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<document id="6D4EA605CBC8E07A3D870C2EEDFE5725" ID-DOI="10.11646/zootaxa.5562.1.9" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595744" ID-ZooBank="A9BB7BE0-4F71-4CE3-B494-34CB50A67F42" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1735932181654" checkinUser="plazi" docAuthor="Álvarez-Parra, Sergio &amp; Huang, Di-Ying" docDate="2024" docId="03D1F769EC62FFF6FF57FF05FD3FFD4D" docLanguage="en" docName="zootaxa.5562.1.9.pdf" docOrigin="Zootaxa 5562 (1)" docSource="https://doi.org/10.11646/zootaxa.5562.1.9" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Manlaya minima Álvarez-Parra &amp; Huang, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="61" masterDocId="FFE88F11EC60FFF3FFC0FF92FFB2FFE4" masterDocTitle="A new species of Manlaya (Hymenoptera: Baissidae) from the Lower Cretaceous Shouchang Formation, Eastern China" masterLastPageNumber="64" masterPageNumber="56" pageNumber="58" updateTime="1736171187634" updateUser="julia" zenodo-license-document="CLOSED">
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<mods:title id="FB34A8BAB9F562B8AD8B9FFF6EAD0CED">A new species of Manlaya (Hymenoptera: Baissidae) from the Lower Cretaceous Shouchang Formation, Eastern China</mods:title>
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<mods:namePart id="6BD77B706CB4CEA092F48CEE5431F592">Álvarez-Parra, Sergio</mods:namePart>
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<mods:namePart id="7902D2D19C08BEE9C34C540040B83AF9">Huang, Di-Ying</mods:namePart>
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<mods:date id="AFDF8177CF5214BD6EE13585E75BFE54">2024</mods:date>
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<subSubSection id="C36215F4EC62FFF1FF57FF05FE7EFF55" box="[151,460,151,178]" pageId="2" pageNumber="58" type="nomenclature">
<paragraph id="8BC7467FEC62FFF1FF57FF05FE7EFF55" blockId="2.[151,995,151,249]" box="[151,460,151,178]" pageId="2" pageNumber="58">
<heading id="D08FF113EC62FFF1FF57FF05FE7EFF55" bold="true" box="[151,460,151,178]" fontSize="11" level="1" pageId="2" pageNumber="58" reason="1">
<emphasis id="B90C9A6DEC62FFF1FF57FF05FE7EFF55" bold="true" box="[151,460,151,178]" pageId="2" pageNumber="58">
<taxonomicName id="4C783DFCEC62FFF1FF57FF05FED5FF55" authorityName="Álvarez-Parra &amp; Huang" authorityYear="2024" box="[151,359,151,178]" class="Insecta" family="Baissidae" genus="Manlaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B90C9A6DEC62FFF1FF57FF05FED5FF55" bold="true" box="[151,359,151,178]" italics="true" pageId="2" pageNumber="58">Manlaya minima</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23F2716EC62FFF1FEAEFF05FE7EFF55" box="[366,460,151,177]" pageId="2" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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urn:lsid:zoobank.org:act:
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</uri>
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<paragraph id="8BC7467FEC62FFF1FF57FF4DFEB9FF1D" blockId="2.[151,995,151,249]" box="[151,267,223,249]" pageId="2" pageNumber="58">
(
<figureCitation id="13435AFAEC62FFF1FF5FFF4DFEB0FF1D" box="[159,258,223,249]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1963,1987]" captionStartId-1="3.[151,250,1876,1900]" captionStartId-2="4.[151,250,1707,1731]" captionTargetBox-0="[346,1241,451,1914]" captionTargetBox-1="[151,1436,597,1853]" captionTargetBox-2="[151,1436,181,1656]" captionTargetId-0="figure-66@2.[346,1241,451,1938]" captionTargetId-1="figure-188@3.[151,1436,597,1853]" captionTargetId-2="figure-17@4.[151,1436,181,1683]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Photograph of the habitus. B, Interpretative drawing of the habitus. Both at the same scale." captionText-1="FIGURE 2. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Head. B, Mesosoma and head. C, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead). D, Legs. E, Metasoma with indication of metasomal segments." captionText-2="FIGURE 3. Forewing of Manlaya minima sp. nov., holotype NIGP206130. A, Photograph of the forewing. B, Name of veins. C, Name of cells. B and C at the same scale." figureDoi-0="http://doi.org/10.5281/zenodo.14595746" figureDoi-1="http://doi.org/10.5281/zenodo.14595748" figureDoi-2="http://doi.org/10.5281/zenodo.14595750" httpUri-0="https://zenodo.org/record/14595746/files/figure.png" httpUri-1="https://zenodo.org/record/14595748/files/figure.png" httpUri-2="https://zenodo.org/record/14595750/files/figure.png" pageId="2" pageNumber="58">Figs 13</figureCitation>
)
</paragraph>
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<subSubSection id="C36215F4EC62FFF1FF57FEB5FAB0FE49" pageId="2" pageNumber="58" type="materials_examined">
<paragraph id="8BC7467FEC62FFF1FF57FEB5FAB0FE49" blockId="2.[151,1436,295,429]" pageId="2" pageNumber="58">
<emphasis id="B90C9A6DEC62FFF1FF57FEB5FEF9FEA5" bold="true" box="[151,331,295,321]" pageId="2" pageNumber="58">Type material.</emphasis>
<materialsCitation id="3B104C22EC62FFF1FE97FEB5FB4CFE49" collectionCode="NIGP" country="China" county="Jiande City" location="Lower Cretaceous Shouchang Formation" municipality="Shouchang Township" pageId="2" pageNumber="58" specimenCode="NIGP206130" specimenCount="1" stateProvince="Zhejiang Province" typeStatus="holotype">
<typeStatus id="54C3F8DDEC62FFF1FE97FEB5FE0DFEA5" box="[343,447,295,321]" pageId="2" pageNumber="58" type="holotype">Holotype</typeStatus>
<specimenCode id="DBDEEE04EC62FFF1FE0CFEB5FDD7FEA5" box="[460,613,295,321]" collectionCode="NIGP" country="China" httpUri="http://grbio.org/cool/j51y-avsb" name="Naking Institute of Geology and Palaeontology" pageId="2" pageNumber="58">NIGP206130</specimenCode>
, a female specimen from lateral view, distalmost parts of wings and ovipositor not preserved.
<emphasis id="B90C9A6DEC62FFF1FF07FEFDFDBEFE6D" bold="true" box="[199,524,367,393]" pageId="2" pageNumber="58">Type locality and horizon.</emphasis>
<location id="8EA710A4EC62FFF1FDD8FEE2FC40FE6D" LSID="urn:lsid:plazi:treatment:03D1F769EC62FFF6FF57FF05FD3FFD4D:8EA710A4EC62FFF1FDD8FEE2FC40FE6D" box="[536,1010,367,394]" country="China" county="Jiande City" municipality="Shouchang Township" name="Lower Cretaceous Shouchang Formation" pageId="2" pageNumber="58" stateProvince="Zhejiang Province">Lower Cretaceous Shouchang Formation</location>
(
<location id="8EA710A4EC62FFF1FBC4FEFDFA22FE6D" LSID="urn:lsid:plazi:treatment:03D1F769EC62FFF6FF57FF05FD3FFD4D:8EA710A4EC62FFF1FBC4FEFDFA22FE6D" box="[1028,1424,367,394]" country="China" county="Jiande City" municipality="Shouchang Township" name="upper Barremian to lower Aptian" pageId="2" pageNumber="58" stateProvince="Zhejiang Province">upper Barremian to lower Aptian</location>
);
<location id="8EA710A4EC62FFF1FF57FE01FEDFFE49" LSID="urn:lsid:plazi:treatment:03D1F769EC62FFF6FF57FF05FD3FFD4D:8EA710A4EC62FFF1FF57FE01FEDFFE49" box="[151,365,403,429]" country="China" county="Jiande City" municipality="Shouchang Township" name="Shanghujia Village" pageId="2" pageNumber="58" stateProvince="Zhejiang Province">Shanghujia Village</location>
(Dongcun section),
<collectingMunicipality id="6BA3DC05EC62FFF1FD91FE01FCF7FE49" box="[593,837,403,429]" pageId="2" pageNumber="58">Shouchang Township</collectingMunicipality>
,
<collectingCounty id="62A63EF3EC62FFF1FC90FE01FC7DFE49" box="[848,975,403,429]" pageId="2" pageNumber="58">Jiande City</collectingCounty>
,
<collectingRegion id="49BC889DEC62FFF1FC1BFE01FB19FE49" box="[987,1195,403,429]" country="China" name="Zhejiang" pageId="2" pageNumber="58">Zhejiang Province</collectingRegion>
,
<collectingCountry id="F36F06EFEC62FFF1FB77FE01FB4CFE49" box="[1207,1278,403,429]" name="China" pageId="2" pageNumber="58">China</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="8BC7467FEC62FFF1FF57F839FBFCF80C" blockId="2.[151,1437,1963,2024]" pageId="2" pageNumber="58">
<emphasis id="B90C9A6DEC62FFF1FF57F839FEA1F827" bold="true" box="[151,275,1963,1987]" pageId="2" pageNumber="58">FIGURE 1.</emphasis>
<taxonomicName id="4C783DFCEC62FFF1FED9F83EFE77F827" authorityName="Álvarez-Parra &amp; Huang" authorityYear="2024" box="[281,453,1964,1988]" class="Insecta" family="Baissidae" genus="Manlaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="58" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B90C9A6DEC62FFF1FED9F83EFE77F827" box="[281,453,1964,1988]" italics="true" pageId="2" pageNumber="58">Manlaya minima</emphasis>
</taxonomicName>
<emphasis id="B90C9A6DEC62FFF1FE0BF83EFDA9F820" bold="true" box="[459,539,1964,1988]" pageId="2" pageNumber="58">
<taxonomicNameLabel id="A23F2716EC62FFF1FE0BF83EFDA9F820" box="[459,539,1964,1988]" pageId="2" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China).
<emphasis id="B90C9A6DEC62FFF1FF57F85DFF1BF803" bold="true" box="[151,169,1999,2023]" pageId="2" pageNumber="58">A</emphasis>
, Photograph of the habitus.
<emphasis id="B90C9A6DEC62FFF1FE08F842FE6BF80C" bold="true" box="[456,473,2000,2024]" pageId="2" pageNumber="58">B</emphasis>
, Interpretative drawing of the habitus. Both at the same scale.
</paragraph>
</caption>
<subSubSection id="C36215F4EC63FFF0FF07FF05FE6FFF31" pageId="3" pageNumber="59" type="etymology">
<paragraph id="8BC7467FEC63FFF0FF07FF05FE6FFF31" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="59">
<emphasis id="B90C9A6DEC63FFF0FF07FF05FEFCFF55" bold="true" box="[199,334,151,177]" pageId="3" pageNumber="59">Etymology.</emphasis>
From the Latin word
<emphasis id="B90C9A6DEC63FFF0FD83FF05FD10FF54" box="[579,674,151,176]" italics="true" pageId="3" pageNumber="59">minimus</emphasis>
, meaning very small, referring to the small body size in comparison to other species of the genus.
</paragraph>
</subSubSection>
<subSubSection id="C36215F4EC63FFF0FF07FF4CFE03FE6D" pageId="3" pageNumber="59" type="diagnosis">
<paragraph id="8BC7467FEC63FFF0FF07FF4CFE03FE6D" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="59">
<emphasis id="B90C9A6DEC63FFF0FF07FF4CFEF0FF1C" bold="true" box="[199,322,222,248]" pageId="3" pageNumber="59">Diagnosis.</emphasis>
Body size small (about
<quantity id="4C80EB9AEC63FFF0FD9CFF4DFD2DFF1D" box="[604,671,223,249]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="3" pageNumber="59" unit="mm" value="2.0">2 mm</quantity>
long); pedicel thicker than scape; pterostigma broad, with anterior margin straight and posterior margin curved; r-rs slightly inclivous, half longer than pterostigma width; 1Rs separated from pterostigma base by length 2× pterostigma basal width, slightly longer than 1M; cell 1mcu strongly transverse, with 1Rs+M and 1Cu parallel; 2Rs+M as long as 1m-cu; 1cu-a slightly postfurcal; cell 2mcu elbowed; vein A sigmoidal basally.
</paragraph>
</subSubSection>
<subSubSection id="C36215F4EC63FFF6FF07FE00FD3FFD4D" lastPageId="5" lastPageNumber="61" pageId="3" pageNumber="59" type="description">
<paragraph id="8BC7467FEC63FFF0FF07FE00FE03FE35" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="59">
<emphasis id="B90C9A6DEC63FFF0FF07FE00FEEBFE48" bold="true" box="[199,345,402,428]" pageId="3" pageNumber="59">Description.</emphasis>
Female. Body stout,
<quantity id="4C80EB9AEC63FFF0FD86FE01FD18FE49" box="[582,682,403,429]" metricMagnitude="-3" metricUnit="m" metricValue="2.07" pageId="3" pageNumber="59" unit="mm" value="2.07">2.07 mm</quantity>
long (
<figureCitation id="13435AFAEC63FFF0FD30FE01FC83FE49" box="[752,817,403,429]" captionStart="FIGURE 1" captionStartId="2.[151,250,1963,1987]" captionTargetBox="[346,1241,451,1914]" captionTargetId="figure-66@2.[346,1241,451,1938]" captionTargetPageId="2" captionText="FIGURE 1. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Photograph of the habitus. B, Interpretative drawing of the habitus. Both at the same scale." figureDoi="http://doi.org/10.5281/zenodo.14595746" httpUri="https://zenodo.org/record/14595746/files/figure.png" pageId="3" pageNumber="59">Fig. 1</figureCitation>
). Colour dark brown, head and mesonotum darker than rest of body. No setation.
</paragraph>
<paragraph id="8BC7467FEC63FFF0FF07FE49FB9CFDD9" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="59">
<emphasis id="B90C9A6DEC63FFF0FF07FE49FEB6FE11" box="[199,260,475,501]" italics="true" pageId="3" pageNumber="59">Head</emphasis>
:
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long and
<quantity id="4C80EB9AEC63FFF0FE2BFE49FDE3FE11" box="[491,593,475,501]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="59" unit="mm" value="0.43">0.43 mm</quantity>
high (
<figureCitation id="13435AFAEC63FFF0FD5BFE49FD44FE11" box="[667,758,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1876,1900]" captionTargetBox="[151,1436,597,1853]" captionTargetId="figure-188@3.[151,1436,597,1853]" captionTargetPageId="3" captionText="FIGURE 2. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Head. B, Mesosoma and head. C, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead). D, Legs. E, Metasoma with indication of metasomal segments." figureDoi="http://doi.org/10.5281/zenodo.14595748" httpUri="https://zenodo.org/record/14595748/files/figure.png" pageId="3" pageNumber="59">Fig. 2A</figureCitation>
); eyes large, rounded, occupying most of side of head,
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in diameter; preserved part of antenna about
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long, pedicel
<quantity id="4C80EB9AEC63FFF0FC09FE6DFB9FFDFD" box="[969,1069,511,537]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="59" unit="mm" value="0.08">0.08 mm</quantity>
long and
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wide, thicker than scape (number of flagellomeres uncertain); mouthparts apparently protruded.
</paragraph>
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<paragraph id="8BC7467FEC63FFF0FF57F8C6FD0DF83D" blockId="3.[151,1437,1876,2009]" pageId="3" pageNumber="59">
<emphasis id="B90C9A6DEC63FFF0FF57F8C6FEA6F888" bold="true" box="[151,276,1876,1900]" pageId="3" pageNumber="59">FIGURE 2.</emphasis>
<taxonomicName id="4C783DFCEC63FFF0FEDCF8C6FE75F888" authorityName="Álvarez-Parra &amp; Huang" authorityYear="2024" box="[284,455,1876,1900]" class="Insecta" family="Baissidae" genus="Manlaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B90C9A6DEC63FFF0FEDCF8C6FE75F888" box="[284,455,1876,1900]" italics="true" pageId="3" pageNumber="59">Manlaya minima</emphasis>
</taxonomicName>
<emphasis id="B90C9A6DEC63FFF0FE0EF8C6FDADF888" bold="true" box="[462,543,1876,1900]" pageId="3" pageNumber="59">
<taxonomicNameLabel id="A23F2716EC63FFF0FE0EF8C6FDADF888" box="[462,543,1876,1900]" pageId="3" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China).
<emphasis id="B90C9A6DEC63FFF0FF57F8EAFF1BF874" bold="true" box="[151,169,1912,1936]" pageId="3" pageNumber="59">A</emphasis>
, Head.
<emphasis id="B90C9A6DEC63FFF0FF3BF8EAFEBEF874" bold="true" box="[251,268,1912,1936]" pageId="3" pageNumber="59">B</emphasis>
, Mesosoma and head.
<emphasis id="B90C9A6DEC63FFF0FE38F8EAFDB8F874" bold="true" box="[504,522,1912,1936]" pageId="3" pageNumber="59">C</emphasis>
, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead).
<emphasis id="B90C9A6DEC63FFF0FA84F80EFAE4F850" bold="true" box="[1348,1366,1948,1972]" pageId="3" pageNumber="59">D</emphasis>
, Legs.
<emphasis id="B90C9A6DEC63FFF0FF57F852FF1AF83C" bold="true" box="[151,168,1984,2008]" pageId="3" pageNumber="59">E</emphasis>
, Metasoma with indication of metasomal segments.
</paragraph>
</caption>
<caption id="DF0716F7EC64FFF7FF57F939FDEEF903" ID-DOI="http://doi.org/10.5281/zenodo.14595750" ID-Zenodo-Dep="14595750" httpUri="https://zenodo.org/record/14595750/files/figure.png" pageId="4" pageNumber="60" startId="4.[151,250,1707,1731]" targetBox="[151,1436,181,1656]" targetPageId="4" targetType="figure">
<paragraph id="8BC7467FEC64FFF7FF57F939FDEEF903" blockId="4.[151,1437,1707,1767]" pageId="4" pageNumber="60">
<emphasis id="B90C9A6DEC64FFF7FF57F939FEA1F927" bold="true" box="[151,275,1707,1731]" pageId="4" pageNumber="60">FIGURE 3.</emphasis>
Forewing of
<taxonomicName id="4C783DFCEC64FFF7FE5DF939FDFBF927" authorityName="Álvarez-Parra &amp; Huang" authorityYear="2024" box="[413,585,1707,1731]" class="Insecta" family="Baissidae" genus="Manlaya" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="minima" status="sp. nov.">
<emphasis id="B90C9A6DEC64FFF7FE5DF939FDFBF927" box="[413,585,1707,1731]" italics="true" pageId="4" pageNumber="60">Manlaya minima</emphasis>
</taxonomicName>
<emphasis id="B90C9A6DEC64FFF7FD90F939FD12F927" bold="true" box="[592,672,1707,1731]" pageId="4" pageNumber="60">
<taxonomicNameLabel id="A23F2716EC64FFF7FD90F939FD12F927" box="[592,672,1707,1731]" pageId="4" pageNumber="60" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype NIGP206130.
<emphasis id="B90C9A6DEC64FFF7FC61F939FC01F927" bold="true" box="[929,947,1707,1731]" pageId="4" pageNumber="60">A</emphasis>
, Photograph of the forewing.
<emphasis id="B90C9A6DEC64FFF7FB24F939FB47F927" bold="true" box="[1252,1269,1707,1731]" pageId="4" pageNumber="60">B</emphasis>
, Name of veins.
<emphasis id="B90C9A6DEC64FFF7FF57F95DFF1BF903" bold="true" box="[151,169,1743,1767]" pageId="4" pageNumber="60">C</emphasis>
, Name of cells.
<emphasis id="B90C9A6DEC64FFF7FE8FF95DFED2F903" bold="true" box="[335,352,1743,1767]" pageId="4" pageNumber="60">B</emphasis>
and
<emphasis id="B90C9A6DEC64FFF7FE52F95DFE16F903" bold="true" box="[402,420,1743,1767]" pageId="4" pageNumber="60">C</emphasis>
at the same scale.
</paragraph>
</caption>
<paragraph id="8BC7467FEC64FFF7FF07F8A8FAB6F87F" blockId="4.[151,1436,1849,2020]" pageId="4" pageNumber="60">
<emphasis id="B90C9A6DEC64FFF7FF07F8A8FE8DF8B7" box="[199,319,1850,1875]" italics="true" pageId="4" pageNumber="60">Mesosoma</emphasis>
: stout (
<figureCitation id="13435AFAEC64FFF7FE59F8ABFE46F8B7" box="[409,500,1849,1875]" captionStart="FIGURE 2" captionStartId="3.[151,250,1876,1900]" captionTargetBox="[151,1436,597,1853]" captionTargetId="figure-188@3.[151,1436,597,1853]" captionTargetPageId="3" captionText="FIGURE 2. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Head. B, Mesosoma and head. C, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead). D, Legs. E, Metasoma with indication of metasomal segments." figureDoi="http://doi.org/10.5281/zenodo.14595748" httpUri="https://zenodo.org/record/14595748/files/figure.png" pageId="4" pageNumber="60">Fig. 2B</figureCitation>
),
<quantity id="4C80EB9AEC64FFF7FDCBF8ABFDC0F8B0" box="[523,626,1849,1876]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="60" unit="mm" value="0.7">0.70 mm</quantity>
long, as long as high (height measured from just above the procoxa); pronotum about
<quantity id="4C80EB9AEC64FFF7FE93F8CFFE0BF89C" box="[339,441,1885,1912]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="60" unit="mm" value="0.15">0.15 mm</quantity>
long; mesonotum humped antero-dorsally (
<figureCitation id="13435AFAEC64FFF7FC6CF8CFFBB4F893" box="[940,1030,1885,1911]" captionStart="FIGURE 2" captionStartId="3.[151,250,1876,1900]" captionTargetBox="[151,1436,597,1853]" captionTargetId="figure-188@3.[151,1436,597,1853]" captionTargetPageId="3" captionText="FIGURE 2. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Head. B, Mesosoma and head. C, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead). D, Legs. E, Metasoma with indication of metasomal segments." figureDoi="http://doi.org/10.5281/zenodo.14595748" httpUri="https://zenodo.org/record/14595748/files/figure.png" pageId="4" pageNumber="60">Fig. 2C</figureCitation>
),
<quantity id="4C80EB9AEC64FFF7FBDCF8CFFB33F89C" box="[1052,1153,1885,1912]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="4" pageNumber="60" unit="mm" value="0.35">0.35 mm</quantity>
long and
<quantity id="4C80EB9AEC64FFF7FB35F8CFFAE8F89C" box="[1269,1370,1885,1912]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="4" pageNumber="60" unit="mm" value="0.22">0.22 mm</quantity>
high; metanotum narrow,
<quantity id="4C80EB9AEC64FFF7FEB8F813FE6EF878" box="[376,476,1921,1948]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="60" unit="mm" value="0.1">0.10 mm</quantity>
long; propodeum with extended surface, slightly protruding posteriorly.
</paragraph>
<paragraph id="8BC7467FEC64FFF6FF07F834FC67FE49" blockId="4.[151,1436,1849,2020]" lastBlockId="5.[151,1437,151,681]" lastPageId="5" lastPageNumber="61" pageId="4" pageNumber="60">
<emphasis id="B90C9A6DEC64FFF7FF07F834FE81F85B" box="[199,307,1958,1983]" italics="true" pageId="4" pageNumber="60">Forewing</emphasis>
: incomplete, at least
<quantity id="4C80EB9AEC64FFF7FDF5F837FD2FF824" box="[565,669,1957,1984]" metricMagnitude="-3" metricUnit="m" metricValue="1.55" pageId="4" pageNumber="60" unit="mm" value="1.55">1.55 mm</quantity>
long (
<figureCitation id="13435AFAEC64FFF7FD2FF837FC85F85B" box="[751,823,1957,1983]" captionStart="FIGURE 3" captionStartId="4.[151,250,1707,1731]" captionTargetBox="[151,1436,181,1656]" captionTargetId="figure-17@4.[151,1436,181,1683]" captionTargetPageId="4" captionText="FIGURE 3. Forewing of Manlaya minima sp. nov., holotype NIGP206130. A, Photograph of the forewing. B, Name of veins. C, Name of cells. B and C at the same scale." figureDoi="http://doi.org/10.5281/zenodo.14595750" httpUri="https://zenodo.org/record/14595750/files/figure.png" pageId="4" pageNumber="60">Fig. 3</figureCitation>
); costal cell narrower than pterostigma; pterostigma broad, 1.8× longer than wide; crossvein r-rs arising medially from pterostigma, slightly inclivous, half longer than pterostigma width; vein 1Rs originating proximad pterostigma base, separated from pterostigma base by length 2× pterostigma basal width; veins 1Rs and 1M aligned, curved; 1Rs 1.3× longer than 1M; 1Rs+M 2.3× longer than 2Rs+M; cell 1mcu strongly transverse, parallel-sided, 3× longer than wide; 2Rs+M about as long as crossvein 1mcu; crossveins 2r-m and 3r-m completely absent (no stubs); distal abscissa of Rs strongly bent anteriorly at point where distal r-m should be present, virtually extending straight towards nearly wing apex; cell 2+3rm narrower than cell 3r at level of pterostigma apex; distal abscissa of M strongly bent posteriorly at point where distal r-m should be present, straight; vein M+Cu mostly spectral; crossvein 1cu-a slightly postfurcal; cell 2mcu elbowed; crossvein 2m-cu distinctly distad r-rs, as long as r-rs level; vein A sigmoidal basally.
</paragraph>
<paragraph id="8BC7467FEC65FFF6FF07FE25FE7BFE35" blockId="5.[151,1437,151,681]" box="[199,457,439,465]" pageId="5" pageNumber="61">
<emphasis id="B90C9A6DEC65FFF6FF07FE25FE8EFE34" box="[199,316,439,465]" italics="true" pageId="5" pageNumber="61">Hind wing</emphasis>
: not visible.
</paragraph>
<paragraph id="8BC7467FEC65FFF6FF07FE49FE3CFDD9" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="61">
<emphasis id="B90C9A6DEC65FFF6FF07FE49FF4EFE10" box="[199,252,475,500]" italics="true" pageId="5" pageNumber="61">Legs</emphasis>
: partly preserved, thin (
<figureCitation id="13435AFAEC65FFF6FDD4FE49FDC3FE11" box="[532,625,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1876,1900]" captionTargetBox="[151,1436,597,1853]" captionTargetId="figure-188@3.[151,1436,597,1853]" captionTargetPageId="3" captionText="FIGURE 2. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Head. B, Mesosoma and head. C, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead). D, Legs. E, Metasoma with indication of metasomal segments." figureDoi="http://doi.org/10.5281/zenodo.14595748" httpUri="https://zenodo.org/record/14595748/files/figure.png" pageId="5" pageNumber="61">Fig. 2D</figureCitation>
); hind femur nearly as long as hind tibia; hind femur about 5× longer than wide; hind tibia about 6× longer than wide; tarsi pentamerous; hind basitarsus much longer than remainder tarsomeres combined.
</paragraph>
<paragraph id="8BC7467FEC65FFF6FF07FDD5FD3FFD4D" blockId="5.[151,1437,151,681]" pageId="5" pageNumber="61">
<emphasis id="B90C9A6DEC65FFF6FF07FDD5FE8EFD84" box="[199,316,583,608]" italics="true" pageId="5" pageNumber="61">Metasoma</emphasis>
: ovoid from lateral view (
<figureCitation id="13435AFAEC65FFF6FDAAFDD5FD70FD85" box="[618,706,583,609]" captionStart="FIGURE 2" captionStartId="3.[151,250,1876,1900]" captionTargetBox="[151,1436,597,1853]" captionTargetId="figure-188@3.[151,1436,597,1853]" captionTargetPageId="3" captionText="FIGURE 2. Manlaya minima sp. nov., holotype NIGP206130, female, from Lower Cretaceous Shouchang Formation (China). A, Head. B, Mesosoma and head. C, Dorsal part of the mesosoma with indication of the mesonotum humped antero-dorsally (black arrowhead) and the propodeum with extended surface and slightly protruding posteriorly (white arrowhead). D, Legs. E, Metasoma with indication of metasomal segments." figureDoi="http://doi.org/10.5281/zenodo.14595748" httpUri="https://zenodo.org/record/14595748/files/figure.png" pageId="5" pageNumber="61">Fig. 2E</figureCitation>
),
<quantity id="4C80EB9AEC65FFF6FD18FDD5FC8CFD85" box="[728,830,583,609]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="5" pageNumber="61" unit="mm" value="1.03">1.03 mm</quantity>
long and
<quantity id="4C80EB9AEC65FFF6FC71FDD5FBA5FD85" box="[945,1047,583,609]" metricMagnitude="-4" metricUnit="m" metricValue="7.6" pageId="5" pageNumber="61" unit="mm" value="0.76">0.76 mm</quantity>
wide; metasoma 1.5× longer than mesosoma; seven metasomal segments distinguished, first segment the longest; ovipositor incomplete, thin, slightly bent upwards, length as preserved:
<quantity id="4C80EB9AEC65FFF6FDE2FD1DFD3BFD4D" box="[546,649,655,681]" metricMagnitude="-4" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="61" unit="mm" value="0.64">0.64 mm</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>