AD155FFCE575FFF1B0672" box="[120,217,1077,1096]" pageId="50" pageNumber="50" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8BC936827D2AD155FFEB573BFE4A0787" blockId="50.[93,620,705,1683]" pageId="50" pageNumber="50">
<materialsCitation id="3B1E3CDF7D2AD155FFEB573BFDA606D3" collectingDate="1926-09" collectorName="da Serra &amp; Naturschutzpark &amp; Santos &amp; Kamme &amp; Serra &amp; Sehr &amp; Regenwald &amp; Oedichrus" country="Brazil" county="Designated" determinerName="Wendeler" elevation="700" location="Alto da Serra" municipality="Female" pageId="50" pageNumber="50" specimenCount="1" stateProvince="Sao Paulo" typeStatus="lectotype">
<typeStatus id="54CD88207D2AD155FFEB573BFF130659" box="[93,209,1105,1124]" pageId="50" pageNumber="50" type="lectotype">LECTOTYPE</typeStatus>
:
<collectingCounty id="62A84E0E7D2AD155FF5E573BFE9F065E" box="[232,349,1105,1124]" pageId="50" pageNumber="50">Designated</collectingCounty>
here.
<collectingMunicipality id="6BADACF87D2AD155FE06573BFE3E065E" box="[432,508,1105,1124]" pageId="50" pageNumber="50">Female</collectingMunicipality>
with the following labels: ‘‘
<collectingCountry id="F36176127D2AD155FE805701FE570644" box="[310,405,1131,1150]" name="Brazil" pageId="50" pageNumber="50">Brasilien</collectingCountry>
<collectingRegion id="49B2F8607D2AD155FE2B5706FDCF0644" box="[413,525,1131,1151]" country="Brazil" name="Sao Paulo" pageId="50" pageNumber="50">Sao Paulo</collectingRegion>
,
<location id="8EA960597D2AD155FDAF5701FF7006A3" LSID="urn:lsid:plazi:treatment:03DF87947D2AD153FF02517EFDF20689:8EA960597D2AD155FDAF5701FF7006A3" country="Brazil" county="Designated" municipality="Female" name="Alto da Serra" pageId="50" pageNumber="50" stateProvince="Sao Paulo">
Alto
<collectorName id="268353547D2AD155FDE75701FF7006A3" pageId="50" pageNumber="50">da Serra</collectorName>
</location>
,
<quantity id="4C8E9B677D2AD155FF7357ECFECF06A3" box="[197,269,1158,1177]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="50" pageNumber="50" unit="m" value="700.0">
<elevation id="005BD1B17D2AD155FF7357ECFECF06A3" box="[197,269,1158,1177]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="50" pageNumber="50" unit="m" value="700.0">700 m</elevation>
</quantity>
<location id="8EA960597D2AD155FEAB57ECFE0A06A3" LSID="urn:lsid:plazi:treatment:03DF87947D2AD153FF02517EFDF20689:8EA960597D2AD155FEAB57ECFE0A06A3" box="[285,456,1158,1177]" country="Brazil" county="Designated" municipality="Female" name="Biol Stat. Ohs." pageId="50" pageNumber="50" stateProvince="Sao Paulo">Biol Stat. Ohs.</location>
<date id="FFC810427D2AD155FE6157ECFDF706A3" box="[471,565,1158,1177]" pageId="50" pageNumber="50" value="1926-09">
<collectingDate id="EF8CE9AA7D2AD155FE6157ECFDF706A3" box="[471,565,1158,1177]" pageId="50" pageNumber="50" value="1926-09">IX 1926</collectingDate>
</date>
/
<collectorName id="268353547D2AD155FD8B57ECFEC3068E" pageId="50" pageNumber="50">Naturschutzpark</collectorName>
zw.
<collectorName id="268353547D2AD155FEF357CBFE49068E" box="[325,395,1185,1204]" pageId="50" pageNumber="50">Santos</collectorName>
&amp;
<collectingRegion id="49B2F8607D2AD155FE0957CBFDF4068E" box="[447,566,1185,1204]" country="Brazil" name="Sao Paulo" pageId="50" pageNumber="50">Sao Paulo</collectingRegion>
, am
<collectorName id="268353547D2AD155FFCE57D6FF0906F5" box="[120,203,1212,1231]" pageId="50" pageNumber="50">Kamme</collectorName>
d.
<collectorName id="268353547D2AD155FF5F57D1FEE606F4" box="[233,292,1211,1230]" pageId="50" pageNumber="50">Serra</collectorName>
doMar/
<collectorName id="268353547D2AD155FE3757D1FE7306F4" box="[385,433,1211,1230]" pageId="50" pageNumber="50">Sehr</collectorName>
feuchter
<collectorName id="268353547D2AD155FD9757D6FF6B06D3" pageId="50" pageNumber="50">Regenwald</collectorName>
/
<collectorName id="268353547D2AD155FF0657BCFED806D3" box="[176,282,1238,1257]" pageId="50" pageNumber="50">Oedichrus</collectorName>
brunneus
<taxonomicNameLabel id="A23157EB7D2AD155FE2457BCFE0606D3" box="[402,452,1238,1257]" pageId="50" pageNumber="50" rank="species">n.sp.</taxonomicNameLabel>
<determinerName id="7E15DA467D2AD155FE7957BCFDF006D3" box="[463,562,1238,1257]" pageId="50" pageNumber="50">Wendeler</determinerName>
det.
</materialsCitation>
/
<typeStatus id="54CD88207D2AD155FFCE579AFF210739" box="[120,227,1264,1283]" pageId="50" pageNumber="50" type="holotype">Holotypus</typeStatus>
/
<typeStatus id="54CD88207D2AD155FF5D579AFE960739" box="[235,340,1264,1283]" pageId="50" pageNumber="50" type="lectotype">Lectotype</typeStatus>
<taxonomicName id="4C764D017D2AD155FEEA579AFF690724" authority="Wendeler" authorityName="Wendeler" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FEEA579AFDE8073E" box="[348,554,1264,1284]" italics="true" pageId="50" pageNumber="50">Oedichirus brunneus</emphasis>
Wendeler
</taxonomicName>
des. L. Herman, 2011.’’ Deposited in the Museum für Naturkunde, Berlin. (Left antennomeres 8–11, right antennomeres 3–11, left mesotarsus, and left metatibia and metatarsus are missing. The head and prothorax are separated from the pterothorax and abdomen; both are glued to a card.)
</paragraph>
<paragraph id="8BC936827D2AD155FFEB56ACFDEA04A9" blockId="50.[93,620,705,1683]" pageId="50" pageNumber="50">
TYPE LOCALITY:
<emphasis id="B902EA907D2AD155FEA556AFFE9207E2" bold="true" box="[275,336,1477,1496]" pageId="50" pageNumber="50">
<collectingCountry id="F36176127D2AD155FEA556AFFE9207E2" box="[275,336,1477,1496]" name="Brazil" pageId="50" pageNumber="50">Brazil</collectingCountry>
</emphasis>
:
<emphasis id="B902EA907D2AD155FED756ACFE0F07E3" box="[353,461,1477,1497]" italics="true" pageId="50" pageNumber="50">
<collectingRegion id="49B2F8607D2AD155FED756ACFE0F07E3" box="[353,461,1477,1497]" country="Brazil" name="Sao Paulo" pageId="50" pageNumber="50">São Paulo</collectingRegion>
</emphasis>
: Alto da Serra (5 Paranapiacaba),
<quantity id="4C8E9B677D2AD155FEE2568AFE5F07CE" box="[340,413,1504,1524]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="50" pageNumber="50" unit="m" value="700.0">700 m</quantity>
, biological station, Nature reserve, crest of Serra do Mar (23
<emphasis id="B902EA907D2AD155FD865691FDFB0434" bold="true" box="[560,569,1531,1550]" pageId="50" pageNumber="50">°</emphasis>
479S, 46
<emphasis id="B902EA907D2AD155FF39557FFF5A0412" bold="true" box="[143,152,1557,1576]" pageId="50" pageNumber="50">°</emphasis>
199W). (Alto da Serra is equivalent to Paranapiacaba [
<bibRefCitation id="EFE74B737D2AD155FF58555AFDEF0479" author="Paynter, R. A. &amp; M. A. Traylor" box="[238,557,1584,1603]" pageId="50" pageNumber="50" refId="ref77365" refString="Paynter, R. A., and M. A. Traylor. 1991. Ornithological gazetteer of Brazil, iii - vii + 788 pp. Cambridge, MA: Harvard University, [published in 2 volumes, pagination continuous]" type="book" year="1991">Paynter and Traylor, 1991: 14</bibRefCitation>
, 439]. Serra do Mar is
<quantity id="4C8E9B677D2AD155FEAC5521FE9A0467" box="[282,344,1610,1630]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="50" pageNumber="50" unit="km" value="40.0">40 km</quantity>
southeast of the city of
<collectingRegion id="49B2F8607D2AD155FDF35521FF700442" country="Brazil" name="Sao Paulo" pageId="50" pageNumber="50">São Paulo</collectingRegion>
between that city and Santos. The coordinates are from
<bibRefCitation id="EFE74B737D2AD155FEA755EAFDDC04A9" author="Paynter, R. A. &amp; M. A. Traylor" box="[273,542,1664,1683]" pageId="50" pageNumber="50" refId="ref77365" refString="Paynter, R. A., and M. A. Traylor. 1991. Ornithological gazetteer of Brazil, iii - vii + 788 pp. Cambridge, MA: Harvard University, [published in 2 volumes, pagination continuous]" type="book" year="1991">Paynter and Traylor, 1991</bibRefCitation>
.)
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D2AD154FFCC55C0FE7C073D" lastPageId="51" lastPageNumber="51" pageId="50" pageNumber="50" type="diagnosis">
<paragraph id="8BC936827D2AD155FFCC55C0FDA904E5" blockId="50.[93,620,1706,1759]" pageId="50" pageNumber="50">
DIAGNOSIS: The species is known only from the female
<typeStatus id="54CD88207D2AD155FEAC55A3FE4504E5" box="[282,391,1737,1759]" pageId="50" pageNumber="50" type="lectotype">lectotype</typeStatus>
. The position of the
</paragraph>
<paragraph id="8BC936827D2AD155FD2D53BFFCB304BC" blockId="50.[667,1193,213,1759]" pageId="50" pageNumber="50">
vulvar lobes of the median gonocoxal plate relative to one another and the surface microstructures (fig. 69) and the steeply sloped median gonocoxal plate anteriad of the vulvar plate (fig. 68) will separate
<taxonomicName id="4C764D017D2AD155FB3A5220FCC20344" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FB3A5220FCC20344" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
from most other species. Species with the most similar form and adornment of the vulvar lobes include
<emphasis id="B902EA907D2AD155FC4F52C8FBB003EF" italics="true" pageId="50" pageNumber="50">
<taxonomicName id="4C764D017D2AD155FC4F52C8FBB80382" box="[1017,1146,417,440]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="batillus">O. batillus</taxonomicName>
,
<taxonomicName id="4C764D017D2AD155FB3A52C8FCDE03EF" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="bullaglaber">O. bullaglaber</taxonomicName>
,
<taxonomicName id="4C764D017D2AD155FC9D52D5FC7503EF" box="[811,951,446,469]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="distortus">O. distortus</taxonomicName>
,
<taxonomicName id="4C764D017D2AD155FC7052D5FBAC03EF" box="[966,1134,446,469]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="geniculatus">O. geniculatus</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C764D017D2AD155FD2D52B7FC8F03C8" box="[667,845,477,498]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="sparsipennis">
<emphasis id="B902EA907D2AD155FD2D52B7FC8F03C8" box="[667,845,477,498]" italics="true" pageId="50" pageNumber="50">O. sparsipennis</emphasis>
</taxonomicName>
.
<taxonomicName id="4C764D017D2AD155FCD752B6FB8E03C9" box="[865,1100,476,499]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FCD752B6FB8E03C9" box="[865,1100,476,499]" italics="true" pageId="50" pageNumber="50">Oedichirus brunneus</emphasis>
</taxonomicName>
can be separated from
<taxonomicName id="4C764D017D2AD155FCE95290FC18002A" box="[863,986,505,528]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="boehmi">
<emphasis id="B902EA907D2AD155FCE95290FC18002A" box="[863,986,505,528]" italics="true" pageId="50" pageNumber="50">O. boehmi</emphasis>
</taxonomicName>
by the pronotal punctation and form of the posterior and anterior vulvar lobes (cf. fig. 69 with 67). The anteroventral angles of tergum IX of the female of
<taxonomicName id="4C764D017D2AD155FCB85105FC5100BF" box="[782,915,622,645]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="distortus">
<emphasis id="B902EA907D2AD155FCB85105FC5100BF" box="[782,915,622,645]" italics="true" pageId="50" pageNumber="50">O. distortus</emphasis>
</taxonomicName>
are fused (fig. 116), but separated in
<taxonomicName id="4C764D017D2AD155FC8751E7FC790099" box="[817,955,652,675]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FC8751E7FC790099" box="[817,955,652,675]" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
(fig. 68).
<taxonomicName id="4C764D017D2AD155FB8651E6FD2C00FA" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="batillus">
<emphasis id="B902EA907D2AD155FB8651E6FD2C00FA" italics="true" pageId="50" pageNumber="50">Oedichirus batillus</emphasis>
</taxonomicName>
has paratergite III, which is replaced by a paratergal carina (see fig. 23) in
<taxonomicName id="4C764D017D2AD155FB3A51ADFCC200C0" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FB3A51ADFCC200C0" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
. Tergum III of
<taxonomicName id="4C764D017D2AD155FC09518EFBA000C0" box="[959,1122,739,762]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="bullaglaber">
<emphasis id="B902EA907D2AD155FC09518EFBA000C0" box="[959,1122,739,762]" italics="true" pageId="50" pageNumber="50">O. bullaglaber</emphasis>
</taxonomicName>
has a median point (as in sternum of fig. 73), which is lacking in
<taxonomicName id="4C764D017D2AD155FC985075FC75010F" box="[814,951,798,821]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FC985075FC75010F" box="[814,951,798,821]" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
. The anterior vulvar lobe of
<taxonomicName id="4C764D017D2AD155FD445051FC150168" box="[754,983,827,850]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FD445051FC150168" box="[754,983,827,850]" italics="true" pageId="50" pageNumber="50">Oedichirus brunneus</emphasis>
</taxonomicName>
has short spicules (fig. 69) which are absent in
<taxonomicName id="4C764D017D2AD155FBB75030FB6B014A" box="[1025,1193,857,880]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="B902EA907D2AD155FBB75030FB6B014A" box="[1025,1193,857,880]" italics="true" pageId="50" pageNumber="50">O. geniculatus</emphasis>
</taxonomicName>
(fig. 151) and the anterolateral margin of the median gonocoxal plate of
<taxonomicName id="4C764D017D2AD155FBA450FEFB6B0190" box="[1042,1193,915,938]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FBA450FEFB6B0190" box="[1042,1193,915,938]" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
anteriad of the vulvar plate is steeply sloped (fig. 68), but more gradually sloped for
<taxonomicName id="4C764D017D2AD155FB3A50A5FCDB0638" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="B902EA907D2AD155FB3A50A5FCDB0638" italics="true" pageId="50" pageNumber="50">O. geniculatus</emphasis>
</taxonomicName>
(figs. 148–150). The females of
<taxonomicName id="4C764D017D2AD155FB3A5086FCC20625" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FB3A5086FCC20625" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C764D017D2AD155FC8F5763FC270625" box="[825,997,1033,1055]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="sparsipennis">
<emphasis id="B902EA907D2AD155FC8F5763FC270625" box="[825,997,1033,1055]" italics="true" pageId="50" pageNumber="50">O. sparsipennis</emphasis>
</taxonomicName>
are similar; both have bicolored legs, a paratergal carina; the median gonocoxal plate anteriad of the vulvar plate of both species is triangular, but the apex is wider in
<taxonomicName id="4C764D017D2AD155FC445715FB6B06AE" box="[1010,1193,1151,1172]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="sparsipennis">
<emphasis id="B902EA907D2AD155FC445715FB6B06AE" box="[1010,1193,1151,1172]" italics="true" pageId="50" pageNumber="50">O. sparsipennis</emphasis>
</taxonomicName>
(fig. 206) than
<taxonomicName id="4C764D017D2AD155FCFC57F6FC160688" box="[842,980,1179,1202]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FCFC57F6FC160688" box="[842,980,1179,1202]" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
(fig. 68). The two species differ slightly as follows: for
<taxonomicName id="4C764D017D2AD155FB3A57D3FCC206D6" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FB3A57D3FCC206D6" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
the lateral margin of the paratergal carina is nearly straight in dorsal view, the anterior vulvar lobe has small, scattered cuticular processes (fig. 69), and it is from the state of
<collectingRegion id="49B2F8607D2AD155FCFE5621FC0C075B" box="[840,974,1355,1377]" country="Brazil" name="Sao Paulo" pageId="50" pageNumber="50">São Paulo</collectingRegion>
, whereas for
<taxonomicName id="4C764D017D2AD155FB3A5626FCE60744" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="sparsipennis">
<emphasis id="B902EA907D2AD155FB3A5626FCE60744" italics="true" pageId="50" pageNumber="50">O. sparsipennis</emphasis>
</taxonomicName>
the lateral margin of the paratergal carina is convex in dorsal view, the anterior vulvar lobe lacks cuticular processes or any other adornment (fig. 207), and the species is from the state of
<taxonomicName id="4C764D017D2AD155FC5B56B4FCC5042B" authority="Catarina. However" authorityName="Catarina. However" class="Insecta" family="Hesperiidae" genus="Santa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="genus">Santa Catarina. However</taxonomicName>
, the two species differ only slightly; it is likely that the question will remain unresolved until males and more females of
<taxonomicName id="4C764D017D2AD155FD2D5539FCE40453" box="[667,806,1618,1641]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2AD155FD2D5539FCE40453" box="[667,806,1618,1641]" italics="true" pageId="50" pageNumber="50">O. brunneus</emphasis>
</taxonomicName>
have been found and compared to
<taxonomicName id="4C764D017D2AD155FD0A551BFCA804BC" box="[700,874,1649,1670]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="50" phylum="Arthropoda" rank="species" species="sparsipennis">
<emphasis id="B902EA907D2AD155FD0A551BFCA804BC" box="[700,874,1649,1670]" italics="true" pageId="50" pageNumber="50">O. sparsipennis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BC936827D2AD154FD0E55E5FE7C073D" blockId="50.[667,1193,213,1759]" lastBlockId="51.[128,654,1148,1759]" lastPageId="51" lastPageNumber="51" pageId="50" pageNumber="50">Externally, diagnostic features include the bicolored yellowish-brown legs with a dark reddish-brown spot at the femorotibial joint, the quadridentate labrum, the equal length and width of the elytra, the short, welldeveloped paratergal carina of segment III reaches the spiracle, and the dense, coarse, dorsal cephalic punctation.</paragraph>
</subSubSection>
<caption id="DF09660A7D2BD154FF20574EFEFD066B" ID-DOI="http://doi.org/10.5281/zenodo.5401286" ID-Zenodo-Dep="5401286" httpUri="https://zenodo.org/record/5401286/files/figure.png" pageId="51" pageNumber="51" startId="51.[150,200,1060,1079]" targetBox="[131,1224,212,1029]" targetPageId="51">
<paragraph id="8BC936827D2BD154FF20574EFEFD066B" blockId="51.[128,1227,1059,1105]" pageId="51" pageNumber="51">
Figs. 68–69.
<taxonomicName id="4C764D017D2BD154FE8D5749FDDE060D" box="[315,540,1059,1079]" class="Insecta" family="Staphylinidae" genus="Oedichirus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="51" phylum="Arthropoda" rank="species" species="brunneus">
<emphasis id="B902EA907D2BD154FE8D5749FDDE060D" box="[315,540,1059,1079]" italics="true" pageId="51" pageNumber="51">Oedichirus brunneus</emphasis>
</taxonomicName>
.
<emphasis id="B902EA907D2BD154FD82574EFD96060D" bold="true" box="[564,596,1060,1079]" pageId="51" pageNumber="51">68.</emphasis>
Segment IX, female.
<emphasis id="B902EA907D2BD154FCD0574EFC44060D" bold="true" box="[870,902,1060,1079]" pageId="51" pageNumber="51">69.</emphasis>
Segment IX, vulvar plate, enlarged (CLSM).
</paragraph>
</caption>
<subSubSection id="C36C65097D2BD153FF2A5664FE0D063B" lastPageId="52" lastPageNumber="52" pageId="51" pageNumber="51" type="description">
<paragraph id="8BC936827D2BD154FF2A5664FEF107A0" blockId="51.[128,654,1148,1759]" pageId="51" pageNumber="51">
DESCRIPTION: Length:
<quantity id="4C8E9B677D2BD154FE1C567AFDD0071F" box="[426,530,1296,1317]" metricMagnitude="-2" metricUnit="m" metricValue="1.02" pageId="51" pageNumber="51" unit="mm" value="10.2">10.2 mm</quantity>
. Length of head:
<quantity id="4C8E9B677D2BD154FF675647FEF50778" box="[209,311,1325,1346]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="51" pageNumber="51" unit="mm" value="1.2">1.2 mm</quantity>
. Width of head:
<quantity id="4C8E9B677D2BD154FD925646FD480778" box="[548,650,1324,1346]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="51" unit="mm" value="1.5">1.5 mm</quantity>
. Pronotal length: 2.0 mm. Pronotal width:
<quantity id="4C8E9B677D2BD154FF36560DFF240747" box="[128,230,1383,1405]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="51" unit="mm" value="1.5">1.5 mm</quantity>
. Elytral length:
<quantity id="4C8E9B677D2BD154FE77560DFDEA0747" box="[449,552,1383,1405]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="51" unit="mm" value="1.5">1.5 mm</quantity>
. Elytral width:
<quantity id="4C8E9B677D2BD154FF6456EEFEED07A0" box="[210,303,1412,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="51" unit="mm" value="1.5">1.5 mm</quantity>
.
</paragraph>
<paragraph id="8BC936827D2BD154FF2B56C9FE6907C9" blockId="51.[128,654,1148,1759]" pageId="51" pageNumber="51">Body concolorous reddish brown. Legs bicolored, yellowish brown with femorotibial spot dark reddish brown.</paragraph>
<paragraph id="8BC936827D2BD154FF2B5696FDF104E5" blockId="51.[128,654,1148,1759]" pageId="51" pageNumber="51">Head wider than long (HW/HL: 1.2). Frontoclypeal ridge complete, not separated medially. Dorsal surface without V-shaped depression; surface uniformly, densely, and coarsely punctate medially, punctation less dense posteriorly and less coarse lateroposteriorly. Labrum quadridentate; surface without tubercle near submedial denticle.</paragraph>
<paragraph id="8BC936827D2BD154FD6D5716FC860747" blockId="51.[701,1228,1148,1759]" pageId="51" pageNumber="51">Pronotum longer than wide (PL/PW: 1.3). Pronotal surface with scattered polished, impunctate spots and with coarse punctation arranged in scattered clusters and irregular rows; submedial punctate groove extending posteriorly from near middle. Elytra with length and width equal (EW/EL: 1.0); surface of disc feebly concave and with coarse punctation.</paragraph>
<paragraph id="8BC936827D2BD153FD6D56ECFE1D0347" blockId="51.[701,1228,1148,1759]" lastBlockId="52.[93,620,213,1203]" lastPageId="52" lastPageNumber="52" pageId="51" pageNumber="51">Abdomen irregularly punctate dorsally, punctures not arranged in transverse rows; punctation of III to VI coarse and dense; punctation of VII and VIII less dense and less coarse. Segment III without paratergite; paratergal carina present and short, reaching to spiracle and with lateral margin nearly straight in dorsal view. Tergum III without median point extending from transverse basal ridge. Tergum VIII with posterior margin truncate or feebly emarginate; transverse basal ridge with apically open median point. Tergum IX with lateroapical process moderately long, nearly twice as long as midbasal length (LLaP/L9 5 1.96), slightly bent ventrally, and approximately parallel to other process; ventromedial margin without posteriorly directed spur (cf. fig. 158).</paragraph>
<paragraph id="8BC936827D2CD153FFCC52E9FE9303A1" blockId="52.[93,620,213,1203]" box="[122,337,387,411]" pageId="52" pageNumber="52">MALE: Unknown.</paragraph>
<paragraph id="8BC936827D2CD153FFCC52CBFE0D063B" blockId="52.[93,620,213,1203]" pageId="52" pageNumber="52">FEMALE: Tergum IX with anteroventral angles separated from each other (fig. 68). Median gonocoxal plate anteriad of vulvar plate large, strongly tapered, trianguliform, and with anterior margin narrowly rounded and strongly sloped lateroposteriorly; gonocoxal plate posteriad of vulvar plate gradually tapered posteriorly to broadly rounded posterior margin. Vulvar plate embedded in anterior half of median gonocoxal plate. Anterior vulvar lobe (fig. 69) transverse and extending across anterior margin of posterior vulvar lobe, with curved posterior margin, and surface moderately densely covered with short cuticular processes. Posterior vulvar lobe with most of surface densely covered with cobble, but with scattered cuticular processes anteriorly on each side of vulvar depression. Vulvar depression membranous and unadorned; vulva longitudinally oriented near midsagittal plane (fig. 69).</paragraph>
</subSubSection>
<subSubSection id="C36C65097D2CD153FFCC5762FE140607" pageId="52" pageNumber="52" type="distribution">
<paragraph id="8BC936827D2CD153FFCC5762FE140607" blockId="52.[93,620,213,1203]" pageId="52" pageNumber="52">
DISTRIBUTION: Known from southeastern
<collectingRegion id="49B2F8607D2CD153FFEB574DFED50606" box="[93,279,1063,1085]" country="Brazil" name="Sao Paulo" pageId="52" pageNumber="52">São Paulo state</collectingRegion>
,
<collectingCountry id="F36176127D2CD153FE93574DFEA80607" box="[293,362,1063,1085]" name="Brazil" pageId="52" pageNumber="52">Brazil</collectingCountry>
(fig. 40).
</paragraph>
</subSubSection>
<subSubSection id="C36C65097D2CD153FFCC5729FDF20689" pageId="52" pageNumber="52" type="discussion">
<paragraph id="8BC936827D2CD153FFCC5729FDF20689" blockId="52.[93,620,213,1203]" pageId="52" pageNumber="52">
REMARKS:
<bibRefCitation id="EFE74B737D2CD153FEB5572FFE3C0661" author="Wendeler, H." box="[259,510,1093,1115]" pageId="52" pageNumber="52" pagination="181 - 192" refId="ref78244" refString="Wendeler, H. 1930. Neue exotische Staphyliniden. Neue Beitrage zur systematischen Insektenkunde 4: 181 - 192." type="journal article" year="1930">Wendeler (1930: 185)</bibRefCitation>
reported the specimen was collected in November, but according to the label attached to the specimen it was collected in September.
</paragraph>
</subSubSection>
</treatment>
</document>
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