287 lines
38 KiB
XML
287 lines
38 KiB
XML
<document id="7C272541E29785AA33CF28B6F3C50905" ID-CLB-Dataset="110541" ID-DOI="10.11646/zootaxa.5168.4.8" ID-GBIF-Dataset="5d8c7b19-0478-46eb-b191-7a09000ba2b3" ID-ISSN="1175-5326" ID-Zenodo-Dep="6899833" ID-ZooBank="3BE34F1B-3F95-4155-99BD-C38B55F72A63" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658733331983" checkinUser="plazi" docAuthor="Munari, Lorenzo" docDate="2022" docId="03CDAC560A0BFFA0A8A6D00F87EEF8F4" docLanguage="en" docName="zootaxa.5168.4.8.pdf" docOrigin="Zootaxa 5168 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Procanace amnoni Munari 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="482" masterDocId="FFF4D42E0A0AFFA3A831D414845BFFDD" masterDocTitle="A new species of Procanace Hendel from the Philippines (Diptera: Canacidae: Canacinae)" masterLastPageNumber="484" masterPageNumber="479" pageNumber="480" updateTime="1699361772291" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="0E037502EEBD1459C8EB0038B889D5F1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="5811F586DE4AE8A1E76BDA41137B8148">
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<mods:title id="4D77E2547E615537AC901AD92E3717F3">A new species of Procanace Hendel from the Philippines (Diptera: Canacidae: Canacinae)</mods:title>
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<mods:name id="DBEE120F254E0E935E20F40B85C4544E" type="personal">
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<mods:roleTerm id="1A6BD87684A9C23E467EF1DCF0DB3D2B">Author</mods:roleTerm>
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<mods:namePart id="ED2EE9A27C392E85348BCBA19426B18B">Munari, Lorenzo</mods:namePart>
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<mods:title id="029FB70DAFD2B95E50B56F929DA51342">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A6B4EB1223CA830559BECE64778494AF">
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<mods:date id="F41978B8AD5014D125948D5527F05B70">2022</mods:date>
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<mods:number id="657469169CD4C27AA347F4698201E0AB">2022-07-25</mods:number>
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<mods:number id="9DB91A0BE60ACF993DDDFB9CBAD62B6A">5168</mods:number>
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<mods:detail id="3AC18C88B54D534C99CBAE739DF305AD" type="issue">
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<mods:number id="63648CDC0975416D21AAD70017E6200D">4</mods:number>
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<mods:start id="B36C1EFE035F4A2618C4D0D4EE668DAE">479</mods:start>
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<mods:classification id="8A6F883A878BE14E00DED5099D9563B5">journal article</mods:classification>
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<mods:identifier id="9100E723CEAC84A1B28A1C51F30DA313" type="DOI">10.11646/zootaxa.5168.4.8</mods:identifier>
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<mods:identifier id="1CE55C25C7DD87161B9966CCC8EF3B53" type="GBIF-Dataset">5d8c7b19-0478-46eb-b191-7a09000ba2b3</mods:identifier>
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<mods:identifier id="BC2018E0EE9D26617ADB03CA8215603A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03CDAC560A0BFFA0A8A6D00F87EEF8F4" ID-DOI="http://doi.org/10.5281/zenodo.6899835" ID-GBIF-Taxon="198141165" ID-Zenodo-Dep="6899835" LSID="urn:lsid:plazi:treatment:03CDAC560A0BFFA0A8A6D00F87EEF8F4" httpUri="http://treatment.plazi.org/id/03CDAC560A0BFFA0A8A6D00F87EEF8F4" lastPageId="3" lastPageNumber="482" pageId="1" pageNumber="480">
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<subSubSection id="C37E4ECB0A0BFFA2A8A6D00F85BAFBE8" box="[151,481,1051,1077]" pageId="1" pageNumber="480" type="nomenclature">
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<paragraph id="8BDB1D400A0BFFA2A8A6D00F85BAFBE8" blockId="1.[151,481,1051,1113]" box="[151,481,1051,1077]" pageId="1" pageNumber="480">
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<heading id="D093AA2C0A0BFFA2A8A6D00F85BAFBE8" bold="true" box="[151,481,1051,1077]" fontSize="11" level="1" pageId="1" pageNumber="480" reason="1">
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<emphasis id="B910C1520A0BFFA2A8A6D00F85BAFBE8" bold="true" box="[151,481,1051,1077]" pageId="1" pageNumber="480">
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<taxonomicName id="4C6466C30A0BFFA2A8A6D00F8527FBE8" ID-CoL="BDCPR" authority="Munari, 2022" authorityName="Munari" authorityYear="2022" box="[151,380,1051,1077]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="species" species="amnoni" status="sp. nov.">
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<emphasis id="B910C1520A0BFFA2A8A6D00F8527FBE8" bold="true" box="[151,380,1051,1077]" italics="true" pageId="1" pageNumber="480">Procanace amnoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2237C290A0BFFA2A9B2D00F85BAFBE8" box="[387,481,1051,1077]" pageId="1" pageNumber="480" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E4ECB0A0BFFA2A8A6D02B84AFFB84" box="[151,244,1087,1113]" pageId="1" pageNumber="480" type="description">
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<paragraph id="8BDB1D400A0BFFA2A8A6D02B84AFFB84" blockId="1.[151,481,1051,1113]" box="[151,244,1087,1113]" pageId="1" pageNumber="480">
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<figureCitation id="135F01C50A0BFFA2A8A6D02B84AFFB84" box="[151,244,1087,1113]" captionStart-0="FIGURES 1‒2" captionStart-1="FIGURES 3‒6" captionStartId-0="2.[151,264,1631,1655]" captionStartId-1="4.[151,264,1882,1906]" captionTargetBox-0="[159,1428,715,1606]" captionTargetBox-1="[179,1423,199,1845]" captionTargetId-0="figure-289@2.[151,1436,705,1608]" captionTargetId-1="figure-38@4.[151,1436,181,1858]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1‒2. Procanace amnoni sp. nov., 1. Head lateral view of a paratype ♀ 2. Left foretibia posterior view of the holotype (♂). Scale bars: = 0.3 mm." captionText-1="FIGURES 3‒6. Procanace amnoni sp. nov., 3. ♂ terminalia caudal view of the holotype; 4. Ditto, left lateral view; 5. Postabdomen ventral view of a paratype ♀ (with spermatheca slightly magnified); 6. Postabdomen lateral view of a paratype ♀. Scale bars: = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6899837" figureDoi-1="http://doi.org/10.5281/zenodo.6899841" httpUri-0="https://zenodo.org/record/6899837/files/figure.png" httpUri-1="https://zenodo.org/record/6899841/files/figure.png" pageId="1" pageNumber="480">Figs 1-6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E4ECB0A0BFFA2A8A6D09380A6FB18" pageId="1" pageNumber="480" type="etymology">
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<paragraph id="8BDB1D400A0BFFA2A8A6D09380A6FB18" blockId="1.[151,1437,1159,2013]" pageId="1" pageNumber="480">
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<emphasis id="B910C1520A0BFFA2A8A6D0938543FB7C" bold="true" box="[151,280,1159,1185]" pageId="1" pageNumber="480">Etymology</emphasis>
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. The species epithet,
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<taxonomicName id="4C6466C30A0BFFA2AA20D093863EFB7D" authorityName="Munari" authorityYear="2022" box="[529,613,1159,1184]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="480" phylum="Arthropoda" rank="species" species="amnoni">
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<emphasis id="B910C1520A0BFFA2AA20D093863EFB7D" box="[529,613,1159,1184]" italics="true" pageId="1" pageNumber="480">amnoni</emphasis>
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</taxonomicName>
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, is a genitive first name to recognize the immense contribution made to Dipterology by the late Dr. Amnon Freidberg, who also collected the
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<typeStatus id="54DFA3E20A0BFFA2AB91D0BF878BFB18" box="[928,976,1195,1221]" pageId="1" pageNumber="480">type</typeStatus>
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specimens described here.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37E4ECB0A0BFFA0A8F6D0DA808EFDE0" lastPageId="3" lastPageNumber="482" pageId="1" pageNumber="480" type="description">
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<paragraph id="8BDB1D400A0BFFA2A8F6D0DA8502FB34" blockId="1.[151,1437,1159,2013]" box="[199,345,1230,1257]" pageId="1" pageNumber="480">
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<emphasis id="B910C1520A0BFFA2A8F6D0DA850AFB35" bold="true" box="[199,337,1230,1256]" pageId="1" pageNumber="480">Description</emphasis>
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.
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</paragraph>
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<paragraph id="8BDB1D400A0BFFA2A8F6D0E7810CFAEC" blockId="1.[151,1437,1159,2013]" pageId="1" pageNumber="480">
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<emphasis id="B910C1520A0BFFA2A8F6D0E7852DFAD0" bold="true" box="[199,374,1267,1293]" pageId="1" pageNumber="480">Measurements</emphasis>
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: body length (measurements carried out on
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<specimenCount id="9D62D6C90A0BFFA2AB6FD0E7804FFAD0" box="[862,1044,1267,1293]" pageId="1" pageNumber="480" type="female" typeStatus="paratypes">two paratypes ♀</specimenCount>
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, since the
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<typeStatus id="54DFA3E20A0BFFA2ACB8D0E780B0FAD0" box="[1161,1259,1267,1293]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
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<specimenCount id="9D62D6C90A0BFFA2ACC0D0E7815DFAD0" box="[1265,1286,1267,1293]" pageId="1" pageNumber="480" type="male">♂</specimenCount>
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and the other
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<typeStatus id="54DFA3E20A0BFFA2A8A6D10384ACFAEC" box="[151,247,1303,1329]" pageId="1" pageNumber="480" type="paratype">paratype</typeStatus>
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<specimenCount id="9D62D6C90A0BFFA2A8CED103854FFAEC" box="[255,276,1303,1329]" pageId="1" pageNumber="480" type="female">♀</specimenCount>
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had the abdomen previously detached for dissection)
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<quantity id="4C9CB0A50A0BFFA2AB40D102804DFAEC" box="[881,1046,1302,1329]" metricMagnitude="-3" metricUnit="m" metricValue="2.735" metricValueMax="2.89" metricValueMin="2.58" pageId="1" pageNumber="480" unit="mm" value="2.735" valueMax="2.89" valueMin="2.58">2.58–2.89 mm</quantity>
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, wing length
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<quantity id="4C9CB0A50A0BFFA2AC9CD1038108FAEC" box="[1197,1363,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="2.48" metricValueMax="2.72" metricValueMin="2.24" pageId="1" pageNumber="480" unit="mm" value="2.48" valueMax="2.72" valueMin="2.24">2.24–2.72 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BDB1D400A0BFFA2A8F6D12F85A4FAA4" blockId="1.[151,1437,1159,2013]" pageId="1" pageNumber="480">
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<emphasis id="B910C1520A0BFFA2A8F6D12F857BFA88" box="[199,288,1339,1365]" italics="true" pageId="1" pageNumber="480">Habitus</emphasis>
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. A small fly with glossy dark brown thorax, dull brown abdomen, and yellow legs; wing moderately infuscated; setal vestiture black.
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</paragraph>
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<paragraph id="8BDB1D400A0BFFA2A8F6D19780BBF864" blockId="1.[151,1437,1159,2013]" pageId="1" pageNumber="480">
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<emphasis id="B910C1520A0BFFA2A8F6D197855FFA40" box="[199,260,1411,1437]" italics="true" pageId="1" pageNumber="480">Head</emphasis>
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(
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<figureCitation id="135F01C50A0BFFA2A925D197850CFA40" box="[276,343,1411,1437]" captionStart="FIGURES 1‒2" captionStartId="2.[151,264,1631,1655]" captionTargetBox="[159,1428,715,1606]" captionTargetId="figure-289@2.[151,1436,705,1608]" captionTargetPageId="2" captionText="FIGURES 1‒2. Procanace amnoni sp. nov., 1. Head lateral view of a paratype ♀ 2. Left foretibia posterior view of the holotype (♂). Scale bars: = 0.3 mm." figureDoi="http://doi.org/10.5281/zenodo.6899837" httpUri="https://zenodo.org/record/6899837/files/figure.png" pageId="1" pageNumber="480">Fig. 1</figureCitation>
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). Subquadrangular, sharp-cornered in lateral view; frons progressively inclined toward the front half; head about as high as or slightly higher than long; frons (including mesofrons of authors) and ocellar area pale brown, sometimes with olivaceous shades anteriorly; occipital area dark brown to black; fronto-orbital plate pale olivaceous to concolorous with frons; face pale cinereous yellow, with very weak carina medially; gena yellow, with postero-dorsal surface narrowly triangular, velvety black; postgena microscopically raspy, dark brown to black; parafacial yellowish, narrower than half the height of postpedicel (basal flagellomere); postocellar setae lacking, or at least not visible; medial and lateral setae subequal in length; postocular setae inconspicuous; gena bearing thin vibrissa and two strong anaclinate setae arising just above peristomal margin (fig. 1); ocellar setae long; ocellar tubercle with 2 divergent thin setulae just behind posterior ocelli; 3 large lateroclinate fronto-orbital setae intermixed with small setulae; frons bare, without interfrontal setae, with sparse, minute setulae only on anterior portion; antenna porrect (both antennae lacking in
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<typeStatus id="54DFA3E20A0BFFA2AAE9D2FF8761F8D8" box="[728,826,1771,1797]" pageId="1" pageNumber="480" type="holotype">holotype</typeStatus>
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(
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<specimenCount id="9D62D6C90A0BFFA2AB7AD2FF873AF8D8" box="[843,865,1771,1797]" pageId="1" pageNumber="480" type="male">♂</specimenCount>
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) and in a
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<typeStatus id="54DFA3E20A0BFFA2ABE7D2FF806DF8D8" box="[982,1078,1771,1797]" pageId="1" pageNumber="480" type="paratype">paratype</typeStatus>
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(
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<specimenCount id="9D62D6C90A0BFFA2AC78D2FF8001F8D8" box="[1097,1114,1771,1797]" pageId="1" pageNumber="480" type="female">♀</specimenCount>
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)), with postpedicel (= basal flagellomere) densely pubescent, dark brown to blackish; arista dark grey, with dorsal and ventral rows of short branches (rays), tip sector bare; eye bare, without interfacetal ommatrichia, its anterior corner with distinctly larger ommatidia; eye large, oblique, its vertical diameter 2.45–2.85 times as long as genal height; clypeus moderately large, dark brown, invested with golden microtomentum, especially obvious on ventral margin; palpus yellow, long, rod-like; ventral and buccal cavities wide; mouthparts strongly sclerotised ventrally (prementum).
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</paragraph>
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<paragraph id="8BDB1D400A0BFFA1A8F6D3D780ABFE54" blockId="1.[151,1437,1159,2013]" lastBlockId="2.[151,1437,151,681]" lastPageId="2" lastPageNumber="481" pageId="1" pageNumber="480">
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<emphasis id="B910C1520A0BFFA2A8F6D3D78543F800" box="[199,280,1987,2013]" italics="true" pageId="1" pageNumber="480">Thorax</emphasis>
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. Dark brown, smooth, rather glossy; mesonotum with major setae only, lacking obvious setulae, invested with thin grey microtomentum; 1 + 3 large dorsocentral setae; acrostichal setulae, including prescutellars, absent; 1 postpronotal seta along with a few weak small setulae on median surface of postpronotal lobe; 1 anterior, 1 posterior notopleural setae; 1 supra-alar seta; 1 lateral postalar seta (medial postalar seta lacking); postalar callus poorly developed; scutellum concolorous with scutum, bearing 2 pairs of setae, one apical pair, one latero-basal pair; both proepisternal and proepimeral setae not visible, apparently lacking; pleura dark brown, subshiny, densely micropubescent (especially on anepisternum); anepisternum with strong seta backwardly orientated on posterior margin; katepisternum lacking setae; anepimeron and meron matt, mostly entirely microtomentose.
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</paragraph>
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<paragraph id="8BDB1D400A08FFA1A8F6D58781DAFD74" blockId="2.[151,1437,151,681]" pageId="2" pageNumber="481">
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<emphasis id="B910C1520A08FFA1A8F6D58784A7FE71" box="[199,252,403,428]" italics="true" pageId="2" pageNumber="481">Legs</emphasis>
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. Entirely yellow, excepting dark grey to brown coxae and dark brown to blackish distal 2 tarsomeres; forecoxa dark grey, bearing long yellow villosity on antero-distal surface; forefemur with row of 3–5 long dorsal setae on distal half, bearing dense, long, fine villosity ventrally; foretibia (fig. 2) with posterior and postero-dorsal rows of fine setae, very characteristic comb of 8–14 (
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<quantity id="4C9CB0A50A08FFA1AAC1D5EB8771FDC4" box="[752,810,511,537]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="2" pageNumber="481" unit="in" value="14.0">14 in</quantity>
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<typeStatus id="54DFA3E20A08FFA1AB00D5EB87CEFDC4" box="[817,917,511,537]" pageId="2" pageNumber="481" type="holotype">holotype</typeStatus>
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) antero-ventral strong spine-like setae present in both sexes, distinctly stronger in
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<typeStatus id="54DFA3E20A08FFA1AA12D63786DEFDE0" box="[547,645,547,573]" pageId="2" pageNumber="481" type="holotype">holotype</typeStatus>
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||
(
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<specimenCount id="9D62D6C90A08FFA1AAA4D63786F7FDE0" box="[661,684,547,573]" pageId="2" pageNumber="481" type="male">♂</specimenCount>
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); foretarsus with rows of setulae; mid femur with numerous anteroventral and anterior fine setae, 5 strong postero-ventral distal setae present in male, mid tibia with long row of 7–10 fine postero-dorsal setae; rest of tibia evenly setulose; setulation of mid tarsus similar to that of foretarsus; hind leg evenly setulose, with longer antero-dorsal, fine setae on tibia; setulation of hind tarsus like that of the other legs.
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</paragraph>
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<caption id="DF1B4DC80A08FFA1A8A6D24B865EF947" ID-DOI="http://doi.org/10.5281/zenodo.6899837" ID-Zenodo-Dep="6899837" httpUri="https://zenodo.org/record/6899837/files/figure.png" pageId="2" pageNumber="481" startId="2.[151,264,1631,1655]" targetBox="[159,1428,715,1606]" targetPageId="2">
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<paragraph id="8BDB1D400A08FFA1A8A6D24B865EF947" blockId="2.[151,1436,1631,1691]" pageId="2" pageNumber="481">
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<emphasis id="B910C1520A08FFA1A8A6D24B8565F9AA" bold="true" box="[151,318,1631,1655]" pageId="2" pageNumber="481">FIGURES 1‒2.</emphasis>
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<taxonomicName id="4C6466C30A08FFA1A979D2748650F9AA" authority="Munari, 2022" authorityName="Munari" authorityYear="2022" box="[328,523,1632,1655]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="481" phylum="Arthropoda" rank="species" species="amnoni" status="sp. nov.">
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<emphasis id="B910C1520A08FFA1A979D2748650F9AA" box="[328,523,1632,1655]" italics="true" pageId="2" pageNumber="481">Procanace amnoni</emphasis>
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</taxonomicName>
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<emphasis id="B910C1520A08FFA1AA24D24B8632F9AA" bold="true" box="[533,617,1631,1655]" pageId="2" pageNumber="481">
|
||
<taxonomicNameLabel id="A2237C290A08FFA1AA24D24B8632F9AA" box="[533,617,1631,1655]" pageId="2" pageNumber="481" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="B910C1520A08FFA1AA4BD24B86D6F9AA" bold="true" box="[634,653,1631,1655]" pageId="2" pageNumber="481">1.</emphasis>
|
||
Head lateral view of a paratype ♀
|
||
<emphasis id="B910C1520A08FFA1AC3FD24B807AF9AA" bold="true" box="[1038,1057,1631,1655]" pageId="2" pageNumber="481">2.</emphasis>
|
||
Left foretibia posterior view of the holotype (♂). Scale bars: = 0.3 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDB1D400A08FFA1A8F6D2DE8780F8DA" blockId="2.[151,1437,1737,2016]" pageId="2" pageNumber="481">
|
||
<emphasis id="B910C1520A08FFA1A8F6D2DE855AF93E" box="[199,257,1738,1763]" italics="true" pageId="2" pageNumber="481">Wing</emphasis>
|
||
. Membrane infuscated; venation dark brown to blackish; Costal vein ratio = 0.17–0.20;
|
||
<emphasis id="B910C1520A08FFA1ACC6D2DE8155F93E" box="[1271,1294,1738,1763]" italics="true" pageId="2" pageNumber="481">M</emphasis>
|
||
vein ratio = 1.07–1.58; haltere yellow, with distinctly triangular knob, tapered apically.
|
||
</paragraph>
|
||
<paragraph id="8BDB1D400A08FFA1A8F6D30580C9F892" blockId="2.[151,1437,1737,2016]" pageId="2" pageNumber="481">
|
||
<emphasis id="B910C1520A08FFA1A8F6D305856AF8F6" box="[199,305,1809,1835]" italics="true" pageId="2" pageNumber="481">Abdomen</emphasis>
|
||
. Dark brown, dull, sparsely setulose; syntergite 1+2 large, about as long as total length of tergites 3–5 as a whole; pregenital tergite of female (tergite 8) bearing very long marginal setae (fig. 6).
|
||
</paragraph>
|
||
<paragraph id="8BDB1D400A08FFA0A8F6D34D8536FEC0" blockId="2.[151,1437,1737,2016]" lastBlockId="3.[151,1437,151,1833]" lastPageId="3" lastPageNumber="482" pageId="2" pageNumber="481">
|
||
<emphasis id="B910C1520A08FFA1A8F6D34D85DBF8AE" box="[199,384,1881,1907]" italics="true" pageId="2" pageNumber="481">Male terminalia</emphasis>
|
||
. (
|
||
<figureCitation id="135F01C50A08FFA1A9A8D34D8658F8AE" box="[409,515,1881,1907]" captionStart="FIGURES 3‒6" captionStartId="4.[151,264,1882,1906]" captionTargetBox="[179,1423,199,1845]" captionTargetId="figure-38@4.[151,1436,181,1858]" captionTargetPageId="4" captionText="FIGURES 3‒6. Procanace amnoni sp. nov., 3. ♂ terminalia caudal view of the holotype; 4. Ditto, left lateral view; 5. Postabdomen ventral view of a paratype ♀ (with spermatheca slightly magnified); 6. Postabdomen lateral view of a paratype ♀. Scale bars: = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6899841" httpUri="https://zenodo.org/record/6899841/files/figure.png" pageId="2" pageNumber="481">Figs 3‒4</figureCitation>
|
||
). Epandrium large, bearing numerous, moderately long to short setae; surstylus rather complex, made up of two large parts, lateral one roughly bilobed apically, with antero-ventral half (finely punctuated in fig. 3) extended almost the full length of posterior half, densely setulose apically, posterior half elongate, lobe-shaped, bearing thin setulae on mesial margin, medial part of surstylus elongate, rounded apically, bearing numerous long setae on mesial margin; epandrium, just below anal membrane, with very strong, black, mesial spine-like setae conspicuously tapered apically, forming a very thin apex (4 of these setae present on left margin, 2 others on right margin); cercus inconspicuous; surstylus lateral view very peculiar, strongly composite as illustrated in fig. 4.
|
||
</paragraph>
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D53386D1FE54" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="482">
|
||
The genitalia of the sole available male (
|
||
<typeStatus id="54DFA3E20A09FFA0AAA7D53386A0FE9C" box="[662,763,295,321]" pageId="3" pageNumber="482" type="holotype">holotype</typeStatus>
|
||
) show a deep laceration of the left epandrial margin due to an accident that occurred during the dissection procedure. The line drawings presented in figs 3‒4 show the genital structure as it appears under the microscope.
|
||
</paragraph>
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D587808EFDE0" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="482">
|
||
<emphasis id="B910C1520A09FFA0A8F6D58785C7FE70" box="[199,412,403,429]" italics="true" pageId="3" pageNumber="482">Female terminalia</emphasis>
|
||
. (
|
||
<figureCitation id="135F01C50A09FFA0A987D5878679FE70" box="[438,546,402,429]" captionStart="FIGURES 3‒6" captionStartId="4.[151,264,1882,1906]" captionTargetBox="[179,1423,199,1845]" captionTargetId="figure-38@4.[151,1436,181,1858]" captionTargetPageId="4" captionText="FIGURES 3‒6. Procanace amnoni sp. nov., 3. ♂ terminalia caudal view of the holotype; 4. Ditto, left lateral view; 5. Postabdomen ventral view of a paratype ♀ (with spermatheca slightly magnified); 6. Postabdomen lateral view of a paratype ♀. Scale bars: = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.6899841" httpUri="https://zenodo.org/record/6899841/files/figure.png" pageId="3" pageNumber="482">Figs 5‒6</figureCitation>
|
||
). Pregenital tergite (tergite 8) with long, strong setae backwards orientated; cercus (fig.6) long, narrow, bearing strong, apical spine curved upward and a distinctly smaller dorsal spine arising anteriorly at some distance from apical spine; sternite 8 undivided, without clusters of curved to hooked spine-like setae (fig. 5), but with few extremely minute spiculae hidden behind posterior margin (not visible in fig. 5); atrial sclerotisation broadly oval in shape; 2 discoid-shaped spermathecae, depressed on upper surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E4ECB0A09FFA0A8F6D6528779FAD0" pageId="3" pageNumber="482" type="diagnosis">
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D652812AFBE8" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="482">
|
||
<emphasis id="B910C1520A09FFA0A8F6D6528591FDBD" bold="true" box="[199,458,582,608]" pageId="3" pageNumber="482">Differential diagnosis</emphasis>
|
||
. This species is distinguished from congeners mainly by the following combination of characters: dark brown fly, with thorax smooth, mesonotum and pleura shiny to subshiny, without obvious setulae; legs entirely yellow, excepting coxae (dark grey) and distal two tarsomeres (dark brown to blackish); body length
|
||
<quantity id="4C9CB0A50A09FFA0A8D5D6A685D2FD10" box="[228,393,690,717]" metricMagnitude="-3" metricUnit="m" metricValue="2.735" metricValueMax="2.89" metricValueMin="2.58" pageId="3" pageNumber="482" unit="mm" value="2.735" valueMax="2.89" valueMin="2.58">2.58‒2.89 mm</quantity>
|
||
; head without postocellar setae; gena yellow, with postero-dorsal surface narrowly triangular, velvety black; vertical diameter of eye 2.45–2.85 times as long as genal height; proepisternal and proepimeral setae (apparently) lacking; katepisternal seta absent; membrane of wing infuscated; venation dark brown to blackish; Costal vein ratio = 0.17–0.20;
|
||
<emphasis id="B910C1520A09FFA0AA30D70B8643FCE5" box="[513,536,799,824]" italics="true" pageId="3" pageNumber="482">M</emphasis>
|
||
vein ratio = 1.07–1.58; haltere yellow, distinctly triangular, tapered apically; surstylus of male terminalia (fig. 3) divided in two parts, lateral part roughly bilobed apically in caudal view, medial part long, relatively simple, strongly setulose on mesial margin; very strong, black spine-like setae arising from mesial margin of epandrium, just below the anal membrane; cercus of male genitalia rudimentary, barely visible; female postabdomen (fig. 5) with sternite 8 undivided, without typical clusters of curved to hooked spine-like setae, although a few extremely minute spiculae, hidden behind the posterior edge (not visible in fig. 5), can be observed at high magnification after flattening the sternite by gentle pressure. These microscopic spiculae could presumably be rudiments of the clusters of recurved bristles that are commonly observed in many species of canacine flies.
|
||
</paragraph>
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D02B8779FAD0" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="482">
|
||
The new species appears to be related to two species with disjointed distributions,
|
||
<taxonomicName id="4C6466C30A09FFA0AC73D02B81C7FB84" authority="Wirth, 1951" authorityName="Wirth" authorityYear="1951" box="[1090,1436,1086,1113]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="townesi">
|
||
<emphasis id="B910C1520A09FFA0AC73D02B814EFB85" box="[1090,1301,1087,1112]" italics="true" pageId="3" pageNumber="482">Procanace townesi</emphasis>
|
||
<bibRefCitation id="EFF560B10A09FFA0AD28D02B81C7FB84" author="Wirth, W. W." box="[1305,1436,1086,1113]" pageId="3" pageNumber="482" pagination="245 - 275" refId="ref3463" refString="Wirth, W. W. (1951) A revision of the dipterous family Canaceidae. Occasional Papers of Bernice P. Bishop Museum, 20 (14), 245 - 275." type="journal article" year="1951">Wirth, 1951</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F3735DD00A09FFA0A8AED0778506FBA0" box="[159,349,1123,1149]" name="Marshall Islands" pageId="3" pageNumber="482">Marshall Islands</collectingCountry>
|
||
) and
|
||
<taxonomicName id="4C6466C30A09FFA0A9ACD077876FFBA0" authority="Mathis, 1988" authorityName="Mathis" authorityYear="1988" box="[413,820,1123,1149]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="cogani">
|
||
<emphasis id="B910C1520A09FFA0A9ACD077865EFBA1" box="[413,517,1123,1148]" italics="true" pageId="3" pageNumber="482">P. cogani</emphasis>
|
||
<bibRefCitation id="EFF560B10A09FFA0AA3CD07786F9FBA0" author="Mathis, W. N." box="[525,674,1123,1149]" pageId="3" pageNumber="482" pagination="22 - 29" refId="ref3255" refString="Mathis, W. N. (1988) Beach flies of the Republic of Seychelles (Diptera: Canacidae). Bulletin of the Washington Biological Society, 8, 22 - 29." type="journal article" year="1988">Mathis, 1988</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F3735DD00A09FFA0AA80D0778777FBA0" box="[689,812,1123,1149]" name="Seychelles" pageId="3" pageNumber="482">Seychelles</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
in that it shares a similar morphological pattern of the bipartite surstylus (with semidetached lateral and mesial parts), as well as the same strong spine-like setae on mesial margin of epandrium, just below the anal membrane. The latter peculiar feature is in all respect a synapomorphic character state with respect to the groundplan of the genus. Obviously, the shape of the surstylus of the new species is quite dissimilar to that of both
|
||
<emphasis id="B910C1520A09FFA0AA3BD0E78740FAD1" box="[522,795,1267,1293]" italics="true" pageId="3" pageNumber="482">
|
||
<taxonomicName id="4C6466C30A09FFA0AA3BD0E78627FAD1" authorityName="Wirth" authorityYear="1951" box="[522,636,1267,1292]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="townesi">P. townesi</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C6466C30A09FFA0AA82D0E78740FAD1" authorityName="Mathis" authorityYear="1988" box="[691,795,1267,1292]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="cogani">P. cogani</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E4ECB0A09FFA0A8F6D103853CF93C" pageId="3" pageNumber="482" type="materials_examined">
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D103853CF93C" blockId="3.[151,1437,151,1833]" pageId="3" pageNumber="482">
|
||
<emphasis id="B910C1520A09FFA0A8F6D103852BFAED" bold="true" box="[199,368,1302,1329]" pageId="3" pageNumber="482">Type material</emphasis>
|
||
.
|
||
<materialsCitation id="3B0C171D0A09FFA0A9B0D1038672F9A8" ID-GBIF-Occurrence="3874246301" collectingDate="2006-09-09" collectingDateMax="2006-10-14" collectingDateMin="2006-09-09" collectionCode="SMNH" collectorName="A. Freidberg & L. Munari" country="Philippines" elevation="3" latitude="11.958333" location="Boracay Island" longLatPrecision="129" longitude="121.93" municipality="Panay" pageId="3" pageNumber="482" specimenCount="2" specimenCount-male="2" stateProvince="Aklan" typeStatus="holotype">
|
||
<collectingCountry id="F3735DD00A09FFA0A9B0D1038671FAEC" box="[385,554,1303,1329]" name="Philippines" pageId="3" pageNumber="482">PHILIPPINES</collectingCountry>
|
||
:
|
||
<typeStatus id="54DFA3E20A09FFA0AA06D10386C2FAEC" box="[567,665,1303,1329]" pageId="3" pageNumber="482" type="holotype">holotype</typeStatus>
|
||
<specimenCount id="9D62D6C90A09FFA0AA92D10386E3FAEC" box="[675,696,1303,1329]" pageId="3" pageNumber="482" type="male">♂</specimenCount>
|
||
″
|
||
<collectingMunicipality id="6BBF873A0A09FFA0AAFCD10C8749FAEE" box="[717,786,1304,1331]" pageId="3" pageNumber="482">Panay</collectingMunicipality>
|
||
/
|
||
<collectingRegion id="49A0D3A20A09FFA0AB1AD1038786FAEC" box="[811,989,1303,1329]" country="Philippines" name="Aklan" pageId="3" pageNumber="482">Aklan Province</collectingRegion>
|
||
/
|
||
<location id="8EBB4B9B0A09FFA0ABC6D10380F9FAEC" LSID="urn:lsid:plazi:treatment:03CDAC560A0BFFA0A8A6D00F87EEF8F4:8EBB4B9B0A09FFA0ABC6D10380F9FAEC" box="[1015,1186,1303,1329]" country="Philippines" latitude="11.958333" longLatPrecision="129" longitude="121.93" municipality="Panay" name="Boracay Island" pageId="3" pageNumber="482" stateProvince="Aklan">Boracay Island</location>
|
||
/
|
||
<collectingRegion id="49A0D3A20A09FFA0AC8CD103814AFAEC" box="[1213,1297,1303,1329]" country="Philippines" name="Laguna" pageId="3" pageNumber="482">Laguna</collectingRegion>
|
||
de Boracay [printed] //
|
||
<geoCoordinate id="EE507B870A09FFA0A911D12885C6FA88" box="[288,413,1339,1367]" degrees="11" direction="north" minutes="57.5" orientation="latitude" pageId="3" pageNumber="482" precision="92" value="11.958333">11°57.5′ N</geoCoordinate>
|
||
<geoCoordinate id="EE507B870A09FFA0A999D128866AFA88" box="[424,561,1339,1367]" degrees="121" direction="east" minutes="55.8" orientation="longitude" pageId="3" pageNumber="482" precision="92" value="121.93">121°55.8′ E</geoCoordinate>
|
||
/
|
||
<quantity id="4C9CB0A50A09FFA0AA61D12E86CBFA88" box="[592,656,1338,1365]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="3" pageNumber="482" unit="m" value="2.5" valueMax="5.0" valueMin="0.0">
|
||
<elevation id="0049FA730A09FFA0AA61D12E86CBFA88" box="[592,656,1338,1365]" metricMagnitude="0" metricUnit="m" metricValue="2.5" metricValueMax="5.0" metricValueMin="0.0" pageId="3" pageNumber="482" unit="m" value="2.5" valueMax="5.0" valueMin="0.0">0‒5m</elevation>
|
||
</quantity>
|
||
<date id="FFDA3B800A09FFA0AAAAD12F8773FA88" box="[667,808,1339,1365]" pageId="3" pageNumber="482" value="2006-10-09" valueMax="2006-10-14" valueMin="2006-10-09">
|
||
<collectingDate id="EF9EC2680A09FFA0AAAAD12F8773FA88" box="[667,808,1339,1365]" pageId="3" pageNumber="482" value="2006-10-09" valueMax="2006-10-14" valueMin="2006-10-09">9‒14.x.2006</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="269178960A09FFA0AB74D12F878CFA88" box="[837,983,1339,1365]" pageId="3" pageNumber="482">A. Freidberg</collectorName>
|
||
[printed] // 365281 [digitalised record number], Philippines: /
|
||
<collectingRegion id="49A0D3A20A09FFA0A99BD14B85A5FAA4" box="[426,510,1375,1401]" country="Philippines" name="Laguna" pageId="3" pageNumber="482">Laguna</collectingRegion>
|
||
de Boracay / 11°57.5′ N 121°55.8′ E /
|
||
<date id="FFDA3B800A09FFA0AB8DD14B802EFAA4" box="[956,1141,1375,1401]" pageId="3" pageNumber="482" value="2006-09-09" valueMax="2006-10-14" valueMin="2006-09-09">
|
||
<collectingDate id="EF9EC2680A09FFA0AB8DD14B802EFAA4" box="[956,1141,1375,1401]" pageId="3" pageNumber="482" value="2006-09-09" valueMax="2006-10-14" valueMin="2006-09-09">9.ix ‒ 14.x.2006</collectingDate>
|
||
</date>
|
||
/
|
||
<collectorName id="269178960A09FFA0ACBAD14B8141FAA4" box="[1163,1306,1375,1401]" pageId="3" pageNumber="482">A. Freidberg</collectorName>
|
||
[printed] // [plastic microtube, with stopper, including terminalia stored in glycerine] // HOLOTYPUS /
|
||
<taxonomicName id="4C6466C30A09FFA0AC86D19781D7FA41" authorityName="Munari" authorityYear="2022" box="[1207,1420,1411,1436]" class="Insecta" family="Canacidae" genus="Procanace" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="species" species="amnoni">
|
||
<emphasis id="B910C1520A09FFA0AC86D19781D7FA41" box="[1207,1420,1411,1436]" italics="true" pageId="3" pageNumber="482">Procanace amnoni</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<emphasis id="B910C1520A09FFA0A8A6D1B384ABFA1C" bold="true" box="[151,240,1447,1473]" pageId="3" pageNumber="482">sp. nov.</emphasis>
|
||
<specimenCount id="9D62D6C90A09FFA0A8C7D1B38550FA1C" box="[246,267,1447,1473]" pageId="3" pageNumber="482" type="male">♂</specimenCount>
|
||
/
|
||
<collectorName id="269178960A09FFA0A92ED1B385D4FA1C" box="[287,399,1447,1473]" pageId="3" pageNumber="482">L. Munari</collectorName>
|
||
des. 2022 [printed red label]″ (
|
||
<collectionCode id="ED7585850A09FFA0AAD9D1B38764FA1C" box="[744,831,1447,1473]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="3" pageNumber="482">SMNH</collectionCode>
|
||
). The holotype is side-pinned by micropin and doublemounted in small block of Plastazote®. In rather poor condition (setal vestiture of frons strongly depauperate, left compound eye and left postocular region largely devoured by
|
||
<taxonomicName id="4C6466C30A09FFA0AB5DD1FB8785F9D4" box="[876,990,1519,1545]" class="Insecta" family="Dermestidae" genus="Anthrenus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="482" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B910C1520A09FFA0AB5DD1FB8785F9D4" box="[876,990,1519,1545]" italics="true" pageId="3" pageNumber="482">Anthrenus</emphasis>
|
||
</taxonomicName>
|
||
, antennae lost, right side of scutellum deeply lacerated, halteres devoured, right mid leg and tarsus of right hind leg missing, left hind leg with tibia, tarsus and apical part of femur missing). Abdomen removed and terminalia dissected and stored in genitalia vial in glycerine pinned beneath specimen
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B0C171D0A09FFA0AA04D24F8539F93C" ID-GBIF-Occurrence="3874246302" collectingDate="2006-09-09" collectingDateMax="2006-10-14" collectingDateMin="2006-09-09" collectionCode="SMNH, LMC" collectorName="A. Freidberg & L. Munari" country="Philippines" elevation="3" latitude="11.958333" location="Boracay Island" longLatPrecision="129" longitude="121.93" municipality="Panay" pageId="3" pageNumber="482" specimenCount="3" specimenCount-female="3" stateProvince="Aklan" typeStatus="paratype">
|
||
<collectingCountry id="F3735DD00A09FFA0AA04D24F8685F9A8" box="[565,734,1627,1653]" name="Philippines" pageId="3" pageNumber="482">PHILIPPINES</collectingCountry>
|
||
:
|
||
<specimenCount id="9D62D6C90A09FFA0AADBD24F8724F9A8" box="[746,895,1627,1653]" pageId="3" pageNumber="482" type="female" typeStatus="paratype">3 paratype ♀</specimenCount>
|
||
, same label data as holotype, digitalised record numbers 365288, 365287, 365282 (
|
||
<collectionCode id="ED7585850A09FFA0AA13D26B862CF944" box="[546,631,1663,1689]" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="3" pageNumber="482">SMNH</collectionCode>
|
||
,
|
||
<collectionCode id="ED7585850A09FFA0AAB3D26B869FF944" box="[642,708,1663,1689]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="482" type="Herbarium">LMC</collectionCode>
|
||
). The
|
||
<typeStatus id="54DFA3E20A09FFA0AB37D26B872AF944" box="[774,881,1663,1689]" pageId="3" pageNumber="482" type="paratype">paratypes</typeStatus>
|
||
are side-pinned by micropin and double-mounted in small block of Plastazote®. One paratype with abdomen removed and stored in genitalia vial in glycerine pinned beneath specimen
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37E4ECB0A09FFA0A8F6D2FE87EEF8F4" pageId="3" pageNumber="482" type="distribution">
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D2FE85B6F8D8" blockId="3.[151,1437,151,1833]" box="[199,493,1770,1797]" pageId="3" pageNumber="482">
|
||
<emphasis id="B910C1520A09FFA0A8F6D2FE8502F8D9" bold="true" box="[199,345,1770,1796]" pageId="3" pageNumber="482">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="F3735DD00A09FFA0A956D2FF85B2F8D8" box="[359,489,1771,1797]" name="Philippines" pageId="3" pageNumber="482">Philippines</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BDB1D400A09FFA0A8F6D31A87EEF8F4" blockId="3.[151,1437,151,1833]" box="[199,949,1806,1833]" pageId="3" pageNumber="482">
|
||
<emphasis id="B910C1520A09FFA0A8F6D31A8518F8F5" bold="true" box="[199,323,1806,1832]" pageId="3" pageNumber="482">Bionomics</emphasis>
|
||
. In all probability maritime habitat; specifics unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |